treatments-xml/data/03/FB/87/03FB87A4FFE9FC7866EFA2B1FD870076.xml
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<document id="3263F4CAE7AC979C1209A2543AE9F9AB" ID-CLB-Dataset="36257" ID-DOI="10.15553/c2016v712a1" ID-GBIF-Dataset="236a38ef-6345-4460-ad5f-ee55cc13eaa1" ID-Zenodo-Dep="400358" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1489701763825" checkinUser="admin" docAuthor="Tefy H. Andriamihajarivo, Porter P. Lowry Ii &amp; George E. Schatz" docDate="2016" docId="03FB87A4FFE9FC7866EFA2B1FD870076" docLanguage="en" docName="Candollea.71.2.167-172.pdf.imf" docOrigin="Candollea 71 (2)" docSource="http://www.bioone.org/doi/full/10.15553/c2016v712a1" docStyle="DocumentStyle:1E5CC43F38D0DBC531513809D1267F7B.3:Candollea.2015-.journal_article" docStyleId="1E5CC43F38D0DBC531513809D1267F7B" docStyleName="Candollea.2015-.journal_article" docStyleVersion="3" docTitle="Pentachlaena pestita Andriam., Lowry &amp; G.E. Schatz, spec. nova" docType="treatment" docVersion="14" lastPageNumber="172" masterDocId="FFC2FFDCFFEBFC7E6667A77FFFA0033A" masterDocTitle="Endemic Families of Madagascar. XIV. a New Restricted Range Species of Pentachlaena H. Perrier (Sarcolaenaceae) from Central Madagascar" masterLastPageNumber="172" masterPageNumber="166" pageNumber="168" updateTime="1698438074985" updateUser="ExternalLinkService">
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<mods:title id="FAE81EAB8557DF709482417E072C170C">Endemic Families of Madagascar. XIV. a New Restricted Range Species of Pentachlaena H. Perrier (Sarcolaenaceae) from Central Madagascar</mods:title>
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<mods:namePart id="65B7168DC139F5361BB965C827824A45">Tefy H. Andriamihajarivo</mods:namePart>
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<mods:namePart id="6FCFA6B5048DB4690CF640411FA4E56B">Porter P. Lowry Ii</mods:namePart>
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<mods:namePart id="322BD5C3C2A2D9E815FD5B78CB7E07D9">George E. Schatz</mods:namePart>
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<mods:title id="534E572F2F17E1DC6EA211BD00F5132E">Candollea</mods:title>
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<mods:date id="E0F4EC788381A62607BC73F30202A9D8">2016</mods:date>
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<treatment id="03FB87A4FFE9FC7866EFA2B1FD870076" ID-DOI="http://doi.org/10.5281/zenodo.5689724" ID-GBIF-Taxon="127847732" ID-Zenodo-Dep="5689724" LSID="urn:lsid:plazi:treatment:03FB87A4FFE9FC7866EFA2B1FD870076" httpUri="http://treatment.plazi.org/id/03FB87A4FFE9FC7866EFA2B1FD870076" lastPageId="6" lastPageNumber="172" pageId="2" pageNumber="168">
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<taxonomicName id="4C524D31FFE9FC7C66EFA2B1FD6F06DC" authority="Andriam., Lowry &amp; G.E. Schatz" authorityName="Andriam., Lowry &amp; G.E. Schatz" box="[136,719,1486,1511]" class="Magnoliopsida" family="Sarcolaenaceae" genus="Pentachlaena" kingdom="Plantae" order="Huerteales" pageId="2" pageNumber="168" phylum="Tracheophyta" rank="species" species="pestita" status="sp. nov.">
<emphasis id="B926EAA0FFE9FC7C66EFA2B1FEC306DC" bold="true" box="[136,355,1486,1510]" italics="true" pageId="2" pageNumber="168">Pentachlaena pestita</emphasis>
Andriam., Lowry &amp; G.E. Schatz
</taxonomicName>
,
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<emphasis id="B926EAA0FFE9FC7C64BCA2B1FF1E053C" bold="true" pageId="2" pageNumber="168">spec. nova</emphasis>
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</paragraph>
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<subSubSection id="C3486539FFE9FC7C66A2A291FE92053C" box="[197,306,1518,1542]" pageId="2" pageNumber="168" type="description">
<paragraph id="8BED36B2FFE9FC7C66A2A291FE92053C" blockId="2.[136,787,1437,1746]" box="[197,306,1518,1542]" pageId="2" pageNumber="168">
(
<figureCitation id="13692A37FFE9FC7C66ABA291FEA8053C" box="[204,264,1518,1542]" captionStart="Fig. 1" captionStartId="3.[174,201,1867,1886]" captionTargetBox="[171,1451,296,1840]" captionTargetId="figure@3.[171,1451,296,1840]" captionText="Fig. 1. Pentochloeno vestito Andriam., Lowry &amp; G. E. Schatz. A. Flowering branch; B. Detail of indument on lower leaf surface; C. Detail of upper leaf surface; D. Flower; E. Detail of androecium; F. Detail of gynoecium; G. Fruit." httpUri="https://zenodo.org/record/400375/files/figure.png" pageId="2" pageNumber="168">Fig. 1</figureCitation>
,
<figureCitation id="13692A37FFE9FC7C6772A291FE85053C" box="[277,293,1518,1542]" captionStart="Fig. 2" captionStartId="4.[136,163,1900,1919]" captionTargetBox="[133,1477,299,1832]" captionText="Fig. 2. Photographs of Pentochloeno vestito Andriam., Lowry &amp; G. E. Schatz. A. Flowering branch; B. Close-up of flower (note dense indument on lower surface of leaf); C. Adult plant; D. Habitat of type locality, with open wooded grassland and bushland / Tapia woodland and rocky quartzite substrate. [Photos: T. Andriamihajarivo]" httpUri="https://zenodo.org/record/400376/files/figure.png" pageId="2" pageNumber="168">2</figureCitation>
).
</paragraph>
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<subSubSection id="C3486539FFE9FC7C66C9A166FF5D05E8" pageId="2" pageNumber="168" type="materials_examined">
<paragraph id="8BED36B2FFE9FC7C66C9A166FF5D05E8" blockId="2.[136,787,1437,1746]" pageId="2" pageNumber="168">
<materialsCitation id="3B3A3CEFFFE9FC7C66C9A166FF5D05E8" ID-GBIF-Occurrence="1440667651" collectingDate="2014-11-04" collectionCode="G, P, TAN" collectorName="Andriamihajarivo et al." country="Madagascar" latitude="-20.445278" location="Ambositra" longitude="47.06167" municipality="Mania Region" pageId="2" pageNumber="168" specimenCode="MO-6684230" specimenCount="1303" specimenCount-male="1303" stateProvince="Fianarantsoa" typeStatus="holotypus">
<emphasis id="B926EAA0FFE9FC7C66C9A166FD00050B" bold="true" box="[174,672,1561,1585]" pageId="2" pageNumber="168">
<typeStatus id="54E98810FFE9FC7C66C9A166FF56050B" box="[174,246,1561,1585]" pageId="2" pageNumber="168">Typus</typeStatus>
:
<collectingCountry id="F3457622FFE9FC7C676DA166FE0C0515" box="[266,428,1561,1585]" name="Madagascar" pageId="2" pageNumber="168">Madagascar</collectingCountry>
. Prov.
<collectingRegion id="4996F850FFE9FC7C679AA166FD3A050B" box="[509,666,1561,1585]" country="Madagascar" name="Fianarantsoa" pageId="2" pageNumber="168">Fianarantsoa</collectingRegion>
:
</emphasis>
Amoroni
<collectingMunicipality id="6B89ACC8FFE9FC7C66C9A146FEF7056B" box="[174,343,1593,1617]" pageId="2" pageNumber="168">Mania Region</collectingMunicipality>
,
<location id="8E8D6069FFE9FC7C6702A146FE42056B" LSID="urn:lsid:plazi:treatment:03FB87A4FFE9FC7866EFA2B1FD870076:8E8D6069FFE9FC7C6702A146FE42056B" box="[357,482,1593,1617]" country="Madagascar" municipality="Mania Region" name="Ambositra" pageId="2" pageNumber="168" stateProvince="Fianarantsoa">Ambositra</location>
, commune rurale
<location id="8E8D6069FFE9FC7C64D2A146FDAB0548" LSID="urn:lsid:plazi:treatment:03FB87A4FFE9FC7866EFA2B1FD870076:8E8D6069FFE9FC7C64D2A146FDAB0548" country="Madagascar" municipality="Mania Region" name="Ihadilagnana, Mt. Ambatolahinanahary" pageId="2" pageNumber="168" stateProvince="Fianarantsoa">Ihadilagnana, Mt. Ambatolahinanahary</location>
,
<geoCoordinate id="EE665075FFE9FC7C6471A126FCAE054B" box="[534,782,1625,1649]" pageId="2" pageNumber="168">20̊2643S 47̊0342E</geoCoordinate>
,
<quantity id="4CAA9B57FFE9FC7C66C9A105FF5F05AB" box="[174,255,1658,1682]" metricMagnitude="3" metricUnit="m" metricValue="1.303" pageId="2" pageNumber="168" unit="m" value="1303.0">
<specimenCount id="9D54FD3BFFE9FC7C66C9A105FF5F05AB" box="[174,255,1658,1682]" pageId="2" pageNumber="168" type="male">1303 m</specimenCount>
</quantity>
,
<date id="FFEC1072FFE9FC7C676EA105FED405A8" box="[265,372,1658,1682]" pageId="2" pageNumber="168" value="2014-11-04">
<collectingDate id="EFA8E99AFFE9FC7C676EA105FED405A8" box="[265,372,1658,1682]" pageId="2" pageNumber="168" value="2014-11-04">4.XI.2014</collectingDate>
</date>
, fl.,
<emphasis id="B926EAA0FFE9FC7C67F9A105FD6305A8" box="[414,707,1658,1682]" italics="true" pageId="2" pageNumber="168">
<collectorName id="26A75364FFE9FC7C67F9A105FD2705A8" box="[414,647,1658,1682]" pageId="2" pageNumber="168">Ħndriamiðajariυo et al</collectorName>
.
<collectingDate id="EFA8E99AFFE9FC7C64E8A105FD6305A8" box="[655,707,1658,1682]" pageId="2" pageNumber="168">1907</collectingDate>
</emphasis>
(holo-:
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!; iso-:
<collectionCode id="ED43AE77FFE9FC7C67E8A1E5FE070588" box="[399,423,1690,1714]" pageId="2" pageNumber="168">G</collectionCode>
!, MO-6685745!,
<collectionCode id="ED43AE77FFE9FC7C6407A1E5FDD0058B" box="[608,624,1690,1713]" pageId="2" pageNumber="168">P</collectionCode>
[
<collectionCode id="ED43AE77FFE9FC7C641BA1E5FD2C0588" box="[636,652,1690,1714]" pageId="2" pageNumber="168">P</collectionCode>
01066023]!;
<collectionCode id="ED43AE77FFE9FC7C66C9A1C5FF4E05E8" box="[174,238,1722,1746]" collectionName="TAN" pageId="2" pageNumber="168">TAN</collectionCode>
!).
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</paragraph>
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<subSubSection id="C3486539FFE9FC7C66C9A184FACE06AA" pageId="2" pageNumber="168" type="description">
<paragraph id="8BED36B2FFE9FC7C66C9A184FA050290" blockId="2.[174,787,1787,1972]" lastBlockId="2.[862,1474,305,426]" pageId="2" pageNumber="168">
<emphasis id="B926EAA0FFE9FC7C66C9A184FA050290" italics="true" pageId="2" pageNumber="168">Haec species Pentacðlaenae latifoliae H. Perrier simillima, sed ab ea foliis majoribus (7.7­13.8 × 5.8­11.8 υs. 4.5­ 8.8 × 3.5­7 cm) υenis secundariis in paribus (10 ad) 14 ad 16 (υs. 7 ad 10) plerumque craspedodromis interdum distaliter subbrocðidodromis (υs. semper brocðidodromis), foliis abaxialiter necnon petiolis caulibus juυenilibus inflorescentiae axibus bracteis sepalis et fructu tricðomatibus stellato­strigosis c. 0.2­0.3 mm in diam. dense υestitis (υs. tricðomatibus stellato­punctatis &lt;0.1 mm in diam. sparsim modiceυe indutis) atque petalis majoribus (1.6­2.2 × 1­1.3) υs. 1.2­2.1 × 0.8­1.4 cm) distinguitur.</emphasis>
</paragraph>
<paragraph id="8BED36B2FFE9FC7C6539A6ADFACE06AA" blockId="2.[824,1475,466,1424]" pageId="2" pageNumber="168">
<emphasis id="B926EAA0FFE9FC7C6539A6ADFC3F02D0" box="[862,927,466,490]" italics="true" pageId="2" pageNumber="168">Sðrub</emphasis>
, 0.5-1.5 m tall.
<emphasis id="B926EAA0FFE9FC7C6203A6ACFB0E02D0" box="[1124,1198,467,490]" italics="true" pageId="2" pageNumber="168">Leaυes</emphasis>
ovate to oblong, rarely orbicular, coriaceous, (5.1-)10-13.8 × (4.3-)8-11.8 cm, weakly discolorous, with dense stellate-strigose trichomes c. 0.2- 0.3 mm in diam. on abaxial surFace, glabrous on adaxial surFace, base usually subcordate, rarely rounded, margin entire, apex retuse to emarginate, midvein channeled above, raised below, venation craspedodromous, occasionally approaching brochidodromous in distal 2 to 5 major secondary veins, with (10 to) 14 to 16 pairs oF secondary veins, weakly impressed above, raised below; petiole (1.3-)1.8-2.2(-2.8) cm long, 2-3 mm in diam., densely stellate-strigose, adaxially canaliculate.
<emphasis id="B926EAA0FFE9FC7C62A0A46BFAE40016" box="[1223,1348,788,812]" italics="true" pageId="2" pageNumber="168">Inflorescences</emphasis>
axillary and terminal, usually 2-flowered (occasionally 1-flowered), peduncles 1-3 mm long, densely stellate-strigose; flowers subsessile, bracts (3-)4-6(-8) × 1 mm, narrowly lanceolate, margin entire to slightly dentate, sparsely stellate-strigose; involucre scale-like, densely stellate-strigose;
<emphasis id="B926EAA0FFE9FC7C65B9A4CAFBB100F7" box="[990,1041,949,973]" italics="true" pageId="2" pageNumber="168">sepals</emphasis>
5, quincuncial, unequal in size, united at the base, 0.4-0.6(-0.8) × 0.8-1 cm, stellate-strigose on outer surFace, glabrous and with evident reticulate veins on inner surFace;
<emphasis id="B926EAA0FFE9FC7C63EAA489FA610734" box="[1421,1473,1014,1038]" italics="true" pageId="2" pageNumber="168">petals</emphasis>
5, quincuncial in bud, white, obovate, glabrous, Free, with evident actinodromous and reticulate venation, 1.6-1.7(-2.2) × 1-1.1 (-1.3) cm, base cuneate, apex truncate, extrastaminal disc 1-1.2 mm high, with a sinuate margin;
<emphasis id="B926EAA0FFE9FC7C623AA308FB0707B4" box="[1117,1191,1143,1166]" italics="true" pageId="2" pageNumber="168">stamens</emphasis>
40 to 70 or more, filaments (0.5-)0.7-0.8(-0.9) mm long, anthers ellipsoid, 0.7-1 mm long, introrse, basifixed; ovary globose, densely stellate-strigose, 2.4-2.7 × 3-3.5 mm, locules 5, style glabrous, ovules 6 per locule, 1.5- 1.6 mm long, 0.5 mm in diam., stigma capitate, light green, 5-lobed, margins oF the lobes distinctly undulate.
<emphasis id="B926EAA0FFE9FC7C62A2A267FB5F0615" box="[1221,1279,1304,1327]" italics="true" pageId="2" pageNumber="168">Fruits</emphasis>
obloid, light brown to orangish, 3 × 2-2.5 cm, flattened at the apex and the base, ridged, dehiscent, exocarp thin, densely stellate-strigose, endocarp thin,
<emphasis id="B926EAA0FFE9FC7C655FA207FCC306AA" box="[824,867,1400,1424]" italics="true" pageId="2" pageNumber="168">seeds</emphasis>
1 per locule, brown, slightly flattened, c. 1.3 cm long.
</paragraph>
</subSubSection>
<subSubSection id="C3486539FFE9FC7C6539A2C6FC08050B" pageId="2" pageNumber="168" type="etymology">
<paragraph id="8BED36B2FFE9FC7C6539A2C6FC08050B" blockId="2.[824,1474,1465,1585]" pageId="2" pageNumber="168">
<emphasis id="B926EAA0FFE9FC7C6539A2C6FC6906EB" box="[862,969,1465,1489]" italics="true" pageId="2" pageNumber="168">Etymology</emphasis>
. The epithet vestita reFers to the dense stellate strigose indument on the abaxial surFace oF the leaves as well as the petioles, young stems, inflorescence axes, bracts, sepals, and Fruits.
</paragraph>
</subSubSection>
<subSubSection id="C3486539FFE9FC7D6539A125FC7904AB" lastPageId="3" lastPageNumber="169" pageId="2" pageNumber="168" type="distribution">
<paragraph id="8BED36B2FFE9FC7C6539A125FB27048E" blockId="2.[824,1474,1626,1972]" pageId="2" pageNumber="168">
<emphasis id="B926EAA0FFE9FC7C6539A125FBF40548" box="[862,1108,1626,1650]" italics="true" pageId="2" pageNumber="168">Distribution and ecology</emphasis>
.
<emphasis id="B926EAA0FFE9FC7C6214A125FAE1054B" box="[1139,1345,1626,1650]" italics="true" pageId="2" pageNumber="168">Pentacðlaena υestita</emphasis>
is restricted to Mt. Ambatolahinanahary, located near the Mania River in the north-western part oF Ambositra district in central Madagascar, some 30 km to the south oF the Ibity massiF, where
<taxonomicName id="4C524D31FFE9FC7C65E3A1A4FC5505C9" box="[900,1013,1755,1779]" class="Magnoliopsida" family="Sarcolaenaceae" genus="Pentacdlaena" kingdom="Plantae" order="Huerteales" pageId="2" pageNumber="168" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis id="B926EAA0FFE9FC7C65E3A1A4FC5505C9" box="[900,1013,1755,1779]" italics="true" pageId="2" pageNumber="168">P. latifolia</emphasis>
</taxonomicName>
, which it most closely resembles, occurs (
<figureCitation id="13692A37FFE9FC7C6527A184FCDE0429" box="[832,894,1787,1811]" captionStart="Fig. 3" captionStartId="5.[847,874,887,906]" captionTargetBox="[846,1494,301,869]" captionTargetId="figure@5.[843,1497,298,873]" captionTargetPageId="5" captionText="Fig. 3. Geographic distribution of Pentochloeno vestito Andriam., Lowry &amp; G. E. Schatz (stars) and P. lotifolio H. Perrier (circles)." httpUri="https://zenodo.org/record/400377/files/figure.png" pageId="2" pageNumber="168">Fig. 3</figureCitation>
). It grows on quartzite substrate in an open matrix oF wooded grassland and bushland/Tapia woodland at elevations between 1250 and 1500 m.
<emphasis id="B926EAA0FFE9FC7C623FA044FA830469" box="[1112,1315,1851,1875]" italics="true" pageId="2" pageNumber="168">Pentacðlaena υestita</emphasis>
is known From only seven collections. Material in bud and flower has been gathered in October and November, and the single Fruiting collection was made in January.
</paragraph>
<caption id="DF2D663AFFE8FC7D66C9A034FCD50442" httpUri="https://zenodo.org/record/400375/files/figure.png" pageId="3" pageNumber="169" targetBox="[171,1451,296,1840]" targetPageId="3">
<paragraph id="8BED36B2FFE8FC7D66C9A034FCD50442" blockId="3.[174,1440,1867,1937]" pageId="3" pageNumber="169">
<emphasis id="B926EAA0FFE8FC7D66C9A034FF4D0464" bold="true" box="[174,237,1867,1886]" pageId="3" pageNumber="169">Fig. 1. </emphasis>
<emphasis id="B926EAA0FFE8FC7D6695A034FE0C0464" box="[242,428,1867,1886]" italics="true" pageId="3" pageNumber="169">Pentochloeno vestito</emphasis>
Andriam., Lowry &amp; G.E. Schatz.
<emphasis id="B926EAA0FFE8FC7D64A3A034FD770464" bold="true" box="[708,727,1867,1886]" pageId="3" pageNumber="169">A.</emphasis>
Flowering branch;
<emphasis id="B926EAA0FFE8FC7D65E6A034FC330464" bold="true" box="[897,915,1867,1886]" pageId="3" pageNumber="169">B.</emphasis>
Detail of indument on lower leaf surface;
<emphasis id="B926EAA0FFE8FC7D6362A034FAB80464" bold="true" box="[1285,1304,1867,1886]" pageId="3" pageNumber="169">C.</emphasis>
Detail of upper leaf surface;
<emphasis id="B926EAA0FFE8FC7D6745A01AFE960442" bold="true" box="[290,310,1893,1912]" pageId="3" pageNumber="169">D.</emphasis>
Flower;
<emphasis id="B926EAA0FFE8FC7D67E5A01AFE330442" bold="true" box="[386,403,1893,1912]" pageId="3" pageNumber="169">E.</emphasis>
Detail of androecium;
<emphasis id="B926EAA0FFE8FC7D643AA01AFDCD0442" bold="true" box="[605,621,1893,1912]" pageId="3" pageNumber="169">F.</emphasis>
Detail of gynoecium;
<emphasis id="B926EAA0FFE8FC7D6549A01AFCE10442" bold="true" box="[814,833,1893,1912]" pageId="3" pageNumber="169">G.</emphasis>
Fruit.
</paragraph>
</caption>
<paragraph id="8BED36B2FFE8FC7D66C9A001FC7904AB" blockId="3.[174,1440,1867,1937]" box="[174,985,1918,1937]" pageId="3" pageNumber="169">
<emphasis id="B926EAA0FFE8FC7D66C9A001FF7804AB" bold="true" box="[174,216,1918,1937]" pageId="3" pageNumber="169">[A-F:</emphasis>
<emphasis id="B926EAA0FFE8FC7D66BAA001FE3404AB" box="[221,404,1918,1937]" italics="true" pageId="3" pageNumber="169">Andriomihojorivo 1907</emphasis>
, G,
<emphasis id="B926EAA0FFE8FC7D67D2A001FDD304AB" box="[437,627,1918,1937]" italics="true" pageId="3" pageNumber="169">Rovololomonono et ol</emphasis>
.
<emphasis id="B926EAA0FFE8FC7D641AA001FD3404AB" box="[637,660,1918,1937]" italics="true" pageId="3" pageNumber="169">57</emphasis>
, TAN] [Drawing: R.L. Andriamiarisoa]
</paragraph>
</subSubSection>
<caption id="DF2D663AFFEFFC7A66EFA013FE280488" httpUri="https://zenodo.org/record/400376/files/figure.png" pageId="4" pageNumber="170" targetBox="[133,1477,299,1832]" targetPageId="4">
<paragraph id="8BED36B2FFEFFC7A66EFA013FE280488" blockId="4.[136,1468,1900,1970]" pageId="4" pageNumber="170">
<emphasis id="B926EAA0FFEFFC7A66EFA013FF680445" bold="true" box="[136,200,1900,1919]" pageId="4" pageNumber="170">Fig. 2. </emphasis>
Photographs of
<emphasis id="B926EAA0FFEFFC7A673DA013FDB40445" box="[346,532,1900,1919]" italics="true" pageId="4" pageNumber="170">Pentochloeno vestito</emphasis>
Andriam., Lowry &amp; G.E. Schatz.
<emphasis id="B926EAA0FFEFFC7A654BA013FC9F0445" bold="true" box="[812,831,1900,1919]" pageId="4" pageNumber="170">A.</emphasis>
Flowering branch;
<emphasis id="B926EAA0FFEFFC7A658EA013FC5B0445" bold="true" box="[1001,1019,1900,1919]" pageId="4" pageNumber="170">B.</emphasis>
Close-up of flower (note dense indument on lower surface of leaf);
<emphasis id="B926EAA0FFEFFC7A677EA0F9FE8B04A3" bold="true" box="[281,299,1926,1945]" pageId="4" pageNumber="170">C.</emphasis>
Adult plant;
<emphasis id="B926EAA0FFEFFC7A67FAA0F9FE1004A3" bold="true" box="[413,432,1926,1945]" pageId="4" pageNumber="170">D.</emphasis>
Habitat of type locality, with open wooded grassland and bushland/Tapia woodland and rocky quartzite substrate. [Photos: T. Andriamihajarivo]
</paragraph>
</caption>
<subSubSection id="C3486539FFEEFC7B66A2A64EFDBC0273" box="[197,540,305,329]" pageId="5" pageNumber="171" type="vernacular_names">
<paragraph id="8BED36B2FFEEFC7B66A2A64EFDBC0273" blockId="5.[197,540,305,329]" box="[197,540,305,329]" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B66A2A64EFECE0273" box="[197,366,305,329]" italics="true" pageId="5" pageNumber="171">Vernacular name</emphasis>
. Mananamba.
</paragraph>
</subSubSection>
<subSubSection id="C3486539FFEEFC7B66A2A60CFED20155" pageId="5" pageNumber="171" type="conservation">
<paragraph id="8BED36B2FFEEFC7B66A2A60CFED20155" blockId="5.[159,809,370,623]" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B66A2A60CFE3802B0" box="[197,408,371,394]" italics="true" pageId="5" pageNumber="171">Conserυation status</emphasis>
. With an extent oF occurrence oF &lt;100 km2, an area oF occupancy oF &lt;10 km2, and two locations with respect to the most serious threat, which is fire, along with an estimated population size oF more than 250 but probably less than 2,500 mature individuals, we assign a preliminary risk oF extinction status oF “Endangered” [EN B1a b(i,ii,iii,iv,v)+2ab(i,ii,iii,iv,v); C2a(ii)] using the
<bibRefCitation id="EFC34B43FFEEFC7B64CAA549FECE0155" author="IUCN Red List Categories" journalOrPublisher="IUCN Species Survival Commission, Gland &amp; Cambridge" pageId="5" pageNumber="171" refString="IUCN (2012). IUCN Red List Categories and Criteria: Version 3.1. 2 nd ed. IUCN Species Survival Commission, Gland &amp; Cambridge" type="book" year="2012">IUCN Red List (IUCN, 2012)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3486539FFEEFC7866A2A5E6FD9D0133" lastPageId="6" lastPageNumber="172" pageId="5" pageNumber="171" type="discussion">
<paragraph id="8BED36B2FFEEFC7B66A2A5E6FC8A008F" blockId="5.[159,810,664,1439]" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B66A2A5E6FEA2018A" box="[197,258,665,688]" italics="true" pageId="5" pageNumber="171">Notes</emphasis>
.
<emphasis id="B926EAA0FFEEFC7B674EA5E7FDB1018A" box="[297,529,664,688]" italics="true" pageId="5" pageNumber="171">Pentacðlaena υestita</emphasis>
most closely resembles
<taxonomicName id="4C524D31FFEEFC7B66F8A5C6FEA601EB" box="[159,262,697,721]" class="Magnoliopsida" family="Sarcolaenaceae" genus="Pentacdlaena" kingdom="Plantae" order="Huerteales" pageId="5" pageNumber="171" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis id="B926EAA0FFEEFC7B66F8A5C6FEA601EB" box="[159,262,697,721]" italics="true" pageId="5" pageNumber="171">P. latifolia</emphasis>
</taxonomicName>
, which also occurs in central Madagascar, primarily on and around the quartzitic Ibity MassiF some 30 km to the north (
<figureCitation id="13692A37FFEEFC7B668CA585FE8A0028" box="[235,298,762,786]" captionStart="Fig. 3" captionStartId="5.[847,874,887,906]" captionTargetBox="[846,1494,301,869]" captionTargetId="figure@5.[843,1497,298,873]" captionTargetPageId="5" captionText="Fig. 3. Geographic distribution of Pentochloeno vestito Andriam., Lowry &amp; G. E. Schatz (stars) and P. lotifolio H. Perrier (circles)." httpUri="https://zenodo.org/record/400377/files/figure.png" pageId="5" pageNumber="171">Fig. 3</figureCitation>
). These two species both have ovate to oblong, weakly discolorous leaves with a retuse to emarginate apex, axillary and terminal inflorescences with one or two flowers subtended by entire or slightly dentate bracts, and occur on quartzite substrates. Our new species can, however, be distinguished by a suite oF characters, as summarized in
<tableCitation id="C6D00309FFEEFC7B64B3A4E2FC84008F" box="[724,804,925,949]" captionStart="Table 1" captionStartId="5.[159,213,1503,1523]" captionTargetBox="[159,1497,1556,1976]" captionText="Table 1. Key characters distinguishing Pentochloeno vestito Andriam., Lowry &amp; G. E. Schatz from P. lotifolio H. Perrier" httpUri="http://table.plazi.org/id/DF2D663AFFEEFC7B66F8A2A0FB4B06C8" pageId="5" pageNumber="171" tableUuid="DF2D663AFFEEFC7B66F8A2A0FB4B06C8">Table 1</tableCitation>
.
</paragraph>
<caption id="DF2D663AFFEEFC7B6528A408FA8B0099" httpUri="https://zenodo.org/record/400377/files/figure.png" pageId="5" pageNumber="171" targetBox="[846,1494,301,869]" targetPageId="5">
<paragraph id="8BED36B2FFEEFC7B6528A408FA8B0099" blockId="5.[847,1485,887,931]" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B6528A408FC2E00B0" bold="true" box="[847,910,887,906]" pageId="5" pageNumber="171">Fig. 3. </emphasis>
Geographic distribution of
<emphasis id="B926EAA0FFEEFC7B6218A408FA9800B0" box="[1151,1336,887,906]" italics="true" pageId="5" pageNumber="171">Pentochloeno vestito</emphasis>
Andriam., Lowry &amp; G.E. Schatz (stars) and
<emphasis id="B926EAA0FFEEFC7B624AA4EFFB240099" box="[1069,1156,912,931]" italics="true" pageId="5" pageNumber="171">P. lotifolio</emphasis>
H. Perrier (circles).
</paragraph>
</caption>
<paragraph id="8BED36B2FFEEFC7B66A2A4C1FD470780" blockId="5.[159,810,664,1439]" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B66A2A4C1FE3D00EF" box="[197,413,958,982]" italics="true" pageId="5" pageNumber="171">Pentacðlaena υestita</emphasis>
, like
<taxonomicName id="4C524D31FFEEFC7B67BDA4C1FDE800EC" box="[474,584,958,982]" class="Magnoliopsida" family="Sarcolaenaceae" genus="Pentacdlaena" kingdom="Plantae" order="Huerteales" pageId="5" pageNumber="171" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis id="B926EAA0FFEEFC7B67BDA4C1FDE800EC" box="[474,584,958,982]" italics="true" pageId="5" pageNumber="171">P. latifolia</emphasis>
</taxonomicName>
, is clearly fire tolerant, as are most oF the other species that have been able to survive in Madagascars wooded grassland and bushland/Tapia woodland habitats. The increased Frequency oF fires in these areas above historical pre-human levels has likely reduced the number oF sites where individuals oF
<emphasis id="B926EAA0FFEEFC7B644FA31DFD240743" box="[552,644,1122,1145]" italics="true" pageId="5" pageNumber="171">P. υestita</emphasis>
can persist and has probably also stunted their growth, preventing them From reaching their maximum potential height (
<figureCitation id="13692A37FFEEFC7B6402A3DDFD030780" box="[613,675,1186,1210]" captionStart="Fig. 2" captionStartId="4.[136,163,1900,1919]" captionTargetBox="[133,1477,299,1832]" captionText="Fig. 2. Photographs of Pentochloeno vestito Andriam., Lowry &amp; G. E. Schatz. A. Flowering branch; B. Close-up of flower (note dense indument on lower surface of leaf); C. Adult plant; D. Habitat of type locality, with open wooded grassland and bushland / Tapia woodland and rocky quartzite substrate. [Photos: T. Andriamihajarivo]" httpUri="https://zenodo.org/record/400376/files/figure.png" pageId="5" pageNumber="171">Fig. 2</figureCitation>
C, D).
</paragraph>
<paragraph id="8BED36B2FFEEFC7866A2A3BCFD9D0133" blockId="5.[159,810,664,1439]" lastBlockId="6.[136,786,305,521]" lastPageId="6" lastPageNumber="172" pageId="5" pageNumber="171">
<bibRefCitation id="EFC34B43FFEEFC7B66A2A3BCFE0C07E1" author="Lowry II, P. P. &amp; T. Haevermans &amp; J. - N. Labat &amp; G. E. Schatz &amp; J. - F. Leroy &amp; A. - E. Wolf" box="[197,428,1219,1243]" journalOrPublisher="Hdansonia" pageId="5" pageNumber="171" pagination="11 - 31" part="22" refString="Lowry II, P. P., T. Haevermans, J. - N. Labat, G. E. Schatz, J. - F. Leroy &amp; A. - E. Wolf (2000). Endemic Families oF Madagascar. V. A synoptic revision oF Eremolaena, Pentachlaena and Perrierodendron (Sarcolaenaceae). Hdansonia ser. 3, 22: 11 - 31." title="Endemic Families oF Madagascar. V. A synoptic revision oF Eremolaena, Pentachlaena and Perrierodendron (Sarcolaenaceae)" type="journal article" year="2000">Lowry et al. (2000)</bibRefCitation>
, in their revision oF
<emphasis id="B926EAA0FFEEFC7B64F0A3BCFC8407E1" box="[663,804,1219,1243]" italics="true" pageId="5" pageNumber="171">Pentacðlaena</emphasis>
, assigned
<emphasis id="B926EAA0FFEEFC7B6764A39BFDAE07C6" box="[259,526,1251,1276]" italics="true" pageId="5" pageNumber="171">Perrier de la Bâtðie 2112</emphasis>
From the Mania River to
<taxonomicName id="4C524D31FFEEFC7B66F8A27AFEAE0626" box="[159,270,1284,1308]" class="Magnoliopsida" family="Sarcolaenaceae" genus="Pentacdlaena" kingdom="Plantae" order="Huerteales" pageId="5" pageNumber="171" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis id="B926EAA0FFEEFC7B66F8A27AFEAE0626" box="[159,270,1284,1308]" italics="true" pageId="5" pageNumber="171">P. latifolia</emphasis>
</taxonomicName>
, having overlooked the Fact that this collection exhibits the distinctive Features that characterize
<emphasis id="B926EAA0FFEEFC7B64CDA259FCA50606" box="[682,773,1317,1341]" italics="true" pageId="5" pageNumber="171">P. υestita</emphasis>
. In re-examining the holdings oF other species oF
<emphasis id="B926EAA0FFEEFC7B64F8A23AFC890667" box="[671,809,1349,1373]" italics="true" pageId="5" pageNumber="171">Pentacðlaena</emphasis>
in the Paris herbarium to compare them with the material oF our new species, we Found that another collection,
<emphasis id="B926EAA0FFEEFC7B6486A2F8FBAD00CE" box="[737,1037,988,1438]" italics="true" pageId="5" pageNumber="171">Perrier de la Bâtðie 1998,</emphasis>
made in January 1913, comprises a mixed gathering oF material clearly belonging to
<emphasis id="B926EAA0FFEEFC7B634DA482FA29072E" box="[1322,1417,1021,1044]" italics="true" pageId="5" pageNumber="171">P. υestita</emphasis>
and to
<taxonomicName id="4C524D31FFEEFC7B6528A362FC15070E" box="[847,949,1052,1076]" class="Magnoliopsida" family="Sarcolaenaceae" genus="Pentacdlaena" kingdom="Plantae" order="Huerteales" pageId="5" pageNumber="171" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis id="B926EAA0FFEEFC7B6528A362FC15070E" box="[847,949,1052,1076]" italics="true" pageId="5" pageNumber="171">P. latifolia</emphasis>
</taxonomicName>
. The label data accompanying the two specimens oF this gathering indicate that it was collected on Mt. Ibity, where
<taxonomicName id="4C524D31FFEEFC7B6528A322FC1C074E" box="[847,956,1116,1140]" class="Magnoliopsida" family="Sarcolaenaceae" genus="Pentacdlaena" kingdom="Plantae" order="Huerteales" pageId="5" pageNumber="171" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis id="B926EAA0FFEEFC7B6528A322FC1C074E" box="[847,956,1116,1140]" italics="true" pageId="5" pageNumber="171">P. latifolia</emphasis>
</taxonomicName>
is abundant, suggesting that some Fragments oF
<emphasis id="B926EAA0FFEEFC7B6528A302FC0907AE" box="[847,937,1149,1172]" italics="true" pageId="5" pageNumber="171">P. υestita</emphasis>
, almost surely belonging to
<emphasis id="B926EAA0FFEEFC7B62B3A302FA7307AE" box="[1236,1491,1148,1172]" italics="true" pageId="5" pageNumber="171">Perrier de la Bâtðie 2112</emphasis>
, were accidentally mounted along with the material oF
<emphasis id="B926EAA0FFEEFC7B63F5A3E2FBAE07EE" italics="true" pageId="5" pageNumber="171">Perrier de la Bâtðie 1998</emphasis>
. While it is not out oF the question that the two taxa were growing in sympatry at Ibity, this seems highly unlikely given that no individuals resembling
<emphasis id="B926EAA0FFEEFC7B6319A382FA79062E" box="[1406,1497,1277,1300]" italics="true" pageId="5" pageNumber="171">P. υestita</emphasis>
have ever been observed intermingled in the population oF
<taxonomicName id="4C524D31FFEEFC7B6528A242FC17066E" box="[847,951,1340,1364]" class="Magnoliopsida" family="Sarcolaenaceae" genus="Pentacdlaena" kingdom="Plantae" order="Huerteales" pageId="5" pageNumber="171" phylum="Tracheophyta" rank="species" species="latifolia">
<emphasis id="B926EAA0FFEEFC7B6528A242FC17066E" box="[847,951,1340,1364]" italics="true" pageId="5" pageNumber="171">P. latifolia</emphasis>
</taxonomicName>
and that no plants oF
<emphasis id="B926EAA0FFEEFC7B62C0A242FAA3066E" box="[1191,1283,1341,1364]" italics="true" pageId="5" pageNumber="171">P. υestita</emphasis>
have ever been collected at Ibity, which Further supports our interpretation that they represent two geographically distinct taxa. A similar case oF closely related but morphologically distinct species restricted to wooded grassland and bushland/Tapia woodland on quartzite and marble substrates in central Madagascar can be Found in the genus
<emphasis id="B926EAA0FFEDFC786714A6EEFE7B0293" box="[371,475,401,425]" italics="true" pageId="6" pageNumber="172">Pðilgamia</emphasis>
Baill.
<emphasis id="B926EAA0FFEDFC786478A6EEFD690293" box="[543,713,401,425]" italics="true" pageId="6" pageNumber="172">(Malpigðiaceae),</emphasis>
whose Four members all occur in this same type oF habitat and have clearly diversified within a limited geographic area (
<bibRefCitation id="EFC34B43FFEDFC7864D1A6AEFF1D0133" author="Arenes, J." editor="Humbert, H." journalOrPublisher="Madagascar Comores" pageId="6" pageNumber="172" pagination="108" refString="Arenes, J. (1950). Malpighiacees. In: Humbert, H. (ed.), Fl. Madagascar Comores 108." title="Malpighiacees" type="book chapter" volumeTitle="Fl" year="1950">Arènes, 1950</bibRefCitation>
;
<bibRefCitation id="EFC34B43FFEDFC7866AAA68EFD910133" author="Madagascar Catalogue" box="[205,561,497,521]" journalOrPublisher="Missouri Botanical Garden, St. Louis &amp; Antananarivo [www. tropicos. org / project / mada]" pageId="6" pageNumber="172" refString="Madagascar Catalogue (2016). Catalogue of tde Vascular Plants of Madagascar. Missouri Botanical Garden, St. Louis &amp; Antananarivo [www. tropicos. org / project / mada]." title="Catalogue of tde Vascular Plants of Madagascar" type="book" year="2016">Madagascar Catalogue, 2016</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF2D663AFFEEFC7B66F8A2A0FB4B06C8" ID-Table-UUID="DF2D663AFFEEFC7B66F8A2A0FB4B06C8" box="[159,1259,1502,1523]" httpUri="http://table.plazi.org/id/DF2D663AFFEEFC7B66F8A2A0FB4B06C8" pageId="5" pageNumber="171" targetBox="[159,1497,1556,1976]" targetIsTable="true" targetPageId="5">
<paragraph id="8BED36B2FFEEFC7B66F8A2A0FB4B06C8" blockId="5.[159,1259,1502,1523]" box="[159,1259,1502,1523]" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B66F8A2A0FF5C06C9" bold="true" box="[159,252,1503,1523]" pageId="5" pageNumber="171">Table 1. </emphasis>
Key characters distinguishing
<emphasis id="B926EAA0FFEEFC7B647CA2A1FD4006C8" box="[539,736,1502,1522]" italics="true" pageId="5" pageNumber="171">Pentochloeno vestito</emphasis>
Andriam., Lowry &amp; G.E. Schatz from
<emphasis id="B926EAA0FFEEFC7B6253A2A1FB2E06C8" box="[1076,1166,1502,1522]" italics="true" pageId="5" pageNumber="171">P. lotifolio</emphasis>
H. Perrier
</paragraph>
</caption>
<paragraph id="8BED36B2FFEEFC7B6478A16BFB72048D" pageId="5" pageNumber="171">
<table id="F952C412FFEE038166F8A16BFA790482" box="[159,1497,1556,1976]" gridcols="3" gridrows="16" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B6478A16BFEEE0579" bold="true" pageId="5" pageNumber="171">
<tr id="356234F0FFEE038166F8A16BFA790512" box="[159,1497,1556,1576]" gridrow="0" pageId="5" pageNumber="171" rowspan-0="1">
<emphasis id="B926EAA0FFEEFC7B6478A16BFB220512" bold="true" box="[543,1154,1556,1576]" italics="true" pageId="5" pageNumber="171">
<th id="76B35D8CFFEE03816478A16BFC510512" box="[543,1009,1556,1576]" gridcol="1" gridrow="0" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B6478A16BFDD90512" bold="true" box="[543,633,1556,1576]" italics="true" pageId="5" pageNumber="171">P. vestitơ</emphasis>
</th>
<th id="76B35D8CFFEE03816246A16BFA790512" box="[1057,1497,1556,1576]" gridcol="2" gridrow="0" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B6246A16BFB220512" bold="true" box="[1057,1154,1556,1576]" italics="true" pageId="5" pageNumber="171">P. lơtifoliơ</emphasis>
</th>
</emphasis>
</tr>
<tr id="356234F0FFEE038166F8A150FA790579" box="[159,1497,1583,1603]" gridrow="1" pageId="5" pageNumber="171" rowspan-1="1" rowspan-2="1">
<th id="76B35D8CFFEE038166F8A150FE450579" box="[159,485,1583,1603]" gridcol="0" gridrow="1" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B66F8A150FEEE0579" bold="true" box="[159,334,1583,1603]" pageId="5" pageNumber="171">Largest leaf blade</emphasis>
</th>
</tr>
</emphasis>
<tr id="356234F0FFEE038166F8A136FA790567" box="[159,1497,1609,1629]" gridrow="2" pageId="5" pageNumber="171">
<th id="76B35D8CFFEE038166F8A136FE450567" box="[159,485,1609,1629]" gridcol="0" gridrow="2" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B66D2A136FE840567" bold="true" box="[181,292,1609,1629]" pageId="5" pageNumber="171">length [cm]</emphasis>
</th>
<td id="76B35D8CFFEE03816478A136FC510567" box="[543,1009,1609,1629]" gridcol="1" gridrow="2" pageId="5" pageNumber="171">(7.7-)9.5-13.8</td>
<td id="76B35D8CFFEE03816246A136FA790567" box="[1057,1497,1609,1629]" gridcol="2" gridrow="2" pageId="5" pageNumber="171">4.5-7.5(-8.8)</td>
</tr>
<tr id="356234F0FFEE038166F8A11CFA790542" box="[159,1497,1635,1656]" gridrow="3" pageId="5" pageNumber="171">
<th id="76B35D8CFFEE038166F8A11CFE450542" box="[159,485,1635,1656]" gridcol="0" gridrow="3" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B66D2A11BFEBD0542" bold="true" box="[181,285,1636,1656]" pageId="5" pageNumber="171">width [cm]</emphasis>
</th>
<td id="76B35D8CFFEE03816478A11CFC510542" box="[543,1009,1635,1656]" gridcol="1" gridrow="3" pageId="5" pageNumber="171">(5.8-)6.5-11.8</td>
<td id="76B35D8CFFEE03816246A11CFA790542" box="[1057,1497,1635,1656]" gridcol="2" gridrow="3" pageId="5" pageNumber="171">3.5-5.7(-7)</td>
</tr>
<tr id="356234F0FFEE038166F8A101FA7905A9" box="[159,1497,1662,1683]" gridrow="4" pageId="5" pageNumber="171">
<th id="76B35D8CFFEE038166F8A101FE4505A9" box="[159,485,1662,1683]" gridcol="0" gridrow="4" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B66C1A100FE4505A9" bold="true" box="[166,485,1663,1683]" pageId="5" pageNumber="171">Indument on abaxial leaf surface</emphasis>
</th>
<td id="76B35D8CFFEE03816478A101FC5105A9" box="[543,1009,1662,1683]" gridcol="1" gridrow="4" pageId="5" pageNumber="171">Densely stellate-strigose, trichomes c. 0.2-</td>
<td id="76B35D8CFFEE03816246A101FA7905A9" box="[1057,1497,1662,1683]" gridcol="2" gridrow="4" pageId="5" pageNumber="171">Sparsely stellate-punctate, trichomes &lt;0.1 mm</td>
</tr>
<tr id="356234F0FFEE038166F8A1E6FA790597" box="[159,1497,1689,1709]" gridrow="5" pageId="5" pageNumber="171" rowspan-0="1">
<td id="76B35D8CFFEE03816478A1E6FC510597" box="[543,1009,1689,1709]" gridcol="1" gridrow="5" pageId="5" pageNumber="171">0.3 mm in diam., on all veins (incl. tertiary),</td>
<td id="76B35D8CFFEE03816246A1E6FA790597" box="[1057,1497,1689,1709]" gridcol="2" gridrow="5" pageId="5" pageNumber="171">in diam., along midvein and major secondary</td>
</tr>
<tr id="356234F0FFEE038166F8A1CCFA7905FD" box="[159,1497,1715,1735]" gridrow="6" pageId="5" pageNumber="171" rowspan-0="1">
<td id="76B35D8CFFEE03816478A1CCFC5105FD" box="[543,1009,1715,1735]" gridcol="1" gridrow="6" pageId="5" pageNumber="171">almost completely obscuring leaf surface</td>
<td id="76B35D8CFFEE03816246A1CCFA7905FD" box="[1057,1497,1715,1735]" gridcol="2" gridrow="6" pageId="5" pageNumber="171">veins, occasionally on inter-secondary veins,</td>
</tr>
<tr id="356234F0FFEE038166F8A1B1FA7905D8" box="[159,1497,1742,1762]" gridrow="7" pageId="5" pageNumber="171" rowspan-0="1" rowspan-1="1">
<td id="76B35D8CFFEE03816246A1B1FA7905D8" box="[1057,1497,1742,1762]" gridcol="2" gridrow="7" pageId="5" pageNumber="171">not obscuring leaf surface</td>
</tr>
<tr id="356234F0FFEE038166F8A196FA7905C7" box="[159,1497,1769,1789]" gridrow="8" pageId="5" pageNumber="171">
<th id="76B35D8CFFEE038166F8A196FE4505C7" box="[159,485,1769,1789]" gridcol="0" gridrow="8" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B66C1A196FE8A05C7" bold="true" box="[166,298,1769,1789]" pageId="5" pageNumber="171">Leaf venation</emphasis>
</th>
<td id="76B35D8CFFEE03816478A196FC5105C7" box="[543,1009,1769,1789]" gridcol="1" gridrow="8" pageId="5" pageNumber="171">Craspedodromous, occasionally approaching</td>
<td id="76B35D8CFFEE03816246A196FA7905C7" box="[1057,1497,1769,1789]" gridcol="2" gridrow="8" pageId="5" pageNumber="171">Brochidodromous</td>
</tr>
<tr id="356234F0FFEE038166F8A07CFA79042D" box="[159,1497,1795,1815]" gridrow="9" pageId="5" pageNumber="171" rowspan-0="1" rowspan-2="1">
<td id="76B35D8CFFEE03816478A07CFC51042D" box="[543,1009,1795,1815]" gridcol="1" gridrow="9" pageId="5" pageNumber="171">brochidodromous in distal 2 to 5 major secondary</td>
</tr>
<tr id="356234F0FFEE038166F8A061FA790408" box="[159,1497,1822,1842]" gridrow="10" pageId="5" pageNumber="171" rowspan-0="1" rowspan-2="1">
<td id="76B35D8CFFEE03816478A061FC510408" box="[543,1009,1822,1842]" gridcol="1" gridrow="10" pageId="5" pageNumber="171">veins</td>
</tr>
<tr id="356234F0FFEE038166F8A046FA790477" box="[159,1497,1849,1869]" gridrow="11" pageId="5" pageNumber="171">
<th id="76B35D8CFFEE038166F8A046FE450477" box="[159,485,1849,1869]" gridcol="0" gridrow="11" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B66C1A046FE680477" bold="true" box="[166,456,1849,1869]" pageId="5" pageNumber="171">Number of pairs of secondary</emphasis>
</th>
<td id="76B35D8CFFEE03816478A046FC510477" box="[543,1009,1849,1869]" gridcol="1" gridrow="11" pageId="5" pageNumber="171">(10-)14-16</td>
<td id="76B35D8CFFEE03816246A046FA790477" box="[1057,1497,1849,1869]" gridcol="2" gridrow="11" pageId="5" pageNumber="171">7-9(-10)</td>
</tr>
<tr id="356234F0FFEE038166F8A02BFA790452" box="[159,1497,1876,1896]" gridrow="12" pageId="5" pageNumber="171" rowspan-1="1" rowspan-2="1">
<th id="76B35D8CFFEE038166F8A02BFE450452" box="[159,485,1876,1896]" gridcol="0" gridrow="12" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B66C1A02BFE8A0452" bold="true" box="[166,298,1876,1896]" pageId="5" pageNumber="171">veins per leaf</emphasis>
</th>
</tr>
<tr id="356234F0FFEE038166F8A011FA7904B9" box="[159,1497,1902,1923]" gridrow="13" pageId="5" pageNumber="171">
<th id="76B35D8CFFEE038166F8A011FE4504B9" box="[159,485,1902,1923]" gridcol="0" gridrow="13" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B66C1A010FE6D04B9" bold="true" box="[166,461,1903,1923]" pageId="5" pageNumber="171">Indument on twigs and petiole</emphasis>
s
</th>
<td id="76B35D8CFFEE03816478A011FC5104B9" box="[543,1009,1902,1923]" gridcol="1" gridrow="13" pageId="5" pageNumber="171">Densely stellate-strigose, almost totally</td>
<td id="76B35D8CFFEE03816246A011FA7904B9" box="[1057,1497,1902,1923]" gridcol="2" gridrow="13" pageId="5" pageNumber="171">Sparsely to moderately densely stellate-</td>
</tr>
<tr id="356234F0FFEE038166F8A0F6FA7904A7" box="[159,1497,1929,1949]" gridrow="14" pageId="5" pageNumber="171" rowspan-0="1">
<td id="76B35D8CFFEE03816478A0F6FC5104A7" box="[543,1009,1929,1949]" gridcol="1" gridrow="14" pageId="5" pageNumber="171">obscuring surface</td>
<td id="76B35D8CFFEE03816246A0F6FA7904A7" box="[1057,1497,1929,1949]" gridcol="2" gridrow="14" pageId="5" pageNumber="171">punctate, covering &lt;50% of the surface</td>
</tr>
<tr id="356234F0FFEE038166F8A0DCFA790482" box="[159,1497,1955,1976]" gridrow="15" pageId="5" pageNumber="171">
<th id="76B35D8CFFEE038166F8A0DCFE450482" box="[159,485,1955,1976]" gridcol="0" gridrow="15" pageId="5" pageNumber="171">
<emphasis id="B926EAA0FFEEFC7B66C1A0DBFE950482" bold="true" box="[166,309,1956,1976]" pageId="5" pageNumber="171">Petal size [cm]</emphasis>
</th>
<td id="76B35D8CFFEE03816478A0DCFC510482" box="[543,1009,1955,1976]" gridcol="1" gridrow="15" pageId="5" pageNumber="171">1.6-1.7(-2.2) X 1-1.1(-1.3)</td>
<td id="76B35D8CFFEE03816246A0DCFA790482" box="[1057,1497,1955,1976]" gridcol="2" gridrow="15" pageId="5" pageNumber="171">1.2(-2.1) X 0.8(-1.4)</td>
</tr>
</table>
</paragraph>
<subSubSection id="C3486539FFEDFC7866C9A550FD870076" pageId="6" pageNumber="172" type="materials_examined">
<paragraph id="8BED36B2FFEDFC7866C9A550FD870076" blockId="6.[136,787,558,845]" pageId="6" pageNumber="172">
<emphasis id="B926EAA0FFEDFC7866C9A550FF52017B" box="[174,242,559,577]" italics="true" pageId="6" pageNumber="172">Paratypi</emphasis>
.
<emphasis id="B926EAA0FFEDFC78676DA550FD9D017B" bold="true" box="[266,573,558,578]" pageId="6" pageNumber="172">Madagascar. Prov. Fianarantsoa:</emphasis>
Région Amoroni Mania, Ambositra, Comm. Ihadilagnana, à lW de Volondoha, 20̊2549S 47̊0449E, 1254 m, 19.I.2005, Fr.,
<emphasis id="B926EAA0FFEDFC786730A51BFDBC014D" box="[343,540,612,631]" italics="true" pageId="6" pageNumber="172">Ħndriamiðajariυo et al</emphasis>
.
<emphasis id="B926EAA0FFEDFC786440A51BFDE8014D" box="[551,584,612,631]" italics="true" pageId="6" pageNumber="172">566</emphasis>
(MO, TAN); Ihadilagnana, W dAmbositra, Mt. Ambatolahinanahary, 20̊2605S 47̊0338E, 1491 m, 25.III.2006, bud,
<emphasis id="B926EAA0FFEDFC786738A5E6FDB10196" box="[351,529,665,684]" italics="true" pageId="6" pageNumber="172">Ħndriamiðajariυo 874</emphasis>
(TAN);
<emphasis id="B926EAA0FFEDFC78643BA5E6FD010196" box="[604,673,665,684]" italics="true" pageId="6" pageNumber="172">ibid. loc.,</emphasis>
same date, fl.,
<emphasis id="B926EAA0FFEDFC7866EFA5CBFEE901FD" box="[136,329,692,711]" italics="true" pageId="6" pageNumber="172">Ħndriamiðajariυo 875</emphasis>
(TAN); Ambohipiandrianana-Ambohibary, Ihadilagnana, Ambositra, 20̊2643S 47̊0342E, 1303 m, 04.XI.2014, bud &amp; fl.,
<emphasis id="B926EAA0FFEDFC7866EFA596FEE301C6" box="[136,323,745,764]" italics="true" pageId="6" pageNumber="172">Ħndriamiðajariυo et al</emphasis>
.
<emphasis id="B926EAA0FFEDFC78672AA596FED801C6" box="[333,376,745,764]" italics="true" pageId="6" pageNumber="172">1908</emphasis>
(G, MO, P, TAN); sur la Mania, bois à Tapia, c. 1200 m, VI.1912, fl.,
<emphasis id="B926EAA0FFEDFC786725A47AFDB3002D" box="[322,531,772,791]" italics="true" pageId="6" pageNumber="172">Perrier de la Bâtðie 2112</emphasis>
(P [3 sheets]); Ihadilagnana, massiF dAmbatolahinanahary, 20̊2626S 47̊0331E, 1346 m, 20.I.2006, bud, fl.,
<emphasis id="B926EAA0FFEDFC7866AAA446FEDE0076" box="[205,382,825,844]" italics="true" pageId="6" pageNumber="172">Raυololomanana et al</emphasis>
.
<emphasis id="B926EAA0FFEDFC7867EFA445FE3D0077" box="[392,413,826,845]" italics="true" pageId="6" pageNumber="172">57</emphasis>
(MO, P, TAN).
</paragraph>
</subSubSection>
</treatment>
</document>