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<document id="42764C78A8D1E5319D4522DF54A3DDE2" ID-DOI="10.11646/zootaxa.5039.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5506250" ID-ZooBank="FB5EDBC1-B2DB-4091-ACDF-C3858F6FCD62" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631598984208" checkinUser="plazi" docAuthor="Lee, Mao-Ying &amp; Munroe, Thomas A." docDate="2021" docId="094B87CCFFF35E3FB5913D8DFA295AAF" docLanguage="en" docName="zootaxa.5039.1.1.pdf" docOrigin="Zootaxa 5039 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Symphurus , Munroe 1992" docType="key" docVersion="3" lastPageNumber="53" masterDocId="F572FFB4FFC75E0BB5063D1AFFB55F28" masterDocTitle="Unraveling cryptic diversity among shallow-water tonguefishes (Pleuronectiformes: Cynoglossidae: Symphurus) from the Indo-West Pacific region, with descriptions of five new species" masterLastPageNumber="55" masterPageNumber="1" pageNumber="53" updateTime="1699414315014" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="33DC627AB1949C77A87AACD7127822F6">Unraveling cryptic diversity among shallow-water tonguefishes (Pleuronectiformes: Cynoglossidae: Symphurus) from the Indo-West Pacific region, with descriptions of five new species</mods:title>
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<mods:namePart id="8B190765AA3B63D348B3F01A80429619">Lee, Mao-Ying</mods:namePart>
<mods:affiliation id="92994C16029D2AE955037D7A9A03023E">Marine Fisheries Division, Fisheries Research Institute, Council of Agriculture, Executive Yuen, 199 Hou-Ih Road, Keelung 20246, Taiwan. &amp; Institute of Oceanography, National Taiwan University, No. 1 Sec. 4 Roosevelt 295 Road, Taipei 10617, Taiwan</mods:affiliation>
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<mods:namePart id="7254E6B63C4F2BAC924D91BA0AE5B822">Munroe, Thomas A.</mods:namePart>
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<mods:affiliation id="4ABB059F44797F93B24198085129DF8C">National Marine Fisheries Service National Systematics Laboratory, NOAA, Smithsonian Institution, P. O. Box 37012, National Museum of Natural History, WC- 57, MRC- 153, Washington, DC 20013 - 7012. munroet @ si. edu; https: // orcid. org / 0000 - 0002 - 6317 - 0985 Corresponding author. coleopetera @ gmail. com; https: // orcid. org / 0000 - 0002 - 4643 - 3173</mods:affiliation>
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<subSubSection id="C9F86551FFF35E3FB5913D8DFBFD5F99" box="[151,1096,151,178]" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="815D36DAFFF35E3FB5913D8DFBFD5F99" blockId="52.[151,1096,151,178]" box="[151,1096,151,178]" pageId="52" pageNumber="53">
<heading id="DA1581B6FFF35E3FB5913D8DFBFD5F99" bold="true" box="[151,1096,151,178]" fontSize="11" level="1" pageId="52" pageNumber="53" reason="1">
<emphasis id="B396EAC8FFF35E3FB5913D8DFBFD5F99" bold="true" box="[151,1096,151,178]" pageId="52" pageNumber="53">
Key to the shallow-water species of
<taxonomicName id="46E24D59FFF35E3FB7553D8DFD6B5F9A" ID-CoL="6464B" ID-ENA="195651" authorityName=", Munroe" authorityYear="1992" box="[595,734,151,178]" class="Actinopterygii" family="Cynoglossidae" genus="Symphurus" kingdom="Animalia" order="Pleuronectiformes" pageId="52" pageNumber="53" phylum="Chordata" rank="genus">
<emphasis id="B396EAC8FFF35E3FB7553D8DFD6B5F9A" bold="true" box="[595,734,151,178]" italics="true" pageId="52" pageNumber="53">Symphurus</emphasis>
</taxonomicName>
from the Indo-Pacific region
</emphasis>
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</paragraph>
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<subSubSection id="C9F86551FFF35E3FB5913DC6FA295AAF" pageId="52" pageNumber="53" type="key">
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<paragraph id="815D36DAFFF35E3FB5913DC6FA295E27" blockId="52.[151,1437,220,1415]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913DC6FA295E27" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913DC6FF005FD9" bold="true" box="[151,181,220,241]" pageId="52" pageNumber="53">1a.</emphasis>
Conspicuous pigmented spot on anterior ocular-side abdomen; ocular-side with conspicuous freckled color pattern; 14 caudalfin rays; five hypurals...........................................................
<taxonomicName id="46E24D59FFF35E3FB1783DE0FA295E27" ID-CoL="53PSS" authority="Munroe 2006" authorityName="Munroe" authorityYear="2006" box="[1150,1436,249,271]" class="Actinopterygii" family="Cynoglossidae" genus="Symphurus" kingdom="Animalia" order="Pleuronectiformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="monostigmus">
<emphasis id="B396EAC8FFF35E3FB1783DE0FAA05E27" box="[1150,1301,250,271]" italics="true" pageId="52" pageNumber="53">S. monostigmus</emphasis>
<bibRefCitation id="E5734B2BFFF35E3FB01C3DE0FA295E27" author="Munroe, T. A." box="[1306,1436,249,271]" pageId="52" pageNumber="53" pagination="230 - 234" refId="ref43051" refString="Munroe, T. A. (2006) New western Indian Ocean tonguefish (Pleuronectiformes: Cynoglossidae, Symphurus). Copeia, 2006 (2), 230 - 234. https: // doi. org / 10.1643 / 0045 - 8511 (2006) 6 [230: NWIOTP] 2.0. CO; 2" type="journal article" year="2006">Munroe 2006</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="815D36DAFFF35E3FB5913C0DFA295E61" blockId="52.[151,1437,220,1415]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913C0DFA295E61" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913C0DFF035E04" bold="true" box="[151,182,279,300]" pageId="52" pageNumber="53">1b.</emphasis>
No pigment spot on anterior ocular-side abdomen; ocular-side without conspicuous freckled pattern; 12 caudal-fin rays; four hypurals............................................................................................
<emphasis id="B396EAC8FFF35E3FB0963C2EFA295E61" bold="true" box="[1424,1436,308,329]" pageId="52" pageNumber="53">2</emphasis>
</keyLead>
</paragraph>
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<keyStep id="3A162E7FFFF35E3FB5913C4BFA295ED1" pageId="52" pageNumber="53">
<paragraph id="815D36DAFFF35E3FB5913C4BFA295E89" blockId="52.[151,1437,220,1415]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913C4BFA295E89" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913C4BFF035E4E" bold="true" box="[151,182,337,358]" pageId="52" pageNumber="53">2a.</emphasis>
Fleshy ridge present on ocular-side lower jaw; black dermal spots present at bases of anterior dorsal-fin rays and on the isthmus; anterior margin of upper eye in advance, or equal with, anterior margin of lower eye; eyes contiguous; a membrane covering both eyes usually present; anterior nostril connected to eyes by fleshy membrane..................................
<emphasis id="B396EAC8FFF35E3FB0963C96FA295E89" bold="true" box="[1424,1436,396,417]" pageId="52" pageNumber="53">3</emphasis>
</keyLead>
</paragraph>
<paragraph id="815D36DAFFF35E3FB5913CB3FA295ED1" blockId="52.[151,1437,220,1415]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913CB3FA295ED1" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913CB3FF025E96" bold="true" box="[151,183,425,446]" pageId="52" pageNumber="53">2b.</emphasis>
No fleshy ridge on ocular-side lower jaw; no dermal spots at bases of anterior dorsal-fin rays or on the isthmus; anterior margins of eyes usually equal in position; eyes separated from each other by narrow interorbital space; no membrane covering eyes; anterior ocular-side nostril not connected to eyes by fleshy membrane....................
<taxonomicName id="46E24D59FFF35E3FB1873CFEFA295ED1" ID-CoL="7B6YJ" authority="Lee et al. 2014" authorityName="Lee" authorityYear="2014" box="[1153,1436,484,506]" class="Actinopterygii" family="Cynoglossidae" genus="Symphurus" kingdom="Animalia" order="Pleuronectiformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="leucochilus">
<emphasis id="B396EAC8FFF35E3FB1873CFEFAB35ED1" box="[1153,1286,484,505]" italics="true" pageId="52" pageNumber="53">S. leucochilus</emphasis>
<bibRefCitation id="E5734B2BFFF35E3FB00A3CFFFA295ED1" author="Lee, M. - Y. &amp; Munroe, T. A. &amp; Shao, K. - T." box="[1292,1436,484,506]" pageId="52" pageNumber="53" pagination="563 - 585" refId="ref42740" refString="Lee, M. - Y., Munroe, T. A. &amp; Shao, K. - T. (2014) Description of a new cryptic, shallow-water tonguefish (Pleuronectiformes: Cynoglossidae: Symphurus) from the western North Pacific Ocean. Journal of Fish Biology, 85 (3), 563 - 585. https: // doi. org / 10.1111 / jfb. 12440" type="journal article" year="2014">
Lee
<emphasis id="B396EAC8FFF35E3FB0303CFFFAD45ED1" box="[1334,1377,484,506]" italics="true" pageId="52" pageNumber="53">et al</emphasis>
. 2014
</bibRefCitation>
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<keyStep id="3A162E7FFFF35E3FB5913F1BFA295D79" pageId="52" pageNumber="53">
<paragraph id="815D36DAFFF35E3FB5913F1BFA295D1C" blockId="52.[151,1437,220,1415]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913F1BFA295D1C" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913F1BFF035D3E" bold="true" box="[151,182,513,534]" pageId="52" pageNumber="53">3a.</emphasis>
Transverse scale rows 2126; longitudinal scale rows 5666; head width usually equal to, or shorter than, head length (HW/HL ≤ 1)................................................................................................
<emphasis id="B396EAC8FFF35E3FB0963F05FA295D1C" bold="true" box="[1424,1436,543,564]" pageId="52" pageNumber="53">4</emphasis>
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<paragraph id="815D36DAFFF35E3FB5913F26FA295D79" blockId="52.[151,1437,220,1415]" box="[151,1436,572,594]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913F26FA295D79" box="[151,1436,572,594]" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913F26FF025D79" bold="true" box="[151,183,572,593]" pageId="52" pageNumber="53">3b.</emphasis>
Transverse scale rows 2637; longitudinal scale rows 7294; head width greater than head length (HW/HL&gt; 1)..........
<emphasis id="B396EAC8FFF35E3FB0963F26FA295D79" bold="true" box="[1424,1436,572,593]" pageId="52" pageNumber="53">6</emphasis>
</keyLead>
</paragraph>
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<keyStep id="3A162E7FFFF35E3FB5913F43FA295DEF" pageId="52" pageNumber="53">
<paragraph id="815D36DAFFF35E3FB5913F43FA295DA4" blockId="52.[151,1437,220,1415]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913F43FA295DA4" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913F43FF035D46" bold="true" box="[151,182,601,622]" pageId="52" pageNumber="53">4a.</emphasis>
Scale rows on head posterior to lower orbit 1516; postorbital length 75.976.4% of HL; predorsal length 18.119.7% of HL; snout length 10.414.3% of HL; SNL/ED= 1.251.45; head length 24.024.2% of SL..............................
<emphasis id="B396EAC8FFF35E3FB0963F6DFA295DA4" bold="true" box="[1424,1436,631,652]" pageId="52" pageNumber="53">5</emphasis>
</keyLead>
</paragraph>
<paragraph id="815D36DAFFF35E3FB5913F8EFA295DEF" blockId="52.[151,1437,220,1415]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913F8EFA295DEF" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913F8EFF025D81" bold="true" box="[151,183,660,681]" pageId="52" pageNumber="53">4b.</emphasis>
Scale rows on head posterior to lower orbit 1114; postorbital length 62.867.5% of SL; predorsal length 23.130.4% of HL; snout length 18.523.4% of HL; SNL/ED= 1.66 2.24; head length 20.522.6% of SL.......
<taxonomicName id="46E24D59FFF35E3FB1793FA8FA295DEF" authority="Lee et al. 2016" authorityName="Lee" authorityYear="2016" box="[1151,1436,689,711]" class="Actinopterygii" family="Cynoglossidae" genus="Symphurus" kingdom="Animalia" order="Pleuronectiformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="longirostris">
<emphasis id="B396EAC8FFF35E3FB1793FA8FAB35DEE" box="[1151,1286,689,711]" italics="true" pageId="52" pageNumber="53">S. longirostris</emphasis>
Lee
<emphasis id="B396EAC8FFF35E3FB0303FA8FAD45DEE" box="[1334,1377,689,711]" italics="true" pageId="52" pageNumber="53">et al</emphasis>
. 2016
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3A162E7FFFF35E3FB5913FD5FA295C14" pageId="52" pageNumber="53">
<paragraph id="815D36DAFFF35E3FB5913FD5FA295C2A" blockId="52.[151,1437,220,1415]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913FD5FA295C2A" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913FD5FF035DCC" bold="true" box="[151,182,719,740]" pageId="52" pageNumber="53">5a.</emphasis>
Lower head lobe much larger than upper head lobe (UHL/LHL=0.78); ID pattern 12222; total vertebrae 48....................................................................................
<taxonomicName id="46E24D59FFF35E3FB16A3FF6FA295C2A" ID-CoL="53PSQ" authority="(Weber 1913)" baseAuthorityName="Weber" baseAuthorityYear="1913" box="[1132,1436,748,770]" class="Actinopterygii" family="Cynoglossidae" genus="Symphurus" kingdom="Animalia" order="Pleuronectiformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="microrhynchus">
<emphasis id="B396EAC8FFF35E3FB16A3FF6FAA65C29" box="[1132,1299,748,769]" italics="true" pageId="52" pageNumber="53">S. microrhynchus</emphasis>
(
<bibRefCitation id="E5734B2BFFF35E3FB0183FF6FA205C2A" author="Weber, M." box="[1310,1429,748,770]" pageId="52" pageNumber="53" refId="ref43796" refString="Weber, M. (1913) Die Fische der Siboga-Expedition. E. J. Brill, Leiden, 710 pp." type="book" year="1913">Weber 1913</bibRefCitation>
)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="815D36DAFFF35E3FB5913E10FA295C14" blockId="52.[151,1437,220,1415]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913E10FA295C14" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913E10FF025C37" bold="true" box="[151,183,778,799]" pageId="52" pageNumber="53">5b.</emphasis>
Lower head lobe smaller than upper head lobe (UHL/LHL=1.01); ID pattern 12322 total vertebrae 46.........................................................................................
<taxonomicName id="46E24D59FFF35E3FB16A3E3DFA295C14" ID-CoL="53PS4" authority="Chabanaud 1948" authorityName="Chabanaud" authorityYear="1948" box="[1132,1436,807,829]" class="Actinopterygii" family="Cynoglossidae" genus="Symphurus" kingdom="Animalia" order="Pleuronectiformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="holothuriae">
<emphasis id="B396EAC8FFF35E3FB16A3E3DFB415C14" box="[1132,1268,807,828]" italics="true" pageId="52" pageNumber="53">S. holothuriae</emphasis>
<bibRefCitation id="E5734B2BFFF35E3FB1FC3E3DFA295C14" author="Chabanaud, P." box="[1274,1436,807,829]" pageId="52" pageNumber="53" pagination="508 - 511" refId="ref41903" refString="Chabanaud, P. (1948) Description de quatre espe`ces ine`dites du genre Symphurus. Bulletin du Museum National d'Histoire Naturelle, Serie 2, 20 (1), 508 - 511." type="journal article" year="1948">Chabanaud 1948</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3A162E7FFFF35E3FB5913E5EFA295C5E" pageId="52" pageNumber="53">
<paragraph id="815D36DAFFF35E3FB5913E5EFA295C71" blockId="52.[151,1437,220,1415]" box="[151,1436,836,858]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913E5EFA295C71" box="[151,1436,836,858]" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913E5EFF035C71" bold="true" box="[151,182,836,857]" pageId="52" pageNumber="53">6a.</emphasis>
Blind-side uniformly white; no pepper-dots; dermal melanophores only present at bases of anterior dorsal-fin rays........
<emphasis id="B396EAC8FFF35E3FB0963E5EFA295C71" bold="true" box="[1424,1436,836,857]" pageId="52" pageNumber="53">7</emphasis>
</keyLead>
</paragraph>
<paragraph id="815D36DAFFF35E3FB5913E7BFA295C5E" blockId="52.[151,1437,220,1415]" box="[151,1436,865,887]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913E7BFA295C5E" box="[151,1436,865,887]" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913E7BFF025C5E" bold="true" box="[151,183,865,886]" pageId="52" pageNumber="53">6b.</emphasis>
Blind side with well-developed pepper-dots and with dermal melanophores along the bases of dorsal-and anal-fin rays....
<emphasis id="B396EAC8FFF35E3FB0963E7BFA295C5E" bold="true" box="[1424,1436,865,886]" pageId="52" pageNumber="53">8</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3A162E7FFFF35E3FB5913E65FA295CC4" pageId="52" pageNumber="53">
<paragraph id="815D36DAFFF35E3FB5913E65FA295C9A" blockId="52.[151,1437,220,1415]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913E65FA295C9A" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913E65FF035CBC" bold="true" box="[151,182,895,916]" pageId="52" pageNumber="53">7a.</emphasis>
Transverse scale rows 2630; longitudinal scale rows 7282; head length 17.320.8% of SL; postorbital length 12.615.2% of SL; peritoneum reddish or bluish-black only on dorsalmost surface...................
<taxonomicName id="46E24D59FFF35E3FB15E3E86FA295C9A" authority="Lee &amp; Munroe" authorityName="Lee &amp; Munroe" authorityYear="2021" box="[1112,1436,924,946]" class="Actinopterygii" family="Cynoglossidae" genus="Symphurus" kingdom="Animalia" order="Pleuronectiformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="brachycephalus">
<emphasis id="B396EAC8FFF35E3FB15E3E86FABD5C99" box="[1112,1288,924,945]" italics="true" pageId="52" pageNumber="53">S. brachycephalus</emphasis>
Lee &amp; Munroe
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="815D36DAFFF35E3FB5913EA0FA295CC4" blockId="52.[151,1437,220,1415]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913EA0FA295CC4" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913EA0FF025CE7" bold="true" box="[151,183,954,975]" pageId="52" pageNumber="53">7b.</emphasis>
Transverse scale rows 3234; longitudinal scale rows 8894; head length 21.923.8% of SL; postorbital length 15.616.6% of SL; entire peritoneum bluish-black...................................................
<taxonomicName id="46E24D59FFF35E3FB19F3ECDFA295CC4" authority="Lee &amp; Munroe" authorityName="Lee &amp; Munroe" authorityYear="2021" box="[1177,1436,983,1004]" class="Actinopterygii" family="Cynoglossidae" genus="Symphurus" kingdom="Animalia" order="Pleuronectiformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="polylepis">
<emphasis id="B396EAC8FFF35E3FB19F3ECDFABD5CC4" box="[1177,1288,983,1004]" italics="true" pageId="52" pageNumber="53">S. polylepis</emphasis>
Lee &amp; Munroe
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3A162E7FFFF35E3FB5913EEEFA295B49" pageId="52" pageNumber="53">
<paragraph id="815D36DAFFF35E3FB5913EEEFA295B0F" blockId="52.[151,1437,220,1415]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913EEEFA295B0F" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913EEEFF005B21" bold="true" box="[151,181,1012,1033]" pageId="52" pageNumber="53">8a.</emphasis>
Transverse scale rows 37; PAL longer than BD; body depth 24.6% of SL; HW/HL= 1.02; ocular side speckled, with irregular blotches.....................................................................
<taxonomicName id="46E24D59FFF35E3FB1823908FA295B0F" authority="Lee &amp; Munroe" authorityName="Lee &amp; Munroe" authorityYear="2021" box="[1156,1436,1041,1063]" class="Actinopterygii" family="Cynoglossidae" genus="Symphurus" kingdom="Animalia" order="Pleuronectiformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="leptosomus">
<emphasis id="B396EAC8FFF35E3FB1823908FABD5B0E" box="[1156,1288,1041,1063]" italics="true" pageId="52" pageNumber="53">S. leptosomus</emphasis>
Lee &amp; Munroe
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="815D36DAFFF35E3FB5913935FA295B49" blockId="52.[151,1437,220,1415]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913935FA295B49" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913935FF025B6C" bold="true" box="[151,183,1071,1092]" pageId="52" pageNumber="53">8b.</emphasis>
Transverse scale rows 2834; PAL shorter than BD; body depth 25.530.3% of SL; HW/HL= 1.061.19; ocular side uniformly light-yellow to brown, sometimes with 57 crossbands.......................................................
<emphasis id="B396EAC8FFF35E3FB0963956FA295B49" bold="true" box="[1424,1436,1100,1121]" pageId="52" pageNumber="53">9</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="3A162E7FFFF35E3FB5913973FA295AAF" pageId="52" pageNumber="53">
<paragraph id="815D36DAFFF35E3FB5913973FA295BDC" blockId="52.[151,1437,220,1415]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB5913973FA295BDC" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB5913973FF1A5B56" bold="true" box="[151,175,1129,1150]" pageId="52" pageNumber="53">9a</emphasis>
. Transverse scale rows 2831; longitudinal scale rows 7281; body thin, TH 15.719.6% of BD; upper jaw length 17.920.4% of HL; snout length 11.716.2% of HL; SNL/ED= 0.861.45; postorbital length 14.516.6% of SL, 71.074.6% of HL; ocular side uniformly light yellow to brown, with well-developed dermal spots at bases of dorsal- and anal-fin rays; eyes oval; anterior margin of upper eye greatly in advance of anterior margin of lower eye, usually extending anteriorly by about 50% of ED of lower eye........................................................................
<taxonomicName id="46E24D59FFF35E3FB1AF39C5FA295BDC" authority="Lee &amp; Munroe" authorityName="Lee &amp; Munroe" authorityYear="2021" box="[1193,1436,1247,1268]" class="Actinopterygii" family="Cynoglossidae" genus="Symphurus" kingdom="Animalia" order="Pleuronectiformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="hongae">
<emphasis id="B396EAC8FFF35E3FB1AF39C5FABD5BDC" box="[1193,1288,1247,1268]" italics="true" pageId="52" pageNumber="53">S. hongae</emphasis>
Lee &amp; Munroe
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="815D36DAFFF35E3FB59139E6FA295AAF" blockId="52.[151,1437,220,1415]" pageId="52" pageNumber="53">
<keyLead id="3A1395EFFFF35E3FB59139E6FA295AAF" pageId="52" pageNumber="53">
<emphasis id="B396EAC8FFF35E3FB59139E6FF025A39" bold="true" box="[151,183,1276,1297]" pageId="52" pageNumber="53">9b.</emphasis>
Transverse scale rows 3134; longitudinal scale rows 7888; body thick, TH 22.130.0% of BD; upper jaw length 21.124.4% of HL; snout length 16.821.2% of HL; SNL/ED= 1.522.00; postorbital length 13.015.0% of SL, 65.870.2% of HL; ocular side with 57 crossbands and scattered dermal spots; eyes slightly round, usually equal in position, or with anterior margin of upper eye slightly in advance of that of lower eye (usually extending beyond lower eye by less than 25% of ED)........................................................................................
<taxonomicName id="46E24D59FFF35E3FB1983868FA295AAF" authority="Lee &amp; Munroe" authorityName="Lee &amp; Munroe" authorityYear="2021" box="[1182,1436,1393,1415]" class="Actinopterygii" family="Cynoglossidae" genus="Symphurus" kingdom="Animalia" order="Pleuronectiformes" pageId="52" pageNumber="53" phylum="Chordata" rank="species" species="robustus">
<emphasis id="B396EAC8FFF35E3FB1983868FABD5AAE" box="[1182,1288,1393,1415]" italics="true" pageId="52" pageNumber="53">S. robustus</emphasis>
Lee &amp; Munroe
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>