173 lines
28 KiB
XML
173 lines
28 KiB
XML
<document id="9C949B910FD5C7844C05582B2CE094BF" ID-CLB-Dataset="8118" ID-DOI="10.5252/z2010n1a2" ID-GBIF-Dataset="4cecd899-5900-4411-8446-0d9552b1b098" ID-ISSN="1638-9387" ID-Zenodo-Dep="4520753" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1612808309234" checkinUser="felipe" docAuthor="Randriamanantsoa, Richard, Aberlenc, Henri-Pierre, Ralisoa, Olga Bakoly, Ratnadass, Alain & Vercambre, Bernard" docDate="2010" docId="901F8794FF857473FD52FC9CFBFEFD26" docLanguage="fr" docName="Zoosystema.32.1.19-72.pdf" docOrigin="Zoosystema 32 (1)" docSource="http://www.bioone.org/doi/abs/10.5252/z2010n1a2" docStyle="DocumentStyle:E638A3C99FA728B529662B46F84D75FD.3:Zoosystema.2009-2010.journal_article" docStyleId="E638A3C99FA728B529662B46F84D75FD" docStyleName="Zoosystema.2009-2010.journal_article" docStyleVersion="3" docTitle="Apicencya waterloti" docType="treatment" docVersion="11" lastPageNumber="37" masterDocId="6C26FFECFF947461FFB5FFEDFF94FFE8" masterDocTitle="Les larves des Scarabaeoidea (Insecta, Coleoptera) en riziculture pluviale des régions de haute et moyenne altitudes du Centre de Madagascar" masterLastPageNumber="72" masterPageNumber="19" pageNumber="36" updateTime="1698928337295" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="68A09F155B446233A5C21027E70B23E9">Les larves des Scarabaeoidea (Insecta, Coleoptera) en riziculture pluviale des régions de haute et moyenne altitudes du Centre de Madagascar</mods:title>
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<mods:namePart id="7B472BC47330956D0DFE38FBD05033C2">Randriamanantsoa, Richard</mods:namePart>
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<mods:affiliation id="24BADFB82D6784EDB3C66AE69180BDF0">Fofifa - URP Scrid, 110 Antsirabe (Madagascar) r _ randriamanantsoa @ yahoo. fr</mods:affiliation>
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<mods:namePart id="0934CD24E3A270E83A286D124AB05AAC">Aberlenc, Henri-Pierre</mods:namePart>
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<mods:affiliation id="3EFD1EFEA908B4D8DC6B7C2BEBEFFD9B">Cirad Bios, UMR CBGP (INRA / IRD / Cirad / Montpellier SupAgro), Campus international de Baillarguet, Ta A- 55 / L, F- 34398 Montpellier cedex 5 (France) henri-pierre. aberlenc @ cirad. fr</mods:affiliation>
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<mods:nameIdentifier id="97D2355EBEA73C8E03CC7B27F89363A2" type="email">aberlenc@cirad.fr</mods:nameIdentifier>
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<mods:namePart id="E17BD4C720E3A6748B6A2728C9D377CC">Ralisoa, Olga Bakoly</mods:namePart>
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<mods:affiliation id="1F1CBFB1028126C144C5C4545FE7B160">Département d’Entomologie, Faculté des Sciences, Université d’Antananarivo, Antananarivo 101 (Madagascar)</mods:affiliation>
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<mods:namePart id="DD552D52FB0551AAAEE48FDA3A201E14">Ratnadass, Alain</mods:namePart>
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<mods:affiliation id="652BB761ED0A0936F92884F1F6FB0CB8">Cirad, UPR HortSys - ICRISAT BP 12404, Niamey (Niger) a. ratnadass @ cgiar. org</mods:affiliation>
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<mods:nameIdentifier id="D9AC05CE18E86C9FE5A57C8193BFE73B" type="email">ratnadass@cgiar.org</mods:nameIdentifier>
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<mods:namePart id="0D0070ECC63D2E5B64219F7D3B2FCD1A">Vercambre, Bernard</mods:namePart>
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<mods:affiliation id="521457076DDAE850B7A38164428594B5">Cirad, UPR Systèmes de Cultures annuels, avenue Agropolis, F- 34398 Montpellier cedex 5 (France) bernard. vercambre @ cirad. fr</mods:affiliation>
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<mods:nameIdentifier id="B30470CEDB70B824846545ED3604941E" type="email">vercambre@cirad.fr</mods:nameIdentifier>
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<mods:title id="DD5F411FBDE5B18AA6C620157AB0BACD">Zoosystema</mods:title>
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<mods:date id="977C4AEC6961E338502763DED5D60A9D">2010</mods:date>
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<mods:number id="0E60F05150C6E2D861F674618ACCCD91">2010-03-31</mods:number>
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<mods:url id="6B4F1EFE32672AF8271DA248218CC2C9">http://www.bioone.org/doi/abs/10.5252/z2010n1a2</mods:url>
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<treatment id="901F8794FF857473FD52FC9CFBFEFD26" ID-DOI="http://doi.org/10.5281/zenodo.4549542" ID-GBIF-Taxon="177425640" ID-Zenodo-Dep="4549542" LSID="urn:lsid:plazi:treatment:901F8794FF857473FD52FC9CFBFEFD26" httpUri="http://treatment.plazi.org/id/901F8794FF857473FD52FC9CFBFEFD26" lastPageId="18" lastPageNumber="37" pageId="17" pageNumber="36">
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<subSubSection id="50AC6509FF857470FD52FC9CFCE1FBE1" pageId="17" pageNumber="36" type="nomenclature">
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<paragraph id="18093682FF857470FD52FC9CFBE3FC64" blockId="17.[743,1143,881,971]" box="[743,1143,881,908]" pageId="17" pageNumber="36">
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<taxonomicName id="DFB64D01FF857470FD52FC9CFBE3FC64" authority="(Dewailly, 1950)" baseAuthorityName="Dewailly" baseAuthorityYear="1950" box="[743,1143,881,908]" class="Insecta" family="Melolonthidae" genus="Apicencya" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="36" phylum="Arthropoda" rank="species" species="waterloti">
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<emphasis id="2AC2EA90FF857470FD52FC9CFC2EFC64" bold="true" box="[743,954,881,908]" italics="true" pageId="17" pageNumber="36">Apicencya waterloti</emphasis>
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(Dewailly, 1950)
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</taxonomicName>
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</paragraph>
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<paragraph id="18093682FF857470FD5EFC7CFBD0FC23" blockId="17.[743,1143,881,971]" pageId="17" pageNumber="36">
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(
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<figureCitation id="808D2A07FF857470FD47FC7CFCD7FC44" box="[754,835,913,940]" captionStart="FIG" captionStartId="29.[113,124,1567,1584]" captionTargetBox="[152,1172,222,1545]" captionTargetId="figure-18@29.[148,709,692,1169]" captionTargetPageId="29" captionText="FIG. 2. — Larves de Scarabaeoidea de Madagascar, habitus: A, Bricoptis variolosa (Gory & Percheron, 1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paramorphochelus cornutus (Nonfried,1892); K, Encya sikorai (Brenske,1891); L, Apicencya waterloti (Dewailly, 1950); M, Empecta scutata (Fairmaire,1901); N, Hoplochelus betanimena (Künckel,1887); O, Triodontus nitidulus (Guérin, 1844). Échelles: A-I, K, L, N, 5 mm; J, O, 2 mm; M, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4520757" httpUri="https://zenodo.org/record/4520757/files/figure.png" pageId="17" pageNumber="36">Figs 2L</figureCitation>
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;
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<figureCitation id="808D2A07FF857470FCE0FC7CFCE2FC44" box="[853,886,913,940]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="17" pageNumber="36">5C</figureCitation>
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;
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<figureCitation id="808D2A07FF857470FC3DFC7CFC33FC44" box="[904,935,913,940]" captionStart="FIG" captionStartId="36.[113,124,1635,1652]" captionTargetBox="[134,1187,219,1587]" captionTargetId="figure-21@36.[131,1191,216,1588]" captionTargetPageId="36" captionText="FIG. 9.— Larves de Scarabaeoidea de Madagascar,épipharynx: A, Apicencya waterloti (Dewailly,1950); B, Empecta scutata (Fairmaire, 1901); C, Hoplochelus betanimena (Künckel, 1887); D, sp. M3; E, Triodontus nitidulus (Guérin, 1844); F, sp. S1; G, sp. S2. Échelles: D, 0,5 mm; E-G, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520777" httpUri="https://zenodo.org/record/4520777/files/figure.png" pageId="17" pageNumber="36">9A</figureCitation>
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;
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<figureCitation id="808D2A07FF857470FC0CFC7CFB9EFC44" box="[953,1034,913,940]" captionStart="FIG" captionStartId="39.[113,124,1567,1584]" captionTargetBox="[118,1207,216,1486]" captionTargetId="figure-14@39.[118,1207,216,1486]" captionTargetPageId="39" captionText="FIG. 12. — Larves de Scarabaeoidea de Madagascar: A-E, hypopharynx; A, Enaria melanictera (Klug, 1833); B, Apicencya waterloti (Dewailly, 1950); C, Empecta scutata (Fairmaire, 1901); D, Hoplochelus betanimena (Künckel, 1887); E, Triodontus nitidulus (Guérin, 1844); F-O, antenne; F, Anochilia bifida (Olivier, 1789); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K-M, Encya sikorai (Brenske,1891); L, M, plages sensorielles de l’article apical; N, Apicencya waterloti (Dewailly, 1950); O, Empecta scutata (Fairmaire, 1901). Échelles: A, F, G, I, J, O, 0,5 mm; E, L, M, 0,2 mm; H, K, N, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520787" httpUri="https://zenodo.org/record/4520787/files/figure.png" pageId="17" pageNumber="36">12B, N</figureCitation>
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;
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<figureCitation id="808D2A07FF857470FBAEFC7CFBA1FC44" box="[1051,1077,913,940]" captionStart="FIG" captionStartId="42.[113,124,1589,1606]" captionTargetBox="[135,1177,220,1549]" captionTargetId="figure-21@42.[136,1178,214,1549]" captionTargetPageId="42" captionText="FIG. 15. — Larves de Scarabaeoidea de Madagascar, mandibule: A, E, I, M, Q, U, Y, gauche, face dorsale; B, F, J, N, R, V, Z1, gauche, face ventrale; C, G, K, O, S, W, Z2, droite,face dorsale; D, H, L, P, T, X, Z3, droite,face ventrale; A-D, Paranodon coquerelii (Fairmaire, 1871); E-H, Paramorphochelus cornutus (Nonfried, 1892); I-L, Encya sikorai (Brenske, 1891); M-P, Enaria melanictera (Klug, 1833); Q-T, Apicencya waterloti (Dewailly, 1950); U-X, Empecta scutata (Fairmaire, 1901); Y-Z3, Hoplochelus betanimena (Künckel, 1887). Échelles: A-D, M-P, 1 mm; E-H, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520799" httpUri="https://zenodo.org/record/4520799/files/figure.png" pageId="17" pageNumber="36">15</figureCitation>
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Q-T; 17E; 20C; 22O-Q; 24A-I)
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</paragraph>
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<paragraph id="18093682FF857470FD10FC1DFCE1FBE1" blockId="17.[674,1211,1008,1225]" box="[677,885,1008,1035]" pageId="17" pageNumber="36">MATÉRIEL EXAMINÉ</paragraph>
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</subSubSection>
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<subSubSection id="50AC6509FF857473FD10FBFDFBFEFD26" lastPageId="18" lastPageNumber="37" pageId="17" pageNumber="36" type="description">
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<paragraph id="18093682FF857470FD10FBFDFB08FB21" blockId="17.[674,1211,1008,1225]" pageId="17" pageNumber="36">
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Larves provenant d’Andranomanelatra, dans une culture de Riz pluvial située en haut de pente. Elles coupaient les racines des plants, avec une destruction à plus de 90 %.
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<typeStatus id="C70D8820FF857470FCF7FB82FCE2FB62" box="[834,886,1135,1162]" pageId="17" pageNumber="36">Type</typeStatus>
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de sol: ferrallitique. Spécimens conservés dans les collections du Fofifa (14 larves), du Cirad-CBGP (
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<specimenCount id="0EB0FD0BFF857470FCF6FB43FC1FFB21" box="[835,907,1198,1225]" pageId="17" pageNumber="36" type="larva">1 larve</specimenCount>
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) et du MNHN (
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<specimenCount id="0EB0FD0BFF857470FBF3FB43FB05FB21" box="[1094,1169,1198,1225]" pageId="17" pageNumber="36" type="larva">1 larve</specimenCount>
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).
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</paragraph>
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<paragraph id="18093682FF857470FD10FB00FCA1FAEE" blockId="17.[677,966,1261,1320]" box="[677,821,1261,1288]" pageId="17" pageNumber="36">DESCRIPTION</paragraph>
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<paragraph id="18093682FF857470FD10FAE0FC52FAC0" blockId="17.[677,966,1261,1320]" box="[677,966,1293,1320]" pageId="17" pageNumber="36">
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Couleur blanche (
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<figureCitation id="808D2A07FF857470FCD2FAE0FC2DFAC0" box="[871,953,1293,1320]" captionStart="FIG" captionStartId="29.[113,124,1567,1584]" captionTargetBox="[152,1172,222,1545]" captionTargetId="figure-18@29.[148,709,692,1169]" captionTargetPageId="29" captionText="FIG. 2. — Larves de Scarabaeoidea de Madagascar, habitus: A, Bricoptis variolosa (Gory & Percheron, 1833); B, Celidota parvula (Janson, 1881); C, Anochilia bifida (Olivier, 1789); D, Euryomia argentea (Olivier, 1789); E, Heteronychus arator rugifrons (Fairmaire, 1871); F, Heteronychus bituberculatus (Kolbe, 1900); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paramorphochelus cornutus (Nonfried,1892); K, Encya sikorai (Brenske,1891); L, Apicencya waterloti (Dewailly, 1950); M, Empecta scutata (Fairmaire,1901); N, Hoplochelus betanimena (Künckel,1887); O, Triodontus nitidulus (Guérin, 1844). Échelles: A-I, K, L, N, 5 mm; J, O, 2 mm; M, 3 mm." figureDoi="http://doi.org/10.5281/zenodo.4520757" httpUri="https://zenodo.org/record/4520757/files/figure.png" pageId="17" pageNumber="36">Fig. 2L</figureCitation>
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).
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</paragraph>
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<paragraph id="18093682FF857470FD10FAA0FD5BFA8F" blockId="17.[675,1212,1357,1605]" box="[677,719,1357,1383]" pageId="17" pageNumber="36">
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<emphasis id="2AC2EA90FF857470FD10FAA0FD5BFA8F" box="[677,719,1357,1383]" italics="true" pageId="17" pageNumber="36">Tête</emphasis>
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</paragraph>
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<paragraph id="18093682FF857470FD10FA81FCBAF9AD" blockId="17.[675,1212,1357,1605]" pageId="17" pageNumber="36">Couleur jaunâtre. Épicrâne avec symétriquement 7 à 9 soies latéro-externes; 2 soies dorso-épicraniales. Front avec 9 soies frontales antérieures, 2 soies frontales externes et 4 soies frontales postérieures, disposées symétriquement. Clypéus avec 1 soie clypéale externe et 2 soies clypéales antérieures symétriques.</paragraph>
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<paragraph id="18093682FF857473FD10F984FE93FE79" blockId="17.[677,1211,1641,1700]" lastBlockId="18.[111,648,216,401]" lastPageId="18" lastPageNumber="37" pageId="17" pageNumber="36">
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<emphasis id="2AC2EA90FF857470FD10F984FC9BF96B" bold="true" box="[677,783,1641,1667]" pageId="17" pageNumber="36">Antennes</emphasis>
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(
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<figureCitation id="808D2A07FF857470FCAAF984FC1DF96C" box="[799,905,1641,1668]" captionStart="FIG" captionStartId="39.[113,124,1567,1584]" captionTargetBox="[118,1207,216,1486]" captionTargetId="figure-14@39.[118,1207,216,1486]" captionTargetPageId="39" captionText="FIG. 12. — Larves de Scarabaeoidea de Madagascar: A-E, hypopharynx; A, Enaria melanictera (Klug, 1833); B, Apicencya waterloti (Dewailly, 1950); C, Empecta scutata (Fairmaire, 1901); D, Hoplochelus betanimena (Künckel, 1887); E, Triodontus nitidulus (Guérin, 1844); F-O, antenne; F, Anochilia bifida (Olivier, 1789); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K-M, Encya sikorai (Brenske,1891); L, M, plages sensorielles de l’article apical; N, Apicencya waterloti (Dewailly, 1950); O, Empecta scutata (Fairmaire, 1901). Échelles: A, F, G, I, J, O, 0,5 mm; E, L, M, 0,2 mm; H, K, N, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520787" httpUri="https://zenodo.org/record/4520787/files/figure.png" pageId="17" pageNumber="36">Fig. 12N</figureCitation>
|
||
). Les trois premiers articles subégaux, le quatrième plus court (longueurs respectives des articles: I = III = 1,93; II = 2; IV = 1). Dernier article antennaire avec 4 plages sensorielles: 2 ventrales, 1 petite aire apicale ronde et 1 dorsale, l’article I portant 4 soies et l’article II portant 2 soies. Avant-dernier article avec une saillie ventrale apico-latérale.
|
||
</paragraph>
|
||
<paragraph id="18093682FF867473FFC4FE5BFE9BFD06" blockId="18.[111,649,438,750]" pageId="18" pageNumber="37">
|
||
<emphasis id="2AC2EA90FF867473FFC4FE5BFF2CFE38" bold="true" box="[113,184,438,464]" pageId="18" pageNumber="37">Labre.</emphasis>
|
||
Un peu plus large que long, symétrique. Zone médiane du bord antérieur en pointe arrondie, avec 2 + 2 soies externes et quatre soies internes rapprochées à l’intérieur d’une zone pigmentée subcirculaire. 2 soies latéro-antérieures externes et 2 + 1 soies latéro-antérieures internes, 1 soie latérale et 5 à 7 courtes soies latéro-postérieures. Soies de l’acanthoparia bien visibles latéralement, de l’aire latéro-antérieure à l’aire latéro-postérieure. 2 soies labro-internes.
|
||
</paragraph>
|
||
<paragraph id="18093682FF867473FFC4FCFFFDEAFC03" blockId="18.[111,647,786,1003]" pageId="18" pageNumber="37">
|
||
<emphasis id="2AC2EA90FF867473FFC4FCFFFF60FCC5" bold="true" box="[113,244,786,813]" pageId="18" pageNumber="37">Épypharynx</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF867473FEB1FCFFFECDFCC5" box="[260,345,786,813]" captionStart="FIG" captionStartId="36.[113,124,1635,1652]" captionTargetBox="[134,1187,219,1587]" captionTargetId="figure-21@36.[131,1191,216,1588]" captionTargetPageId="36" captionText="FIG. 9.— Larves de Scarabaeoidea de Madagascar,épipharynx: A, Apicencya waterloti (Dewailly,1950); B, Empecta scutata (Fairmaire, 1901); C, Hoplochelus betanimena (Künckel, 1887); D, sp. M3; E, Triodontus nitidulus (Guérin, 1844); F, sp. S1; G, sp. S2. Échelles: D, 0,5 mm; E-G, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520777" httpUri="https://zenodo.org/record/4520777/files/figure.png" pageId="18" pageNumber="37">Fig. 9A</figureCitation>
|
||
). Haptomère constitué d’un groupe de très fortes épines. Proplegmatum réduit, formé de 12 à 15 bourrelets parallèles. Plegmatum développé. Acanthoparia avec 13 à 16 courtes soies lamelliformes incurvées vers l’avant, de taille croissant d’arrière en avant.Chaetoparia dissymétriques,à fortes soies convergeant vers l’aire pédiale, qui est glabre.
|
||
</paragraph>
|
||
<paragraph id="18093682FF867473FFC4FBE2FD9BFAE0" blockId="18.[112,648,1039,1288]" pageId="18" pageNumber="37">
|
||
<emphasis id="2AC2EA90FF867473FFC4FBE2FF60FBC1" bold="true" box="[113,244,1039,1065]" pageId="18" pageNumber="37">Mandibules</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF867473FEB7FBFDFED9FBC3" box="[258,333,1040,1067]" captionStart="FIG" captionStartId="42.[113,124,1589,1606]" captionTargetBox="[135,1177,220,1549]" captionTargetId="figure-21@42.[136,1178,214,1549]" captionTargetPageId="42" captionText="FIG. 15. — Larves de Scarabaeoidea de Madagascar, mandibule: A, E, I, M, Q, U, Y, gauche, face dorsale; B, F, J, N, R, V, Z1, gauche, face ventrale; C, G, K, O, S, W, Z2, droite,face dorsale; D, H, L, P, T, X, Z3, droite,face ventrale; A-D, Paranodon coquerelii (Fairmaire, 1871); E-H, Paramorphochelus cornutus (Nonfried, 1892); I-L, Encya sikorai (Brenske, 1891); M-P, Enaria melanictera (Klug, 1833); Q-T, Apicencya waterloti (Dewailly, 1950); U-X, Empecta scutata (Fairmaire, 1901); Y-Z3, Hoplochelus betanimena (Künckel, 1887). Échelles: A-D, M-P, 1 mm; E-H, 0,5 mm." figureDoi="http://doi.org/10.5281/zenodo.4520799" httpUri="https://zenodo.org/record/4520799/files/figure.png" pageId="18" pageNumber="37">Fig. 15</figureCitation>
|
||
Q-T). Pas d’aire stridulatoire. Aire scissoriale de la mandibule gauche allongée, avec une forte dent apicale émoussée, la deuxième dent arrondie et la troisième dent fine peu saillante. Aire scissoriale de la mandibule droite allongée, avec une forte dent apicale émoussée, une seconde dent effacée et une troisième dent arrondie. 8 soies latéro-externes et 1 soie dorso-externe.
|
||
</paragraph>
|
||
<paragraph id="18093682FF867473FFC4FAC1FDCAF9EE" blockId="18.[112,649,1324,1542]" pageId="18" pageNumber="37">
|
||
<emphasis id="2AC2EA90FF867473FFC4FAC1FF5AFAAE" bold="true" box="[113,206,1324,1350]" pageId="18" pageNumber="37">Maxilles</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF867473FF6BFAC0FED5FAA0" box="[222,321,1325,1352]" captionStart="FIG" captionStartId="44.[113,124,1612,1629]" captionTargetBox="[117,1206,214,1559]" captionTargetId="figure-21@44.[117,1206,214,1559]" captionTargetPageId="44" captionText="FIG. 17. — Larves de Scarabaeoidea de Madagascar, maxille, face dorsale: A, Paranodon coquerelii (Fairmaire, 1871); B, Paramorphochelus cornutus (Nonfried, 1892); C, Encya sikorai (Brenske, 1891); D, Enaria melanictera (Klug, 1833); E, Apicencya waterloti (Dewailly, 1950); F, Empecta scutata (Fairmaire, 1901); G, Hoplochelus betanimena (Künckel, 1887); H, Triodontus nitidulus (Guérin, 1844). Échelles: A, 0,5 mm; B, 0,1 mm; D, H, 0,2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520807" httpUri="https://zenodo.org/record/4520807/files/figure.png" pageId="18" pageNumber="37">Fig. 17E</figureCitation>
|
||
). Galea et lacinia soudées à la base, séparées à l’apex. Lacinia à uncus avec 3 dents sclérifiées développées, entourées par une rangée de 4 + 2 épines de chaque côté. Galea avec uncus sclérifié, armée de 6 + 2 fortes épines sclérifiées. Sur la face dorsale, aire stridulatoire composée de 16 à 17 dents séparées en forme de cônes sclérifiés.
|
||
</paragraph>
|
||
<paragraph id="18093682FF867473FFC4F9C7FE25F94C" blockId="18.[112,647,1578,1700]" pageId="18" pageNumber="37">
|
||
<emphasis id="2AC2EA90FF867473FFC4F9C7FE92F9AC" bold="true" box="[113,262,1578,1604]" pageId="18" pageNumber="37">Hypopharynx</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF867473FEA7F9C7FEFAF9AD" box="[274,366,1578,1605]" captionStart="FIG" captionStartId="39.[113,124,1567,1584]" captionTargetBox="[118,1207,216,1486]" captionTargetId="figure-14@39.[118,1207,216,1486]" captionTargetPageId="39" captionText="FIG. 12. — Larves de Scarabaeoidea de Madagascar: A-E, hypopharynx; A, Enaria melanictera (Klug, 1833); B, Apicencya waterloti (Dewailly, 1950); C, Empecta scutata (Fairmaire, 1901); D, Hoplochelus betanimena (Künckel, 1887); E, Triodontus nitidulus (Guérin, 1844); F-O, antenne; F, Anochilia bifida (Olivier, 1789); G, Heteronychus plebeius (Klug, 1833); H, Hexodon unicolor (Olivier, 1789); I, Heteroconus paradoxus (Endrödi,1968); J, Paranodon coquerelii (Fairmaire,1871); K-M, Encya sikorai (Brenske,1891); L, M, plages sensorielles de l’article apical; N, Apicencya waterloti (Dewailly, 1950); O, Empecta scutata (Fairmaire, 1901). Échelles: A, F, G, I, J, O, 0,5 mm; E, L, M, 0,2 mm; H, K, N, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520787" httpUri="https://zenodo.org/record/4520787/files/figure.png" pageId="18" pageNumber="37">Fig. 12B</figureCitation>
|
||
). Glosses densément recouvertes de longues soies à l’apex et de très fortes soies moins denses orientées vers l’arrière et formant une zone subtriangulaire à la base.
|
||
</paragraph>
|
||
<paragraph id="18093682FF867473FD10FF35FD79FF1A" blockId="18.[675,1212,216,465]" box="[677,749,216,242]" pageId="18" pageNumber="37">
|
||
<emphasis id="2AC2EA90FF867473FD10FF35FD79FF1A" box="[677,749,216,242]" italics="true" pageId="18" pageNumber="37">Thorax</emphasis>
|
||
</paragraph>
|
||
<paragraph id="18093682FF867473FD10FF15FBB2FE39" blockId="18.[675,1212,216,465]" pageId="18" pageNumber="37">
|
||
<emphasis id="2AC2EA90FF867473FD10FF15FD73FEFA" bold="true" box="[677,743,248,274]" pageId="18" pageNumber="37">Pattes</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF867473FD46FF15FCC1FEFB" box="[755,853,248,275]" captionStart="FIG" captionStartId="47.[113,124,1635,1652]" captionTargetBox="[136,1188,217,1589]" captionTargetId="figure-14@47.[136,1188,217,1589]" captionTargetPageId="47" captionText="FIG. 20. — Larves de Scarabaeoidea de Madagascar, pattes: A, Encya sikorai (Brenske, 1891); B, Enaria melanictera (Klug, 1833); C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel,1887); F, Triodontus nitidulus (Guérin, 1844). Échelles: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4520817" httpUri="https://zenodo.org/record/4520817/files/figure.png" pageId="18" pageNumber="37">Fig. 20C</figureCitation>
|
||
). Longues, de longueur croissante de P1 à P3, toutes de structure analogue: coxa, trochanter inséré latéralement sur la coxa, fémur, tibiotarse. P1 et P2 à griffe très longue, très forte à la base, incurvée, sclérifiée à l’apex. P3 à griffe courte, à peine plus longue que large, à apex pointu sclérifié, portant deux longues soies.
|
||
</paragraph>
|
||
<paragraph id="18093682FF867473FD10FE18FC93FDE7" blockId="18.[674,1210,501,718]" box="[677,775,501,527]" pageId="18" pageNumber="37">
|
||
<emphasis id="2AC2EA90FF867473FD10FE18FC93FDE7" box="[677,775,501,527]" italics="true" pageId="18" pageNumber="37">Abdomen</emphasis>
|
||
</paragraph>
|
||
<paragraph id="18093682FF867473FD10FDF8FBFEFD26" blockId="18.[674,1210,501,718]" pageId="18" pageNumber="37">
|
||
<emphasis id="2AC2EA90FF867473FD10FDF8FD7CFDC7" bold="true" box="[677,744,533,559]" pageId="18" pageNumber="37">Raster</emphasis>
|
||
(
|
||
<figureCitation id="808D2A07FF867473FD47FDF8FCD5FDD8" box="[754,833,533,560]" captionStart="FIG" captionStartId="32.[113,124,1635,1652]" captionTargetBox="[118,1198,220,1565]" captionTargetId="figure-21@32.[118,1207,220,1565]" captionTargetPageId="32" captionText="FIG. 5. — Larves de Scarabaeoidea de Madagascar, raster: A, Enaria melanictera (Klug, 1833); B, sp. M2; C, Apicencya waterloti (Dewailly,1950); D, Empecta scutata (Fairmaire,1901); E, Hoplochelus betanimena (Künckel, 1887); F, sp.M3; G, Triodontus nitidulus (Guérin, 1844); H, sp. S1; I, sp. S2. Échelles: A, B, F-I, 0,5 mm; C, D, 1 mm; E, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4520765" httpUri="https://zenodo.org/record/4520765/files/figure.png" pageId="18" pageNumber="37">Fig.5C</figureCitation>
|
||
).Fente anale en «Y». Palidia en forme de fuseau,formés de deux rangées symétriques de 13-15 fortes soies orientées vers l’intérieur. Septula glabre. Teges formés de soies concentriques,denses,épaisses et longues à l’intérieur,plus courtes et éparses à l’extérieur. Barbula avec de nombreuses soies allongées.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |