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<document id="1D01D29E0257958EEB5A24078A9B47AA" ID-CLB-Dataset="4854" ID-DOI="10.5852/ejt.2021.763.1463" ID-GBIF-Dataset="31f73be9-4a90-4c0a-937e-784437eff4bd" ID-ISSN="2118-9773" ID-Zenodo-Dep="5235968" ID-ZooBank="9823AAD7-1113-434B-9882-1CF885DE7CED" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629723143846" checkinUser="felipe" docAuthor="Murao, Ryuki" docDate="2021" docId="941087F1CB42FFE4DCAF4991FAABCCFB" docLanguage="en" docName="ejt-763_1-74.pdf" docOrigin="European Journal of Taxonomy 763" docSource="http://dx.doi.org/10.5852/ejt.2021.763.1463" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Lasioglossum (Hemihalictus) ikudomei Murao 2021, sp. nov." docType="treatment" docUuid="2F66A01B-2176-4764-8DB7-59D53A335FB5" docUuidSource="ZooBank" docVersion="10" lastPageNumber="18" masterDocId="6829FF89CB4FFFF5DD544D6AF87ECE05" masterDocTitle="Redefinition of the sexstrigatus group of Lasioglossum (Hemihalictus) Cockerell, 1897 (Hymenoptera, Apoidea, Halictidae), with a revision of Japanese species" masterLastPageNumber="74" masterPageNumber="1" pageNumber="14" updateTime="1699140989011" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="3D127FAAAF67057A638DD940E8AB13B7">Redefinition of the sexstrigatus group of Lasioglossum (Hemihalictus) Cockerell, 1897 (Hymenoptera, Apoidea, Halictidae), with a revision of Japanese species</mods:title>
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<mods:namePart id="DA5D13392058007C73E1C92136C09FFC">Murao, Ryuki</mods:namePart>
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<mods:affiliation id="FB72890D528DF174A6CEC7D1AD651E3C">Regional Environmental Planning Co., Ltd, 1 - 10 - 12, Muromi, Sawara-ku, Fukuoka, 814 – 0015 Japan.</mods:affiliation>
<mods:nameIdentifier id="7127E6AE62AE02F7AD4C154A54E8266E" type="email">r.murao@mbr.nifty.com</mods:nameIdentifier>
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<mods:title id="211E2C7DEC4DE183744BCAD604206A95">European Journal of Taxonomy</mods:title>
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<mods:part id="168DAEB66C14F9EAC39C18C70E7A75A1">
<mods:date id="DA16B9CDF8069BB74A488FF833DFBB8F">2021</mods:date>
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<mods:number id="B95A9AD5AD45AC3D3EE1BAC9E0E48CDA">2021-08-18</mods:number>
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<treatment id="941087F1CB42FFE4DCAF4991FAABCCFB" ID-DOI="http://doi.org/10.5281/zenodo.5236035" ID-GBIF-Taxon="184930201" ID-Zenodo-Dep="5236035" LSID="urn:lsid:zoobank.org:act:2F66A01B-2176-4764-8DB7-59D53A335FB5" httpUri="http://treatment.plazi.org/id/941087F1CB42FFE4DCAF4991FAABCCFB" lastPageId="17" lastPageNumber="18" pageId="13" pageNumber="14">
<subSubSection id="54A3656CCB42FFF8DCAF4991FC47CB12" box="[507,1081,1275,1303]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="1C0636E7CB42FFF8DCAF4991FC47CB12" blockId="13.[371,1217,1275,1378]" box="[507,1081,1275,1303]" pageId="13" pageNumber="14">
<heading id="474E818BCB42FFF8DCAF4991FC47CB12" box="[507,1081,1275,1303]" centered="true" fontSize="11" level="2" pageId="13" pageNumber="14" reason="2">
<taxonomicName id="DBB94D64CB42FFF8DCAF4991FBA8CB13" authority="Murao, 2021" authorityName="Murao" authorityYear="2021" box="[507,982,1275,1302]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ikudomei" status="sp. nov." subGenus="Hemihalictus">
<emphasis id="2ECDEAF5CB42FFF8DCAF4991FBA8CB13" bold="true" box="[507,982,1275,1302]" pageId="13" pageNumber="14">
<emphasis id="2ECDEAF5CB42FFF8DCAF4991FADFCB13" bold="true" box="[507,673,1275,1302]" italics="true" pageId="13" pageNumber="14">Lasioglossum</emphasis>
(
<emphasis id="2ECDEAF5CB42FFF8DFE64991FB2BCB13" bold="true" box="[690,853,1275,1302]" italics="true" pageId="13" pageNumber="14">Hemihalictus</emphasis>
)
<emphasis id="2ECDEAF5CB42FFF8DE324991FBA8CB13" bold="true" box="[870,982,1275,1302]" italics="true" pageId="13" pageNumber="14">ikudomei</emphasis>
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35FE578ECB42FFF8DE8A4997FC47CB12" box="[990,1081,1277,1303]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="54A3656CCB42FFF8DC27484BFBE0CB64" pageId="13" pageNumber="14" type="description">
<paragraph id="1C0636E7CB42FFF8DC27484BFCBFCB39" blockId="13.[371,1217,1275,1378]" box="[371,1217,1313,1340]" pageId="13" pageNumber="14">
<uri id="68283AE5CB42FFF8DC27484BFCBFCB39" box="[371,1217,1313,1340]" pageId="13" pageNumber="14">
urn:lsid:zoobank.org:act:
<uuid id="681F0C32CB42FFF8DFC0484BFCBFCB39" box="[660,1217,1313,1340]" pageId="13" pageNumber="14">2F66A01B-2176-4764-8DB7-59D53A335FB5</uuid>
</uri>
</paragraph>
<paragraph id="1C0636E7CB42FFF8DFC1482DFBE0CB64" blockId="13.[371,1217,1275,1378]" box="[661,926,1351,1378]" pageId="13" pageNumber="14">
<figureCitation id="84822A62CB42FFF8DFC1482DFAA1CB67" box="[661,735,1351,1378]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[189,1398,502,1892]" captionTargetId="figure-130@14.[189,1398,502,1892]" captionTargetPageId="14" captionText="Fig. 5. Lasioglossum (Hemihalictus) ikudomei sp. nov. A. ♀, lateral habitus. B. ♀, head in frontal view. C. ♂, head in frontal view. D. ♀, labrum. E. ♀, mesoscutum. F. ♀, metapostnotum. Scale bars: A = 2 mm; B–C, E = 1 mm; D = 0.2 mm; F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5235978" httpUri="https://zenodo.org/record/5235978/files/figure.png" pageId="13" pageNumber="14">Figs 5</figureCitation>
,
<figureCitation id="84822A62CB42FFF8DFB8482DFB5ECB67" box="[748,800,1351,1378]" captionStart="Fig" captionStartId="53.[189,232,1859,1885]" captionTargetBox="[206,1391,272,1780]" captionTargetId="figure-71@53.[194,1399,265,1787]" captionTargetPageId="53" captionText="Fig. 15. 1st metasomal tergum, ♀. A. Lasioglossum (Hemihalictus) frigidum Sakagami &amp; Ebmer, 1996. B. L. (H.) ikudomei sp. nov. C. L. (H.) kiautschouense (Strand, 1910). D. L. (H.) ohei Hirashima &amp; Sakagami, 1966. E. L. (H.) simplicior (Cockerell, 1931). F. L. (H.) smilodon Ebmer &amp; Sakagami, 1994. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5236000" httpUri="https://zenodo.org/record/5236000/files/figure.png" pageId="13" pageNumber="14">15B</figureCitation>
,
<figureCitation id="84822A62CB42FFF8DE78482DFB1FCB64" box="[812,865,1351,1377]" captionStart="Fig" captionStartId="56.[189,232,1612,1638]" captionTargetBox="[188,1400,413,1548]" captionTargetId="figure-79@56.[189,1398,414,1546]" captionTargetPageId="56" captionText="Fig. 18. Distribution maps. A. Lasioglossum (Hemihalictus) sphecodicolor Sakagami &amp; Tadauchi, 1995. B. L. (H.) subsimplicior sp. nov. C. L. (H.) taeniolellum (Vachal, 1903). D. L. (H.) amamiense Ebmer &amp; Sakagami, 1994 (red circle), L. (H.) ikudomei sp. nov. (purple circle), L. (H.) smilodon Ebmer &amp; Sakagami, 1994 (light blue circle) and L. (H.) tadauchii Murao, 2012 (yellow circle)." figureDoi="http://doi.org/10.5281/zenodo.5236010" httpUri="https://zenodo.org/record/5236010/files/figure.png" pageId="13" pageNumber="14">18D</figureCitation>
,
<figureCitation id="84822A62CB42FFF8DE39482DFBE0CB64" box="[877,926,1351,1377]" captionStart="Fig" captionStartId="60.[189,232,1855,1881]" captionTargetBox="[352,1240,266,1820]" captionTargetId="figure-108@60.[340,1248,257,1826]" captionTargetPageId="60" captionText="Fig. 20. Female inner hind tibial spur. A. Lasioglossum (Hemihalictus) frigidum Sakagami &amp; Ebmer, 1996. B. L. (H.) ikudomei sp. nov. C. L. (H.) kiautschouense (Strand, 1910). D. L. (H.) ohei Hirashima &amp; Sakagami, 1966. E. L. (H.) smilodon Ebmer &amp; Sakagami, 1994. F. L. (H.) spectrum sp. nov. G. L. (H.) speculinum (Cockerell, 1925). H. L. (H.) sphecodicolor Sakagami &amp; Tadauchi, 1995. I. L. (H.) taeniolellum (Vachal, 1903). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5236014" httpUri="https://zenodo.org/record/5236014/files/figure.png" pageId="13" pageNumber="14">20B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="54A3656CCB42FFF8DDE948E0FC6BC86F" pageId="13" pageNumber="14" type="diagnosis">
<paragraph id="1C0636E7CB42FFF8DDE948E0F948CBA1" blockId="13.[189,310,1418,1444]" box="[189,310,1418,1444]" pageId="13" pageNumber="14">
<heading id="474E818BCB42FFF8DDE948E0F948CBA1" bold="true" box="[189,310,1418,1444]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="2ECDEAF5CB42FFF8DDE948E0F948CBA1" bold="true" box="[189,310,1418,1444]" pageId="13" pageNumber="14">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="1C0636E7CB42FFF8DDE948D1FC6BC86F" blockId="13.[187,1399,1467,1643]" pageId="13" pageNumber="14">
Females are similar to
<taxonomicName id="DBB94D64CB42FFF8DC8748D1FC99CBD0" authority="Ebmer &amp; Sakagami, 1994" authorityName="Ebmer &amp; Sakagami" authorityYear="1994" box="[467,1255,1467,1494]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="smilodon" subGenus="Hemihalictus">
<emphasis id="2ECDEAF5CB42FFF8DC8748D1FA0ACBD0" box="[467,628,1467,1493]" italics="true" pageId="13" pageNumber="14">Lasioglossum</emphasis>
(
<emphasis id="2ECDEAF5CB42FFF8DFDC48D1FB5ACBD0" box="[648,804,1467,1493]" italics="true" pageId="13" pageNumber="14">Hemihalictus</emphasis>
)
<emphasis id="2ECDEAF5CB42FFF8DE6C48D1FBDACBD0" box="[824,932,1467,1493]" italics="true" pageId="13" pageNumber="14">smilodon</emphasis>
<bibRefCitation id="78284B16CB42FFF8DEFA48D1FC99CBD0" author="Ebmer A. W. &amp; Maeta Y. &amp; Sakagami S. F." box="[942,1255,1467,1493]" pageId="13" pageNumber="14" pagination="23 - 36" refId="ref40317" refString="Ebmer A. W., Maeta Y. &amp; Sakagami S. F. 1994. Six new halictine bee species from southwest archipelago, Japan (Hymenoptera, Halictidae). Bulletin of the Faculty of Agriculture, Shimane University 28: 23 - 36." type="journal article" year="1994">Ebmer &amp; Sakagami, 1994</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="64AE7677CB42FFF8D86048D6FD09CBD3" box="[1332,1399,1468,1494]" name="Japan" pageId="13" pageNumber="14">Japan</collectingCountry>
but are separated from them by the supraclypeal area shinier (IS weakly tessellate), the PP on the mesoscutum sparser (IS=3 d in maximum) (
<figureCitation id="84822A62CB42FFF8DF924B6CFB5EC825" box="[710,800,1541,1568]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[189,1398,502,1892]" captionTargetId="figure-130@14.[189,1398,502,1892]" captionTargetPageId="14" captionText="Fig. 5. Lasioglossum (Hemihalictus) ikudomei sp. nov. A. ♀, lateral habitus. B. ♀, head in frontal view. C. ♂, head in frontal view. D. ♀, labrum. E. ♀, mesoscutum. F. ♀, metapostnotum. Scale bars: A = 2 mm; B–C, E = 1 mm; D = 0.2 mm; F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5235978" httpUri="https://zenodo.org/record/5235978/files/figure.png" pageId="13" pageNumber="14">Fig. 5E</figureCitation>
), and T1 with very weak lineolation. In contrast, in
<taxonomicName id="DBB94D64CB42FFF8DDE94B46F902C840" authorityName="Ebmer &amp; Sakagami" authorityYear="1994" box="[189,380,1579,1606]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="59" phylum="Arthropoda" rank="species" species="smilodon" subGenus="Hemihalictus">
<emphasis id="2ECDEAF5CB42FFF8DDE94B46F8B3C840" box="[189,205,1580,1605]" italics="true" pageId="13" pageNumber="14">L</emphasis>
. (
<emphasis id="2ECDEAF5CB42FFF8DDB04B46F887C840" box="[228,249,1580,1605]" italics="true" pageId="13" pageNumber="14">H</emphasis>
.)
<emphasis id="2ECDEAF5CB42FFF8DC444B41F902C840" box="[272,380,1579,1605]" italics="true" pageId="13" pageNumber="14">smilodon</emphasis>
</taxonomicName>
, the supraclypeal area is dimly shiny (IS distinctly tessellate), the PP on the mesoscutum denser (IS = 2 d at maximum) (
<figureCitation id="84822A62CB42FFF8DF724B3AFAFFC86E" box="[550,641,1616,1643]" captionStart="Fig" captionStartId="28.[189,232,1928,1954]" captionTargetBox="[189,1399,500,1890]" captionTargetId="figure-81@28.[189,1399,500,1890]" captionTargetPageId="28" captionText="Fig. 9. Lasioglossum (Hemihalictus) smilodon Ebmer &amp; Sakagami, 1994. A. ♀, lateral habitus. B. ♀, head in frontal view. C. ♂, head in frontal view. D. ♀, labrum. E. ♀, mesoscutum. F. ♀, metapostnotum. Scale bars: A = 2 mm; B = 1 mm; C, E–F = 0.5 mm; D = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5235988" httpUri="https://zenodo.org/record/5235988/files/figure.png" pageId="13" pageNumber="14">Fig. 9E</figureCitation>
), and the lineolation of T1 clearer.
</paragraph>
</subSubSection>
<subSubSection id="54A3656CCB42FFF8DDE94BFEFA9CC900" pageId="13" pageNumber="14" type="etymology">
<paragraph id="1C0636E7CB42FFF8DDE94BFEF93AC8AB" blockId="13.[189,324,1684,1710]" box="[189,324,1684,1710]" pageId="13" pageNumber="14">
<heading id="474E818BCB42FFF8DDE94BFEF93AC8AB" bold="true" box="[189,324,1684,1710]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="2ECDEAF5CB42FFF8DDE94BFEF93AC8AB" bold="true" box="[189,324,1684,1710]" pageId="13" pageNumber="14">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="1C0636E7CB42FFF8DDE94BAFFA9CC900" blockId="13.[189,1399,1733,1797]" pageId="13" pageNumber="14">
The specific name is dedicated to Dr Shuichi Ikudome (KWC), who contributed greatly to clarify the bee fauna of Ryukyus Islands, southwestern
<collectingCountry id="64AE7677CB42FFF8DFCC4B81FAA1C900" box="[664,735,1771,1797]" name="Japan" pageId="13" pageNumber="14">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="54A3656CCB42FFFADDE94A47FB9CCF6F" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="1C0636E7CB42FFF8DDE94A47F9D4C942" blockId="13.[189,426,1837,1863]" box="[189,426,1837,1863]" pageId="13" pageNumber="14">
<heading id="474E818BCB42FFF8DDE94A47F9D4C942" bold="true" box="[189,426,1837,1863]" fontSize="11" level="3" pageId="13" pageNumber="14" reason="3">
<emphasis id="2ECDEAF5CB42FFF8DDE94A47F9D4C942" bold="true" box="[189,426,1837,1863]" pageId="13" pageNumber="14">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="1C0636E7CB42FFF8DDBE4A34FB76C9EC" blockId="13.[185,1399,1886,2025]" pageId="13" pageNumber="14">
<materialsCitation id="ACD13CBACB42FFF8DDBE4A34FB76C9EC" ID-GBIF-Occurrence="3337333704" collectingDate="1992-07-03" collectionCode="ELKU" collectorName="S. Ikudome" country="Japan" county="Ryukyus" location="Henna-zaki" municipality="Miyako-jima" pageId="13" pageNumber="14" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa Pref." typeStatus="holotype">
<emphasis id="2ECDEAF5CB42FFF8DDBE4A34F922C97D" bold="true" box="[234,348,1886,1912]" pageId="13" pageNumber="14">
<typeStatus id="C3028845CB42FFF8DDBE4A34F922C97D" box="[234,348,1886,1912]" pageId="13" pageNumber="14" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="64AE7677CB42FFF8DDE94AEEF968C99B" box="[189,278,1924,1950]" name="Japan" pageId="13" pageNumber="14">JAPAN</collectingCountry>
–
<emphasis id="2ECDEAF5CB42FFF8DC644AEEF9E0C99B" bold="true" box="[304,414,1924,1950]" pageId="13" pageNumber="14">
<collectingCounty id="F5674E6BCB42FFF8DC644AEEF9E0C99B" box="[304,414,1924,1950]" pageId="13" pageNumber="14">Ryukyus</collectingCounty>
</emphasis>
•
<specimenCount id="0ABFFD6ECB42FFF8DCE24AEEF9B3C99B" box="[438,461,1924,1950]" pageId="13" pageNumber="14" type="female">♀</specimenCount>
;
<collectingRegion id="DE7DF805CB42FFF8DC8E4AEEFAF8C99B" box="[474,646,1924,1950]" country="Japan" name="Okinawa" pageId="13" pageNumber="14">Okinawa Pref.</collectingRegion>
,
<collectingMunicipality id="FC62AC9DCB42FFF8DFC44AEEFB52C99B" box="[656,812,1924,1950]" pageId="13" pageNumber="14">Miyako-jima</collectingMunicipality>
,
<location id="1966603CCB42FFF8DE634AEEFBBCC99B" LSID="urn:lsid:plazi:treatment:941087F1CB42FFE4DCAF4991FAABCCFB:1966603CCB42FFF8DE634AEEFBBCC99B" box="[823,962,1924,1950]" country="Japan" county="Ryukyus" municipality="Miyako-jima" name="Henna-zaki" pageId="13" pageNumber="14" stateProvince="Okinawa Pref.">Henna-zaki</location>
;
<date id="68071027CB42FFF8DE994AEEFC2EC99B" box="[973,1104,1924,1950]" pageId="13" pageNumber="14" value="1992-07-03">
<collectingDate id="7843E9CFCB42FFF8DE994AEEFC2EC99B" box="[973,1104,1924,1950]" pageId="13" pageNumber="14" value="1992-07-03">3 Jul. 1992</collectingDate>
</date>
;
<collectorName id="B14C5331CB42FFF8D90F4AEEFC9FC99B" box="[1115,1249,1924,1950]" pageId="13" pageNumber="14">S. Ikudome</collectorName>
leg.;
<collectionCode id="7AA8AE22CB42FFF8D8754AEEFD0DC99B" box="[1313,1395,1924,1950]" pageId="13" pageNumber="14">ELKU</collectionCode>
. [Verbatim label:MIYAKO-JIMA/ Henna-zaki/Okinawa Pref./
<date id="68071027CB42FFF8DED24AC3FC74C9C6" box="[902,1034,1961,1988]" pageId="13" pageNumber="14" value="1992-07-03">3.VII.1992</date>
/S.Ikudome leg.//HOLOTYPE //
<emphasis id="2ECDEAF5CB42FFF8DD814AA4F901C9ED" box="[213,383,1998,2024]" italics="true" pageId="13" pageNumber="14">Lasiioglossum</emphasis>
<taxonomicName id="DBB94D64CB42FFF8DCD14AA4FB7FC9EC" authority="Murao" authorityName="Murao" authorityYear="2021" box="[389,769,1998,2025]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ikudomei" subGenus="Hemihalictus">
(
<emphasis id="2ECDEAF5CB42FFF8DCDB4AA4FA55C9ED" box="[399,555,1998,2024]" italics="true" pageId="13" pageNumber="14">Hemihalictus</emphasis>
)
<emphasis id="2ECDEAF5CB42FFF8DF694AA4FAD7C9ED" box="[573,681,1998,2024]" italics="true" pageId="13" pageNumber="14">ikudomei</emphasis>
Murao
</taxonomicName>
]
</materialsCitation>
</paragraph>
<paragraph id="1C0636E7CB41FFFADDBE4C63FB9CCF6F" blockId="14.[189,1399,265,432]" lastBlockId="15.[189,1399,265,363]" lastPageId="15" lastPageNumber="16" pageId="14" pageNumber="15">
<materialsCitation id="ACD13CBACB41FFFBDDBE4C63FB91CF43" ID-GBIF-Occurrence="3337333359" collectingDate="1989-06-04" collectionCode="KWC" collectorName="Yoron Is." country="Japan" location="Yoron Is" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="1" stateProvince="Ryukyus" typeStatus="paratype">
<emphasis id="2ECDEAF5CB41FFFBDDBE4C63F919CF26" bold="true" box="[234,359,265,291]" pageId="14" pageNumber="15">
<typeStatus id="C3028845CB41FFFBDDBE4C63F919CF26" box="[234,359,265,291]" pageId="14" pageNumber="15" type="paratype">Paratypes</typeStatus>
</emphasis>
<collectingCountry id="64AE7677CB41FFFBDDE94C46F968CF43" box="[189,278,300,326]" name="Japan" pageId="14" pageNumber="15">JAPAN</collectingCountry>
–
<emphasis id="2ECDEAF5CB41FFFBDC664C46F9DECF43" bold="true" box="[306,416,300,326]" pageId="14" pageNumber="15">
<collectingRegion id="DE7DF805CB41FFFBDC664C46F9DECF43" box="[306,416,300,326]" pageId="14" pageNumber="15">Ryukyus</collectingRegion>
</emphasis>
•
<specimenCount id="0ABFFD6ECB41FFFBDCEC4C46F99BCF43" box="[440,485,300,326]" pageId="14" pageNumber="15" type="female">1 ♀</specimenCount>
;
<location id="1966603CCB41FFFBDCA64C46FA24CF43" LSID="urn:lsid:plazi:treatment:941087F1CB42FFE4DCAF4991FAABCCFB:1966603CCB41FFFBDCA64C46FA24CF43" box="[498,602,300,326]" country="Japan" name="Yoron Is" pageId="14" pageNumber="15" stateProvince="Ryukyus">Yoron Is</location>
.,
<location id="1966603CCB41FFFBDF3F4C46FB74CF43" LSID="urn:lsid:plazi:treatment:941087F1CB42FFE4DCAF4991FAABCCFB:1966603CCB41FFFBDF3F4C46FB74CF43" box="[619,778,300,326]" country="Japan" name="Maeno-hama" pageId="14" pageNumber="15" stateProvince="Ryukyus">Maeno-hama</location>
;
<date id="68071027CB41FFFBDE414C46FBDECF43" box="[789,928,300,326]" pageId="14" pageNumber="15" value="1989-06-04">
<collectingDate id="7843E9CFCB41FFFBDE414C46FBDECF43" box="[789,928,300,326]" pageId="14" pageNumber="15" value="1989-06-04">4 Jun. 1989</collectingDate>
</date>
;
<collectionCode id="7AA8AE22CB41FFFBDEFF4C46FB91CF43" box="[939,1007,300,326]" pageId="14" pageNumber="15">KWC</collectionCode>
</materialsCitation>
•
<materialsCitation id="ACD13CBACB41FFFBD95C4C46FAA8CF6C" ID-GBIF-Occurrence="3337333743" collectingDate="1979-04-05" collectionCode="ELKU" collectorName="K. Kusigemati" country="Japan" location="Hedo" municipality="Kunigami" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa Pref." typeStatus="paratype">
<specimenCount id="0ABFFD6ECB41FFFBD95C4C46FC4ACF43" box="[1032,1076,300,326]" pageId="14" pageNumber="15" type="female">1 ♀</specimenCount>
;
<collectingRegion id="DE7DF805CB41FFFBD9164C46FC91CF43" box="[1090,1263,300,326]" country="Japan" name="Okinawa" pageId="14" pageNumber="15">Okinawa Pref.</collectingRegion>
,
<collectingMunicipality id="FC62AC9DCB41FFFBD9AE4C46FD0CCF43" box="[1274,1394,300,326]" pageId="14" pageNumber="15">Kunigami</collectingMunicipality>
,
<location id="1966603CCB41FFFBDDE94C25F97ECF6C" LSID="urn:lsid:plazi:treatment:941087F1CB42FFE4DCAF4991FAABCCFB:1966603CCB41FFFBDDE94C25F97ECF6C" box="[189,256,335,361]" country="Japan" municipality="Kunigami" name="Hedo" pageId="14" pageNumber="15" stateProvince="Okinawa Pref.">Hedo</location>
;
<date id="68071027CB41FFFBDC5F4C25F9EBCF6C" box="[267,405,335,362]" pageId="14" pageNumber="15" value="1979-04-05">
<collectingDate id="7843E9CFCB41FFFBDC5F4C25F9EBCF6C" box="[267,405,335,362]" pageId="14" pageNumber="15" value="1979-04-05">5 Apr. 1979</collectingDate>
</date>
;
<collectorName id="B14C5331CB41FFFBDCF44C25FA37CF6C" box="[416,585,335,361]" pageId="14" pageNumber="15">K. Kusigemati</collectorName>
leg.;
<collectionCode id="7AA8AE22CB41FFFBDFDC4C25FAA8CF6C" box="[648,726,335,361]" pageId="14" pageNumber="15">ELKU</collectionCode>
</materialsCitation>
•
<materialsCitation id="ACD13CBACB41FFFBDFB94C25F975CF88" ID-GBIF-Occurrence="3337333763" collectingDate="1979-04-05" collectionCode="ELKU" collectorName="K. Ohara" country="Japan" location="Hedo" pageId="14" pageNumber="15" specimenCount="2" specimenCount-female="2" stateProvince="Okinawa Pref." typeStatus="paratype">
<specimenCount id="0ABFFD6ECB41FFFBDFB94C25FB50CF6C" box="[749,814,335,361]" pageId="14" pageNumber="15" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="DE7DF805CB41FFFBDE6F4C25FB99CF6C" box="[827,999,335,361]" country="Japan" name="Okinawa" pageId="14" pageNumber="15">Okinawa Pref.</collectingRegion>
,
<location id="1966603CCB41FFFBDEA54C25FC4ACF6C" LSID="urn:lsid:plazi:treatment:941087F1CB42FFE4DCAF4991FAABCCFB:1966603CCB41FFFBDEA54C25FC4ACF6C" box="[1009,1076,335,361]" country="Japan" name="Hedo" pageId="14" pageNumber="15" stateProvince="Okinawa Pref.">Hedo</location>
;
<date id="68071027CB41FFFBD96A4C25FCB6CF6C" box="[1086,1224,335,362]" pageId="14" pageNumber="15" value="1979-04-05">
<collectingDate id="7843E9CFCB41FFFBD96A4C25FCB6CF6C" box="[1086,1224,335,362]" pageId="14" pageNumber="15" value="1979-04-05">5 Apr. 1979</collectingDate>
</date>
;
<collectorName id="B14C5331CB41FFFBD9874C25FD40CF6C" box="[1235,1342,335,361]" pageId="14" pageNumber="15">K. Ohara</collectorName>
leg.;
<collectionCode id="7AA8AE22CB41FFFBDDE94C19F975CF88" box="[189,267,371,397]" pageId="14" pageNumber="15">ELKU</collectionCode>
</materialsCitation>
•
<materialsCitation id="ACD13CBACB41FFFBDC704C18F988CFB5" ID-GBIF-Occurrence="3337333595" collectingDate="2017-04-02" collectorName="K. Otsui" country="Japan" county="Kunigami-gun" latitude="26.866667" location="Cape Hedo" longLatPrecision="1239" longitude="128.25" municipality="Kunigami-son" pageId="14" pageNumber="15" specimenCount="2" specimenCount-female="2" stateProvince="Okinawa Pref." typeStatus="paratype">
<specimenCount id="0ABFFD6ECB41FFFBDC704C18F919CF88" box="[292,359,370,397]" pageId="14" pageNumber="15" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="DE7DF805CB41FFFBDC224C18FA5DCF89" box="[374,547,370,396]" country="Japan" name="Okinawa" pageId="14" pageNumber="15">Okinawa Pref.</collectingRegion>
,
<collectingCounty id="F5674E6BCB41FFFBDF7A4C18FAA2CF89" box="[558,732,370,396]" pageId="14" pageNumber="15">Kunigami-gun</collectingCounty>
,
<collectingMunicipality id="FC62AC9DCB41FFFBDFBC4C18FBEDCF89" box="[744,915,370,396]" pageId="14" pageNumber="15">Kunigami-son</collectingMunicipality>
,
<location id="1966603CCB41FFFBDECB4C18FC58CF89" LSID="urn:lsid:plazi:treatment:941087F1CB42FFE4DCAF4991FAABCCFB:1966603CCB41FFFBDECB4C18FC58CF89" box="[927,1062,370,396]" country="Japan" county="Kunigami-gun" latitude="26.866667" longLatPrecision="1239" longitude="128.25" municipality="Kunigami-son" name="Cape Hedo" pageId="14" pageNumber="15" stateProvince="Okinawa Pref.">Cape Hedo</location>
;
<geoCoordinate id="798D5020CB41FFFBD9664C1AFCE0CF88" box="[1074,1182,368,397]" degrees="26" direction="north" minutes="52" orientation="latitude" pageId="14" pageNumber="15" precision="925" value="26.866667">26°52′ N</geoCoordinate>
,
<geoCoordinate id="798D5020CB41FFFBD9FE4C1AFD5CCF88" box="[1194,1314,368,397]" degrees="128" direction="east" minutes="15" orientation="longitude" pageId="14" pageNumber="15" precision="925" value="128.25">128°15′ E</geoCoordinate>
;
<date id="68071027CB41FFFBD87B4C18F885CFAA" pageId="14" pageNumber="15" value="2017-04-02">
<collectingDate id="7843E9CFCB41FFFBD87B4C18F885CFAA" pageId="14" pageNumber="15" value="2017-04-02">2 Apr. 2017</collectingDate>
</date>
;
<collectorName id="B14C5331CB41FFFBDC5E4CFCF90ECFAA" box="[266,368,405,432]" pageId="14" pageNumber="15">K. Otsui</collectorName>
leg.; cMur
</materialsCitation>
•
<materialsCitation id="ACD13CBACB41FFFADF404CFFFAEDCF26" ID-GBIF-Occurrence="3337333524" collectingDate="2007-08-22" collectorName="Y. Nishimura" country="Japan" county="Okinawa-jima" lastPageId="15" lastPageNumber="16" location="Kunigami-son" municipality="Kunigami-gun" pageId="14" pageNumber="15" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa Pref." typeStatus="paratype">
<specimenCount id="0ABFFD6ECB41FFFBDF404CFFFA3DCFB5" box="[532,579,405,432]" pageId="14" pageNumber="15" type="female">1 ♀</specimenCount>
;
<collectingRegion id="DE7DF805CB41FFFBDF004CFFFB7DCFAA" box="[596,771,405,431]" country="Japan" name="Okinawa" pageId="14" pageNumber="15">Okinawa Pref.</collectingRegion>
,
<collectingCounty id="F5674E6BCB41FFFBDE454CFFFBC2CFAA" box="[785,956,405,431]" pageId="14" pageNumber="15">Okinawa-jima</collectingCounty>
,
<collectingMunicipality id="FC62AC9DCB41FFFBDE9E4CFFFC06CFAA" box="[970,1144,405,431]" pageId="14" pageNumber="15">Kunigami-gun</collectingMunicipality>
,
<location id="1966603CCB41FFFBD9D24CFFFD4FCFAA" LSID="urn:lsid:plazi:treatment:941087F1CB42FFE4DCAF4991FAABCCFB:1966603CCB41FFFBD9D24CFFFD4FCFAA" box="[1158,1329,405,431]" country="Japan" county="Okinawa-jima" municipality="Kunigami-gun" name="Kunigami-son" pageId="14" pageNumber="15" stateProvince="Okinawa Pref.">Kunigami-son</location>
,
<location id="1966603CCB41FFFBD8694CFFFD0CCFAA" LSID="urn:lsid:plazi:treatment:941087F1CB42FFE4DCAF4991FAABCCFB:1966603CCB41FFFBD8694CFFFD0CCFAA" box="[1341,1394,405,431]" country="Japan" county="Okinawa-jima" municipality="Kunigami-gun" name="Ada" pageId="14" pageNumber="15" stateProvince="Okinawa Pref.">Ada</location>
;
<date id="68071027CB40FFFADDE94C63F91DCF26" box="[189,355,265,291]" pageId="15" pageNumber="16" value="2007-08-22">
<collectingDate id="7843E9CFCB40FFFADDE94C63F91DCF26" box="[189,355,265,291]" pageId="15" pageNumber="16" value="2007-08-22">22 Aug. 2007</collectingDate>
</date>
;
<collectorName id="B14C5331CB40FFFADC244C63FA70CF26" box="[368,526,265,291]" pageId="15" pageNumber="16">Y. Nishimura</collectorName>
leg.; cMur
</materialsCitation>
•
<materialsCitation id="ACD13CBACB40FFFADFFB4C63F93BCF42" ID-GBIF-Occurrence="3337333574" collectingDate="1995-03-27" collectionCode="ELKU" collectorName="T. Matsumura" country="Japan" location="Miyako Is" pageId="15" pageNumber="16" specimenCount="1" specimenCount-female="1" stateProvince="Ryukyus" typeStatus="paratype">
<specimenCount id="0ABFFD6ECB40FFFADFFB4C63FAA0CF26" box="[687,734,265,291]" pageId="15" pageNumber="16" type="female">1 ♀</specimenCount>
;
<location id="1966603CCB40FFFADFBA4C63FB13CF26" LSID="urn:lsid:plazi:treatment:941087F1CB42FFE4DCAF4991FAABCCFB:1966603CCB40FFFADFBA4C63FB13CF26" box="[750,877,265,291]" country="Japan" name="Miyako Is" pageId="15" pageNumber="16" stateProvince="Ryukyus">Miyako Is</location>
.,
<location id="1966603CCB40FFFADED44C63FC6CCF26" LSID="urn:lsid:plazi:treatment:941087F1CB42FFE4DCAF4991FAABCCFB:1966603CCB40FFFADED44C63FC6CCF26" box="[896,1042,265,291]" country="Japan" name="Nishi-heana" pageId="15" pageNumber="16" stateProvince="Ryukyus">Nishi-heana</location>
;
<date id="68071027CB40FFFAD94B4C63FCBDCF21" box="[1055,1219,265,292]" pageId="15" pageNumber="16" value="1995-03-27">
<collectingDate id="7843E9CFCB40FFFAD94B4C63FCBDCF21" box="[1055,1219,265,292]" pageId="15" pageNumber="16" value="1995-03-27">27 Mar. 1995</collectingDate>
</date>
;
<collectorName id="B14C5331CB40FFFAD9854C63FD09CF26" box="[1233,1399,265,291]" pageId="15" pageNumber="16">T. Matsumura</collectorName>
leg.;
<collectionCode id="7AA8AE22CB40FFFADDA34C47F93BCF42" box="[247,325,301,327]" pageId="15" pageNumber="16">ELKU</collectionCode>
</materialsCitation>
•
<materialsCitation id="ACD13CBACB40FFFADC084C47FD52CF42" ID-GBIF-Occurrence="3337333348" box="[348,1324,301,327]" collectingDate="1992-07-03" collectionCode="ELKU" collectorName="S. Ikudome" country="Japan" location="Henna-zaki" municipality="Miyako-jima" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa Pref." typeStatus="paratype">
<specimenCount id="0ABFFD6ECB40FFFADC084C47F9F4CF42" box="[348,394,301,327]" pageId="15" pageNumber="16" type="male">1 ♂</specimenCount>
;
<collectingRegion id="DE7DF805CB40FFFADCC24C47FA3CCF42" box="[406,578,301,327]" country="Japan" name="Okinawa" pageId="15" pageNumber="16">Okinawa Pref.</collectingRegion>
,
<collectingMunicipality id="FC62AC9DCB40FFFADF184C47FA96CF42" box="[588,744,301,327]" pageId="15" pageNumber="16">Miyako-jima</collectingMunicipality>
,
<location id="1966603CCB40FFFADFA74C47FB00CF42" LSID="urn:lsid:plazi:treatment:941087F1CB42FFE4DCAF4991FAABCCFB:1966603CCB40FFFADFA74C47FB00CF42" box="[755,894,301,327]" country="Japan" municipality="Miyako-jima" name="Henna-zaki" pageId="15" pageNumber="16" stateProvince="Okinawa Pref.">Henna-zaki</location>
;
<date id="68071027CB40FFFADEDD4C47FC72CF42" box="[905,1036,301,327]" pageId="15" pageNumber="16" value="1992-07-03">
<collectingDate id="7843E9CFCB40FFFADEDD4C47FC72CF42" box="[905,1036,301,327]" pageId="15" pageNumber="16" value="1992-07-03">3 Jul. 1992</collectingDate>
</date>
;
<collectorName id="B14C5331CB40FFFAD9434C47FCE3CF42" box="[1047,1181,301,327]" pageId="15" pageNumber="16">S. Ikudome</collectorName>
leg.;
<collectionCode id="7AA8AE22CB40FFFAD98A4C47FD52CF42" box="[1246,1324,301,327]" pageId="15" pageNumber="16">ELKU</collectionCode>
</materialsCitation>
•
<materialsCitation id="ACD13CBACB40FFFAD8174C47FBA3CF6F" ID-GBIF-Occurrence="3337333660" collectingDate="2005-03-16" collectingDateMax="2005-03-17" collectingDateMin="2005-03-16" collectionCode="SULE" collectorName="Y. Maeta" country="Japan" location="Iriomote Is." pageId="15" pageNumber="16" specimenCount="1" specimenCount-female="1" stateProvince="Okinawa Pref." typeStatus="paratype">
<specimenCount id="0ABFFD6ECB40FFFAD8174C47FD11CF42" box="[1347,1391,301,327]" pageId="15" pageNumber="16" type="female">1 ♀</specimenCount>
;
<collectingRegion id="DE7DF805CB40FFFADDE94C3AF914CF6F" box="[189,362,336,362]" country="Japan" name="Okinawa" pageId="15" pageNumber="16">Okinawa Pref.</collectingRegion>
,
<location id="1966603CCB40FFFADC214C3AFA7FCF6E" LSID="urn:lsid:plazi:treatment:941087F1CB42FFE4DCAF4991FAABCCFB:1966603CCB40FFFADC214C3AFA7FCF6E" box="[373,513,336,363]" country="Japan" name="Iriomote Is." pageId="15" pageNumber="16" stateProvince="Okinawa Pref.">Iriomote Is.</location>
;
<date id="68071027CB40FFFADF594C3AFAA4CF6E" box="[525,730,336,363]" pageId="15" pageNumber="16" value="2005-03-16" valueMax="2005-03-17" valueMin="2005-03-16">
<collectingDate id="7843E9CFCB40FFFADF594C3AFAA4CF6E" box="[525,730,336,363]" pageId="15" pageNumber="16" value="2005-03-16" valueMax="2005-03-17" valueMin="2005-03-16">16–17 Mar. 2005</collectingDate>
</date>
;
<collectorName id="B14C5331CB40FFFADFB14C3BFB30CF6E" box="[741,846,337,363]" pageId="15" pageNumber="16">Y. Maeta</collectorName>
leg.;
<collectionCode id="7AA8AE22CB40FFFADEC44C3AFBA3CF6F" box="[912,989,336,362]" pageId="15" pageNumber="16">SULE</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="48C6666FCB41FFFBDDE94AE2FB79C9EC" ID-DOI="http://doi.org/10.5281/zenodo.5235978" ID-Zenodo-Dep="5235978" httpUri="https://zenodo.org/record/5235978/files/figure.png" pageId="14" pageNumber="15" startId="14.[189,232,1928,1954]" targetBox="[189,1398,502,1892]" targetPageId="14">
<paragraph id="1C0636E7CB41FFFBDDE94AE2FB79C9EC" blockId="14.[189,1399,1928,2026]" pageId="14" pageNumber="15">
<emphasis id="2ECDEAF5CB41FFFBDDE94AE2F974C9A6" bold="true" box="[189,266,1928,1955]" pageId="14" pageNumber="15">Fig. 5.</emphasis>
<taxonomicName id="DBB94D64CB41FFFBDC454AE2FAA3C9A7" authority="Murao, 2021" authorityName="Murao" authorityYear="2021" box="[273,733,1928,1955]" class="Insecta" family="Halictidae" genus="Lasioglossum" kingdom="Animalia" order="Hymenoptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="ikudomei" status="sp. nov." subGenus="Hemihalictus">
<emphasis id="2ECDEAF5CB41FFFBDC454AE2F9CCC9A7" box="[273,434,1928,1954]" italics="true" pageId="14" pageNumber="15">Lasioglossum</emphasis>
(
<emphasis id="2ECDEAF5CB41FFFBDC974AE2FA21C9A7" box="[451,607,1928,1954]" italics="true" pageId="14" pageNumber="15">Hemihalictus</emphasis>
)
<emphasis id="2ECDEAF5CB41FFFBDF254AE2FAA3C9A7" box="[625,733,1928,1954]" italics="true" pageId="14" pageNumber="15">ikudomei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="35FE578ECB41FFFBDFB74AE3FB40C9A6" box="[739,830,1929,1955]" pageId="14" pageNumber="15" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="2ECDEAF5CB41FFFBDE104AE2FB27C9A7" bold="true" box="[836,857,1928,1954]" pageId="14" pageNumber="15">A</emphasis>
. ♀, lateral habitus.
<emphasis id="2ECDEAF5CB41FFFBD9144AE3FC2AC9A6" bold="true" box="[1088,1108,1929,1955]" pageId="14" pageNumber="15">B</emphasis>
. ♀, head in frontal view.
<emphasis id="2ECDEAF5CB41FFFBDDE94AC1F8ACC9C0" bold="true" box="[189,210,1963,1989]" pageId="14" pageNumber="15">C</emphasis>
. ♂, head in frontal view.
<emphasis id="2ECDEAF5CB41FFFBDF584AC6FA5FC9C3" bold="true" box="[524,545,1964,1990]" pageId="14" pageNumber="15">D</emphasis>
. ♀, labrum.
<emphasis id="2ECDEAF5CB41FFFBDFEA4AC6FAACC9C3" bold="true" box="[702,722,1964,1990]" pageId="14" pageNumber="15">E</emphasis>
. ♀, mesoscutum.
<emphasis id="2ECDEAF5CB41FFFBDEFA4AC6FBBEC9C3" bold="true" box="[942,960,1964,1990]" pageId="14" pageNumber="15">F</emphasis>
. ♀, metapostnotum. Scale bars: A = 2 mm; B–C, E = 1 mm; D = 0.2 mm; F = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="54A3656CCB40FFE4DDE94CF8FBC7CF43" lastPageId="17" lastPageNumber="18" pageId="15" pageNumber="16" type="description">
<paragraph id="1C0636E7CB40FFFADDE94CF8F930CFA9" blockId="15.[189,334,402,428]" box="[189,334,402,428]" pageId="15" pageNumber="16">
<heading id="474E818BCB40FFFADDE94CF8F930CFA9" bold="true" box="[189,334,402,428]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
<emphasis id="2ECDEAF5CB40FFFADDE94CF8F930CFA9" bold="true" box="[189,334,402,428]" pageId="15" pageNumber="16">Description</emphasis>
</heading>
</paragraph>
<paragraph id="1C0636E7CB40FFFADDBE4CABF93BCFDE" blockId="15.[185,1399,449,797]" box="[234,325,449,475]" pageId="15" pageNumber="16">
<emphasis id="2ECDEAF5CB40FFFADDBE4CABF93BCFDE" bold="true" box="[234,325,449,475]" pageId="15" pageNumber="16">Female</emphasis>
</paragraph>
<paragraph id="1C0636E7CB40FFFADDE94C8CFBE6CD18" blockId="15.[185,1399,449,797]" pageId="15" pageNumber="16">MEASUREMENTS (n=5, unit mm). BL =5.13–5.63 (5.43± 0.23), WL= 4.25–4.88 (4.65 ± 0.26), HL=1.55– 1.65 (1.61 ±0.04), HW= 1.48–1.65 (1.57± 0.06), IOD =0.26–0.29 (0.28 ± 0.02), OOD=0.24–0.31 (0.26 ± 0.02), OCD =0.18–0.19 (0.19± 0.01, n=4), UOD =0.94–1.03 (0.98 ±0.04), MOD =1.10– 1.23 (1.15± 0.05), LOD =0.84–0.94 (0.88± 0.04), IAD =0.15–0.16 (0.16 ± 0.01), AOD=0.26–0.31 (0.28 ± 0.02), CAL =0.29–0.32 (0.31± 0.01), CPL=0.34–0.37 (0.35 ±0.01), EL=1.75–1.85 (1.80± 0.04), EW =0.39–0.45 (0.41± 0.04), GW=0.29–0.39 (0.35 ± 0.04), SPL=0.61–0.66 (0.64 ± 0.02), F1L=0.10 (0.10 ± 0.00), F2L =0.08 (0.08± 0.00), F3L= 0.08 (0.08± 0.00), F2W=0.13–0.15 (0.14 ± 0.01), MsW = 1.80–2.05 (1.93 ±0.09), SCL= 0.38–0.45 (0.43 ± 0.03), MNL=0.25–0.30 (0.27± 0.02), MPL =0.25–0.33 (0.28 ±0.03), MtW=1.75–2.10 (1.97 ± 0.13).</paragraph>
<paragraph id="1C0636E7CB40FFFADDE94E2FFD4FCDA3" blockId="15.[189,1399,836,935]" pageId="15" pageNumber="16">COLORATION. Body black except for the following parts: mandible reddish brown apically; flagellum brown or blackish brown ventrally; tegula yellowish brown translucent; tibial spur yellow; metasomal terga narrowly yellowish brown translucent apically. Wings transparent, veins and stigma brown.</paragraph>
<paragraph id="1C0636E7CB40FFFADDE94EA4F9CCCA51" blockId="15.[189,1399,974,1108]" pageId="15" pageNumber="16">PUBESCENCE. Body hairs whitish, and covered with erect and sparse straight or fine branched hairs except for the following parts: pronotum moderately densely tomentose dorsally and around lobe; hind trochanter, femur, and tibia mixed with plumose hairs, forming scopa. Disc of T1 without distinct short hairs on medial area.</paragraph>
<paragraph id="1C0636E7CB40FFFADDE94911FC4BCBFF" blockId="15.[188,1399,1147,1531]" pageId="15" pageNumber="16">
STRUCTURE AND SCULPTURE HEAD. Slightly longer than wide or nearly as long as wide; HW:HL=1:1.03. Vertex rounded in frontal view. MOD:UOD:LOD=1:0.85:0.76. IOD:OOD:OCD =1:0.95:0.69. IAD:AOD = 1:1.86. Ocellocular area with moderately dense PP, IS smooth (IS =0.5–3 d). Paraocular area and frons weakly shiny, with shallow reticulate PP. Supraclypeal area slightly convex, weakly shiny, with sparse PP, IS weakly tessellate (IS =1–4 d). CPL:CAL= 1:0.87. Clypeus nearly flat, with sparse PP over entire surface, IS very weakly tessellate on upper half or ⅓ and smooth on lower half or ⅔ (IS =1–5 d). EW:GW =1:0.86. Genal area to postgena with distinct straight ridges. Malar space linear. Occiput not carinate. Hypostomal carinae nearly parallel. Mandible bidentate. Labrum (
<figureCitation id="84822A62CB40FFFAD85E481FFD17CB95" box="[1290,1385,1397,1424]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[189,1398,502,1892]" captionTargetId="figure-130@14.[189,1398,502,1892]" captionTargetPageId="14" captionText="Fig. 5. Lasioglossum (Hemihalictus) ikudomei sp. nov. A. ♀, lateral habitus. B. ♀, head in frontal view. C. ♂, head in frontal view. D. ♀, labrum. E. ♀, mesoscutum. F. ♀, metapostnotum. Scale bars: A = 2 mm; B–C, E = 1 mm; D = 0.2 mm; F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5235978" httpUri="https://zenodo.org/record/5235978/files/figure.png" pageId="15" pageNumber="16">Fig. 5D</figureCitation>
): basal area approximately 1.9 × as wide as long; distal process approximately 0.6× as long as basal area, tongue-like, and without lateral projection; distal keel rounded, pointed apically. Antenna short, not reaching metasoma. F2L:F2W=1:1.72; flagellum nearly flattened ventrally.
</paragraph>
<paragraph id="1C0636E7CB40FFE5DDE94B48FAF1CF4D" blockId="15.[186,1400,1570,2025]" lastBlockId="16.[189,1398,265,329]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
THORAX. Dorsolateral angle of pronotum obtuse; lateral surface without ridges; lateral lobe rounded. Tegula ovoid, nearly smooth. Mesoscutum (
<figureCitation id="84822A62CB40FFFADF804B2CFB4FC865" box="[724,817,1605,1632]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[189,1398,502,1892]" captionTargetId="figure-130@14.[189,1398,502,1892]" captionTargetPageId="14" captionText="Fig. 5. Lasioglossum (Hemihalictus) ikudomei sp. nov. A. ♀, lateral habitus. B. ♀, head in frontal view. C. ♂, head in frontal view. D. ♀, labrum. E. ♀, mesoscutum. F. ♀, metapostnotum. Scale bars: A = 2 mm; B–C, E = 1 mm; D = 0.2 mm; F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5235978" httpUri="https://zenodo.org/record/5235978/files/figure.png" pageId="15" pageNumber="16">Fig. 5E</figureCitation>
) with moderately dense PP over entire surface; IS weakly tessellate over entire surface (particularly very weak on posterior area) (IS =0.5–3 d); parapsidal line a narrow groove. Mesoscutellum with 2–4 PP on submedian area and denser PP on marginal area; IS nearly smooth on submedian area and very weakly tessellate on marginal area (IS =2–5 d on submedian area, =0.5–2 d on marginal area). Metanotum weakly rugulose. Mesepisternum weakly shiny, with dense shallow PP on upper area and reticulate PP on lower area; IS nearly smooth on upper area (IS=0.5–1 d on upper area). SCL:MNL:MPL=1:0.62:0.66. Propodeum: metapostnotum (
<figureCitation id="84822A62CB40FFFAD8424A76FD13C932" box="[1302,1389,1820,1847]" captionStart="Fig" captionStartId="14.[189,232,1928,1954]" captionTargetBox="[189,1398,502,1892]" captionTargetId="figure-130@14.[189,1398,502,1892]" captionTargetPageId="14" captionText="Fig. 5. Lasioglossum (Hemihalictus) ikudomei sp. nov. A. ♀, lateral habitus. B. ♀, head in frontal view. C. ♂, head in frontal view. D. ♀, labrum. E. ♀, mesoscutum. F. ♀, metapostnotum. Scale bars: A = 2 mm; B–C, E = 1 mm; D = 0.2 mm; F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5235978" httpUri="https://zenodo.org/record/5235978/files/figure.png" pageId="15" pageNumber="16">Fig. 5F</figureCitation>
) gently inclined, with irregular sinuate ridges on anterior half in
<typeStatus id="C3028845CB40FFFADEFE4A55FC6FC95C" box="[938,1041,1855,1881]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
and
<specimenCount id="0ABFFD6ECB40FFFAD91E4A2AFC90C95F" box="[1098,1262,1856,1882]" pageId="15" pageNumber="16" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
(remaining
<specimenCount id="0ABFFD6ECB40FFFADDE94A09F918C97B" box="[189,358,1891,1918]" pageId="15" pageNumber="16" type="generic" typeStatus="paratypes">five paratypes</specimenCount>
with short longitudinal ridges), with weak tessellation on posterior half, and nearly smooth among ridges; junction between metapostnotum and posterior surface not carinate, with weak tessellation; lateral surface weakly rugulose; posterior surface with lateral carina on lower half, without oblique carina. Coxae usual shape, without tubercle. Fore trochanter narrow, longer than wide. Basitibial plate of hind leg carinate marginally. Inner hind tibial spur with slender 2–4 teeth as in
<figureCitation id="84822A62CB5FFFE5D9ED4C63FD61CF26" box="[1209,1311,265,291]" captionStart="Fig" captionStartId="60.[189,232,1855,1881]" captionTargetBox="[352,1240,266,1820]" captionTargetId="figure-108@60.[340,1248,257,1826]" captionTargetPageId="60" captionText="Fig. 20. Female inner hind tibial spur. A. Lasioglossum (Hemihalictus) frigidum Sakagami &amp; Ebmer, 1996. B. L. (H.) ikudomei sp. nov. C. L. (H.) kiautschouense (Strand, 1910). D. L. (H.) ohei Hirashima &amp; Sakagami, 1966. E. L. (H.) smilodon Ebmer &amp; Sakagami, 1994. F. L. (H.) spectrum sp. nov. G. L. (H.) speculinum (Cockerell, 1925). H. L. (H.) sphecodicolor Sakagami &amp; Tadauchi, 1995. I. L. (H.) taeniolellum (Vachal, 1903). Scale bars: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.5236014" httpUri="https://zenodo.org/record/5236014/files/figure.png" pageId="16" pageNumber="17">Fig. 20B</figureCitation>
(n= 7). Fore wing with three submarginal cells.
</paragraph>
<paragraph id="1C0636E7CB5FFFE5DDE94C18FC3CCFFE" blockId="16.[188,1401,369,507]" pageId="16" pageNumber="17">
ABDOMEN. Disc of T1 without distinct PP on medial area, and with very weak lineolation interrupted on medial area (
<figureCitation id="84822A62CB5FFFE5DCD74CFCF98ECFB4" box="[387,496,406,433]" captionStart="Fig" captionStartId="53.[189,232,1859,1885]" captionTargetBox="[206,1391,272,1780]" captionTargetId="figure-71@53.[194,1399,265,1787]" captionTargetPageId="53" captionText="Fig. 15. 1st metasomal tergum, ♀. A. Lasioglossum (Hemihalictus) frigidum Sakagami &amp; Ebmer, 1996. B. L. (H.) ikudomei sp. nov. C. L. (H.) kiautschouense (Strand, 1910). D. L. (H.) ohei Hirashima &amp; Sakagami, 1966. E. L. (H.) simplicior (Cockerell, 1931). F. L. (H.) smilodon Ebmer &amp; Sakagami, 1994. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5236000" httpUri="https://zenodo.org/record/5236000/files/figure.png" pageId="16" pageNumber="17">Fig. 15B</figureCitation>
). Disc of T2 nearly smooth on anterior to medial area in
<typeStatus id="C3028845CB5FFFE5D9FC4CFCFD71CFB5" box="[1192,1295,406,432]" pageId="16" pageNumber="17" type="holotype">holotype</typeStatus>
and
<specimenCount id="0ABFFD6ECB5FFFE5D8194CFDF94CCFD3" pageId="16" pageNumber="17" type="generic" typeStatus="paratypes">two paratypes</specimenCount>
, and with weak lineolation on posterior area (anterior and posterior area with weak lineolation in
<specimenCount id="0ABFFD6ECB5FFFE5DD8E4C8BF9F6CFFE" box="[218,392,481,507]" pageId="16" pageNumber="17" type="generic" typeStatus="paratypes">four paratypes</specimenCount>
). Discs of T3–T4 with weak lineolation over entire surface.
</paragraph>
<paragraph id="1C0636E7CB5FFFE5DDBE4F4EF954CC3B" blockId="16.[185,1399,548,723]" box="[234,298,548,574]" pageId="16" pageNumber="17">
<emphasis id="2ECDEAF5CB5FFFE5DDBE4F4EF954CC3B" bold="true" box="[234,298,548,574]" pageId="16" pageNumber="17">Male</emphasis>
</paragraph>
<paragraph id="1C0636E7CB5FFFE5DDE94F20FBDBCCD6" blockId="16.[185,1399,548,723]" pageId="16" pageNumber="17">MEASUREMENTS (n=1, unit mm). BL=4.23, WL=3.92, HL=1.56, HW=1.51, IOD =0.29, OOD =0.29, OCD =0.16, UOD=0.98, MOD =1.04, LOD =0.84, IAD =0.20, AOD= 0.24, CAL =0.31, CPL=0.33, EL = 1.07, EW= 0.40, GW=0.44, SPL=0.44, F1L=0.11, F2L=0.18, F3L= 0.16, F2W=0.13, MsW = 1.55, SCL=0.36, MNL=0.22, MPL=0.27, MtW=1.35.</paragraph>
<paragraph id="1C0636E7CB5FFFE5DDE94F96FB96CD80" blockId="16.[188,1399,764,902]" pageId="16" pageNumber="17">COLORATION. Body black except for the following parts: mandible yellowish brown except for apically reddish brown; labrum dark yellow; pronotal lobe yellowish brown; tegula yellowish brown translucent; legs brown, without distinct yellow marks; tibial spur yellow; metasomal terga broadly yellowish brown translucent apically. Wings transparent, veins and stigma pale brown.</paragraph>
<paragraph id="1C0636E7CB5FFFE5DDE94EC5FBD1CA3D" blockId="16.[189,1400,943,1080]" pageId="16" pageNumber="17">PUBESCENCE. Body hairs whitish, and covered with erect and sparse straight or fine branched hairs except for the following parts: lower paraocular area and pronotal dorsum to lobe sparsely tomentose. Disc of T1 with sparse hairs only on marginal area. Disc of T2–T4 with sparse short hairs over entire surface. T2–T3 with thin apical fimbriae, less distinct than in the female.</paragraph>
<paragraph id="1C0636E7CB5FFFE5DDE9490BFA21CBA1" blockId="16.[185,1401,1121,1444]" pageId="16" pageNumber="17">STRUCTURE AND SCULPTURE HEAD. Nearly as long as wide; HW:HL=1:1.03. Vertex rounded in frontal view. MOD:UOD:LOD =1:0.94:0.81. IOD:OOD:OCD =1:1:0.54. IAD:AOD =1:1.22. Ocellocular area with moderately dense PP, IS smooth (IS =1–4 d). Paraocular area and frons weakly shiny, with shallow reticulate PP. Supraclypeal area slightly convex, weakly shiny, with moderately dense PP, IS weakly tessellate (IS= 1–3 d). CPL:CAL= 1:0.93. Clypeus nearly flat, with sparse shallow PP over entire sutface; IS smooth (IS =1–6 d). EW:GW=1:1.11. Genal area on lower margin and postgena with straight ridges. Malar space linear. Hypostomal carinae nearly parallel. Mandible edentate. Labrum with basal elevation, but not examined in detail. Antenna short, not reaching metasoma. F2L:F2W =1:0.75; flagellum nearly flattened ventrally.</paragraph>
<paragraph id="1C0636E7CB5FFFE5DDE948A4FAF1C95E" blockId="16.[189,1400,1485,1883]" pageId="16" pageNumber="17">THORAX. Dorsolateral angle of pronotum obtuse; lateral surface without ridges; lateral lobe rounded. Tegula ovoid, nearly smooth. Mesoscutum with moderately dense PP over entire surface; IS weakly tessellate on anterior margin, nearly smooth on rest parts (IS =1.5–3 d); parapsidal line a narrow groove. Mesoscutellum with sparse PP over entire surface; IS smooth (IS=2.5–6 d). Metanotum weakly rugulose. Mesepisternum with moderately dense PP over entire surface; IS smooth (IS=1–3 d). SCL:MNL:MPL=1:0.63:0.75. Propodeum: metapostnotum weakly shiny and gently inclined, with short straight ridges occupying anterior ⅔; junction between metapostnotum and posterior surface not carinate, nearly smooth; lateral surface weakly reticulate; posterior surface nearly smooth, with lateral carina on lower ⅓, and without oblique carina. Coxae usual shape, without tubercle. Fore trochanter narrow, longer than wide. Basitibial plate of hind leg carinate marginally. Inner hind tibial spur serrate. Fore wing with three submarginal cells.</paragraph>
<paragraph id="1C0636E7CB5FFFE5DDE94AEEFB68C9EC" blockId="16.[189,1398,1924,2025]" pageId="16" pageNumber="17">ABDOMEN. Disc of T1 nearly smooth. Disc of T2–T3 with sparse fine PP; T2 weakly lineolate on apical margin; T3 weakly lioneolate on apical half. Disc of T4 weakly lineolate over entire surface. S7 with moderately long, apically rounded median process.</paragraph>
<paragraph id="1C0636E7CB5EFFE4DDE94C63FBC7CF43" blockId="17.[189,1398,265,327]" pageId="17" pageNumber="18">GENITALIA. Gonobase flat at bottom; gonocoxite smooth; ventral retrorse lobe tongue-like, moderately long but not reaching gonobase, with sparse short hairs ventrally.</paragraph>
</subSubSection>
<subSubSection id="54A3656CCB5EFFE4DDE94C04F892CFDE" pageId="17" pageNumber="18" type="distribution">
<paragraph id="1C0636E7CB5EFFE4DDE94C04F928CF8D" blockId="17.[189,342,366,392]" box="[189,342,366,392]" pageId="17" pageNumber="18">
<heading id="474E818BCB5EFFE4DDE94C04F928CF8D" bold="true" box="[189,342,366,392]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="2ECDEAF5CB5EFFE4DDE94C04F928CF8D" bold="true" box="[189,342,366,392]" pageId="17" pageNumber="18">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="1C0636E7CB5EFFE4DDE94CF4F892CFDE" blockId="17.[189,1399,413,475]" pageId="17" pageNumber="18">
<collectingCountry id="64AE7677CB5EFFE4DDE94CF4F97ECFBD" box="[189,256,414,440]" name="Japan" pageId="17" pageNumber="18">Japan</collectingCountry>
(central to southern Ryukyus: Yoron-jima Is., Okinawa-jima Is., Miyako-jima Is., Iriomote-jima Is.).
</paragraph>
</subSubSection>
<subSubSection id="54A3656CCB5EFFE4DDE94F68FAABCCFB" pageId="17" pageNumber="18" type="description">
<paragraph id="1C0636E7CB5EFFE4DDE94F68F91ECC19" blockId="17.[189,352,514,540]" box="[189,352,514,540]" pageId="17" pageNumber="18">
<heading id="474E818BCB5EFFE4DDE94F68F91ECC19" bold="true" box="[189,352,514,540]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="2ECDEAF5CB5EFFE4DDE94F68F91ECC19" bold="true" box="[189,352,514,540]" pageId="17" pageNumber="18">Flight period</emphasis>
</heading>
</paragraph>
<paragraph id="1C0636E7CB5EFFE4DDE94F5BFA04CC49" blockId="17.[189,634,561,588]" box="[189,634,561,588]" pageId="17" pageNumber="18">Female: middle March to late August.</paragraph>
<paragraph id="1C0636E7CB5EFFE4DDE94F19F93ECC88" blockId="17.[189,320,627,653]" box="[189,320,627,653]" pageId="17" pageNumber="18">Male: July.</paragraph>
<paragraph id="1C0636E7CB5EFFE4DDE94FDFF904CCCA" blockId="17.[189,378,693,719]" box="[189,378,693,719]" pageId="17" pageNumber="18">
<heading id="474E818BCB5EFFE4DDE94FDFF904CCCA" bold="true" box="[189,378,693,719]" fontSize="11" level="3" pageId="17" pageNumber="18" reason="3">
<emphasis id="2ECDEAF5CB5EFFE4DDE94FDFF904CCCA" bold="true" box="[189,378,693,719]" pageId="17" pageNumber="18">Flower records</emphasis>
</heading>
</paragraph>
<paragraph id="1C0636E7CB5EFFE4DDE94F8FFAABCCFB" blockId="17.[189,725,740,767]" box="[189,725,740,767]" pageId="17" pageNumber="18">
<taxonomicName id="DBB94D64CB5EFFE4DDE94F8FF910CCFB" box="[189,366,741,766]" class="Magnoliopsida" family="Asteraceae" genus="Ixeris" kingdom="Plantae" order="Asterales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="japonica">
<emphasis id="2ECDEAF5CB5EFFE4DDE94F8FF910CCFB" box="[189,366,741,766]" italics="true" pageId="17" pageNumber="18">Ixeris japonica</emphasis>
</taxonomicName>
(Burm.f.) Nakai (
<taxonomicName id="DBB94D64CB5EFFE4DF154F8EFAB7CCFB" box="[577,713,740,766]" class="Magnoliopsida" family="Asteraceae" kingdom="Plantae" order="Asterales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="family">Asteraceae</taxonomicName>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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