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<document id="FF683B81D770E3F920C707A5A162EF9B" ID-DOI="10.5281/zenodo.175869" ID-GBIF-Dataset="ef14a2a6-57bf-4a25-94ff-5a94baeec224" ID-ISSN="1175-5326" ID-Zenodo-Dep="175869" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459822160839" checkinUser="plazi" docAuthor="Bartsch, Ilse" docDate="2007" docId="03FB87B1CE29B634088BFCFCB413A6F2" docLanguage="en" docName="zt01435p049.pdf" docOrigin="Zootaxa 1435" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Copidognathus angusticeps Bartsch, 2007, spec. nov." docType="treatment" docVersion="8" lastPageNumber="45" masterDocId="FFC2FFC9CE28B630081CFFABB54EA508" masterDocTitle="Halacarid mites (Acari: Halacaridae) from Esperance (Western Australia), description of two new Copidognathus species" masterLastPageNumber="49" masterPageNumber="41" pageNumber="42" updateTime="1698221939282" updateUser="plazi">
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<mods:title id="A280DD45A1AF4802FA9DC2ADDED9DB78">Halacarid mites (Acari: Halacaridae) from Esperance (Western Australia), description of two new Copidognathus species</mods:title>
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<mods:namePart id="48E8B04F3D823067E2D8622F5248A534">Bartsch, Ilse</mods:namePart>
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<mods:date id="2DC708AC5ED47FA5BC28EB0F6C766C2A">2007</mods:date>
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<treatment id="03FB87B1CE29B634088BFCFCB413A6F2" ID-DOI="http://doi.org/10.5281/zenodo.6247701" ID-GBIF-Taxon="119356015" ID-Zenodo-Dep="6247701" LSID="urn:lsid:plazi:treatment:03FB87B1CE29B634088BFCFCB413A6F2" httpUri="http://treatment.plazi.org/id/03FB87B1CE29B634088BFCFCB413A6F2" lastPageId="4" lastPageNumber="45" pageId="1" pageNumber="42">
<subSubSection id="C348652CCE29B631088BFCFCB445A69C" pageId="1" pageNumber="42" type="nomenclature">
<paragraph id="8BED36A7CE29B631088BFCFCB72CA679" blockId="1.[151,610,855,916]" box="[151,610,855,881]" pageId="1" pageNumber="42">
<heading id="D0A581CBCE29B631088BFCFCB72CA679" bold="true" box="[151,610,855,881]" fontSize="11" level="1" pageId="1" pageNumber="42" reason="1">
<emphasis id="B926EAB5CE29B631088BFCFCB72CA679" bold="true" box="[151,610,855,881]" pageId="1" pageNumber="42">
<taxonomicName id="4C524D24CE29B631088BFCFCB4AAA679" box="[151,484,855,881]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="1" pageNumber="42" phylum="Arthropoda" rank="species" species="angusticeps" status="sp. nov.">
<emphasis id="B926EAB5CE29B631088BFCFCB4AAA679" bold="true" box="[151,484,855,881]" italics="true" pageId="1" pageNumber="42">Copidognathus angusticeps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557CECE29B63109F7FCFCB72CA679" box="[491,610,855,881]" pageId="1" pageNumber="42" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BED36A7CE29B631088BFCD1B445A69C" blockId="1.[151,610,855,916]" box="[151,267,890,916]" pageId="1" pageNumber="42">
<figureCitation id="13692A22CE29B631088BFCD1B445A69C" box="[151,267,890,916]" captionStart-0="FIGURES 1 7" captionStart-1="FIGURES 8 13" captionStartId-0="2.[151,269,1456,1480]" captionStartId-1="3.[151,269,1532,1556]" captionTargetBox-0="[183,1397,396,1455]" captionTargetBox-1="[152,1425,442,1531]" captionTargetId-0="figure@2.[151,1436,351,1512]" captionTargetId-1="figure@3.[151,1436,394,1564]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 7. Copidognathus angusticeps spec. nov., male, 1, idiosoma, dorsal; 2, idiosoma, ventral; 3, ocular plate; 4, part of AE with epimeral pore; 5, gnathosoma, ventral; 6, gnathosoma, dorsal; 7, palp. (glp, gland pores; pa, porose areola; pc, pore canaliculus) Scale = 50 µm." captionText-1="FIGURES 8 13. Copidognathus angusticeps spec. nov., male, 8, leg I, medial (seta in dotted line absent on this leg); 9, basifemur to tarsus II, medial; 10, leg III, medial; 11, leg IV, medial; 12, tarsus II, lateral (medial setae and claw in broken line). Scale = 50 µm. 13, Suctoria from Copidognathus angusticeps spec. nov." httpUri-0="https://zenodo.org/record/175870/files/figure.png" httpUri-1="https://zenodo.org/record/175871/files/figure.png" pageId="1" pageNumber="42">Figs 112</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C348652CCE29B631088BFC61B046A104" pageId="1" pageNumber="42" type="materials_examined">
<paragraph id="8BED36A7CE29B631088BFC61B046A104" blockId="1.[151,1437,970,2036]" pageId="1" pageNumber="42">
<emphasis id="B926EAB5CE29B631088BFC61B4C3A6EC" bold="true" box="[151,397,970,996]" pageId="1" pageNumber="42">Material examined.</emphasis>
<typeStatus id="54E98805CE29B631098AFC61B74DA6EC" box="[406,515,970,996]" pageId="1" pageNumber="42" type="holotype">Holotype</typeStatus>
female,
<collectionCode id="ED43AE62CE29B6310A71FC61B7FBA6EC" LSID="urn:lsid:biocol.org:col:34981" box="[621,693,970,996]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="1" pageNumber="42">WAM</collectionCode>
T78346, Western
<collectingCountry id="F3457637CE29B6310B8FFC61B14FA6EC" box="[915,1025,970,996]" name="Australia" pageId="1" pageNumber="42">Australia</collectingCountry>
, Esperance, Duke of Orleans Bay,
<geoCoordinate id="EE665060CE29B631088BFC59B5B4A104" box="[151,250,1010,1036]" direction="south" orientation="latitude" pageId="1" pageNumber="42" precision="925" value="-33.916668">33°55S</geoCoordinate>
,
<geoCoordinate id="EE665060CE29B631091AFC59B437A104" box="[262,377,1010,1036]" direction="east" orientation="longitude" pageId="1" pageNumber="42" precision="925" value="122.583336">122°35E</geoCoordinate>
, from seagrass (
<taxonomicName id="4C524D24CE29B6310A24FC59B7F0A103" box="[568,702,1010,1035]" class="Liliopsida" family="Cymodoceaceae" genus="Amphibolis" kingdom="Plantae" order="Alismatales" pageId="1" pageNumber="42" phylum="Tracheophyta" rank="genus">
<emphasis id="B926EAB5CE29B6310A24FC59B7F0A103" box="[568,702,1010,1035]" italics="true" pageId="1" pageNumber="42">Amphibolis</emphasis>
</taxonomicName>
spec.) with epiflora and fauna,
<date id="FFEC1067CE29B6310C2CFC59B04AA104" box="[1072,1284,1010,1036]" pageId="1" pageNumber="42" value="2003-02-17">17 February 2003</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C348652CCE29B63108DAFBB1B476A1F4" pageId="1" pageNumber="42" type="diagnosis">
<paragraph id="8BED36A7CE29B63108DAFBB1B476A1F4" blockId="1.[151,1437,970,2036]" pageId="1" pageNumber="42">
<emphasis id="B926EAB5CE29B63108DAFBB1B408A13C" bold="true" box="[198,326,1050,1076]" pageId="1" pageNumber="42">Diagnosis.</emphasis>
Length 490 µm. Dorsal plates reticulate and densely punctate, with delicately raised areolae. PD with single pair of costae. Anterior pair of gland pores on PD in margin of plate level with insertion of leg IV. Epimeral pores large. Gnathosoma 0.36 of length of idiosoma. Rostrum extending to end of P-3, basal pair of maxillary setae in basal half of gnathosomal base. Legs slender. Telofemora with coarse porosity. Tibiae I to IV with 2, 2, 1, 1 bipectinate setae. Tarsi with 4, 4, 4, 3 dorsal setae. Pectines of paired claws with numerous delicate tines.
</paragraph>
</subSubSection>
<subSubSection id="C348652CCE29B63308DAFAA1B4D3A45A" lastPageId="3" lastPageNumber="44" pageId="1" pageNumber="42" type="description">
<paragraph id="8BED36A7CE29B63108DAFAA1B078A3BC" blockId="1.[151,1437,970,2036]" pageId="1" pageNumber="42">
<emphasis id="B926EAB5CE29B63108DAFAA1B411A02C" bold="true" box="[198,351,1290,1316]" pageId="1" pageNumber="42">Description.</emphasis>
Length 490 µm, width 277 µm. Surface of dorsal plates with delicate pits, 12 µm in diameter, which are arranged within polygons and at low magnification demonstrate a faint reticulation (
<figureCitation id="13692A22CE29B6310D5EFA99B0C7A044" box="[1346,1417,1330,1356]" captionStart="FIGURES 1 7" captionStartId="2.[151,269,1456,1480]" captionTargetBox="[183,1397,396,1455]" captionTargetId="figure@2.[151,1436,351,1512]" captionTargetPageId="2" captionText="FIGURES 1 7. Copidognathus angusticeps spec. nov., male, 1, idiosoma, dorsal; 2, idiosoma, ventral; 3, ocular plate; 4, part of AE with epimeral pore; 5, gnathosoma, ventral; 6, gnathosoma, dorsal; 7, palp. (glp, gland pores; pa, porose areola; pc, pore canaliculus) Scale = 50 µm." httpUri="https://zenodo.org/record/175870/files/figure.png" pageId="1" pageNumber="42">Fig. 1</figureCitation>
); deeper integumental layers with numerous canaliculi giving the plates an intense punctate ornamentation. Within slightly raised areolae reticulation more conspicuous and polygons smaller than in remainder of plate. Length of AD 157 µm, width 117 µm, with elongate pair of porose areolae, gland pores in lateral margin at the level of insertion of legs I. Length of OC 105 µm, width 80 µm. Lateral margin with two gland pores and a pore canaliculus (
<figureCitation id="13692A22CE29B6310979FA51B4E2A31C" box="[357,428,1530,1556]" captionStart="FIGURES 1 7" captionStartId="2.[151,269,1456,1480]" captionTargetBox="[183,1397,396,1455]" captionTargetId="figure@2.[151,1436,351,1512]" captionTargetPageId="2" captionText="FIGURES 1 7. Copidognathus angusticeps spec. nov., male, 1, idiosoma, dorsal; 2, idiosoma, ventral; 3, ocular plate; 4, part of AE with epimeral pore; 5, gnathosoma, ventral; 6, gnathosoma, dorsal; 7, palp. (glp, gland pores; pa, porose areola; pc, pore canaliculus) Scale = 50 µm." httpUri="https://zenodo.org/record/175870/files/figure.png" pageId="1" pageNumber="42">Fig. 3</figureCitation>
); raised area slightly oblique. PD almost twice length of AD, its length 305 µm, width 179 µm; anterior margin ovate. With pair of slightly raised costae, about one or two polygons wide. One pair of gland pores in lateral margin, at 0.59 relative to length of PD or level with insertion of leg IV. Following pair of gland pores in posterior margin of plate. Dorsal setae short, slender. Pair of ds-1 on AD posterior to the level of gland pores; ds-
<quantity id="4CAA9B42CE29B63109AEF931B491A3BC" box="[434,479,1690,1716]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="42" unit="in" value="2.0">2 in</quantity>
anterior margin of OC; ds-3 to ds-5 on PD immediately lateral to costae.
</paragraph>
<paragraph id="8BED36A7CE29B63108DAF969B115A2C4" blockId="1.[151,1437,970,2036]" pageId="1" pageNumber="42">
Ventral plates almost evenly punctate and faintly reticulate. Length of AE 164 µm, width 245 µm, without epimeral processes (
<figureCitation id="13692A22CE29B6310998F941B485A20C" box="[388,459,1770,1796]" captionStart="FIGURES 1 7" captionStartId="2.[151,269,1456,1480]" captionTargetBox="[183,1397,396,1455]" captionTargetId="figure@2.[151,1436,351,1512]" captionTargetPageId="2" captionText="FIGURES 1 7. Copidognathus angusticeps spec. nov., male, 1, idiosoma, dorsal; 2, idiosoma, ventral; 3, ocular plate; 4, part of AE with epimeral pore; 5, gnathosoma, ventral; 6, gnathosoma, dorsal; 7, palp. (glp, gland pores; pa, porose areola; pc, pore canaliculus) Scale = 50 µm." httpUri="https://zenodo.org/record/175870/files/figure.png" pageId="1" pageNumber="42">Fig. 2</figureCitation>
). Pair of epimeral pores within slight cavity; pores enlarged, inner diameter about 6 µm, outer diameter 15 µm. Opening of pores concealed by wide tines and densely arranged filaments (
<figureCitation id="13692A22CE29B6310D59F8B9B0C0A224" box="[1349,1422,1810,1836]" captionStart="FIGURES 1 7" captionStartId="2.[151,269,1456,1480]" captionTargetBox="[183,1397,396,1455]" captionTargetId="figure@2.[151,1436,351,1512]" captionTargetPageId="2" captionText="FIGURES 1 7. Copidognathus angusticeps spec. nov., male, 1, idiosoma, dorsal; 2, idiosoma, ventral; 3, ocular plate; 4, part of AE with epimeral pore; 5, gnathosoma, ventral; 6, gnathosoma, dorsal; 7, palp. (glp, gland pores; pa, porose areola; pc, pore canaliculus) Scale = 50 µm." httpUri="https://zenodo.org/record/175870/files/figure.png" pageId="1" pageNumber="42">Fig. 4</figureCitation>
). AE with three pairs of setae. PE extending distinctly beyond insertion of leg IV; its length 244 µm. PE with four to five long setae, one dorsal seta and three to four ventral setae. Length of GA 260 µm, width 184 µm, with three and four pgs. Length of GO 104 µm, width 55 µm. Distance from anterior margin of GO to that of GA slightly more than length of GO. Ovipositor extending somewhat beyond GO.
</paragraph>
<paragraph id="8BED36A7CE29B63208DAF871B61BA45A" blockId="1.[151,1437,970,2036]" lastBlockId="2.[151,1436,152,338]" lastPageId="2" lastPageNumber="43" pageId="1" pageNumber="42">
Gnathosoma slender, 2.8 times longer than wide, its length 174 µm, width 63 µm. Lateral and ventral flank of gnathosomal base porose (
<figureCitation id="13692A22CE2AB6320A2CFF33B739A5BA" box="[560,631,152,178]" captionStart="FIGURES 1 7" captionStartId="2.[151,269,1456,1480]" captionTargetBox="[183,1397,396,1455]" captionTargetId="figure@2.[151,1436,351,1512]" captionTargetPageId="2" captionText="FIGURES 1 7. Copidognathus angusticeps spec. nov., male, 1, idiosoma, dorsal; 2, idiosoma, ventral; 3, ocular plate; 4, part of AE with epimeral pore; 5, gnathosoma, ventral; 6, gnathosoma, dorsal; 7, palp. (glp, gland pores; pa, porose areola; pc, pore canaliculus) Scale = 50 µm." httpUri="https://zenodo.org/record/175870/files/figure.png" pageId="2" pageNumber="43">Fig. 5</figureCitation>
); dorsal flank with delicate pits. Tectum triangular (
<figureCitation id="13692A22CE2AB6320CC4FF33B06EA5BA" box="[1240,1312,152,178]" captionStart="FIGURES 1 7" captionStartId="2.[151,269,1456,1480]" captionTargetBox="[183,1397,396,1455]" captionTargetId="figure@2.[151,1436,351,1512]" captionTargetPageId="2" captionText="FIGURES 1 7. Copidognathus angusticeps spec. nov., male, 1, idiosoma, dorsal; 2, idiosoma, ventral; 3, ocular plate; 4, part of AE with epimeral pore; 5, gnathosoma, ventral; 6, gnathosoma, dorsal; 7, palp. (glp, gland pores; pa, porose areola; pc, pore canaliculus) Scale = 50 µm." httpUri="https://zenodo.org/record/175870/files/figure.png" pageId="2" pageNumber="43">Fig. 6</figureCitation>
). Rostrum almost as long as gnathosomal base. Apical pair of maxillary setae at about 0.41 relative to length rostrum, basal pair of setae in basal half of gnathosomal base or 0.23 relative to length of gnathosoma (from basis to tip of gnathosoma). Rostral sulcus extending to apical pair of setae. P-2 with dorsal seta at 0.78; P-4 with three setae in basal whorl; apically with two spurs and one setula.
</paragraph>
<caption id="DF2D662FCE2AB632088BFA1BB723A305" httpUri="https://zenodo.org/record/175870/files/figure.png" pageId="2" pageNumber="43" targetBox="[183,1397,396,1455]" targetPageId="2">
<paragraph id="8BED36A7CE2AB632088BFA1BB723A305" blockId="2.[151,1436,1456,1549]" pageId="2" pageNumber="43">
<emphasis id="B926EAB5CE2AB632088BFA1BB40DA0C0" bold="true" box="[151,323,1456,1480]" pageId="2" pageNumber="43">FIGURES 17.</emphasis>
<taxonomicName id="4C524D24CE2AB6320957FA1AB73AA0C0" box="[331,628,1457,1480]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="2" pageNumber="43" phylum="Arthropoda" rank="species" species="angusticeps" status="sp. nov.">
<emphasis id="B926EAB5CE2AB6320957FA1AB73AA0C0" box="[331,628,1457,1480]" italics="true" pageId="2" pageNumber="43">Copidognathus angusticeps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557CECE2AB6320A67FA1BB7A7A0C0" box="[635,745,1456,1480]" pageId="2" pageNumber="43" rank="species">
<emphasis id="B926EAB5CE2AB6320A67FA1BB7A7A0C0" bold="true" box="[635,745,1456,1480]" pageId="2" pageNumber="43">spec. nov.</emphasis>
</taxonomicNameLabel>
, male, 1, idiosoma, dorsal; 2, idiosoma, ventral; 3, ocular plate; 4, part of AE with epimeral pore; 5, gnathosoma, ventral; 6, gnathosoma, dorsal; 7, palp. (glp, gland pores; pa, porose areola; pc, pore canaliculus) Scale = 50 µm.
</paragraph>
</caption>
<paragraph id="8BED36A7CE2AB63308DAF9EAB4D3A45A" blockId="2.[151,1437,1601,2027]" lastBlockId="3.[151,1436,152,338]" lastPageId="3" lastPageNumber="44" pageId="2" pageNumber="43">
Legs slender; without lamellae or fossa membranes. Lateral flanks of telofemora and tibiae I and II with coarse porosity; porosity of lateral flanks of trochanters, telofemora and tibiae III and IV present but somewhat more delicate. Medial flanks of telofemora reticulate, accordingly ventral and dorsal margin crenulate; each polygon punctate and enclosing pits. Length:height ratio of telofemora I to III 2.52.6:1 (
<figureCitation id="13692A22CE2AB6320D0DF912B0C0A3DB" box="[1297,1422,1721,1747]" captionStart="FIGURES 8 13" captionStartId="3.[151,269,1532,1556]" captionTargetBox="[152,1425,442,1531]" captionTargetId="figure@3.[151,1436,394,1564]" captionTargetPageId="3" captionText="FIGURES 8 13. Copidognathus angusticeps spec. nov., male, 8, leg I, medial (seta in dotted line absent on this leg); 9, basifemur to tarsus II, medial; 10, leg III, medial; 11, leg IV, medial; 12, tarsus II, lateral (medial setae and claw in broken line). Scale = 50 µm. 13, Suctoria from Copidognathus angusticeps spec. nov." httpUri="https://zenodo.org/record/175871/files/figure.png" pageId="2" pageNumber="43">Figs 810</figureCitation>
); telofemur IV 2.8 times longer than high (
<figureCitation id="13692A22CE2AB6320A67F94AB79CA3F3" box="[635,722,1761,1787]" captionStart="FIGURES 8 13" captionStartId="3.[151,269,1532,1556]" captionTargetBox="[152,1425,442,1531]" captionTargetId="figure@3.[151,1436,394,1564]" captionTargetPageId="3" captionText="FIGURES 8 13. Copidognathus angusticeps spec. nov., male, 8, leg I, medial (seta in dotted line absent on this leg); 9, basifemur to tarsus II, medial; 10, leg III, medial; 11, leg IV, medial; 12, tarsus II, lateral (medial setae and claw in broken line). Scale = 50 µm. 13, Suctoria from Copidognathus angusticeps spec. nov." httpUri="https://zenodo.org/record/175871/files/figure.png" pageId="2" pageNumber="43">Fig. 11</figureCitation>
). Telofemora I and II longer than the legs tibiae, telofemora of legs III and IV shorter than their tibiae. Leg chaetotaxy: leg I, 1, 2, 5, 4, 7, 67; leg II, 1, 2, 5, 4, 7, 4; leg III, 1, 2, 2, 3, 5, 4; leg IV, 0, 2, 2, 3, 5, 3. Tibiae I to IV with 2, 2, 1, 1 bipectinate, bluntly ending ventromedial setae and 1, 1, 1, 1 slender, smooth ventral setae. Tarsus I with four dorsal setae, included the long, slender solenidion; ventrally on one tarsus three on the other tarsus two setae (
<figureCitation id="13692A22CE2AB6320BC6F82AB16FA293" box="[986,1057,1921,1947]" captionStart="FIGURES 8 13" captionStartId="3.[151,269,1532,1556]" captionTargetBox="[152,1425,442,1531]" captionTargetId="figure@3.[151,1436,394,1564]" captionTargetPageId="3" captionText="FIGURES 8 13. Copidognathus angusticeps spec. nov., male, 8, leg I, medial (seta in dotted line absent on this leg); 9, basifemur to tarsus II, medial; 10, leg III, medial; 11, leg IV, medial; 12, tarsus II, lateral (medial setae and claw in broken line). Scale = 50 µm. 13, Suctoria from Copidognathus angusticeps spec. nov." httpUri="https://zenodo.org/record/175871/files/figure.png" pageId="2" pageNumber="43">Fig. 8</figureCitation>
). Tarsus II with four long setae, solenidion seta-like, 43 µm long, in dorsolateral position (
<figureCitation id="13692A22CE2AB6320B49F802B6FEA2CB" box="[853,944,1961,1987]" captionStart="FIGURES 8 13" captionStartId="3.[151,269,1532,1556]" captionTargetBox="[152,1425,442,1531]" captionTargetId="figure@3.[151,1436,394,1564]" captionTargetPageId="3" captionText="FIGURES 8 13. Copidognathus angusticeps spec. nov., male, 8, leg I, medial (seta in dotted line absent on this leg); 9, basifemur to tarsus II, medial; 10, leg III, medial; 11, leg IV, medial; 12, tarsus II, lateral (medial setae and claw in broken line). Scale = 50 µm. 13, Suctoria from Copidognathus angusticeps spec. nov." httpUri="https://zenodo.org/record/175871/files/figure.png" pageId="2" pageNumber="43">Fig. 12</figureCitation>
). Dorsomedial fossary seta much shorter than conspicuously long lateral fossary seta. On tarsi I and II two distal fossary setae situated unpaired, on tarsi III and IV paired. Tarsi I and II each with pair of doubled pas; at tip of tarsus III medial pas slender, setalike, lateral pas short, bristle-like; at tip of tarsus IV both medial and lateral pas short, bristle-like. Claws slender, all with accessory process. Pectines of claws II to IV with numerous delicate tines. Ventral plates AE, PE and GA with suctorians, their length about 59 µm; theca annulated (
<figureCitation id="13692A22CE2BB6330CF8FEBBB077A422" box="[1252,1337,272,298]" captionStart="FIGURES 8 13" captionStartId="3.[151,269,1532,1556]" captionTargetBox="[152,1425,442,1531]" captionTargetId="figure@3.[151,1436,394,1564]" captionTargetPageId="3" captionText="FIGURES 8 13. Copidognathus angusticeps spec. nov., male, 8, leg I, medial (seta in dotted line absent on this leg); 9, basifemur to tarsus II, medial; 10, leg III, medial; 11, leg IV, medial; 12, tarsus II, lateral (medial setae and claw in broken line). Scale = 50 µm. 13, Suctoria from Copidognathus angusticeps spec. nov." httpUri="https://zenodo.org/record/175871/files/figure.png" pageId="3" pageNumber="44">Fig. 13</figureCitation>
), 39 µm long, and stalk 19 µm.
</paragraph>
</subSubSection>
<caption id="DF2D662FCE2BB633088BFA57B146A351" httpUri="https://zenodo.org/record/175871/files/figure.png" pageId="3" pageNumber="44" targetBox="[152,1425,442,1531]" targetPageId="3">
<paragraph id="8BED36A7CE2BB633088BFA57B146A351" blockId="3.[151,1436,1532,1625]" pageId="3" pageNumber="44">
<emphasis id="B926EAB5CE2BB633088BFA57B41EA31C" bold="true" box="[151,336,1532,1556]" pageId="3" pageNumber="44">FIGURES 813.</emphasis>
<taxonomicName id="4C524D24CE2BB633094BFA57B730A31B" box="[343,638,1532,1555]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="angusticeps" status="sp. nov.">
<emphasis id="B926EAB5CE2BB633094BFA57B730A31B" box="[343,638,1532,1555]" italics="true" pageId="3" pageNumber="44">Copidognathus angusticeps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557CECE2BB6330A99FA57B7BDA31C" box="[645,755,1532,1556]" pageId="3" pageNumber="44" rank="species">
<emphasis id="B926EAB5CE2BB6330A99FA57B7BDA31C" bold="true" box="[645,755,1532,1556]" pageId="3" pageNumber="44">spec. nov.</emphasis>
</taxonomicNameLabel>
, male, 8, leg I, medial (seta in dotted line absent on this leg); 9, basifemur to tarsus II, medial; 10, leg III, medial; 11, leg IV, medial; 12, tarsus II, lateral (medial setae and claw in broken line). Scale = 50 µm. 13, Suctoria from
<taxonomicName id="4C524D24CE2BB6330A71F9E9B6DAA351" box="[621,916,1602,1625]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="angusticeps" status="sp. nov.">
<emphasis id="B926EAB5CE2BB6330A71F9E9B6DAA351" box="[621,916,1602,1625]" italics="true" pageId="3" pageNumber="44">Copidognathus angusticeps</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21557CECE2BB6330B87F9EAB146A351" box="[923,1032,1601,1625]" pageId="3" pageNumber="44" rank="species">spec. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="C348652CCE2BB63308DAF927B72DA3C6" pageId="3" pageNumber="44" type="etymology">
<paragraph id="8BED36A7CE2BB63308DAF927B72DA3C6" blockId="3.[151,1437,1676,2022]" pageId="3" pageNumber="44">
<emphasis id="B926EAB5CE2BB63308DAF927B41DA3AE" bold="true" box="[198,339,1676,1702]" pageId="3" pageNumber="44">Etymology.</emphasis>
Because of its narrow gnathosoma the species is called
<taxonomicName id="4C524D24CE2BB6330BEBF926B1CFA3AE" box="[1015,1153,1677,1702]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="B926EAB5CE2BB6330BEBF926B1CFA3AE" box="[1015,1153,1677,1702]" italics="true" pageId="3" pageNumber="44">angusticeps</emphasis>
</taxonomicName>
, derived from angustus (Latin), narrow, and ceps (Latin), head.
</paragraph>
</subSubSection>
<subSubSection id="C348652CCE2BB63408DAF977B02EA5D2" lastPageId="4" lastPageNumber="45" pageId="3" pageNumber="44" type="discussion">
<paragraph id="8BED36A7CE2BB63408DAF977B02EA5D2" blockId="3.[151,1437,1676,2022]" lastBlockId="4.[151,1437,152,1018]" lastPageId="4" lastPageNumber="45" pageId="3" pageNumber="44">
<emphasis id="B926EAB5CE2BB63308DAF977B40EA3FE" bold="true" box="[198,320,1756,1782]" pageId="3" pageNumber="44">Remarks.</emphasis>
A character, not found elsewhere in the genus
<taxonomicName id="4C524D24CE2BB6330B9CF976B17DA3FE" box="[896,1075,1757,1782]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB5CE2BB6330B9CF976B17DA3FE" box="[896,1075,1757,1782]" italics="true" pageId="3" pageNumber="44">Copidognathus</emphasis>
</taxonomicName>
, is the presence of two gland pores on the OC. In general
<taxonomicName id="4C524D24CE2BB63309F9F8AEB7D6A216" box="[485,664,1797,1822]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB5CE2BB63309F9F8AEB7D6A216" box="[485,664,1797,1822]" italics="true" pageId="3" pageNumber="44">Copidognathus</emphasis>
</taxonomicName>
species have four pairs of pores in all, one pair of pores is on the AD, one pair on the OC and two pairs are on the PD. The gland pores may be almost vestigial or very conspicuous, opening on raised cones. Five pairs, as present in many other genera, is expected to be the plesiomorphic state, and the presence of two gland pores on the OC is a common phenomenon in several halacarid genera, amongst others in
<taxonomicName id="4C524D24CE2BB63309CDF80EB7FFA2B6" authority="Viets" authorityName="Viets" box="[465,689,1956,1982]" class="Arachnida" family="Halacaridae" genus="Halacarellus" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB5CE2BB63309CDF80EB727A2B6" box="[465,617,1957,1982]" italics="true" pageId="3" pageNumber="44">Halacarellus</emphasis>
Viets
</taxonomicName>
,
<taxonomicName id="4C524D24CE2BB6330AA2F80EB6D6A2B6" authority="Newell" authorityName="Newell" box="[702,920,1956,1982]" class="Arachnida" family="Halacaridae" genus="Isobactrus" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB5CE2BB6330AA2F80EB674A2B6" box="[702,826,1957,1982]" italics="true" pageId="3" pageNumber="44">Isobactrus</emphasis>
Newell
</taxonomicName>
and
<taxonomicName id="4C524D24CE2BB6330BC8F80EB01DA2B6" authority="Trouessart. The" authorityName="Trouessart. The" box="[980,1363,1956,1982]" class="Arachnida" family="Halacaridae" genus="Rhombognathus" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB5CE2BB6330BC8F80EB1DCA2B6" box="[980,1170,1957,1982]" italics="true" pageId="3" pageNumber="44">Rhombognathus</emphasis>
Trouessart. The
</taxonomicName>
pores are rarely adjacent, as in
<taxonomicName id="4C524D24CE2BB63309A7F866B729A2EE" box="[443,615,1997,2022]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="B926EAB5CE2BB63309A7F866B729A2EE" box="[443,615,1997,2022]" italics="true" pageId="3" pageNumber="44">C. angusticeps</emphasis>
</taxonomicName>
and in
<taxonomicName id="4C524D24CE2BB6330AA2F866B131A2EE" authority="Bartsch, 1992" authorityName="Bartsch" authorityYear="1992" box="[702,1151,1996,2022]" class="Arachnida" family="Halacaridae" genus="Mictognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Trombidiformes" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="secundus">
<emphasis id="B926EAB5CE2BB6330AA2F866B69CA2EE" box="[702,978,1997,2022]" italics="true" pageId="3" pageNumber="44">Mictognathus secundus</emphasis>
<bibRefCitation id="EFC34B56CE2BB6330BC5F867B131A2EE" author="Bartsch" box="[985,1151,1996,2022]" pageId="3" pageNumber="49" refString="Bartsch, I. (1992) Mictognathus secundus, a psammophilous halacarid mite (Acari: Halacaridae). Acarologia, 33, 85 - 89." type="journal article" year="1992">Bartsch, 1992</bibRefCitation>
</taxonomicName>
. Two gland pores on the OC of
<taxonomicName id="4C524D24CE2CB63408FAFF33B4DCA5B9" box="[230,402,152,177]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="B926EAB5CE2CB63408FAFF33B4DCA5B9" box="[230,402,152,177]" italics="true" pageId="4" pageNumber="45">C. angusticeps</emphasis>
</taxonomicName>
may be an anomaly of this specimen or species, but might also represent a situation otherwise reduced in the genus
<taxonomicName id="4C524D24CE2CB63409F8FF6BB7D9A5D1" box="[484,663,192,217]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B926EAB5CE2CB63409F8FF6BB7D9A5D1" box="[484,663,192,217]" italics="true" pageId="4" pageNumber="45">Copidognathus</emphasis>
</taxonomicName>
or a duplication of the gland pore always present on the OC.
</paragraph>
</subSubSection>
<subSubSection id="C348652CCE2CB63408DAFF43B076A6A2" pageId="4" pageNumber="45" type="reference_group">
<paragraph id="8BED36A7CE2CB63408DAFF43B156A7EA" blockId="4.[151,1437,152,1018]" pageId="4" pageNumber="45">
In
<taxonomicName id="4C524D24CE2CB63408FBFF43B762A409" box="[231,556,232,257]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="B926EAB5CE2CB63408FBFF43B762A409" box="[231,556,232,257]" italics="true" pageId="4" pageNumber="45">Copidognathus angusticeps</emphasis>
</taxonomicName>
the PD bears the anterior pair of gland pores in the margin of the plate. A similar position is found in, e.g.,
<taxonomicName id="4C524D24CE2CB6340A31FEBBB614A422" authority="Bartsch, 1999" authorityName="Bartsch" authorityYear="1999" box="[557,858,272,298]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="amalus">
<emphasis id="B926EAB5CE2CB6340A31FEBBB7E6A421" box="[557,680,272,297]" italics="true" pageId="4" pageNumber="45">C. amalus</emphasis>
<bibRefCitation id="EFC34B56CE2CB6340AAEFEBBB614A422" author="Bartsch" box="[690,858,272,298]" pageId="4" pageNumber="49" refString="Bartsch, I. (1999) Halacaridae (Acari) from Western Australia. Four species of Copidognathus. In D. I. Walker and F. E. Wells (eds), Proceedings of the Ninth International Marine Biological Workshop: The Seagrass Flora and Fauna of Rottnest Island, Western Australia, Western Australian Museum, Perth, 333 - 357." type="book chapter" year="1999">Bartsch, 1999</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C524D24CE2CB6340B76FEBBB18BA422" authority="Viets, 1940" authorityName="Viets" authorityYear="1940" box="[874,1221,272,298]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="brachystomus">
<emphasis id="B926EAB5CE2CB6340B76FEBBB17CA421" box="[874,1074,272,297]" italics="true" pageId="4" pageNumber="45">C. brachystomus</emphasis>
<bibRefCitation id="EFC34B56CE2CB6340C20FEBBB18BA422" author="Viets" box="[1084,1221,272,298]" pageId="4" pageNumber="49" refString="Viets, K. (1940) Meeresmilben aus der Adria (Halacaridae und Hydrachnellae, Acari). Archiv fur Naturgeschichte, (Neue Folge) 9, 1 - 135." type="journal article" year="1940">Viets, 1940</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C524D24CE2CB6340D18FEBBB40AA45A" authority="Bartsch, 1997" authorityName="Bartsch" authorityYear="1997" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="biodomus">
<emphasis id="B926EAB5CE2CB6340D18FEBBB0D2A421" box="[1284,1436,272,297]" italics="true" pageId="4" pageNumber="45">C. biodomus</emphasis>
<bibRefCitation id="EFC34B56CE2CB634088BFE93B40AA45A" author="Bartsch" box="[151,324,312,338]" pageId="4" pageNumber="49" refString="Bartsch, I. (1997) Copidognathus biodomus (Halacaride: Acari), a new species from eastern Canada. Mitteilungen aus dem hamburgischen zoologischen Museum und Institut, 94, 153 - 159." type="journal article" year="1997">Bartsch, 1997</bibRefCitation>
</taxonomicName>
, species recorded from southwestern
<collectingCountry id="F3457637CE2CB6340B3DFE93B6DEA45A" box="[801,912,312,338]" name="Australia" pageId="4" pageNumber="45">Australia</collectingCountry>
, Europe and eastern
<collectingCountry id="F3457637CE2CB6340CBDFE93B1B2A45A" box="[1185,1276,312,338]" name="Canada" pageId="4" pageNumber="45">Canada</collectingCountry>
, respectively (
<bibRefCitation id="EFC34B56CE2CB6340883FECBB40AA472" author="Bartsch" box="[159,324,352,378]" pageId="4" pageNumber="49" refString="Bartsch, I. (1997) Copidognathus biodomus (Halacaride: Acari), a new species from eastern Canada. Mitteilungen aus dem hamburgischen zoologischen Museum und Institut, 94, 153 - 159." type="journal article" year="1997">Bartsch 1997</bibRefCitation>
,
<bibRefCitation id="EFC34B56CE2CB6340949FECBB4DCA472" author="Bartsch" box="[341,402,352,378]" pageId="4" pageNumber="49" refString="Bartsch, I. (1999) Halacaridae (Acari) from Western Australia. Four species of Copidognathus. In D. I. Walker and F. E. Wells (eds), Proceedings of the Ninth International Marine Biological Workshop: The Seagrass Flora and Fauna of Rottnest Island, Western Australia, Western Australian Museum, Perth, 333 - 357." type="book chapter" year="1999">1999</bibRefCitation>
,
<bibRefCitation id="EFC34B56CE2CB63409B8FECBB4ACA472" author="Bartsch" box="[420,482,352,378]" pageId="4" pageNumber="49" refString="Bartsch, I. (2001) Black Sea Copidognathinae (Arachnida: Acari: Halacaridae): A review. Mitteilungen aus dem Museum fur Naturkunde in Berlin, Zoologische Reihe 77, 247 - 275." type="journal article" year="2001">2001</bibRefCitation>
;
<bibRefCitation id="EFC34B56CE2CB63409E8FECBB7CDA472" author="Viets" box="[500,643,352,378]" pageId="4" pageNumber="49" refString="Viets, K. (1940) Meeresmilben aus der Adria (Halacaridae und Hydrachnellae, Acari). Archiv fur Naturgeschichte, (Neue Folge) 9, 1 - 135." type="journal article" year="1940">Viets, 1940</bibRefCitation>
).
<taxonomicName id="4C524D24CE2CB6340A80FECBB6A8A471" box="[668,998,352,377]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="B926EAB5CE2CB6340A80FECBB6A8A471" box="[668,998,352,377]" italics="true" pageId="4" pageNumber="45">Copidognathus angusticeps</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D24CE2CB6340C37FECBB1E9A471" box="[1067,1191,352,377]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="amalus">
<emphasis id="B926EAB5CE2CB6340C37FECBB1E9A471" box="[1067,1191,352,377]" italics="true" pageId="4" pageNumber="45">C. amalus</emphasis>
</taxonomicName>
are similar in their general shape but differ in the ornamentation of the dorsal plates, the shape of the OC and the size of the epimeral pores. The dorsal plates of
<taxonomicName id="4C524D24CE2CB6340A54FE1BB7B8A4C1" box="[584,758,432,457]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="B926EAB5CE2CB6340A54FE1BB7B8A4C1" box="[584,758,432,457]" italics="true" pageId="4" pageNumber="45">C. angusticeps</emphasis>
</taxonomicName>
demonstrate a faint reticulation due to minute pits which are arranged in polygons, the width of the OC is less than 1.5 times the length and the epimeral pores are more than 10 µm in width, in contrast, the dorsal plates of
<taxonomicName id="4C524D24CE2CB6340B11FDABB6CBA711" box="[781,901,512,537]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="amalus">
<emphasis id="B926EAB5CE2CB6340B11FDABB6CBA711" box="[781,901,512,537]" italics="true" pageId="4" pageNumber="45">C. amalus</emphasis>
</taxonomicName>
are characterized by the wide porose areolae with rosette pores (
<bibRefCitation id="EFC34B56CE2CB6340961FD83B796A74A" author="Bartsch" box="[381,728,552,578]" pageId="4" pageNumber="49" refString="Bartsch, I. (1999) Halacaridae (Acari) from Western Australia. Four species of Copidognathus. In D. I. Walker and F. E. Wells (eds), Proceedings of the Ninth International Marine Biological Workshop: The Seagrass Flora and Fauna of Rottnest Island, Western Australia, Western Australian Museum, Perth, 333 - 357." type="book chapter" year="1999">Bartsch 1999: fig. 6A and C</bibRefCitation>
), the OC, with a tail-like posterior process, are three times longer than wide, and the epimeral pores are 3 µm in diameter. The dorsal plates of both
<taxonomicName id="4C524D24CE2CB6340CAEFDFBB009A761" box="[1202,1351,592,617]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="biodomus">
<emphasis id="B926EAB5CE2CB6340CAEFDFBB009A761" box="[1202,1351,592,617]" italics="true" pageId="4" pageNumber="45">C. biodomus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C524D24CE2CB6340D9DFDFBB474A799" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="brachystomus">
<emphasis id="B926EAB5CE2CB6340D9DFDFBB474A799" italics="true" pageId="4" pageNumber="45">C. brachystomus</emphasis>
</taxonomicName>
bear areolae with rosette pores, the length of the gnathosoma is less than 1.5 times the width (versus 2.8 in
<taxonomicName id="4C524D24CE2CB6340922FD0BB4A5A7B1" box="[318,491,672,697]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="B926EAB5CE2CB6340922FD0BB4A5A7B1" box="[318,491,672,697]" italics="true" pageId="4" pageNumber="45">C. angusticeps</emphasis>
</taxonomicName>
), the epimeral pores 35 µm in diameter, and the length of telofemur I less than twice the height (versus a length:height ratio of about 2.5 in
<taxonomicName id="4C524D24CE2CB6340B47FD63B149A7E9" box="[859,1031,712,737]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="B926EAB5CE2CB6340B47FD63B149A7E9" box="[859,1031,712,737]" italics="true" pageId="4" pageNumber="45">C. angusticeps</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8BED36A7CE2CB63408DAFD5BB076A6A2" blockId="4.[151,1437,152,1018]" pageId="4" pageNumber="45">
The epimeral pores of
<taxonomicName id="4C524D24CE2CB63409CFFD5BB7CFA601" box="[467,641,752,777]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="angusticeps">
<emphasis id="B926EAB5CE2CB63409CFFD5BB7CFA601" box="[467,641,752,777]" italics="true" pageId="4" pageNumber="45">C. angusticeps</emphasis>
</taxonomicName>
are enlarged and obscured by tufts of filaments. Enlarged epimeral pores are known in
<taxonomicName id="4C524D24CE2CB634099DFCB3B67DA63A" ID-CoL="Y4J4" authority="Benfatti et al., 1989" authorityName="Benfatti et al." authorityYear="1989" box="[385,819,792,818]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="dactyloporus">
<emphasis id="B926EAB5CE2CB634099DFCB3B773A639" box="[385,573,792,817]" italics="true" pageId="4" pageNumber="45">C. dactyloporus</emphasis>
<bibRefCitation id="EFC34B56CE2CB6340A58FCB3B67DA63A" author="Benfatti" box="[580,819,792,818]" pageId="4" pageNumber="49" refString="Benfatti, D., Mari, M. &amp; Morselli, I. (1989) Copidognathus dactyloporus, a new freshwater species (Halacaridae, Acari). Bollettino di Zoologia, 56, 99 - 104." type="journal article" year="1989">
Benfatti
<emphasis id="B926EAB5CE2CB6340AB7FCB3B791A639" box="[683,735,792,817]" italics="true" pageId="4" pageNumber="45">et al</emphasis>
., 1989
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C524D24CE2CB6340B23FCB3B132A63A" ID-CoL="Y4J8" authority="Viets, 1935" authorityName="Viets" authorityYear="1935" box="[831,1148,792,818]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="tectiporus">
<emphasis id="B926EAB5CE2CB6340B23FCB3B699A639" box="[831,983,792,817]" italics="true" pageId="4" pageNumber="45">C. tectiporus</emphasis>
(
<bibRefCitation id="EFC34B56CE2CB6340BF4FCB3B13CA63A" author="Viets" box="[1000,1138,792,818]" pageId="4" pageNumber="49" refString="Viets, K. (1935) Wassermilben aus Bulgarien. Zoologischer Anzeiger, 109, 33 - 39." type="journal article" year="1935">Viets, 1935</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="4C524D24CE2CB6340C94FCB3B453A652" ID-CoL="Y4J5" authority="Benfatti et al., 1992" authorityName="Benfatti et al." authorityYear="1992" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="hephaestios">
<emphasis id="B926EAB5CE2CB6340C94FCB3B07BA639" box="[1160,1333,792,817]" italics="true" pageId="4" pageNumber="45">C. hephaestios</emphasis>
<bibRefCitation id="EFC34B56CE2CB6340D21FCB3B453A652" author="Benfatti" pageId="4" pageNumber="49" refString="Benfatti, D., Mari, M. &amp; Morselli, I. (1992) Halacaroidea (Acari, Actinedida) from four lakes of volcanic origin in Lazio (Central Italy). Bollettino di Zoologia, 59, 105 - 111." type="journal article" year="1992">
Benfatti
<emphasis id="B926EAB5CE2CB634088BFCEBB584A651" box="[151,202,832,857]" italics="true" pageId="4" pageNumber="45">et al</emphasis>
., 1992
</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C524D24CE2CB6340935FCEBB72DA652" ID-CoL="Y4J6" authority="Bartsch, 1979" authorityName="Bartsch" authorityYear="1979" box="[297,611,832,858]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="poriferus">
<emphasis id="B926EAB5CE2CB6340935FCEBB4F9A651" box="[297,439,832,857]" italics="true" pageId="4" pageNumber="45">C. poriferus</emphasis>
<bibRefCitation id="EFC34B56CE2CB63409A2FCEBB72DA652" author="Bartsch" box="[446,611,832,858]" pageId="4" pageNumber="49" refString="Bartsch, I. (1979) Halacaridae (Acari) von der Atlantikkuste Nordamerikas. Beschreibung der Arten. Mikrofauna des Meeresbodens, 79, 1 - 62." type="journal article" year="1979">Bartsch, 1979</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C524D24CE2CB6340ABDFCEBB686A652" ID-CoL="Y4J7" authority="Viets, 1936" authorityName="Viets" authorityYear="1936" box="[673,968,832,858]" class="Arachnida" family="Halacaridae" genus="Copidognathus" kingdom="Animalia" order="Trombidiformes" pageId="4" pageNumber="45" phylum="Arthropoda" rank="species" species="profundus">
<emphasis id="B926EAB5CE2CB6340ABDFCEBB677A651" box="[673,825,832,857]" italics="true" pageId="4" pageNumber="45">C. profundus</emphasis>
<bibRefCitation id="EFC34B56CE2CB6340B5CFCEBB686A652" author="Viets" box="[832,968,832,858]" pageId="4" pageNumber="49" refString="Viets, K. (1936) Hydracarinen aus Jugoslawien (Systematische, okologische, faunistische und tiergeographische Untersuchungen uber die Hydrachnellae und Halacaridae des Sußwassers). Archiv fur Hydrobiologie, 29, 351 - 409." type="journal article" year="1936">Viets, 1936</bibRefCitation>
</taxonomicName>
, all living in low salinity habitats (
<bibRefCitation id="EFC34B56CE2CB6340D42FCEBB59AA68A" author="Viets" pageId="4" pageNumber="49" refString="Viets, K. (1935) Wassermilben aus Bulgarien. Zoologischer Anzeiger, 109, 33 - 39." type="journal article" year="1935">Viets 1935</bibRefCitation>
,
<bibRefCitation id="EFC34B56CE2CB63408FFFCC3B46FA68A" author="Viets" box="[227,289,872,898]" pageId="4" pageNumber="49" refString="Viets, K. (1936) Hydracarinen aus Jugoslawien (Systematische, okologische, faunistische und tiergeographische Untersuchungen uber die Hydrachnellae und Halacaridae des Sußwassers). Archiv fur Hydrobiologie, 29, 351 - 409." type="journal article" year="1936">1936</bibRefCitation>
;
<bibRefCitation id="EFC34B56CE2CB6340933FCC3B49EA68A" author="Bartsch" box="[303,464,872,898]" pageId="4" pageNumber="49" refString="Bartsch, I. (1979) Halacaridae (Acari) von der Atlantikkuste Nordamerikas. Beschreibung der Arten. Mikrofauna des Meeresbodens, 79, 1 - 62." type="journal article" year="1979">Bartsch 1979</bibRefCitation>
;
<bibRefCitation id="EFC34B56CE2CB63409C3FCC3B785A68A" author="Benfatti" box="[479,715,872,898]" pageId="4" pageNumber="49" refString="Benfatti, D., Mari, M. &amp; Morselli, I. (1989) Copidognathus dactyloporus, a new freshwater species (Halacaridae, Acari). Bollettino di Zoologia, 56, 99 - 104." type="journal article" year="1989">
Benfatti
<emphasis id="B926EAB5CE2CB6340A5BFCC3B732A689" box="[583,636,872,897]" italics="true" pageId="4" pageNumber="45">et al</emphasis>
. 1989
</bibRefCitation>
,
<bibRefCitation id="EFC34B56CE2CB6340AC6FCC3B657A68A" author="Benfatti" box="[730,793,872,898]" pageId="4" pageNumber="49" refString="Benfatti, D., Mari, M. &amp; Morselli, I. (1992) Halacaroidea (Acari, Actinedida) from four lakes of volcanic origin in Lazio (Central Italy). Bollettino di Zoologia, 59, 105 - 111." type="journal article" year="1992">1992</bibRefCitation>
), and in these species the enlargement of the pores is expected to facilitate the osmoregulation and thus be of competitive advantage in a life in freshwater.
</paragraph>
</subSubSection>
<subSubSection id="C348652CCE2CB63408DAFC1CB413A6F2" pageId="4" pageNumber="45" type="distribution">
<paragraph id="8BED36A7CE2CB63408DAFC1CB413A6F2" blockId="4.[151,1437,152,1018]" pageId="4" pageNumber="45">
<emphasis id="B926EAB5CE2CB63408DAFC1CB429A6D9" bold="true" box="[198,359,951,977]" pageId="4" pageNumber="45">Distribution.</emphasis>
Esperance, Western
<collectingCountry id="F3457637CE2CB6340A7DFC13B79EA6DA" box="[609,720,952,978]" name="Australia" pageId="4" pageNumber="45">Australia</collectingCountry>
. The single individual recorded was extracted from the epibios on seagrass.
</paragraph>
</subSubSection>
</treatment>
</document>