297 lines
38 KiB
XML
297 lines
38 KiB
XML
<document id="E4D848777645E1FD424DD60A92E28989" ID-DOI="10.5852/ejt.2012.5" ID-GBIF-Dataset="e809cd3a-36bb-413d-9f19-70837d7db34b" ID-Zenodo-Dep="3715167" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1584542927971" checkinUser="plazi" docAuthor="Grootaert, Patrick & Shamshev, Igor V." docDate="2012" docId="03BC87A25D29FFEDFD9DBE96330C844C" docLanguage="en" docName="ejt-2011-5.pdf.imf" docOrigin="European Journal of Taxonomy 5" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Nanodromia Grootaert 1994" docType="treatment" docVersion="15" lastPageNumber="113" masterDocId="FF85FFDA5D46FF9DFFDEBF61367B815E" masterDocTitle="The fast-running flies (Diptera, Hybotidae, Tachydromiinae) of Singapore and adjacent regions" masterLastPageNumber="162" masterPageNumber="1" pageNumber="112" updateTime="1698729919212" updateUser="plazi">
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<mods:title id="1C8E95E9C3BAC8C0FF3DF1B93ED2951B">The fast-running flies (Diptera, Hybotidae, Tachydromiinae) of Singapore and adjacent regions</mods:title>
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<mods:namePart id="40C92FD89302C2239ED7FB54C0EEFB1E">Grootaert, Patrick</mods:namePart>
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<mods:namePart id="762DD6EDC7ACF57580C08FACE6A10DA8">Shamshev, Igor V.</mods:namePart>
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<mods:title id="159BE222811BA4B062D77C9126DF5A20">European Journal of Taxonomy</mods:title>
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<mods:part id="F2EED75C3A43301717DF5CB6035743BE">
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<mods:date id="5F33A8939EB44AC17D4928492D904B05">2012</mods:date>
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<mods:number id="7818C84A0137FE0AE0F3546A2DA45AB0">2012-01-06</mods:number>
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<mods:number id="78B1C9BBA1F9D54363C9A8F619CDFE2D">5</mods:number>
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<mods:identifier id="3A8D287F89BE700154BF698DF150AE45" type="DOI">10.5852/ejt.2012.5</mods:identifier>
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<treatment id="03BC87A25D29FFEDFD9DBE96330C844C" ID-DOI="http://doi.org/10.5281/zenodo.3716654" ID-GBIF-Taxon="162630272" ID-Zenodo-Dep="3716654" LSID="urn:lsid:plazi:treatment:03BC87A25D29FFEDFD9DBE96330C844C" httpUri="http://treatment.plazi.org/id/03BC87A25D29FFEDFD9DBE96330C844C" lastPageId="112" lastPageNumber="113" pageId="111" pageNumber="112">
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<subSubSection id="C30F653F5D29FFF2FD9DBE96358B834F" box="[579,1008,503,530]" pageId="111" pageNumber="112" type="nomenclature">
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<paragraph id="8BAA36B45D29FFF2FD9DBE96358B834F" blockId="111.[189,1388,503,596]" box="[579,1008,503,530]" pageId="111" pageNumber="112">
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<heading id="D0E281D85D29FFF2FD9DBE96358B834F" box="[579,1008,503,530]" centered="true" fontSize="11" level="2" pageId="111" pageNumber="112" reason="2">
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Genus
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<taxonomicName id="4C154D375D29FFF2FD48BE96358B834F" ID-CoL="63CJM" ID-ENA="626626" authority="Grootaert, 1994" authorityName="Grootaert" authorityYear="1994" box="[662,1008,503,530]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EAA65D29FFF2FD48BE963554834C" bold="true" box="[662,815,503,530]" italics="true" pageId="111" pageNumber="112">Nanodromia</emphasis>
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<bibRefCitation id="EF844B455D29FFF2FCE8BE96358B834F" author="Grootaert" box="[822,1008,503,529]" firstAuthor="Grootaert" pageId="111" pageNumber="112" pagination="141 - 156" refId="ref68922" refString="Grootaert P. 1994. Two new genera of empidoid flies (Diptera, Empidoidea, Hybotidae, Tachydromiinae) from Melanesia. Studia dipterologica 1 (2): 141 - 156." type="journal article" year="1994">Grootaert, 1994</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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||
</subSubSection>
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<treatmentCitationGroup id="AB05119A5D29FFF2FF63BD5B3317830A" box="[189,1388,570,596]" pageId="111" pageNumber="112">
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<subSubSection id="C30F653F5D29FFF2FF63BD5B342F830A" box="[189,596,570,596]" pageId="111" pageNumber="112" type="reference_group">
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<paragraph id="8BAA36B45D29FFF2FF63BD5B342F830A" blockId="111.[189,1388,503,596]" box="[189,596,570,596]" pageId="111" pageNumber="112">
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<treatmentCitation id="0AB410A55D29FFF2FF63BD5B342B830A" author="Grootaert P." box="[189,592,570,596]" page="142" pageId="111" pageNumber="112" year="1994">
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<taxonomicName id="4C154D375D29FFF2FF63BD5B342B830A" ID-CoL="63CJM" authority="Grootaert, 1994: 142" authorityName="Grootaert" authorityPageNumber="142" authorityYear="1994" box="[189,592,570,596]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EAA65D29FFF2FF63BD5B372B830A" box="[189,336,570,596]" italics="true" pageId="111" pageNumber="112">Nanodromia</emphasis>
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<bibRefCitation id="EF844B455D29FFF2FE86BD5B342B830A" author="Grootaert" box="[344,592,570,596]" firstAuthor="Grootaert" pageId="111" pageNumber="112" pagination="141 - 156" refId="ref68922" refString="Grootaert P. 1994. Two new genera of empidoid flies (Diptera, Empidoidea, Hybotidae, Tachydromiinae) from Melanesia. Studia dipterologica 1 (2): 141 - 156." type="journal article" year="1994">Grootaert, 1994: 142</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F653F5D29FFF2FD85BD5B3317830A" box="[603,1388,570,596]" pageId="111" pageNumber="112" type="type_taxon">
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<paragraph id="8BAA36B45D29FFF2FD85BD5B3317830A" blockId="111.[189,1388,503,596]" box="[603,1388,570,596]" pageId="111" pageNumber="112">
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Type-species:
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<taxonomicName id="4C154D375D29FFF2FCDDBD5A3232830A" ID-CoL="45MYV" authority="Grootaert, 1994" authorityName="Grootaert" authorityYear="1994" box="[771,1097,570,596]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="cryptica">
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<emphasis id="B961EAA65D29FFF2FCDDBD5A35FE830A" box="[771,901,571,596]" italics="true" pageId="111" pageNumber="112">N. cryptica</emphasis>
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<bibRefCitation id="EF844B455D29FFF2FC53BD5B3232830A" author="Grootaert" box="[909,1097,570,596]" firstAuthor="Grootaert" pageId="111" pageNumber="112" pagination="141 - 156" refId="ref68922" refString="Grootaert P. 1994. Two new genera of empidoid flies (Diptera, Empidoidea, Hybotidae, Tachydromiinae) from Melanesia. Studia dipterologica 1 (2): 141 - 156." type="journal article" year="1994">Grootaert, 1994</bibRefCitation>
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</taxonomicName>
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, by original designation.
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</paragraph>
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</subSubSection>
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</treatmentCitationGroup>
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<subSubSection id="C30F653F5D29FFF2FF63BD1D37418209" pageId="111" pageNumber="112" type="discussion">
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<paragraph id="8BAA36B45D29FFF2FF63BD1D372883C8" blockId="111.[189,339,636,662]" box="[189,339,636,662]" pageId="111" pageNumber="112">
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<heading id="D0E281D85D29FFF2FF63BD1D372883C8" bold="true" box="[189,339,636,662]" fontSize="11" level="3" pageId="111" pageNumber="112" reason="3">
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<emphasis id="B961EAA65D29FFF2FF63BD1D372883C8" bold="true" box="[189,339,636,662]" pageId="111" pageNumber="112">Recognition</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAA36B45D29FFF2FF63BDCD37418209" blockId="111.[189,1399,684,855]" pageId="111" pageNumber="112">
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The genus
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<taxonomicName id="4C154D375D29FFF2FE96BDCD37A08398" authorityName="Grootaert" authorityYear="1994" box="[328,475,684,710]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EAA65D29FFF2FE96BDCD37A08398" box="[328,475,684,710]" italics="true" pageId="111" pageNumber="112">Nanodromia</emphasis>
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</taxonomicName>
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can be distinguished from other genera of the
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<taxonomicName id="4C154D375D29FFF2FBF0BDCD32E68398" authorityName="Collin" authorityYear="1961" box="[1070,1181,684,710]" class="Insecta" family="Hybotidae" kingdom="Animalia" order="Diptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="tribe" tribe="Drapetini">Drapetini</taxonomicName>
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by the following combination of characters: postpedicel with dorsoapical extension, wing with Rs relatively short and originating closer to tip of R
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<subScript id="179134F15D29FFF2FDCABC673466824A" attach="left" box="[532,541,774,788]" fontSize="6" pageId="111" pageNumber="112">1</subScript>
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than humeral crossvein, cells br and bm of subequal length, abdominal tergites lacking squamiform setae, intersegmental modifications present, two rod-shaped genitalic apodemes.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F653F5D29FFF2FF63BC1E35F687B9" pageId="111" pageNumber="112" type="diagnosis">
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<paragraph id="8BAA36B45D29FFF2FF63BC1E374D82C7" blockId="111.[189,310,895,921]" box="[189,310,895,921]" pageId="111" pageNumber="112">
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<heading id="D0E281D85D29FFF2FF63BC1E374D82C7" bold="true" box="[189,310,895,921]" fontSize="11" level="3" pageId="111" pageNumber="112" reason="3">
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<emphasis id="B961EAA65D29FFF2FF63BC1E374D82C7" bold="true" box="[189,310,895,921]" pageId="111" pageNumber="112">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAA36B45D29FFF2FF63BCCE35F687B9" blockId="111.[189,1399,943,1768]" pageId="111" pageNumber="112">
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Small flies with body length about
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<quantity id="4CED9B515D29FFF2FD96BCCE34D88294" box="[584,675,943,970]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="111" pageNumber="112" unit="mm" value="1.5">1.5 mm</quantity>
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. Frons very broad, with sides divergent above. Eyes contiguous on face, lower ommatidia enlarged or ommatidia of subequal size. Two pairs of vertical bristles, but outer pair (sometimes both pairs) often hardly prominent. Ocellar tubercle with 2 pairs of bristles. Antenna with pedicel bearing long ventral subapical seta; postpedicel with dorsoapical extension; stylus arising dorsoapically, arista-like, long. Gena barely extended below eye. Palpus small, rounded, usually with short subapical seta. Thorax brown to black; sometimes modified (
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<taxonomicName id="4C154D375D29FFF2FB88BB04374085FA" authority="Grootaert & Shamshev" authorityName="Grootaert & Shamshev" authorityYear="2003" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="narmkroi">
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<emphasis id="B961EAA65D29FFF2FB88BB0432988521" box="[1110,1251,1125,1151]" italics="true" pageId="111" pageNumber="112">N. narmkroi</emphasis>
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Grootaert & Shamshev
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</taxonomicName>
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). Postpronotal bristle usually well prominent (except
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<taxonomicName id="4C154D375D29FFF2FC69BBEB323C85FD" authorityName="Grootaert & Shamshev" authorityYear="2003" box="[951,1095,1161,1187]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="narmkroi">
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<emphasis id="B961EAA65D29FFF2FC69BBEB323C85FD" box="[951,1095,1161,1187]" italics="true" pageId="111" pageNumber="112">N. narmkroi</emphasis>
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</taxonomicName>
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). Mesonotal bristles very prominent but sometimes acrostichals and dorsocentrals weakly differentiated. Anepisternum bare, rarely setose (
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<taxonomicName id="4C154D375D29FFF2FEB2BBB3347B85B5" authorityName="Grootaert & Shamshev" authorityYear="2003" box="[364,512,1233,1259]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="narmkroi">
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<emphasis id="B961EAA65D29FFF2FEB2BBB3347B85B5" box="[364,512,1233,1259]" italics="true" pageId="111" pageNumber="112">N. narmkroi</emphasis>
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</taxonomicName>
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). Legs short; fore femur usually very thickened; mid legs usually with secondary sexual characters in male; hind tibia lacking prominent bristles. Wings usually normally developed or sometimes modified (non Oriental species); often with distinct pattern; Rs relatively short and originating closer to tip of R
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<subScript id="179134F15D29FFF2FD9CBA3134308400" attach="left" box="[578,587,1360,1374]" fontSize="6" pageId="111" pageNumber="112">1</subScript>
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than humeral crossvein; R
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<subScript id="179134F15D29FFF2FC56BA3135D88400" attach="left" box="[904,931,1360,1374]" fontSize="6" pageId="111" pageNumber="112">2+3</subScript>
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complete, meeting costa before middle of wing; R
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<subScript id="179134F15D29FFF2FEE2BA15372D84DC" attach="left" box="[316,342,1396,1410]" fontSize="6" pageId="111" pageNumber="112">4+5</subScript>
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and M
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<subScript id="179134F15D29FFF2FE75BA1537BD84DC" attach="left" box="[427,454,1396,1410]" fontSize="6" pageId="111" pageNumber="112">1+2</subScript>
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strongly divergent; A
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<subScript id="179134F15D29FFF2FD19BA1534AB84DC" attach="left" box="[711,720,1396,1410]" fontSize="6" pageId="111" pageNumber="112">1</subScript>
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absent; crossvein CuA
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<subScript id="179134F15D29FFF2FC3FBA15359184DC" attach="left" box="[993,1002,1396,1410]" fontSize="6" pageId="111" pageNumber="112">2</subScript>
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absent; crossvein bm-cu oblique; cells br and bm of subequal length; Halter with knob yellow to black. Abdomen with tergite 1 entirely pale; no squamiform setae; gland-like intersegmental structures present. Terminalia rotated 90° to right; epandrium completely divided; left epandrial lamella usually very narrow, separated from hypandrium, sometimes completely reduced (
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<taxonomicName id="4C154D375D29FFF2FD9ABA9434AD8750" authorityName="Grootaert & Shamshev" authorityYear="2003" box="[580,726,1524,1550]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="species" species="narmkroi">
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<emphasis id="B961EAA65D29FFF2FD9ABA9434AD8750" box="[580,726,1524,1550]" italics="true" pageId="111" pageNumber="112">N. narmkroi</emphasis>
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</taxonomicName>
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); left surstylus undifferentiated from epandrial lamella, undivided, with unmodified setation; right epandrial lamella usually large, positioned ventrally; right surstylus present, slightly differentiated from epandrial lamella; cerci fused (sometimes incompletely), often bearing spines or papilla-like setae; hypandrium usually with 2 bristles on apical portion; phallus very short; two rod-shaped apodemes (i.e. ejaculatory and ventral apodemes) present.
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<emphasis id="B961EAA65D29FFF2FB1AB9E4336787C1" box="[1220,1308,1669,1695]" italics="true" pageId="111" pageNumber="112">Female</emphasis>
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similar to male, but legs with unmodified structure and vestiture; terminalia short; tergite 8 fused laterally with sternite 8; sternite 8 without folded apex; cercus broad-ovate.
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</paragraph>
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</subSubSection>
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<subSubSection id="C30F653F5D29FFEDFF63B871358280CD" lastPageId="112" lastPageNumber="113" pageId="111" pageNumber="112" type="discussion">
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<paragraph id="8BAA36B45D29FFF2FF63B87137548674" blockId="111.[189,303,1808,1834]" box="[189,303,1808,1834]" pageId="111" pageNumber="112">
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<heading id="D0E281D85D29FFF2FF63B87137548674" bold="true" box="[189,303,1808,1834]" fontSize="11" level="3" pageId="111" pageNumber="112" reason="3">
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<emphasis id="B961EAA65D29FFF2FF63B87137548674" bold="true" box="[189,303,1808,1834]" pageId="111" pageNumber="112">Remarks</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BAA36B45D29FFF2FF63B85E34B186B5" blockId="111.[189,1399,1855,2028]" pageId="111" pageNumber="112">
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The genus
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<taxonomicName id="4C154D375D29FFF2FE99B82137A18604" authorityName="Grootaert" authorityYear="1994" box="[327,474,1856,1882]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="genus">
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<emphasis id="B961EAA65D29FFF2FE99B82137A18604" box="[327,474,1856,1882]" italics="true" pageId="111" pageNumber="112">Nanodromia</emphasis>
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</taxonomicName>
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is known only from the Australasian (3 species from
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<collectingCountry id="F30276245D29FFF2FB5EB82133148604" box="[1152,1391,1855,1882]" name="Papua New Guinea" pageId="111" pageNumber="112">Papua New Guinea</collectingCountry>
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) and Oriental (6 species) regions (
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<bibRefCitation id="EF844B455D29FFF2FD95B805357E8620" author="Grootaert P." box="[587,773,1892,1918]" journalOrPublisher="Bulletin et Annales de la Societe Royale Belge d'Entomologie" pageId="111" pageNumber="112" pagination="235 - 244" part="128" refId="ref68881" refString="Grootaert P. 1992. New species of the genus Chersodromia Walker (Diptera Empidoidea Hybotidae) from the northern coast of Papua New Guinea. Bulletin et Annales de la Societe Royale Belge d'Entomologie 128: 235 - 244." title="New species of the genus Chersodromia Walker (Diptera Empidoidea Hybotidae) from the northern coast of Papua New Guinea" type="journal article" year="1992">Grootaert 1992</bibRefCitation>
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;
|
||
<bibRefCitation id="EF844B455D29FFF2FCCDB80532148620" author="Grootaert & Shamshev" box="[787,1135,1892,1918]" firstAuthor="Grootaert" pageId="111" pageNumber="112" pagination="41 - 52" refId="ref69082" refString="Grootaert P. & Shamshev I. 2003. New species of the genus Nanodromia Grootaert (Diptera: Empidoidea, Hybotidae) from Thailand and Singapore. The Natural History Journal of Chulalongkorn University 3 (2): 41 - 52." type="journal article" year="2003">Grootaert & Shamshev 2003</bibRefCitation>
|
||
). Here
|
||
<taxonomicName id="4C154D375D29FFF2FB18B80533228620" authorityName="Grootaert" authorityYear="1994" box="[1222,1369,1892,1918]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="111" pageNumber="112" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961EAA65D29FFF2FB18B80533228620" box="[1222,1369,1892,1918]" italics="true" pageId="111" pageNumber="112">Nanodromia</emphasis>
|
||
</taxonomicName>
|
||
is recorded for the first time from
|
||
<collectingCountry id="F30276245D29FFF2FDF7B8E934DB86FC" box="[553,672,1928,1954]" name="Singapore" pageId="111" pageNumber="112">Singapore</collectingCountry>
|
||
with three new species described below. There is an additional species with black legs known only from a single female (
|
||
<collectingCountry id="F30276245D29FFF2FCA0B8CD35818698" box="[894,1018,1964,1990]" name="Singapore" pageId="111" pageNumber="112">Singapore</collectingCountry>
|
||
, Sime forest
|
||
<date id="FFAB10745D29FFF2FB44B8CC33088698" box="[1178,1395,1964,1991]" pageId="111" pageNumber="112" value="2006-02-17">17 February 2006</date>
|
||
, forest, reg. 26034) that remains undescribed.
|
||
</paragraph>
|
||
<paragraph id="8BAA36B45D36FFEDFF63BE6A358280CD" blockId="112.[189,1399,267,404]" pageId="112" pageNumber="113">
|
||
<taxonomicName id="4C154D375D36FFEDFF63BE6A372B807B" authorityName="Grootaert" authorityYear="1994" box="[189,336,267,293]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961EAA65D36FFEDFF63BE6A372B807B" box="[189,336,267,293]" italics="true" pageId="112" pageNumber="113">Nanodromia</emphasis>
|
||
</taxonomicName>
|
||
as its name indicates is a genus of very small flies, and hence identification is quite difficult. In addition many species resemble one another and leg colouration is sometimes variable. The position of the abdominal glands can be helpful in alcohol preserved specimens, but dissection of the male genitalia is often mandatory to confirm an unambiguous identification.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30F653F5D36FFEDFF63BEDD330C844C" pageId="112" pageNumber="113" type="key">
|
||
<paragraph id="8BAA36B45D36FFEDFF63BEDD350A8088" blockId="112.[189,881,444,471]" box="[189,881,444,471]" pageId="112" pageNumber="113">
|
||
<heading id="D0E281D85D36FFEDFF63BEDD350A8088" bold="true" box="[189,881,444,471]" fontSize="11" level="3" pageId="112" pageNumber="113" reason="3">
|
||
<emphasis id="B961EAA65D36FFEDFF63BEDD350A8088" bold="true" box="[189,881,444,471]" pageId="112" pageNumber="113">
|
||
Key to species of
|
||
<taxonomicName id="4C154D375D36FFEDFE4ABEDD34568089" ID-CoL="63CJM" authorityName="Grootaert" authorityYear="1994" box="[404,557,444,471]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B961EAA65D36FFEDFE4ABEDD34568089" bold="true" box="[404,557,444,471]" italics="true" pageId="112" pageNumber="113">Nanodromia</emphasis>
|
||
</taxonomicName>
|
||
from the Oriental Region
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<key id="FF8461855D36FFEDFF63BE8D330C844C" pageId="112" pageNumber="113">
|
||
<keyStep id="30E12E115D36FFEDFF63BE8D330C832A" pageId="112" pageNumber="113">
|
||
<paragraph id="8BAA36B45D36FFEDFF63BE8D330C8375" blockId="112.[189,1399,492,628]" pageId="112" pageNumber="113">
|
||
<keyLead id="30E495815D36FFEDFF63BE8D330C8375" pageId="112" pageNumber="113">1. Scutum uniformly covered with hair-like setae (acrostichal and dorsocentral bristles undifferentiated)...................................................................................................................................2</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36B45D36FFEDFF1BBD57330C832A" blockId="112.[189,1399,492,628]" pageId="112" pageNumber="113">
|
||
<keyLead id="30E495815D36FFEDFF1BBD57330C832A" pageId="112" pageNumber="113">
|
||
– Acrostichal and dorsocentral bristles well differentiated in rows (
|
||
<collectingCountry id="F30276245D36FFEDFC33BD5432208311" box="[1005,1115,565,591]" name="Thailand" pageId="112" pageNumber="113">Thailand</collectingCountry>
|
||
;
|
||
<collectingCountry id="F30276245D36FFEDFBB7BD5432A38311" box="[1129,1240,565,591]" name="Malaysia" pageId="112" pageNumber="113">Malaysia</collectingCountry>
|
||
;
|
||
<collectingCountry id="F30276245D36FFEDFB39BD543318830E" box="[1255,1379,565,592]" name="Singapore" pageId="112" pageNumber="113">Singapore</collectingCountry>
|
||
).. ..........................................................................................
|
||
<taxonomicName id="4C154D375D36FFEDFCB1BD3B330C832A" ID-CoL="45MYY" authority="Grootaert & Shamshev, 2003" authorityName="Grootaert & Shamshev" authorityYear="2003" box="[879,1399,601,628]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="narmkroi">
|
||
<emphasis id="B961EAA65D36FFEDFCB1BD3B3272832A" bold="true" box="[879,1033,601,628]" italics="true" pageId="112" pageNumber="113">N. narmkroi</emphasis>
|
||
<bibRefCitation id="EF844B455D36FFEDFBCBBD3B330C832A" author="Grootaert & Shamshev" box="[1045,1399,602,628]" firstAuthor="Grootaert" pageId="112" pageNumber="113" pagination="41 - 52" refId="ref69082" refString="Grootaert P. & Shamshev I. 2003. New species of the genus Nanodromia Grootaert (Diptera: Empidoidea, Hybotidae) from Thailand and Singapore. The Natural History Journal of Chulalongkorn University 3 (2): 41 - 52." type="journal article" year="2003">Grootaert & Shamshev, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E12E115D36FFEDFF63BDFC330C827A" pageId="112" pageNumber="113">
|
||
<paragraph id="8BAA36B45D36FFEDFF63BDFC330C8385" blockId="112.[189,1399,668,805]" pageId="112" pageNumber="113">
|
||
<keyLead id="30E495815D36FFEDFF63BDFC330C8385" pageId="112" pageNumber="113">
|
||
2. Legs uniformly yellow; mid femur with basal swelling (
|
||
<figureCitation id="132E2A315D36FFEDFC10BDFD326683E9" box="[974,1053,668,695]" captionStart="Fig" captionStartId="3.[189,232,1930,1956]" captionTargetBox="[211,1382,1105,1876]" captionTargetId="figure@3.[189,1399,1083,1890]" captionTargetPageId="3" captionText="Fig. 1. Singapore in the Oriental Region (after Grootaert, 2009). STMF = subtropical mixed forest; STD = subtropical desert and semidesert; TDFS = tropical dry forest and savannah; TWF = tropical wet forest. The Weber line is considered to form the southeast boundary of the Oriental Region." figureDoi="http://doi.org/10.5281/zenodo.3715169" httpUri="https://zenodo.org/record/3715169/files/figure.png" pageId="112" pageNumber="113">Fig. 1</figureCitation>
|
||
) and mid tibia curved in dorsoventral plane (
|
||
<collectingCountry id="F30276245D36FFEDFE12BDA0344E8382" box="[460,565,705,732]" name="Thailand" pageId="112" pageNumber="113">Thailand</collectingCountry>
|
||
).........................................
|
||
<taxonomicName id="4C154D375D36FFEDFCB2BDA3330C8385" ID-CoL="45MYX" authority="Grootaert & Shamshev, 2003" authorityName="Grootaert & Shamshev" authorityYear="2003" box="[876,1399,705,732]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="narmjeud">
|
||
<emphasis id="B961EAA65D36FFEDFCB2BDA332728382" bold="true" box="[876,1033,705,732]" italics="true" pageId="112" pageNumber="113">N. narmjeud</emphasis>
|
||
<bibRefCitation id="EF844B455D36FFEDFBCBBDA0330C8385" author="Grootaert & Shamshev" box="[1045,1399,705,731]" firstAuthor="Grootaert" pageId="112" pageNumber="113" pagination="41 - 52" refId="ref69082" refString="Grootaert P. & Shamshev I. 2003. New species of the genus Nanodromia Grootaert (Diptera: Empidoidea, Hybotidae) from Thailand and Singapore. The Natural History Journal of Chulalongkorn University 3 (2): 41 - 52." type="journal article" year="2003">Grootaert & Shamshev, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36B45D36FFEDFF1BBD87330C827A" blockId="112.[189,1399,668,805]" pageId="112" pageNumber="113">
|
||
<keyLead id="30E495815D36FFEDFF1BBD87330C827A" pageId="112" pageNumber="113">– Legs with hind femur and (or) tibia brownish near middle (sometimes only faintly); mid femur slender or with basal swelling, but mid tibia always straight..........................................................3</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E12E115D36FFEDFF63BC2C330C82EF" pageId="112" pageNumber="113">
|
||
<paragraph id="8BAA36B45D36FFEDFF63BC2C330E82D3" blockId="112.[189,1399,845,945]" pageId="112" pageNumber="113">
|
||
<keyLead id="30E495815D36FFEDFF63BC2C330E82D3" pageId="112" pageNumber="113">
|
||
3. Hind tibia unmodified. Gland-like structures present between tergites 4 and 5 and 5 and 6 (
|
||
<figureCitation id="132E2A315D36FFEDFA9CBC2C374382D3" captionStart="Figs 172-177" captionStartId="115.[189,241,1824,1850]" captionTargetBox="[288,1262,278,1700]" captionTargetId="figure@115.[256,1331,265,1784]" captionText="Figs 172-177. Nanodromia, ♂. 172. N. hutan sp. nov., hind leg, anterior view. 173. N. hutan sp. nov., tip of hind tibia, dorsal view. 174. N. phukhao Grootaert & Shamshev, 2003, hind tibia, dorsal view. 175. N. narmjeud Grootaert & Shamshev, 2003, bladder-like gland between tergite 4 and 5, dorsal view. 176. N. narmkroi Grootaert & Shamshev, 2003, gland-like fields between tergites 4-5 and 5-6, dorsal view. 177. N. phukhao Grootaert & Shamshev, 2003, gland-like fields between tergites 4-5 and 5-6, dorsal view. (Figs 174-177 after Grootaert & Shamshev 2003)." figureDoi="http://doi.org/10.5281/zenodo.3723758" httpUri="https://zenodo.org/record/3723758/files/figure.png" pageId="112" pageNumber="113">Figs 176-177</figureCitation>
|
||
)........................................................................................................................................4
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36B45D36FFEDFF1BBCF6330C82EF" blockId="112.[189,1399,845,945]" box="[197,1399,918,945]" pageId="112" pageNumber="113">
|
||
<keyLead id="30E495815D36FFEDFF1BBCF6330C82EF" box="[197,1399,918,945]" pageId="112" pageNumber="113">
|
||
– Hind tibia somewhat curved. Bladder-like structure opening between tergite 4 and 5 (
|
||
<figureCitation id="132E2A315D36FFEDFB0BBCF7334182EF" box="[1237,1338,918,945]" captionStart="Figs 172-177" captionStartId="115.[189,241,1824,1850]" captionTargetBox="[288,1262,278,1700]" captionTargetId="figure@115.[256,1331,265,1784]" captionText="Figs 172-177. Nanodromia, ♂. 172. N. hutan sp. nov., hind leg, anterior view. 173. N. hutan sp. nov., tip of hind tibia, dorsal view. 174. N. phukhao Grootaert & Shamshev, 2003, hind tibia, dorsal view. 175. N. narmjeud Grootaert & Shamshev, 2003, bladder-like gland between tergite 4 and 5, dorsal view. 176. N. narmkroi Grootaert & Shamshev, 2003, gland-like fields between tergites 4-5 and 5-6, dorsal view. 177. N. phukhao Grootaert & Shamshev, 2003, gland-like fields between tergites 4-5 and 5-6, dorsal view. (Figs 174-177 after Grootaert & Shamshev 2003)." figureDoi="http://doi.org/10.5281/zenodo.3723758" httpUri="https://zenodo.org/record/3723758/files/figure.png" pageId="112" pageNumber="113">Fig. 175</figureCitation>
|
||
).....5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E12E115D36FFEDFF63BCB8330C853C" pageId="112" pageNumber="113">
|
||
<paragraph id="8BAA36B45D36FFEDFF63BCB8330C8546" blockId="112.[189,1399,985,1122]" pageId="112" pageNumber="113">
|
||
<keyLead id="30E495815D36FFEDFF63BCB8330C8546" pageId="112" pageNumber="113">
|
||
4. Mid femur with strong black bristle in anterior fourth; some curved spine-like ventral bristles near tip (
|
||
<collectingCountry id="F30276245D36FFEDFED3BC9F370E8547" box="[269,373,1022,1049]" name="Thailand" pageId="112" pageNumber="113">Thailand</collectingCountry>
|
||
)....................................................................
|
||
<taxonomicName id="4C154D375D36FFEDFCAABC9F330C8546" ID-CoL="45MYZ" authority="Grootaert & Shamshev, 2003" authorityName="Grootaert & Shamshev" authorityYear="2003" box="[884,1399,1021,1048]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="phukhao">
|
||
<emphasis id="B961EAA65D36FFEDFCAABC9F32738546" bold="true" box="[884,1032,1021,1048]" italics="true" pageId="112" pageNumber="113">N. phukhao</emphasis>
|
||
<bibRefCitation id="EF844B455D36FFEDFBCABC9F330C8546" author="Grootaert & Shamshev" box="[1044,1399,1022,1048]" firstAuthor="Grootaert" pageId="112" pageNumber="113" pagination="41 - 52" refId="ref69082" refString="Grootaert P. & Shamshev I. 2003. New species of the genus Nanodromia Grootaert (Diptera: Empidoidea, Hybotidae) from Thailand and Singapore. The Natural History Journal of Chulalongkorn University 3 (2): 41 - 52." type="journal article" year="2003">Grootaert & Shamshev, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36B45D36FFEDFF1BBB42330C853C" blockId="112.[189,1399,985,1122]" pageId="112" pageNumber="113">
|
||
<keyLead id="30E495815D36FFEDFF1BBB42330C853C" pageId="112" pageNumber="113">
|
||
– Mid femur with yellow anterior preapical bristle only; without curved bristles ventrally (
|
||
<collectingCountry id="F30276245D36FFEDFB2ABB4333148563" box="[1268,1391,1058,1085]" name="Singapore" pageId="112" pageNumber="113">Singapore</collectingCountry>
|
||
) ....................................................................................................................................
|
||
<taxonomicName id="4C154D375D36FFEDFB7DBB29336A853C" ID-CoL="BCVHV" authority="Grootaert & Shamshev, 2012" authorityName="Grootaert & Shamshev" authorityYear="2012" box="[1187,1297,1095,1122]" class="Insecta" family="Hybotidae" genus="Drapetis" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="hutan" status="sp. nov.">
|
||
<emphasis id="B961EAA65D36FFEDFB7DBB29336A853C" bold="true" box="[1187,1297,1095,1122]" italics="true" pageId="112" pageNumber="113">N. hutan</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25257DD5D36FFEDFAC4BB29330C853C" box="[1306,1399,1096,1122]" pageId="112" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="30E12E115D36FFEDFF63BBEB330C844C" pageId="112" pageNumber="113">
|
||
<paragraph id="8BAA36B45D36FFEDFF63BBEB330C8597" blockId="112.[189,1399,1162,1299]" pageId="112" pageNumber="113">
|
||
<keyLead id="30E495815D36FFEDFF63BBEB330C8597" pageId="112" pageNumber="113">
|
||
5. Hind tibia somewhat curved at tip; bearing squamiform setae on apical half (
|
||
<figureCitation id="132E2A315D36FFEDFBA5BBEB329A85FA" box="[1147,1249,1162,1188]" captionStart="Figs 172-177" captionStartId="115.[189,241,1824,1850]" captionTargetBox="[288,1262,278,1700]" captionTargetId="figure@115.[256,1331,265,1784]" captionText="Figs 172-177. Nanodromia, ♂. 172. N. hutan sp. nov., hind leg, anterior view. 173. N. hutan sp. nov., tip of hind tibia, dorsal view. 174. N. phukhao Grootaert & Shamshev, 2003, hind tibia, dorsal view. 175. N. narmjeud Grootaert & Shamshev, 2003, bladder-like gland between tergite 4 and 5, dorsal view. 176. N. narmkroi Grootaert & Shamshev, 2003, gland-like fields between tergites 4-5 and 5-6, dorsal view. 177. N. phukhao Grootaert & Shamshev, 2003, gland-like fields between tergites 4-5 and 5-6, dorsal view. (Figs 174-177 after Grootaert & Shamshev 2003)." figureDoi="http://doi.org/10.5281/zenodo.3723758" httpUri="https://zenodo.org/record/3723758/files/figure.png" pageId="112" pageNumber="113">Fig. 174</figureCitation>
|
||
) (
|
||
<collectingCountry id="F30276245D36FFEDFB25BBEB331185FB" box="[1275,1386,1162,1189]" name="Thailand" pageId="112" pageNumber="113">Thailand</collectingCountry>
|
||
). ................................................................................................
|
||
<taxonomicName id="4C154D375D36FFEDFC45BBCE330C8597" ID-CoL="45MZ3" authority="Grootaert & Shamshev, 2003" authorityName="Grootaert & Shamshev" authorityYear="2003" box="[923,1399,1198,1225]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="taksin">
|
||
<emphasis id="B961EAA65D36FFEDFC45BBCE32708597" bold="true" box="[923,1035,1198,1225]" italics="true" pageId="112" pageNumber="113">N. taksin</emphasis>
|
||
<bibRefCitation id="EF844B455D36FFEDFBC8BBCE330C8597" author="Grootaert & Shamshev" box="[1046,1399,1198,1225]" firstAuthor="Grootaert" pageId="112" pageNumber="113" pagination="41 - 52" refId="ref69082" refString="Grootaert P. & Shamshev I. 2003. New species of the genus Nanodromia Grootaert (Diptera: Empidoidea, Hybotidae) from Thailand and Singapore. The Natural History Journal of Chulalongkorn University 3 (2): 41 - 52." type="journal article" year="2003">Grootaert & Shamshev, 2003</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="8BAA36B45D36FFEDFF1BBBB5330C844C" blockId="112.[189,1399,1162,1299]" pageId="112" pageNumber="113">
|
||
<keyLead id="30E495815D36FFEDFF1BBBB5330C844C" pageId="112" pageNumber="113">
|
||
– Hind tibia only faintly twisted; lacking squamiform setae on apical half, but with single line of short spinules (
|
||
<figureCitation id="132E2A315D36FFEDFE42BB99347C844C" box="[412,519,1272,1298]" captionStart="Figs 172-177" captionStartId="115.[189,241,1824,1850]" captionTargetBox="[288,1262,278,1700]" captionTargetId="figure@115.[256,1331,265,1784]" captionText="Figs 172-177. Nanodromia, ♂. 172. N. hutan sp. nov., hind leg, anterior view. 173. N. hutan sp. nov., tip of hind tibia, dorsal view. 174. N. phukhao Grootaert & Shamshev, 2003, hind tibia, dorsal view. 175. N. narmjeud Grootaert & Shamshev, 2003, bladder-like gland between tergite 4 and 5, dorsal view. 176. N. narmkroi Grootaert & Shamshev, 2003, gland-like fields between tergites 4-5 and 5-6, dorsal view. 177. N. phukhao Grootaert & Shamshev, 2003, gland-like fields between tergites 4-5 and 5-6, dorsal view. (Figs 174-177 after Grootaert & Shamshev 2003)." figureDoi="http://doi.org/10.5281/zenodo.3723758" httpUri="https://zenodo.org/record/3723758/files/figure.png" pageId="112" pageNumber="113">Fig. 173</figureCitation>
|
||
) (
|
||
<collectingCountry id="F30276245D36FFEDFDF8BB9934E7844D" box="[550,668,1272,1299]" name="Singapore" pageId="112" pageNumber="113">Singapore</collectingCountry>
|
||
).............................................................
|
||
<taxonomicName id="4C154D375D36FFEDFBB6BB99337C844C" ID-CoL="BDCKM" authority="Grootaert & Shamshev, 2012" authorityName="Grootaert & Shamshev" authorityYear="2012" box="[1128,1287,1271,1298]" class="Insecta" family="Hybotidae" genus="Nanodromia" kingdom="Animalia" order="Diptera" pageId="112" pageNumber="113" phylum="Arthropoda" rank="species" species="spinulosa" status="sp. nov.">
|
||
<emphasis id="B961EAA65D36FFEDFBB6BB99337C844C" bold="true" box="[1128,1287,1271,1298]" italics="true" pageId="112" pageNumber="113">N. spinulosa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25257DD5D36FFEDFACBBB99330C844C" box="[1301,1399,1272,1298]" pageId="112" pageNumber="113" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |