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<document id="D5F6627179EE704E988DA549DB2796CF" ID-CLB-Dataset="2487" ID-DOI="10.21248/contrib.entomol.67.1.063-106" ID-GBIF-Dataset="9d4f2ec6-ef3a-4e00-9a24-f7b8370d4dc3" ID-ISSN="0005-805X" ID-Zenodo-Dep="5742262" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638267231974" checkinUser="felipe" docAuthor="Assing, Volker" docDate="2017" docId="08707B47FFCA397EFCFFE92AFB21DE3B" docLanguage="en" docName="ContribEntomol.67.1.63-106.pdf" docOrigin="Beiträge Zur Entomologie = Contributions to Entomology 67 (1)" docStyle="DocumentStyle:02FB06864C20681C17549899CC34643B.2:ContribEntomol.2017-.journal_article" docStyleId="02FB06864C20681C17549899CC34643B" docStyleName="ContribEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Amaurodera longisetosa Assing 2017, spec. nov." docType="treatment" docUuid="887533D9-985B-41FE-B3C8-F1FF8938DC8B" docUuidSource="ZooBank" docVersion="12" lastPageNumber="97" masterDocId="F449033FFFEB395CFFC0EE59FFCFD83A" masterDocTitle="On the Lomechusini fauna of the East Palaearctic and Oriental regions, with a focus on the genera Orphnebius and Amaurodera (Coleoptera: Staphylinidae: Aleocharinae)" masterLastPageNumber="106" masterPageNumber="63" pageNumber="96" updateTime="1699471393470" updateUser="ExternalLinkService">
<mods:mods id="BE55A95377FD597638A7615D69EE0482" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DAF6BB7441581F3F4AEED46956E6C685">
<mods:title id="68E746D8FAC8FACE4ED2A0812AC191BA">On the Lomechusini fauna of the East Palaearctic and Oriental regions, with a focus on the genera Orphnebius and Amaurodera (Coleoptera: Staphylinidae: Aleocharinae)</mods:title>
</mods:titleInfo>
<mods:name id="068AEDC2B37C6EB2E849E09A688D34AF" type="personal">
<mods:role id="5D388F63E9777F175AFB2584F59280C3">
<mods:roleTerm id="09DE3E18DC36893A822F6BD42763B0F9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A4B0AB503C94D0D2A7D4FA73A64AE5C9">Assing, Volker</mods:namePart>
</mods:name>
<mods:typeOfResource id="CE94FA6DD9494549C75496DD26982C73">text</mods:typeOfResource>
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<mods:titleInfo id="B300D5EBE7A757EA0D73F79FCA1EDE69">
<mods:title id="E59BDB7CEDAF1A6769B435EE40F8ADE1">Beiträge Zur Entomologie = Contributions to Entomology</mods:title>
</mods:titleInfo>
<mods:part id="8D8667318676E3EAF0A80304C98998F6">
<mods:date id="B174E26D59874EF08AD723BCC16BE926">2017</mods:date>
<mods:detail id="881E05F4A8D96A7BA91EF77569F2A248" type="pubDate">
<mods:number id="8087BC9FB7553CCA488F58297AC56297">2017-06-30</mods:number>
</mods:detail>
<mods:detail id="250184E52F836243782B94E337E1A798" type="volume">
<mods:number id="DFA97DC6E3EA17A0D147A2F92BB14F97">67</mods:number>
</mods:detail>
<mods:detail id="FD6C84FD2303F9E6CDCF2F634A359ED6" type="issue">
<mods:number id="2ABDED8E80B8E9B969C42EF22E5B26A7">1</mods:number>
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<mods:start id="7E945F37FE2D4D0F211E40B2C5C29285">63</mods:start>
<mods:end id="90D2E0AFE573829AE9C7228D23E95F65">106</mods:end>
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<mods:classification id="2773F30B2D4ED457A790A8A9A289A07A">journal article</mods:classification>
<mods:identifier id="E8A0597C5882C3FF343E67E31947E112" type="CLB-Dataset">2487</mods:identifier>
<mods:identifier id="FD405EA8B19908FDC937D51A88565ED7" type="DOI">10.21248/contrib.entomol.67.1.063-106</mods:identifier>
<mods:identifier id="DA31B5A74AA5CB23F34772EF03F4DFA7" type="GBIF-Dataset">9d4f2ec6-ef3a-4e00-9a24-f7b8370d4dc3</mods:identifier>
<mods:identifier id="5E4E991CEC2623194DB0B1383CAE60B2" type="ISSN">0005-805X</mods:identifier>
<mods:identifier id="38D4A2E2E76E93676E2804284BE8565F" type="Zenodo-Dep">5742262</mods:identifier>
<mods:identifier id="B9CE07D53916E04F3D9DBE502855DBCE" type="ZooBank">0C30D2D6-63EC-49ED-8FF6-49417D1019B2</mods:identifier>
</mods:mods>
<treatment id="08707B47FFCA397EFCFFE92AFB21DE3B" ID-DOI="http://doi.org/10.5281/zenodo.5885112" ID-GBIF-Taxon="192459948" ID-Zenodo-Dep="5885112" LSID="urn:lsid:zoobank.org:act:887533D9-985B-41FE-B3C8-F1FF8938DC8B" httpUri="http://treatment.plazi.org/id/08707B47FFCA397EFCFFE92AFB21DE3B" lastPageId="34" lastPageNumber="97" pageId="33" pageNumber="96">
<subSubSection id="C8C399DAFFCA397DFCFFE92AFA8EDF95" pageId="33" pageNumber="96" type="nomenclature">
<paragraph id="8066CA51FFCA397DFCFFE92AFB66DFB4" blockId="33.[831,1345,1907,1999]" box="[831,1193,1907,1934]" pageId="33" pageNumber="96">
<heading id="DB2E7D3DFFCA397DFCFFE92AFB66DFB4" box="[831,1193,1907,1934]" fontSize="12" level="2" pageId="33" pageNumber="96" reason="2">
<taxonomicName id="47D9B1D2FFCA397DFCFFE92AFBF5DFB4" ID-CoL="CJLQ" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[831,1082,1907,1934]" class="Insecta" family="Staphylinidae" genus="Amaurodera" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="96" phylum="Arthropoda" rank="species" species="longisetosa" status="sp. nov.">
<emphasis id="B2AD1643FFCA397DFCFFE92AFBF5DFB4" box="[831,1082,1907,1934]" italics="true" pageId="33" pageNumber="96">Amaurodera longisetosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A99EAB38FFCA397DFB80E92AFB66DFB4" box="[1088,1193,1907,1934]" pageId="33" pageNumber="96" rank="species">spec. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph id="8066CA51FFCA397DFCFFE9C2FA8EDF95" blockId="33.[831,1345,1907,1999]" box="[831,1345,1947,1967]" pageId="33" pageNumber="96">
<uri id="F448C653FFCA397DFCFFE9C2FA8EDF95" box="[831,1345,1947,1967]" pageId="33" pageNumber="96">
urn:lsid:zoobank.org:act:
<uuid id="F47FF084FFCA397DFC30E9C2FA8EDF95" box="[1008,1345,1947,1967]" pageId="33" pageNumber="96">887533D9-985B-41FE-B3C8-F1FF8938DC8B</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C8C399DAFFCA397DFCFFE9E2FC30DFF5" box="[831,1023,1979,1999]" pageId="33" pageNumber="96" type="description">
<paragraph id="8066CA51FFCA397DFCFFE9E2FC30DFF5" blockId="33.[831,1345,1907,1999]" box="[831,1023,1979,1999]" pageId="33" pageNumber="96">
<figureCitation id="18E2D6D4FFCA397DFCFFE9E2FCB8DFF5" box="[831,887,1979,1999]" captionStart="Figs 1–30" captionStartId="9.[151,191,1935,1957]" captionTargetBox="[151,1397,244,1921]" captionTargetId="figure-202@9.[151,1398,234,1933]" captionTargetPageId="9" captionText="Figs 1–30: Antenna (1–19), forebody (20, 23, 26), abdomen (21, 24, 27, 29), antero-lateral portion of abdomen in lateral-dorsal view (22, 25, 28), and abdominal segment VII (30) of Orphnebius triacuminatus (1, 20–22), O. gracilis (2, 23–25), O. protuberatus (3, 26–28), O. pertortus (4, 29–30), O. reticulipennis (5), Zyrastilbus almorensis (6), Pheidologitonetes bursata (7), Rabdotodrusilla vara (8), Amaurodera bicarinata (9), A. gilvicornis (10), A. latisulcata (11), A. brevipes (12), A. migritheca (13), A. longisetosa (14), A. calicitheca (15), Drusilla smetanai (16), D. nepalensis (17), D. lativentris (18), and Witteia tensa (19). Scale bars: 1–29: 1.0 mm; 30: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742266" httpUri="https://zenodo.org/record/5742266/files/figure.png" pageId="33" pageNumber="96">Figs 14</figureCitation>
,
<figureCitation id="18E2D6D4FFCA397DFCBEE9E2FC7EDFF5" box="[894,945,1979,1999]" captionStart="Figs 43–57" captionStartId="13.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1963]" captionTargetId="figure-141@13.[151,1398,232,1971]" captionText="Figs 43–57: Amaurodera bicarinata (43–46), A. gilvicornis (47–48), A. latisulcata (49–50), A. brevipes (51), A. migritheca (52–54), A. longisetosa (55–56), and A. calicitheca (57): forebody (43, 47, 49, 51, 52, 55, 57); pronotum in dorsal view (44, 50, 53); pronotum in dorso-lateral view (45, 56); abdomen (46, 48, 54). Scale bars: 43, 46–49, 51–52, 54–55, 57: 1.0 mm; 44–45, 50, 53, 56: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5885053" httpUri="https://zenodo.org/record/5885053/files/figure.png" pageId="33" pageNumber="96">55–56</figureCitation>
,
<figureCitation id="18E2D6D4FFCA397DFC78E9E2FC34DFF5" box="[952,1019,1979,1999]" captionStart="Figs 150–172" captionStartId="19.[151,191,1942,1964]" captionTargetBox="[168,1384,241,1927]" captionTargetId="figure-160@19.[165,1384,231,1931]" captionText="Figs 150–172: Amaurodera frondium (150–151), A calicitheca (152–158), A. discoidea, paratype (159), A. longisetosa (160–163), A. bulbosa (paratypes of A. discoidea from Sabah) (164–165), Rabdotodrusilla vara (166–169), and Witteia tensa (170–172): median lobe of aedeagus in lateral and in ventral view (150–155, 160–162, 164–165, 168–169, 170–171); apical portion of median lobe in ventral view (156); spermatheca (157–159, 163, 172); male tergite VIII (166); male sternite VIII (167). Scale bars: 150–155, 157–165: 0.5 mm; 166–172: 0.2 mm; 156: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5885063" httpUri="https://zenodo.org/record/5885063/files/figure.png" pageId="33" pageNumber="96">160–163</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C8C399DAFFCA397EFCDAE9B0FE98D9DB" lastPageId="34" lastPageNumber="97" pageId="33" pageNumber="96" type="materials_examined">
<paragraph id="8066CA51FFCA397EFCDAE9B0FE98D9DB" blockId="33.[794,1398,2025,2081]" lastBlockId="34.[189,794,233,481]" lastPageId="34" lastPageNumber="97" pageId="33" pageNumber="96">
<emphasis id="B2AD1643FFCA397DFCDAE9B0FC74D03B" bold="true" box="[794,955,2025,2049]" pageId="33" pageNumber="96">Type material</emphasis>
:
<materialsCitation id="30B1C00CFFCA397EFC12E9B0FD97D91B" ID-GBIF-Occurrence="3449505445" collectingDate="1987-05-17" collectingDateMax="2007-02-16" collectingDateMin="1987-05-09" collectionCode="MHNG" collectorName="Burckhardt - Lobl" county="Kota Kinabalu - Tambunan" determinerName="V. Assing" elevation="1350" lastPageId="34" lastPageNumber="97" location="km 60 Kota Kinabalu - Tambunan" municipality="Crocker Ra" pageId="33" pageNumber="96" specimenCount="2" stateProvince="SABAH" typeStatus="holotype">
<typeStatus id="5F6274F3FFCA397DFC12E9B0FBFAD03B" box="[978,1077,2025,2049]" pageId="33" pageNumber="96" type="holotype">Holotype</typeStatus>
<specimenCount id="96DF01D8FFCA397DFB86E9B3FB94D03B" box="[1094,1115,2026,2049]" pageId="33" pageNumber="96">♂</specimenCount>
: “
<collectingRegion id="421D04B3FFCA397DFBBAE9B0FB1CD03B" box="[1146,1235,2025,2049]" country="Malaysia" name="Sabah" pageId="33" pageNumber="96">SABAH</collectingRegion>
:
<collectingMunicipality id="6002502BFFCA397DFB26E9B0FAA0D03B" box="[1254,1391,2025,2049]" county="Kota Kinabalu - Tambunan" municipality="Burckhardt - Lobl" name="Crocker Ra" pageId="33" pageNumber="96" stateProvince="SABAH">Crocker Ra</collectingMunicipality>
.,
<quantity id="472167B4FFCA397DFCDAE650FCA4D01B" box="[794,875,2057,2081]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="33" pageNumber="96" unit="m" value="1350.0">
<elevation id="0BF42D62FFCA397DFCDAE650FCA4D01B" box="[794,875,2057,2081]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="33" pageNumber="96" unit="m" value="1350.0">1350 m</elevation>
</quantity>
,
<location id="85069C8AFFCA397DFCBEE650FB39D01B" LSID="urn:lsid:plazi:treatment:08707B47FFCA397EFCFFE92AFB21DE3B:85069C8AFFCA397DFCBEE650FB39D01B" box="[894,1270,2057,2081]" county="Kota Kinabalu - Tambunan" municipality="Crocker Ra" name="km 60 Kota Kinabalu - Tambunan" pageId="33" pageNumber="96" stateProvince="SABAH">km 60 Kota Kinabalu–Tambunan</location>
,
<date id="F467EC91FFCA397DFAC8E650FABDD01B" box="[1288,1394,2057,2081]" pageId="33" pageNumber="96" value="1987-05-17">
<collectingDate id="E4231579FFCA397DFAC8E650FABDD01B" box="[1288,1394,2057,2081]" pageId="33" pageNumber="96" value="1987-05-17">17.V.1987</collectingDate>
</date>
<collectorName id="2D2CAF87FFCA397EFAB2E650FEB4D93B" lastPageId="34" lastPageNumber="97" pageId="33" pageNumber="96">Burckhardt - Löbl</collectorName>
/
<typeStatus id="5F6274F3FFC9397EFE4FEEB0FE31D93B" box="[399,510,233,257]" pageId="34" pageNumber="97" type="holotype">Holotypus</typeStatus>
♂
<taxonomicName id="47D9B1D2FFC9397EFDDEEEB0FCD6D93B" authority="Assing, 2017" authorityName="Assing" authorityYear="2017" box="[542,793,233,257]" class="Insecta" family="Staphylinidae" genus="Amaurodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="97" phylum="Arthropoda" rank="species" species="longisetosa" status="sp. nov.">
<emphasis id="B2AD1643FFC9397EFDDEEEB0FCD6D93B" box="[542,793,233,257]" italics="true" pageId="34" pageNumber="97">
<location id="85069C8AFFC9397EFDDEEEB0FD6DD93B" LSID="urn:lsid:plazi:treatment:08707B47FFCA397EFCFFE92AFB21DE3B:85069C8AFFC9397EFDDEEEB0FD6DD93B" box="[542,674,233,257]" county="Kota Kinabalu - Tambunan" municipality="Crocker Ra" name="Amaurodera" pageId="34" pageNumber="97" stateProvince="SABAH">Amaurodera</location>
longisetosa
</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A99EAB38FFC9397EFF7DEF53FF3AD91B" box="[189,245,266,289]" pageId="34" pageNumber="97" rank="species">
<specimenCount id="96DF01D8FFC9397EFF7DEF53FF15D91B" box="[189,218,266,289]" pageId="34" pageNumber="97" type="generic">sp.</specimenCount>
n.
</taxonomicNameLabel>
, det.
<determinerName id="75BA2695FFC9397EFEF7EF53FE56D91B" box="[311,409,265,289]" pageId="34" pageNumber="97">V. Assing</determinerName>
2016” (
<collectionCode id="E6C85294FFC9397EFE34EF50FD82D91B" box="[500,589,265,289]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="34" pageNumber="97" type="Museum">MHNG</collectionCode>
).
</materialsCitation>
<materialsCitation id="30B1C00CFFC9397EFDA2EF53FF32D9BB" ID-GBIF-Occurrence="3449505364" collectingDate="1987-05-17" collectingDateMax="2007-02-16" collectingDateMin="1987-05-09" collectionCode="MHNG" collectorName="Crocker Ra. &amp; Burckhardt - Lobl" county="Kota Kinabalu - Tambunan" determinerName="V. Assing" elevation="1270" location="km 60 rte Kota Kinabalu - Tambunan" municipality="Burckhardt - Lobl" pageId="34" pageNumber="97" specimenCount="8" stateProvince="SABAH" typeStatus="paratype">
<typeStatus id="5F6274F3FFC9397EFDA2EF53FD03D918" box="[610,716,266,290]" pageId="34" pageNumber="97" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="96DF01D8FFC9397EFD19EF50FCD8D91B" box="[729,791,265,289]" pageId="34" pageNumber="97" type="generic">8 exs.</specimenCount>
: “
<collectingRegion id="421D04B3FFC9397EFF06EF70FED0D97B" box="[198,287,297,321]" country="Malaysia" name="Sabah" pageId="34" pageNumber="97">SABAH</collectingRegion>
:
<collectorName id="2D2CAF87FFC9397EFEEEEF70FE79D97B" box="[302,438,297,321]" pageId="34" pageNumber="97">Crocker Ra.</collectorName>
,
<quantity id="472167B4FFC9397EFE05EF70FDD9D97B" box="[453,534,297,321]" metricMagnitude="3" metricUnit="m" metricValue="1.27" pageId="34" pageNumber="97" unit="m" value="1270.0">
<elevation id="0BF42D62FFC9397EFE05EF70FDD9D97B" box="[453,534,297,321]" metricMagnitude="3" metricUnit="m" metricValue="1.27" pageId="34" pageNumber="97" unit="m" value="1270.0">1270 m</elevation>
</quantity>
,
<location id="85069C8AFFC9397EFDE6EF70FEA1D95B" LSID="urn:lsid:plazi:treatment:08707B47FFCA397EFCFFE92AFB21DE3B:85069C8AFFC9397EFDE6EF70FEA1D95B" county="Kota Kinabalu - Tambunan" municipality="Burckhardt - Lobl" name="km 60 rte Kota Kinabalu - Tambunan" pageId="34" pageNumber="97" stateProvince="SABAH">km 60 rte Kota Kinabalu–Tambunan</location>
,
<date id="F467EC91FFC9397EFEBDEF10FE02D95B" box="[381,461,329,353]" pageId="34" pageNumber="97" value="1987-05-17">
<collectingDate id="E4231579FFC9397EFEBDEF10FE02D95B" box="[381,461,329,353]" pageId="34" pageNumber="97" value="1987-05-17">17.V.87</collectingDate>
</date>
,
<collectorName id="2D2CAF87FFC9397EFE1BEF10FD6BD95B" box="[475,676,329,353]" pageId="34" pageNumber="97">Burckhardt - Löbl</collectorName>
” (
<collectionCode id="E6C85294FFC9397EFD7FEF10FCD9D95B" box="[703,790,329,353]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="34" pageNumber="97" type="Museum">MHNG</collectionCode>
, cAss);
</materialsCitation>
<materialsCitation id="30B1C00CFFC9397EFEC9EF30FDB8D99B" ID-GBIF-Occurrence="3449505439" collectingDate="1987-05-09" collectingDateMax="2007-02-16" collectingDateMin="1987-05-09" collectionCode="MHNG" collectorName="Burckhardt - Lobl" county="Kota Kinabalu - Tambunan" determinerName="V. Assing" elevation="575" location="Poring Hot Springs" municipality="Burckhardt - Lobl" pageId="34" pageNumber="97" specimenCount="1" stateProvince="SABAH" typeStatus="holotype">
<specimenCount id="96DF01D8FFC9397EFEC9EF30FEE1D9BB" box="[265,302,361,385]" pageId="34" pageNumber="97">1 ♀</specimenCount>
: “
<collectingRegion id="421D04B3FFC9397EFE8AEF30FE6CD9BB" box="[330,419,361,385]" country="Malaysia" name="Sabah" pageId="34" pageNumber="97">SABAH</collectingRegion>
:
<location id="85069C8AFFC9397EFE72EF30FD42D9BB" LSID="urn:lsid:plazi:treatment:08707B47FFCA397EFCFFE92AFB21DE3B:85069C8AFFC9397EFE72EF30FD42D9BB" box="[434,653,361,385]" county="Kota Kinabalu - Tambunan" municipality="Burckhardt - Lobl" name="Poring Hot Springs" pageId="34" pageNumber="97" stateProvince="SABAH">Poring Hot Springs</location>
,
<quantity id="472167B4FFC9397EFD5DEF30FCD9D9BB" box="[669,790,361,385]" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="34" pageNumber="97" unit="m" value="575.0" valueMax="600.0" valueMin="550.0">
<elevation id="0BF42D62FFC9397EFD5DEF30FCD9D9BB" box="[669,790,361,385]" metricMagnitude="2" metricUnit="m" metricValue="5.75" metricValueMax="6.0" metricValueMin="5.5" pageId="34" pageNumber="97" unit="m" value="575.0" valueMax="600.0" valueMin="550.0">550–600 m</elevation>
</quantity>
,
<date id="F467EC91FFC9397EFF7DEFD0FED5D99B" box="[189,282,393,417]" pageId="34" pageNumber="97" value="1987-05-09">
<collectingDate id="E4231579FFC9397EFF7DEFD0FED5D99B" box="[189,282,393,417]" pageId="34" pageNumber="97" value="1987-05-09">9.V.1987</collectingDate>
</date>
,
<collectorName id="2D2CAF87FFC9397EFEEAEFD0FE39D99B" box="[298,502,393,417]" pageId="34" pageNumber="97">Burckhardt - Löbl</collectorName>
” (
<collectionCode id="E6C85294FFC9397EFDD3EFD0FDA4D99B" box="[531,619,393,417]" collectionName="Switzerland, Geneva, Museum d'Histoire Naturelle" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="34" pageNumber="97" type="Museum">MHNG</collectionCode>
);
</materialsCitation>
<materialsCitation id="30B1C00CFFC9397EFD44EFD0FE9CD9DB" ID-GBIF-Occurrence="3449505421" collectingDate="2007-02-14" collectingDateMax="2007-02-16" collectingDateMin="2007-02-14" collectionCode="MHNG" collectorName="Danum Valley, B. R. L. &amp; G. de Rougemont" county="Kota Kinabalu - Tambunan" determinerName="V. Assing" elevation="1350" location="SABAH" municipality="Burckhardt - Lobl" pageId="34" pageNumber="97" specimenCount="1" stateProvince="SABAH" typeStatus="holotype">
1 ♀: “
<collectingRegion id="421D04B3FFC9397EFD04EFD0FCD5D99B" box="[708,794,393,417]" country="Malaysia" name="Sabah" pageId="34" pageNumber="97">SABAH</collectingRegion>
<collectorName id="2D2CAF87FFC9397EFF7DEFF3FE64D9FB" box="[189,427,425,449]" pageId="34" pageNumber="97">Danum Valley, B.R.L.</collectorName>
, f.i.t.
<date id="F467EC91FFC9397EFE2BEFF0FD4FD9FB" box="[491,640,425,449]" pageId="34" pageNumber="97" value="2007-02-14" valueMax="2007-02-16" valueMin="2007-02-14">
<collectingDate id="E4231579FFC9397EFE2BEFF0FD4FD9FB" box="[491,640,425,449]" pageId="34" pageNumber="97" value="2007-02-14" valueMax="2007-02-16" valueMin="2007-02-14">14–16.II.2007</collectingDate>
</date>
,
<collectorName id="2D2CAF87FFC9397EFD4EEFF0FF38D9DB" pageId="34" pageNumber="97">G. de Rougemont</collectorName>
” (cRou)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C8C399DAFFC9397EFF7DEC50FE26DA7B" pageId="34" pageNumber="97" type="etymology">
<paragraph id="8066CA51FFC9397EFF7DEC50FE26DA7B" blockId="34.[189,794,521,577]" pageId="34" pageNumber="97">
<emphasis id="B2AD1643FFC9397EFF7DEC50FEFBDA1B" bold="true" box="[189,308,521,545]" pageId="34" pageNumber="97">Etymology</emphasis>
: The specific epithet (adjective) alludes to the long setae on the pronotum.
</paragraph>
</subSubSection>
<subSubSection id="C8C399DAFFC9397EFF7DEC30FC64DD5B" pageId="34" pageNumber="97" type="description">
<paragraph id="8066CA51FFC9397EFF7DEC30FDCEDCBB" blockId="34.[189,795,617,2081]" pageId="34" pageNumber="97">
<emphasis id="B2AD1643FFC9397EFF7DEC30FEF0DABB" bold="true" box="[189,319,617,641]" pageId="34" pageNumber="97">Description</emphasis>
: Body length
<quantity id="472167B4FFC9397EFE25EC30FDA9DABB" box="[485,614,617,641]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" metricValueMax="5.5" metricValueMin="4.4" pageId="34" pageNumber="97" unit="mm" value="4.95" valueMax="5.5" valueMin="4.4">4.4–5.5 mm</quantity>
; length of forebody
<quantity id="472167B4FFC9397EFEC2ECD0FE4CDA9B" box="[258,387,649,673]" metricMagnitude="-3" metricUnit="m" metricValue="2.4" metricValueMax="2.6" metricValueMin="2.2" pageId="34" pageNumber="97" unit="mm" value="2.4" valueMax="2.6" valueMin="2.2">2.2–2.6 mm</quantity>
. Other measurements: head width:
<quantity id="472167B4FFC9397EFF7DECF0FE97DAFB" box="[189,344,681,705]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" metricValueMax="7.8" metricValueMin="6.0" pageId="34" pageNumber="97" unit="mm" value="0.69" valueMax="0.78" valueMin="0.6">0.60–0.78 mm</quantity>
; length of pronotum:
<quantity id="472167B4FFC9397EFDF9ECF0FD1BDAFB" box="[569,724,681,705]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" metricValueMax="9.8" metricValueMin="7.8" pageId="34" pageNumber="97" unit="mm" value="0.88" valueMax="0.98" valueMin="0.78">0.78–0.98 mm</quantity>
; width of pronotum:
<quantity id="472167B4FFC9397EFEA1EC90FE34DADB" box="[353,507,713,737]" metricMagnitude="-4" metricUnit="m" metricValue="6.45" metricValueMax="7.1" metricValueMin="5.8" pageId="34" pageNumber="97" unit="mm" value="0.645" valueMax="0.71" valueMin="0.58">0.58–0.71 mm</quantity>
; elytral length at suture:
<quantity id="472167B4FFC9397EFF7DECB0FE97DB3B" box="[189,344,745,769]" metricMagnitude="-4" metricUnit="m" metricValue="5.3" metricValueMax="5.8" metricValueMin="4.8" pageId="34" pageNumber="97" unit="mm" value="0.53" valueMax="0.58" valueMin="0.48">0.48–0.58 mm</quantity>
; elytral width:
<quantity id="472167B4FFC9397EFE3EECB0FD56DB3B" box="[510,665,745,769]" metricMagnitude="-4" metricUnit="m" metricValue="9.65" metricValueMax="10.5" metricValueMin="8.8" pageId="34" pageNumber="97" unit="mm" value="0.965" valueMax="1.05" valueMin="0.88">0.88–1.05 mm</quantity>
. Coloration (
<figureCitation id="18E2D6D4FFC9397EFF04ED50FEDCDB1B" box="[196,275,777,801]" captionStart="Figs 1–30" captionStartId="9.[151,191,1935,1957]" captionTargetBox="[151,1397,244,1921]" captionTargetId="figure-202@9.[151,1398,234,1933]" captionTargetPageId="9" captionText="Figs 1–30: Antenna (1–19), forebody (20, 23, 26), abdomen (21, 24, 27, 29), antero-lateral portion of abdomen in lateral-dorsal view (22, 25, 28), and abdominal segment VII (30) of Orphnebius triacuminatus (1, 20–22), O. gracilis (2, 23–25), O. protuberatus (3, 26–28), O. pertortus (4, 29–30), O. reticulipennis (5), Zyrastilbus almorensis (6), Pheidologitonetes bursata (7), Rabdotodrusilla vara (8), Amaurodera bicarinata (9), A. gilvicornis (10), A. latisulcata (11), A. brevipes (12), A. migritheca (13), A. longisetosa (14), A. calicitheca (15), Drusilla smetanai (16), D. nepalensis (17), D. lativentris (18), and Witteia tensa (19). Scale bars: 1–29: 1.0 mm; 30: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742266" httpUri="https://zenodo.org/record/5742266/files/figure.png" pageId="34" pageNumber="97">Figs 14</figureCitation>
,
<figureCitation id="18E2D6D4FFC9397EFEDEED50FEF4DB1B" box="[286,315,777,801]" captionStart="Figs 43–57" captionStartId="13.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1963]" captionTargetId="figure-141@13.[151,1398,232,1971]" captionText="Figs 43–57: Amaurodera bicarinata (43–46), A. gilvicornis (47–48), A. latisulcata (49–50), A. brevipes (51), A. migritheca (52–54), A. longisetosa (55–56), and A. calicitheca (57): forebody (43, 47, 49, 51, 52, 55, 57); pronotum in dorsal view (44, 50, 53); pronotum in dorso-lateral view (45, 56); abdomen (46, 48, 54). Scale bars: 43, 46–49, 51–52, 54–55, 57: 1.0 mm; 44–45, 50, 53, 56: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5885053" httpUri="https://zenodo.org/record/5885053/files/figure.png" pageId="34" pageNumber="97">55</figureCitation>
): head dark–brown to blackish; pronotum dark–brown to blackish-brown; elytra dark reddish to reddish-brown with the postero-lateral portions extensively darker; abdomen blackish-brown with the anterior segments usually slightly – or partly – paler; legs palebrown, usually with the profemora and the apical halves of the meso- and metafemora, sometimes also the basal portions of the meso- and metatibiae, somewhat darker; antennae dark-brown to blackish-brown with the apical antennomeres more or less extensively yellow to darkyellow; maxillary palpi brown to dark-brown with the terminal palpomere yellowish.
</paragraph>
<paragraph id="8066CA51FFC9397EFF7DEAD0FE1BDEBB" blockId="34.[189,795,617,2081]" pageId="34" pageNumber="97">
Head (
<figureCitation id="18E2D6D4FFC9397EFEC9EAD0FE9BDC9B" box="[265,340,1161,1185]" captionStart="Figs 43–57" captionStartId="13.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1963]" captionTargetId="figure-141@13.[151,1398,232,1971]" captionText="Figs 43–57: Amaurodera bicarinata (43–46), A. gilvicornis (47–48), A. latisulcata (49–50), A. brevipes (51), A. migritheca (52–54), A. longisetosa (55–56), and A. calicitheca (57): forebody (43, 47, 49, 51, 52, 55, 57); pronotum in dorsal view (44, 50, 53); pronotum in dorso-lateral view (45, 56); abdomen (46, 48, 54). Scale bars: 43, 46–49, 51–52, 54–55, 57: 1.0 mm; 44–45, 50, 53, 56: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5885053" httpUri="https://zenodo.org/record/5885053/files/figure.png" pageId="34" pageNumber="97">Fig. 55</figureCitation>
) posteriorly nearly truncate and of very variable shape (see measurements), in larger specimens relatively larger and more transverse than in smaller specimens; postero-lateral outline between eyes and posterior constriction more or less strongly convex in dorsal view; median dorsal portion not impressed; punctation moderately sparse and fine; interstices with or without extremely shallow traces of microreticulation visible only at high magnification (100 x). Eyes moderately large and convex, 0.5–0.7 times as long as distance from posterior margin of eye to posterior constriction (relative size dependent on relative head size, smaller in large-headed than in smallheaded specimens). Antenna (
<figureCitation id="18E2D6D4FFC9397EFDC5E850FD9FDE1B" box="[517,592,1545,1569]" captionStart="Figs 1–30" captionStartId="9.[151,191,1935,1957]" captionTargetBox="[151,1397,244,1921]" captionTargetId="figure-202@9.[151,1398,234,1933]" captionTargetPageId="9" captionText="Figs 1–30: Antenna (1–19), forebody (20, 23, 26), abdomen (21, 24, 27, 29), antero-lateral portion of abdomen in lateral-dorsal view (22, 25, 28), and abdominal segment VII (30) of Orphnebius triacuminatus (1, 20–22), O. gracilis (2, 23–25), O. protuberatus (3, 26–28), O. pertortus (4, 29–30), O. reticulipennis (5), Zyrastilbus almorensis (6), Pheidologitonetes bursata (7), Rabdotodrusilla vara (8), Amaurodera bicarinata (9), A. gilvicornis (10), A. latisulcata (11), A. brevipes (12), A. migritheca (13), A. longisetosa (14), A. calicitheca (15), Drusilla smetanai (16), D. nepalensis (17), D. lativentris (18), and Witteia tensa (19). Scale bars: 1–29: 1.0 mm; 30: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5742266" httpUri="https://zenodo.org/record/5742266/files/figure.png" pageId="34" pageNumber="97">Fig. 14</figureCitation>
) long and slender,
<quantity id="472167B4FFC9397EFF7DE870FEF3DE7B" box="[189,316,1577,1601]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="34" pageNumber="97" unit="mm" value="2.5" valueMax="2.7" valueMin="2.3">2.3–2.7 mm</quantity>
long; antennomere IX approximately twice as long as broad and antennomere X approximately 1.5 times as long as broad.
</paragraph>
<paragraph id="8066CA51FFC9397EFF7DE8D3FD17DF5B" blockId="34.[189,795,617,2081]" pageId="34" pageNumber="97">
Pronotum (
<figureCitation id="18E2D6D4FFC9397EFEFFE8D0FE46DE9B" box="[319,393,1673,1697]" captionStart="Figs 43–57" captionStartId="13.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1963]" captionTargetId="figure-141@13.[151,1398,232,1971]" captionText="Figs 43–57: Amaurodera bicarinata (43–46), A. gilvicornis (47–48), A. latisulcata (49–50), A. brevipes (51), A. migritheca (52–54), A. longisetosa (55–56), and A. calicitheca (57): forebody (43, 47, 49, 51, 52, 55, 57); pronotum in dorsal view (44, 50, 53); pronotum in dorso-lateral view (45, 56); abdomen (46, 48, 54). Scale bars: 43, 46–49, 51–52, 54–55, 57: 1.0 mm; 44–45, 50, 53, 56: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5885053" httpUri="https://zenodo.org/record/5885053/files/figure.png" pageId="34" pageNumber="97">Fig. 56</figureCitation>
) moderately slender, approximately 1.36 times as long as broad, without sexual dimorphism; dorsal surface with very dense microgranules and opaque; midline with narrow median sulcus reaching neither anterior nor posterior margins; lateral portions with 4–6 long, stout, and erect black setae on either side; anterolateral portions additionally with some short setae.
</paragraph>
<paragraph id="8066CA51FFC9397EFF7DE930FE34D01B" blockId="34.[189,795,617,2081]" pageId="34" pageNumber="97">
Elytra (
<figureCitation id="18E2D6D4FFC9397EFECAE930FE9ADFBB" box="[266,341,1897,1921]" captionStart="Figs 43–57" captionStartId="13.[151,191,2001,2023]" captionTargetBox="[151,1398,232,1963]" captionTargetId="figure-141@13.[151,1398,232,1971]" captionText="Figs 43–57: Amaurodera bicarinata (43–46), A. gilvicornis (47–48), A. latisulcata (49–50), A. brevipes (51), A. migritheca (52–54), A. longisetosa (55–56), and A. calicitheca (57): forebody (43, 47, 49, 51, 52, 55, 57); pronotum in dorsal view (44, 50, 53); pronotum in dorso-lateral view (45, 56); abdomen (46, 48, 54). Scale bars: 43, 46–49, 51–52, 54–55, 57: 1.0 mm; 44–45, 50, 53, 56: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5885053" httpUri="https://zenodo.org/record/5885053/files/figure.png" pageId="34" pageNumber="97">Fig. 55</figureCitation>
) approximately 0.6 times as long as pronotum; punctation extremely fine (barely noticeable even at high magnification) and moderately dense; microreticulation absent. Hind wings fully developed. Legs including tarsi long and slender; metatarsomere I shorter than the combined length of II and III.
</paragraph>
<paragraph id="8066CA51FFC9397EFC80EEB0FC73D9DB" blockId="34.[832,1437,233,609]" pageId="34" pageNumber="97">Abdomen narrower than elytra; tergites III–V with moderately deep anterior impressions; tergites III–VII with sparse punctures on disc (especially laterally) and at posterior margin; tergal surfaces with nearly obsolete microreticulation visible only at high magnification (100 x); posterior margin of tergite VII with palisade fringe; posterior margin of tergite VIII weakly convex and serrate.</paragraph>
<paragraph id="8066CA51FFC9397EFC80EFB3FAA2DA7B" blockId="34.[832,1437,233,609]" pageId="34" pageNumber="97">
♂: median lobe of aedeagus (
<figureCitation id="18E2D6D4FFC9397EFB5CEFB0FAE3DA3B" box="[1180,1324,489,513]" captionStart="Figs 150–172" captionStartId="19.[151,191,1942,1964]" captionTargetBox="[168,1384,241,1927]" captionTargetId="figure-160@19.[165,1384,231,1931]" captionText="Figs 150–172: Amaurodera frondium (150–151), A calicitheca (152–158), A. discoidea, paratype (159), A. longisetosa (160–163), A. bulbosa (paratypes of A. discoidea from Sabah) (164–165), Rabdotodrusilla vara (166–169), and Witteia tensa (170–172): median lobe of aedeagus in lateral and in ventral view (150–155, 160–162, 164–165, 168–169, 170–171); apical portion of median lobe in ventral view (156); spermatheca (157–159, 163, 172); male tergite VIII (166); male sternite VIII (167). Scale bars: 150–155, 157–165: 0.5 mm; 166–172: 0.2 mm; 156: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5885063" httpUri="https://zenodo.org/record/5885063/files/figure.png" pageId="34" pageNumber="97">Figs 160–162</figureCitation>
) approximately
<quantity id="472167B4FFC9397EFC4DEC50FC10DA1B" box="[909,991,521,545]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="34" pageNumber="97" unit="mm" value="0.6">0.6 mm</quantity>
long; ventral process of distinctive shape, basally broad, lateral margins angled in ventral view.
</paragraph>
<paragraph id="8066CA51FFC9397EFC80EC13FB1EDA5B" blockId="34.[832,1437,233,609]" box="[832,1233,585,609]" pageId="34" pageNumber="97">
♀: spermatheca shaped as in
<figureCitation id="18E2D6D4FFC9397EFBB6EC10FB02DA5B" box="[1142,1229,585,609]" captionStart="Figs 150–172" captionStartId="19.[151,191,1942,1964]" captionTargetBox="[168,1384,241,1927]" captionTargetId="figure-160@19.[165,1384,231,1931]" captionText="Figs 150–172: Amaurodera frondium (150–151), A calicitheca (152–158), A. discoidea, paratype (159), A. longisetosa (160–163), A. bulbosa (paratypes of A. discoidea from Sabah) (164–165), Rabdotodrusilla vara (166–169), and Witteia tensa (170–172): median lobe of aedeagus in lateral and in ventral view (150–155, 160–162, 164–165, 168–169, 170–171); apical portion of median lobe in ventral view (156); spermatheca (157–159, 163, 172); male tergite VIII (166); male sternite VIII (167). Scale bars: 150–155, 157–165: 0.5 mm; 166–172: 0.2 mm; 156: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5885063" httpUri="https://zenodo.org/record/5885063/files/figure.png" pageId="34" pageNumber="97">Fig. 163</figureCitation>
.
</paragraph>
<paragraph id="8066CA51FFC9397EFC80ECD0FBE0DB5B" blockId="34.[832,1437,649,865]" pageId="34" pageNumber="97">
<emphasis id="B2AD1643FFC9397EFC80ECD0FBFBDA9B" bold="true" box="[832,1076,649,673]" pageId="34" pageNumber="97">Intraspecific variation</emphasis>
: Body size, head shape, and relative head size are remarkably variable. Moreover, the spermatheca of the female from Poring Hot Springs differs by shorter length (
<quantity id="472167B4FFC9397EFB91ECB0FB7CDB3B" box="[1105,1203,745,769]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="34" pageNumber="97" unit="mm" value="0.3">0.30 mm</quantity>
) and the shape of the distal portion. However, since no additional differences were observed, this is attributed to intra- rather than interspecific variation.
</paragraph>
<paragraph id="8066CA51FFC9397EFC80EDD0FC64DD5B" blockId="34.[832,1437,905,1377]" pageId="34" pageNumber="97">
<emphasis id="B2AD1643FFC9397EFC80EDD0FBD6DB9B" bold="true" box="[832,1049,905,929]" pageId="34" pageNumber="97">Comparative notes</emphasis>
:
<taxonomicName id="47D9B1D2FFC9397EFBEEEDD0FAFBDB9B" authorityName="Assing" authorityYear="2017" box="[1070,1332,905,929]" class="Insecta" family="Staphylinidae" genus="Amaurodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="97" phylum="Arthropoda" rank="species" species="longisetosa">
<emphasis id="B2AD1643FFC9397EFBEEEDD0FAFBDB9B" box="[1070,1332,905,929]" italics="true" pageId="34" pageNumber="97">Amaurodera longisetosa</emphasis>
</taxonomicName>
is characterized particularly by the variability of the size and shape of the posteriorly truncate head, the chaetotaxy of the pronotum, and the primary sexual characters. Using the key to the
<taxonomicName id="47D9B1D2FFC9397EFBECEA50FACADC1B" box="[1068,1285,1033,1057]" class="Insecta" family="Staphylinidae" genus="Amaurodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="97" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B2AD1643FFC9397EFBECEA50FB7FDC1B" box="[1068,1200,1033,1057]" italics="true" pageId="34" pageNumber="97">Amaurodera</emphasis>
species
</taxonomicName>
of Borneo in
<bibRefCitation id="E448B7A0FFC9397EFC80EA70FC00DC7B" author="PACE, R." box="[832,975,1065,1089]" pageId="34" pageNumber="97" pagination="107 - 156" refId="ref34350" refString="PACE, R. 2008 a: Generi e specie della tribu Lomechusini del Borneo (Coleoptera, Staphylinidae). - Revue Suisse de Zoologie 115 (1): 107 - 156." type="journal article" year="2008">PACE (2008a)</bibRefCitation>
, it would key out at couplet 4, together with
<taxonomicName id="47D9B1D2FFC9397EFC80EA10FB0BDC5B" authority="CAMERON, 1928" authorityName="CAMERON" authorityYear="1928" box="[832,1220,1097,1121]" class="Insecta" family="Staphylinidae" genus="Amaurodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="97" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="B2AD1643FFC9397EFC80EA10FC6FDC5B" box="[832,928,1097,1121]" italics="true" pageId="34" pageNumber="97">A. similis</emphasis>
CAMERON, 1928 (
<collectingCountry id="F8CE8AC1FFC9397EFB9FEA10FB71DC5B" box="[1119,1214,1097,1121]" name="Malaysia" pageId="34" pageNumber="97">Sarawak</collectingCountry>
)
</taxonomicName>
, from which it additionally differs by larger body size and darker coloration. Regarding the presence of long setae in the antero-lateral portions of the pronotum and the long proximal portion of the spermathecal capsule,
<taxonomicName id="47D9B1D2FFC9397EFBBFEA90FAC1DCDB" authorityName="Assing" authorityYear="2017" box="[1151,1294,1225,1249]" class="Insecta" family="Staphylinidae" genus="Amaurodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="97" phylum="Arthropoda" rank="species" species="longisetosa">
<emphasis id="B2AD1643FFC9397EFBBFEA90FAC1DCDB" box="[1151,1294,1225,1249]" italics="true" pageId="34" pageNumber="97">A. longisetosa</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="47D9B1D2FFC9397EFC80EAB0FC79DD3B" authorityName="PACE" authorityYear="2008" box="[832,950,1257,1281]" class="Insecta" family="Staphylinidae" genus="Amaurodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="97" phylum="Arthropoda" rank="species" species="amabilis">
<emphasis id="B2AD1643FFC9397EFC80EAB0FC79DD3B" box="[832,950,1257,1281]" italics="true" pageId="34" pageNumber="97">A. amabilis</emphasis>
</taxonomicName>
, from which it is distinguished by much longer and broader elytra, a posteriorly nearly truncate head, slightly larger and more bulging eyes, and a darker abdomen.
</paragraph>
</subSubSection>
<subSubSection id="C8C399DAFFC9397EFC80EBD0FB21DE3B" pageId="34" pageNumber="97" type="distribution">
<paragraph id="8066CA51FFC9397EFC80EBD0FB21DE3B" blockId="34.[832,1437,1417,1538]" pageId="34" pageNumber="97">
<emphasis id="B2AD1643FFC9397EFC80EBD0FB60DD9B" bold="true" box="[832,1199,1417,1441]" pageId="34" pageNumber="97">Distribution and natural history</emphasis>
: The type specimens were collected in four close localities in
<collectingRegion id="421D04B3FFC9397EFB23EBF0FAECDDFB" box="[1251,1315,1449,1473]" country="Malaysia" name="Sabah" pageId="34" pageNumber="97">Sabah</collectingRegion>
at altitudes of
<quantity id="472167B4FFC9397EFC9AEB90FC10DDDB" box="[858,991,1481,1505]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="13.5" metricValueMin="5.5" pageId="34" pageNumber="97" unit="m" value="950.0" valueMax="1350.0" valueMin="550.0">550–1350 m</quantity>
, in one locality together with
<taxonomicName id="47D9B1D2FFC9397EFAD6EB90FA59DDDB" authorityName="PACE" authorityYear="2008" box="[1302,1430,1481,1505]" class="Insecta" family="Staphylinidae" genus="Amaurodera" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="97" phylum="Arthropoda" rank="species" species="frondium">
<emphasis id="B2AD1643FFC9397EFAD6EB90FA59DDDB" box="[1302,1430,1481,1505]" italics="true" pageId="34" pageNumber="97">A. frondium</emphasis>
</taxonomicName>
. One
<typeStatus id="5F6274F3FFC9397EFCB3EBB3FC1FDE38" box="[883,976,1514,1538]" pageId="34" pageNumber="97" type="paratype">paratype</typeStatus>
was collected on the wing.
</paragraph>
</subSubSection>
</treatment>
</document>
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