204 lines
22 KiB
XML
204 lines
22 KiB
XML
<document id="F8A46E0B88DBF5CB7AFD26DDBAC34BF1" ID-DOI="10.11646/zootaxa.3999.1.5" ID-GBIF-Dataset="3ef5a58e-b058-4c33-a79d-f768fdc73427" ID-ISSN="1175-5326" ID-Zenodo-Dep="289212" ID-ZooBank="4DB46BCD-D37E-4B27-8C15-6B43057E7FFD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461279862226" checkinUser="plazi" docAuthor="Dinesh, K. P., Vijayakumar, S. P., Channakeshavamurthy, B. H., Torsekar, Varun R., Kulkarni, Nirmal U. & Shanker, Kartik" docDate="2015" docId="2B7BF863FFAF880AFF23FDAE026CFE53" docLanguage="en" docName="zt03999p094.pdf" docOrigin="Zootaxa 3999 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fejervarya" docType="treatment" docVersion="7" lastPageNumber="82" masterDocId="D742801BFFAC880EFFB4FF8F0055FF9C" masterDocTitle="Systematic status of Fejervarya (( Amphibia, Anura, Dicroglossidae) from South and SE Asia with the description of a new species from the Western Ghats of Peninsular India" masterLastPageNumber="94" masterPageNumber="79" pageNumber="82" updateTime="1698613705382" updateUser="plazi">
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<mods:titleInfo id="05D6FFE3EFB0694F1221515F0CC9409B">
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<mods:title id="C627F52E5EF17B7200BE76B19E1AAB44">Systematic status of Fejervarya (( Amphibia, Anura, Dicroglossidae) from South and SE Asia with the description of a new species from the Western Ghats of Peninsular India</mods:title>
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</mods:titleInfo>
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<mods:name id="46DF1C35CADB0FE647F3167BFEBEAFBB" type="personal">
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<mods:namePart id="C9D159768352F3E9C3DBBDD48AB9E2BD">Dinesh, K. P.</mods:namePart>
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<mods:namePart id="5DF2E3E053006E5ED3F84F50DBF27212">Vijayakumar, S. P.</mods:namePart>
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<mods:name id="868F2366DF6052DEC0BDB3C272F74880" type="personal">
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<mods:roleTerm id="1894A3B543DF028E1BA19B8550504501">Author</mods:roleTerm>
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<mods:namePart id="9622539E24EF099A7F162AF5B8227C5C">Channakeshavamurthy, B. H.</mods:namePart>
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<mods:name id="1BCACDFE2FB64E2AA63829543D996662" type="personal">
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<mods:roleTerm id="97E13A7BFCF7718295A489523C075B84">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="9E9FBB69073D5B4930981751A480EBF2">Torsekar, Varun R.</mods:namePart>
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</mods:name>
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<mods:name id="22AE01922995F9B491B2F126B85258ED" type="personal">
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<mods:roleTerm id="D61DCE15513664655E080498BDCF4BCB">Author</mods:roleTerm>
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<mods:namePart id="42D2CD57636815B21C5CFA9F838674B8">Kulkarni, Nirmal U.</mods:namePart>
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<mods:name id="D3062B557F1637766B86D04B6EE509B0" type="personal">
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<mods:namePart id="9AEE55299CB6974E471DC2C2F43C028E">Shanker, Kartik</mods:namePart>
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<mods:typeOfResource id="A33FE7C2446FB96D196492A6B0DAD6CE">text</mods:typeOfResource>
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<mods:title id="8A1EAC2D737346787D3EF54A6F0EDE1E">Zootaxa</mods:title>
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<mods:part id="A1CB50C3C075618FC45AD92FAF9FC012">
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<mods:date id="D111A7363CD4ED1149D92B49EC420645">2015</mods:date>
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<mods:detail id="704C8506A09EA6F9686B38C07F1BF88C" type="volume">
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<mods:number id="99ABC61D8D24AE1F768C6871D7A5A2AE">3999</mods:number>
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<mods:detail id="CB9304D1020C82BBEBE33133B19E8502" type="issue">
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<mods:classification id="844A8AB6FCAFE815E4FE05BD20EC18EC">journal article</mods:classification>
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<mods:identifier id="602625420E648C6C8E3053E6BD70A42B" type="DOI">10.11646/zootaxa.3999.1.5</mods:identifier>
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<mods:identifier id="2E299887DDC65CE86BE0730C077BAA04" type="ISSN">1175-5326</mods:identifier>
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<treatment id="2B7BF863FFAF880AFF23FDAE026CFE53" ID-DOI="http://doi.org/10.5281/zenodo.5672931" ID-GBIF-Taxon="127692704" ID-Zenodo-Dep="5672931" LSID="urn:lsid:plazi:treatment:2B7BF863FFAF880AFF23FDAE026CFE53" httpUri="http://treatment.plazi.org/id/2B7BF863FFAF880AFF23FDAE026CFE53" lastPageId="4" lastPageNumber="82" pageId="3" pageNumber="82">
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<subSubSection id="EBC81AFEFFAF880DFF23FDAE0191FDA6" box="[151,452,544,571]" pageId="3" pageNumber="82" type="nomenclature">
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<paragraph id="A36D4975FFAF880DFF23FDAE0191FDA6" blockId="3.[151,452,544,571]" box="[151,452,544,571]" pageId="3" pageNumber="82">
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<heading id="F825FE19FFAF880DFF23FDAE0191FDA6" bold="true" box="[151,452,544,571]" fontSize="11" level="1" pageId="3" pageNumber="82" reason="1">
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<emphasis id="91A69567FFAF880DFF23FDAE0191FDA6" bold="true" box="[151,452,544,571]" pageId="3" pageNumber="82">
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Phylogeny of
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<taxonomicName id="64D232F6FFAF880DFEF4FDAF0191FDA6" ID-CoL="4JPG" box="[320,452,544,570]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
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<emphasis id="91A69567FFAF880DFEF4FDAF0191FDA6" bold="true" box="[320,452,544,570]" italics="true" pageId="3" pageNumber="82">Fejervarya</emphasis>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="EBC81AFEFFAF880AFF23FDE8026CFE53" lastPageId="4" lastPageNumber="83" pageId="3" pageNumber="82" type="description">
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<paragraph id="A36D4975FFAF880DFF23FDE80558FCE1" blockId="3.[151,1436,615,893]" pageId="3" pageNumber="82">
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The phylogeny recovered a well supported clade consisting of all
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<taxonomicName id="64D232F6FFAF880DFCCEFDE703A3FDE3" box="[890,1014,616,639]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
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<emphasis id="91A69567FFAF880DFCCEFDE703A3FDE3" box="[890,1014,616,639]" italics="true" pageId="3" pageNumber="82">Fejervarya</emphasis>
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</taxonomicName>
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from South and South
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<collectingCountry id="DBC509E5FFAF880DFB4FFDE70533FD1C" box="[1275,1382,615,640]" name="China" pageId="3" pageNumber="82">East Asia</collectingCountry>
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(
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<figureCitation id="3BE955F0FFAF880DFAC1FDE800FDFD38" captionStart="FIGURE 2" captionStartId="3.[151,250,1889,1911]" captionTargetBox="[175,1424,957,1852]" captionTargetId="figure@3.[163,1442,949,1859]" captionTargetPageId="3" captionText="FIGURE 2. Maximum Likelihood tree for 33 dicroglossid taxa based on 6879 bp of mitochondrial (16 S and 12 S) and nuclear genes (BDNF, Rhod, Tyr, RAG- 1, RAG- 2, NCX 1, and CXCR 4)." httpUri="https://zenodo.org/record/289214/files/figure.png" pageId="3" pageNumber="82">Fig 2</figureCitation>
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). We also recovered two sister clades within
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<taxonomicName id="64D232F6FFAF880DFD0AFD02036FFD38" box="[702,826,653,676]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
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<emphasis id="91A69567FFAF880DFD0AFD02036FFD38" box="[702,826,653,676]" italics="true" pageId="3" pageNumber="82">Fejervarya</emphasis>
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</taxonomicName>
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. The geographical ranges of the two clades broadly overlap around the Northeast Indian regions and
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<collectingCountry id="DBC509E5FFAF880DFD09FD200374FD54" box="[701,801,687,712]" name="Thailand" pageId="3" pageNumber="82">Thailand</collectingCountry>
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(
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<figureCitation id="3BE955F0FFAF880DFC84FD20033BFD54" box="[816,878,687,712]" captionStart="FIGURE 2" captionStartId="3.[151,250,1889,1911]" captionTargetBox="[175,1424,957,1852]" captionTargetId="figure@3.[163,1442,949,1859]" captionTargetPageId="3" captionText="FIGURE 2. Maximum Likelihood tree for 33 dicroglossid taxa based on 6879 bp of mitochondrial (16 S and 12 S) and nuclear genes (BDNF, Rhod, Tyr, RAG- 1, RAG- 2, NCX 1, and CXCR 4)." httpUri="https://zenodo.org/record/289214/files/figure.png" pageId="3" pageNumber="82">Fig 2</figureCitation>
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). The
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<taxonomicName id="64D232F6FFAF880DFC02FD3F0466FD5B" box="[950,1075,688,711]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
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<emphasis id="91A69567FFAF880DFC02FD3F0466FD5B" box="[950,1075,688,711]" italics="true" pageId="3" pageNumber="82">Fejervarya</emphasis>
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</taxonomicName>
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clade was recovered as sister to
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<taxonomicName id="64D232F6FFAF880DFF23FD5B0165FD70" box="[151,304,724,748]" class="Amphibia" family="Dicroglossidae" genus="Sphaerotheca" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
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<emphasis id="91A69567FFAF880DFF23FD5B0165FD70" box="[151,304,724,748]" italics="true" pageId="3" pageNumber="82">Sphaerotheca</emphasis>
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</taxonomicName>
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. The unidentified individuals in our collection formed a monophyletic group and showed a sister relationship with
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<taxonomicName id="64D232F6FFAF880DFED9FD77023AFC8C" box="[365,623,760,784]" class="Amphibia" family="Dicroglossidae" genus="Minervarya" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="species" species="sahyadris">
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<emphasis id="91A69567FFAF880DFED9FD77023AFC8C" box="[365,623,760,784]" italics="true" pageId="3" pageNumber="82">Minervarya sahyadris</emphasis>
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</taxonomicName>
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(
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<figureCitation id="3BE955F0FFAF880DFD33FD78029AFC8C" box="[647,719,759,784]" captionStart="FIGURE 2" captionStartId="3.[151,250,1889,1911]" captionTargetBox="[175,1424,957,1852]" captionTargetId="figure@3.[163,1442,949,1859]" captionTargetPageId="3" captionText="FIGURE 2. Maximum Likelihood tree for 33 dicroglossid taxa based on 6879 bp of mitochondrial (16 S and 12 S) and nuclear genes (BDNF, Rhod, Tyr, RAG- 1, RAG- 2, NCX 1, and CXCR 4)." httpUri="https://zenodo.org/record/289214/files/figure.png" pageId="3" pageNumber="82">Fig 2</figureCitation>
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). The clade consisting of the unidentified individuals and
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<taxonomicName id="64D232F6FFAF880DFF23FC9201C0FCA8" box="[151,405,796,820]" class="Amphibia" family="Dicroglossidae" genus="Minervarya" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="species" species="sahyadris">
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<emphasis id="91A69567FFAF880DFF23FC9201C0FCA8" box="[151,405,796,820]" italics="true" pageId="3" pageNumber="82">Minervarya sahyadris</emphasis>
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</taxonomicName>
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was deeply nested within the larger
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<taxonomicName id="64D232F6FFAF880DFCECFC920380FCA8" box="[856,981,797,820]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
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<emphasis id="91A69567FFAF880DFCECFC920380FCA8" box="[856,981,797,820]" italics="true" pageId="3" pageNumber="82">Fejervarya</emphasis>
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</taxonomicName>
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clade. Based on this, we synonymize
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<taxonomicName id="64D232F6FFAF880DFF23FCCF022FFCC4" authority="Dubois, Ohler and Biju, 2001" authorityName="Dubois, Ohler and Biju" authorityYear="2001" box="[151,634,831,856]" class="Amphibia" family="Dicroglossidae" genus="Minervarya" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
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<emphasis id="91A69567FFAF880DFF23FCCF014EFCCB" box="[151,283,832,855]" italics="true" pageId="3" pageNumber="82">Minervarya</emphasis>
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<bibRefCitation id="C7433484FFAF880DFE90FCB0022FFCC4" author="Dubois" box="[292,634,831,856]" pageId="3" pageNumber="82" refString="Dubois, A., Ohler, A. & Biju, S. D. (2001) A new genus and species of Ranidae (Amphibia; Anura) from southwestern India. Alytes, 19, 53 - 79." type="journal article" year="2001">Dubois, Ohler and Biju, 2001</bibRefCitation>
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</taxonomicName>
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under
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<taxonomicName id="64D232F6FFAF880DFD78FCCF03BCFCC4" authority="Bolkay, 1915" authorityName="Bolkay" authorityYear="1915" box="[716,1001,831,856]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="genus">
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<emphasis id="91A69567FFAF880DFD78FCCF031DFCCB" box="[716,840,832,855]" italics="true" pageId="3" pageNumber="82">Fejervarya</emphasis>
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Bolkay, 1915
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</taxonomicName>
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(see discussion below) and provide a diagnosis and describe the unidentified individuals in our collection as
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<taxonomicName id="64D232F6FFAF880DFC07FCEA04F9FCE0" box="[947,1196,868,892]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="3" pageNumber="82" phylum="Chordata" rank="species" species="gomantaki" status="sp. nov.">
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<emphasis id="91A69567FFAF880DFC07FCEA04F9FCE0" box="[947,1196,868,892]" italics="true" pageId="3" pageNumber="82">Fejervarya gomantaki</emphasis>
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</taxonomicName>
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<emphasis id="91A69567FFAF880DFB00FCEB0558FCE1" bold="true" box="[1204,1293,868,893]" pageId="3" pageNumber="82">
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<taxonomicNameLabel id="8A95281CFFAF880DFB00FCEB0558FCE1" box="[1204,1293,868,893]" pageId="3" pageNumber="82" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<caption id="F7AD19FDFFAF880DFF23F8EE0365F809" httpUri="https://zenodo.org/record/289214/files/figure.png" pageId="3" pageNumber="82" targetBox="[175,1424,957,1852]" targetPageId="3">
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<paragraph id="A36D4975FFAF880DFF23F8EE0365F809" blockId="3.[151,1436,1889,1941]" pageId="3" pageNumber="82">
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<emphasis id="91A69567FFAF880DFF23F8EE0146F8EA" bold="true" box="[151,275,1889,1911]" pageId="3" pageNumber="82">FIGURE 2.</emphasis>
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Maximum Likelihood tree for 33 dicroglossid taxa based on 6879 bp of mitochondrial (16S and 12S) and nuclear genes (BDNF, Rhod, Tyr, RAG-1, RAG-2, NCX1, and CXCR4).
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</paragraph>
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</caption>
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<paragraph id="A36D4975FFAF880AFF73F84D026CFE53" blockId="3.[199,1436,1986,2011]" lastBlockId="4.[151,1436,151,464]" lastPageId="4" lastPageNumber="83" pageId="3" pageNumber="82">
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<emphasis id="91A69567FFAF880DFF73F84D01E5F847" bold="true" box="[199,432,1986,2011]" pageId="3" pageNumber="82">Generic allocation.</emphasis>
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There is considerable confusion in generic level taxonomy and systematics within dicroglossid frogs, specifically the distinction between
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<taxonomicName id="64D232F6FFA8880AFC95FF1703C8FF33" box="[801,925,152,175]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="genus">
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<emphasis id="91A69567FFA8880AFC95FF1703C8FF33" box="[801,925,152,175]" italics="true" pageId="4" pageNumber="83">Fejervarya</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="64D232F6FFA8880AFC05FF170460FF33" box="[945,1077,152,175]" class="Amphibia" family="Dicroglossidae" genus="Minervarya" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="genus">
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<emphasis id="91A69567FFA8880AFC05FF170460FF33" box="[945,1077,152,175]" italics="true" pageId="4" pageNumber="83">Minervarya</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="64D232F6FFA8880AFBC2FF1704B4FF2C" box="[1142,1249,152,176]" class="Amphibia" family="Dicroglossidae" genus="Zakerana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="genus">
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<emphasis id="91A69567FFA8880AFBC2FF1704B4FF2C" box="[1142,1249,152,176]" italics="true" pageId="4" pageNumber="83">Zakerana</emphasis>
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</taxonomicName>
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(see discussion below). Apart from the strong phylogenetic support (
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<figureCitation id="3BE955F0FFA8880AFD5EFF33037DFF48" box="[746,808,188,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1889,1911]" captionTargetBox="[175,1424,957,1852]" captionTargetId="figure@3.[163,1442,949,1859]" captionTargetPageId="3" captionText="FIGURE 2. Maximum Likelihood tree for 33 dicroglossid taxa based on 6879 bp of mitochondrial (16 S and 12 S) and nuclear genes (BDNF, Rhod, Tyr, RAG- 1, RAG- 2, NCX 1, and CXCR 4)." httpUri="https://zenodo.org/record/289214/files/figure.png" pageId="4" pageNumber="83">Fig 2</figureCitation>
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), the individuals are assignable to the genus
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|
<taxonomicName id="64D232F6FFA8880AFAABFF3205C9FF48" box="[1311,1436,189,212]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="genus">
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<emphasis id="91A69567FFA8880AFAABFF3205C9FF48" box="[1311,1436,189,212]" italics="true" pageId="4" pageNumber="83">Fejervarya</emphasis>
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</taxonomicName>
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|
based on the diagnosable generic morphological characters assigned by
|
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<bibRefCitation id="C7433484FFA8880AFC63FF500495FF64" box="[983,1216,223,248]" pageId="4" pageNumber="83" refString="Dubois, A., Ohler, A. & Biju, S. D. (2001) A new genus and species of Ranidae (Amphibia; Anura) from southwestern India. Alytes, 19, 53 - 79." type="journal article">
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Dubois
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|
<emphasis id="91A69567FFA8880AFB85FF6E0439FF64" box="[1073,1132,224,248]" italics="true" pageId="4" pageNumber="83">et al.</emphasis>
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|
(2001)
|
|
</bibRefCitation>
|
|
for the then genus
|
|
<taxonomicName id="64D232F6FFA8880AFF23FE8A0224FE80" authority="Dubois, Ohler and Biju, 2001" authorityName="Dubois, Ohler and Biju" authorityYear="2001" box="[151,625,260,285]" class="Amphibia" family="Dicroglossidae" genus="Minervarya" higherTaxonomySource="CoL" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="genus">
|
|
<emphasis id="91A69567FFA8880AFF23FE8A014EFE80" box="[151,283,261,284]" italics="true" pageId="4" pageNumber="83">Minervarya</emphasis>
|
|
<bibRefCitation id="C7433484FFA8880AFE97FE8B0224FE80" author="Dubois" box="[291,625,260,285]" pageId="4" pageNumber="83" refString="Dubois, A., Ohler, A. & Biju, S. D. (2001) A new genus and species of Ranidae (Amphibia; Anura) from southwestern India. Alytes, 19, 53 - 79." type="journal article" year="2001">Dubois, Ohler and Biju, 2001</bibRefCitation>
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|
</taxonomicName>
|
|
and
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|
<taxonomicName id="64D232F6FFA8880AFD1DFE8A0393FE81" authority="Bolkay, 1915" authorityName="Bolkay" authorityYear="1915" box="[681,966,260,285]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="genus">
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|
<emphasis id="91A69567FFA8880AFD1DFE8A0373FE80" box="[681,806,261,284]" italics="true" pageId="4" pageNumber="83">Fejervarya</emphasis>
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|
Bolkay, 1915
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|
</taxonomicName>
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|
; namely to the group of small sized frogs having snout not greatly pointed or slightly rounded; relatively small tympanum; small rounded or laterally compressed internal metatarsal tubercles; rudimentary webbing on feet and small tibia; presence of rictal glands, fejervaryan lines, and horizontal band on the upper lip. Within
|
|
<taxonomicName id="64D232F6FFA8880AFCD1FEFF03B4FE1B" box="[869,993,368,391]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="genus">
|
|
<emphasis id="91A69567FFA8880AFCD1FEFF03B4FE1B" box="[869,993,368,391]" italics="true" pageId="4" pageNumber="83">Fejervarya</emphasis>
|
|
</taxonomicName>
|
|
, presence of a rictal gland is a derived character unique to the clade composed of
|
|
<taxonomicName id="64D232F6FFA8880AFD2CFE1A03D8FE30" box="[664,909,404,428]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="species" species="sahyadris">
|
|
<emphasis id="91A69567FFA8880AFD2CFE1A03D8FE30" box="[664,909,404,428]" italics="true" pageId="4" pageNumber="83">Fejervarya sahyadris</emphasis>
|
|
</taxonomicName>
|
|
,
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|
<taxonomicName id="64D232F6FFA8880AFC14FE1A04CBFE30" box="[928,1182,404,428]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="species" species="gomantaki" status="sp. nov.">
|
|
<emphasis id="91A69567FFA8880AFC14FE1A04CBFE30" box="[928,1182,404,428]" italics="true" pageId="4" pageNumber="83">Fejervarya gomantaki</emphasis>
|
|
</taxonomicName>
|
|
<emphasis id="91A69567FFA8880AFB1EFE1B055CFE31" bold="true" box="[1194,1289,404,429]" pageId="4" pageNumber="83">
|
|
<taxonomicNameLabel id="8A95281CFFA8880AFB1EFE1B055CFE31" box="[1194,1289,404,429]" pageId="4" pageNumber="83" rank="species">sp. nov.</taxonomicNameLabel>
|
|
</emphasis>
|
|
and a north eastern species
|
|
<taxonomicName id="64D232F6FFA8880AFEF1FE370267FE4C" box="[325,562,440,464]" class="Amphibia" family="Dicroglossidae" genus="Fejervarya" kingdom="Animalia" order="Anura" pageId="4" pageNumber="83" phylum="Chordata" rank="species" species="chilapata">
|
|
<emphasis id="91A69567FFA8880AFEF1FE370267FE4C" box="[325,562,440,464]" italics="true" pageId="4" pageNumber="83">Fejervarya chilapata</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |