157 lines
17 KiB
XML
157 lines
17 KiB
XML
<document id="92B7F7E8B01C7012B6D5388B08FB666A" ID-DOI="http://dx.doi.org/10.3897/zookeys.719.21253" ID-GBIF-Dataset="b92acfc0-9d67-42c4-9e37-ebd5a53d24b5" ID-Pensoft-Pub="1313-2970-719-75" ID-ZooBank="36C4E2C8E07D4CC9A1D696B0FCE92CCF" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-719-75" ModsDocOrigin="ZooKeys 719" ModsDocTitle="A taxonomic revision of the subfamily Tillinae Leach sensu lato (Coleoptera, Cleridae) in the New World" checkinTime="1555319501509" checkinUser="pensoft" docAuthor="Burke, Alan & Zolnerowich, Gregory" docDate="2017" docId="23ED0E09A6E946AE093785EE2400A4D0" docLanguage="en" docName="ZooKeys 719: 75-157" docOrigin="ZooKeys 719" docSource="http://dx.doi.org/10.3897/zookeys.719.21253" docTitle="Onychotillus Chapin 1945" docType="treatment" docVersion="5" lastPageNumber="137" masterDocId="5C2D8118751DD235197A6569FFE59A6E" masterDocTitle="A taxonomic revision of the subfamily Tillinae Leach sensu lato (Coleoptera, Cleridae) in the New World" masterLastPageNumber="157" masterPageNumber="75" pageNumber="135" updateTime="1701317054126" updateUser="plazi">
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<mods:title id="7729F33CABF95731A807320BBB51F9D6">A taxonomic revision of the subfamily Tillinae Leach sensu lato (Coleoptera, Cleridae) in the New World</mods:title>
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<mods:namePart id="DD6ECCD93F4BB63AFD6338086421DE52">Burke, Alan</mods:namePart>
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<mods:namePart id="17E23EAF59352130E1DC2A7F716D85C9">Zolnerowich, Gregory</mods:namePart>
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<mods:date id="E13F12207653FEB870C4755478AEF86E">2017</mods:date>
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<mods:number id="779AFDC4920850066AE783D345FB6D9F">719</mods:number>
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<mods:identifier id="E85BFBB74355CBBB78B125478B840A54" type="DOI">http://dx.doi.org/10.3897/zookeys.719.21253</mods:identifier>
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<treatment id="23ED0E09A6E946AE093785EE2400A4D0" ID-GBIF-Taxon="156200940" LSID="urn:lsid:plazi:treatment:23ED0E09A6E946AE093785EE2400A4D0" httpUri="http://treatment.plazi.org/id/23ED0E09A6E946AE093785EE2400A4D0" lastPageId="62" lastPageNumber="137" pageId="60" pageNumber="135">
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<subSubSection id="616B561DE8AFCA0CB8811D7F45CE32DB" pageId="60" pageNumber="135" type="nomenclature">
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<paragraph id="8F97716C5D21362C1DC8CC393A067E22" pageId="60" pageNumber="135">
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<taxonomicName id="45226D1ED702723E5779B845526E3217" authority="Chapin, 1945" authorityName="Chapin" authorityYear="1945" class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus" order="Coleoptera" pageId="60" pageNumber="135" phylum="Arthropoda" rank="genus">Onychotillus Chapin, 1945</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="9C9A16D914485BB43260E89E23C727D2" pageId="60" pageNumber="135" type="type species">
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<paragraph id="0F9BECBB912B1A4F08E9F68DC3477218" pageId="60" pageNumber="135">Type species.</paragraph>
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<paragraph id="28A97215FD4986DF1F9019C326B8517B" pageId="60" pageNumber="135">
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<taxonomicName id="FF9FEE4CE9BC8621876B94C69AD126E1" class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus vittatus" order="Coleoptera" pageId="60" pageNumber="135" phylum="Arthropoda" rank="species" species="vittatus">Onychotillus vittatus</taxonomicName>
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Chapin, original designation.
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</paragraph>
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</subSubSection>
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<subSubSection id="6B5F13247FABFE93BAD6E3D9E3FCC8D9" pageId="60" pageNumber="135" type="distribution">
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<paragraph id="10D18BB471394CB0BD04F3EF44CC83C7" pageId="60" pageNumber="135">Distribution.</paragraph>
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<paragraph id="4460A83288626E84159554B19F559F24" pageId="60" pageNumber="135">Shown in Fig. 21K.</paragraph>
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</subSubSection>
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<subSubSection id="D7BCB5EF1FC7E7C3BC39795A5D6B1CD6" pageId="61" pageNumber="136" type="differential diagnosis">
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<paragraph id="A8F4744D3C68E785C45A0538BAD5B5FE" pageId="61" pageNumber="136">
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<pageBreakToken id="E8EB68EC1FA1F1AECA2D4C52060AC5EF" pageId="61" pageNumber="136" start="start">Differential</pageBreakToken>
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diagnosis.
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</paragraph>
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<paragraph id="214BE9098CE1EBB30BCA89B5C933210C" pageId="61" pageNumber="136">
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Species of
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<taxonomicName id="E30168F73C352DCD374D7C5C746E9882" class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Onychotillus</taxonomicName>
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somewhat resemble small-sized members of
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<taxonomicName id="A4D664DBF8B6F6A9842355344FCA29E6" class="Insecta" family="Cleridae" genus="Cymatodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatodera" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Cymatodera</taxonomicName>
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and species of
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<taxonomicName id="85695C291530865AE5848B54061ABE03" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Cymatoderella</taxonomicName>
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. These genera can be differentiated without difficulty based on the number of tarsal claws.
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<taxonomicName id="6CD24E76049517E4C9C1D170326C72EE" class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Onychotillus</taxonomicName>
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species have a single tarsal claw, while members of
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<taxonomicName id="4E34B39BD3D6619FDEDE56FF9437E591" class="Insecta" family="Cleridae" genus="Cymatodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatodera" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Cymatodera</taxonomicName>
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and
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<taxonomicName id="82B23CFE01F0E89319BEAFB717ADCF47" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Cymatoderella</taxonomicName>
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have two tarsal claws (Fig. 7B). Additionally, the distribution of
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<taxonomicName id="85595FCEC900ACE03E58A4126F0B1C82" class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Onychotillus</taxonomicName>
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does not overlap with that of
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<taxonomicName id="901D28475854AF1B68EA6AD0BCEDFC6A" class="Insecta" family="Cleridae" genus="Cymatodera" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatodera" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Cymatodera</taxonomicName>
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and
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<taxonomicName id="99C1D5F27F7ECEB4182ACB77CF9F6982" class="Insecta" family="Cleridae" genus="Cymatoderella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cymatoderella" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Cymatoderella</taxonomicName>
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, with the former restricted to the West Indies, while the latter two have a continental distribution.
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</paragraph>
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</subSubSection>
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<subSubSection id="34CFC0EE4A6581553876A9D86D6DAB49" pageId="61" pageNumber="136" type="redescription">
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<paragraph id="35F0AE9CFF0DEE0608E632317F352A35" pageId="61" pageNumber="136">Redescription.</paragraph>
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<paragraph id="CF8908260E57E345B0097A8F31F950DA" pageId="61" pageNumber="136">
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Size: 3-12 mm. Color: Body ranging from light fuscous to ferrugineous; some individuals may possess one pale, median fascia on each elytron. Body: Winged species; short to moderately long, slightly robust, somewhat subparallel specimens (Fig. 5
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<normalizedToken id="3193605FC86D8C383C3CC86FB327542F" originalValue="A–B">A-B</normalizedToken>
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).
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</paragraph>
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<paragraph id="35BDE4AC36B5448C5B7C221FEA3B1DE6" pageId="61" pageNumber="136">
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Head: Including eye slightly narrower than pronotum; integument coarsely punctate; punctations vary from narrow to wide; eyes large, finely faceted, feebly emarginate posteriorly, bulging laterally; antennae with 11 antennomeres, extending beyond posterior margin of pronotum; last antennomere as long as or conspicuously longer than ninth antennomere (Fig. 10
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<normalizedToken id="3DCED67C8329FC1E92E628D5EDF27351" originalValue="E–F">E-F</normalizedToken>
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); frons feebly bi-impressed; terminal labial palpi securiform; terminal maxillary palpi cylindrical, distally compressed.
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</paragraph>
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<paragraph id="D4D888A1681EA0CBEC2B20857C8B2492" pageId="61" pageNumber="136">Thorax: Pronotum shiny, smooth to feebly punctate; punctation may range from narrow to slightly wide; lateral margins subparallel, slightly constricted anteriorly, somewhat wider posteriorly. Prosternum: Much wider than long, smooth to rugose, variously punctate. Mesoventrite: Wider than long, smooth, feebly to moderately punctate. Metaventrite: Convex, wider than long, surface conspicuously punctate to almost smooth. Metaventral process compressed anteriorly. Metepisterna largely exposed, the elytra do not coverer these plates in lateral view.</paragraph>
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<paragraph id="57A55682B02F3C8FBBD06F8C68E8ACDD" pageId="61" pageNumber="136">Elytra: Short to elongate, subparallel, widest on middle half, surface moderately to coarsely punctate; punctations arranged in regular striae; punctations do not extend to apex; epipleural fold complete, narrowing toward apex.</paragraph>
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<paragraph id="2E439DB7AB0D388EEF66B371EF6298AD" pageId="61" pageNumber="136">Legs: Femora smooth to rugose; rather swollen distally; tibiae rugose to rugulose; pulvillar formula 4-4-4; one tarsal denticle, tarsal denticle subtriangular in shape.</paragraph>
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<paragraph id="BE87188C320AB59C9FAED2D8D7060ED2" pageId="61" pageNumber="136">Abdomen: Six visible ventrites. Ventrites 1-5 not impressed laterally. Sixth ventrite of males feebly differentiated from that of females; females with sixth ventrite broadly rounded, pygidium simple.</paragraph>
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</subSubSection>
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<subSubSection id="A6DC2A5C411516FA60B2AF5F00082CEC" lastPageId="62" lastPageNumber="137" pageId="61" pageNumber="136" type="remarks">
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<paragraph id="BCA82ED34BDFC415F14C6FD81A344137" pageId="61" pageNumber="136">Remarks.</paragraph>
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<paragraph id="AF366B5E7BD70A9F1BAF0D2B18A24D37" lastPageId="62" lastPageNumber="137" pageId="61" pageNumber="136">
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<taxonomicName id="782473BB8FDB02110FFB12D23BB2D250" class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Onychotillus</taxonomicName>
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is currently composed of five species, all inhabiting the West Indies. Of the five valid species, three are thought to be restricted to Cuba (
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<bibRefCitation id="B06207C6FD59F93F11DD1686D33BE55D" pageId="61" pageNumber="136">De Zayas 1988</bibRefCitation>
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). Based on the limited descriptive work given by
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<bibRefCitation id="AB9D2B87A9E77FD5CC8C33FF9042050F" pageId="61" pageNumber="136">De Zayas (1988)</bibRefCitation>
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, where no key for identification was provided, specimens of what appears to be
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<taxonomicName id="2485CA571C7025C9C693732842D1EFB7" class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus cubana" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="species" species="cubana">Onychotillus cubana</taxonomicName>
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were examined (Fig. 5A); however, none of these individuals were collected in Cuba, so these specimens are tentatively assigned to
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<taxonomicName id="8AE57687B3A5E5252225A9A764C96566" lsidName="O. cubana" pageId="61" pageNumber="136" rank="species" species="cubana">O. cubana</taxonomicName>
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, pending further material to be examined. It is important to note that De
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<normalizedToken id="8816B46FE607568B5DD75734BC02C703" originalValue="Zayas’">Zayas'</normalizedToken>
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descriptions were based, in most cases, on a single specimen, and access to that material has been particularly difficult. Therefore, due to the lack of availability of material and the poorly detailed descriptions given by De Zayas, three
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<taxonomicName id="5F0BF60A962BF2DE652E42031479FF6D" class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus" order="Coleoptera" pageId="61" pageNumber="136" phylum="Arthropoda" rank="genus">Onychotillus</taxonomicName>
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species are excluded from this revision,
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<taxonomicName id="6E6C908BF8339CE815F107316BC43A35" lsidName="O. trinitatis" pageId="61" pageNumber="136" rank="species" species="trinitatis">O. trinitatis</taxonomicName>
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,
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<taxonomicName id="AC5DF595AB6D9E36EC0AF52B1835D6DC" lsidName="O. minuta" pageId="61" pageNumber="136" rank="species" species="minuta">O. minuta</taxonomicName>
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and
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<taxonomicName id="5C327CE8D23149480CDE66015AF92F2C" lsidName="O. dimidiata" pageId="61" pageNumber="136" rank="species" species="dimidiata">O. dimidiata</taxonomicName>
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, all endemic to Cuba. On the other hand, we examined
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<pageBreakToken id="CC70F204DF54C4D4150072E7E5235453" pageId="62" pageNumber="137" start="start">material</pageBreakToken>
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of
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<taxonomicName id="91C0066DECF2B9E71BAAB3DE5FC9A285" class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus vittatus" order="Coleoptera" pageId="62" pageNumber="137" phylum="Arthropoda" rank="species" species="vittatus">Onychotillus vittatus</taxonomicName>
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collected in the Dominican Republic and the Cayman Islands. These collecting localities represent new distribution records for
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<taxonomicName id="3F6BAFB021ADEBB95A4EE4E9084F331B" lsidName="O. vittatus" pageId="62" pageNumber="137" rank="species" species="vittatus">O. vittatus</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="83C4FB5E860CB6B067059E29CB8A14C0" pageId="62" pageNumber="137" type="key">
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<paragraph id="645C4C449B51914F32A05F8C7164B363" pageId="62" pageNumber="137">
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Key to species of
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<taxonomicName id="7A16D1C47EDC7D7808735980C75ABBDF" class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus" order="Coleoptera" pageId="62" pageNumber="137" phylum="Arthropoda" rank="genus">Onychotillus</taxonomicName>
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</paragraph>
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<paragraph id="E9D02C448B307259FEC29782D6AB187C" pageId="62" pageNumber="137">
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<table id="9A5DA9BC6D6E7DD7639B422008C454DE" pageId="62" pageNumber="137">
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<tr id="8553C0CCD1248C1FE7931FEE1A744FEF" pageId="62" pageNumber="137">
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<td id="C6C5C64EA6881C0CBD1A7DEF7A1CB1CE" colspan="1" pageId="62" pageNumber="137" rowspan="1">5B10F</td>
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<td id="619C05E26F4D1C04CA411B4F7FB2BDBB" colspan="1" pageId="62" pageNumber="137" rowspan="1">
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<taxonomicName id="693DF37C2ED89D1151F2B58F36BC6308" class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus vittatus" order="Coleoptera" pageId="62" pageNumber="137" phylum="Arthropoda" rank="species" species="vittatus">Onychotillus vittatus</taxonomicName>
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</td>
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</tr>
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<tr id="299D128EA08BD53DB1308957CA8F5EE1" pageId="62" pageNumber="137">
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<td id="7E38DD7BC6A9C5E4469980B31AB57F69" colspan="1" pageId="62" pageNumber="137" rowspan="1">5A10E</td>
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<td id="2B619B4E2A657C3535B746031CB8B9A4" colspan="1" pageId="62" pageNumber="137" rowspan="1">
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<taxonomicName id="6CCF024C252D1DC074691B1C6552E272" class="Insecta" family="Cleridae" genus="Onychotillus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Onychotillus cubana" order="Coleoptera" pageId="62" pageNumber="137" phylum="Arthropoda" rank="species" species="cubana">Onychotillus cubana</taxonomicName>
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</td>
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</tr>
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</table>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |