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<document id="823E9D24CDEBA9A36BCF5065FA2B869B" ID-DOI="10.5281/zenodo.215396" ID-GBIF-Dataset="62b67fc1-89a3-4bb2-bc29-531913e5b2ff" ID-ISSN="1175-5326" ID-Zenodo-Dep="215396" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460556383699" checkinUser="plazi" docAuthor="Rakotoarison, Andolalao, Glaw, Frank, Vieites, David R., Raminosoa, Noromalala R. &amp; Vences, Miguel" docDate="2012" docId="039387E38905FFB6FF7539CAFF0CFCEC" docLanguage="en" docName="zt03563p025.pdf" docOrigin="Zootaxa 3563" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Platypelis olgae Rakotoarison, Glaw, Vieites, Raminosoa &amp; Vences, 2012, sp. nov." docType="treatment" docVersion="7" lastPageNumber="20" masterDocId="FFAAFF9B8915FFA5FFE23C2BFFE3FFC0" masterDocTitle="Taxonomy and natural history of arboreal microhylid frogs (Platypelis) from the Tsaratanana Massif in northern Madagascar, with description of a new species" masterLastPageNumber="25" masterPageNumber="1" pageNumber="17" updateTime="1698319345731" updateUser="plazi">
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<mods:title id="FB3EE50CC7FF8EFCEB52B4D8C5C71BC1">Taxonomy and natural history of arboreal microhylid frogs (Platypelis) from the Tsaratanana Massif in northern Madagascar, with description of a new species</mods:title>
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<mods:namePart id="1362B9F16B5C3427FF3166D3C61C11E8">Rakotoarison, Andolalao</mods:namePart>
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<mods:namePart id="DFA8E634064C143CC8CE333436FFF6C7">Glaw, Frank</mods:namePart>
</mods:name>
<mods:name id="7C273775F9AB2AF5A304A7EA620A7D4B" type="personal">
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<mods:namePart id="89244D8BD41803C06E8E5167C7DAB994">Vieites, David R.</mods:namePart>
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<mods:namePart id="AEC82C5A8016732505DD91491032B76A">Raminosoa, Noromalala R.</mods:namePart>
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<mods:namePart id="08F81C4343D402A9966DD6D826746B51">Vences, Miguel</mods:namePart>
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<mods:typeOfResource id="3C7F3CAB9229493F723C8C61A362BAD4">text</mods:typeOfResource>
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<mods:title id="E22DC9B7623DAF37426B56EB63B9473B">Zootaxa</mods:title>
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<mods:date id="81A9463BBDC7B76332EFF88BD5B797C9">2012</mods:date>
<mods:detail id="F19C16D312E82EFC796AC3CDFEA23324" type="volume">
<mods:number id="E486DA8C7FCA184C13BD14DCD4963609">3563</mods:number>
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<mods:identifier id="3DE4A0469028DA4057C0EE40BBEE2222" type="DOI">10.5281/zenodo.215396</mods:identifier>
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<mods:identifier id="5037FDD187498870C782DCF35D517BF8" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="10175B044DD81D1526E419916481FF8F" type="Zenodo-Dep">215396</mods:identifier>
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<treatment id="039387E38905FFB6FF7539CAFF0CFCEC" ID-DOI="http://doi.org/10.5281/zenodo.6176526" ID-GBIF-Taxon="119555838" ID-Zenodo-Dep="6176526" LSID="urn:lsid:plazi:treatment:039387E38905FFB6FF7539CAFF0CFCEC" httpUri="http://treatment.plazi.org/id/039387E38905FFB6FF7539CAFF0CFCEC" lastPageId="19" lastPageNumber="20" pageId="16" pageNumber="17">
<subSubSection id="C320657E8905FFB5FF7539CAFE5AFA3B" box="[151,441,1505,1531]" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="8B8536F58905FFB5FF7539CAFE5AFA3B" blockId="16.[151,441,1505,1531]" box="[151,441,1505,1531]" pageId="16" pageNumber="17">
<heading id="D0CD81998905FFB5FF7539CAFE5AFA3B" bold="true" box="[151,441,1505,1531]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="B94EEAE78905FFB5FF7539CAFE5AFA3B" bold="true" box="[151,441,1505,1531]" pageId="16" pageNumber="17">
<taxonomicName id="4C3A4D768905FFB5FF7539CAFEB7FA3B" ID-CoL="4JVGK" box="[151,340,1505,1531]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="species" species="olgae" status="sp. nov.">
<emphasis id="B94EEAE78905FFB5FF7539CAFEB7FA3B" bold="true" box="[151,340,1505,1531]" italics="true" pageId="16" pageNumber="17">Platypelis olgae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27D579C8905FFB5FEBE39CAFE5AFA3B" box="[348,441,1505,1531]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C320657E8905FFB5FF753A03FE70F8D8" pageId="16" pageNumber="17" type="materials_examined">
<paragraph id="8B8536F58905FFB5FF753A03FB31F96C" blockId="16.[151,1436,1576,2032]" pageId="16" pageNumber="17">
<emphasis id="B94EEAE78905FFB5FF753A03FEE8F980" bold="true" box="[151,267,1576,1600]" pageId="16" pageNumber="17">
<typeStatus id="548188578905FFB5FF753A03FEE5F980" box="[151,262,1576,1600]" pageId="16" pageNumber="17" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED2BAE308905FFB5FEF73A03FEACF980" box="[277,335,1576,1600]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="16" pageNumber="17">ZSM</collectionCode>
1666/2010 (
<collectionCode id="ED2BAE308905FFB5FE003A03FDD6F980" box="[482,565,1576,1600]" pageId="16" pageNumber="17">ZCMV</collectionCode>
12402), adult male, collected at the Tsaratanana Massif, between Camps 3 (Bepia) and 4 (Andranomadio), without precise geographical coordinates or altitude data, but located in between these two campsites (coordinates given in Materials and Methods) with altitudes of
<date id="FF8410358905FFB5FBA93A5BFB10F948" box="[1099,1267,1648,1672]" pageId="16" pageNumber="17" value="2294" valueMax="2503">
2294 and
<quantity id="4CC29B108905FFB5FB593A5BFB10F948" box="[1211,1267,1648,1672]" metricMagnitude="3" metricUnit="m" metricValue="2.503" pageId="16" pageNumber="17" unit="m" value="2503.0">2503</quantity>
</date>
m a.s.l., on
<date id="FF8410358905FFB5FA623A5BFEEFF96C" pageId="16" pageNumber="17" value="2010-06-16">16 June 2010</date>
, by M. Vences, D. R. Vieites, R.-D. Randrianiaina, S. Rasamison, and E. Rajeriarison.
</paragraph>
<paragraph id="8B8536F58905FFB5FF253A93FE70F8D8" blockId="16.[151,1436,1576,2032]" pageId="16" pageNumber="17">
<emphasis id="B94EEAE78905FFB5FF253A93FEA5F910" bold="true" box="[199,326,1720,1744]" pageId="16" pageNumber="17">
<typeStatus id="548188578905FFB5FF253A93FEA2F910" box="[199,321,1720,1744]" pageId="16" pageNumber="17" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED2BAE308905FFB5FEAE3A93FE65F910" box="[332,390,1720,1744]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="16" pageNumber="17">ZSM</collectionCode>
1667/2010 (
<collectionCode id="ED2BAE308905FFB5FDF03A93FD86F910" box="[530,613,1720,1744]" pageId="16" pageNumber="17">ZCMV</collectionCode>
12428, female) and
<collectionCode id="ED2BAE308905FFB5FCA83A93FC60F910" box="[842,899,1720,1744]" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="16" pageNumber="17">ZSM</collectionCode>
1668/2010 (DRV 6225, male), both collected at Tsaratanana Camp 5 (Amboditsaratanana),
<geoCoordinate id="EE0E50328905FFB5FD703AF7FCFBF934" box="[658,792,1756,1780]" direction="south" orientation="latitude" pageId="16" pageNumber="17" precision="1" value="-14.08019">14.08019°S</geoCoordinate>
,
<geoCoordinate id="EE0E50328905FFB5FCCB3AF7FC52F934" box="[809,945,1756,1780]" direction="east" orientation="longitude" pageId="16" pageNumber="17" precision="1" value="48.98536">48.98536°E</geoCoordinate>
,
<quantity id="4CC29B108905FFB5FC203AF7FBFDF934" box="[962,1054,1756,1780]" metricMagnitude="3" metricUnit="m" metricValue="2.457" pageId="16" pageNumber="17" unit="m" value="2457.0">2457 m</quantity>
, on
<date id="FF8410358905FFB5FBBA3AF7FB18F934" box="[1112,1275,1756,1780]" pageId="16" pageNumber="17" value="2010-06-18">18 June 2010</date>
, by the same collectors as
<typeStatus id="548188578905FFB5FECB3B2BFE6CF8D8" box="[297,399,1792,1816]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C320657E8905FFB4FF253B0FFE36FBD0" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="8B8536F58905FFB4FF253B0FFB4FFF14" blockId="16.[151,1436,1576,2032]" lastBlockId="17.[151,1437,151,2013]" lastPageId="17" lastPageNumber="18" pageId="16" pageNumber="17">
<emphasis id="B94EEAE78905FFB5FF253B0FFDF5F8FD" bold="true" box="[199,534,1828,1853]" pageId="16" pageNumber="17">Diagnosis and comparisons.</emphasis>
Assigned to the genus
<taxonomicName id="4C3A4D768905FFB5FCC23B0FFC73F8FC" box="[800,912,1828,1852]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="genus">
<emphasis id="B94EEAE78905FFB5FCC23B0FFC73F8FC" box="[800,912,1828,1852]" italics="true" pageId="16" pageNumber="17">Platypelis</emphasis>
</taxonomicName>
in the
<taxonomicName id="4C3A4D768905FFB5FC033B0FFB81F8FC" box="[993,1122,1828,1852]" class="Amphibia" family="Microhylidae" kingdom="Animalia" order="Anura" pageId="16" pageNumber="17" phylum="Chordata" rank="subFamily" subFamily="Cophylinae">Cophylinae</taxonomicName>
based on enlarged terminal discs on fingers and toes, absence of finger-like prepollex, absence of nuptial pads, occurrence in
<collectingCountry id="F32D76658905FFB5FB3D3B63FA8BF8A0" box="[1247,1384,1864,1888]" name="Madagascar" pageId="16" pageNumber="17">Madagascar</collectingCountry>
, and molecular phylogenetic relationships. Distinguished from all other cophyline species with enlarged discs on fingers and toes by combination of the following character states: small body size (adult SVL
<quantity id="4CC29B108905FFB5FB8B3BBBFB04F868" box="[1129,1255,1936,1960]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="16" pageNumber="17" unit="mm" value="21.0" valueMax="22.0" valueMin="20.0">20–22 mm</quantity>
), ventral side of body and limbs with greenish or yellowish color especially on throat, dorsally without a conspicuous pattern of three symmetrical dark patches, no sharp border between dorsal and lateral color, toe 3 as long or slightly longer than toe 5, hindlimb reaching between forelimb and tympanum when adpressed along body, vomerine teeth absent, males with prepollical tubercle but lacking a finger-like prepollex as typical for
<taxonomicName id="4C3A4D768904FFB4FBF43C97FB46FF14" box="[1046,1189,188,212]" class="Amphibia" family="Microhylidae" genus="Anodonthyla" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B94EEAE78904FFB4FBF43C97FB46FF14" box="[1046,1189,188,212]" italics="true" pageId="17" pageNumber="18">Anodonthyla</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B8536F58904FFB4FF253CF4FADEFE47" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">
Distinguished from other species of
<taxonomicName id="4C3A4D768904FFB4FD853CCBFD34FF38" box="[615,727,224,248]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B94EEAE78904FFB4FD853CCBFD34FF38" box="[615,727,224,248]" italics="true" pageId="17" pageNumber="18">Platypelis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3A4D768904FFB4FCF33CCBFC93FF38" box="[785,880,224,248]" class="Amphibia" family="Microhylidae" genus="Cophyla" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B94EEAE78904FFB4FCF33CCBFC93FF38" box="[785,880,224,248]" italics="true" pageId="17" pageNumber="18">Cophyla</emphasis>
</taxonomicName>
as follows: from
<emphasis id="B94EEAE78904FFB4FBDF3CCBFC42FEDC" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="4C3A4D768904FFB4FBDF3CCBFB18FF38" box="[1085,1275,224,248]" class="Amphibia" family="Microhylidae" genus="Cophyla" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="phyllodactyla">C. phyllodactyla</taxonomicName>
,
<taxonomicName id="4C3A4D768904FFB4FAEA3CCBFA99FF38" box="[1288,1402,224,248]" class="Amphibia" family="Microhylidae" genus="Cophyla" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="berara">C. berara</taxonomicName>
,
<taxonomicName id="4C3A4D768904FFB4FA653CCAFF0CFEDC" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="alticola">P. alticola</taxonomicName>
,
<taxonomicName id="4C3A4D768904FFB4FF193D2EFE91FEDC" box="[251,370,261,284]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="cowanii">P. cowanii</taxonomicName>
,
<taxonomicName id="4C3A4D768904FFB4FE9C3D2EFE12FEDC" box="[382,497,260,284]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="grandis">P. grandis</taxonomicName>
,
<taxonomicName id="4C3A4D768904FFB4FE1F3D2EFD76FEDC" box="[509,661,261,284]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="mavomavo">P. mavomavo</taxonomicName>
,
<taxonomicName id="4C3A4D768904FFB4FD423D2EFCEBFEDC" box="[672,776,260,284]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="milloti">P. milloti</taxonomicName>
,
<taxonomicName id="4C3A4D768904FFB4FCF13D2EFC7EFEDC" box="[787,925,260,284]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="pollicaris">P. pollicaris</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C3A4D768904FFB4FC3B3D2EFB86FEDC" box="[985,1125,260,284]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="tuberifera">
<emphasis id="B94EEAE78904FFB4FC3B3D2EFB86FEDC" box="[985,1125,260,284]" italics="true" pageId="17" pageNumber="18">P. tuberifera</emphasis>
</taxonomicName>
by smaller body size (adult SVL
<quantity id="4CC29B108904FFB4FF303D0CFEA9FEFF" box="[210,330,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="17" pageNumber="18" unit="mm" value="21.0" valueMax="22.0" valueMin="20.0">20–22 mm</quantity>
vs.
<quantity id="4CC29B108904FFB4FE953D0CFDE2FEFF" box="[375,513,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="6.4" metricValueMax="10.5" metricValueMin="2.3" pageId="17" pageNumber="18" unit="mm" value="64.0" valueMax="105.0" valueMin="23.0">23–105 mm</quantity>
); and from all these species plus
<taxonomicName id="4C3A4D768904FFB4FC8C3D02FBAAFE80" box="[878,1097,297,320]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="tsaratananaensis">
<emphasis id="B94EEAE78904FFB4FC8C3D02FBAAFE80" box="[878,1097,297,320]" italics="true" pageId="17" pageNumber="18">P. tsaratananaensis</emphasis>
</taxonomicName>
and
<emphasis id="B94EEAE78904FFB4FB9F3D02FB2AFE80" box="[1149,1225,297,320]" italics="true" pageId="17" pageNumber="18">P. t e t r a</emphasis>
, by the absence or rudimentary state of vomerine teeth (vs. presence). Furthermore distinguished from all these species except
<taxonomicName id="4C3A4D768904FFB4FA653D66FEF3FE48" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="mavomavo">
<emphasis id="B94EEAE78904FFB4FA653D66FEF3FE48" italics="true" pageId="17" pageNumber="18">P. mavomavo</emphasis>
</taxonomicName>
and juvenile
<taxonomicName id="4C3A4D768904FFB4FE4A3D5AFDFAFE48" box="[424,537,368,392]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B94EEAE78904FFB4FE4A3D5AFDFAFE48" box="[424,537,368,392]" italics="true" pageId="17" pageNumber="18">P. grandis</emphasis>
</taxonomicName>
by the presence of green-yellow color on the ventral side (vs. absence).
</paragraph>
<paragraph id="8B8536F58904FFB4FF253DBFFBFFFD67" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">
Three other species are small-sized and lack vomerine teeth as
<taxonomicName id="4C3A4D768904FFB4FC603DBEFC39FE6C" box="[898,986,404,428]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="olgae">
<emphasis id="B94EEAE78904FFB4FC603DBEFC39FE6C" box="[898,986,404,428]" italics="true" pageId="17" pageNumber="18">P. olgae</emphasis>
</taxonomicName>
, i.e.,
<taxonomicName id="4C3A4D768904FFB4FBFA3DBEFB75FE6C" box="[1048,1174,404,428]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="barbouri">
<emphasis id="B94EEAE78904FFB4FBFA3DBEFB75FE6C" box="[1048,1174,404,428]" italics="true" pageId="17" pageNumber="18">P. barbouri</emphasis>
</taxonomicName>
,
<emphasis id="B94EEAE78904FFB4FB413DBEFB1BFE6C" box="[1187,1272,405,428]" italics="true" pageId="17" pageNumber="18">P. r a v u s</emphasis>
, and probably
<taxonomicName id="4C3A4D768904FFB4FF753D93FEC7FE10" box="[151,292,440,464]" class="Amphibia" family="Microhylidae" genus="Cophyla" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="occultans">
<emphasis id="B94EEAE78904FFB4FF753D93FEC7FE10" box="[151,292,440,464]" italics="true" pageId="17" pageNumber="18">C. occultans</emphasis>
</taxonomicName>
(state of vomerine teeth requires confirmation). The new species differs from
<taxonomicName id="4C3A4D768904FFB4FB443D92FACBFE10" box="[1190,1320,440,464]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="barbouri">
<emphasis id="B94EEAE78904FFB4FB443D92FACBFE10" box="[1190,1320,440,464]" italics="true" pageId="17" pageNumber="18">P. barbouri</emphasis>
</taxonomicName>
by greenyellow color on the ventral side (vs. reddish), from
<emphasis id="B94EEAE78904FFB4FD393DF6FCD2FE34" box="[731,817,477,500]" italics="true" pageId="17" pageNumber="18">P. r a v u s</emphasis>
by absence of a dorsal pattern consisting of three dark symmetrical patches (vs. presence), and from
<taxonomicName id="4C3A4D768904FFB4FD483E2AFCD0FDD8" box="[682,819,512,536]" class="Amphibia" family="Microhylidae" genus="Cophyla" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="occultans">
<emphasis id="B94EEAE78904FFB4FD483E2AFCD0FDD8" box="[682,819,512,536]" italics="true" pageId="17" pageNumber="18">P. occultans</emphasis>
</taxonomicName>
by a green-yellow venter (vs. uniform grey). Further distinguished from these three species as well as all other microhylids by molecular relationships and genetic divergence: the morphologically similar small-sized species
<taxonomicName id="4C3A4D768904FFB4FCA33E62FC22FDA0" box="[833,961,584,608]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="barbouri">
<emphasis id="B94EEAE78904FFB4FCA33E62FC22FDA0" box="[833,961,584,608]" italics="true" pageId="17" pageNumber="18">P. barbouri</emphasis>
</taxonomicName>
and
<emphasis id="B94EEAE78904FFB4FC1A3E62FBB3FDA0" box="[1016,1104,585,608]" italics="true" pageId="17" pageNumber="18">P. r av u s</emphasis>
are placed with high support into different clades, not closely related to
<taxonomicName id="4C3A4D768904FFB4FD663E46FD03FD44" box="[644,736,620,644]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="olgae">
<emphasis id="B94EEAE78904FFB4FD663E46FD03FD44" box="[644,736,620,644]" italics="true" pageId="17" pageNumber="18">P. olgae</emphasis>
</taxonomicName>
(
<figureCitation id="13012A708904FFB4FD103E47FCDAFD44" box="[754,825,620,644]" captionStart="FIGURE 7. 50" captionStartId="18.[151,250,1903,1926]" captionTargetBox="[234,1345,481,1864]" captionTargetId="figure@18.[234,1347,481,1864]" captionTargetPageId="18" captionText="FIGURE 7. 50 % majority-rule consensus tree from a Bayesian analysis of partial DNA sequences of the 16 S rRNA gene of Platypelis. Values on nodes are posterior probabilities (not shown if &lt;0.95). Cophyla berara was used as the outgroup (not shown)." httpUri="https://zenodo.org/record/215403/files/figure.png" pageId="17" pageNumber="18">Fig. 7</figureCitation>
), and the uncorrected pairwise divergence in the 16S fragment studied herein is&gt;10% to all other cophylines included in the analysis.
</paragraph>
<paragraph id="8B8536F58904FFB4FF253E98FE36FBD0" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">
<emphasis id="B94EEAE78904FFB4FF253E98FE04FD0C" bold="true" box="[199,487,691,716]" pageId="17" pageNumber="18">Available earlier names.</emphasis>
Few junior synonyms of nominal species of
<taxonomicName id="4C3A4D768904FFB4FC033E9FFBB2FD0C" box="[993,1105,692,716]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B94EEAE78904FFB4FC033E9FFBB2FD0C" box="[993,1105,692,716]" italics="true" pageId="17" pageNumber="18">Platypelis</emphasis>
</taxonomicName>
or
<taxonomicName id="4C3A4D768904FFB4FB943E9FFB36FD0C" box="[1142,1237,692,716]" class="Amphibia" family="Microhylidae" genus="Cophyla" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="genus">
<emphasis id="B94EEAE78904FFB4FB943E9FFB36FD0C" box="[1142,1237,692,716]" italics="true" pageId="17" pageNumber="18">Cophyla</emphasis>
</taxonomicName>
exist that need to be considered as possible earlier names for
<taxonomicName id="4C3A4D768904FFB4FD983EF2FB98FD2F" authority="Blommers-Schlosser &amp; Blanc 1991" authorityName="Blommers-Schlosser &amp; Blanc" authorityYear="1991" box="[634,1147,727,752]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="olgae">
<emphasis id="B94EEAE78904FFB4FD983EF2FD30FD30" box="[634,723,728,752]" italics="true" pageId="17" pageNumber="18">P. olgae</emphasis>
(
<bibRefCitation id="EFAB4B048904FFB4FD033EFCFB91FD2F" author="Blommers-Schlosser" box="[737,1138,727,751]" pageId="17" pageNumber="18" refString="Blommers-Schlosser, R. M. A. &amp; Blanc, C. P. (1991) Amphibiens (premiere partie). Faune de Madagascar, 75, 1 - 379." type="journal article" year="1991">Blommers-Schlösser &amp; Blanc 1991</bibRefCitation>
)
</taxonomicName>
.
<taxonomicName id="4C3A4D768904FFB4FB673EF3FEABFCD4" authority="Millot &amp; Guibe" authorityName="Millot &amp; Guibe" class="Amphibia" family="Microhylidae" genus="Paracophyla" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="tuberculata">
<emphasis id="B94EEAE78904FFB4FB673EF3FA7FFD30" box="[1157,1436,728,752]" italics="true" pageId="17" pageNumber="18">Paracophyla tuberculata</emphasis>
Millot &amp; Guibé
</taxonomicName>
is morphologically similar according to the original description (small size, absence of vomerine teeth) but has a longer hindlimb (tibiotarsal articulation reaching the eye), and due to its
<typeStatus id="548188578904FFB4FB963F0AFB47FCF9" box="[1140,1188,801,825]" pageId="17" pageNumber="18">type</typeStatus>
locality (Andasibe, at mid-altitude in the northern central east of
<collectingCountry id="F32D76658904FFB4FD9F3F6FFCE5FC9C" box="[637,774,836,860]" name="Madagascar" pageId="17" pageNumber="18">Madagascar</collectingCountry>
) its established synonymy with
<taxonomicName id="4C3A4D768904FFB4FB963F6FFAB3FC9C" box="[1140,1360,836,860]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="barbouri">
<emphasis id="B94EEAE78904FFB4FB963F6FFAB3FC9C" box="[1140,1360,836,860]" italics="true" pageId="17" pageNumber="18">Platypelis barbouri</emphasis>
</taxonomicName>
which is common at this site is very plausible. Other available names are
<taxonomicName id="4C3A4D768904FFB4FC5B3F43FAC2FCBF" authority="Mocquard" authorityName="Mocquard" box="[953,1313,871,896]" class="Amphibia" family="Microhylidae" genus="Platyhyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="verrucosa">
<emphasis id="B94EEAE78904FFB4FC5B3F43FB7CFC40" box="[953,1183,872,896]" italics="true" pageId="17" pageNumber="18">Platyhyla verrucosa</emphasis>
Mocquard
</taxonomicName>
,
<taxonomicName id="4C3A4D768904FFB4FAD33F43FE9BFC65" authority="Boettger" authorityName="Boettger" class="Amphibia" family="Microhylidae" genus="Platyhyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="voeltzkowi">
<emphasis id="B94EEAE78904FFB4FAD33F43FEEEFC64" italics="true" pageId="17" pageNumber="18">Platyhyla voeltzkowi</emphasis>
Boettger
</taxonomicName>
, and
<taxonomicName id="4C3A4D768904FFB4FE5B3FA7FD3BFC64" authority="Ahl" authorityName="Ahl" box="[441,728,908,932]" class="Amphibia" family="Microhylidae" genus="Cophyla" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="tuberculata">
<emphasis id="B94EEAE78904FFB4FE5B3FA7FD41FC64" box="[441,674,908,932]" italics="true" pageId="17" pageNumber="18">Cophyla tuberculata</emphasis>
Ahl
</taxonomicName>
, all junior synonyms of
<taxonomicName id="4C3A4D768904FFB4FC173FA7FB26FC64" box="[1013,1221,908,932]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B94EEAE78904FFB4FC173FA7FB26FC64" box="[1013,1221,908,932]" italics="true" pageId="17" pageNumber="18">Platypelis grandis</emphasis>
</taxonomicName>
; the name-bearing
<typeStatus id="548188578904FFB4FF753F9AFF31FC09" box="[151,210,945,969]" pageId="17" pageNumber="18">types</typeStatus>
of these nomina are large-sized and have strongly tuberculated dorsal skin (even the probably juvenile
<typeStatus id="548188578904FFB4FA833F9AFA7FFC09" box="[1377,1436,945,969]" pageId="17" pageNumber="18">types</typeStatus>
of
<taxonomicName id="4C3A4D768904FFB4FF5D3FFEFE88FC2C" box="[191,363,980,1004]" class="Amphibia" family="Microhylidae" genus="Cophyla" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="tuberculata">
<emphasis id="B94EEAE78904FFB4FF5D3FFEFE88FC2C" box="[191,363,980,1004]" italics="true" pageId="17" pageNumber="18">C. tuberculata</emphasis>
</taxonomicName>
measures
<quantity id="4CC29B108904FFB4FE143FFFFDACFC2C" box="[502,591,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="17" pageNumber="18" unit="mm" value="26.0">26 mm</quantity>
SVL;
<bibRefCitation id="EFAB4B048904FFB4FD4C3FFFFBBEFC2C" author="Blommers-Schlosser" box="[686,1117,980,1004]" pageId="17" pageNumber="18" refString="Blommers-Schlosser, R. M. A. &amp; Blanc, C. P. (1991) Amphibiens (premiere partie). Faune de Madagascar, 75, 1 - 379." type="journal article" year="1991">Blommers-Schlösser &amp; Blanc 1991</bibRefCitation>
), excluding their possible conspecificity with
<emphasis id="B94EEAE78904FFB4FE963FD2FE36FBD0" box="[372,469,1016,1040]" italics="true" pageId="17" pageNumber="18">
<taxonomicName id="4C3A4D768904FFB4FE963FD2FE32FBD0" box="[372,465,1016,1040]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="17" pageNumber="18" phylum="Chordata" rank="species" species="olgae">P. olgae</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C320657E8904FFB7FF253830FE35FEDC" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="8B8536F58904FFB4FF253830FBCCF990" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">
<emphasis id="B94EEAE78904FFB4FF253830FDF3FBF4" bold="true" box="[199,528,1051,1076]" pageId="17" pageNumber="18">
Description of the
<typeStatus id="548188578904FFB4FE403837FDE8FBF4" box="[418,523,1052,1076]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male in good state of preservation, some muscle tissue removed from right thigh. Snout-vent length
<quantity id="4CC29B108904FFB4FE493814FDF1FB97" box="[427,530,1087,1111]" metricMagnitude="-2" metricUnit="m" metricValue="2.03" pageId="17" pageNumber="18" unit="mm" value="20.3">20.3 mm</quantity>
. Body slender; head slightly longer than wide, not wider than body; snout bluntly rounded in dorsal and lateral views; nostrils directed almost laterally, not protuberant, nearer to tip of snout than to eye; canthus rostralis indistinct, very slightly concave; loreal region plain; tympanum moderately distinct, 69% of eye diameter; supratympanic fold moderately distinct, almost straight; tongue ovoid, not bifid or notched; weakly expressed maxillary teeth present; vomerine teeth not visible; choanae rounded. Forelimbs slender; subarticular tubercles on all fingers single, flat, but relatively poorly recognizable; outer metacarpal tubercle relatively large and flat, but poorly distinct; inner metacarpal tubercle large, forming distinct protuberance at base of first finger; hand with traces of webbing only recognizable between fingers 3 and 4; fingers distinctly flattened and broad along entire length; relative length of fingers 1&lt;2&lt;4&lt;3, fourth finger slightly longer than second; finger discs distinctly enlarged, slightly triangular; nuptial pads absent. Hindlimbs slender; tibiotarsal articulation reaching between forelimb and tympanum when hindlimb adpressed along body; tibia length, 41% of SVL; lateral metatarsalia strongly connected; inner metatarsal tubercle small and flat, difficult to recognize; outer metatarsal tubercle absent; webbing between toes very weakly developed, with traces of webbing between third and fourth toe; subarticular tubercles on toes hardly recognizable; toes flattened and relatively broad along their entire length; relative length of toes 1&lt;2&lt;5&lt;3&lt;4; third toe very slightly (left) to distinctly (right) longer than fifth.
</paragraph>
<paragraph id="8B8536F58904FFB4FF253A77FEBFF957" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">Dorsal skin smooth, without dorsolateral folds. Ventral skin smooth on throat and chest and moderately granular on belly.</paragraph>
<paragraph id="8B8536F58904FFB4FF253A8FFF3EF88C" blockId="17.[151,1437,151,2013]" pageId="17" pageNumber="18">After two years in 70% ethanol, dorsum almost uniformly brown with distinctly lighter brown on anterior head (anterior to eyes). Forelimbs and hindlimbs dorsally brown with moderately distinct white dots, becoming more evident on hands and feet. No recognizable dark crossbands on arms and limbs. Ventrally, marbled brown and white on throat, brown and yellowish on chest, dark brown and whitish on belly, and brown and yellowish on limbs.</paragraph>
<paragraph id="8B8536F58904FFB7FF253B7CFE35FEDC" blockId="17.[151,1437,151,2013]" lastBlockId="18.[151,1436,151,428]" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18">
<emphasis id="B94EEAE78904FFB4FF253B7CFED8F8B0" bold="true" box="[199,315,1879,1904]" pageId="17" pageNumber="18">Variation.</emphasis>
The two known males (
<typeStatus id="548188578904FFB4FD843B7CFD29F8AF" box="[614,714,1879,1903]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
and
<typeStatus id="548188578904FFB4FCE83B7CFC8FF8AF" box="[778,876,1879,1903]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
) have small tubercles scattered over the dorsal surface, while the female was more smooth. Both
<typeStatus id="548188578904FFB4FD2D3B56FCD9F855" box="[719,826,1917,1941]" pageId="17" pageNumber="18" type="paratype">paratypes</typeStatus>
had a yellow ventral ground color and less amount of dark pattern as compared to the
<typeStatus id="548188578904FFB4FDE73B8BFD88F878" box="[517,619,1952,1976]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
. The female
<typeStatus id="548188578904FFB4FD1D3B8AFCBCF879" box="[767,863,1953,1977]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
(ZSM 1667/2010) showed no differentiation from the
<typeStatus id="548188578904FFB4FF233BEFFEC0F81C" box="[193,291,1988,2012]" pageId="17" pageNumber="18" type="holotype">holotype</typeStatus>
in the mitochondrial DNA fragment studied (
<figureCitation id="13012A708904FFB4FCC83BEFFC93F81C" box="[810,880,1988,2012]" captionStart="FIGURE 7. 50" captionStartId="18.[151,250,1903,1926]" captionTargetBox="[234,1345,481,1864]" captionTargetId="figure@18.[234,1347,481,1864]" captionTargetPageId="18" captionText="FIGURE 7. 50 % majority-rule consensus tree from a Bayesian analysis of partial DNA sequences of the 16 S rRNA gene of Platypelis. Values on nodes are posterior probabilities (not shown if &lt;0.95). Cophyla berara was used as the outgroup (not shown)." httpUri="https://zenodo.org/record/215403/files/figure.png" pageId="17" pageNumber="18">Fig. 7</figureCitation>
; male
<typeStatus id="548188578904FFB4FC5E3BEEFBFFF81D" box="[956,1052,1989,2013]" pageId="17" pageNumber="18" type="paratype">paratype</typeStatus>
not sequenced). In all specimens, the third toe is as long as or slightly longer than toe five. Finger three is as long as finger four in all three specimens. In none of the specimens are vomerine teeth visible, but under the oral mucosa posteromedial bony ridges can be noticed when carefully probing with a needle, bearing denticulations that possibly could represent rudimentary vomerine teeth.
</paragraph>
</subSubSection>
<subSubSection id="C320657E8907FFB6FF253D03FF0CFCEC" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="etymology">
<paragraph id="8B8536F58907FFB7FF253D03FEA2FE6C" blockId="18.[151,1436,151,428]" pageId="18" pageNumber="19">
<emphasis id="B94EEAE78907FFB7FF253D03FEADFE80" bold="true" box="[199,334,296,320]" pageId="18" pageNumber="19">Etymology.</emphasis>
This species is dedicated to the late Prof. Olga Ramilijaona, former head of the zoology department at UADBA, aunt and mentor of the first author (AR), and friend and collaborator of MV, FG, NR, and DRV. The name is a patronym in recognition of Olga's tireless and successful efforts to develop zoological science in
<collectingCountry id="F32D76658907FFB7FF513DBFFEDEFE6C" box="[179,317,404,428]" name="Madagascar" pageId="18" pageNumber="19">Madagascar</collectingCountry>
.
</paragraph>
<caption id="DF45667D8907FFB7FF753B44FF0AF803" httpUri="https://zenodo.org/record/215403/files/figure.png" pageId="18" pageNumber="19" targetBox="[234,1345,481,1864]" targetPageId="18">
<paragraph id="8B8536F58907FFB7FF753B44FF0AF803" blockId="18.[151,1436,1903,1987]" pageId="18" pageNumber="19">
<emphasis id="B94EEAE78907FFB7FF753B44FEF7F845" bold="true" box="[151,276,1903,1926]" pageId="18" pageNumber="19">FIGURE 7.</emphasis>
50% majority-rule consensus tree from a Bayesian analysis of partial DNA sequences of the 16S rRNA gene of
<taxonomicName id="4C3A4D768907FFB7FF753BA5FF1FF864" box="[151,252,1934,1956]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="genus">
<emphasis id="B94EEAE78907FFB7FF753BA5FF1FF864" box="[151,252,1934,1956]" italics="true" pageId="18" pageNumber="19">Platypelis</emphasis>
</taxonomicName>
. Values on nodes are posterior probabilities (not shown if &lt;0.95).
<taxonomicName id="4C3A4D768907FFB7FC543BA5FBBAF864" box="[950,1113,1934,1956]" class="Amphibia" family="Microhylidae" genus="Cophyla" kingdom="Animalia" order="Anura" pageId="18" pageNumber="19" phylum="Chordata" rank="species" species="berara">
<emphasis id="B94EEAE78907FFB7FC543BA5FBBAF864" box="[950,1113,1934,1956]" italics="true" pageId="18" pageNumber="19">Cophyla berara</emphasis>
</taxonomicName>
was used as the outgroup (not shown).
</paragraph>
</caption>
<caption id="DF45667D8906FFB6FF753E67FB81FD41" httpUri="https://zenodo.org/record/215404/files/figure.png" pageId="19" pageNumber="20" targetBox="[151,1436,181,566]" targetPageId="19">
<paragraph id="8B8536F58906FFB6FF753E67FB81FD41" blockId="19.[151,1436,588,641]" pageId="19" pageNumber="20">
<emphasis id="B94EEAE78906FFB6FF753E67FEF1FDA1" bold="true" box="[151,274,588,611]" pageId="19" pageNumber="20">FIGURE 8.</emphasis>
Lateral and dorsal views of various embryonic developmental stages of
<taxonomicName id="4C3A4D768906FFB6FC0F3E67FAE5FDA2" box="[1005,1286,588,610]" class="Amphibia" family="Microhylidae" genus="Platypelis" kingdom="Animalia" order="Anura" pageId="19" pageNumber="20" phylum="Chordata" rank="species" species="tsaratananaensis">
<emphasis id="B94EEAE78906FFB6FC0F3E67FAE5FDA2" box="[1005,1286,588,610]" italics="true" pageId="19" pageNumber="20">Platypelis tsaratananaensis</emphasis>
</taxonomicName>
: (a–b) stage 8; (c–d) stage 11; (e–f) stage 12; (g–h) stage 13; (i–j) stage 15. Stages according to Gosner (1960).
</paragraph>
</caption>
<paragraph id="8B8536F58906FFB6FF253E82FF0CFCEC" blockId="19.[151,1436,680,812]" pageId="19" pageNumber="20">
<emphasis id="B94EEAE78906FFB6FF253E82FE64FD01" bold="true" box="[199,391,680,705]" pageId="19" pageNumber="20">Natural history.</emphasis>
The
<typeStatus id="548188578906FFB6FE263E83FDC5FD00" box="[452,550,680,704]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
was found at daytime on a small log that had been felled and where it might have emerged from a tiny water filled tree hole. The two
<typeStatus id="548188578906FFB6FCF43EE6FC62FD25" box="[790,897,717,741]" pageId="19" pageNumber="20" type="paratype">paratypes</typeStatus>
were found hidden among dense layers of moss and leaf litter on the ground. No further information is available on the natural history, reproduction, or calls of this species.
</paragraph>
</subSubSection>
</treatment>
</document>
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