137 lines
18 KiB
XML
137 lines
18 KiB
XML
<document id="6062A96909922589D2C52C70E4851289" ID-DOI="10.5281/zenodo.4532981" ID-ISSN="1942-1354" ID-Zenodo-Dep="4532981" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612994740723" checkinUser="felipe" docAuthor="Penz, Carla M." docDate="2008" docId="0F1331235507230CFF4FD89450F57A4B" docLanguage="en" docName="InsectaMundi.2008.0035.1-25.pdf" docOrigin="Insecta Mundi 2008 (35)" docStyle="DocumentStyle:A87B5EC635B7D5A430FC1D805C706BD1.3:InsectaMundi.2007-2008.journal_article" docStyleId="A87B5EC635B7D5A430FC1D805C706BD1" docStyleName="InsectaMundi.2007-2008.journal_article" docStyleVersion="3" docTitle="Eryphanis zolzivora" docType="treatment" docVersion="4" lastPageNumber="14" masterDocId="F32A495B55082303FF8FDA5653137D50" masterDocTitle="Phylogenetic Revision of Eryphanis Boisduval, with a Description of a New Species from Ecuador (Lepidoptera, Nymphalidae)" masterLastPageNumber="25" masterPageNumber="1" pageNumber="14" updateTime="1698870476287" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
|
|
<mods:mods id="64C18631DA4BFFE0C41F734F5AC2E087" xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo id="D58412B18C56CBE21E11FFE65704258F">
|
|
<mods:title id="2FB2CDEB49E8414BA5DA8AA19EEA7257">Phylogenetic Revision of Eryphanis Boisduval, with a Description of a New Species from Ecuador (Lepidoptera, Nymphalidae)</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name id="EA7CEA0F4E762E90809AF3E2656FD891" type="personal">
|
|
<mods:role id="751B6FA029A037642DDF2036E4824E0F">
|
|
<mods:roleTerm id="AA0DA412AA3A722FA802A62A3885BAE1">Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart id="44EFFC69406438E0F2EC833F9281B224">Penz, Carla M.</mods:namePart>
|
|
<mods:affiliation id="F3598A94CB47AEBA7070B0CC25489E26">Department of Biological Sciences University of New Orleans 2000 Lakeshore Drive New Orleans, Louisiana, 70148, USA cpenz @ uno. edu</mods:affiliation>
|
|
</mods:name>
|
|
<mods:typeOfResource id="60A72A7F005B5CEF4C99EBFABF03CCD1">text</mods:typeOfResource>
|
|
<mods:relatedItem id="0BA39232BB48F00C77A58639F5DAB595" type="host">
|
|
<mods:titleInfo id="CC65CE678EB99098EE8D9DD22BC20643">
|
|
<mods:title id="AFEEA36CDFCE89758648B075729A8081">Insecta Mundi</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part id="364DC049AEE7EECB093113E7871C75D5">
|
|
<mods:date id="0F52B06DF205C09C39398330AE58BC72">2008</mods:date>
|
|
<mods:detail id="2919FE690E02944934D398F33EB04B3E" type="pubDate">
|
|
<mods:number id="430308748D3F289ECFC4C1C0029947BC">2008-06-06</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="3ED59DC0ABB1683EAC3AEB9A708C34D4" type="volume">
|
|
<mods:number id="191299DFB14E466F43DCF9FEE8F21DA1">2008</mods:number>
|
|
</mods:detail>
|
|
<mods:detail id="C8DA59BD5D4A92AE9BB8D01AC7F20BC7" type="issue">
|
|
<mods:number id="46D613E6D5B45787113B950F1E73D4ED">35</mods:number>
|
|
</mods:detail>
|
|
<mods:extent id="A1F6D42CA84C999A89A1B06252AF4F67" unit="page">
|
|
<mods:start id="581718E56AAD53CB507CDF04BE2D4A22">1</mods:start>
|
|
<mods:end id="1FCBD63B712D5DF61D46AB879C07386B">25</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:classification id="EFE1F261E50183C20B27866B3FC55E1B">journal article</mods:classification>
|
|
<mods:identifier id="586023A53B827153C955B5A59A376D3C" type="DOI">10.5281/zenodo.4532981</mods:identifier>
|
|
<mods:identifier id="3015779FB1B74404EDB854BA871EF23F" type="ISSN">1942-1354</mods:identifier>
|
|
<mods:identifier id="30EAB106F86F7D5C1EBDA91D3B86A046" type="Zenodo-Dep">4532981</mods:identifier>
|
|
</mods:mods>
|
|
<treatment id="0F1331235507230CFF4FD89450F57A4B" ID-DOI="http://doi.org/10.5281/zenodo.4533097" ID-Zenodo-Dep="4533097" LSID="urn:lsid:plazi:treatment:0F1331235507230CFF4FD89450F57A4B" httpUri="http://treatment.plazi.org/id/0F1331235507230CFF4FD89450F57A4B" lastPageNumber="14" pageId="15" pageNumber="14">
|
|
<subSubSection id="CFA0D3BE5507230CFF4FD89451A77F8B" box="[192,692,706,731]" pageId="15" pageNumber="14" type="nomenclature">
|
|
<paragraph id="870580355507230CFF4FD89451A77F8B" blockId="15.[192,692,706,731]" box="[192,692,706,731]" pageId="15" pageNumber="14">
|
|
<heading id="DC4D37595507230CFF4FD89451A77F8B" bold="true" box="[192,692,706,731]" fontSize="10" level="2" pageId="15" pageNumber="14" reason="4">
|
|
<taxonomicName id="40BAFBB65507230CFF4FD89451A77F8B" authority="(Hewitson, 1877)" baseAuthorityName="Hewitson" baseAuthorityYear="1877" box="[192,692,706,731]" class="Insecta" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="zolzivora">
|
|
<emphasis id="B5CE5C275507230CFF4FD89451A77F8B" bold="true" box="[192,692,706,731]" pageId="15" pageNumber="14">
|
|
<emphasis id="B5CE5C275507230CFF4FD89452DC7F8B" bold="true" box="[192,463,706,731]" italics="true" pageId="15" pageNumber="14">Eryphanis zolzivora</emphasis>
|
|
(Hewitson, 1877)
|
|
</emphasis>
|
|
</taxonomicName>
|
|
</heading>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="CFA0D3BE5507230CFF4FD8B451117FAB" box="[192,514,738,763]" pageId="15" pageNumber="14" type="description">
|
|
<paragraph id="870580355507230CFF4FD8B451117FAB" blockId="15.[192,514,738,763]" box="[192,514,738,763]" pageId="15" pageNumber="14">
|
|
<heading id="DC4D37595507230CFF4FD8B451117FAB" bold="true" box="[192,514,738,763]" fontSize="10" level="2" pageId="15" pageNumber="14" reason="4">
|
|
<emphasis id="B5CE5C275507230CFF4FD8B451117FAB" bold="true" box="[192,514,738,763]" pageId="15" pageNumber="14">
|
|
(
|
|
<figureCitation id="1F819CB05507230CFF46D8B4523F7FAB" box="[201,300,738,763]" captionStart="Figure 1" captionStartId="3.[192,275,1387,1409]" captionTargetBox="[198,1437,201,1373]" captionTargetId="figure-132@3.[192,1443,195,1374]" captionTargetPageId="3" captionText="Figure 1. Eryphanis adult males, dorsal side on the left, ventral side on the right. Scale bar next to A = 1cm. A) E. automedon, Colombia, Cali. B) E. lycomedon, Colombia, Cali. C) E. reevesii, Brazil, São Paulo, Pinhal. D) E. bubocula, Costa Rica, Guanacaste, Rio San Lorenzo. E) E. aesacus, [Mexico], Catemaco. F) E. gerhardi, Bolivia. G) E. zolzivora, Bolivia. H) E. greeneyi, Ecuador, Napo, Yanayacu Biological Station, HOLOTYPE. I) E. opimus, Colombia, Cali." figureDoi="http://doi.org/10.5281/zenodo.4532985" httpUri="https://zenodo.org/record/4532985/files/figure.png" pageId="15" pageNumber="14">Fig. 1G</figureCitation>
|
|
,
|
|
<figureCitation id="1F819CB05507230CFEB6D8B552737FAB" box="[313,352,739,763]" captionStart="Figure 2" captionStartId="4.[192,276,1405,1427]" captionTargetBox="[195,1438,197,1377]" captionTargetId="figure-144@4.[192,1444,194,1380]" captionTargetPageId="4" captionText="Figure 2. Eryphanis adult females, dorsal side on the left, ventral side on the right. Scale bar next to A = 1cm. A) E. automedon, Paraguay, Pedro Juan Caballero. B) E. lycomedon, Colombia, Cali. C) E. reevesii, Brazil, Rio de Janeiro, Nova Friburgo. D) E. bubocula, Costa Rica, Cartago, Tapanti. E) E. aesacus, Mexico, Oaxaca, Tuxtepec.F) E. gerhardi,Brazil, Rondonia, Jaru.G) E.zolzivora, Bolivia, Cochabamba. H) E. greeneyi,Ecuador, Napo, Yanayacu Biological Station, PARATYPE." figureDoi="http://doi.org/10.5281/zenodo.4532987" httpUri="https://zenodo.org/record/4532987/files/figure.png" pageId="15" pageNumber="14">2G</figureCitation>
|
|
,
|
|
<figureCitation id="1F819CB05507230CFEE3D8B552807FAB" box="[364,403,739,763]" captionStart="Figure 4" captionStartId="7.[193,276,1750,1772]" captionTargetBox="[261,1378,190,1718]" captionTargetId="figure-15@7.[257,1382,189,1721]" captionTargetPageId="7" captionText="Figure 4. Eryphanis male genitalia in lateral view, with separate drawings of the phallus. Scale bar next to A = 1mm. Note that the phallus was not removed from the genitalic capsule, and the drawing angles are thus not homogeneous across species. A) E. automedon, Colombia, Cali. B) E. lycomedon, Colombia, Cali. C) E. reevesii, Brazil, São Paulo, Pinhal. D) E. bubocula, Costa Rica, Guanacaste, Rio San Lorenzo. E) E. aesacus, Mexico, San Luis Potosi. F) E. gerhardi, Bolivia, Cochabamba. G) E. zolzivora, Bolivia, Cochabamba. H) E. greeneyi, Ecuador, near Baños, PARATYPE. I) E. opimus, Colombia, Cali." figureDoi="http://doi.org/10.5281/zenodo.4532989" httpUri="https://zenodo.org/record/4532989/files/figure.png" pageId="15" pageNumber="14">4G</figureCitation>
|
|
,
|
|
<figureCitation id="1F819CB05507230CFE10D8B552D57FAB" box="[415,454,739,763]" captionStart="Figure 5" captionStartId="8.[194,277,1776,1798]" captionTargetBox="[259,1375,204,1738]" captionTargetId="figure-21@8.[255,1383,201,1740]" captionTargetPageId="8" captionText="Figure 5. Eryphanis male genitalia in ventral view, with separate drawings of the juxta and dorsal view of tegumen+uncus. Scale bar next to A = 1mm. A) E. automedon, Colombia, Cali. B) E. lycomedon, Colombia, Cali. C) E. reevesii, Brazil, São Paulo, Pinhal. D) E. bubocula, Costa Rica, Guanacaste, Rio San Lorenzo. E) E. aesacus, Mexico, San Luis Potosi. F) E. gerhardi, Bolivia, Cochabamba. G) E. zolzivora, Bolivia, Cochabamba. H) E. greeneyi, Ecuador, near Baños, PARATYPE. I) E. opimus, Colombia, Cali." figureDoi="http://doi.org/10.5281/zenodo.4532991" httpUri="https://zenodo.org/record/4532991/files/figure.png" pageId="15" pageNumber="14">5G</figureCitation>
|
|
,
|
|
<figureCitation id="1F819CB05507230CFE5DD8B452EA7FAB" box="[466,505,738,763]" captionStart="Figure 6" captionStartId="9.[192,276,1236,1258]" captionTargetBox="[200,1439,199,1223]" captionTargetId="figure-197@9.[192,1443,194,1244]" captionTargetPageId="9" captionText="Figure 6. Eryphanis female sterigma and corpus bursa. Ribs of the intersegmental sac were omitted. Scale bars next to F = 1mm, note that sterigma and corpus bursa are in different magnifications. A) E. automedon, Peru, Puerto Maldonado. B) E. lycomedon, Costa Rica, Puntarenas. C) E. reevesii, Brazil, São Paulo, Pinhal. D) E. bubocula, Costa Rica, Cartago, Tapanti. E) E. aesacus,Mexico. F) E. gerhardi, Brazil, Rondonia, Jaru. G) E. zolzivora, Bolivia, Cochabamba. H) E. greeneyi, Ecuador, Napo, Yanayacu Biological Station, PARATYPE." figureDoi="http://doi.org/10.5281/zenodo.4532993" httpUri="https://zenodo.org/record/4532993/files/figure.png" pageId="15" pageNumber="14">6G</figureCitation>
|
|
)
|
|
</emphasis>
|
|
</heading>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="CFA0D3BE5507230CFF4FD97552CC7E6B" box="[192,479,803,827]" pageId="15" pageNumber="14" type="materials_examined">
|
|
<paragraph id="870580355507230CFF4FD97552CC7E6B" blockId="15.[192,479,803,827]" box="[192,479,803,827]" pageId="15" pageNumber="14">
|
|
<materialsCitation id="37D28A685507230CFF4FD97552C87E6B" box="[192,475,803,827]" country="Bolivia" location="Bolivia" pageId="15" pageNumber="14" specimenCount="1" typeStatus="holotype">
|
|
<emphasis id="B5CE5C275507230CFF4FD97552697E6B" bold="true" box="[192,378,803,827]" pageId="15" pageNumber="14">
|
|
<typeStatus id="58013E975507230CFF4FD97552167E6B" box="[192,261,803,827]" pageId="15" pageNumber="14">Type</typeStatus>
|
|
locality.
|
|
</emphasis>
|
|
<collectingCountry id="FFADC0A55507230CFE0FD97552C87E6B" box="[384,475,803,827]" name="Bolivia" pageId="15" pageNumber="14">Bolivia</collectingCountry>
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="CFA0D3BE5507230CFF4FD93551FD7BEB" pageId="15" pageNumber="14" type="diagnosis">
|
|
<paragraph id="870580355507230CFF4FD93551777EAB" blockId="15.[192,1440,867,1019]" pageId="15" pageNumber="14">
|
|
<emphasis id="B5CE5C275507230CFF4FD935525A7E2B" bold="true" box="[192,329,867,891]" pageId="15" pageNumber="14">Diagnosis</emphasis>
|
|
. Males and females reddish-brown with conspicuous FW orange submarginal line. Distinguished from
|
|
<taxonomicName id="40BAFBB65507230CFEE6D9D252F77ECB" baseAuthorityName="Staudinger" baseAuthorityYear="1887" box="[361,484,900,923]" class="Insecta" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="opimus">
|
|
<emphasis id="B5CE5C275507230CFEE6D9D252F77ECB" box="[361,484,900,923]" italics="true" pageId="15" pageNumber="14">E. opimus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="40BAFBB65507230CFDAFD9D251B57ECB" authorityName="Penz and DeVries" authorityYear="2008" box="[544,678,900,923]" class="Insecta" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="greeneyi">
|
|
<emphasis id="B5CE5C275507230CFDAFD9D251B57ECB" box="[544,678,900,923]" italics="true" pageId="15" pageNumber="14">E. greeneyi</emphasis>
|
|
</taxonomicName>
|
|
by the dorsal FW orange submarginal line forming a thin, yet conspicuous distal orange crescent mark that is unique to this species. Here the postmedial branch of this line extends anteriorly to the wing margin, and the HW projection at Cu1 is vestigial or small, not extending beyond the anal margin.
|
|
</paragraph>
|
|
<paragraph id="870580355507230CFF4FDE7550BC78CB" blockId="15.[192,1441,1059,1723]" pageId="15" pageNumber="14">
|
|
<emphasis id="B5CE5C275507230CFF4FDE75517B796B" bold="true" box="[192,616,1059,1083]" pageId="15" pageNumber="14">Annotated redescription. Male.</emphasis>
|
|
FW length
|
|
<quantity id="40422DD05507230CFD7CDE75507F796B" box="[755,876,1059,1083]" metricMagnitude="-2" metricUnit="m" metricValue="5.85" metricValueMax="6.0" metricValueMin="5.7" pageId="15" pageNumber="14" unit="mm" value="58.5" valueMax="60.0" valueMin="57.0">57-60 mm</quantity>
|
|
(n=3). Dorsal FW and HW with reddish-brown background and very faint pinkish iridescent sheen from submedial to postmedial areas of the FW, starting approximately at midlength of the discal cell, but barely visible on the HW. Dorsal FW with a well-defined orange submarginal line interrupted by small gaps below M3, and forming a V-shaped mark with the arms directed distally in cell M3. Submarginal branch of FW submarginal line starts in cell M2, and postmedial branch starts at wing margin, being clearly visible below the R veins. Wing apex with a thin, orange crescent mark. Dorsal FW and HW with well-developed orange marginal outline. HW with a small projection at vein Cu1 that does not extend beyond anal margin. Ventral FW and HW main color brown. Ventral HW white postmedial transverse band well developed, continuous. Ventral HW with brown wavy lines across discal cell. Ventral HW eyespot in cell Sc+Rs not outlined by a pale halo. Ventral HW eyespot in cell Cu1 very large, expanding to cell Cu2. Inner dark circle larger than in
|
|
<taxonomicName id="40BAFBB65507230CFB7CDF32567F782B" baseAuthorityName="Staudinger" baseAuthorityYear="1887" box="[1267,1388,1380,1403]" class="Insecta" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="opimus">
|
|
<emphasis id="B5CE5C275507230CFB7CDF32567F782B" box="[1267,1388,1380,1403]" italics="true" pageId="15" pageNumber="14">E. opimus</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName id="40BAFBB65507230CFF4FDFD2525578CB" authorityName="Penz and DeVries" authorityYear="2008" box="[192,326,1412,1435]" class="Insecta" family="Nymphalidae" genus="Eryphanis" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="greeneyi">
|
|
<emphasis id="B5CE5C275507230CFF4FDFD2525578CB" box="[192,326,1412,1435]" italics="true" pageId="15" pageNumber="14">E. greeneyi</emphasis>
|
|
</taxonomicName>
|
|
. Uncus humped, valva with subterminal swelling.
|
|
</paragraph>
|
|
<paragraph id="870580355507230CFF7FDFF551FD7BEB" blockId="15.[192,1441,1059,1723]" pageId="15" pageNumber="14">
|
|
<emphasis id="B5CE5C275507230CFF7FDFF5524F78EB" bold="true" box="[240,348,1443,1467]" pageId="15" pageNumber="14">Female.</emphasis>
|
|
FW length
|
|
<quantity id="40422DD05507230CFE6ADFF5512478EB" box="[485,567,1443,1467]" metricMagnitude="-2" metricUnit="m" metricValue="6.6" pageId="15" pageNumber="14" unit="mm" value="66.0">66 mm</quantity>
|
|
(n=1). Dorsal FW and HW with reddish- brown background that is not as dense as in male, producing a semi-transparent effect. Both wings with pinkish iridescent sheen in the medial area, faint overall, but stronger in the HW. Dorsal FW with a well defined, thick orange submarginal line, barely interrupted at the wing veins below M3, and forming a V-shaped mark with the arms directed distally in cell M3. Submarginal branch of FW submarginal line starts in cell M2. Postmedial branch of submarginal line starts at wing margin, clearly visible in the cell below the R stem. Wing apex with a thin, orange crescent mark, as in male. HW projection at Cu1 can be less distinct than in male, and very small in some specimens. Ventral surface of wings as in male. Sterigma projection with an elongated arm, and multiple points anteriorly.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="CFA0D3BE5507230CFF4FDCB550F57A4B" pageId="15" pageNumber="14" type="distribution">
|
|
<paragraph id="870580355507230CFF4FDCB550F57A4B" blockId="15.[192,1440,1763,1819]" pageId="15" pageNumber="14">
|
|
<emphasis id="B5CE5C275507230CFF4FDCB5527E7BAB" bold="true" box="[192,365,1763,1787]" pageId="15" pageNumber="14">Distribution</emphasis>
|
|
.
|
|
<collectingCountry id="FFADC0A55507230CFEF4DCB552C57BAB" box="[379,470,1763,1787]" name="Bolivia" pageId="15" pageNumber="14">Bolivia</collectingCountry>
|
|
, perhaps reaching southern
|
|
<collectingCountry id="FFADC0A55507230CFCBEDCB2507E7BAB" box="[817,877,1764,1787]" name="Peru" pageId="15" pageNumber="14">Peru</collectingCountry>
|
|
(
|
|
<bibRefCitation id="E32BFDC45507230CFCF4DCB557417BAB" author="Fruhstorfer, H." box="[891,1106,1763,1787]" pageId="15" pageNumber="14" pagination="285 - 332" refId="ref12225" refString="Fruhstorfer, H. 1912. Family: Brassolidae, p 285 - 332. In: A. Seitz (ed.). Die Gross-Schmetterlinge der Erde, Volume 5. Alfred Kernen; Stuttgart. 1024 p." type="book chapter" year="1912">Frühstorfer 1912</bibRefCitation>
|
|
, Appendix 1). Collection labels list an elevation of
|
|
<quantity id="40422DD05507230CFE54DD5551287A4B" box="[475,571,1795,1819]" metricMagnitude="0" metricUnit="m" metricValue="1.65" pageId="15" pageNumber="14" unit="m" value="1.65">1,650 m</quantity>
|
|
(El Palmar,
|
|
<collectingRegion id="457E4ED75507230CFD5CDD5550667A4B" box="[723,885,1795,1819]" country="Bolivia" name="Cochabamba" pageId="15" pageNumber="14">Cochabamba</collectingRegion>
|
|
,
|
|
<collectingCountry id="FFADC0A55507230CFCF0DD5550C97A4B" box="[895,986,1795,1819]" name="Bolivia" pageId="15" pageNumber="14">Bolivia</collectingCountry>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |