treatments-xml/data/BA/03/15/BA0315B01ED05A1DB3B02229F76A78EE.xml

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<mods:title id="E54A99DAF0A61D260FD2EE33969784E4">A new, widespread genus of Baetidae from South Asia (Insecta, Ephemeroptera)</mods:title>
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<mods:namePart id="5BC4A4CA83B58C07226DCC83554DDFED">Kaltenbach, Thomas</mods:namePart>
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<mods:affiliation id="74130F469095D877FC981794B711E120">Museum cantonal des Sciences Naturelles, Departement de zoologie, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
<mods:nameIdentifier id="77E0EA34F06F0D322AC73B1349D5A3EB" type="email">thomas.kaltenbach@bluewin.ch</mods:nameIdentifier>
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<mods:namePart id="9C6F73FFABE5524CD6777E3F312F492F">Kluge, Nikita J.</mods:namePart>
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<mods:affiliation id="C0A9D7968A3CACE42C90E04D46E6B52E">Department of Entomology, Biological Faculty, Saint-Petersburg State University, Universitetskaya nab., 7 / 9, Saint Petersburg, 199034, Russia</mods:affiliation>
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<mods:namePart id="21451045BC61848809A0A8B98CBF7F82">Gattolliat, Jean-Luc</mods:namePart>
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<mods:affiliation id="2314FBD268B92114E7558D12D0244891">Museum cantonal des Sciences Naturelles, Departement de zoologie, Palais de Rumine, Place Riponne 6, CH- 1005 Lausanne, Switzerland &amp; University of Lausanne (UNIL), Department of Ecology and Evolution, CH- 1015 Lausanne, Switzerland</mods:affiliation>
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<mods:date id="052CFE879DC3297E82F8EAA9BF8F278E">2023</mods:date>
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<paragraph id="12216206845D051A1511B2CB415D76FD" pageId="0" pageNumber="231">
<taxonomicName id="2B02AFDCF6C0CDE36A2CE176324F5FF3" LSID="https://zoobank.org/F4E4945E-62ED-4E71-80A0-B993DA84EB57" authority="Kaltenbach &amp; Kluge &amp; Gattolliat, 2023" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Arcobaetis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="genus" status="gen. nov.">Arcobaetis</taxonomicName>
<taxonomicNameLabel id="109C8424215A25ED8A2976AE53B2D286" pageId="0" pageNumber="231">gen. nov.</taxonomicNameLabel>
</paragraph>
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<figureCitation id="D1A4406941DD307BA844D6495A023369" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Arcobaetis sumbawensis sp. nov., larva, habitus (holotype) a dorsal view b lateral view c ventral view. Scale bars: 1 mm" figureDoi="10.3897/zookeys.1168.104844.figure1" httpUri="https://binary.pensoft.net/fig/872203" pageId="0" pageNumber="231">Figs 1</figureCitation>
<figureCitation id="CA283BDBC30B23FEE7D6CEC3BC0F23AC" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">, 2</figureCitation>
<figureCitation id="0685EB67ECF8DBD3B980C7E114A7C59B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct." figureDoi="10.3897/zookeys.1168.104844.figure3" httpUri="https://binary.pensoft.net/fig/872205" pageId="0" pageNumber="231">, 3</figureCitation>
<figureCitation id="F7F149AFEB5F3D40C6FC0817A217FCCE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Arcobaetis sumbawensis sp. nov., larva a foreleg b middle leg c hind leg d subimaginal gonostylus developing under cuticle of last instar male larva e head. Scale bars: 0.05 mm (e); 0.1 mm (d)." figureDoi="10.3897/zookeys.1168.104844.figure4" httpUri="https://binary.pensoft.net/fig/872206" pageId="0" pageNumber="231">, 4</figureCitation>
<figureCitation id="931119DA214306AD984F8D43BF9351A0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Arcobaetis sumbawensis sp. nov., larva, SEM pictures a fore claw, lateroventral view b middle claw, anterior view c middle claw, posterior view. Arrows: subapical setae. Scale bars: 0.01 mm." figureDoi="10.3897/zookeys.1168.104844.figure5" httpUri="https://binary.pensoft.net/fig/872207" pageId="0" pageNumber="231">, 5</figureCitation>
<figureCitation id="BCD784FE30BA271AE1CA2589A00FE75B" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Arcobaetis sumatrensis sp. nov., larva, habitus (holotype) a dorsal view b lateral view c ventral view. Scale bar: 1 mm" figureDoi="10.3897/zookeys.1168.104844.figure6" httpUri="https://binary.pensoft.net/fig/872208" pageId="0" pageNumber="231">, 6</figureCitation>
<figureCitation id="D4097647691C2E1C954378BA6AFA536E" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Arcobaetis sumatrensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure7" httpUri="https://binary.pensoft.net/fig/872209" pageId="0" pageNumber="231">, 7</figureCitation>
<figureCitation id="DCAB5837310B48A81CE33F30E676759C" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Arcobaetis sumatrensis sp. nov., larva a foreleg b fore claw c tergalius II d tergalius III e tergalius IV f tergalius V g tergalius VI h tergalius VII i tergum IV j scale on tergum IV k paraproct." figureDoi="10.3897/zookeys.1168.104844.figure8" httpUri="https://binary.pensoft.net/fig/872210" pageId="0" pageNumber="231">, 8</figureCitation>
<figureCitation id="86AC1482702BB053DE08D63CECE1B665" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Arcobaetis sumatrensis sp. nov., larva a foreleg b middle leg c hind leg d tergalius IV e head f fragment of cercus (lateral view). Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1168.104844.figure9" httpUri="https://binary.pensoft.net/fig/872211" pageId="0" pageNumber="231">, 9</figureCitation>
<figureCitation id="459B1F4BF278CBE3A3267D9A05EFBE1B" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Arcobaetis bornensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h l abium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure10" httpUri="https://binary.pensoft.net/fig/872212" pageId="0" pageNumber="231">, 10</figureCitation>
<figureCitation id="F34CE5DD3D898138D39ECC979D95CC66" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Arcobaetis bornensis sp. nov., larva a foreleg b fore claw c tergum IV d tergalius IV e paraproct f antenna (right: dorsal view; left: section in lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure11" httpUri="https://binary.pensoft.net/fig/872213" pageId="0" pageNumber="231">, 11</figureCitation>
<figureCitation id="039F30165639620270F258C97D855525" captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Arcobaetis bornensis sp. nov., larva a foreleg b hind leg c habitat (photo Kate Baker, University Exeter, UK)." figureDoi="10.3897/zookeys.1168.104844.figure12" httpUri="https://binary.pensoft.net/fig/872214" pageId="0" pageNumber="231">, 12</figureCitation>
<figureCitation id="23E4E31683866DCAA8DD1B8F60066CF8" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Arcobaetis sripadai sp. nov., larval exuviae (with same magnification): a-c fore, middle and hind legs (triangles show points where patella-tibial suture crosses inner margin) d half of pronotum and mesonotum e sternum and tergum V f sterna and terga VII-VIII g sterna and terga IX-X." figureDoi="10.3897/zookeys.1168.104844.figure13" httpUri="https://binary.pensoft.net/fig/872215" pageId="0" pageNumber="231">, 13</figureCitation>
<figureCitation id="3C791B0631911D2B6C25DEB2AD5ADCD5" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Arcobaetis sripadai sp. nov., larval exuviae (with same magnification) a antenna b labrum (dorsal side) c right mandible d left mandible e hypopharynx and superlinguae f maxilla g labium (dorsal focus) h labium (ventral focus) i labial palp." figureDoi="10.3897/zookeys.1168.104844.figure14" httpUri="https://binary.pensoft.net/fig/872216" pageId="0" pageNumber="231">, 14</figureCitation>
<figureCitation id="2FB6C0EDB61C4B63E239DBEDB709697E" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Arcobaetis sripadai sp. nov., larval exuviae a, b fore and middle legs (triangles show points where patella-tibial suture crosses inner margin c claw d-k posterior margins of abdominal terga II-VI and VIII-X l-m posterior margin of abdominal sterna VIII and IX of male larva n paraproct. o-p fragments of cerci (lateral view)." figureDoi="10.3897/zookeys.1168.104844.figure15" httpUri="https://binary.pensoft.net/fig/872217" pageId="0" pageNumber="231">, 15</figureCitation>
<figureCitation id="5D2F98EFDDF05C7BC1E6C6220EB9F8B7" captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. Arcobaetis sripadai sp. nov., subimaginal exuviae a hind tarsus b apex of fore tarsus c wing membrane d fragment of mesonotum e fragment of abdominal tergum VII with two right submedian sigilla." figureDoi="10.3897/zookeys.1168.104844.figure16" httpUri="https://binary.pensoft.net/fig/872218" pageId="0" pageNumber="231">, 16</figureCitation>
<figureCitation id="E4E84478D8FDAFC58FC8A688E876C38A" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Arcobaetis sripadai sp. nov. a male imago b head and thorax c apex of abdomen d subimaginal exuviae of meso- and metapleura e subimaginal exuviae of half of mesonotum f, g fore and middle legs h tarsus of middle leg I, j subimaginal exuviae of fore and middle legs k cercus." figureDoi="10.3897/zookeys.1168.104844.figure17" httpUri="https://binary.pensoft.net/fig/872219" pageId="0" pageNumber="231">, 17</figureCitation>
<figureCitation id="E20DDB1751330760832B5320D287B205" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Arcobaetis sripadai sp. nov. a genitalia of male imago b, c imaginal genitalia and their subimaginal exuviae with the same magnification. Abbreviation: m. s asymmetric sterno-styligeral muscle (arrows show its base and apex)" figureDoi="10.3897/zookeys.1168.104844.figure18" httpUri="https://binary.pensoft.net/fig/872220" pageId="0" pageNumber="231">, 18</figureCitation>
<figureCitation id="592B8C27364C0F025A91C7386F35523B" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Arcobaetis gen. nov., distribution." figureDoi="10.3897/zookeys.1168.104844.figure19" httpUri="https://binary.pensoft.net/fig/872221" pageId="0" pageNumber="231">, 19</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="622A32730F149547B90B51C14722996F" pageId="0" pageNumber="231" type="type species">
<paragraph id="2A5EE27D6EF45C30BE1C60CA3D22C3B5" pageId="0" pageNumber="231">Type species.</paragraph>
<paragraph id="C776431984AE9EA069D0AC6CA699337B" pageId="0" pageNumber="231">
<taxonomicName id="DC4138E79F32F43647CD629B2E4F7C7A" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis sripadai" order="Ephemeroptera" pageId="0" pageNumber="231" rank="species" species="sripadai">
<emphasis id="783BEF7995A989FB0746A6DE8DE327AB" italics="true" pageId="0" pageNumber="231">Arcobaetis sripadai</emphasis>
</taxonomicName>
sp. nov., by present designation.
</paragraph>
<paragraph id="03534854E5639A10FA6EB8214670945D" pageId="0" pageNumber="231">
Species included in 
<taxonomicName id="62900651889D261FC8021EF3C0284633" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="genus">
<emphasis id="CF7656AEE6394157764A1C40942DC84B" italics="true" pageId="0" pageNumber="231">Arcobaetis</emphasis>
</taxonomicName>
gen. nov.
</paragraph>
<paragraph id="94F028F81EC7563934E747000C68615A" pageId="0" pageNumber="231">
1. 
<taxonomicName id="2613C1B3E2CC2B28E166BA9747058716" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis sumbawensis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="species" species="sumbawensis">
<emphasis id="048F02D51B414F8BE2E5DC0617EE7ED7" italics="true" pageId="0" pageNumber="231">Arcobaetis sumbawensis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
<paragraph id="B48AEC9C1A4D7C1E72D8932D47E61CCF" pageId="0" pageNumber="231">
2. 
<taxonomicName id="C58D28B36290F7B309252739A862CCD1" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis sumatrensis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="species" species="sumatrensis">
<emphasis id="9DF89D6F7978A6604E78A2D30666C2A9" italics="true" pageId="0" pageNumber="231">Arcobaetis sumatrensis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
<paragraph id="E58D8EFD11C64F7CD3A24F3CD1A929BD" pageId="0" pageNumber="231">
3. 
<taxonomicName id="A4CA253C6881B3572C5B96E4A286F21E" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis bornensis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="species" species="bornensis">
<emphasis id="97CB432873CD0C9BCF68896FFD2E8990" italics="true" pageId="0" pageNumber="231">Arcobaetis bornensis</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
<paragraph id="26B3E1C2AC1234AC84615CC1629585B2" pageId="0" pageNumber="231">
4. 
<taxonomicName id="1F2F7D7D4A3B3D4AE9882DF4CD128ED9" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis sripadai" order="Ephemeroptera" pageId="0" pageNumber="231" rank="species" species="sripadai">
<emphasis id="4B6C3ABCFBD58CF4CC9E8E65AFB27267" italics="true" pageId="0" pageNumber="231">Arcobaetis sripadai</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
<paragraph id="48C6007A82F8D4EB3C7705E1F7194589" pageId="0" pageNumber="231">
5. 
<taxonomicName id="74FCBB63799ED92865DE54514F73F328" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" baseAuthorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" baseAuthorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis gracilentus" order="Ephemeroptera" pageId="0" pageNumber="231" rank="species" species="gracilentus">
<emphasis id="6165261AED36631E5FD06999BFF01CF2" italics="true" pageId="0" pageNumber="231">Arcobaetis gracilentus</emphasis>
</taxonomicName>
(Chang &amp; Yang, in 
<bibRefCitation id="4532056FCF7B465F2417BFA05BB015CF" author="Kang, S-C" journalOrPublisher="Journal of Taiwan Museum" pageId="0" pageNumber="231" pagination="9 - 44" refId="B13" refString="Kang, S-C, Chang, H-C, Yang, C-T, 1994. A revision of the genus Baetis in Taiwan (Ephemeroptera, Baetidae). Journal of Taiwan Museum 47: 9 - 44" title="A revision of the genus Baetis in Taiwan (Ephemeroptera, Baetidae)." volume="47" year="1994">Kang et al. 1994</bibRefCitation>
), comb. nov.
</paragraph>
</subSubSection>
<subSubSection id="733DC9994E888D0FB45C0A198890BC10" pageId="0" pageNumber="231" type="diagnosis">
<paragraph id="BDC72985102ADA19E564B53BD321A435" pageId="0" pageNumber="231">Diagnosis.</paragraph>
<paragraph id="F6E699226125D0FE161247E9F9376357" pageId="0" pageNumber="231">
<emphasis id="ED31C641DB6F9291F8975653DF0CFE47" bold="true" pageId="0" pageNumber="231">Larva.</emphasis>
The following combination of characters differentiate 
<taxonomicName id="66F5DDF0FAB0290D0F36CB374224693A" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="genus">
<emphasis id="76F0D624A09647309A8FD8B9A74E1232" italics="true" pageId="0" pageNumber="231">Arcobaetis</emphasis>
</taxonomicName>
gen. nov. from all other genera of 
<taxonomicName id="3853A84C04B346DF165F8B10130C3C81" authorityName="Leach" authorityYear="1815" class="Insecta" family="Baetidae" kingdom="Animalia" lsidName="" order="Ephemeroptera" pageId="0" pageNumber="231" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
: A) frons with carina-like elevation, slightly overlapping antennal base (Fig. 
<figureCitation id="38A83368253DAC9D81ACBF20A02D0FE5" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Arcobaetis sumbawensis sp. nov., larva a foreleg b middle leg c hind leg d subimaginal gonostylus developing under cuticle of last instar male larva e head. Scale bars: 0.05 mm (e); 0.1 mm (d)." figureDoi="10.3897/zookeys.1168.104844.figure4" httpUri="https://binary.pensoft.net/fig/872206" pageId="0" pageNumber="231">4e</figureCitation>
); B) maxillary palp with two segments (Fig. 
<figureCitation id="D625D729BCD8498DA8A36D41D52F2190" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2g</figureCitation>
); C) both mandibles with outermost denticle in ventral position, set apart from other denticles; both mandibles with long, slightly feathered setae between prostheca and mola (Fig. 
<figureCitation id="755153CE55E0DDB065E5347332EFDF31" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Arcobaetis sumatrensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure7" httpUri="https://binary.pensoft.net/fig/872209" pageId="0" pageNumber="231">7b, d</figureCitation>
); D) paraglossae dorsally with an arc of long, spine-like, simple setae in distal area (Fig. 
<figureCitation id="5549DE80EE98B909E7CDCAEB14FEBFB4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2h, j</figureCitation>
); E) labial palp segment II with poorly developed distomedial protuberance (Fig. 
<figureCitation id="A21618D298E00D38DC74610E5357B072" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2h</figureCitation>
); F) legs very slender; femur length 4-6 
<normalizedToken id="E8CE70256D155331069A8C5D6981FD2E" originalValue="×">x</normalizedToken>
maximum width; outer margin of femora with row of short, spine-like setae (Fig. 
<figureCitation id="06FA406428BCA005D3EDC191C1BB4439" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Arcobaetis sumbawensis sp. nov., larva a foreleg b middle leg c hind leg d subimaginal gonostylus developing under cuticle of last instar male larva e head. Scale bars: 0.05 mm (e); 0.1 mm (d)." figureDoi="10.3897/zookeys.1168.104844.figure4" httpUri="https://binary.pensoft.net/fig/872206" pageId="0" pageNumber="231">4a-c</figureCitation>
); G) claw with single row of denticles, distal denticles larger and directed distad, proximal denticles minute (Figs 
<figureCitation id="B7183A15074A4943689741816D0889CD" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct." figureDoi="10.3897/zookeys.1168.104844.figure3" httpUri="https://binary.pensoft.net/fig/872205" pageId="0" pageNumber="231">3b</figureCitation>
, 
<figureCitation id="35ED2A50BDD971A25C068069646E3F29" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Arcobaetis sumbawensis sp. nov., larva, SEM pictures a fore claw, lateroventral view b middle claw, anterior view c middle claw, posterior view. Arrows: subapical setae. Scale bars: 0.01 mm." figureDoi="10.3897/zookeys.1168.104844.figure5" httpUri="https://binary.pensoft.net/fig/872207" pageId="0" pageNumber="231">5a, b</figureCitation>
); H) surface of abdominal terga with fine, longitudinally striated scales situated in angulate nests, whose angles bear opercula (Fig. 
<figureCitation id="25433344C3242155576013FCF6A6C3E3" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct." figureDoi="10.3897/zookeys.1168.104844.figure3" httpUri="https://binary.pensoft.net/fig/872205" pageId="0" pageNumber="231">3c, d</figureCitation>
); H) folding of subimaginal gonostyli developing under cuticle of last instar male larva of the &quot; 
<taxonomicName id="4A4303031EC6454A536E35CF7CA305CF" authorityName="Novikova &amp; Kluge" authorityYear="1987" class="Insecta" family="Baetidae" genus="Nigrobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nigrobaetis" order="Ephemeroptera" pageId="0" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="A29E810992F86E8E46417DE96D884288" italics="true" pageId="0" pageNumber="231">Nigrobaetis</emphasis>
</taxonomicName>
-type&quot;.
</paragraph>
<caption id="6387E992CDF1100DD8DA20ACA3D35373" doi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231" start="Figure 2" startId="F2">
<paragraph id="0CFBC2118A55D987A575A867306877B5" pageId="0" pageNumber="231">
<emphasis id="CB87879D18657D83DA7F02B2DCE8F8E6" bold="true" pageId="0" pageNumber="231">Figure 2.</emphasis>
<taxonomicName id="03BB4FD2AABD419EC8FCD4026195669A" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis sumbawensis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="species" species="sumbawensis">
<emphasis id="5773F78E02DFEF100FACB76F5424441D" italics="true" pageId="0" pageNumber="231">Arcobaetis sumbawensis</emphasis>
</taxonomicName>
sp. nov., larva 
<emphasis id="CB810260A5FC4315C6C758FDA46240A2" bold="true" pageId="0" pageNumber="231">a</emphasis>
labrum (left: ventral view; right: dorsal view) 
<emphasis id="0BA9B06EDCEC16B85898C657AE313636" bold="true" pageId="0" pageNumber="231">b</emphasis>
right mandible 
<emphasis id="41357E4B62C1F7CA35CCEC129C9232D7" bold="true" pageId="0" pageNumber="231">c</emphasis>
right prostheca 
<emphasis id="896CD7C62527BC5AA59DB2269B947093" bold="true" pageId="0" pageNumber="231">d</emphasis>
left mandible 
<emphasis id="7330F15C722A40AD7424D5F3CC05F313" bold="true" pageId="0" pageNumber="231">e</emphasis>
left prostheca 
<emphasis id="5786CC2844F57263B5545A78DEB199D7" bold="true" pageId="0" pageNumber="231">f</emphasis>
hypopharynx and superlinguae 
<emphasis id="A2AFA268F306E721759D67548620AF05" bold="true" pageId="0" pageNumber="231">g</emphasis>
maxilla 
<emphasis id="2808CFDCEC80CC1AA3865484D77CFBB4" bold="true" pageId="0" pageNumber="231">h</emphasis>
labium (left: ventral view; right: dorsal view) 
<emphasis id="EF1AA888503D8D13D5C3B4016577CFF9" bold="true" pageId="0" pageNumber="231">i</emphasis>
apex of paraglossa (ventral view) 
<emphasis id="849AB7A6AAB10AB6B1B565A59A5B3A30" bold="true" pageId="0" pageNumber="231">j</emphasis>
apex of paraglossa (dorsal view).
</paragraph>
</caption>
<caption id="0C97ACC08EF04A841F828C64B3D57A72" doi="10.3897/zookeys.1168.104844.figure3" httpUri="https://binary.pensoft.net/fig/872205" pageId="0" pageNumber="231" start="Figure 3" startId="F3">
<paragraph id="BE3C8FD554E399589E44251098A8DE31" pageId="0" pageNumber="231">
<emphasis id="EA84E5DD9FE934E2AF963A0A65242E55" bold="true" pageId="0" pageNumber="231">Figure 3.</emphasis>
<taxonomicName id="404802C1A87035BFE4E91DFF20D305F5" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis sumbawensis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="species" species="sumbawensis">
<emphasis id="B5AF98FDFB06876429D1A2F77AA80211" italics="true" pageId="0" pageNumber="231">Arcobaetis sumbawensis</emphasis>
</taxonomicName>
sp. nov., larva 
<emphasis id="0EFAD9B9921F6BE00C5CBC0C77A45023" bold="true" pageId="0" pageNumber="231">a</emphasis>
foreleg 
<emphasis id="F3D02E5DDE044C300C410B0E71E1DAFC" bold="true" pageId="0" pageNumber="231">b</emphasis>
fore claw 
<emphasis id="B71DC54A33A1DBFD9C03FC344E963A36" bold="true" pageId="0" pageNumber="231">c</emphasis>
tergum IV 
<emphasis id="D15DAA68D64FB83DCF3D3D3520BD27AC" bold="true" pageId="0" pageNumber="231">d</emphasis>
scale on tergum IV 
<emphasis id="131625E7CEB9166F54BAEEAC2D568DA1" bold="true" pageId="0" pageNumber="231">e</emphasis>
tergalius I 
<emphasis id="0B9AD328AFBEDD67CEDD564CB54C74F9" bold="true" pageId="0" pageNumber="231">f</emphasis>
paraproct.
</paragraph>
</caption>
<caption id="42E2C96D07E62A9AD70A2FEBFFC1F64A" doi="10.3897/zookeys.1168.104844.figure4" httpUri="https://binary.pensoft.net/fig/872206" pageId="0" pageNumber="231" start="Figure 4" startId="F4">
<paragraph id="1FC042106869569E3C1816642AA1129F" pageId="0" pageNumber="231">
<emphasis id="60ECB8236E5E54477614CF797A1CAA12" bold="true" pageId="0" pageNumber="231">Figure 4.</emphasis>
<taxonomicName id="AF965DF5B5D7D7438F83AFE43F25A4E5" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis sumbawensis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="species" species="sumbawensis">
<emphasis id="EC62423784049861DA81C8CF3365C7A0" italics="true" pageId="0" pageNumber="231">Arcobaetis sumbawensis</emphasis>
</taxonomicName>
sp. nov., larva 
<emphasis id="F80DCD7EB906CCAFDC7175A394F820A5" bold="true" pageId="0" pageNumber="231">a</emphasis>
foreleg 
<emphasis id="0B5B1769B75A0C2620033309D4ADB5EE" bold="true" pageId="0" pageNumber="231">b</emphasis>
middle leg 
<emphasis id="DE15EA19CA52D4E560F7E4791954961F" bold="true" pageId="0" pageNumber="231">c</emphasis>
hind leg 
<emphasis id="FDC445214F8374EA009EFB4FC36206C8" bold="true" pageId="0" pageNumber="231">d</emphasis>
subimaginal gonostylus developing under cuticle of last instar male larva 
<emphasis id="D9DE251DD93E91D51EC661B09DF12AB1" bold="true" pageId="0" pageNumber="231">e</emphasis>
head. Scale bars: 0.05 mm (
<emphasis id="9F5FB25F052FD895F129EE84CF73F81E" bold="true" pageId="0" pageNumber="231">e</emphasis>
); 0.1 mm (
<emphasis id="1F925493B57D78F2B0DC9F5D46A124A7" bold="true" pageId="0" pageNumber="231">d</emphasis>
).
</paragraph>
</caption>
<caption id="D3A421A838F4A3A336BD68A8E0FBF55F" doi="10.3897/zookeys.1168.104844.figure5" httpUri="https://binary.pensoft.net/fig/872207" pageId="0" pageNumber="231" start="Figure 5" startId="F5">
<paragraph id="13C0602D227477B698928FC737D0A523" pageId="0" pageNumber="231">
<emphasis id="A0F2C1D9402DA287E6E7ED9475405891" bold="true" pageId="0" pageNumber="231">Figure 5.</emphasis>
<taxonomicName id="244DB4EFB2D488FE5431B5A0DEED1913" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis sumbawensis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="species" species="sumbawensis">
<emphasis id="FE47DDF3CF661D069DD26145C295A8D3" italics="true" pageId="0" pageNumber="231">Arcobaetis sumbawensis</emphasis>
</taxonomicName>
sp. nov., larva, SEM pictures 
<emphasis id="A3C6BC38530D2B7ACCD941B47279328C" bold="true" pageId="0" pageNumber="231">a</emphasis>
fore claw, lateroventral view 
<emphasis id="C02742517B15AD27B68312A655928084" bold="true" pageId="0" pageNumber="231">b</emphasis>
middle claw, anterior view 
<emphasis id="DD3DEED5890AEF0D413970796E765696" bold="true" pageId="0" pageNumber="231">c</emphasis>
middle claw, posterior view. Arrows: subapical setae. Scale bars: 0.01 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="9EC0A9EE317578B9F050AE101336A09B" pageId="0" pageNumber="231" type="description">
<paragraph id="1D8ACC6BA26499B4D4455E1EF4CB287C" pageId="0" pageNumber="231">Description.</paragraph>
<paragraph id="FD06AF08FB04092481C094767E402F08" pageId="0" pageNumber="231">
<emphasis id="26374D7F5A907AEBE4DABCD5D6AC92D2" bold="true" pageId="0" pageNumber="231">
Larva. 
<emphasis id="CDA93FB33C293EA476F0E424D1B44509" italics="true" pageId="0" pageNumber="231">Head</emphasis>
</emphasis>
(Fig. 
<figureCitation id="1DAE8A7143516E02159144512C621189" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Arcobaetis sumbawensis sp. nov., larva a foreleg b middle leg c hind leg d subimaginal gonostylus developing under cuticle of last instar male larva e head. Scale bars: 0.05 mm (e); 0.1 mm (d)." figureDoi="10.3897/zookeys.1168.104844.figure4" httpUri="https://binary.pensoft.net/fig/872206" pageId="0" pageNumber="231">4e</figureCitation>
). Frons with carina-like elevation, slightly overlapping antennal base.
</paragraph>
<paragraph id="DA94B099930B898A3961FE2C8E7C6C96" pageId="0" pageNumber="231">
<emphasis id="9AD5FCB947E1A3AC93FC2FF2EAC20E08" bold="true" italics="true" pageId="0" pageNumber="231">Labrum</emphasis>
(Fig. 
<figureCitation id="FC9AB5CC85B33A71E82BD9E0048AA34E" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2a</figureCitation>
). Sub-rectangular, wider than long. Distal margin with medial, shallow emargination and small process. Dorsally with pair of long, simple, sub-median setae, and on each side partial sub-marginal arc of long, stout, simple setae; sometimes with setae between sub-median seta and partial sub-marginal arc; surface scattered with medium to long, simple setae. Ventrally with long long-feathered setae on anterolateral margin and medium short-feathered setae on medial margin; several small, stout setae near anterolateral and sometimes also midlateral margin.
</paragraph>
<paragraph id="0860BF6628731EB11532930B75351940" pageId="0" pageNumber="231">
<emphasis id="169A60B52BE802DFE56B981DD89EF72D" bold="true" italics="true" pageId="0" pageNumber="231">Right mandible</emphasis>
(Fig. 
<figureCitation id="7BB7236B525E0CA8954F408CEE2C9597" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2b, c</figureCitation>
). Incisor and kinetodontium fused; incisor with denticles, outer denticle turned ventrally and much lower than other denticles; kinetodontium with denticles; inner margin of innermost denticle of kinetodontium with row of thin setae; prostheca stick-like, distolaterally denticulate; margin between prostheca and mola straight, with long, slightly feathered setae; apex of mola with tuft of setae.
</paragraph>
<paragraph id="8CCC97B712A60DC7E5675E261A25B5EE" pageId="0" pageNumber="231">
<emphasis id="EFC2B4D2F966D97C3601F8E83FB37B02" bold="true" italics="true" pageId="0" pageNumber="231">Left mandible</emphasis>
(Fig. 
<figureCitation id="C9203C74A32252C6A4CF37B095EDBAD9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2d, e</figureCitation>
). Incisor and kinetodontium fused; incisor with denticles, outer denticle turned ventrally and set apart from other denticles; kinetodontium with denticles; prostheca robust, distally denticulate; margin between prostheca and mola straight, with row of long, slightly feathered setae; apex of mola without tuft of setae.
</paragraph>
<paragraph id="CD0307C3D171D6EC67A9970DF6355219" pageId="0" pageNumber="231">Both mandibles with outer lateral margins almost straight.</paragraph>
<paragraph id="4B75D0EC377BDF3ACE001B0FB3A1EF0E" pageId="0" pageNumber="231">
<emphasis id="C1B926EC52CD5EBC73C06C54FB082D09" bold="true" italics="true" pageId="0" pageNumber="231">Hypopharynx and superlinguae</emphasis>
(Fig. 
<figureCitation id="75E88ABC0C4630528D2CA37484DBDE85" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2f</figureCitation>
). Apex of lingua with poorly developed tuft of setae. Distolateral margin of superlinguae with fine setae.
</paragraph>
<paragraph id="72F95BA2E720C6EE175F596A37D8019A" pageId="0" pageNumber="231">
<emphasis id="DD2519D49B78ABAB9AC9DB81F259621F" bold="true" italics="true" pageId="0" pageNumber="231">Maxilla</emphasis>
(Fig. 
<figureCitation id="C60A30D85A7D964D31BE35DE0E7F64F9" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2g</figureCitation>
). Apically with three slender, pointed canines and three denti-setae; distal denti-seta tooth-like, bent in the same direction as canines; other denti-setae slender, bifid, and pectinate; maxillary palp with two segments, apex rounded.
</paragraph>
<paragraph id="A9E3E9A451B0F7E2651C2AD134396EAB" pageId="0" pageNumber="231">
<emphasis id="D8D75AECF63262EFB6603DC590CCAF0A" bold="true" italics="true" pageId="0" pageNumber="231">Labium</emphasis>
(Fig. 
<figureCitation id="B894545A5497F074C3DD09484BDBA4FA" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Arcobaetis sumbawensis sp. nov., larva a labrum (left: ventral view; right: dorsal view) b right mandible c right prostheca d left mandible e left prostheca f hypopharynx and superlinguae g maxilla h labium (left: ventral view; right: dorsal view) i apex of paraglossa (ventral view) j apex of paraglossa (dorsal view)." figureDoi="10.3897/zookeys.1168.104844.figure2" httpUri="https://binary.pensoft.net/fig/872204" pageId="0" pageNumber="231">2h</figureCitation>
). Glossae basally broad, narrowing towards apex, shorter than paraglossae; inner margin with row of spine-like setae, increasing in length distally; apex with several medium to long, robust setae; outer margin with row of spine-like setae; ventral surface with numerous medium to long fine, simple scattered setae. Paraglossae laterally straight, distally slightly bent inwards; apex with three rows of long, robust, distally pectinate setae; ventrally usually with several short, simple setae in distomedial area; dorsally with distal arc of long, spine-like setae. Labial palp with three segments, segment II with slightly developed distomedial protuberance.
</paragraph>
<paragraph id="054A89F64068FC30CB90E90662B9B9F8" pageId="0" pageNumber="231">
<emphasis id="EFB54C721C5A3D3913ADD8C76B582F86" bold="true" pageId="0" pageNumber="231">
<emphasis id="A74C4D109A163012482B8B0ED54962B0" italics="true" pageId="0" pageNumber="231">Thorax</emphasis>
. 
<emphasis id="91439B9D6E81CBBA5DC4B9D262609587" italics="true" pageId="0" pageNumber="231">Legs</emphasis>
</emphasis>
(Figs 
<figureCitation id="C3F93F421CE7241A89BF54A436036DE9" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct." figureDoi="10.3897/zookeys.1168.104844.figure3" httpUri="https://binary.pensoft.net/fig/872205" pageId="0" pageNumber="231">3a, b</figureCitation>
, 
<figureCitation id="B78C62A2A33BB584EA84F2896B3E1581" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Arcobaetis sumbawensis sp. nov., larva a foreleg b middle leg c hind leg d subimaginal gonostylus developing under cuticle of last instar male larva e head. Scale bars: 0.05 mm (e); 0.1 mm (d)." figureDoi="10.3897/zookeys.1168.104844.figure4" httpUri="https://binary.pensoft.net/fig/872206" pageId="0" pageNumber="231">4a-c</figureCitation>
). Long and very slender. 
<emphasis id="8CF1FCD9734D86E6A78B3FB157682843" bold="true" italics="true" pageId="0" pageNumber="231">Femur.</emphasis>
Length approx. 4-6 
<normalizedToken id="1B575E5B397FBAF51DD89C0381F89CE9" originalValue="×">x</normalizedToken>
maximum width; outer margin with row of curved, spine-like setae; apex rounded, with pair of curved, spine-like setae; ventrally with stout, pointed, pectinate setae along margin; femoral patch absent. 
<emphasis id="A7A2E61E7AB1DE6783AD560339195F8E" bold="true" italics="true" pageId="0" pageNumber="231">Tibia.</emphasis>
Outer margin of fore legs almost bare, with one curved, spine-like seta at apex; middle and hind legs additionally with row of few spine-like setae. 
<emphasis id="0D22D97A7E77758C66E9C3A16F6E902E" bold="true" italics="true" pageId="0" pageNumber="231">Tarsus.</emphasis>
Dorsal margin almost bare. 
<emphasis id="CF0B085589B245BC4DAB7B25E7F87799" bold="true" italics="true" pageId="0" pageNumber="231">Claw.</emphasis>
Slender, with long point, and with one row of denticles. Distal denticles larger and directed distad, proximal denticles are minute.
</paragraph>
<paragraph id="0A58FBA5C0747928BF4FD0FC0BE178E2" pageId="0" pageNumber="231">
<emphasis id="7CB9D3BCC107A633DB33367101EA20B0" bold="true" pageId="0" pageNumber="231">
Abdomen. 
<emphasis id="91481D34F1C162F348AFB2A28136805B" italics="true" pageId="0" pageNumber="231">Terga</emphasis>
</emphasis>
(Fig. 
<figureCitation id="E3E8089CC5D85F0033554BFBDE27ADB7" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct." figureDoi="10.3897/zookeys.1168.104844.figure3" httpUri="https://binary.pensoft.net/fig/872205" pageId="0" pageNumber="231">3c, d</figureCitation>
). Surface with irregular rows of scale bases and with fine, longitudinally striated scales situated in angulate nests, whose angles bear opercula (see also 
<bibRefCitation id="154B5A94D32F0A25F2F25ECC0C11534C" DOI="https://doi.org/10.15298/rusentj.20.4.02" author="Kluge, NJ" journalOrPublisher="Russian Entomological Journal" pageId="0" pageNumber="231" pagination="361 - 376" refId="B17" refString="Kluge, NJ, 2012. Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae). Russian Entomological Journal 20 (4): 361 - 376, DOI: https://doi.org/10.15298/rusentj.20.4.02" title="Non-African representatives of the plesiomorphon Protopatellata (Ephemeroptera: Baetidae)." url="https://doi.org/10.15298/rusentj.20.4.02" volume="20" year="2012">Kluge 2012</bibRefCitation>
: 365, fig. 6).
</paragraph>
<paragraph id="98913AD4D4CC7C152AFE02B76B00A32E" pageId="0" pageNumber="231">
<emphasis id="D9E7B6D2971EA47B4FA2C38D50A978BB" bold="true" italics="true" pageId="0" pageNumber="231">Tergalii.</emphasis>
Present on segments I-VII or II-VII.
</paragraph>
<paragraph id="39ABDFCECE9F9E08F302E76A7742284B" pageId="0" pageNumber="231">
<emphasis id="D5B1D03DF447F3B6696C770C9E7143AA" bold="true" italics="true" pageId="0" pageNumber="231">Paraproct</emphasis>
(Fig. 
<figureCitation id="C61884A861A05E99FF7DBB8ECB0A1AE0" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Arcobaetis sumbawensis sp. nov., larva a foreleg b fore claw c tergum IV d scale on tergum IV e tergalius I f paraproct." figureDoi="10.3897/zookeys.1168.104844.figure3" httpUri="https://binary.pensoft.net/fig/872205" pageId="0" pageNumber="231">3f</figureCitation>
). Posterior margin with stout spines. Cercotractor with numerous, small, marginal spines.
</paragraph>
<paragraph id="0BCD27F5DACB9E5456812BEEECC3E32D" pageId="0" pageNumber="231">
<emphasis id="DE0905B29751D64A7DB0945279F80235" bold="true" italics="true" pageId="0" pageNumber="231">Larval protogonostyli</emphasis>
(Fig. 
<figureCitation id="BC91CEEBE70830EC9A8B11E397CF16C3" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Arcobaetis sumbawensis sp. nov., larva a foreleg b middle leg c hind leg d subimaginal gonostylus developing under cuticle of last instar male larva e head. Scale bars: 0.05 mm (e); 0.1 mm (d)." figureDoi="10.3897/zookeys.1168.104844.figure4" httpUri="https://binary.pensoft.net/fig/872206" pageId="0" pageNumber="231">4d</figureCitation>
). Subimaginal gonostyli developing under cuticle of last instar male larvae folded in &quot; 
<taxonomicName id="13B9135004FA751B063A553AEAB0AE70" authorityName="Novikova &amp; Kluge" authorityYear="1987" class="Insecta" family="Baetidae" genus="Nigrobaetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nigrobaetis" order="Ephemeroptera" pageId="0" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="775F708CEAC8D45B2210F496CD196127" italics="true" pageId="0" pageNumber="231">Nigrobaetis</emphasis>
</taxonomicName>
-type&quot;: all segments (1st, 2nd, and 3rd) directed caudally and compressed in longitudinal direction, base of 2nd segment deeply inserted into 1st segment, and 3rd segment inserted into 2nd segment (as in 
<bibRefCitation id="15721A69A9A83A7C9639BE6FCE7D8095" author="Novikova, EA" journalOrPublisher="Entomologicheskoe Obozrenie" pageId="0" pageNumber="231" pagination="623 - 644" refId="B26" refString="Novikova, EA, Kluge, NJ, 1994. Mayflies of the subgenus Nigrobaetis (Ephemeroptera, Baetidae, Baetis). Entomologicheskoe Obozrenie 73: 623 - 644" title="Mayflies of the subgenus Nigrobaetis (Ephemeroptera, Baetidae, Baetis)." volume="73" year="1994">Novikova and Kluge 1994</bibRefCitation>
: fig. 2: 1, 5, 20).
</paragraph>
</subSubSection>
<subSubSection id="CF1B32C620BFCA1DE0C34E309C1CC5D0" pageId="0" pageNumber="231" type="diagnoses">
<paragraph id="24FC619975CE8B9BF9CA6EE4B17A9143" pageId="0" pageNumber="231">Diagnoses.</paragraph>
<paragraph id="6F250C6019D018DE2736EB79700E9BB0" pageId="0" pageNumber="231">
<emphasis id="F5D4F6ABE971462AB028E3DC324CAC17" bold="true" pageId="0" pageNumber="231">Imago.</emphasis>
Following combination of characters differentiate 
<taxonomicName id="8123C0837F1436823983AE8F005F7B9B" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="genus">
<emphasis id="9D75EB4E5AECA0208DF4C4871C1D949C" italics="true" pageId="0" pageNumber="231">Arcobaetis</emphasis>
</taxonomicName>
gen. nov. from other genera of 
<taxonomicName id="7AB8AB143C126B8BBBC08EA8DFAE8A4B" authorityName="Leach" authorityYear="1815" class="Insecta" family="Baetidae" kingdom="Animalia" lsidName="" order="Ephemeroptera" pageId="0" pageNumber="231" phylum="Arthropoda" rank="family">Baetidae</taxonomicName>
A) forewing with double intercalary veins (Fig. 
<figureCitation id="2E7EEB7E47157FE793F9744170F0BA1E" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Arcobaetis sripadai sp. nov. a male imago b head and thorax c apex of abdomen d subimaginal exuviae of meso- and metapleura e subimaginal exuviae of half of mesonotum f, g fore and middle legs h tarsus of middle leg I, j subimaginal exuviae of fore and middle legs k cercus." figureDoi="10.3897/zookeys.1168.104844.figure17" httpUri="https://binary.pensoft.net/fig/872219" pageId="0" pageNumber="231">17a</figureCitation>
); B) gonostylus segment III extraordinary small, much narrower than apex of 2nd segment (Fig. 
<figureCitation id="BED2A3A117BA7067A79839B5902A4BA3" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Arcobaetis sripadai sp. nov. a genitalia of male imago b, c imaginal genitalia and their subimaginal exuviae with the same magnification. Abbreviation: m. s asymmetric sterno-styligeral muscle (arrows show its base and apex)" figureDoi="10.3897/zookeys.1168.104844.figure18" httpUri="https://binary.pensoft.net/fig/872220" pageId="0" pageNumber="231">18a</figureCitation>
); C) gonovectes sharply bent (Fig. 
<figureCitation id="EF7DD16655BBD57F8DC72E4B9F2CFFAC" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Arcobaetis sripadai sp. nov. a genitalia of male imago b, c imaginal genitalia and their subimaginal exuviae with the same magnification. Abbreviation: m. s asymmetric sterno-styligeral muscle (arrows show its base and apex)" figureDoi="10.3897/zookeys.1168.104844.figure18" httpUri="https://binary.pensoft.net/fig/872220" pageId="0" pageNumber="231">18a</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="09B82CFA6E5295BD047CFAF12270598F" pageId="0" pageNumber="231" type="description">
<paragraph id="8156DA8102570059B948CA71D3AA6DE9" pageId="0" pageNumber="231">Description.</paragraph>
<paragraph id="9F458015E535A5AED2D9800815810B19" pageId="0" pageNumber="231">
<emphasis id="254FFD4F5ADDDDCC18F1EB66514FDF6B" bold="true" pageId="0" pageNumber="231">Male Imago.</emphasis>
See description of male imago under 
<taxonomicName id="33D8601339E714FDA0209AB15857CA4C" lsidName="A. sripadai" pageId="0" pageNumber="231" rank="species" species="sripadai">
<emphasis id="B77F3CE1E37CD1B09C41313777E4E132" italics="true" pageId="0" pageNumber="231">A. sripadai</emphasis>
</taxonomicName>
sp. nov. below.
</paragraph>
</subSubSection>
<subSubSection id="E4021B2ECDC405454A04D07161DE8BE1" pageId="0" pageNumber="231" type="etymology">
<paragraph id="5F0D94555A952AA2DBDE7F5A8F70B77E" pageId="0" pageNumber="231">Etymology.</paragraph>
<paragraph id="2ED6B70DBC2A21C55020E796B5161AD3" pageId="0" pageNumber="231">
<taxonomicName id="CDD731B695ECC86B80C6E286BCFA2982" authorityName="Kaltenbach &amp; Kluge &amp; Gattolliat" authorityYear="2023" family="Baetidae" genus="Arcobaetis" kingdom="Animalia" lsidName="Arcobaetis" order="Ephemeroptera" pageId="0" pageNumber="231" rank="genus">
<emphasis id="A981DE71942813A7BD920E3ABDD53A93" italics="true" pageId="0" pageNumber="231">Arcobaetis</emphasis>
</taxonomicName>
is a combination of 
<emphasis id="BEC950AD35E623662A1CBCB382DBBEBE" italics="true" pageId="0" pageNumber="231">Arco</emphasis>
-, in reference to the Latin word arcus for arc and the arc of long, simple setae dorsodistally on paraglossae, and 
<emphasis id="18443675A04F7DAA013D806CDA24E0E6" italics="true" pageId="0" pageNumber="231">baetis</emphasis>
, to highlight the similarities with the genus 
<taxonomicName id="5A97E2AAD7EBB02D4A359817317E0123" authorityName="Leach" authorityYear="1815" class="Insecta" family="Baetidae" genus="Baetis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Baetis" order="Ephemeroptera" pageId="0" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="A4784D800BD4E5B1C3B90FD1741CABB0" italics="true" pageId="0" pageNumber="231">Baetis</emphasis>
</taxonomicName>
. The gender is masculine.
</paragraph>
</subSubSection>
<subSubSection id="1EBD83BCAA6F8924E294E05833B5C0D9" pageId="0" pageNumber="231" type="distribution">
<paragraph id="9FFEA193128163234711C48EAA88A239" pageId="0" pageNumber="231">Distribution</paragraph>
<paragraph id="45BC825B885E11FFE77869AF8B44F523" pageId="0" pageNumber="231">
<emphasis id="460624D3BC520398E9C33592FF3678D5" bold="true" pageId="0" pageNumber="231">
(Fig. 
<figureCitation id="CDCFD3A18DA2B760498337D5E0399E81" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Arcobaetis gen. nov., distribution." figureDoi="10.3897/zookeys.1168.104844.figure19" httpUri="https://binary.pensoft.net/fig/872221" pageId="0" pageNumber="231">19</figureCitation>
).
</emphasis>
Indonesia (Sumatra, Sumbawa), Brunei, Sri Lanka, Taiwan.
</paragraph>
</subSubSection>
</treatment>
</document>