346 lines
34 KiB
XML
346 lines
34 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.259.2970" ID-GBIF-Dataset="37fcf036-b7b9-407b-a090-85e2cea21080" ID-PMC="PMC3591765" ID-Pensoft-Pub="1313-2970-259-1" ID-PubMed="23653501" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-259-1" ModsDocOrigin="ZooKeys 259" ModsDocTitle="A taxonomic revision of the Cymindis (Pinacodera) limbata species group (Coleoptera, Carabidae, Lebiini), including description of a new species from Florida, U.S.A." checkinTime="1451247765886" checkinUser="pensoft" docAuthor="Hunting, Wesley M." docDate="2013" docId="3906B31F8FC94F53EFA31BF26D7E2C52" docLanguage="en" docName="ZooKeys 259: 1-73" docOrigin="ZooKeys 259" docSource="http://dx.doi.org/10.3897/zookeys.259.2970" docTitle="Cymindis platicollis subsp. platicollis, stat. n." docType="treatment" docVersion="4" lastPageNumber="32" masterDocId="FF90904E7A57FFE9CD13EE7858701A20" masterDocTitle="A taxonomic revision of the Cymindis (Pinacodera) limbata species group (Coleoptera, Carabidae, Lebiini), including description of a new species from Florida, U. S. A." masterLastPageNumber="73" masterPageNumber="1" pageNumber="27" updateTime="1668155161763" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A taxonomic revision of the Cymindis (Pinacodera) limbata species group (Coleoptera, Carabidae, Lebiini), including description of a new species from Florida, U. S. A.</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hunting, Wesley M.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>259</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>73</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.259.2970</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.259.2970</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-259-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152039707" LSID="urn:lsid:plazi:treatment:3906B31F8FC94F53EFA31BF26D7E2C52" httpUri="http://treatment.plazi.org/id/3906B31F8FC94F53EFA31BF26D7E2C52" lastPageId="31" lastPageNumber="32" pageId="26" pageNumber="27">
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<subSubSection pageId="26" pageNumber="27" type="nomenclature">
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName LSID="http://species-id.net/wiki/Cymindis_platicollis_platicollis" authority="Say" class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis platicollis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="platicollis" subSpecies="platicollis">
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<pageBreakToken pageId="26" pageNumber="27" start="start">Cymindis</pageBreakToken>
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platicollis platicollis (Say)
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</taxonomicName>
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<taxonomicNameLabel pageId="26" pageNumber="27">stat. n.</taxonomicNameLabel>
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Figs 12-15, 17
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="27" type="reference_group">
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName class="Insecta" family="Carabidae" genus="Lebia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lebia platicollis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="platicollis">Lebia platicollis</taxonomicName>
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<bibRefCitation pageId="26" pageNumber="27">Say 1823</bibRefCitation>
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: 14. NEOTYPE male; TYPE LOCALITY - Allegheny, Allegheny County, Pennsylvania, U.S.A.; designated by
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<bibRefCitation author="Lindroth, CH" journalOrPublisher="Psyche" pageId="57" pageNumber="58" pagination="326 - 361" title="North American ground-beetles (Coleoptera, Carabidae, excluding Cicindelinae) described by Thomas Say: designation of lectotypes and neotypes." volume="76" year="1969">Lindroth and Freitag 1969</bibRefCitation>
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: 350.
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</paragraph>
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis platicollis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="platicollis">Cymindis platicollis</taxonomicName>
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;
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<bibRefCitation author="LeConte, JL" journalOrPublisher="Annals of the Lyceum of Natural History of New York" pageId="57" pageNumber="58" pagination="173 - 474" title="A descriptive catalogue of the geodephagous Coleoptera inhabiting the United States east of the Rocky Mountains." url="10.1111/j.1749-6632.1848.tb00277.x" volume="4" year="1848">LeConte 1848</bibRefCitation>
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: 189.
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</paragraph>
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis fuscata" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fuscata">Cymindis fuscata</taxonomicName>
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<bibRefCitation pageId="26" pageNumber="27">Dejean 1831</bibRefCitation>
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: 321. HOLOTYPE (probably), male in Chaudoir/
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<normalizedToken originalValue="Oberthür">Oberthuer</normalizedToken>
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collection, labeled:
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<normalizedToken originalValue="“Latreille”">"Latreille"</normalizedToken>
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[
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<normalizedToken originalValue="handwritten–">handwritten-</normalizedToken>
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green paper] [MNHP] -
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<bibRefCitation author="LeConte, JL" journalOrPublisher="Annals of the Lyceum of Natural History of New York" pageId="57" pageNumber="58" pagination="173 - 474" title="A descriptive catalogue of the geodephagous Coleoptera inhabiting the United States east of the Rocky Mountains." url="10.1111/j.1749-6632.1848.tb00277.x" volume="4" year="1848">LeConte 1848</bibRefCitation>
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: 189.
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</paragraph>
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera fuscata" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="fuscata">Pinacodera fuscata</taxonomicName>
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;
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<bibRefCitation author="Schaum, HR" journalOrPublisher="Naturgesichte der Insecten Deutschlands. Erste Abtheilung. Coleoptera Erste Band. Nicolaische Verlagsbuchhandlung, Berlin" pageId="57" pageNumber="58" title="(1857)." year="1856 - 1860">Schaum 1857</bibRefCitation>
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: 294. -
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<bibRefCitation pageId="26" pageNumber="27">Chaudoir 1875</bibRefCitation>
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: 3. -
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<bibRefCitation pageId="26" pageNumber="27">Horn 1882</bibRefCitation>
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: 162. -
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<bibRefCitation author="Leng, CW" journalOrPublisher="Bulletin of the Amerian Museum of Natural History" pageId="57" pageNumber="58" pagination="555 - 601" title="List of the Carabidae of Florida." volume="34" year="1915">Leng 1915</bibRefCitation>
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: 588. -
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<bibRefCitation pageId="26" pageNumber="27">Casey 1920</bibRefCitation>
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: 280. -
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<bibRefCitation pageId="26" pageNumber="27">Notman 1928</bibRefCitation>
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: 240. -
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<bibRefCitation author="Lindroth, CH" journalOrPublisher="Opuscula Entomologica Supplementum" pageId="57" pageNumber="58" pagination="1 - 160" title="The carabid beetles of Newfoundland including the French islands St. Pierre and Miquelon." volume="11" year="1955">Lindroth 1955</bibRefCitation>
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: 24. -
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<bibRefCitation author="Lindroth, CH" journalOrPublisher="Opuscula Entomologica, Supplementum" pageId="57" pageNumber="58" pagination="945 - 1192" title="The ground beetles (Carabidae excl. Cicindelinae) of Canada and Alaska. Part 6." volume="35" year="1969">1969</bibRefCitation>
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: 1068. -
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<bibRefCitation author="Kirk, VM" journalOrPublisher="South Carolina Agricultural Experiment Station Technical Bulletin" pageId="57" pageNumber="58" pagination="124 - 148" title="A list of the beetles of South Carolina Part 1 - northern coastal plain." volume="1033" year="1969">Kirk 1969</bibRefCitation>
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: 16. -
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<bibRefCitation author="Kirk, VM" journalOrPublisher="South Carolina Agricultural Experiment Station Technical Bulletin" pageId="57" pageNumber="58" pagination="1 - 117" title="A list of the beetles of South Carolina. Part 2 - mountain, piedmont, and southern coastal plain." volume="1038" year="1970">1970</bibRefCitation>
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: 17.
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</paragraph>
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera platycollis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="platycollis">Pinacodera platycollis</taxonomicName>
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;
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<bibRefCitation author="Schaum, HR" journalOrPublisher="Naturgesichte der Insecten Deutschlands. Erste Abtheilung. Coleoptera Erste Band. Nicolaische Verlagsbuchhandlung, Berlin" pageId="57" pageNumber="58" title="(1857)." year="1856 - 1860">Schaum 1857</bibRefCitation>
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: 294 (unjustified emendation) -
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<bibRefCitation pageId="26" pageNumber="27">Brimley 1938</bibRefCitation>
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:125.
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</paragraph>
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera platicollis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="platicollis">Pinacodera platicollis</taxonomicName>
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;
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<bibRefCitation pageId="26" pageNumber="27">Chaudoir 1875</bibRefCitation>
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: 2. -
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<bibRefCitation pageId="26" pageNumber="27">Horn 1882</bibRefCitation>
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: 147. -
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<bibRefCitation pageId="26" pageNumber="27">Blatchley 1910</bibRefCitation>
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: 152, 153. - Notman, 1928: 239. - Lindroth, 1969: 1068-1069. -
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<bibRefCitation author="Lindroth, CH" journalOrPublisher="Psyche" pageId="57" pageNumber="58" pagination="326 - 361" title="North American ground-beetles (Coleoptera, Carabidae, excluding Cicindelinae) described by Thomas Say: designation of lectotypes and neotypes." volume="76" year="1969">Lindroth and Freitag 1969</bibRefCitation>
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: 350. -
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<bibRefCitation author="Kirk, VM" journalOrPublisher="South Carolina Agricultural Experiment Station Technical Bulletin" pageId="57" pageNumber="58" pagination="124 - 148" title="A list of the beetles of South Carolina Part 1 - northern coastal plain." volume="1033" year="1969">Kirk 1969</bibRefCitation>
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: 16. -
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<bibRefCitation author="Kirk, VM" journalOrPublisher="South Carolina Agricultural Experiment Station Technical Bulletin" pageId="57" pageNumber="58" pagination="1 - 117" title="A list of the beetles of South Carolina. Part 2 - mountain, piedmont, and southern coastal plain." volume="1038" year="1970">1970</bibRefCitation>
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: 17. -
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<bibRefCitation pageId="26" pageNumber="27">Ciegler 2000</bibRefCitation>
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: 119.
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</paragraph>
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera punctigera" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="punctigera">Pinacodera punctigera</taxonomicName>
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(
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<bibRefCitation pageId="26" pageNumber="27">Dejean 1831</bibRefCitation>
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)
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<bibRefCitation author="Wickham, HF" journalOrPublisher="Bulletin from the Laboratories of Natural History of the State University of Iowa" pageId="57" pageNumber="58" pagination="96 - 115" title="The Coleoptera of the lower Rio Grande Valley." volume="4" year="1897">Wickham 1897</bibRefCitation>
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: 112 [not
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<bibRefCitation author="LeConte, JL" journalOrPublisher="Annals of the Lyceum of Natural History of New York" pageId="57" pageNumber="58" pagination="125 - 184" title="Descriptions of new species of Coleoptera, from California." url="10.1111/j.1749-6632.1852.tb00123.x" volume="5" year="1851">LeConte 1851</bibRefCitation>
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: 178].
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</paragraph>
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis planipennis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="planipennis">Cymindis planipennis</taxonomicName>
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;
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<bibRefCitation pageId="26" pageNumber="27">Casey 1913</bibRefCitation>
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: 189 [not
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<bibRefCitation author="LeConte, JL" journalOrPublisher="Smithsonian Miscellaneous Collections" pageId="57" pageNumber="58" pagination="1 - 92" title="New species of North American Coleoptera. Prepared for the Smithsonian Institution." volume="167" year="1863">LeConte 1863</bibRefCitation>
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: 6].
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</paragraph>
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera abbreviata" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="abbreviata">Pinacodera abbreviata</taxonomicName>
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<bibRefCitation pageId="26" pageNumber="27">Casey 1920</bibRefCitation>
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: 283. HOLOTYPE male labeled:
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<normalizedToken originalValue="“Col”">"Col"</normalizedToken>
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.; "Casey bequest 1925"; "TYPE USNM 47614" [red paper], "abbreviata Csy" [handwritten] [USNM]. TYPE AREA - Colorado, U.S.A. syn. n.
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</paragraph>
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera obscura" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="obscura">Pinacodera obscura</taxonomicName>
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<bibRefCitation pageId="26" pageNumber="27">Casey 1920</bibRefCitation>
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: 284. Female, labeled: "Southern Pines, A H Manee", N.C.; "Casey bequest, 1975"; "TYPE USNM 47612" [red paper]; "obscura Csy" [handwritten] [USNM]. -
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<bibRefCitation pageId="26" pageNumber="27">Brimley 1938</bibRefCitation>
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: 125. -
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<bibRefCitation pageId="26" pageNumber="27">Fattig 1949</bibRefCitation>
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: 40. syn. n.
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</paragraph>
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<paragraph pageId="26" pageNumber="27">
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera ampliata" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ampliata">Pinacodera ampliata</taxonomicName>
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<bibRefCitation pageId="26" pageNumber="27">Casey 1920</bibRefCitation>
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: 282. [=
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera planipennis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="planipennis">Pinacodera planipennis</taxonomicName>
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<bibRefCitation pageId="26" pageNumber="27">Casey 1913</bibRefCitation>
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].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="27" type="holotype">
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<paragraph pageId="26" pageNumber="27">Holotype</paragraph>
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<paragraph pageId="26" pageNumber="27">
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female, labeled:
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<normalizedToken originalValue="“Col”">"Col"</normalizedToken>
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; "Casey bequest 1925"; "TYPE USNM 47611" [red paper]; "ampliata Csy" [handwritten] [USNM].
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="27" type="diagnosis">
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<paragraph pageId="26" pageNumber="27">Diagnosis.</paragraph>
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<paragraph pageId="26" pageNumber="27">Specimens of this subspecies have uniformly colored head, pronotum, and elytra, with translucently bordered pronotum and elytra (Fig. 12).</paragraph>
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<caption pageId="26" pageNumber="27">
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<paragraph pageId="26" pageNumber="27">
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Figure 12. Dorsal habitus and color pattern of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis platicollis subsp. platicollis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="subSpecies" species="platicollis" subSpecies="platicollis">Cymindis platicollis platicollis</taxonomicName>
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(Say) (OBL 9.16 mm).
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection lastPageId="28" lastPageNumber="29" pageId="26" pageNumber="27" type="description">
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<paragraph pageId="26" pageNumber="27">Description.</paragraph>
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<paragraph pageId="26" pageNumber="27">
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With character states of subgenus
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<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">Pinacodera</taxonomicName>
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and species
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis platicollis" order="Coleoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="platicollis">Cymindis platicollis</taxonomicName>
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restricted as follows: OBL. 8.17 - 11.67 mm. Length (n= 20 males, 20 females): head 0.80 - 1.08, pronotum 1.56 - 2.36, elytra 4.92 - 6.83, metepisternum 1.10 - 1.70 mm; width: head 1.60 - 2.28, pronotum 2.00 - 3.32, elytra 3.33 - 5.17, metepisternum 0.60 - 0.84 mm.
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</paragraph>
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<paragraph pageId="26" pageNumber="27">Body proportions. HW/HL 1.83 - 2.26; PWM/PL 1.16 - 1.41; EL/EW 1.29- 1.49; ML/MW 1.71 - 2.07.</paragraph>
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<paragraph pageId="26" pageNumber="27">Color. Dorsal surface of head brown to rufo-piceous; pronotum and elytra brunneo-piceous to rufo-piceous, with pale, somewhat translucent margins. Antennae and palpi rufo-testaceous to brunneous palpi; elytral epipleura testaceous to rufo-testaceous; ventral thoracic sclerites and abdominal sterna testaceous to piceous.</paragraph>
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<paragraph pageId="27" pageNumber="28">
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<pageBreakToken pageId="27" pageNumber="28" start="start">Microsculpture</pageBreakToken>
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. Microlines not visible on dorsum of head capsule and pronotum at 50
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification. Elytra with mesh pattern isodiametric, microlines clearly defined throughout dorsal surface.
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</paragraph>
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<paragraph pageId="27" pageNumber="28">
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Macrosculpture and pilosity. Head capsule dorsally with fine, randomly scattered setigerous punctures (setae not visible or only barely so at 50
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<normalizedToken originalValue="×">x</normalizedToken>
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magnification) from constriction of neck extended anteriorly toward clypeus. Elytra with striae moderately impressed and punctulate throughout length; intervals slightly convex (few with greater convexity in intervals 1, 3 and 5); abdominal sterna with fine pilose punctures throughout.
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</paragraph>
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<paragraph pageId="27" pageNumber="28">Fixed setae. Elytra with 14-15 lateral (umbilical) setae;two setae on each of abdominal sterna III to VI; 4 setae along apical margin of sternum VII (Fig. 3).</paragraph>
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<paragraph pageId="27" pageNumber="28">Luster. Head capsule and pronotum glossyl elytra moderately glossy.</paragraph>
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<paragraph pageId="27" pageNumber="28">Pronotum. Anterior transverse impression shallow (Fig. 13); posterior transverse impression moderately deep; median longitudinal impression moderately shallow; posteriolateral angles almost right angled to obtuse.</paragraph>
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<paragraph pageId="28" pageNumber="29">
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<pageBreakToken pageId="28" pageNumber="29" start="start">Hind</pageBreakToken>
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wings. Macropterous.
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</paragraph>
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<paragraph pageId="28" pageNumber="29">Male genitalia. Phallus (Fig. 14A) length 1.70 - 2.42 mm.</paragraph>
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</subSubSection>
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<subSubSection pageId="28" pageNumber="29" type="variation">
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<paragraph pageId="28" pageNumber="29">Variation.</paragraph>
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<paragraph pageId="28" pageNumber="29">
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Through the range of
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<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis platicollis subsp. platicollis" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="platicollis" subSpecies="platicollis">Cymindis platicollis platicollis</taxonomicName>
|
||
a three-phased cline is observed (Fig. 17, Table 4). Phase 1: northeastern specimens have an average overall body length of 8.83 mm, average phallus length of 1.76 mm (Fig. 15A) and are dorsally glabrous; (~75%) with a single row of ~50-80 (interval 2 with more than 70) punctures in interval 1-7, some (~25%) with one to two rows of punctures in intervals 2, 4 and 6, all others with one row of punctures (except interval 8 with two to four rows). Phase 2: more southern specimens have an average overall body length of 9.22 mm, average phallus length 1.82 mm and have dorsal setation on humeral area of elytra; some specimens with few setae visible on dorsum of head and disc of elytra; (~86 %) with odd intervals bearing one row of scattered setigerous puncture and even intervals bearing two or three rows of scattered setigerous punctures, others (~14%) with interval 2 having two or three rows of setigerous punctures (rarely one row), interval 8 with two or three rows, and all other intervals bearing one row of setigerous punctures. Phase 3: southwestern specimens (extreme south west Oklahoma to mid-western Texas south west to Nuevo Leon) have an average overall body length of 9.92 mm, an average phallus length of 2.08 mm (Fig. 15B) and are dorsally setose, most individuals having one or two rows (mostly two) of moderately deep, randomly spaced, pilose punctures in all intervals, interval 8 with two to three rows. Few with various combinations of above.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="A5505682267A5DAF8A4E443874695F06" httpUri="http://table.plazi.org/id/A5505682267A5DAF8A4E443874695F06" pageId="28" pageNumber="29">
|
||
<paragraph pageId="28" pageNumber="29">
|
||
Table 4. Geographical variation in the extent of elytral punctation in
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis platicollis subsp. platicollis" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="platicollis" subSpecies="platicollis">Cymindis platicollis platicollis</taxonomicName>
|
||
(Say), by state (and regions of Texas) and number of individuals examined. Legend: 1, dorsal surface of elytra glabrous, punctation fine; 2, dorsal surface of elytra setose basally, punctation moderately fine; 3, entire dorsal surface of elytra setose, punctation coarse.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="28" pageNumber="29">
|
||
<table pageId="28" pageNumber="29">
|
||
<tr pageId="28" pageNumber="29">
|
||
<th colspan="1" pageId="28" pageNumber="29" rowspan="2">Locality</th>
|
||
<th colspan="1" pageId="28" pageNumber="29" rowspan="2">N</th>
|
||
<th colspan="3" pageId="28" pageNumber="29" rowspan="1">Punctation states / No. individuals</th>
|
||
</tr>
|
||
<tr pageId="28" pageNumber="29">
|
||
<th colspan="1" pageId="28" pageNumber="29" rowspan="1">1</th>
|
||
<th colspan="1" pageId="28" pageNumber="29" rowspan="1">2</th>
|
||
<th colspan="1" pageId="28" pageNumber="29" rowspan="1">3</th>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<caption pageId="28" pageNumber="29">
|
||
<paragraph pageId="28" pageNumber="29">
|
||
Figure 13. Pronotum, dorsal aspect, of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis platicollis subsp. platicollis" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="platicollis" subSpecies="platicollis">Cymindis platicollis platicollis</taxonomicName>
|
||
(Say).
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="28" pageNumber="29">
|
||
<paragraph pageId="28" pageNumber="29">
|
||
Figure 14. Structural features of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis platicollis subsp. platicollis" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="platicollis" subSpecies="platicollis">Cymindis platicollis platicollis</taxonomicName>
|
||
(Say): A, phallus and everted endophallus, right lateral aspect; B, female reproductive tract and ovipositor, ventral aspect. Legend: aa, apical area; bc, bursa copulatrix; bl, basal lobe; co, common oviduct; ep, endophallic plate; gc1, gonocoxite 1; gc2, gonocoxite 2; lt, lateral tergite; s, shaft; sd, spermathecal diverticulum; sg, spermathecal gland; sgd, spermathecal gland duct; sp, spermatheca.
|
||
</paragraph>
|
||
</caption>
|
||
<caption pageId="28" pageNumber="29">
|
||
<paragraph pageId="28" pageNumber="29">
|
||
Figure 15. Lateral aspect of phallus (sac everted) of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis platicollis subsp. platicollis" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="platicollis" subSpecies="platicollis">Cymindis platicollis platicollis</taxonomicName>
|
||
(Say), showing interpopulation variation of phallic apex texturing and difference in typical phallus size between northeastern (A) and southwestern (B) populations. Legend: aa, apical area; bl, basal lobe; s, shaft; obl, overall body length.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29" type="habitat, habits and seasonal occurrence">
|
||
<paragraph pageId="28" pageNumber="29">Habitat, habits and seasonal occurrence.</paragraph>
|
||
<paragraph pageId="28" pageNumber="29">
|
||
The known elevational range of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis platicollis subsp. platicollis" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="subSpecies" species="platicollis" subSpecies="platicollis">Cymindis platicollis platicollis</taxonomicName>
|
||
extends from 3 to 1935 m. Specimens were collected under stones, under and on bark of trees and associated mosses in forests of buckeye, beech-magnolia, elm, hackberry, hickory, juniper, mesquite, oak, oak-pine, and tamarack. It has been collected from shrub species
|
||
<taxonomicName class="Magnoliopsida" family="Fabaceae" genus="Leucaena" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Leucaena pulver" order="Fabales" pageId="28" pageNumber="29" phylum="Tracheophyta" rank="species" species="pulver">Leucaena pulver</taxonomicName>
|
||
(Schltdl.), from bromeliads associated with oak and also from the nest of woodrats,
|
||
<taxonomicName class="Mammalia" family="Cricetidae" genus="Neotoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neotoma micropus" order="Rodentia" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="micropus">Neotoma micropus</taxonomicName>
|
||
Baird. Adults are crepuscular, and most commonly collected from late February through July.
|
||
</paragraph>
|
||
<paragraph pageId="29" pageNumber="30">
|
||
<pageBreakToken pageId="29" pageNumber="30" start="start">I</pageBreakToken>
|
||
witnessed several pairs in copula over a three-week period of collecting in Georgia and Florida from late February to mid-March of 2008. Of these mated pairs I brought 9 females and 10 males back to the University of Alberta to attempt rearing larvae. I kept them all in a single plastic container with substrate from under the trees they were captured on. A wet ball of tissue provided moisture, and several 3rd to 5th instar larvae of cabbage looper moth species
|
||
<taxonomicName class="Insecta" family="Noctuidae" genus="Trichoplusia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Trichoplusia ni" order="Lepidoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="ni">Trichoplusia ni</taxonomicName>
|
||
(
|
||
<normalizedToken originalValue="Hübner">Huebner</normalizedToken>
|
||
) were introduced each week for food. All individuals (with the exception of one female) survived for the first three months in captivity. By mid-June (three months after capture) males started to die, and within the next two weeks all had expired. All of the remaining eight females lived at least until mid-September (6 months after capture) with the last individual dying in early November (7.5 months after capture). Males lived an average of 82 days after capture and females lived more than twice as long with an average lifespan of 166.5 days after capture.
|
||
</paragraph>
|
||
<paragraph pageId="29" pageNumber="30">
|
||
All beetles were removed from the container every week and the substrate was searched for eggs and larvae. No evidence of oviposition was found. Other attempts to rear larvae from mated
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Pinacodera</taxonomicName>
|
||
adults (Mahar, 1978) were also unsuccessful. Many lebiines are known to have unusual ovipositional habits or needs; that may also be the case in
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Pinacodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pinacodera" order="Coleoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="genus">Pinacodera</taxonomicName>
|
||
and a reason why rearing is problematic.
|
||
</paragraph>
|
||
<paragraph pageId="29" pageNumber="30">
|
||
Collecting methods include asafetida and molasses traps, sugar baits painted on tree trunks, beating and sweeping vegetation, at light and u.v. light, Lindgren funnel traps, Berlese traps, Malaise traps, flight intercept traps (
|
||
<normalizedToken originalValue="FIT’s">FIT's</normalizedToken>
|
||
), pitfall traps, hand collecting, and sticky traps.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30" type="geographical distribution">
|
||
<paragraph pageId="29" pageNumber="30">Geographical distribution.</paragraph>
|
||
<paragraph lastPageId="30" lastPageNumber="31" pageId="29" pageNumber="30">
|
||
The range of this subspecies (Fig. 17) extends in eastern Canada from southern Quebec west to southern Ontario; in the eastern United
|
||
<pageBreakToken pageId="30" pageNumber="31" start="start">States</pageBreakToken>
|
||
from Maine south to mid-Georgia west to eastern Colorado and Nebraska south to southern Texas. In Mexico it is known from Nuevo Leon in the northern portion of the Sierra Madre Oriental.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="30" pageNumber="31" type="evolutionary affinities">
|
||
<paragraph pageId="30" pageNumber="31">Evolutionary affinities.</paragraph>
|
||
<paragraph pageId="30" pageNumber="31">
|
||
This subspecies is, by definition the closest relative of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis platicollis subsp. atripennis" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="platicollis" subSpecies="atripennis">Cymindis platicollis atripennis</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="chorological affinities">
|
||
<paragraph pageId="30" pageNumber="31">Chorological affinities.</paragraph>
|
||
<paragraph lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis platicollis subsp. platicollis" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="platicollis" subSpecies="platicollis">Cymindis platicollis platicollis</taxonomicName>
|
||
is sympatric in portions of its range with
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis limbata" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="limbata">Cymindis limbata</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis complanata" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="complanata">Cymindis complanata</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis punctigera subsp. punctigera" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="punctigera" subSpecies="punctigera">Cymindis punctigera punctigera</taxonomicName>
|
||
, and
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis chevrolati" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="chevrolati">Cymindis chevrolati</taxonomicName>
|
||
. It is al
|
||
<pageBreakToken pageId="31" pageNumber="32" start="start">lopatric</pageBreakToken>
|
||
with
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis platicollis subsp. atripennis" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="platicollis" subSpecies="atripennis">Cymindis platicollis atripennis</taxonomicName>
|
||
,
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis rufostigma" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="rufostigma">Cymindis rufostigma</taxonomicName>
|
||
, and all other taxa in the limbata species group.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="31" pageNumber="32" type="material examined">
|
||
<paragraph pageId="31" pageNumber="32">Material examined.</paragraph>
|
||
<paragraph pageId="31" pageNumber="32">
|
||
I have examined 897 specimens of
|
||
<taxonomicName class="Insecta" family="Carabidae" genus="Cymindis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymindis platicollis subsp. platicollis" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="platicollis" subSpecies="platicollis">Cymindis platicollis platicollis</taxonomicName>
|
||
: 24 males and 17 females were dissected. For details see University of Alberta Strickland Virtual Entomology Museum Database (
|
||
<bibRefCitation pageId="31" pageNumber="32">University of Alberta 2009</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |