403 lines
45 KiB
XML
403 lines
45 KiB
XML
<document id="0B1FD43A52FFDF4F8A291440FC97C95D" ID-CLB-Dataset="7783" ID-DOI="10.5852/ejt.2021.738.1261" ID-GBIF-Dataset="02cf5931-b18e-4d58-a6ff-9c7b1e37e68d" ID-ISSN="2118-9773" ID-Zenodo-Dep="4594541" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615410255512" checkinUser="plazi" docAuthor="Turner, Ian M. & Cheek, Martin" docDate="2021" docId="355087D8FFF6E4711811FB97EEAEC0C0" docLanguage="en" docName="ejt-738_1-60.pdf" docOrigin="European Journal of Taxonomy 738" docSource="http://dx.doi.org/10.5852/ejt.2021.738.1261" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Chassalia calamus T. Y. Yu 2021, sp. nov." docType="treatment" docVersion="9" lastPageNumber="19" masterDocId="C969FFA0FFF9E4631A5DFFA8EF2BC42F" masterDocTitle="Revision of Chassalia (Rubiaceae-Rubioideae-Palicoureeae) in Borneo, with 14 new species" masterLastPageNumber="60" masterPageNumber="1" pageNumber="16" updateTime="1698932934732" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="9897A18ED10FE07564B421E698340FCD">
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<mods:title id="46359921A9B616057106D1288A752A73">Revision of Chassalia (Rubiaceae-Rubioideae-Palicoureeae) in Borneo, with 14 new species</mods:title>
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<mods:name id="BB7A0CE6DD76106FA55FB880D4CE022D" type="personal">
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<mods:roleTerm id="E5814F5CAC12AF139266B184BAAF94FA">Author</mods:roleTerm>
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<mods:namePart id="F074D3FC909C87713CCA0450238A2E34">Turner, Ian M.</mods:namePart>
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<mods:nameIdentifier id="8580C607FD6111F7AE606451E6F0440E" type="email">i.turner@kew.org</mods:nameIdentifier>
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<mods:namePart id="802BE3E7B8655640DC4809EFB6657998">Cheek, Martin</mods:namePart>
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<mods:affiliation id="B0555D9FA08EC741418FEDC8A3B29BC0">Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey TW 9 3 AE, UK. Queen Mary University of London, E 1 4 NS, UK. Singapore Botanical Liaison Officer, Herbarium, Royal Botanic Gardens, Kew, Richmond, Surrey TW 9 3 AE, UK. Singapore Botanic Gardens, National Parks Board, 1 Cluny Road, Singapore 259569, Singapore.</mods:affiliation>
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<mods:nameIdentifier id="8CDCE753F9FE5B8024EB9B26304518D0" type="email">m.cheek@kew.org</mods:nameIdentifier>
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<mods:typeOfResource id="2B7924EE0CEC779E1F1F325AE62B5758">text</mods:typeOfResource>
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<mods:title id="110AA2409BDD67024DEE0EA47BEB9291">European Journal of Taxonomy</mods:title>
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<mods:part id="F34C53102FF8F9255525207AE41DB0A5">
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<mods:date id="4F56FB81844976C815B1DF54AF6B8EC0">2021</mods:date>
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<mods:number id="76057BC82D0CFC29C64D3FFB52DBC80F">2021-03-09</mods:number>
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<mods:number id="D7299B51A1FFA58A331CD2C472C01122">738</mods:number>
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<mods:url id="82FBF60BDFCB483B05ACACF55830BD51">http://dx.doi.org/10.5852/ejt.2021.738.1261</mods:url>
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<mods:classification id="6F243F8468DCE1C01EFD0F0A14037AFA">journal article</mods:classification>
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<mods:identifier id="13EAB9BF89FF43AB0FF6C6ED5A5383D4" type="CLB-Dataset">7783</mods:identifier>
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<treatment id="355087D8FFF6E4711811FB97EEAEC0C0" ID-DOI="http://doi.org/10.5281/zenodo.4599385" ID-GBIF-Taxon="180327051" ID-Zenodo-Dep="4599385" LSID="urn:lsid:plazi:treatment:355087D8FFF6E4711811FB97EEAEC0C0" httpUri="http://treatment.plazi.org/id/355087D8FFF6E4711811FB97EEAEC0C0" lastPageId="18" lastPageNumber="19" pageId="15" pageNumber="16">
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<subSubSection id="F5E36545FFF6E46C1811FB97ECC3C075" box="[588,1000,1087,1114]" pageId="15" pageNumber="16" type="nomenclature">
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<paragraph id="BD4636CEFFF6E46C1811FB97ECC3C075" blockId="15.[583,1005,1087,1184]" box="[588,1000,1087,1114]" pageId="15" pageNumber="16">
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<heading id="E60E81A2FFF6E46C1811FB97ECC3C075" box="[588,1000,1087,1114]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="16" reason="2">
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<taxonomicName id="7AF94D4DFFF6E46C1811FB97ECACC075" ID-CoL="9V3MB" authority="T.Y.Yu" authorityName="T. Y. Yu" authorityYear="2021" box="[588,903,1087,1114]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="calamus" status="sp. nov.">
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<emphasis id="8F8DEADCFFF6E46C1811FB97EC04C075" bold="true" box="[588,815,1087,1114]" italics="true" pageId="15" pageNumber="16">Chassalia calamus</emphasis>
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T.Y.Yu
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</taxonomicName>
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<taxonomicNameLabel id="94BE57A7FFF6E46C19D3FBE8ECC3C075" box="[910,1000,1088,1114]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="F5E36545FFF6E46C181AFBCBEC67C08F" pageId="15" pageNumber="16" type="description">
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<paragraph id="BD4636CEFFF6E46C181AFBCBECC6C051" blockId="15.[583,1005,1087,1184]" box="[583,1005,1123,1150]" pageId="15" pageNumber="16">urn:lsid:ipni.org:names:77215404-1</paragraph>
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<paragraph id="BD4636CEFFF6E46C18BAFB2EEC67C08F" blockId="15.[583,1005,1087,1184]" box="[743,844,1158,1184]" pageId="15" pageNumber="16">
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<figureCitation id="25C22A4BFFF6E46C18BAFB2EEC67C08F" box="[743,844,1158,1184]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="16.[189,232,1894,1920]" captionStartId-1="17.[189,232,1998,2024]" captionTargetBox-0="[203,1342,275,1800]" captionTargetBox-1="[266,1321,265,1960]" captionTargetId-0="figure-14@16.[189,1398,262,1858]" captionTargetId-1="figure-11@17.[266,1321,265,1960]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="Fig. 8. Chassalia calamus T.Y.Yu sp. nov. A. Flowering shoot. B. Inflorescence. C. Longitudinal view of flower showing stamens, band of hairs, stigma, style, disc, calyx and hypanthium. D. Part of infructescence. E. Dry fruit. F. Pyrene ventral surface. From Asik Mantor SAN 118683. Drawn by T.Y. Yu." captionText-1="Fig. 9. Holotype of Chassalia calamus T.Y.Yu sp. nov. (Asik Mantor SAN 118683)." figureDoi-0="http://doi.org/10.5281/zenodo.4594561" figureDoi-1="http://doi.org/10.5281/zenodo.4594563" httpUri-0="https://zenodo.org/record/4594561/files/figure.png" httpUri-1="https://zenodo.org/record/4594563/files/figure.png" pageId="15" pageNumber="16">Figs 8–9</figureCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="F5E36545FFF6E46C1AE0FB6EEBCBC1BB" pageId="15" pageNumber="16" type="diagnosis">
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<paragraph id="BD4636CEFFF6E46C1AE0FB6EEE1DC0CF" blockId="15.[189,310,1222,1248]" box="[189,310,1222,1248]" pageId="15" pageNumber="16">
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||
<heading id="E60E81A2FFF6E46C1AE0FB6EEE1DC0CF" bold="true" box="[189,310,1222,1248]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
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<emphasis id="8F8DEADCFFF6E46C1AE0FB6EEE1DC0CF" bold="true" box="[189,310,1222,1248]" pageId="15" pageNumber="16">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="BD4636CEFFF6E46C1AE0FB5BEBCBC1BB" blockId="15.[188,1400,1267,1428]" pageId="15" pageNumber="16">
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Differs from other species in having a strong and thick, but hollow, stem
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<quantity id="7A019B2BFFF6E46C1E74FB5BEBB1C121" box="[1065,1178,1267,1294]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="15" pageNumber="16" unit="mm" value="8.5" valueMax="12.0" valueMin="5.0">5–12 mm</quantity>
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diameter (vs stem solid, ca
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<quantity id="7A019B2BFFF6E46C1B74FABCEEA0C100" box="[297,395,1300,1327]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="15" pageNumber="16" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
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diameter); leaves large 17–33 ×
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<quantity id="7A019B2BFFF6E46C1949FABCECBAC100" box="[788,913,1300,1327]" metricMagnitude="-2" metricUnit="m" metricValue="8.75" metricValueMax="12.0" metricValueMin="5.5" pageId="15" pageNumber="16" unit="cm" value="8.75" valueMax="12.0" valueMin="5.5">5.5–12 cm</quantity>
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(in most species except
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<taxonomicName id="7AF94D4DFFF6E46C1EE5FABDEA53C101" authority="Turner & Cheek, 2021" authorityName="Turner & Cheek" authorityYear="2021" box="[1208,1400,1301,1327]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="atropurpurea" status="sp. nov.">
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<emphasis id="8F8DEADCFFF6E46C1EE5FABDEA53C101" box="[1208,1400,1301,1327]" italics="true" pageId="15" pageNumber="16">C. atropurpurea</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="94BE57A7FFF6E46C1AE3FA9FEE30C17E" box="[190,283,1335,1361]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
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and
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<taxonomicName id="7AF94D4DFFF6E46C1B05FA9FED2FC17F" authority="Turner & Cheek, 2021" authorityName="Turner & Cheek" authorityYear="2021" box="[344,516,1334,1361]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="involucrata" status="sp. nov.">
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<emphasis id="8F8DEADCFFF6E46C1B05FA9FED2FC17F" box="[344,516,1334,1361]" italics="true" pageId="15" pageNumber="16">C. involucrata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="94BE57A7FFF6E46C1853FA9FED40C17E" box="[526,619,1335,1361]" pageId="15" pageNumber="16" rank="species">sp. nov.</taxonomicNameLabel>
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not reaching these dimensions); inflorescence usually very short with peduncle very short or absent, (<0.1–)
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<quantity id="7A019B2BFFF6E46C18E9FAF0EC13C15D" box="[692,824,1368,1395]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.5" metricValueMin="0.5" pageId="15" pageNumber="16" unit="cm" value="2.0" valueMax="3.5" valueMin="0.5">0.5–3.5 cm</quantity>
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(in other species usually exceeding
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<quantity id="7A019B2BFFF6E46C1E86FAF0EA01C15D" box="[1243,1322,1368,1395]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="15" pageNumber="16" unit="cm" value="3.5">3.5 cm</quantity>
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long); secondary peduncle usually very short to absent making the inflorescence highly compact.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="F5E36545FFF6E46C1AE0FA11EA5DC20D" pageId="15" pageNumber="16" type="etymology">
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<paragraph id="BD4636CEFFF6E46C1AE0FA11EE6FC1FC" blockId="15.[189,324,1465,1491]" box="[189,324,1465,1491]" pageId="15" pageNumber="16">
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||
<heading id="E60E81A2FFF6E46C1AE0FA11EE6FC1FC" bold="true" box="[189,324,1465,1491]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
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<emphasis id="8F8DEADCFFF6E46C1AE0FA11EE6FC1FC" bold="true" box="[189,324,1465,1491]" pageId="15" pageNumber="16">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="BD4636CEFFF6E46C1AE0FA4EEA5DC20D" blockId="15.[189,1398,1510,1570]" pageId="15" pageNumber="16">
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The epithet, treated as a noun in apposition, is derived from the Latin ʻ
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<taxonomicName id="7AF94D4DFFF6E46C19B8FA4EEB6CC22F" authorityName="T. Y. Yu" authorityYear="2021" box="[997,1095,1510,1536]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="species" species="calamus">
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<emphasis id="8F8DEADCFFF6E46C19B8FA4EEB6CC22F" box="[997,1095,1510,1536]" italics="true" pageId="15" pageNumber="16">calamus</emphasis>
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</taxonomicName>
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ʼ, a reed or more generally a hollow-stemmed plant. The name refers to the hollow stem, unusual in Bornean species of
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<taxonomicName id="7AF94D4DFFF6E46C1EA7F9A0EA44C20D" box="[1274,1391,1544,1570]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="genus">
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<emphasis id="8F8DEADCFFF6E46C1EA7F9A0EA44C20D" box="[1274,1391,1544,1570]" italics="true" pageId="15" pageNumber="16">Chassalia</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="F5E36545FFF6E46C1AE0F9E0EE06C3C7" pageId="15" pageNumber="16" type="materials_examined">
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<paragraph id="BD4636CEFFF6E46C1AE0F9E0EFD1C24D" blockId="15.[189,250,1608,1631]" box="[189,250,1608,1634]" pageId="15" pageNumber="16">
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||
<materialsCitation id="0D913C93FFF6E46C1AE0F9E0EFD1C24D" ID-GBIF-Occurrence="3050815328" box="[189,250,1608,1634]" collectingDate="1986-11-18" collectionCode="K" country="Malaysia" location="Borneo" municipality="Nabawan" pageId="15" pageNumber="16" specimenCode="K001129723" specimenCount="2" stateProvince="Sabah" typeStatus="holotype">
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||
<emphasis id="8F8DEADCFFF6E46C1AE0F9E0EFD1C24D" bold="true" box="[189,250,1608,1634]" pageId="15" pageNumber="16">
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||
<typeStatus id="6242886CFFF6E46C1AE0F9E0EFD1C24D" box="[189,250,1608,1634]" pageId="15" pageNumber="16">Type</typeStatus>
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||
</emphasis>
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||
</materialsCitation>
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||
</paragraph>
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<paragraph id="BD4636CEFFF6E46C1AE0F9DDEC9FC29F" blockId="15.[189,1399,1653,1713]" pageId="15" pageNumber="16">
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||
<collectingCountry id="C5EE765EFFF6E46C1AE0F9DDEE7FC2A0" box="[189,340,1653,1679]" name="Malaysia" pageId="15" pageNumber="16">MALAYSIA</collectingCountry>
|
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–
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<emphasis id="8F8DEADCFFF6E46C1B27F9DDED1DC2A1" bold="true" box="[378,566,1652,1679]" pageId="15" pageNumber="16">
|
||
<location id="B8266015FFF6E46C1B27F9DDEEFCC2A0" LSID="urn:lsid:plazi:treatment:355087D8FFF6E4711811FB97EEAEC0C0:B8266015FFF6E46C1B27F9DDEEFCC2A0" box="[378,471,1653,1679]" country="Malaysia" municipality="Nabawan" name="Borneo" pageId="15" pageNumber="16" stateProvince="Sabah">Borneo</location>
|
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,
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<collectingRegion id="7F3DF82CFFF6E46C1BB5F9DCED1DC2A1" box="[488,566,1652,1678]" country="Malaysia" name="Sabah" pageId="15" pageNumber="16">Sabah</collectingRegion>
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||
</emphasis>
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•
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<collectingMunicipality id="5D22ACB4FFF6E46C1805F9DCEDE1C2A1" box="[600,714,1652,1678]" pageId="15" pageNumber="16">Nabawan</collectingMunicipality>
|
||
,
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<location id="B8266015FFF6E46C1887F9DCECBBC2A1" LSID="urn:lsid:plazi:treatment:355087D8FFF6E4711811FB97EEAEC0C0:B8266015FFF6E46C1887F9DCECBBC2A1" box="[730,912,1652,1678]" country="Malaysia" municipality="Nabawan" name="Sungai Millian" pageId="15" pageNumber="16" stateProvince="Sabah">Sungai Millian</location>
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||
,
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<locationDeviation id="ACF42FECFFF6E46C19FCF9DCEB53C2A1" box="[929,1144,1652,1678]" pageId="15" pageNumber="16">along stream-side</locationDeviation>
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||
;
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||
<date id="C947100EFFF6E46C1ED4F9DDEA18C2A1" box="[1161,1331,1652,1679]" pageId="15" pageNumber="16" value="1986-11-18">
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||
<collectingDate id="D903E9E6FFF6E46C1ED4F9DDEA18C2A1" box="[1161,1331,1652,1679]" pageId="15" pageNumber="16" value="1986-11-18">18 Nov. 1986</collectingDate>
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||
</date>
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||
;
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<emphasis id="8F8DEADCFFF6E46C1F19F9DCEE85C29F" italics="true" pageId="15" pageNumber="16">Asik Mantor SAN 118683</emphasis>
|
||
;
|
||
<typeStatus id="6242886CFFF6E46C1BE0F93EED03C29F" box="[445,552,1686,1712]" pageId="15" pageNumber="16" type="holotype">holotype</typeStatus>
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||
: K[
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<specimenCode id="ED5F9EB5FFF6E46C180CF93EEDDCC29E" box="[593,759,1686,1713]" pageId="15" pageNumber="16">K001129723</specimenCode>
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||
]!;
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||
<materialsCitation id="0D913C93FFF6E46C1950F93EEC9BC29F" ID-GBIF-Occurrence="3050815332" box="[781,944,1686,1712]" pageId="15" pageNumber="16" specimenCount="1">isotype: SAN</materialsCitation>
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.
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</paragraph>
|
||
<paragraph id="BD4636CEFFF6E46C1AE0F97EEE9FC2DF" blockId="15.[189,436,1750,1776]" box="[189,436,1750,1776]" pageId="15" pageNumber="16">
|
||
<heading id="E60E81A2FFF6E46C1AE0F97EEE9FC2DF" bold="true" box="[189,436,1750,1776]" fontSize="11" level="3" pageId="15" pageNumber="16" reason="3">
|
||
<emphasis id="8F8DEADCFFF6E46C1AE0F97EEE9FC2DF" bold="true" box="[189,436,1750,1776]" pageId="15" pageNumber="16">Additional material</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="BD4636CEFFF6E46C1AE0F8ABEE06C3C7" blockId="15.[189,1399,1795,2024]" pageId="15" pageNumber="16">
|
||
<materialsCitation id="0D913C93FFF6E46C1AE0F8ABEC1FC310" ID-GBIF-Occurrence="3050815304" collectingDate="1986-08-23" collectionCode="KEP, SAN" collectorName="Sumbing Jimpin" country="Malaysia" county="Keningau" location="Borneo" municipality="Crocker" pageId="15" pageNumber="16" specimenCode="SAN 120566" specimenCount="1" stateProvince="Sabah">
|
||
<collectingCountry id="C5EE765EFFF6E46C1AE0F8ABEE7FC332" box="[189,340,1795,1821]" name="Malaysia" pageId="15" pageNumber="16">MALAYSIA</collectingCountry>
|
||
–
|
||
<emphasis id="8F8DEADCFFF6E46C1B26F8ABED13C332" bold="true" box="[379,568,1795,1821]" pageId="15" pageNumber="16">
|
||
<location id="B8266015FFF6E46C1B26F8ABEEF3C332" LSID="urn:lsid:plazi:treatment:355087D8FFF6E4711811FB97EEAEC0C0:B8266015FFF6E46C1B26F8ABEEF3C332" box="[379,472,1795,1821]" country="Malaysia" county="Keningau" municipality="Crocker" name="Borneo" pageId="15" pageNumber="16" stateProvince="Sabah">Borneo</location>
|
||
,
|
||
<collectingRegion id="7F3DF82CFFF6E46C1BB7F8ABED13C332" box="[490,568,1795,1821]" country="Malaysia" name="Sabah" pageId="15" pageNumber="16">Sabah</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<collectingCounty id="54274E42FFF6E46C1801F8ABEDFAC332" box="[604,721,1795,1821]" pageId="15" pageNumber="16">Keningau</collectingCounty>
|
||
,
|
||
<locationDeviation id="ACF42FECFFF6E46C18BFF8ABEA2FC332" box="[738,1284,1795,1821]" pageId="15" pageNumber="16">Crocker range area, mile 16, Kimanis Road</locationDeviation>
|
||
;
|
||
<date id="C947100EFFF6E46C1F49F8ABEFD0C311" pageId="15" pageNumber="16" value="1986-08-23">
|
||
<collectingDate id="D903E9E6FFF6E46C1F49F8ABEFD0C311" pageId="15" pageNumber="16" value="1986-08-23">23 Aug. 1986</collectingDate>
|
||
</date>
|
||
;
|
||
<emphasis id="8F8DEADCFFF6E46C1B54F88CED4CC311" box="[265,615,1828,1855]" italics="true" pageId="15" pageNumber="16">
|
||
<collectorName id="100C5318FFF6E46C1B54F88CEEE2C311" box="[265,457,1828,1854]" pageId="15" pageNumber="16">Sumbing Jimpin</collectorName>
|
||
<specimenCode id="ED5F9EB5FFF6E46C1B8CF88DED4CC311" box="[465,615,1828,1855]" collectionCode="SAN" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15310" lsid="urn:lsid:biocol.org:col:15310" name="Forest Research Centre" pageId="15" pageNumber="16" type="Herbarium">SAN 120566</specimenCode>
|
||
</emphasis>
|
||
; K!,
|
||
<collectionCode id="DBE8AE0BFFF6E46C18FAF88DEDCCC310" box="[679,743,1829,1855]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15093" lsid="urn:lsid:biocol.org:col:15093" name="Forest Research Institute Malaysia" pageId="15" pageNumber="16" type="Herbarium">KEP</collectionCode>
|
||
!,
|
||
<collectionCode id="DBE8AE0BFFF6E46C18A4F88DEC1FC310" box="[761,820,1829,1855]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15310" lsid="urn:lsid:biocol.org:col:15310" name="Forest Research Centre" pageId="15" pageNumber="16" type="Herbarium">SAN</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="0D913C93FFF6E46C1912F88CEB0EC34F" ID-GBIF-Occurrence="3050815346" collectingDate="1989-08-24" collectionCode="K, SAN" collectorName="Fidilis." country="Malaysia" county="Tambunan" elevation="1400" location="Tambunan" municipality="Sunsuron Km" pageId="15" pageNumber="16" specimenCode="SAN 127894" specimenCount="1" stateProvince="Borneo">
|
||
<collectingCounty id="54274E42FFF6E46C1912F88CECE5C311" box="[847,974,1828,1854]" pageId="15" pageNumber="16">Tambunan</collectingCounty>
|
||
,
|
||
<locationDeviation id="ACF42FECFFF6E46C1986F88DEE61C34E" pageId="15" pageNumber="16">Sunsuron Km 54 Jalan Tambunan/ Penampang</locationDeviation>
|
||
; alt.
|
||
<quantity id="7A019B2BFFF6E46C1BDFF8EEEEF5C34E" box="[386,478,1862,1889]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="15" pageNumber="16" unit="m" value="1400.0">
|
||
<elevation id="36D4D1FDFFF6E46C1BDFF8EEEEF5C34E" box="[386,478,1862,1889]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="15" pageNumber="16" unit="m" value="1400.0">1400 m</elevation>
|
||
</quantity>
|
||
;
|
||
<date id="C947100EFFF6E46C1BB6F8EEEDA5C34F" box="[491,654,1862,1888]" pageId="15" pageNumber="16" value="1989-08-24">
|
||
<collectingDate id="D903E9E6FFF6E46C1BB6F8EEEDA5C34F" box="[491,654,1862,1888]" pageId="15" pageNumber="16" value="1989-08-24">24 Aug. 1989</collectingDate>
|
||
</date>
|
||
;
|
||
<emphasis id="8F8DEADCFFF6E46C18C6F8EEEC86C34F" box="[667,941,1862,1888]" italics="true" pageId="15" pageNumber="16">
|
||
<collectorName id="100C5318FFF6E46C18C6F8EEEDC4C34F" box="[667,751,1862,1888]" pageId="15" pageNumber="16">Fidilis.</collectorName>
|
||
<collectionCode id="DBE8AE0BFFF6E46C18ABF8EFEC26C34F" box="[758,781,1863,1888]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="15" pageNumber="16" type="Herbarium">K</collectionCode>
|
||
.
|
||
<specimenCode id="ED5F9EB5FFF6E46C1944F8EEEC86C34F" box="[793,941,1862,1888]" collectionCode="SAN" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15310" lsid="urn:lsid:biocol.org:col:15310" name="Forest Research Centre" pageId="15" pageNumber="16" type="Herbarium">SAN 127894</specimenCode>
|
||
</emphasis>
|
||
;
|
||
<collectionCode id="DBE8AE0BFFF6E46C19E1F8EEECF3C34F" box="[956,984,1862,1888]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="15" pageNumber="16" type="Herbarium">K</collectionCode>
|
||
!,
|
||
<collectionCode id="DBE8AE0BFFF6E46C19B7F8EEEB0EC34F" box="[1002,1061,1862,1888]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15310" lsid="urn:lsid:biocol.org:col:15310" name="Forest Research Centre" pageId="15" pageNumber="16" type="Herbarium">SAN</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="0D913C93FFF6E46C1E63F8EFEE0DC38C" ID-GBIF-Occurrence="3050815324" collectionCode="SAN" collectorName="Saw" country="Malaysia" county="Ranau" elevation="343" latitude="5.8791666" location="Ulu Tungud Forest Reserve" longLatPrecision="21" longitude="116.92444" municipality="Ulu Tungud Forest Reserve" pageId="15" pageNumber="16" specimenCode="SAN 146079" specimenCount="1" stateProvince="Borneo">
|
||
<collectingCounty id="54274E42FFF6E46C1E63F8EFEBA7C34E" box="[1086,1164,1863,1889]" pageId="15" pageNumber="16">Ranau</collectingCounty>
|
||
,
|
||
<location id="B8266015FFF6E46C1EC5F8EEEE35C3AD" LSID="urn:lsid:plazi:treatment:355087D8FFF6E4711811FB97EEAEC0C0:B8266015FFF6E46C1EC5F8EEEE35C3AD" country="Malaysia" county="Ranau" latitude="5.8791666" longLatPrecision="21" longitude="116.92444" municipality="Ulu Tungud Forest Reserve" name="Ulu Tungud Forest Reserve" pageId="15" pageNumber="16" stateProvince="Borneo">Ulu Tungud Forest Reserve</location>
|
||
,
|
||
<location id="B8266015FFF6E46C1B76F8C0EDBBC3AD" LSID="urn:lsid:plazi:treatment:355087D8FFF6E4711811FB97EEAEC0C0:B8266015FFF6E46C1B76F8C0EDBBC3AD" box="[299,656,1896,1922]" country="Malaysia" county="Ranau" latitude="5.8791666" longLatPrecision="21" longitude="116.92444" municipality="Ulu Tungud Forest Reserve" name="Gunung Monkobo Expedition" pageId="15" pageNumber="16" stateProvince="Borneo">Gunung Monkobo Expedition</location>
|
||
;
|
||
<geoCoordinate id="D8CD5009FFF6E46C18C1F8CEEC0FC3AD" box="[668,804,1894,1922]" degrees="5" direction="north" minutes="52" orientation="latitude" pageId="15" pageNumber="16" precision="15" seconds="45" value="5.8791666">5°52′45″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="D8CD5009FFF6E46C196DF8CEECFAC3AD" box="[816,977,1894,1922]" degrees="116" direction="east" minutes="55" orientation="longitude" pageId="15" pageNumber="16" precision="15" seconds="28" value="116.92444">116°55′28″ E</geoCoordinate>
|
||
; alt.
|
||
<quantity id="7A019B2BFFF6E46C1E57F8C0EB73C3AD" box="[1034,1112,1896,1922]" metricMagnitude="2" metricUnit="m" metricValue="3.4299999999999997" pageId="15" pageNumber="16" unit="m" value="343.0">
|
||
<elevation id="36D4D1FDFFF6E46C1E57F8C0EB73C3AD" box="[1034,1112,1896,1922]" metricMagnitude="2" metricUnit="m" metricValue="3.4299999999999997" pageId="15" pageNumber="16" unit="m" value="343.0">343 m</elevation>
|
||
</quantity>
|
||
;
|
||
<emphasis id="8F8DEADCFFF6E46C1E38F8C0EA45C3AD" box="[1125,1390,1896,1922]" italics="true" pageId="15" pageNumber="16">
|
||
Saw L.G.
|
||
<specimenCode id="ED5F9EB5FFF6E46C1E87F8C0EA45C3AD" box="[1242,1390,1896,1922]" collectionCode="SAN" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15310" lsid="urn:lsid:biocol.org:col:15310" name="Forest Research Centre" pageId="15" pageNumber="16" type="Herbarium">SAN 146079</specimenCode>
|
||
</emphasis>
|
||
; K!,
|
||
<collectionCode id="DBE8AE0BFFF6E46C1AB6F821EE0DC38C" box="[235,294,1929,1955]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15310" lsid="urn:lsid:biocol.org:col:15310" name="Forest Research Centre" pageId="15" pageNumber="16" type="Herbarium">SAN</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="0D913C93FFF6E46C1B1DF821EEFCC3EA" ID-GBIF-Occurrence="3050815372" collectingDate="2001-09-02" collectionCode="SAN" collectorName="Pereira" country="Malaysia" county="Tambunan" location="Rafflesia Information Centre" municipality="Rafflesia Information Centre" pageId="15" pageNumber="16" specimenCount="1" stateProvince="Borneo">
|
||
<collectingCounty id="54274E42FFF6E46C1B1DF821EE94C38C" box="[320,447,1929,1955]" pageId="15" pageNumber="16">Tambunan</collectingCounty>
|
||
,
|
||
<location id="B8266015FFF6E46C1B96F821EC34C38C" LSID="urn:lsid:plazi:treatment:355087D8FFF6E4711811FB97EEAEC0C0:B8266015FFF6E46C1B96F821EC34C38C" box="[459,799,1929,1955]" country="Malaysia" county="Tambunan" municipality="Rafflesia Information Centre" name="Rafflesia Information Centre" pageId="15" pageNumber="16" stateProvince="Borneo">Rafflesia Information Centre</location>
|
||
,
|
||
<location id="B8266015FFF6E46C1976F821EB69C38B" LSID="urn:lsid:plazi:treatment:355087D8FFF6E4711811FB97EEAEC0C0:B8266015FFF6E46C1976F821EB69C38B" box="[811,1090,1929,1956]" country="Malaysia" county="Tambunan" municipality="Rafflesia Information Centre" name="Hill Dipterocarp Forest" pageId="15" pageNumber="16" stateProvince="Borneo">Hill Dipterocarp Forest</location>
|
||
;
|
||
<date id="C947100EFFF6E46C1E12F821EBF4C38C" box="[1103,1247,1929,1955]" pageId="15" pageNumber="16" value="2001-09-02">
|
||
<collectingDate id="D903E9E6FFF6E46C1E12F821EBF4C38C" box="[1103,1247,1929,1955]" pageId="15" pageNumber="16" value="2001-09-02">2 Sep. 2001</collectingDate>
|
||
</date>
|
||
;
|
||
<emphasis id="8F8DEADCFFF6E46C1EB1F822EE74C3EA" italics="true" pageId="15" pageNumber="16">J.T. Pereira et al. JTP 845</emphasis>
|
||
; K!,
|
||
<collectionCode id="DBE8AE0BFFF6E46C1BC1F803EEFCC3EA" box="[412,471,1963,1989]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15310" lsid="urn:lsid:biocol.org:col:15310" name="Forest Research Centre" pageId="15" pageNumber="16" type="Herbarium">SAN</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="0D913C93FFF6E46C1BB3F803EE02C3C7" ID-GBIF-Occurrence="3050815331" collectingDate="1987-02-25" collectionCode="SAN" collectorName="Sumbing Jimpin" country="Malaysia" location="Sungai Pamentarian" municipality="Pandewan" pageId="15" pageNumber="16" specimenCode="SAN 120057" specimenCount="1" stateProvince="Borneo">
|
||
<collectingMunicipality id="5D22ACB4FFF6E46C1BB3F803ED41C3EA" box="[494,618,1963,1989]" pageId="15" pageNumber="16">Pandewan</collectingMunicipality>
|
||
,
|
||
<location id="B8266015FFF6E46C1828F803EC4FC3EA" LSID="urn:lsid:plazi:treatment:355087D8FFF6E4711811FB97EEAEC0C0:B8266015FFF6E46C1828F803EC4FC3EA" box="[629,868,1963,1989]" country="Malaysia" municipality="Pandewan" name="Sungai Pamentarian" pageId="15" pageNumber="16" stateProvince="Borneo">Sungai Pamentarian</location>
|
||
;
|
||
<date id="C947100EFFF6E46C1932F803EB20C3EA" box="[879,1035,1963,1990]" pageId="15" pageNumber="16" value="1987-02-25">
|
||
<collectingDate id="D903E9E6FFF6E46C1932F803EB20C3EA" box="[879,1035,1963,1990]" pageId="15" pageNumber="16" value="1987-02-25">25 Feb. 1987</collectingDate>
|
||
</date>
|
||
;
|
||
<emphasis id="8F8DEADCFFF6E46C1E4AF803EA45C3EA" box="[1047,1390,1963,1989]" italics="true" pageId="15" pageNumber="16">
|
||
<collectorName id="100C5318FFF6E46C1E4AF803EBFEC3EA" box="[1047,1237,1963,1989]" pageId="15" pageNumber="16">Sumbing Jimpin</collectorName>
|
||
<specimenCode id="ED5F9EB5FFF6E46C1E86F803EA45C3EA" box="[1243,1390,1963,1989]" collectionCode="SAN" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15310" lsid="urn:lsid:biocol.org:col:15310" name="Forest Research Centre" pageId="15" pageNumber="16" type="Herbarium">SAN 120057</specimenCode>
|
||
</emphasis>
|
||
; K!,
|
||
<collectionCode id="DBE8AE0BFFF6E46C1AB6F866EE02C3C7" box="[235,297,1998,2024]" country="Malaysia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15310" lsid="urn:lsid:biocol.org:col:15310" name="Forest Research Centre" pageId="15" pageNumber="16" type="Herbarium">SAN</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="E9866646FFE9E4731AE0F8CEEE35C3C5" ID-DOI="http://doi.org/10.5281/zenodo.4594561" ID-Zenodo-Dep="4594561" httpUri="https://zenodo.org/record/4594561/files/figure.png" pageId="16" pageNumber="17" startId="16.[189,232,1894,1920]" targetBox="[203,1342,275,1800]" targetPageId="16">
|
||
<paragraph id="BD4636CEFFE9E4731AE0F8CEEE35C3C5" blockId="16.[189,1399,1894,2026]" pageId="16" pageNumber="17">
|
||
<emphasis id="8F8DEADCFFE9E4731AE0F8CEEE3BC3AF" bold="true" box="[189,272,1894,1920]" pageId="16" pageNumber="17">Fig. 8.</emphasis>
|
||
<taxonomicName id="7AF94D4DFFE9E4731B40F8CEED74C3AF" authority="T.Y.Yu" authorityName="T. Y. Yu" authorityYear="2021" box="[285,607,1894,1920]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="16" pageNumber="17" phylum="Tracheophyta" rank="species" species="calamus" status="sp. nov.">
|
||
<emphasis id="8F8DEADCFFE9E4731B40F8CEED29C3AF" box="[285,514,1894,1920]" italics="true" pageId="16" pageNumber="17">Chassalia calamus</emphasis>
|
||
T.Y.Yu
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="94BE57A7FFE9E4731831F8CEEDE6C3AF" box="[620,717,1894,1920]" pageId="16" pageNumber="17" rank="species">sp. nov.</taxonomicNameLabel>
|
||
<emphasis id="8F8DEADCFFE9E4731884F8CEEDC5C3AF" bold="true" box="[729,750,1894,1920]" pageId="16" pageNumber="17">A</emphasis>
|
||
. Flowering shoot.
|
||
<emphasis id="8F8DEADCFFE9E4731981F8CEECDBC3AF" bold="true" box="[988,1008,1894,1920]" pageId="16" pageNumber="17">B</emphasis>
|
||
. Inflorescence.
|
||
<emphasis id="8F8DEADCFFE9E4731EE8F8CEEBE1C3AF" bold="true" box="[1205,1226,1894,1920]" pageId="16" pageNumber="17">C</emphasis>
|
||
. Longitudinal view of flower showing stamens, band of hairs, stigma, style, disc, calyx and hypanthium.
|
||
<emphasis id="8F8DEADCFFE9E4731F42F821EA1FC38C" bold="true" box="[1311,1332,1929,1955]" pageId="16" pageNumber="17">D</emphasis>
|
||
. Part of infructescence.
|
||
<emphasis id="8F8DEADCFFE9E4731BC6F804EE87C3E9" bold="true" box="[411,428,1964,1990]" pageId="16" pageNumber="17">E</emphasis>
|
||
. Dry fruit.
|
||
<emphasis id="8F8DEADCFFE9E4731867F804ED61C3E9" bold="true" box="[570,586,1964,1990]" pageId="16" pageNumber="17">F</emphasis>
|
||
. Pyrene ventral surface. From
|
||
<emphasis id="8F8DEADCFFE9E47319E3F804EBC4C3E9" box="[958,1263,1964,1991]" italics="true" pageId="16" pageNumber="17">Asik Mantor SAN 118683</emphasis>
|
||
. Drawn by T.Y. Yu.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="E9866646FFE8E4721AE0F866EBA1C3C7" ID-DOI="http://doi.org/10.5281/zenodo.4594563" ID-Zenodo-Dep="4594563" box="[189,1162,1998,2024]" httpUri="https://zenodo.org/record/4594563/files/figure.png" pageId="17" pageNumber="18" startId="17.[189,232,1998,2024]" targetBox="[266,1321,265,1960]" targetPageId="17">
|
||
<paragraph id="BD4636CEFFE8E4721AE0F866EBA1C3C7" blockId="17.[189,1162,1998,2024]" box="[189,1162,1998,2024]" pageId="17" pageNumber="18">
|
||
<emphasis id="8F8DEADCFFE8E4721AE0F866EE21C3C7" bold="true" box="[189,266,1998,2024]" pageId="17" pageNumber="18">Fig. 9.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="7AF94D4DFFE8E4721BFBF866EDF6C3C7" authority="T.Y.Yu" authorityName="T. Y. Yu" authorityYear="2021" box="[422,733,1998,2024]" class="Magnoliopsida" family="Rubiaceae" genus="Chassalia" kingdom="Plantae" order="Gentianales" pageId="17" pageNumber="18" phylum="Tracheophyta" rank="species" species="calamus" status="sp. nov.">
|
||
<emphasis id="8F8DEADCFFE8E4721BFBF866EDAEC3C7" box="[422,645,1998,2024]" italics="true" pageId="17" pageNumber="18">Chassalia calamus</emphasis>
|
||
T.Y.Yu
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="94BE57A7FFE8E47218B9F866EC15C3C7" box="[740,830,1998,2024]" pageId="17" pageNumber="18" rank="species">sp. nov.</taxonomicNameLabel>
|
||
(
|
||
<emphasis id="8F8DEADCFFE8E4721912F866EB52C3C7" box="[847,1145,1998,2024]" italics="true" pageId="17" pageNumber="18">Asik Mantor SAN 118683</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="F5E36545FFEBE4711AE0FEA0EDB0C074" pageId="18" pageNumber="19" type="description">
|
||
<paragraph id="BD4636CEFFEBE4711AE0FEA0EE65C50D" blockId="18.[189,334,264,290]" box="[189,334,264,290]" pageId="18" pageNumber="19">
|
||
<heading id="E60E81A2FFEBE4711AE0FEA0EE65C50D" bold="true" box="[189,334,264,290]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
|
||
<emphasis id="8F8DEADCFFEBE4711AE0FEA0EE65C50D" bold="true" box="[189,334,264,290]" pageId="18" pageNumber="19">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="BD4636CEFFEBE4711AE0FE9FEDB0C074" blockId="18.[188,1401,311,1115]" pageId="18" pageNumber="19">
|
||
Herb, shrub to small treelet, (0.5–)
|
||
<quantity id="7A019B2BFFEBE471180DFE9FED87C57D" box="[592,684,311,338]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="18" pageNumber="19" unit="m" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 m</quantity>
|
||
tall, glabrous. Stem terete, flattened and slightly canaliculate, or sometimes deeply canaliculate, hollow. Lower internodes 3–6 ×
|
||
<quantity id="7A019B2BFFEBE471198DFEF2EB7DC55A" box="[976,1110,346,373]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="18" pageNumber="19" unit="cm" value="0.85" valueMax="1.2" valueMin="0.5">0.5–1.2 cm</quantity>
|
||
, surface usually covered by pustules, internode of flowering stems shorter, 1–2.5 ×
|
||
<quantity id="7A019B2BFFEBE4711930FED6ECDFC5B7" box="[877,1012,382,408]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="8.0" metricValueMin="3.0" pageId="18" pageNumber="19" unit="cm" value="0.55" valueMax="0.8" valueMin="0.3">0.3–0.8 cm</quantity>
|
||
. Leave-blades elliptic to broadly lanceolate, very large, 17–33 ×
|
||
<quantity id="7A019B2BFFEBE471186CFE09ED9AC593" box="[561,689,417,444]" metricMagnitude="-2" metricUnit="m" metricValue="8.75" metricValueMax="12.0" metricValueMin="5.5" pageId="18" pageNumber="19" unit="cm" value="8.75" valueMax="12.0" valueMin="5.5">5.5–12 cm</quantity>
|
||
, midrib raised both adaxially and abaxially, secondary veins 14–18 on each side, petiole flat, canaliculate, (0.5–)
|
||
<quantity id="7A019B2BFFEBE4711964FE6CEC86C5F0" box="[825,941,452,479]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="5.0" metricValueMin="1.5" pageId="18" pageNumber="19" unit="cm" value="3.25" valueMax="5.0" valueMin="1.5">1.5–5 cm</quantity>
|
||
. Stipule large, not sheathing, broadly flabellate, 0.8–1.2 ×
|
||
<quantity id="7A019B2BFFEBE4711BEFFE40ED0FC62D" box="[434,548,488,514]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" metricValueMax="2.0" metricValueMin="0.8" pageId="18" pageNumber="19" unit="cm" value="1.4" valueMax="2.0" valueMin="0.8">0.8–2 cm</quantity>
|
||
, apex usually irregular. Inflorescence a compound cyme, peduncle very short to absent, (<0.1–)
|
||
<quantity id="7A019B2BFFEBE4711B8AFDA3ED4AC609" box="[471,609,523,550]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" metricValueMax="3.5" metricValueMin="0.5" pageId="18" pageNumber="19" unit="cm" value="2.0" valueMax="3.5" valueMin="0.5">0.5–3.5 cm</quantity>
|
||
, rachis and branches usually very short to absent, (<0.1–)
|
||
<quantity id="7A019B2BFFEBE4711F43FDA3EA59C609" box="[1310,1394,523,550]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="19" unit="cm" value="1.5">1.5 cm</quantity>
|
||
. Bracts
|
||
<quantity id="7A019B2BFFEBE4711B4CFD87EE40C667" box="[273,363,558,585]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" metricValueMax="2.54" metricValueMin="2.032" pageId="18" pageNumber="19" unit="in" value="9.0" valueMax="10.0" valueMin="8.0">8–10 in</quantity>
|
||
each capitulum, pale green, 7–8 ×
|
||
<quantity id="7A019B2BFFEBE471194FFD87EC5CC666" box="[786,887,559,585]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="18" pageNumber="19" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
|
||
. Flowers
|
||
<quantity id="7A019B2BFFEBE47119B1FD87EB7EC667" box="[1004,1109,558,585]" metricMagnitude="-1" metricUnit="m" metricValue="4.318" metricValueMax="5.08" metricValueMin="3.556" pageId="18" pageNumber="19" unit="in" value="17.0" valueMax="20.0" valueMin="14.0">14–20 in</quantity>
|
||
each capitulum, pedicel short or absent, hypanthium ellipsoid, 1.5 ×
|
||
<quantity id="7A019B2BFFEBE471188CFDFAEC66C643" box="[721,845,594,621]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="18" pageNumber="19" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.5–2 mm</quantity>
|
||
, calyx tube slightly curved, 1.5–1.8 ×
|
||
<quantity id="7A019B2BFFEBE4711F46FDFAEA5CC643" box="[1307,1399,594,621]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="mm" value="0.5">0.5 mm</quantity>
|
||
in total, corolla whitish green, cream-white or yellowish white, 10–12 ×
|
||
<quantity id="7A019B2BFFEBE4711E43FDDDEB57C6BF" box="[1054,1148,629,656]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="mm" value="0.5">0.5 mm</quantity>
|
||
, corolla tube
|
||
<quantity id="7A019B2BFFEBE4711F43FDDDEA52C6BF" box="[1310,1401,629,656]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="mm" value="0.5">0.5 mm</quantity>
|
||
wide at base, broadening in distal third to
|
||
<quantity id="7A019B2BFFEBE4711896FD30EC02C69C" box="[715,809,664,691]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="19" unit="mm" value="2.5">2.5 mm</quantity>
|
||
wide; corolla lobes triangular, 2–2.5 ×
|
||
<quantity id="7A019B2BFFEBE4711F4EFD30EA5FC69C" box="[1299,1396,664,691]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="19" unit="mm" value="1.5">1.5 mm</quantity>
|
||
. Disc cylindric, 0.8 ×
|
||
<quantity id="7A019B2BFFEBE4711BE7FD14ED32C6F9" box="[442,537,700,727]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="18" pageNumber="19" unit="mm" value="0.5">0.5 mm</quantity>
|
||
. Stamens 2.2 ×
|
||
<quantity id="7A019B2BFFEBE4711885FD14EC1DC6F9" box="[728,822,700,726]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="19" unit="mm" value="0.2">0.2 mm</quantity>
|
||
, anthers placed at the mouth of corolla, filament insertion
|
||
<quantity id="7A019B2BFFEBE4711B77FD77EE46C6D5" box="[298,365,735,762]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="19" unit="mm" value="8.0">8 mm</quantity>
|
||
from base of corolla, filament very short, less than
|
||
<quantity id="7A019B2BFFEBE471199CFD77EB31C6D5" box="[961,1050,735,762]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="18" pageNumber="19" unit="mm" value="0.1">0.1 mm</quantity>
|
||
long, band of hairs
|
||
<quantity id="7A019B2BFFEBE4711F5DFD77EA5CC6D5" box="[1280,1399,735,762]" metricMagnitude="-4" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="18" pageNumber="19" unit="mm" value="0.75" valueMax="1.0" valueMin="0.5">0.5–1 mm</quantity>
|
||
wide, at base of stamens
|
||
<quantity id="7A019B2BFFEBE4711BB6FCABED1AC732" box="[491,561,771,797]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="18" pageNumber="19" unit="mm" value="8.0">8 mm</quantity>
|
||
from mouth of corolla tube, hairs erect, colourless,
|
||
<quantity id="7A019B2BFFEBE4711EFEFCABEA19C732" box="[1187,1330,771,797]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="18" pageNumber="19" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.1–0.2 mm</quantity>
|
||
long. Style
|
||
<quantity id="7A019B2BFFEBE4711B5EFC8EEE7EC76F" box="[259,341,806,833]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="18" pageNumber="19" unit="cm" value="1.5">1.5 cm</quantity>
|
||
long. Fruits ellipsoid, 5–8 × 4–5 ×
|
||
<quantity id="7A019B2BFFEBE471195BFC8EECA9C76F" box="[774,898,806,833]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.0" metricValueMin="3.5" pageId="18" pageNumber="19" unit="mm" value="3.75" valueMax="4.0" valueMin="3.5">3.5–4 mm</quantity>
|
||
, pedicel of fruit persistent, not accrescent or slightly accrescent,
|
||
<quantity id="7A019B2BFFEBE4711B97FCE1ED06C74B" box="[458,557,841,868]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="18" pageNumber="19" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
|
||
long. Pyrenes plano-convex, dorsal surface convex, with two adjacent parallel longitudinal crests, each 0.1 ×
|
||
<quantity id="7A019B2BFFEBE47118C8FCC4EC28C7A8" box="[661,771,876,903]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="18" pageNumber="19" unit="mm" value="0.25">0.25 mm</quantity>
|
||
, curved,
|
||
<quantity id="7A019B2BFFEBE471192FFCC5ECE4C7A8" box="[882,975,877,903]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="18" pageNumber="19" unit="mm" value="0.4">0.4 mm</quantity>
|
||
apart along their length, forming a longitudinal groove from apex to base between the two crests; ventral surface concave, central part slightly elevated, margin with an endocarpal outgrowth
|
||
<quantity id="7A019B2BFFEBE4711918FC1BECB6C7E1" box="[837,925,947,974]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="19" unit="mm" value="0.7">0.7 mm</quantity>
|
||
wide, with a groove between the edge of endocarp and endocarpal outgrowth
|
||
<quantity id="7A019B2BFFEBE471183BFC7FEDEBC7DE" box="[614,704,983,1009]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="19" unit="mm" value="0.2">0.2 mm</quantity>
|
||
wide. Raphal opening at base of the endocarpal outgrowth, round,
|
||
<quantity id="7A019B2BFFEBE4711B52FC52EE42C03A" box="[271,361,1018,1045]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="19" unit="mm" value="0.2">0.2 mm</quantity>
|
||
in diameter,
|
||
<quantity id="7A019B2BFFEBE471185DFC52EDA6C03A" box="[512,653,1018,1045]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="18" pageNumber="19" unit="mm" value="0.25" valueMax="0.3" valueMin="0.2">0.2–0.3 mm</quantity>
|
||
from the edge of the pyrene wall, apex with a bony spine, spine triangular, erect, 0.5 ×
|
||
<quantity id="7A019B2BFFEBE4711B95FBB5ED0EC017" box="[456,549,1053,1080]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="18" pageNumber="19" unit="mm" value="0.2">0.2 mm</quantity>
|
||
. PGS not detected. Seeds light brown, hemi-ellipsoid, slightly crested at central part of dorsal side, 4.5 ×
|
||
<quantity id="7A019B2BFFEBE4711867FBE9EDBCC074" box="[570,663,1089,1115]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="18" pageNumber="19" unit="mm" value="0.7">0.7 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="F5E36545FFEBE4711AE0FB2AEEAEC0C0" pageId="18" pageNumber="19" type="distribution">
|
||
<paragraph id="BD4636CEFFEBE4711AE0FB2AEE94C0B3" blockId="18.[189,447,1154,1180]" box="[189,447,1154,1180]" pageId="18" pageNumber="19">
|
||
<heading id="E60E81A2FFEBE4711AE0FB2AEE94C0B3" bold="true" box="[189,447,1154,1180]" fontSize="11" level="3" pageId="18" pageNumber="19" reason="3">
|
||
<emphasis id="8F8DEADCFFEBE4711AE0FB2AEE94C0B3" bold="true" box="[189,447,1154,1180]" pageId="18" pageNumber="19">Distribution, habitat</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="BD4636CEFFEBE4711AE0FB19EEAEC0C0" blockId="18.[189,1399,1201,1263]" pageId="18" pageNumber="19">
|
||
Borneo:
|
||
<collectingCountry id="C5EE765EFFEBE4711B79FB19EEB8C0E4" box="[292,403,1201,1227]" name="Malaysia" pageId="18" pageNumber="19">Malaysia</collectingCountry>
|
||
,
|
||
<collectingRegion id="7F3DF82CFFEBE4711BC3FB19EEC2C0E4" box="[414,489,1201,1227]" country="Malaysia" name="Sabah" pageId="18" pageNumber="19">Sabah</collectingRegion>
|
||
, around Ranau and Tambunan. Hill forest, stream valley, rocky moist slopes, alt.
|
||
<quantity id="7A019B2BFFEBE4711AB7FB7CEEABC0C0" box="[234,384,1236,1263]" metricMagnitude="2" metricUnit="m" metricValue="7.125" metricValueMax="13.0" metricValueMin="1.25" pageId="18" pageNumber="19" unit="m" value="712.5" valueMax="1300.0" valueMin="125.0">125–1300 m</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |