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<document id="202D40ED284C99C808FFACE4DDAC50EA" ID-DOI="10.11646/zootaxa.3947.3.9" ID-GBIF-Dataset="d5fc6687-0be9-46f7-9f7c-43c383106b31" ID-ISSN="1175-5326" ID-Zenodo-Dep="238969" ID-ZooBank="28501A93-1BBD-4133-A7A3-D6A740121A6E" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461041693295" checkinUser="plazi" docAuthor="Meini, Gianluca" docDate="2015" docId="03BC87B5FFD7FFFEFF23FC68FD3C60EF" docLanguage="en" docName="zt03947p439.pdf" docOrigin="Zootaxa 3947 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Orthoplana lunae Meini, 2015, sp. nov." docType="treatment" docVersion="9" lastPageNumber="433" masterDocId="FF85FFCDFFD5FFF6FFB4FFDAFFF46327" masterDocTitle="Two new Otoplanid species (Platyhelminthes: Rhabditophora: Proseriata) of the genera Orthoplana Steinböck, 1932 and Postbursoplana Ax, 1956 from the Tuscan coast (Italy)" masterLastPageNumber="439" masterPageNumber="425" pageNumber="427" updateTime="1698575742563" updateUser="plazi">
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<mods:title id="53CFFC6984F6B8908BA1B8D26616A823">Two new Otoplanid species (Platyhelminthes: Rhabditophora: Proseriata) of the genera Orthoplana Steinböck, 1932 and Postbursoplana Ax, 1956 from the Tuscan coast (Italy)</mods:title>
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<mods:namePart id="0D1EAE651FE1EE1F005ED246913093CC">Meini, Gianluca</mods:namePart>
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<mods:date id="618668B574CC6D08DC98E65BB3917C0B">2015</mods:date>
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<treatment id="03BC87B5FFD7FFFEFF23FC68FD3C60EF" ID-DOI="http://doi.org/10.5281/zenodo.5624445" ID-GBIF-Taxon="119609855" ID-Zenodo-Dep="5624445" LSID="urn:lsid:plazi:treatment:03BC87B5FFD7FFFEFF23FC68FD3C60EF" httpUri="http://treatment.plazi.org/id/03BC87B5FFD7FFFEFF23FC68FD3C60EF" lastPageId="8" lastPageNumber="433" pageId="2" pageNumber="427">
<subSubSection id="C30F6528FFD7FFF4FF23FC68FEED60C9" pageId="2" pageNumber="427" type="nomenclature">
<paragraph id="8BAA36A3FFD7FFF4FF23FC68FE2760EA" blockId="2.[151,467,946,1006]" box="[151,467,946,973]" pageId="2" pageNumber="427">
<heading id="D0E281CFFFD7FFF4FF23FC68FE2760EA" bold="true" box="[151,467,946,973]" fontSize="11" level="1" pageId="2" pageNumber="427" reason="1">
<emphasis id="B961EAB1FFD7FFF4FF23FC68FE2760EA" bold="true" box="[151,467,946,973]" pageId="2" pageNumber="427">
<taxonomicName id="4C154D20FFD7FFF4FF23FC68FE9A60EB" box="[151,366,946,972]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="2" pageNumber="427" phylum="Platyhelminthes" rank="species" species="lunae" status="sp. nov.">
<emphasis id="B961EAB1FFD7FFF4FF23FC68FE9A60EB" bold="true" box="[151,366,946,972]" italics="true" pageId="2" pageNumber="427">Orthoplana lunae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257CAFFD7FFF4FEC2FC69FE2760EA" box="[374,467,947,973]" pageId="2" pageNumber="427" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAA36A3FFD7FFF4FF23FC0FFEED60C9" blockId="2.[151,467,946,1006]" box="[151,281,981,1006]" pageId="2" pageNumber="427">
(
<figureCitation id="132E2A26FFD7FFF4FF2BFC0FFEE460C9" box="[159,272,981,1006]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURES 3 5" captionStart-3="FIGURES 6 7" captionStart-4="FIGURE 8" captionStart-5="FIGURES 9 10" captionStartId-0="3.[151,250,1953,1975]" captionStartId-1="4.[151,250,1136,1158]" captionStartId-2="5.[151,264,1993,2016]" captionStartId-3="6.[151,264,1953,1976]" captionStartId-4="7.[151,250,1020,1042]" captionStartId-5="7.[151,264,1952,1975]" captionTargetBox-0="[371,1254,210,1903]" captionTargetBox-2="[176,1418,688,1992]" captionTargetBox-3="[397,1183,201,1920]" captionTargetBox-5="[197,1375,1104,1914]" captionTargetId-0="figure@3.[302,1284,193,1932]" captionTargetId-2="figure@5.[151,1436,679,1996]" captionTargetId-3="figure@6.[391,1195,193,1932]" captionTargetId-5="figure@7.[151,1436,1036,1966]" captionTargetPageId-0="3" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-5="7" captionText-0="FIGURE 1. General organi z ation of Orthoplana lunae sp. nov. (a) and superficial dorsal view with the general arrangement of the rhabdites on the body surface (b)." captionText-1="FIGURE 2. Photograph of Orthoplana lunae sp. nov. in vivo." captionText-2="FIGURES 3 5. Photographs of Orthoplana lunae sp. nov., in vivo: anterior end (3, 4) and posterior end (5)." captionText-3="FIGURES 6 7. Photographs of Orthoplana lunae sp. nov., in vivo: pharyngeal zone (6) and post-pharyngeal zone (7)." captionText-4="FIGURE 8. Spines of male sclerotic apparatus of Orthoplana lunae sp. nov." captionText-5="FIGURES 9 10. Reconstruction from camera-lucida drawings of the morphology (9) and spatial distribution (10) of the spines of the male sclerotic apparatus of Orthoplana lunae sp. nov." httpUri-0="https://zenodo.org/record/238970/files/figure.png" httpUri-2="https://zenodo.org/record/238971/files/figure.png" httpUri-3="https://zenodo.org/record/238972/files/figure.png" httpUri-5="https://zenodo.org/record/238973/files/figure.png" pageId="2" pageNumber="427">Figs 110</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F6528FFD7FFF4FF23FBC7FC1E6629" pageId="2" pageNumber="427" type="materials_examined">
<paragraph id="8BAA36A3FFD7FFF4FF23FBC7FBDF677D" blockId="2.[151,1436,1053,1978]" pageId="2" pageNumber="427">
<emphasis id="B961EAB1FFD7FFF4FF23FBC7FEF06711" bold="true" box="[151,260,1053,1078]" pageId="2" pageNumber="427">
<typeStatus id="54AE8801FFD7FFF4FF23FBC7FEF06711" box="[151,260,1053,1078]" pageId="2" pageNumber="427" type="holotype">Holotype</typeStatus>
</emphasis>
. A sagittally-sectioned specimen (EMLC-UP 160) deposited in the Electron Microscopy Laboratory Collection of the Dipartimento di Biologia, Unità di Etologia (Università di Pisa).
</paragraph>
<paragraph id="8BAA36A3FFD7FFF4FF73FBBFFC906785" blockId="2.[151,1436,1053,1978]" pageId="2" pageNumber="427">
<emphasis id="B961EAB1FFD7FFF4FF73FBBFFEA96759" bold="true" box="[199,349,1125,1150]" pageId="2" pageNumber="427">
<typeStatus id="54AE8801FFD7FFF4FF73FBBFFEF66759" box="[199,258,1125,1150]" pageId="2" pageNumber="427">Type</typeStatus>
locality
</emphasis>
.
<collectingCountry id="F3027633FFD7FFF4FED8FBBFFE546759" box="[364,416,1125,1150]" name="Italy" pageId="2" pageNumber="427">Italy</collectingCountry>
, Tuscany, Ligurian Sea: loc. Partaccia (
<taxonomicName id="4C154D20FFD7FFF4FCEEFBBFFC596759" box="[858,941,1125,1150]" class="Magnoliopsida" family="Fabaceae" genus="Marina" kingdom="Plantae" order="Fabales" pageId="2" pageNumber="438" phylum="Tracheophyta" rank="genus">Marina</taxonomicName>
di Massa, Massa) (
<geoCoordinate id="EE215064FFD7FFF4FB33FBBCFAEC6759" box="[1159,1304,1126,1150]" direction="north" orientation="latitude" pageId="2" pageNumber="427" precision="15" value="44.023335">44°0124N</geoCoordinate>
,
<geoCoordinate id="EE215064FFD7FFF4FA97FBBFFF586785" direction="east" orientation="longitude" pageId="2" pageNumber="427" precision="15" value="10.071111">10°0416 E</geoCoordinate>
). Surf-zone characterized by fine sand. Coll.
<date id="FFAB1063FFD7FFF4FD12FB53FC956785" box="[678,865,1161,1186]" pageId="2" pageNumber="427" value="2008-09">September 2008</date>
.
</paragraph>
<paragraph id="8BAA36A3FFD7FFF4FF73FB77FC1E6629" blockId="2.[151,1436,1053,1978]" pageId="2" pageNumber="427">
<emphasis id="B961EAB1FFD7FFF4FF73FB77FE4967E1" bold="true" box="[199,445,1197,1222]" pageId="2" pageNumber="427">Additional material.</emphasis>
At least
<specimenCount id="9D13FD2AFFD7FFF4FD9FFB74FD3267E1" box="[555,710,1197,1222]" pageId="2" pageNumber="427" type="generic">30 specimens</specimenCount>
were studied
<emphasis id="B961EAB1FFD7FFF4FCD8FB74FC4E67E2" box="[876,954,1198,1221]" italics="true" pageId="2" pageNumber="427">in vivo</emphasis>
, including drawings and photographs, all from the
<typeStatus id="54AE8801FFD7FFF4FEBCFB08FECC67CD" box="[264,312,1234,1258]" pageId="2" pageNumber="427">type</typeStatus>
locality.
<typeStatus id="54AE8801FFD7FFF4FE1EFB08FDE367CD" box="[426,535,1234,1258]" pageId="2" pageNumber="427" type="paratype">Paratypes</typeStatus>
(EMLC-UP 160-A,B,C,D): four specimens from the
<typeStatus id="54AE8801FFD7FFF4FB20FB08FB3067CD" box="[1172,1220,1234,1258]" pageId="2" pageNumber="427">type</typeStatus>
locality, sagittally sectioned. Three whole mounts (EMLC-UP 160-E,F) from the
<typeStatus id="54AE8801FFD7FFF4FCEDFB2CFC7D6629" box="[857,905,1270,1294]" pageId="2" pageNumber="427">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C30F6528FFD7FFF2FF73FAC3FB3F655B" lastPageId="4" lastPageNumber="429" pageId="2" pageNumber="427" type="description">
<paragraph id="8BAA36A3FFD7FFF4FF73FAC3FAB66551" blockId="2.[151,1436,1053,1978]" pageId="2" pageNumber="427">
<emphasis id="B961EAB1FFD7FFF4FF73FAC3FEAD6615" bold="true" box="[199,345,1305,1330]" pageId="2" pageNumber="427">Description.</emphasis>
The length of sexually mature animals, alive and in extension, reaches
<quantity id="4CED9B46FFD7FFF4FB35FAC3FAFF6616" box="[1153,1291,1305,1330]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="1.6" metricValueMin="1.4" pageId="2" pageNumber="427" unit="mm" value="1.5" valueMax="1.6" valueMin="1.4">1.41.6 mm</quantity>
. The body is fusiform, dorsally convex, ventrally flat, colourless and transparent (
<figureCitation id="132E2A26FFD7FFF4FC27FAE7FC2E6671" box="[915,986,1341,1366]" captionStart="FIGURE 1" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[371,1254,210,1903]" captionTargetId="figure@3.[302,1284,193,1932]" captionTargetPageId="3" captionText="FIGURE 1. General organi z ation of Orthoplana lunae sp. nov. (a) and superficial dorsal view with the general arrangement of the rhabdites on the body surface (b)." httpUri="https://zenodo.org/record/238970/files/figure.png" pageId="2" pageNumber="427">Figs 1</figureCitation>
,
<figureCitation id="132E2A26FFD7FFF4FC53FAE4FC0C6671" box="[999,1016,1342,1366]" captionStart="FIGURE 2" captionStartId="4.[151,250,1136,1158]" captionText="FIGURE 2. Photograph of Orthoplana lunae sp. nov. in vivo." pageId="2" pageNumber="427">2</figureCitation>
). The anterior end is characterized by a Köpfchen (
<bibRefCitation id="EF844B52FFD7FFF4FE88FAB8FE55665D" author="Ax" box="[316,417,1377,1402]" pageId="2" pageNumber="427" refString="Ax, P. (1956) Monographie der Otoplanidae (Turbellaria): Morphologie und Systematik. Akademie der Wissenschaften und der Literatur, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse, 13, 159 - 278." type="journal article" year="1956">Ax 1956</bibRefCitation>
) or cephalic swelling with numerous tactile hairs (th) and followed by two lateral couples of Tastborsten (
<bibRefCitation id="EF844B52FFD7FFF4FEEDFA5CFE4966B9" author="Ax" box="[345,445,1413,1438]" pageId="2" pageNumber="427" refString="Ax, P. (1956) Monographie der Otoplanidae (Turbellaria): Morphologie und Systematik. Akademie der Wissenschaften und der Literatur, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse, 13, 159 - 278." type="journal article" year="1956">Ax 1956</bibRefCitation>
) or tactile bristles (tb) retractable into the respective wide pockets (
<figureCitation id="132E2A26FFD7FFF4FB0EFA5FFAF666B9" box="[1210,1282,1413,1438]" captionStart="FIGURE 1" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[371,1254,210,1903]" captionTargetId="figure@3.[302,1284,193,1932]" captionTargetPageId="3" captionText="FIGURE 1. General organi z ation of Orthoplana lunae sp. nov. (a) and superficial dorsal view with the general arrangement of the rhabdites on the body surface (b)." httpUri="https://zenodo.org/record/238970/files/figure.png" pageId="2" pageNumber="427">Figs 1</figureCitation>
,
<figureCitation id="132E2A26FFD7FFF4FAA5FA5CFAD566B9" box="[1297,1313,1414,1438]" captionStart="FIGURE 2" captionStartId="4.[151,250,1136,1158]" captionText="FIGURE 2. Photograph of Orthoplana lunae sp. nov. in vivo." pageId="2" pageNumber="427">2</figureCitation>
,
<figureCitation id="132E2A26FFD7FFF4FA9AFA5CFAAF66B9" box="[1326,1371,1414,1438]" captionStart="FIGURES 3 5" captionStartId="5.[151,264,1993,2016]" captionTargetBox="[176,1418,688,1992]" captionTargetId="figure@5.[151,1436,679,1996]" captionTargetPageId="5" captionText="FIGURES 3 5. Photographs of Orthoplana lunae sp. nov., in vivo: anterior end (3, 4) and posterior end (5)." httpUri="https://zenodo.org/record/238971/files/figure.png" pageId="2" pageNumber="427">3, 4</figureCitation>
). The anterior pair of these bristles is thinner and shorter than the posterior pair, which is characterized by a greater length and thickness. The rhabdites (r), a variety of secretions in epidermal cells or in sub-epidermal glands, are present grouped in longitudinal lines along the body areas without cilia, with the exception of the anterior end, where they are scattered with a low concentration (
<figureCitation id="132E2A26FFD7FFF4FDDAF9CFFD406509" box="[622,692,1557,1582]" captionStart="FIGURE 1" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[371,1254,210,1903]" captionTargetId="figure@3.[302,1284,193,1932]" captionTargetPageId="3" captionText="FIGURE 1. General organi z ation of Orthoplana lunae sp. nov. (a) and superficial dorsal view with the general arrangement of the rhabdites on the body surface (b)." httpUri="https://zenodo.org/record/238970/files/figure.png" pageId="2" pageNumber="427">Fig. 1</figureCitation>
). The anterior restricted intestine protuberance, called cephalic intestine (ci), with the statocyst (sta) and the ovoid brain (b) are located behind the cephalic swelling. The ovoid brain (b) is lengthened and provided with two
<emphasis id="B961EAB1FFD7FFF4FD23F987FD546551" bold="true" box="[663,672,1629,1654]" pageId="2" pageNumber="427">-</emphasis>
three pairs of lateral rounded protuberances (
<figureCitation id="132E2A26FFD7FFF4FB23F987FB296551" box="[1175,1245,1629,1654]" captionStart="FIGURE 1" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[371,1254,210,1903]" captionTargetId="figure@3.[302,1284,193,1932]" captionTargetPageId="3" captionText="FIGURE 1. General organi z ation of Orthoplana lunae sp. nov. (a) and superficial dorsal view with the general arrangement of the rhabdites on the body surface (b)." httpUri="https://zenodo.org/record/238970/files/figure.png" pageId="2" pageNumber="427">Figs 1</figureCitation>
,
<figureCitation id="132E2A26FFD7FFF4FB5FF984FB0F6551" box="[1259,1275,1630,1654]" captionStart="FIGURE 2" captionStartId="4.[151,250,1136,1158]" captionText="FIGURE 2. Photograph of Orthoplana lunae sp. nov. in vivo." pageId="2" pageNumber="427">2</figureCitation>
,
<figureCitation id="132E2A26FFD7FFF4FAB3F984FAC06551" box="[1287,1332,1630,1654]" captionStart="FIGURES 3 5" captionStartId="5.[151,264,1993,2016]" captionTargetBox="[176,1418,688,1992]" captionTargetId="figure@5.[151,1436,679,1996]" captionTargetPageId="5" captionText="FIGURES 3 5. Photographs of Orthoplana lunae sp. nov., in vivo: anterior end (3, 4) and posterior end (5)." httpUri="https://zenodo.org/record/238971/files/figure.png" pageId="2" pageNumber="427">3, 4</figureCitation>
).
</paragraph>
<paragraph id="8BAA36A3FFD7FFF4FF73F95BFC026599" blockId="2.[151,1436,1053,1978]" pageId="2" pageNumber="427">The testes (te), situated behind the brain and anterior to the pharynx, consist of two close series of medium-size follicles arranged along the longitudinal axis, 1113 per side, in a single line.</paragraph>
<paragraph id="8BAA36A3FFD7FFF4FF73F910FC326421" blockId="2.[151,1436,1053,1978]" pageId="2" pageNumber="427">
Two longitudinal rows of small vitellaries (vi) stretch externally to the testes from just behind the brain to the penis papilla opening, uniformly distributed in single units (
<figureCitation id="132E2A26FFD7FFF4FC82F937FC896421" box="[822,893,1773,1798]" captionStart="FIGURE 1" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[371,1254,210,1903]" captionTargetId="figure@3.[302,1284,193,1932]" captionTargetPageId="3" captionText="FIGURE 1. General organi z ation of Orthoplana lunae sp. nov. (a) and superficial dorsal view with the general arrangement of the rhabdites on the body surface (b)." httpUri="https://zenodo.org/record/238970/files/figure.png" pageId="2" pageNumber="427">Figs 1</figureCitation>
,
<figureCitation id="132E2A26FFD7FFF4FC3FF934FC6F6421" box="[907,923,1774,1798]" captionStart="FIGURE 2" captionStartId="4.[151,250,1136,1158]" captionText="FIGURE 2. Photograph of Orthoplana lunae sp. nov. in vivo." pageId="2" pageNumber="427">2</figureCitation>
,
<figureCitation id="132E2A26FFD7FFF4FC13F934FC436421" box="[935,951,1774,1798]" captionStart="FIGURES 3 5" captionStartId="5.[151,264,1993,2016]" captionTargetBox="[176,1418,688,1992]" captionTargetId="figure@5.[151,1436,679,1996]" captionTargetPageId="5" captionText="FIGURES 3 5. Photographs of Orthoplana lunae sp. nov., in vivo: anterior end (3, 4) and posterior end (5)." httpUri="https://zenodo.org/record/238971/files/figure.png" pageId="2" pageNumber="427">3</figureCitation>
).
</paragraph>
<paragraph id="8BAA36A3FFD7FFF4FF73F8C8FBB56469" blockId="2.[151,1436,1053,1978]" pageId="2" pageNumber="427">
Two globoid germaries (ge), at some distance from each other, lie in front of the pharynx, posterior to the last testis follicles; they have small dimensions and contain numerous egg cells (
<figureCitation id="132E2A26FFD7FFF4FC5AF8EFFBDB6469" box="[1006,1071,1845,1870]" captionStart="FIGURE 1" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[371,1254,210,1903]" captionTargetId="figure@3.[302,1284,193,1932]" captionTargetPageId="3" captionText="FIGURE 1. General organi z ation of Orthoplana lunae sp. nov. (a) and superficial dorsal view with the general arrangement of the rhabdites on the body surface (b)." httpUri="https://zenodo.org/record/238970/files/figure.png" pageId="2" pageNumber="427">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BAA36A3FFD7FFF4FF73F883FBA7649D" blockId="2.[151,1436,1053,1978]" pageId="2" pageNumber="427">
The pharynx (ph), situated in the rear half of the body, exhibits the so-called bell-shaped organization or Glöckchen (
<bibRefCitation id="EF844B52FFD7FFF4FE87F8A4FE5464B1" author="Ax" box="[307,416,1917,1942]" pageId="2" pageNumber="427" refString="Ax, P. (1956) Monographie der Otoplanidae (Turbellaria): Morphologie und Systematik. Akademie der Wissenschaften und der Literatur, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse, 13, 159 - 278." type="journal article" year="1956">Ax, 1956</bibRefCitation>
), typical of the genus. The sacciform intestine (i) is a caecum at both ends and the anterior restricted protuberance or cephalic intestine (ci) is constituted by turgid cells (
<figureCitation id="132E2A26FFD7FFF4FBB4F87BFBB6649D" box="[1024,1090,1953,1978]" captionStart="FIGURE 1" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[371,1254,210,1903]" captionTargetId="figure@3.[302,1284,193,1932]" captionTargetPageId="3" captionText="FIGURE 1. General organi z ation of Orthoplana lunae sp. nov. (a) and superficial dorsal view with the general arrangement of the rhabdites on the body surface (b)." httpUri="https://zenodo.org/record/238970/files/figure.png" pageId="2" pageNumber="427">Fig. 1</figureCitation>
).
</paragraph>
<caption id="DF6A662BFFD6FFF5FF23F87BFDE564F1" httpUri="https://zenodo.org/record/238970/files/figure.png" pageId="3" pageNumber="428" targetBox="[371,1254,210,1903]" targetPageId="3">
<paragraph id="8BAA36A3FFD6FFF5FF23F87BFDE564F1" blockId="3.[151,1435,1953,2006]" pageId="3" pageNumber="428">
<emphasis id="B961EAB1FFD6FFF5FF23F87BFEE66491" bold="true" box="[151,274,1953,1975]" pageId="3" pageNumber="428">FIGURE 1.</emphasis>
General organi
<emphasis id="B961EAB1FFD6FFF5FE1BF878FE4E6490" bold="true" box="[431,442,1954,1975]" pageId="3" pageNumber="428">z</emphasis>
ation of
<taxonomicName id="4C154D20FFD6FFF5FDBAF87BFD366490" box="[526,706,1953,1975]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="3" pageNumber="428" phylum="Platyhelminthes" rank="species" species="lunae" status="sp. nov.">
<emphasis id="B961EAB1FFD6FFF5FDBAF87BFD366490" box="[526,706,1953,1975]" italics="true" pageId="3" pageNumber="428">Orthoplana lunae</emphasis>
</taxonomicName>
<emphasis id="B961EAB1FFD6FFF5FD7DF878FCED6490" bold="true" box="[713,793,1954,1975]" pageId="3" pageNumber="428">
<taxonomicNameLabel id="A25257CAFFD6FFF5FD7DF878FCED6490" box="[713,793,1954,1975]" pageId="3" pageNumber="428" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(a) and superficial dorsal view with the general arrangement of the rhabdites on the body surface (b).
</paragraph>
</caption>
<caption id="DF6A662BFFD1FFF2FF23FBAAFCF867A1" box="[151,780,1136,1158]" pageId="4" pageNumber="429">
<paragraph id="8BAA36A3FFD1FFF2FF23FBAAFCF867A1" blockId="4.[151,780,1136,1158]" box="[151,780,1136,1158]" pageId="4" pageNumber="429">
<emphasis id="B961EAB1FFD1FFF2FF23FBAAFEE667A2" bold="true" box="[151,274,1136,1158]" pageId="4" pageNumber="429">FIGURE 2.</emphasis>
Photograph of
<taxonomicName id="4C154D20FFD1FFF2FE1BFBAAFD9067A1" box="[431,612,1136,1158]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="4" pageNumber="429" phylum="Platyhelminthes" rank="species" species="lunae" status="sp. nov.">
<emphasis id="B961EAB1FFD1FFF2FE1BFBAAFD9067A1" box="[431,612,1136,1158]" italics="true" pageId="4" pageNumber="429">Orthoplana lunae</emphasis>
</taxonomicName>
<emphasis id="B961EAB1FFD1FFF2FDDEFBAAFD4F67A2" bold="true" box="[618,699,1136,1157]" pageId="4" pageNumber="429">
<taxonomicNameLabel id="A25257CAFFD1FFF2FDDEFBAAFD4F67A2" box="[618,699,1136,1157]" pageId="4" pageNumber="429" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B961EAB1FFD1FFF2FD76FBABFCF267A1" box="[706,774,1137,1158]" italics="true" pageId="4" pageNumber="429">in vivo</emphasis>
.
</paragraph>
</caption>
<paragraph id="8BAA36A3FFD1FFF2FF73FB69FC96667C" blockId="4.[151,1436,1203,1372]" pageId="4" pageNumber="429">
The accessory male pore (amp) is clearly visible laterally to the penis papilla and can be observed in living and semi-squashed specimens (
<figureCitation id="132E2A26FFD1FFF2FE7AFB0CFDED67C8" box="[462,537,1238,1263]" captionStart="FIGURE 1" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[371,1254,210,1903]" captionTargetId="figure@3.[302,1284,193,1932]" captionTargetPageId="3" captionText="FIGURE 1. General organi z ation of Orthoplana lunae sp. nov. (a) and superficial dorsal view with the general arrangement of the rhabdites on the body surface (b)." httpUri="https://zenodo.org/record/238970/files/figure.png" pageId="4" pageNumber="429">Figs 1</figureCitation>
,
<figureCitation id="132E2A26FFD1FFF2FD9FFB0DFDC867C8" box="[555,572,1239,1263]" captionStart="FIGURES 6 7" captionStartId="6.[151,264,1953,1976]" captionTargetBox="[397,1183,201,1920]" captionTargetId="figure@6.[391,1195,193,1932]" captionTargetPageId="6" captionText="FIGURES 6 7. Photographs of Orthoplana lunae sp. nov., in vivo: pharyngeal zone (6) and post-pharyngeal zone (7)." httpUri="https://zenodo.org/record/238972/files/figure.png" pageId="4" pageNumber="429">7</figureCitation>
). The caudal end appears sharpened and narrow, provided with numerous adhesive papillae (ap) (
<figureCitation id="132E2A26FFD1FFF2FE2EFB21FE2B6634" box="[410,479,1275,1300]" captionStart="FIGURE 1" captionStartId="3.[151,250,1953,1975]" captionTargetBox="[371,1254,210,1903]" captionTargetId="figure@3.[302,1284,193,1932]" captionTargetPageId="3" captionText="FIGURE 1. General organi z ation of Orthoplana lunae sp. nov. (a) and superficial dorsal view with the general arrangement of the rhabdites on the body surface (b)." httpUri="https://zenodo.org/record/238970/files/figure.png" pageId="4" pageNumber="429">Figs 1</figureCitation>
,
<figureCitation id="132E2A26FFD1FFF2FE59FB21FE096634" box="[493,509,1275,1299]" captionStart="FIGURE 2" captionStartId="4.[151,250,1136,1158]" captionText="FIGURE 2. Photograph of Orthoplana lunae sp. nov. in vivo." pageId="4" pageNumber="429">2</figureCitation>
,
<figureCitation id="132E2A26FFD1FFF2FDBCFB21FDEC6633" box="[520,536,1275,1300]" captionStart="FIGURES 3 5" captionStartId="5.[151,264,1993,2016]" captionTargetBox="[176,1418,688,1992]" captionTargetId="figure@5.[151,1436,679,1996]" captionTargetPageId="5" captionText="FIGURES 3 5. Photographs of Orthoplana lunae sp. nov., in vivo: anterior end (3, 4) and posterior end (5)." httpUri="https://zenodo.org/record/238971/files/figure.png" pageId="4" pageNumber="429">5</figureCitation>
). In the post-pharyngeal zone, the sacciform
<emphasis id="B961EAB1FFD1FFF2FBB9FB21FB2D6634" box="[1037,1241,1275,1299]" italics="true" pageId="4" pageNumber="429">vesicula seminalis</emphasis>
(vs) is connected distally with a well developed
<emphasis id="B961EAB1FFD1FFF2FE49FAC5FD1D6610" box="[509,745,1311,1335]" italics="true" pageId="4" pageNumber="429">vesicula granulorum</emphasis>
(vg). The male sclerotic apparatus (s) consists of a median stylet and 19 spines of variable shape and length (
<figureCitation id="132E2A26FFD1FFF2FD70FA99FCF8667C" box="[708,780,1347,1372]" captionStart="FIGURE 8" captionStartId="7.[151,250,1020,1042]" captionText="FIGURE 8. Spines of male sclerotic apparatus of Orthoplana lunae sp. nov." pageId="4" pageNumber="429">Figs 8</figureCitation>
,
<figureCitation id="132E2A26FFD1FFF2FCADFA99FCA1667C" box="[793,853,1347,1371]" captionStart="FIGURES 9 10" captionStartId="7.[151,264,1952,1975]" captionTargetBox="[197,1375,1104,1914]" captionTargetId="figure@7.[151,1436,1036,1966]" captionTargetPageId="7" captionText="FIGURES 9 10. Reconstruction from camera-lucida drawings of the morphology (9) and spatial distribution (10) of the spines of the male sclerotic apparatus of Orthoplana lunae sp. nov." httpUri="https://zenodo.org/record/238973/files/figure.png" pageId="4" pageNumber="429">9, 10</figureCitation>
):
</paragraph>
<paragraph id="8BAA36A3FFD1FFF2FF23FA56FC7F66E0" blockId="4.[151,1437,1419,1660]" pageId="4" pageNumber="429">
• a “Trichterrohr” (T) (according to
<bibRefCitation id="EF844B52FFD1FFF2FDD2FA51FD216683" author="Ax" box="[614,725,1419,1444]" pageId="4" pageNumber="429" refString="Ax, P. (1956) Monographie der Otoplanidae (Turbellaria): Morphologie und Systematik. Akademie der Wissenschaften und der Literatur, Abhandlungen der Mathematisch-Naturwissenschaftlichen Klasse, 13, 159 - 278." type="journal article" year="1956">Ax, 1956</bibRefCitation>
) or median stylet, 4849 Μm long, centrally located in the sclerotized complex, club-shaped with pointed tip at both ends;
</paragraph>
<paragraph id="8BAA36A3FFD1FFF2FF23FA0EFDE16528" blockId="4.[151,1437,1419,1660]" pageId="4" pageNumber="429">• 1 pair of straight spines, 44 Μm long, arranged at both sides of the median stylet, with a very slight subterminal prominence and a pointed tip.</paragraph>
<paragraph id="8BAA36A3FFD1FFF2FF23F9C6FB3F655B" blockId="4.[151,1437,1419,1660]" pageId="4" pageNumber="429">• 17 spines, placed on both sides of the previous ones, practically straight, with pointed tips weakly bent outwards and small subterminal cuneiform prominence. Their length becomes progressively reduced starting from a maximum of 40 Μm, to a minimum of 17 Μm. The proximal end is narrow and blunt.</paragraph>
</subSubSection>
<subSubSection id="C30F6528FFD1FFFEFF73F971FA6761D7" lastPageId="8" lastPageNumber="433" pageId="4" pageNumber="429" type="discussion">
<paragraph id="8BAA36A3FFD1FFF2FF73F971FE48642C" blockId="4.[151,1436,1707,2020]" pageId="4" pageNumber="429">
<emphasis id="B961EAB1FFD1FFF2FF73F971FECF65E3" bold="true" box="[199,315,1707,1732]" pageId="4" pageNumber="429">Remarks.</emphasis>
<taxonomicName id="4C154D20FFD1FFF2FEF1F971FDE465E4" box="[325,528,1707,1731]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="4" pageNumber="429" phylum="Platyhelminthes" rank="species" species="lunae" status="sp. nov.">
<emphasis id="B961EAB1FFD1FFF2FEF1F971FDE465E4" box="[325,528,1707,1731]" italics="true" pageId="4" pageNumber="429">Orthoplana lunae</emphasis>
</taxonomicName>
<emphasis id="B961EAB1FFD1FFF2FDAEF971FD8265E3" bold="true" box="[538,630,1707,1732]" pageId="4" pageNumber="429">
<taxonomicNameLabel id="A25257CAFFD1FFF2FDAEF971FD8265E3" box="[538,630,1707,1732]" pageId="4" pageNumber="429" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, is of a similar body length to that observed in
<taxonomicName id="4C154D20FFD1FFF2FB1EF971FAEB65E4" box="[1194,1311,1707,1731]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="4" pageNumber="429" phylum="Platyhelminthes" rank="species" species="borealis">
<emphasis id="B961EAB1FFD1FFF2FB1EF971FAEB65E4" box="[1194,1311,1707,1731]" italics="true" pageId="4" pageNumber="429">
<collectingCountry id="F3027633FFD1FFF2FB1EF971FB4A65E4" box="[1194,1214,1707,1731]" name="Iceland" pageId="4" pageNumber="429">O</collectingCountry>
.borealis
</emphasis>
</taxonomicName>
(
<quantity id="4CED9B46FFD1FFF2FA86F971FA6465E4" box="[1330,1424,1707,1732]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="429" unit="mm" value="1.5">1.5 mm</quantity>
), shorter than in
<taxonomicName id="4C154D20FFD1FFF2FE8BF915FE3C65C0" box="[319,456,1743,1767]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="4" pageNumber="429" phylum="Platyhelminthes" rank="species" species="bregazzii">
<emphasis id="B961EAB1FFD1FFF2FE8BF915FE3C65C0" box="[319,456,1743,1767]" italics="true" pageId="4" pageNumber="429">
<collectingCountry id="F3027633FFD1FFF2FE8BF915FEA265C1" box="[319,342,1743,1766]" name="Iceland" pageId="4" pageNumber="429">O</collectingCountry>
. bregazzii
</emphasis>
</taxonomicName>
(
<quantity id="4CED9B46FFD1FFF2FE62F914FDCD65C0" box="[470,569,1742,1767]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="4" pageNumber="429" unit="mm" value="4.5" valueMax="5.0" valueMin="4.0">45 mm</quantity>
),
<taxonomicName id="4C154D20FFD1FFF2FDF8F915FCF165C0" box="[588,773,1743,1767]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="4" pageNumber="429" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="B961EAB1FFD1FFF2FDF8F915FCF165C0" box="[588,773,1743,1767]" italics="true" pageId="4" pageNumber="429">
<collectingCountry id="F3027633FFD1FFF2FDF8F915FD9765C1" box="[588,611,1743,1766]" name="Iceland" pageId="4" pageNumber="429">O</collectingCountry>
. mediterranea
</emphasis>
</taxonomicName>
(
<quantity id="4CED9B46FFD1FFF2FCA0F914FC8065C0" box="[788,884,1742,1767]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="4" pageNumber="429" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
) and
<taxonomicName id="4C154D20FFD1FFF2FC05F915FBA865C0" box="[945,1116,1743,1767]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="4" pageNumber="429" phylum="Platyhelminthes" rank="species" species="sewardensis">
<emphasis id="B961EAB1FFD1FFF2FC05F915FBA865C0" box="[945,1116,1743,1767]" italics="true" pageId="4" pageNumber="429">
<collectingCountry id="F3027633FFD1FFF2FC05F915FC3C65C1" box="[945,968,1743,1766]" name="Iceland" pageId="4" pageNumber="429">O</collectingCountry>
. sewardensis
</emphasis>
</taxonomicName>
(
<quantity id="4CED9B46FFD1FFF2FBDFF914FB1665C0" box="[1131,1250,1742,1767]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="3.0" metricValueMin="1.5" pageId="4" pageNumber="429" unit="mm" value="2.25" valueMax="3.0" valueMin="1.5">1.53 mm</quantity>
), and larger than in
<taxonomicName id="4C154D20FFD1FFF2FF00F929FEE0642C" box="[180,276,1779,1803]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="4" pageNumber="429" phylum="Platyhelminthes" rank="species" species="kohni">
<emphasis id="B961EAB1FFD1FFF2FF00F929FEE0642C" box="[180,276,1779,1803]" italics="true" pageId="4" pageNumber="429">
<collectingCountry id="F3027633FFD1FFF2FF00F929FF3F642D" box="[180,203,1779,1802]" name="Iceland" pageId="4" pageNumber="429">O</collectingCountry>
. kohni
</emphasis>
</taxonomicName>
(
<quantity id="4CED9B46FFD1FFF2FE97F929FE5B642C" box="[291,431,1779,1803]" metricMagnitude="-3" metricUnit="m" metricValue="1.2999999999999998" metricValueMax="1.4" metricValueMin="1.2" pageId="4" pageNumber="429" unit="mm" value="1.2999999999999998" valueMax="1.4" valueMin="1.2">1.21.4 mm</quantity>
).
</paragraph>
<paragraph id="8BAA36A3FFD1FFF2FF73F8CCFE5D6498" blockId="4.[151,1436,1707,2020]" pageId="4" pageNumber="429">
Starting from the upper end of the testis rows, the body of the new species shows a constant width until the level of germaries, located in front of the pharynx in the initial part of the caudal body half, as in
<taxonomicName id="4C154D20FFD1FFF2FB45F8E1FA686474" box="[1265,1436,1851,1875]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="4" pageNumber="429" phylum="Platyhelminthes" rank="species" species="sewardensis">
<emphasis id="B961EAB1FFD1FFF2FB45F8E1FA686474" box="[1265,1436,1851,1875]" italics="true" pageId="4" pageNumber="429">
<collectingCountry id="F3027633FFD1FFF2FB45F8E1FAFC6475" box="[1265,1288,1851,1874]" name="Iceland" pageId="4" pageNumber="429">O</collectingCountry>
. sewardensis
</emphasis>
</taxonomicName>
and
<taxonomicName id="4C154D20FFD1FFF2FF7DF885FE716450" box="[201,389,1887,1911]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="4" pageNumber="429" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="B961EAB1FFD1FFF2FF7DF885FE716450" box="[201,389,1887,1911]" italics="true" pageId="4" pageNumber="429">
<collectingCountry id="F3027633FFD1FFF2FF7DF885FF146451" box="[201,224,1887,1910]" name="Iceland" pageId="4" pageNumber="429">O</collectingCountry>
. mediterranea
</emphasis>
</taxonomicName>
. The shape of the small cephalic swelling differs from previously described species of the genus: in
<taxonomicName id="4C154D20FFD1FFF2FEB1F859FE9364BC" box="[261,359,1923,1947]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="4" pageNumber="429" phylum="Platyhelminthes" rank="species" species="lunae">
<emphasis id="B961EAB1FFD1FFF2FEB1F859FE9364BC" box="[261,359,1923,1947]" italics="true" pageId="4" pageNumber="429">
<collectingCountry id="F3027633FFD1FFF2FEB1F859FEE864BD" box="[261,284,1923,1946]" name="Iceland" pageId="4" pageNumber="429">O</collectingCountry>
. lunae
</emphasis>
</taxonomicName>
the Köpfchen is ovoid and slightly rounded, while in the other species of the genus the small head is clearly flattened.
</paragraph>
<paragraph id="8BAA36A3FFD1FFF3FF73F811FCB0611A" blockId="4.[151,1436,1707,2020]" lastBlockId="5.[151,1436,151,645]" lastPageId="5" lastPageNumber="430" pageId="4" pageNumber="429">
The distribution and the conspicuous size of the tactile bristles correspond to that described in
<taxonomicName id="4C154D20FFD1FFF2FB57F811FA6864C4" box="[1251,1436,1995,2019]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="4" pageNumber="429" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="B961EAB1FFD1FFF2FB57F811FA6864C4" box="[1251,1436,1995,2019]" italics="true" pageId="4" pageNumber="429">
<collectingCountry id="F3027633FFD1FFF2FB57F811FB0E64C5" box="[1251,1274,1995,2018]" name="Iceland" pageId="4" pageNumber="429">O</collectingCountry>
. mediterranea
</emphasis>
</taxonomicName>
and
<taxonomicName id="4C154D20FFD0FFF3FF7DFF42FEDF6397" box="[201,299,152,176]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="5" pageNumber="430" phylum="Platyhelminthes" rank="species" species="kohni">
<emphasis id="B961EAB1FFD0FFF3FF7DFF42FEDF6397" box="[201,299,152,176]" italics="true" pageId="5" pageNumber="430">
<collectingCountry id="F3027633FFD0FFF3FF7DFF42FF146388" box="[201,224,152,175]" name="Iceland" pageId="5" pageNumber="430">O</collectingCountry>
. kohni
</emphasis>
</taxonomicName>
, while in
<taxonomicName id="4C154D20FFD0FFF3FE16FF42FDBB6397" box="[418,591,152,176]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="5" pageNumber="430" phylum="Platyhelminthes" rank="species" species="sewardensis">
<emphasis id="B961EAB1FFD0FFF3FE16FF42FDBB6397" box="[418,591,152,176]" italics="true" pageId="5" pageNumber="430">
<collectingCountry id="F3027633FFD0FFF3FE16FF42FE4D6388" box="[418,441,152,175]" name="Iceland" pageId="5" pageNumber="430">O</collectingCountry>
. sewardensis
</emphasis>
</taxonomicName>
only one pair is visible on the sides of the cephalic swelling. Unlike any other species of the genus, the brain of the new species shows lateral rounded protuberances. The yolk follicle path, arranged in a single longitudinal row from the anterior end to the penis papilla opening on each side of the body, as well as the medium-small follicle dimensions, are similar to that observed in
<taxonomicName id="4C154D20FFD0FFF3FBB6FEDEFB48623B" box="[1026,1212,260,284]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="5" pageNumber="430" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="B961EAB1FFD0FFF3FBB6FEDEFB48623B" box="[1026,1212,260,284]" italics="true" pageId="5" pageNumber="430">
<collectingCountry id="F3027633FFD0FFF3FBB6FEDEFBED623C" box="[1026,1049,260,283]" name="Iceland" pageId="5" pageNumber="430">O</collectingCountry>
. mediterranea
</emphasis>
</taxonomicName>
. In
<taxonomicName id="4C154D20FFD0FFF3FB5EFEDEFA61623B" box="[1258,1429,260,284]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="5" pageNumber="430" phylum="Platyhelminthes" rank="species" species="sewardensis">
<emphasis id="B961EAB1FFD0FFF3FB5EFEDEFA61623B" box="[1258,1429,260,284]" italics="true" pageId="5" pageNumber="430">
<collectingCountry id="F3027633FFD0FFF3FB5EFEDEFAF5623C" box="[1258,1281,260,283]" name="Iceland" pageId="5" pageNumber="430">O</collectingCountry>
. sewardensis
</emphasis>
</taxonomicName>
, the vitellaries are generally similar to those of the new species, but they are different in their distribution and the presence of gaps between every single follicle. The localization of the testes is generally comparable to that observed in two of the species of the genus, but there is a dissimilarity in their distribution: in
<taxonomicName id="4C154D20FFD0FFF3FB54FEAAFA6862AF" box="[1248,1436,368,392]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="5" pageNumber="430" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="B961EAB1FFD0FFF3FB54FEAAFA6862AF" box="[1248,1436,368,392]" italics="true" pageId="5" pageNumber="430">
<collectingCountry id="F3027633FFD0FFF3FB54FEAAFB0362A0" box="[1248,1271,368,391]" name="Iceland" pageId="5" pageNumber="430">O</collectingCountry>
. mediterranea
</emphasis>
</taxonomicName>
there are many more follicles, that reach the level of the upper vitellaries; they are smaller and have gaps between the follicles. In
<taxonomicName id="4C154D20FFD0FFF3FEFEFE62FE0262F7" box="[330,502,440,464]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="5" pageNumber="430" phylum="Platyhelminthes" rank="species" species="sewardensis">
<emphasis id="B961EAB1FFD0FFF3FEFEFE62FE0262F7" box="[330,502,440,464]" italics="true" pageId="5" pageNumber="430">
<collectingCountry id="F3027633FFD0FFF3FEFEFE62FE9562E8" box="[330,353,440,463]" name="Iceland" pageId="5" pageNumber="430">O</collectingCountry>
. sewardensis
</emphasis>
</taxonomicName>
, the testes have larger dimensions and the presence of gaps between each follicle; which are rounded and few in number. The position of the paired germaries is similar to that observed in
<taxonomicName id="4C154D20FFD0FFF3FA35FE06FEDA613F" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="5" pageNumber="430" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="B961EAB1FFD0FFF3FA35FE06FEDA613F" italics="true" pageId="5" pageNumber="430">
<collectingCountry id="F3027633FFD0FFF3FA35FE06FA6C62D4" box="[1409,1432,476,499]" name="Iceland" pageId="5" pageNumber="430">O</collectingCountry>
. mediterranea
</emphasis>
</taxonomicName>
and
<taxonomicName id="4C154D20FFD0FFF3FED7FDDAFDF9613F" box="[355,525,512,536]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="5" pageNumber="430" phylum="Platyhelminthes" rank="species" species="sewardensis">
<emphasis id="B961EAB1FFD0FFF3FED7FDDAFDF9613F" box="[355,525,512,536]" italics="true" pageId="5" pageNumber="430">
<collectingCountry id="F3027633FFD0FFF3FED7FDDAFE8E6130" box="[355,378,512,535]" name="Iceland" pageId="5" pageNumber="430">O</collectingCountry>
. sewardensis
</emphasis>
</taxonomicName>
, although their dimensions are smaller. In
<taxonomicName id="4C154D20FFD0FFF3FC5DFDDAFBBD613F" box="[1001,1097,512,536]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="5" pageNumber="430" phylum="Platyhelminthes" rank="species" species="kohni">
<emphasis id="B961EAB1FFD0FFF3FC5DFDDAFBBD613F" box="[1001,1097,512,536]" italics="true" pageId="5" pageNumber="430">
<collectingCountry id="F3027633FFD0FFF3FC5DFDDAFBF46130" box="[1001,1024,512,535]" name="Iceland" pageId="5" pageNumber="430">O</collectingCountry>
. kohni
</emphasis>
</taxonomicName>
, the germaries are much more anterior than the pharynx, and they have a larger dimensions.
</paragraph>
<paragraph id="8BAA36A3FFD0FFF3FF73FD9DFC2861A2" blockId="5.[151,1436,151,645]" pageId="5" pageNumber="430">
The sacciform intestine with cephalic protuberance, the pharynx, the position of
<emphasis id="B961EAB1FFD0FFF3FBD6FD92FAC46147" box="[1122,1328,584,608]" italics="true" pageId="5" pageNumber="430">vesicula seminalis</emphasis>
,
<emphasis id="B961EAB1FFD0FFF3FAF4FD92FEE961A3" italics="true" pageId="5" pageNumber="430">vesicula granulorum</emphasis>
and
<emphasis id="B961EAB1FFD0FFF3FEE7FDB7FE1061A3" box="[339,484,620,644]" italics="true" pageId="5" pageNumber="430">penis papilla</emphasis>
coincide with the other species of the genus.
</paragraph>
<caption id="DF6A662BFFD0FFF3FF23F813FB1664C7" box="[151,1250,1993,2016]" httpUri="https://zenodo.org/record/238971/files/figure.png" pageId="5" pageNumber="430" targetBox="[176,1418,688,1992]" targetPageId="5">
<paragraph id="8BAA36A3FFD0FFF3FF23F813FB1664C7" blockId="5.[151,1250,1993,2016]" box="[151,1250,1993,2016]" pageId="5" pageNumber="430">
<emphasis id="B961EAB1FFD0FFF3FF23F813FECD64F8" bold="true" box="[151,313,1993,2016]" pageId="5" pageNumber="430">FIGURES 35.</emphasis>
Photographs of
<taxonomicName id="4C154D20FFD0FFF3FE54F810FD6164C7" box="[480,661,1994,2016]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="5" pageNumber="430" phylum="Platyhelminthes" rank="species" species="lunae" status="sp. nov.">
<emphasis id="B961EAB1FFD0FFF3FE54F810FD6164C7" box="[480,661,1994,2016]" italics="true" pageId="5" pageNumber="430">Orthoplana lunae</emphasis>
</taxonomicName>
<emphasis id="B961EAB1FFD0FFF3FD28F810FD1864F8" bold="true" box="[668,748,1994,2015]" pageId="5" pageNumber="430">
<taxonomicNameLabel id="A25257CAFFD0FFF3FD28F810FD1864F8" box="[668,748,1994,2015]" pageId="5" pageNumber="430" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B961EAB1FFD0FFF3FD4DF811FCC964C7" box="[761,829,1995,2016]" italics="true" pageId="5" pageNumber="430">in vivo</emphasis>
: anterior end (3, 4) and posterior end (5).
</paragraph>
</caption>
<caption id="DF6A662BFFD3FFF0FF23F87BFABB6490" box="[151,1359,1953,1976]" httpUri="https://zenodo.org/record/238972/files/figure.png" pageId="6" pageNumber="431" targetBox="[397,1183,201,1920]" targetPageId="6">
<paragraph id="8BAA36A3FFD3FFF0FF23F87BFABB6490" blockId="6.[151,1359,1953,1976]" box="[151,1359,1953,1976]" pageId="6" pageNumber="431">
<emphasis id="B961EAB1FFD3FFF0FF23F87BFECE6490" bold="true" box="[151,314,1953,1976]" pageId="6" pageNumber="431">FIGURES 67.</emphasis>
Photographs of
<emphasis id="B961EAB1FFD3FFF0FE54F87BFD056490" bold="true" box="[480,753,1953,1976]" pageId="6" pageNumber="431">
<taxonomicName id="4C154D20FFD3FFF0FE54F87BFD6E649F" box="[480,666,1953,1976]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="6" pageNumber="431" phylum="Platyhelminthes" rank="species" species="lunae" status="sp. nov.">
<emphasis id="B961EAB1FFD3FFF0FE54F87BFD6E649F" bold="true" box="[480,666,1953,1976]" italics="true" pageId="6" pageNumber="431">Orthoplana lunae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257CAFFD3FFF0FD15F878FD056490" box="[673,753,1954,1975]" pageId="6" pageNumber="431" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<emphasis id="B961EAB1FFD3FFF0FD4BF878FCB76490" box="[767,835,1954,1975]" italics="true" pageId="6" pageNumber="431">in vivo</emphasis>
: pharyngeal zone (6) and post-pharyngeal zone (7).
</paragraph>
</caption>
<caption id="DF6A662BFFD2FFF1FF23FC26FC546735" box="[151,928,1020,1042]" pageId="7" pageNumber="432">
<paragraph id="8BAA36A3FFD2FFF1FF23FC26FC546735" blockId="7.[151,928,1020,1042]" box="[151,928,1020,1042]" pageId="7" pageNumber="432">
<emphasis id="B961EAB1FFD2FFF1FF23FC26FEE66736" bold="true" box="[151,274,1020,1042]" pageId="7" pageNumber="432">FIGURE 8.</emphasis>
Spines of male sclerotic apparatus of
<taxonomicName id="4C154D20FFD2FFF1FD20FC26FCBD6735" box="[660,841,1020,1042]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="7" pageNumber="432" phylum="Platyhelminthes" rank="species" species="lunae" status="sp. nov.">
<emphasis id="B961EAB1FFD2FFF1FD20FC26FCBD6735" box="[660,841,1020,1042]" italics="true" pageId="7" pageNumber="432">Orthoplana lunae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257CAFFD2FFF1FCFBFC27FC546735" box="[847,928,1021,1042]" pageId="7" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<caption id="DF6A662BFFD2FFF1FF23F87AFD0D64F2" httpUri="https://zenodo.org/record/238973/files/figure.png" pageId="7" pageNumber="432" targetBox="[197,1375,1104,1914]" targetPageId="7">
<paragraph id="8BAA36A3FFD2FFF1FF23F87AFD0D64F2" blockId="7.[151,1435,1952,2006]" pageId="7" pageNumber="432">
<emphasis id="B961EAB1FFD2FFF1FF23F87AFEB26491" bold="true" box="[151,326,1952,1975]" pageId="7" pageNumber="432">FIGURES 910.</emphasis>
Reconstruction from camera-lucida drawings of the morphology (9) and spatial distribution (10) of the spines of the male sclerotic apparatus of
<taxonomicName id="4C154D20FFD2FFF1FE5AF81AFD5664F1" box="[494,674,1984,2006]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="7" pageNumber="432" phylum="Platyhelminthes" rank="species" species="lunae" status="sp. nov.">
<emphasis id="B961EAB1FFD2FFF1FE5AF81AFD5664F1" box="[494,674,1984,2006]" italics="true" pageId="7" pageNumber="432">Orthoplana lunae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257CAFFD2FFF1FD1DF81AFD0D64F2" box="[681,761,1984,2005]" pageId="7" pageNumber="432" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BAA36A3FFDDFFFEFF73FF4DFA7D61A2" blockId="8.[151,1437,151,968]" pageId="8" pageNumber="433">
The spines of the sclerotic apparatus of the new species show a different organization compared to that of all the other species described in this genus. They are constituted by a median stylet pointed at both ends, 4849 Μm long, encircled by 19 bristles, with variable shape and length (1744 Μm). In
<taxonomicName id="4C154D20FFDDFFFEFBBAFF3AFB3D63DF" box="[1038,1225,224,248]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="433" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="B961EAB1FFDDFFFEFBBAFF3AFB3D63DF" box="[1038,1225,224,248]" italics="true" pageId="8" pageNumber="433">
<collectingCountry id="F3027633FFDDFFFEFBBAFF3AFBD163D0" box="[1038,1061,224,247]" name="Iceland" pageId="8" pageNumber="433">O</collectingCountry>
. mediterranea
</emphasis>
</taxonomicName>
, the median stylet has a relatively similar but variable length (4985 Μm) and is funnel-shaped, with a sharp narrowing toward the distal pointed end. The new species
<taxonomicName id="4C154D20FFDDFFFEFD9FFEF2FD7F6267" box="[555,651,296,320]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="433" phylum="Platyhelminthes" rank="species" species="lunae">
<emphasis id="B961EAB1FFDDFFFEFD9FFEF2FD7F6267" box="[555,651,296,320]" italics="true" pageId="8" pageNumber="433">
<collectingCountry id="F3027633FFDDFFFEFD9FFEF2FDB66218" box="[555,578,296,319]" name="Iceland" pageId="8" pageNumber="433">O</collectingCountry>
. lunae
</emphasis>
</taxonomicName>
,
<taxonomicName id="4C154D20FFDDFFFEFD2EFEF2FCA76267" box="[666,851,296,320]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="433" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="B961EAB1FFDDFFFEFD2EFEF2FCA76267" box="[666,851,296,320]" italics="true" pageId="8" pageNumber="433">
<collectingCountry id="F3027633FFDDFFFEFD2EFEF2FD456218" box="[666,689,296,319]" name="Iceland" pageId="8" pageNumber="433">O</collectingCountry>
. mediterranea
</emphasis>
</taxonomicName>
and
<taxonomicName id="4C154D20FFDDFFFEFC3DFEF2FC1D6267" box="[905,1001,296,320]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="433" phylum="Platyhelminthes" rank="species" species="kohni">
<emphasis id="B961EAB1FFDDFFFEFC3DFEF2FC1D6267" box="[905,1001,296,320]" italics="true" pageId="8" pageNumber="433">
<collectingCountry id="F3027633FFDDFFFEFC3DFEF2FC546218" box="[905,928,296,319]" name="Iceland" pageId="8" pageNumber="433">O</collectingCountry>
. kohni
</emphasis>
</taxonomicName>
have a tube-shaped median stylet with a continuous, uninterrupted wall.
<taxonomicName id="4C154D20FFDDFFFEFDA7FE96FD4A6243" box="[531,702,332,356]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="433" phylum="Platyhelminthes" rank="species" species="sewardensis">
<emphasis id="B961EAB1FFDDFFFEFDA7FE96FD4A6243" box="[531,702,332,356]" italics="true" pageId="8" pageNumber="433">
<collectingCountry id="F3027633FFDDFFFEFDA7FE96FDDE6244" box="[531,554,332,355]" name="Iceland" pageId="8" pageNumber="433">O</collectingCountry>
. sewardensis
</emphasis>
</taxonomicName>
and
<taxonomicName id="4C154D20FFDDFFFEFD41FE96FC8B6243" box="[757,895,332,356]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="433" phylum="Platyhelminthes" rank="species" species="bregazzii">
<emphasis id="B961EAB1FFDDFFFEFD41FE96FC8B6243" box="[757,895,332,356]" italics="true" pageId="8" pageNumber="433">
<collectingCountry id="F3027633FFDDFFFEFD41FE96FCF86244" box="[757,780,332,355]" name="Iceland" pageId="8" pageNumber="433">O</collectingCountry>
. bregazzii
</emphasis>
</taxonomicName>
are particularly similar regarding their sclerotic apparatus as both species have a groove-shaped stylet. In
<taxonomicName id="4C154D20FFDDFFFEFC99FEAAFC6462AF" box="[813,912,368,392]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="433" phylum="Platyhelminthes" rank="species" species="lunae">
<emphasis id="B961EAB1FFDDFFFEFC99FEAAFC6462AF" box="[813,912,368,392]" italics="true" pageId="8" pageNumber="433">
<collectingCountry id="F3027633FFDDFFFEFC99FEAAFCB062A0" box="[813,836,368,391]" name="Iceland" pageId="8" pageNumber="433">O</collectingCountry>
. lunae
</emphasis>
</taxonomicName>
and
<taxonomicName id="4C154D20FFDDFFFEFC7EFEAAFB7162AF" box="[970,1157,368,392]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="433" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="B961EAB1FFDDFFFEFC7EFEAAFB7162AF" box="[970,1157,368,392]" italics="true" pageId="8" pageNumber="433">
<collectingCountry id="F3027633FFDDFFFEFC7EFEAAFC1562A0" box="[970,993,368,391]" name="Iceland" pageId="8" pageNumber="433">O</collectingCountry>
. mediterranea
</emphasis>
</taxonomicName>
, the spines are arranged around the median stylet to form a crown, similar in shape and numbers of spines in each species. In fact, in the former species there are 19 spines, while in the latter 1722 spines are present. They show an elongated and thin body, with a sub-terminal prominence and a bent tip. The different peculiarities between the spines in both species are in their length, 5565 Μm in
<taxonomicName id="4C154D20FFDDFFFEFDB5FDDAFD4E613F" box="[513,698,512,536]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="433" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="B961EAB1FFDDFFFEFDB5FDDAFD4E613F" box="[513,698,512,536]" italics="true" pageId="8" pageNumber="433">
<collectingCountry id="F3027633FFDDFFFEFDB5FDDAFDEC6130" box="[513,536,512,535]" name="Iceland" pageId="8" pageNumber="433">O</collectingCountry>
. mediterranea
</emphasis>
</taxonomicName>
and 1844 Μm in
<taxonomicName id="4C154D20FFDDFFFEFC32FDDAFC12613F" box="[902,998,512,536]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="433" phylum="Platyhelminthes" rank="species" species="lunae">
<emphasis id="B961EAB1FFDDFFFEFC32FDDAFC12613F" box="[902,998,512,536]" italics="true" pageId="8" pageNumber="433">
<collectingCountry id="F3027633FFDDFFFEFC32FDDAFC696130" box="[902,925,512,535]" name="Iceland" pageId="8" pageNumber="433">O</collectingCountry>
. lunae
</emphasis>
</taxonomicName>
, and also in their peculiar structures of the distal ends:
<taxonomicName id="4C154D20FFDDFFFEFEFAFDFEFDFE611B" box="[334,522,548,572]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="433" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="B961EAB1FFDDFFFEFEFAFDFEFDFE611B" box="[334,522,548,572]" italics="true" pageId="8" pageNumber="433">
<collectingCountry id="F3027633FFDDFFFEFEFAFDFEFE91611C" box="[334,357,548,571]" name="Iceland" pageId="8" pageNumber="433">O</collectingCountry>
. mediterranea
</emphasis>
</taxonomicName>
has a sub-terminal cuneiform protrusion well distinguishable with an outwards small tip, and the distal extremity is sharply curved to form a hooked tip; the new species displays a poorly developed sub-terminal cuneiform prominence, and the distal end of each spine appears with straight tip upward.
</paragraph>
<paragraph id="8BAA36A3FFDDFFFEFF73FD4AFAD861EB" blockId="8.[151,1437,151,968]" pageId="8" pageNumber="433">
On the base of the data presented,
<taxonomicName id="4C154D20FFDDFFFEFDF9FD4AFD5A618F" box="[589,686,656,680]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="433" phylum="Platyhelminthes" rank="species" species="lunae">
<emphasis id="B961EAB1FFDDFFFEFDF9FD4AFD5A618F" box="[589,686,656,680]" italics="true" pageId="8" pageNumber="433">
<collectingCountry id="F3027633FFDDFFFEFDF9FD4AFD906180" box="[589,612,656,679]" name="Iceland" pageId="8" pageNumber="433">O</collectingCountry>
. lunae
</emphasis>
</taxonomicName>
differs from the previously described species in body dimensions, size and distribution of testes and vitellaries, and, above all, the characteristics of the sclerotic apparatus.
</paragraph>
<paragraph id="8BAA36A3FFDDFFFEFF73FD0DFA6761D7" blockId="8.[151,1437,151,968]" box="[199,1427,727,752]" pageId="8" pageNumber="433">
The new species has many homologies with
<taxonomicName id="4C154D20FFDDFFFEFD0FFD02FC8061D7" box="[699,884,728,752]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="433" phylum="Platyhelminthes" rank="species" species="mediterranea">
<emphasis id="B961EAB1FFDDFFFEFD0FFD02FC8061D7" box="[699,884,728,752]" italics="true" pageId="8" pageNumber="433">
<collectingCountry id="F3027633FFDDFFFEFD0FFD02FD2661C8" box="[699,722,728,751]" name="Iceland" pageId="8" pageNumber="433">O</collectingCountry>
. mediterranea
</emphasis>
</taxonomicName>
, but it constitutes a valid new otoplanid species.
</paragraph>
</subSubSection>
<subSubSection id="C30F6528FFDDFFFEFF73FD26FE506083" pageId="8" pageNumber="433" type="etymology">
<paragraph id="8BAA36A3FFDDFFFEFF73FD26FDBE60A7" blockId="8.[151,1437,151,968]" pageId="8" pageNumber="433">
<emphasis id="B961EAB1FFDDFFFEFF73FD26FEBA6032" bold="true" box="[199,334,764,789]" pageId="8" pageNumber="433">Etymology.</emphasis>
The name
<taxonomicName id="4C154D20FFDDFFFEFE63FD26FDE16033" box="[471,533,764,788]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="433" phylum="Platyhelminthes" rank="species" species="lunae">
<emphasis id="B961EAB1FFDDFFFEFE63FD26FDE16033" box="[471,533,764,788]" italics="true" pageId="8" pageNumber="433">lunae</emphasis>
</taxonomicName>
refers to the
<typeStatus id="54AE8801FFDDFFFEFD0EFD27FD1E6032" box="[698,746,765,789]" pageId="8" pageNumber="433">type</typeStatus>
locality where this species has been collected. The site of sampling is on the coast of the territory called Lunigiana, corresponding to the basin of the Magra River located between Liguria, Emilia-Romagna and Tuscany (Ligurian Sea,
<collectingCountry id="F3027633FFDDFFFEFCD5FC9EFC6C607A" box="[865,920,836,861]" name="Italy" pageId="8" pageNumber="433">Italy</collectingCountry>
), and
<taxonomicName id="4C154D20FFDDFFFEFC68FC9EFBEE607B" box="[988,1050,836,860]" class="Rhabditophora" family="Otoplanidae" genus="Orthoplana" kingdom="Animalia" order="Seriata" pageId="8" pageNumber="433" phylum="Platyhelminthes" rank="species" species="lunae">
<emphasis id="B961EAB1FFDDFFFEFC68FC9EFBEE607B" box="[988,1050,836,860]" italics="true" pageId="8" pageNumber="433">lunae</emphasis>
</taxonomicName>
means “coming from the territory of the ancient Etruscan town of Luna”.
</paragraph>
<paragraph id="8BAA36A3FFDDFFFEFF73FC56FE506083" blockId="8.[151,1437,151,968]" box="[199,420,908,933]" pageId="8" pageNumber="433">
<emphasis id="B961EAB1FFDDFFFEFF73FC56FEDE6082" bold="true" box="[199,298,908,933]" pageId="8" pageNumber="433">Habitat.</emphasis>
Surf zone.
</paragraph>
</subSubSection>
<subSubSection id="C30F6528FFDDFFFEFF73FC75FD3C60EF" box="[199,712,943,968]" pageId="8" pageNumber="433" type="distribution">
<paragraph id="8BAA36A3FFDDFFFEFF73FC75FD3C60EF" blockId="8.[151,1437,151,968]" box="[199,712,943,968]" pageId="8" pageNumber="433">
<emphasis id="B961EAB1FFDDFFFEFF73FC75FE9460EF" bold="true" box="[199,352,943,968]" pageId="8" pageNumber="433">Distribution.</emphasis>
Known only from
<typeStatus id="54AE8801FFDDFFFEFD83FC6AFD9360EF" box="[567,615,944,968]" pageId="8" pageNumber="433">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>