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<document id="6651F7B06A2045DA35F3877BED1CEB6B" ID-CLB-Dataset="25594" ID-DOI="10.11646/zootaxa.4646.3.3" ID-GBIF-Dataset="4a3d31ca-e16b-4ccf-8407-231aa27702f1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3351058" ID-ZooBank="25CA4A18-955F-4EA7-8978-E93032B54A2E" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1564032263204" checkinUser="plazi" docAuthor="Cranston, Peter S." docDate="2019" docId="B33701109B1D1E42C68CFC29FF593672" docLanguage="en" docName="zootaxao.4646.3.3.pdf" docOrigin="Zootaxa 4646 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Riethia Kieffer 1917" docType="treatment" docVersion="12" lastPageNumber="495" masterDocId="4F0E79689B3F1E60C61BFFD0FFA93021" masterDocTitle="Riethia (Kieffer 1917) (Diptera: Chironomidae) revised for the Austro-Pacific region" masterLastPageNumber="500" masterPageNumber="461" pageNumber="495" updateTime="1698747359191" updateUser="ExternalLinkService">
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<mods:title id="9256FDBF10FA808F0BB8E5C08FEB5B26">Riethia (Kieffer 1917) (Diptera: Chironomidae) revised for the Austro-Pacific region</mods:title>
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<subSubSection id="7384E38D9B1D1E42C68CFC29FECF3432" box="[151,358,1017,1044]" pageId="34" pageNumber="495" type="nomenclature">
<paragraph id="3B21B0069B1D1E42C68CFC29FECF3432" blockId="34.[151,358,1017,1080]" box="[151,358,1017,1044]" pageId="34" pageNumber="495">
<heading id="6069076A9B1D1E42C68CFC29FECF3432" bold="true" box="[151,358,1017,1044]" fontSize="11" level="1" pageId="34" pageNumber="495" reason="1">
<emphasis id="09EA6C149B1D1E42C68CFC29FECF3432" bold="true" box="[151,358,1017,1044]" pageId="34" pageNumber="495">
<taxonomicName id="FC9ECB859B1D1E42C68CFC29FF463435" ID-CoL="79ZG" ID-ENA="391820" authorityName="Kieffer" authorityYear="1917" box="[151,239,1017,1044]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="495" phylum="Arthropoda" rank="genus">
<emphasis id="09EA6C149B1D1E42C68CFC29FF463435" bold="true" box="[151,239,1017,1044]" italics="true" pageId="34" pageNumber="495">Riethia</emphasis>
</taxonomicName>
larva B
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="7384E38D9B1D1E42C68CFBCDFF563419" box="[151,255,1053,1080]" pageId="34" pageNumber="495" type="description">
<paragraph id="3B21B0069B1D1E42C68CFBCDFF563419" blockId="34.[151,358,1017,1080]" box="[151,255,1053,1080]" pageId="34" pageNumber="495">
(
<figureCitation id="A3A5AC839B1D1E42C6BBFBCDFF5E3419" box="[160,247,1053,1080]" captionStart="FIGURE 5" captionStartId="14.[151,250,1912,1937]" captionTargetBox="[292,1266,205,1851]" captionTargetId="figure@14.[272,1316,181,1886]" captionTargetPageId="14" captionText="FIGURE 5. Riethia spp. Larva. A. Dorsal head, R. cinctipes Freeman; B. Mentum, R. stictoptera Kieffer; C. Mandible, R. stictoptera. D-I. Antenna, D. R. azeylandica sp.n., E. R. cinctipes Freeman, F. R. donedwardi sp.n., G. R. neocaledonica sp.n., H. R. larval sp.B; I. R. stictoptera. Maxilla, J. R. neocaledonica, K. Pseudochironomus sp., L. Aedokritus coffeatus Trivinho- Strixino, M. Manoa pahayokeensis Jacobsen &amp; Perry, N. putative Megacentron Freeman, O. Manoa obscura Fittkau." httpUri="https://zenodo.org/record/3351068/files/figure.png" pageId="34" pageNumber="495">Fig. 5H</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7384E38D9B1D1E42C68CFBB5FF593672" pageId="34" pageNumber="495" type="materials_examined">
<paragraph id="3B21B0069B1D1E42C68CFBB5FEE034E6" blockId="34.[151,1437,1125,1619]" pageId="34" pageNumber="495">
<emphasis id="09EA6C149B1D1E42C68CFBB5FED334A1" bold="true" box="[151,378,1125,1152]" pageId="34" pageNumber="495">Material examined</emphasis>
, larval head capsules only,
<collectingCountry id="4389F0969B1D1E42C4A9FBB5FCF8345E" box="[690,849,1125,1151]" name="Australia" pageId="34" pageNumber="495">AUSTRALIA</collectingCountry>
,
<specimenCount id="2D987B8F9B1D1E42C57BFBB5FC21345E" box="[864,904,1125,1151]" pageId="34" pageNumber="495" type="generic">2L.</specimenCount>
,
<collectingRegion id="F95A7EE49B1D1E42C58EFBB5FBB5345E" box="[917,1052,1125,1151]" country="Australia" name="Queensland" pageId="34" pageNumber="495">Queensland</collectingRegion>
, nr Mareeba, Davies Ck.,
<geoCoordinate id="5EAAD6C19B1D1E42C35FFBB5FA34345E" box="[1348,1437,1125,1151]" degrees="17" direction="south" minutes="01" orientation="latitude" pageId="34" pageNumber="495" precision="925" value="-17.016666">17°01'S</geoCoordinate>
<geoCoordinate id="5EAAD6C19B1D1E42C68CFB59FEA93485" box="[151,256,1161,1188]" degrees="145" direction="east" minutes="35" orientation="longitude" pageId="34" pageNumber="495" precision="925" value="145.58333">145°35'E</geoCoordinate>
,
<date id="4F2096C69B1D1E42C717FB59FEDF3485" box="[268,374,1161,1188]" pageId="34" pageNumber="495" value="2005-09-03">3.ix.2005</date>
(
<collectionCode id="5D8F28C39B1D1E42C79FFB59FE1B3482" box="[388,434,1161,1187]" country="0" name="University of Montana Museum" pageId="34" pageNumber="495">MV</collectionCode>
<emphasis id="09EA6C149B1D1E42C7A2FB59FDB73485" bold="true" box="[441,542,1161,1188]" pageId="34" pageNumber="495">
<collectionCode id="5D8F28C39B1D1E42C7A2FB59FE5C3485" box="[441,501,1161,1188]" pageId="34" pageNumber="495">FNQ</collectionCode>
7_6
</emphasis>
(
<date id="4F2096C69B1D1E42C435FB59FDEC3485" box="[558,581,1161,1188]" pageId="34" pageNumber="495">3i</date>
), MV
<emphasis id="09EA6C149B1D1E42C495FB59FD5A3485" bold="true" box="[654,755,1161,1188]" pageId="34" pageNumber="495">FNQ7_8</emphasis>
(
<date id="4F2096C69B1D1E42C519FB59FCB23485" box="[770,795,1161,1188]" pageId="34" pageNumber="495">4i</date>
); 2L, Cloudy Ck.,
<geoCoordinate id="5EAAD6C19B1D1E42C5F0FB59FBCC3485" box="[1003,1125,1161,1188]" degrees="18" direction="south" minutes="59.50" orientation="latitude" pageId="34" pageNumber="495" precision="9" value="-18.991667">18°59.50'S</geoCoordinate>
<geoCoordinate id="5EAAD6C19B1D1E42C277FB59FB5F3482" box="[1132,1270,1161,1187]" degrees="146" direction="east" minutes="12.30" orientation="longitude" pageId="34" pageNumber="495" precision="9" value="146.205">146°12.30'E</geoCoordinate>
(
<emphasis id="09EA6C149B1D1E42C31CFB59FAFE3482" box="[1287,1367,1161,1187]" italics="true" pageId="34" pageNumber="495">Krosch</emphasis>
) (
<collectionCode id="5D8F28C39B1D1E42C375FB59FA353482" box="[1390,1436,1161,1187]" country="0" name="University of Montana Museum" pageId="34" pageNumber="495">MV</collectionCode>
<emphasis id="09EA6C149B1D1E42C68CFB7DFE8D34E9" bold="true" box="[151,292,1197,1224]" pageId="34" pageNumber="495">
<collectionCode id="5D8F28C39B1D1E42C68CFB7DFEB834E9" box="[151,273,1197,1224]" pageId="34" pageNumber="495">FNQCCR</collectionCode>
3,
</emphasis>
2).
</paragraph>
<paragraph id="3B21B0069B1D1E42C6DCFB02FD3F3531" blockId="34.[151,1437,1125,1619]" pageId="34" pageNumber="495">
<materialsCitation id="8BF6BA5B9B1D1E42C6DCFB02FD3F3531" ID-GBIF-Occurrence="2382714447" collectingDate="2009-01-03" collectingDateMax="2009-02-16" collectingDateMin="2009-01-03" collectionCode="MV" collectorName="Clyde R. &amp; Brooman &amp; Currowan S. F. &amp; Cabbage Tree Ck." latitude="-35.566666" location="New South Wales" longLatPrecision="1192" longitude="150.03334" pageId="34" pageNumber="495" specimenCode="AU09, CTCR14, 15" specimenCount="1" stateProvince="New South Wales">
<collectingRegion id="F95A7EE49B1D1E42C6DCFB02FE2434CA" box="[199,397,1233,1260]" country="Australia" name="New South Wales" pageId="34" pageNumber="495">New South Wales</collectingRegion>
, 2L (1
<collectingDate id="5F646F2E9B1D1E42C7C1FB01FE5B34CA" box="[474,498,1233,1259]" pageId="34" pageNumber="495">
<date id="4F2096C69B1D1E42C7C1FB01FE5B34CA" box="[474,498,1233,1259]" pageId="34" pageNumber="495">3i</date>
</collectingDate>
, 1
<collectingDate id="5F646F2E9B1D1E42C40AFB01FD8034CA" box="[529,553,1233,1259]" pageId="34" pageNumber="495">
<date id="4F2096C69B1D1E42C40AFB01FD8034CA" box="[529,553,1233,1259]" pageId="34" pageNumber="495">4i</date>
</collectingDate>
,
<collectorName id="966BD5D09B1D1E42C428FB01FD3134CD" box="[563,664,1233,1260]" pageId="34" pageNumber="495">Clyde R.</collectorName>
,
<collectorName id="966BD5D09B1D1E42C4B8FB02FCA734CD" box="[675,782,1234,1260]" pageId="34" pageNumber="495">Brooman</collectorName>
,
<collectingDate id="5F646F2E9B1D1E42C503FB01FC2434CA" box="[792,909,1233,1259]" pageId="34" pageNumber="495" value="2009-02-16">
<date id="4F2096C69B1D1E42C503FB01FC2434CA" box="[792,909,1233,1259]" pageId="34" pageNumber="495" value="2009-02-16">16.ii.2009</date>
</collectingDate>
; 2L,
<collectorName id="966BD5D09B1D1E42C5D8FB01FBCC34CA" box="[963,1125,1233,1259]" pageId="34" pageNumber="495">Currowan S.F.</collectorName>
,
<collectorName id="966BD5D09B1D1E42C26BFB01FA9334CA" box="[1136,1338,1233,1260]" pageId="34" pageNumber="495">Cabbage Tree Ck.</collectorName>
,
<geoCoordinate id="5EAAD6C19B1D1E42C35FFB01FA3534CD" box="[1348,1436,1233,1260]" degrees="35" direction="south" minutes="34" orientation="latitude" pageId="34" pageNumber="495" precision="925" value="-35.566666">35°34'S</geoCoordinate>
<geoCoordinate id="5EAAD6C19B1D1E42C68CFB25FEA83531" box="[151,257,1269,1296]" degrees="150" direction="east" minutes="02" orientation="longitude" pageId="34" pageNumber="495" precision="925" value="150.03334">150°02'E</geoCoordinate>
,
<collectingDate id="5F646F2E9B1D1E42C716FB25FEC0352E" box="[269,361,1269,1295]" pageId="34" pageNumber="495" value="2009-01-03">
<date id="4F2096C69B1D1E42C716FB25FEC0352E" box="[269,361,1269,1295]" pageId="34" pageNumber="495" value="2009-01-03">3.i.2009</date>
</collectingDate>
(
<collectionCode id="5D8F28C39B1D1E42C76CFB25FE0C352E" box="[375,421,1269,1295]" country="0" name="University of Montana Museum" pageId="34" pageNumber="495">MV</collectionCode>
<emphasis id="09EA6C149B1D1E42C7B7FB25FDCA3531" bold="true" box="[428,611,1269,1296]" pageId="34" pageNumber="495">
<specimenCode id="6B38187D9B1D1E42C7B7FB25FDF73531" box="[428,606,1269,1296]" collectionCode="MV" country="0" name="University of Montana Museum" pageId="34" pageNumber="495">AU09CTCR14</specimenCode>
,
</emphasis>
<specimenCode id="6B38187D9B1D1E42C471FB25FD203531" box="[618,649,1269,1296]" collectionCode="MV" country="0" name="University of Montana Museum" pageId="34" pageNumber="495">15</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="3B21B0069B1D1E42C6DCFACAFB7B35E2" blockId="34.[151,1437,1125,1619]" pageId="34" pageNumber="495">
Four larvae form a clade based on molecular data, supported by ordinations of mensural data versus CO1 genetic data (N. Herold
<emphasis id="09EA6C149B1D1E42C7B7FAEDFD983576" box="[428,561,1341,1367]" italics="true" pageId="34" pageNumber="495">unpubl. obs</emphasis>
.). Morphologically these resemble larvae of
<taxonomicName id="FC9ECB859B1D1E42C228FAEEFB113576" box="[1075,1208,1342,1367]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="495" phylum="Arthropoda" rank="species" species="cinctipes">
<emphasis id="09EA6C149B1D1E42C228FAEEFB113576" box="[1075,1208,1342,1367]" italics="true" pageId="34" pageNumber="495">R. cinctipes</emphasis>
</taxonomicName>
in the characteristically thinned cuticle on one side of antennal segment 3, beside, and perhaps associated with, the Lauterborn organs (
<figureCitation id="A3A5AC839B1D1E42C6BBFA55FF533581" box="[160,250,1413,1440]" captionStart="FIGURE 5" captionStartId="14.[151,250,1912,1937]" captionTargetBox="[292,1266,205,1851]" captionTargetId="figure@14.[272,1316,181,1886]" captionTargetPageId="14" captionText="FIGURE 5. Riethia spp. Larva. A. Dorsal head, R. cinctipes Freeman; B. Mentum, R. stictoptera Kieffer; C. Mandible, R. stictoptera. D-I. Antenna, D. R. azeylandica sp.n., E. R. cinctipes Freeman, F. R. donedwardi sp.n., G. R. neocaledonica sp.n., H. R. larval sp.B; I. R. stictoptera. Maxilla, J. R. neocaledonica, K. Pseudochironomus sp., L. Aedokritus coffeatus Trivinho- Strixino, M. Manoa pahayokeensis Jacobsen &amp; Perry, N. putative Megacentron Freeman, O. Manoa obscura Fittkau." httpUri="https://zenodo.org/record/3351068/files/figure.png" pageId="34" pageNumber="495">Fig. 5H</figureCitation>
). The major distinction is in antennal segment 4 being subequal to slightly shorter than the 2
<superScript id="CCEB1D4E9B1D1E42C31BFA54FAB935B3" attach="left" box="[1280,1296,1412,1426]" fontSize="6" pageId="34" pageNumber="495">nd</superScript>
(80100%), whereas in
<taxonomicName id="FC9ECB859B1D1E42C70CFA7AFED235E2" box="[279,379,1450,1475]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="495" phylum="Arthropoda" rank="species" species="cinctipes">
<emphasis id="09EA6C149B1D1E42C70CFA7AFED235E2" box="[279,379,1450,1475]" italics="true" pageId="34" pageNumber="495">cinctipes</emphasis>
</taxonomicName>
it is significantly and consistently shorter than the 2
<superScript id="CCEB1D4E9B1D1E42C5DFFA78FC7D3597" attach="left" box="[964,980,1448,1462]" fontSize="6" pageId="34" pageNumber="495">nd</superScript>
(by c. 60%) (
<tableCitation id="761C85BD9B1D1E42C276FA79FB6C35E2" box="[1133,1221,1449,1475]" captionStart="TABLE 2" captionStartId="32.[149,242,622,649]" captionTargetBox="[164,1976,660,1609]" captionText="TABLE 2. Morphometric data for larvae of Riethia Kieffer" httpUri="http://table.plazi.org/id/6FE1E08E9B1F1E40C68EFDBEFC8432A8" pageId="34" pageNumber="495" tableUuid="6FE1E08E9B1F1E40C68EFDBEFC8432A8">Table 2</tableCitation>
).
</paragraph>
<paragraph id="3B21B0069B1D1E42C68CFA1EFBBE35C6" blockId="34.[151,1437,1125,1619]" box="[151,1047,1485,1512]" pageId="34" pageNumber="495">
Larva
<typeStatus id="E4250EA49B1D1E42C6C4FA1EFEA635C9" box="[223,271,1486,1512]" pageId="34" pageNumber="495">type</typeStatus>
B is unreared, and cannot be associated due to sympatry with others.
</paragraph>
<paragraph id="3B21B0069B1D1E42C6DCFA21FF593672" blockId="34.[151,1437,1125,1619]" pageId="34" pageNumber="495">
Both molecular vouchers from Davies Ck. (
<collectionCode id="5D8F28C39B1D1E42C4B6FA21FD40362A" box="[685,745,1521,1547]" pageId="34" pageNumber="495">FNQ</collectionCode>
76, FNQ78) are from
<bibRefCitation id="5F0FCDF79B1D1E42C5EDFA21FB40362A" author="Cranston, P. S. &amp; Hardy, N. B. &amp; Morse, G. E." box="[1014,1257,1521,1548]" pageId="34" pageNumber="495" pagination="172 - 188" refId="ref39964" refString="Cranston, P. S., Hardy, N. B. &amp; Morse, G. E. (2011) A dated molecular phylogeny for the Chironomidae (Diptera). Systematic Entomology, 37, 172 - 188. https: // doi. org / 10.1111 / j. 1365 - 3113.2011.00603. x" type="journal article" year="2011">
Cranston
<emphasis id="09EA6C149B1D1E42C27AFA22FB3B362A" box="[1121,1170,1521,1547]" italics="true" pageId="34" pageNumber="495">et al</emphasis>
. (2011)
</bibRefCitation>
for
<taxonomicName id="FC9ECB859B1D1E42C30DFA21FACE362A" authorityName="Kieffer" authorityYear="1917" box="[1302,1383,1521,1547]" class="Insecta" family="Chironomidae" genus="Riethia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="495" phylum="Arthropoda" rank="genus">
<emphasis id="09EA6C149B1D1E42C30DFA21FACE362A" box="[1302,1383,1521,1547]" italics="true" pageId="34" pageNumber="495">Riethia</emphasis>
</taxonomicName>
species 3 and 4, separated morphologically due to earlier failure to recognize presence of 3
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and 4
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instars of the same species.
</paragraph>
</subSubSection>
</treatment>
</document>