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<document id="0E3EEB6F7D5C4D0790C4D469D7A54D39" ID-DOI="10.5281/zenodo.5386711" ID-ISSN="2345-7600" ID-Zenodo-Dep="5386711" ID-ZooBank="BAA493D3-8195-417B-ABD8-A36D3FD90533" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630303402995" checkinUser="valdenar" docAuthor="Grinang, Jongkar &amp; Ng, Peter K. L." docDate="2015" docId="039387F0C66AFF9DAEF3F8FF0A2FF7A4" docLanguage="en" docName="63rbz564-582.pdf" docOrigin="Raffles Bulletin of Zoology 63" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Lepidothelphusa sangon Grinang &amp; Ng 2015, new species" docType="treatment" docUuid="EEB8BD33-A7CB-4E22-A5E3-EC8AD1B542B5" docUuidSource="ZooBank" docVersion="4" lastPageNumber="578" masterDocId="FFAAFF88C666FF93AD69FFE40E06FFB1" masterDocTitle="Taxonomy of the semiterrestrial crab Lepidothelphusa cognettii (Nobili, 1903) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of five new species from Sarawak, Malaysia, Borneo" masterLastPageNumber="582" masterPageNumber="564" pageNumber="576" updateTime="1699221322658" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="F3EA5CDE5601D753DC401D6E03BE9CBA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="21198035052CD5A5C4A4B4FD011B24AB">
<mods:title id="FAA313AF25AEED3D3BB10880E4AD9BCA">Taxonomy of the semiterrestrial crab Lepidothelphusa cognettii (Nobili, 1903) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of five new species from Sarawak, Malaysia, Borneo</mods:title>
</mods:titleInfo>
<mods:name id="A0C5F5699A28ADF13C4650E6D32B62A8" type="personal">
<mods:role id="08D67784E8A2DC7213D3EBECDFACCC56">
<mods:roleTerm id="56A42CD0B385F9E9E8B169344190D584">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D05C86846496C3B64E0D78A974D2AEFE">Grinang, Jongkar</mods:namePart>
</mods:name>
<mods:name id="6D4477AF65A03C476F314811533B7A94" type="personal">
<mods:role id="0CBE774D31F82C59FD4C05D9D8199BE9">
<mods:roleTerm id="D84BA63D2303F20F940F2F700B6808B2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="308B92831969E1BDBAC109BD84911659">Ng, Peter K. L.</mods:namePart>
</mods:name>
<mods:typeOfResource id="CA740D8D0BA072470097228C10305D1F">text</mods:typeOfResource>
<mods:relatedItem id="4F3ACF6511F3ED7DF47901F747886B1F" type="host">
<mods:titleInfo id="E6A12D0BBF3F62CCE17C4329538C2BD0">
<mods:title id="90AEFC6474F129334AC42405A8AB1AF7">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="1902DC70E82053501A883B8E43CB9415">
<mods:date id="BB3F7CBE3CAB16EC59736F0AD414D1C9">2015</mods:date>
<mods:detail id="454C2B45F2D24F760CA26E5548807AB2" type="pubDate">
<mods:number id="A15FE42062A5F13289E9F7D36B51EBCB">2015-12-18</mods:number>
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<mods:detail id="370C5ADDA83D791E2B2DE4F090DECA4D" type="volume">
<mods:number id="7FD7F946B2A723CCCAFD8827E25DC988">63</mods:number>
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<mods:start id="998AE8A4A7709E0F4BB6EA544D8B73C8">564</mods:start>
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<mods:classification id="9DDEB0142628AFE295D97A07C56E5067">journal article</mods:classification>
<mods:identifier id="BA7B133F1FA8D692E9F7B7BA54F41289" type="DOI">10.5281/zenodo.5386711</mods:identifier>
<mods:identifier id="A6A80C812D7C625BA543D362C5A6E8FE" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="E519A6767EC638767751B0EAC909B061" type="Zenodo-Dep">5386711</mods:identifier>
<mods:identifier id="DAB9729ECD747D0BD926B9592BAE0483" type="ZooBank">BAA493D3-8195-417B-ABD8-A36D3FD90533</mods:identifier>
</mods:mods>
<treatment id="039387F0C66AFF9DAEF3F8FF0A2FF7A4" LSID="urn:lsid:zoobank.org:act:EEB8BD33-A7CB-4E22-A5E3-EC8AD1B542B5" httpUri="http://treatment.plazi.org/id/039387F0C66AFF9DAEF3F8FF0A2FF7A4" lastPageId="14" lastPageNumber="578" pageId="12" pageNumber="576">
<subSubSection id="C320656DC66AFF9FAEF3F8FF0B39F882" box="[922,1343,1819,1843]" pageId="12" pageNumber="576" type="nomenclature">
<paragraph id="8B8536E6C66AFF9FAEF3F8FF0B39F882" blockId="12.[922,1343,1819,1875]" box="[922,1343,1819,1843]" pageId="12" pageNumber="576">
<heading id="D0CD818AC66AFF9FAEF3F8FF0B39F882" bold="true" box="[922,1343,1819,1843]" centered="true" fontSize="10" level="2" pageId="12" pageNumber="576" reason="2">
<emphasis id="B94EEAF4C66AFF9FAEF3F8FF0B39F882" bold="true" box="[922,1343,1819,1843]" pageId="12" pageNumber="576">
<taxonomicName id="4C3A4D65C66AFF9FAEF3F8FF0AACF882" ID-CoL="3TDTL" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[922,1194,1819,1843]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="576" phylum="Arthropoda" rank="species" species="sangon" status="sp. nov.">
<emphasis id="B94EEAF4C66AFF9FAEF3F8FF0AACF882" bold="true" box="[922,1194,1819,1843]" italics="true" pageId="12" pageNumber="576">Lepidothelphusa sangon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC66AFF9FA9D0F8FF0B39F882" box="[1209,1343,1819,1843]" pageId="12" pageNumber="576" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320656DC66AFF9FAEA0F8DF0B16F8E2" box="[969,1296,1851,1875]" pageId="12" pageNumber="576" type="description">
<paragraph id="8B8536E6C66AFF9FAEA0F8DF0B16F8E2" blockId="12.[922,1343,1819,1875]" box="[969,1296,1851,1875]" pageId="12" pageNumber="576">
(
<figureCitation id="13012A63C66AFF9FAEB8F8DF0A53F8E2" box="[977,1109,1851,1875]" captionStart="Fig" captionStartId="13.[136,171,2006,2028]" captionTargetBox="[176,1431,189,1991]" captionTargetId="figure-90@13.[176,1431,189,1991]" captionTargetPageId="13" captionText="Fig. 11. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386749" httpUri="https://zenodo.org/record/5386749/files/figure.png" pageId="12" pageNumber="576">Figs. 11A–I</figureCitation>
,
<figureCitation id="13012A63C66AFF9FA90BF8DF0AB6F8E2" box="[1122,1200,1851,1875]" captionStart="Fig" captionStartId="14.[113,148,1078,1100]" captionTargetBox="[116,1452,188,1063]" captionTargetId="figure-12@14.[92,1472,172,1070]" captionTargetPageId="14" captionText="Fig. 12. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of left G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386751" httpUri="https://zenodo.org/record/5386751/files/figure.png" pageId="12" pageNumber="576">12A–E</figureCitation>
,
<figureCitation id="13012A63C66AFF9FA9D4F8DF0B0EF8E2" box="[1213,1288,1851,1875]" captionStart="Fig" captionStartId="16.[113,148,1308,1330]" captionTargetBox="[113,1452,189,1293]" captionTargetId="figure-491@16.[113,1452,188,1294]" captionTargetPageId="16" captionText="Fig. 14. Live colours of three Lepidothelphusa species from Kuching Division, southwestern Sarawak. A, B, Lepidothelphusa loi, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak; C, D, Lepidothelphusa padawan, new species, paratype male (15.8 × 13.2 mm) (SBC.C.00440–4), Bung Brungu, Padawan, Sarawak; E, F, Lepidothelphusa sangon, new species, paratype male (8.0 × 7.0 mm) (SBC.C.00476), Nawang Waterfall, Bung Bratak, Bau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5386759" httpUri="https://zenodo.org/record/5386759/files/figure.png" pageId="12" pageNumber="576">14E, F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C320656DC66AFF9DAE44F89F0EF8FA84" lastPageId="14" lastPageNumber="578" pageId="12" pageNumber="576" type="materials_examined">
<paragraph id="8B8536E6C66AFF9DAE44F89F0EF8FA84" blockId="12.[813,1452,1915,2099]" lastBlockId="14.[113,753,1213,1334]" lastPageId="14" lastPageNumber="578" pageId="12" pageNumber="576">
<emphasis id="B94EEAF4C66AFF9FAE44F89F0A0AF822" bold="true" box="[813,1036,1915,1939]" pageId="12" pageNumber="576">Material examined.</emphasis>
<materialsCitation id="3B523CBBC66AFF9FA97DF89F0D60F842" collectingDate="2013-09-14" collectionCode="ZRC" country="Malaysia" county="Ng" elevation="187" location="Nawang Waterfall" municipality="Bau" pageId="12" pageNumber="576" specimenCode="ZRC 2015.0611" specimenCount="1" typeStatus="holotype">
<typeStatus id="54818844C66AFF9FA97DF89F0A7DF822" box="[1044,1147,1915,1939]" pageId="12" pageNumber="576" type="holotype">Holotype</typeStatus>
: male (8.4 ×
<quantity id="4CC29B03C66AFF9FA862F89F0B59F822" box="[1291,1375,1915,1939]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="12" pageNumber="576" unit="mm" value="7.6">7.6 mm</quantity>
) (
<specimenCode id="DB9C9E9DC66AFF9FA81FF89F0DA4F802" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="576">ZRC 2015.0611</specimenCode>
),
<location id="8EE5603DC66AFF9FAEDBF8780A71F802" LSID="urn:lsid:plazi:treatment:039387F0C66AFF9DAEF3F8FF0A2FF7A4:8EE5603DC66AFF9FAEDBF8780A71F802" box="[946,1143,1947,1972]" country="Malaysia" county="Ng" municipality="Bau" name="Nawang Waterfall" pageId="12" pageNumber="576">Nawang Waterfall</location>
,
<elevation id="0017D1D5C66AFF9FA9E9F87F0B6EF802" box="[1152,1384,1947,1971]" metricMagnitude="2" metricUnit="m" metricValue="1.8699999999999999" pageId="12" pageNumber="576" unit="m" value="187.0">
<quantity id="4CC29B03C66AFF9FA9E9F87F0AC4F802" box="[1152,1218,1947,1971]" metricMagnitude="2" metricUnit="m" metricValue="1.8699999999999999" pageId="12" pageNumber="576" unit="m" value="187.0">187 m</quantity>
above sea level
</elevation>
,
<location id="8EE5603DC66AFF9FA81BF8780D71F862" LSID="urn:lsid:plazi:treatment:039387F0C66AFF9DAEF3F8FF0A2FF7A4:8EE5603DC66AFF9FA81BF8780D71F862" country="Malaysia" county="Ng" municipality="Bau" name="Bung Bratak" pageId="12" pageNumber="576">Bung Bratak</location>
,
<collectingMunicipality id="6BE1AC9CC66AFF9FAEEDF8580DB4F862" box="[900,946,1980,2003]" pageId="12" pageNumber="576">Bau</collectingMunicipality>
,
<collectingCountry id="F32D7676C66AFF9FAEA9F85F0A19F862" box="[960,1055,1979,2003]" name="Malaysia" pageId="12" pageNumber="576">Sarawak</collectingCountry>
, coll. PKL
<collectingCounty id="62E44E6AC66AFF9FA9CBF8580AC5F865" box="[1186,1219,1980,2004]" pageId="12" pageNumber="576">Ng</collectingCounty>
et al.,
<date id="FF841026C66AFF9FA87DF85F0D60F842" pageId="12" pageNumber="576" value="2013-09-14">
<collectingDate id="EFC0E9CEC66AFF9FA87DF85F0D60F842" pageId="12" pageNumber="576" value="2013-09-14">14 September 2013</collectingDate>
</date>
</materialsCitation>
.
<typeStatus id="54818844C66AFF9FAE07F8380DDCF845" box="[878,986,2012,2036]" pageId="12" pageNumber="576" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D3CFD6FC66AFF9FAE8AF83F0A44F842" box="[995,1090,2011,2035]" pageId="12" pageNumber="576" type="male">14 males</specimenCount>
(largest 8.0 ×
<quantity id="4CC29B03C66AFF9FA9B0F83F0B2BF842" box="[1241,1325,2011,2035]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="12" pageNumber="576" unit="mm" value="7.1">7.1 mm</quantity>
),
<specimenCount id="9D3CFD6FC66AFF9FA856F8380BAEF842" box="[1343,1448,2011,2035]" pageId="12" pageNumber="576" type="female">7 females</specimenCount>
, 2 ovigerous females (largest 7.4 × 6.0 mm),
<materialsCitation id="3B523CBBC66AFF9FA869F81F0AABF782" collectingDate="2013-09-14" collectionCode="ZRC" country="Malaysia" county="Ng" elevation="187" location="Nawang Waterfall" municipality="Bau" pageId="12" pageNumber="576" specimenCode="ZRC 2015.0612" specimenCount="1" specimenCount-juvenile="1" typeStatus="paratype">
<specimenCount id="9D3CFD6FC66AFF9FA869F81F0B6FF7A2" box="[1280,1385,2043,2067]" pageId="12" pageNumber="576" type="juvenile">1 juvenile</specimenCount>
(
<specimenCode id="DB9C9E9DC66AFF9FA81FF81F0DA5F782" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="12" pageNumber="576">ZRC 2015.0612</specimenCode>
), data same as holotype
</materialsCitation>
;
<materialsCitation id="3B523CBBC66AFF9DA9D3F7FF0C11FB44" collectingDate="2013-09-14" collectionCode="ZRC" collectorName="J Grinang" country="Malaysia" county="Ng" elevation="187" lastPageId="14" lastPageNumber="578" location="Nawang Waterfall" municipality="Bau" pageId="12" pageNumber="576" specimenCount="4" specimenCount-male="4" typeStatus="paratype">
<specimenCount id="9D3CFD6FC66AFF9FA9D3F7FF0B09F782" box="[1210,1295,2075,2099]" pageId="12" pageNumber="576" type="male">4 males</specimenCount>
(largest 9.0 ×
<quantity id="4CC29B03C668FF9DAD18FB590EC0FB64" box="[113,198,1213,1237]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="14" pageNumber="578" unit="mm" value="7.9">7.9 mm</quantity>
) (SBC.00431–34), location same as holotype, coll.
<collectorName id="26CF5330C668FF9DAD18FB3A0EE7FB44" box="[113,225,1245,1269]" pageId="14" pageNumber="578">J Grinang</collectorName>
et al.,
<date id="FF841026C668FF9DAC5FFB390C11FB44" box="[310,535,1245,1269]" pageId="14" pageNumber="578" value="2013-09-14">
<collectingDate id="EFC0E9CEC668FF9DAC5FFB390C11FB44" box="[310,535,1245,1269]" pageId="14" pageNumber="578" value="2013-09-14">14 September 2013</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B523CBBC668FF9DAF4FFB390EFCFA84" collectingDate="2013-05-01" collectionCode="ZRC" collectorName="J Grinang" country="Malaysia" county="Ng" elevation="187" location="Nawang Waterfall" municipality="Bau" pageId="14" pageNumber="578" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D3CFD6FC668FF9DAF4FFB390C75FB44" box="[550,627,1245,1269]" pageId="14" pageNumber="578" type="male">1 male</specimenCount>
(8.0 × 7.0 mm) (SBC.C.00476), location same as holotype,
<collectorName id="26CF5330C668FF9DAFE8FB1A0CEBFAA4" box="[641,749,1277,1301]" pageId="14" pageNumber="578">J Grinang</collectorName>
,
<date id="FF841026C668FF9DAD18FAF90EFCFA84" box="[113,250,1309,1334]" pageId="14" pageNumber="578" value="2013-05-01">
<collectingDate id="EFC0E9CEC668FF9DAD18FAF90EFCFA84" box="[113,250,1309,1334]" pageId="14" pageNumber="578" value="2013-05-01">1 May 2013</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF45666EC66BFF9EADE1F8320A9DF7F2" ID-DOI="http://doi.org/10.5281/zenodo.5386749" ID-Zenodo-Dep="5386749" httpUri="https://zenodo.org/record/5386749/files/figure.png" pageId="13" pageNumber="577" startId="13.[136,171,2006,2028]" targetBox="[176,1431,189,1991]" targetPageId="13">
<paragraph id="8B8536E6C66BFF9EADE1F8320A9DF7F2" blockId="13.[136,1474,2006,2115]" pageId="13" pageNumber="577">
Fig. 11.
<taxonomicName id="4C3A4D65C66BFF9EADB4F8320FCCF85D" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[221,458,2006,2028]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="577" phylum="Arthropoda" rank="species" species="sangon" status="sp. nov.">
<emphasis id="B94EEAF4C66BFF9EADB4F8320FCCF85D" box="[221,458,2006,2028]" italics="true" pageId="13" pageNumber="577">Lepidothelphusa sangon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC66BFF9EACB1F8330C56F85D" box="[472,592,2006,2028]" pageId="13" pageNumber="577" rank="species">new species</taxonomicNameLabel>
, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela.
</paragraph>
</caption>
<caption id="DF45666EC668FF9DAD18FBD20CBEFB37" ID-DOI="http://doi.org/10.5281/zenodo.5386751" ID-Zenodo-Dep="5386751" httpUri="https://zenodo.org/record/5386751/files/figure.png" pageId="14" pageNumber="578" startId="14.[113,148,1078,1100]" targetBox="[116,1452,188,1063]" targetPageId="14">
<paragraph id="8B8536E6C668FF9DAD18FBD20CBEFB37" blockId="14.[113,1451,1078,1158]" pageId="14" pageNumber="578">
Fig. 12.
<taxonomicName id="4C3A4D65C668FF9DADACFBD20FB4FBFD" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[197,434,1078,1100]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="578" phylum="Arthropoda" rank="species" species="sangon" status="sp. nov.">
<emphasis id="B94EEAF4C668FF9DADACFBD20FB4FBFD" box="[197,434,1078,1100]" italics="true" pageId="14" pageNumber="578">Lepidothelphusa sangon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC668FF9DACA8FBD30C3FFBFD" box="[449,569,1078,1100]" pageId="14" pageNumber="578" rank="species">new species</taxonomicNameLabel>
, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of left G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C320656DC668FF9DAD18FAB90A09FAA4" pageId="14" pageNumber="578" type="diagnosis">
<paragraph id="8B8536E6C668FF9DAD18FAB90F4DF844" blockId="14.[113,753,1373,2038]" pageId="14" pageNumber="578">
<emphasis id="B94EEAF4C668FF9DAD18FAB90EE0FAC4" bold="true" box="[113,230,1373,1397]" pageId="14" pageNumber="578">Diagnosis.</emphasis>
Carapace quadrate, surface relatively flat, rugose, anterolateral margins smooth; antero- and posterolateral regions rugose; epibranchial tooth undiscernible; external orbital tooth with outer margin concave, smooth; serrated projection on frontal region distinct; epigastric and postorbital cristae distinct, margins serrated; cervical and H-grooves shallow, not confluent (
<figureCitation id="13012A63C668FF9DAC10F9F90C20F984" box="[377,550,1565,1589]" captionStart="Fig" captionStartId="13.[136,171,2006,2028]" captionTargetBox="[176,1431,189,1991]" captionTargetId="figure-90@13.[176,1431,189,1991]" captionTargetPageId="13" captionText="Fig. 11. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386749" httpUri="https://zenodo.org/record/5386749/files/figure.png" pageId="14" pageNumber="578">Fig. 11A, B, D</figureCitation>
); ischium of third maxilliped rhomboidal, slightly longer than broad sulcus undiscernible (
<figureCitation id="13012A63C668FF9DAC66F9B90F6AF9C4" box="[271,364,1629,1653]" captionStart="Fig" captionStartId="13.[136,171,2006,2028]" captionTargetBox="[176,1431,189,1991]" captionTargetId="figure-90@13.[176,1431,189,1991]" captionTargetPageId="13" captionText="Fig. 11. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386749" httpUri="https://zenodo.org/record/5386749/files/figure.png" pageId="14" pageNumber="578">Fig. 11F</figureCitation>
); inner margin of merus of chelipeds with relatively lower serrated projection (
<figureCitation id="13012A63C668FF9DAF77F9990C78F924" box="[542,638,1661,1685]" captionStart="Fig" captionStartId="13.[136,171,2006,2028]" captionTargetBox="[176,1431,189,1991]" captionTargetId="figure-90@13.[176,1431,189,1991]" captionTargetPageId="13" captionText="Fig. 11. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386749" httpUri="https://zenodo.org/record/5386749/files/figure.png" pageId="14" pageNumber="578">Fig. 11G</figureCitation>
); carpus of chelipeds rugose, inner angle with broad, low triangular tooth (
<figureCitation id="13012A63C668FF9DAD10F9590ED9F964" box="[121,223,1725,1749]" captionStart="Fig" captionStartId="13.[136,171,2006,2028]" captionTargetBox="[176,1431,189,1991]" captionTargetId="figure-90@13.[176,1431,189,1991]" captionTargetPageId="13" captionText="Fig. 11. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386749" httpUri="https://zenodo.org/record/5386749/files/figure.png" pageId="14" pageNumber="578">Fig. 11A</figureCitation>
); fingers of adult major male chela gaping (
<figureCitation id="13012A63C668FF9DAFACF9590EBFF944" captionStart="Fig" captionStartId="13.[136,171,2006,2028]" captionTargetBox="[176,1431,189,1991]" captionTargetId="figure-90@13.[176,1431,189,1991]" captionTargetPageId="13" captionText="Fig. 11. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386749" httpUri="https://zenodo.org/record/5386749/files/figure.png" pageId="14" pageNumber="578">Fig. 11H, I</figureCitation>
); ambulatory legs short, merus of fourth ambulatory legs about half of length of carapace (
<figureCitation id="13012A63C668FF9DAF74F9190CA0F8A4" box="[541,678,1789,1813]" captionStart="Fig" captionStartId="13.[136,171,2006,2028]" captionTargetBox="[176,1431,189,1991]" captionTargetId="figure-90@13.[176,1431,189,1991]" captionTargetPageId="13" captionText="Fig. 11. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386749" httpUri="https://zenodo.org/record/5386749/files/figure.png" pageId="14" pageNumber="578">Fig. 11A, E</figureCitation>
); male abdomen triangular, somite 6 width and length subequal, subequal to length of telson (
<figureCitation id="13012A63C668FF9DACDFF8D90C1DF8E4" box="[438,539,1853,1877]" captionStart="Fig" captionStartId="13.[136,171,2006,2028]" captionTargetBox="[176,1431,189,1991]" captionTargetId="figure-90@13.[176,1431,189,1991]" captionTargetPageId="13" captionText="Fig. 11. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386749" httpUri="https://zenodo.org/record/5386749/files/figure.png" pageId="14" pageNumber="578">Fig. 11C</figureCitation>
); G1 stout, slightly sinuous, basal segment weakly constricted on distal third; terminal segment slightly curving outwards in dorsal view, almost straight in ventral view (
<figureCitation id="13012A63C668FF9DACACF8790C83F804" box="[453,645,1949,1973]" captionStart="Fig" captionStartId="14.[113,148,1078,1100]" captionTargetBox="[116,1452,188,1063]" captionTargetId="figure-12@14.[92,1472,172,1070]" captionTargetPageId="14" captionText="Fig. 12. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of left G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386751" httpUri="https://zenodo.org/record/5386751/files/figure.png" pageId="14" pageNumber="578">Fig. 12A, B, D, E</figureCitation>
); G2 with relatively long distal segment, about half length of basal segment (
<figureCitation id="13012A63C668FF9DADB2F8390F39F844" box="[219,319,2013,2037]" captionStart="Fig" captionStartId="14.[113,148,1078,1100]" captionTargetBox="[116,1452,188,1063]" captionTargetId="figure-12@14.[92,1472,172,1070]" captionTargetPageId="14" captionText="Fig. 12. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of left G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386751" httpUri="https://zenodo.org/record/5386751/files/figure.png" pageId="14" pageNumber="578">Fig. 12C</figureCitation>
).
</paragraph>
<paragraph id="8B8536E6C668FF9DAE44FB590A09FAA4" blockId="14.[813,1451,1213,1301]" pageId="14" pageNumber="578">
<emphasis id="B94EEAF4C668FF9DAE44FB590D82FB64" bold="true" box="[813,900,1213,1237]" pageId="14" pageNumber="578">Colour.</emphasis>
In life, both sexes are red to reddish-brown with yellow chelipeds (
<figureCitation id="13012A63C668FF9DAE93FB390A79FB44" box="[1018,1151,1245,1269]" captionStart="Fig" captionStartId="16.[113,148,1308,1330]" captionTargetBox="[113,1452,189,1293]" captionTargetId="figure-491@16.[113,1452,188,1294]" captionTargetPageId="16" captionText="Fig. 14. Live colours of three Lepidothelphusa species from Kuching Division, southwestern Sarawak. A, B, Lepidothelphusa loi, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak; C, D, Lepidothelphusa padawan, new species, paratype male (15.8 × 13.2 mm) (SBC.C.00440–4), Bung Brungu, Padawan, Sarawak; E, F, Lepidothelphusa sangon, new species, paratype male (8.0 × 7.0 mm) (SBC.C.00476), Nawang Waterfall, Bung Bratak, Bau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5386759" httpUri="https://zenodo.org/record/5386759/files/figure.png" pageId="14" pageNumber="578">Fig. 14E, F</figureCitation>
). The colour in females is slightly less striking.
</paragraph>
</subSubSection>
<subSubSection id="C320656DC668FF9DAE44FAD90B3EFA24" pageId="14" pageNumber="578" type="etymology">
<paragraph id="8B8536E6C668FF9DAE44FAD90B3EFA24" blockId="14.[813,1452,1341,1429]" pageId="14" pageNumber="578">
<emphasis id="B94EEAF4C668FF9DAE44FAD90DA9FAE4" bold="true" box="[813,943,1341,1365]" pageId="14" pageNumber="578">Etymology.</emphasis>
The name is derived from the Bidayuh word “sangon” for beautiful, alluding to the striking colour of the crab. The name is used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C320656DC668FF9DAE44FA590A2FF7A4" pageId="14" pageNumber="578" type="discussion">
<paragraph id="8B8536E6C668FF9DAE44FA590A1DF8C4" blockId="14.[812,1452,1469,1909]" pageId="14" pageNumber="578">
<emphasis id="B94EEAF4C668FF9DAE44FA590D9BFA64" bold="true" box="[813,925,1469,1493]" pageId="14" pageNumber="578">Remarks.</emphasis>
<taxonomicName id="4C3A4D65C668FF9DAEC1FA590ABEFA64" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[936,1208,1469,1493]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="578" phylum="Arthropoda" rank="species" species="sangon" status="sp. nov.">
<emphasis id="B94EEAF4C668FF9DAEC1FA590ABEFA64" box="[936,1208,1469,1493]" italics="true" pageId="14" pageNumber="578">Lepidothelphusa sangon</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC668FF9DA9A0FA5A0B55FA64" box="[1225,1363,1469,1493]" pageId="14" pageNumber="578" rank="species">new species</taxonomicNameLabel>
, can be easily distinguished from
<taxonomicName id="4C3A4D65C668FF9DA920FA3A0AC2FA44" authorityName="Nobili" authorityYear="1903" box="[1097,1220,1502,1525]" class="Malacostraca" family="Potamidae" genus="Potamon" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="578" phylum="Arthropoda" rank="species" sensu="stricto" species="cognettii">
<emphasis id="B94EEAF4C668FF9DA920FA3A0AC2FA44" box="[1097,1220,1502,1525]" italics="true" pageId="14" pageNumber="578">L. cognettii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D578FC668FF9DA9A2FA3A0B0EFA44" box="[1227,1288,1502,1525]" pageId="14" pageNumber="578" sensu="stricto">s. str.</taxonomicNameLabel>
,
<taxonomicName id="4C3A4D65C668FF9DA87CFA3A0B76FA44" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[1301,1392,1501,1525]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="578" phylum="Arthropoda" rank="species" species="limau" status="sp. nov.">
<emphasis id="B94EEAF4C668FF9DA87CFA3A0B76FA44" box="[1301,1392,1501,1525]" italics="true" pageId="14" pageNumber="578">L. limau</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC668FF9DA816FA3A0D7BF9A4" pageId="14" pageNumber="578" rank="species">new species</taxonomicNameLabel>
, and
<taxonomicName id="4C3A4D65C668FF9DAEDBFA1A0A36F9A4" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[946,1072,1533,1557]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="578" phylum="Arthropoda" rank="species" species="padawan" status="sp. nov.">
<emphasis id="B94EEAF4C668FF9DAEDBFA1A0A36F9A4" box="[946,1072,1533,1557]" italics="true" pageId="14" pageNumber="578">L. padawan</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC668FF9DA955FA1A0AB8F9A4" box="[1084,1214,1533,1557]" pageId="14" pageNumber="578" rank="species">new species</taxonomicNameLabel>
, by its relatively small adult size (males and females are already fully mature at 8.0 mm carapace width). Its size range is similar to
<taxonomicName id="4C3A4D65C668FF9DA874F9DA0BA3F9E4" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[1309,1445,1597,1621]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="578" phylum="Arthropoda" rank="species" species="flavochela" status="sp. nov.">
<emphasis id="B94EEAF4C668FF9DA874F9DA0BA3F9E4" box="[1309,1445,1597,1621]" italics="true" pageId="14" pageNumber="578">L. flavochela</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC668FF9DAE44F9BA0DABF9C4" box="[813,941,1629,1653]" pageId="14" pageNumber="578" rank="species">new species</taxonomicNameLabel>
, and
<taxonomicName id="4C3A4D65C668FF9DAE88F9BA0A11F9C4" authority="Grinang &amp; Ng, 2015" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[993,1047,1629,1653]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="578" phylum="Arthropoda" rank="species" species="loi" status="sp. nov.">
<emphasis id="B94EEAF4C668FF9DAE88F9BA0A11F9C4" box="[993,1047,1629,1653]" italics="true" pageId="14" pageNumber="578">L. loi</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A27D578FC668FF9DA94AF9BA0AA5F9C4" box="[1059,1187,1629,1653]" pageId="14" pageNumber="578" rank="species">new species</taxonomicNameLabel>
, but distinguishable from the latter by its striking reddish carapace and ambulatory legs (
<figureCitation id="13012A63C668FF9DAE00F9790DE0F904" box="[873,998,1693,1717]" captionStart="Fig" captionStartId="16.[113,148,1308,1330]" captionTargetBox="[113,1452,189,1293]" captionTargetId="figure-491@16.[113,1452,188,1294]" captionTargetPageId="16" captionText="Fig. 14. Live colours of three Lepidothelphusa species from Kuching Division, southwestern Sarawak. A, B, Lepidothelphusa loi, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak; C, D, Lepidothelphusa padawan, new species, paratype male (15.8 × 13.2 mm) (SBC.C.00440–4), Bung Brungu, Padawan, Sarawak; E, F, Lepidothelphusa sangon, new species, paratype male (8.0 × 7.0 mm) (SBC.C.00476), Nawang Waterfall, Bung Bratak, Bau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5386759" httpUri="https://zenodo.org/record/5386759/files/figure.png" pageId="14" pageNumber="578">Fig. 14E, F</figureCitation>
versus
<figureCitation id="13012A63C668FF9DA955F9790ACDF904" box="[1084,1227,1693,1717]" captionStart="Fig" captionStartId="15.[136,171,1302,1324]" captionTargetBox="[136,1474,189,1286]" captionTargetId="figure-77@15.[136,1474,188,1286]" captionTargetPageId="15" captionText="Fig. 13. Live colours of three Lepidothelphusa species from Kuching Division, southwestern Sarawak. A, B, Lepidothelphusa cognettii (Nobili, 1903), male (12.0 × 10.2 mm) (SBC.C.00382–90), Batu Panggah Trail, Gunung Penrissen, Sarawak; C, D, Lepidothelphusa flavochela, new species, paratype male (8.5 × 7.5 mm) (SBC.C.00392–401), Pagar Besi, Kampung Gumbang, Bau, Sarawak; E, F, Lepidothelphusa limau, new species, paratype male (10.5 × 9.7 mm) (SBC.C.00402–05), Kampung Peros, Bau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5386755" httpUri="https://zenodo.org/record/5386755/files/figure.png" pageId="14" pageNumber="578">Figs. 13C, D</figureCitation>
,
<figureCitation id="13012A63C668FF9DA9B0F9790B2CF904" box="[1241,1322,1693,1717]" captionStart="Fig" captionStartId="16.[113,148,1308,1330]" captionTargetBox="[113,1452,189,1293]" captionTargetId="figure-491@16.[113,1452,188,1294]" captionTargetPageId="16" captionText="Fig. 14. Live colours of three Lepidothelphusa species from Kuching Division, southwestern Sarawak. A, B, Lepidothelphusa loi, holotype male (10.3 × 9.6 mm) (ZRC 2015.0607), Gunung Ampungan, Sadong River Basin, Serian, Sarawak; C, D, Lepidothelphusa padawan, new species, paratype male (15.8 × 13.2 mm) (SBC.C.00440–4), Bung Brungu, Padawan, Sarawak; E, F, Lepidothelphusa sangon, new species, paratype male (8.0 × 7.0 mm) (SBC.C.00476), Nawang Waterfall, Bung Bratak, Bau, Sarawak." figureDoi="http://doi.org/10.5281/zenodo.5386759" httpUri="https://zenodo.org/record/5386759/files/figure.png" pageId="14" pageNumber="578">14A, B</figureCitation>
). It is close to
<taxonomicName id="4C3A4D65C668FF9DAE22F95A0DA1F964" authorityName="Grinang &amp; Ng" authorityYear="2015" box="[843,935,1725,1749]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" kingdom="Animalia" order="Decapoda" pageId="14" pageNumber="578" phylum="Arthropoda" rank="species" species="limau">
<emphasis id="B94EEAF4C668FF9DAE22F95A0DA1F964" box="[843,935,1725,1749]" italics="true" pageId="14" pageNumber="578">L. limau</emphasis>
</taxonomicName>
, but can be distinguished by its more squarish and flat carapace (
<figureCitation id="13012A63C668FF9DA96EF9390A6BF944" box="[1031,1133,1757,1781]" captionStart="Fig" captionStartId="13.[136,171,2006,2028]" captionTargetBox="[176,1431,189,1991]" captionTargetId="figure-90@13.[176,1431,189,1991]" captionTargetPageId="13" captionText="Fig. 11. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386749" httpUri="https://zenodo.org/record/5386749/files/figure.png" pageId="14" pageNumber="578">Fig. 11B</figureCitation>
versus
<figureCitation id="13012A63C668FF9DA9A5F9390B21F944" box="[1228,1319,1757,1781]" captionStart="Fig" captionStartId="7.[136,171,2006,2028]" captionTargetBox="[177,1434,189,1991]" captionTargetId="figure-90@7.[177,1434,189,1991]" captionTargetPageId="7" captionText="Fig. 5. Lepidothelphusa limau, new species, holotype male (12.1 × 10.8 mm) (ZRC 2015.0605), Kampung Peros, Krokong, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386729" httpUri="https://zenodo.org/record/5386729/files/figure.png" pageId="14" pageNumber="578">Fig. 5B</figureCitation>
), relatively smaller projection on the merus of the cheliped (
<figureCitation id="13012A63C668FF9DA820F9190BAAF8A4" box="[1353,1452,1789,1813]" captionStart="Fig" captionStartId="13.[136,171,2006,2028]" captionTargetBox="[176,1431,189,1991]" captionTargetId="figure-90@13.[176,1431,189,1991]" captionTargetPageId="13" captionText="Fig. 11. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386749" httpUri="https://zenodo.org/record/5386749/files/figure.png" pageId="14" pageNumber="578">Fig. 11G</figureCitation>
versus
<figureCitation id="13012A63C668FF9DAE13F8F90DD6F884" box="[890,976,1821,1845]" captionStart="Fig" captionStartId="7.[136,171,2006,2028]" captionTargetBox="[177,1434,189,1991]" captionTargetId="figure-90@7.[177,1434,189,1991]" captionTargetPageId="7" captionText="Fig. 5. Lepidothelphusa limau, new species, holotype male (12.1 × 10.8 mm) (ZRC 2015.0605), Kampung Peros, Krokong, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386729" httpUri="https://zenodo.org/record/5386729/files/figure.png" pageId="14" pageNumber="578">Fig. 5G</figureCitation>
), narrower gap of the fingers of chela (
<figureCitation id="13012A63C668FF9DA8E9F8F90D5CF8E4" captionStart="Fig" captionStartId="13.[136,171,2006,2028]" captionTargetBox="[176,1431,189,1991]" captionTargetId="figure-90@13.[176,1431,189,1991]" captionTargetPageId="13" captionText="Fig. 11. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386749" httpUri="https://zenodo.org/record/5386749/files/figure.png" pageId="14" pageNumber="578">Fig. 11H</figureCitation>
versus
<figureCitation id="13012A63C668FF9DAEDAF8D90A0FF8E4" box="[947,1033,1853,1877]" captionStart="Fig" captionStartId="7.[136,171,2006,2028]" captionTargetBox="[177,1434,189,1991]" captionTargetId="figure-90@7.[177,1434,189,1991]" captionTargetPageId="7" captionText="Fig. 5. Lepidothelphusa limau, new species, holotype male (12.1 × 10.8 mm) (ZRC 2015.0605), Kampung Peros, Krokong, Bau, Sarawak. A, overall dorsal view of habitus; B, dorsal view of carapace; C, ventral view showing anterior thoracic sternum and male abdomen; D, frontal view of carapace; E, outer view of right fourth ambulatory leg; F, left third maxilliped showing exopod; G, left cheliped showing serrated inner margin of merus; H, outer view of left chela; I, outer view of right chela." figureDoi="http://doi.org/10.5281/zenodo.5386729" httpUri="https://zenodo.org/record/5386729/files/figure.png" pageId="14" pageNumber="578">Fig. 5H</figureCitation>
) and relatively shorter G1 (
<figureCitation id="13012A63C668FF9DA828F8D90D46F8C4" captionStart="Fig" captionStartId="14.[113,148,1078,1100]" captionTargetBox="[116,1452,188,1063]" captionTargetId="figure-12@14.[92,1472,172,1070]" captionTargetPageId="14" captionText="Fig. 12. Lepidothelphusa sangon, new species, holotype male (8.4 × 7.6 mm) (ZRC 2015.0611), Nawang Waterfall, Bung Bratak, Bau, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of left G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386751" httpUri="https://zenodo.org/record/5386751/files/figure.png" pageId="14" pageNumber="578">Fig. 12A, D</figureCitation>
versus
<figureCitation id="13012A63C668FF9DAEFCF8B90A08F8C4" box="[917,1038,1885,1909]" captionStart="Fig" captionStartId="8.[113,148,958,980]" captionTargetBox="[114,1452,188,944]" captionTargetId="figure-12@8.[100,1462,177,951]" captionTargetPageId="8" captionText="Fig. 6. Lepidothelphusa limau, new species, holotype male (12.1 × 10.8 mm) (ZRC 2015.0605), Kampung Peros, Krokong, Bau, Sarawak. A, dorsal view of left G1; B, dorsal view of distal part of left G1; C, dorsal view of right G2; D, ventral view of left G1; E, ventral view of distal part of left G1. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5386733" httpUri="https://zenodo.org/record/5386733/files/figure.png" pageId="14" pageNumber="578">Fig. 6A, D</figureCitation>
).
</paragraph>
<paragraph id="8B8536E6C668FF9DAE44F8790A2FF7A4" blockId="14.[813,1452,1949,2069]" pageId="14" pageNumber="578">
<emphasis id="B94EEAF4C668FF9DAE44F8790D8DF804" bold="true" box="[813,907,1949,1973]" pageId="14" pageNumber="578">Habitat.</emphasis>
The habitat is moist rocks and leaf litter, in partially exposed or shaded, small streams in old primary dipterocarp forests. The substrate is mudstone and the site is less than
<quantity id="4CC29B03C668FF9DAE44F8190D74F7A4" box="[813,882,2045,2069]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="578" unit="m" value="200.0">200 m</quantity>
above sea level.
</paragraph>
</subSubSection>
</treatment>
</document>
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