375 lines
41 KiB
XML
375 lines
41 KiB
XML
<document id="263506568AB476A1D7D2ABA3F0EAD128" ID-DOI="10.11646/zootaxa.3609.6.3" ID-GBIF-Dataset="0c7c3c11-b26b-43a9-9fe1-237ee136e62b" ID-ISSN="1175-5326" ID-Zenodo-Dep="215563" ID-ZooBank="9269ABC0-246F-407F-8B42-550C1B3F3C59" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460573196693" checkinUser="plazi" docAuthor="Martinez, Mariano I., Thandar, Ahmed S. & Penchaszadeh, Pablo E." docDate="2013" docId="FF3C484D4B53FFAFBDBEFB3AFCD4FB96" docLanguage="en" docName="zt03609p588.pdf" docOrigin="Zootaxa 3609 (6)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Havelockia Pearson 1903" docType="treatment" docVersion="7" lastPageNumber="587" masterDocId="030530354B52FFABBD29FF9CFFACFFEA" masterDocTitle="A new species of Havelockia Pearson, 1903 from the Argentine Sea (Holothuroidea: Dendrochirotida: Sclerodactylidae)" masterLastPageNumber="588" masterPageNumber="583" pageNumber="584" updateTime="1698322627716" updateUser="plazi">
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<mods:mods id="A7EEED9A1E41EF6F49B07D51A60D0631" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BD1733E8C9D7C8B87BE0A350765488C8">
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<mods:title id="D6143B7F07BC2102DD9D2803C9E90FBB">A new species of Havelockia Pearson, 1903 from the Argentine Sea (Holothuroidea: Dendrochirotida: Sclerodactylidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="7F38306F602B4EDD058D1B4CD92B5244" type="personal">
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<mods:role id="620BFF6450FCAB9B2C077ED3BCA4873F">
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<mods:roleTerm id="5A638A2B1B261344AC3C34D28F0ED6AB">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="72D38E8A44569D8FBF88DD09E8295D83">Martinez, Mariano I.</mods:namePart>
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</mods:name>
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<mods:name id="B55AF43575EFEE0C13678FDA4679A235" type="personal">
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<mods:roleTerm id="E9A2B275A3521BB9E08BBD5C9A98CF39">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="C0B64CC36BFE88CA7AF7876FE39599E5">Thandar, Ahmed S.</mods:namePart>
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</mods:name>
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<mods:name id="0EA61E4B3B29A410E3AA12A2ADD9A3CE" type="personal">
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<mods:role id="9A14AC565161E326B0B8EB60AC90B3BB">
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<mods:roleTerm id="1333FD0F503572579F592690C026130A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="32F5A354F0001FCDDA9AE315223F6286">Penchaszadeh, Pablo E.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="623BA144F36F3FFF3D7181FF15DE52E9">text</mods:typeOfResource>
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<mods:titleInfo id="8FF90F9B69CCB2A63D414C0FC47CF7D8">
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<mods:title id="0BFB0C0AA25D165BE2456D6D7505FB19">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="CC64A0B799704E8A6864FB8539F246E5">
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<mods:date id="0C498901E4BF2A2E8D2A374E5E7230AB">2013</mods:date>
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<mods:detail id="17F3D8AB735E1E57DB25562A93BBAF8A" type="volume">
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<mods:number id="C53B612ED238EC039A6FA46F509E92A5">3609</mods:number>
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</mods:detail>
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<mods:detail id="77BA6D515E98315C2D73472F1A9E9E2C" type="issue">
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<mods:number id="F2F6541687858A95E3B0DD80F9D5D7A5">6</mods:number>
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<mods:start id="6C0E4251E5377A3F20A7A688539C51C1">583</mods:start>
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<mods:end id="FB36F1E647C48F7FBC8BC3977AD6D2B4">588</mods:end>
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</mods:extent>
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<mods:classification id="52D9EE19C8573F876E780A2E020E38FD">journal article</mods:classification>
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<mods:identifier id="8FA9419D5E168DC59041A0197DF8B500" type="DOI">10.11646/zootaxa.3609.6.3</mods:identifier>
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<mods:identifier id="738B4A25EDF93B977D0FB390997EB1DB" type="GBIF-Dataset">0c7c3c11-b26b-43a9-9fe1-237ee136e62b</mods:identifier>
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<mods:identifier id="CD4F118566E19EBBF7565321CA43F77F" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="AC2C6FC526C4BB900F8E9F92BF42E114" type="Zenodo-Dep">215563</mods:identifier>
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<mods:identifier id="F45869606EC423ACDCB86E46841A506B" type="ZooBank">9269ABC0-246F-407F-8B42-550C1B3F3C59</mods:identifier>
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</mods:mods>
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<treatment id="FF3C484D4B53FFAFBDBEFB3AFCD4FB96" ID-DOI="http://doi.org/10.5281/zenodo.6150605" ID-GBIF-Taxon="119559248" ID-Zenodo-Dep="6150605" LSID="urn:lsid:plazi:treatment:FF3C484D4B53FFAFBDBEFB3AFCD4FB96" httpUri="http://treatment.plazi.org/id/FF3C484D4B53FFAFBDBEFB3AFCD4FB96" lastPageId="4" lastPageNumber="587" pageId="1" pageNumber="584">
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<subSubSection id="3F8FAAD04B53FFAABDBEFB3AFD80FB2A" box="[151,556,1190,1216]" pageId="1" pageNumber="584" type="nomenclature">
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<paragraph id="772AF95B4B53FFAABDBEFB3AFD80FB2A" blockId="1.[151,556,1190,1216]" box="[151,556,1190,1216]" pageId="1" pageNumber="584">
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<heading id="2C624E374B53FFAABDBEFB3AFD80FB2A" bold="true" box="[151,556,1190,1216]" fontSize="11" level="1" pageId="1" pageNumber="584" reason="1">
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||
<emphasis id="45E125494B53FFAABDBEFB3AFD80FB2A" bold="true" box="[151,556,1190,1216]" pageId="1" pageNumber="584">
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Genus
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<taxonomicName id="B09582D84B53FFAABDC7FB3AFD80FB2A" ID-CoL="7P5HX" authority="Pearson, 1903" authorityName="Pearson" authorityYear="1903" box="[238,556,1190,1216]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="584" phylum="Echinodermata" rank="genus">
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<emphasis id="45E125494B53FFAABDC7FB3AFEDBFB2A" bold="true" box="[238,375,1190,1216]" italics="true" pageId="1" pageNumber="584">Havelockia</emphasis>
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Pearson, 1903
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="3F8FAAD04B53FFAABDBEFB71FC91FA7F" pageId="1" pageNumber="584" type="diagnosis">
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<paragraph id="772AF95B4B53FFAABDBEFB71FD95FAEF" blockId="1.[151,1436,1261,1466]" box="[151,569,1261,1286]" pageId="1" pageNumber="584">
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<emphasis id="45E125494B53FFAABDBEFB71FEA6FAEC" bold="true" box="[151,266,1261,1286]" pageId="1" pageNumber="584">Diagnosis</emphasis>
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(after Thandar, 1989: 292)
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</paragraph>
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<paragraph id="772AF95B4B53FFAABDEEFA8DFC91FA7F" blockId="1.[151,1436,1261,1466]" pageId="1" pageNumber="584">Calcareous ring short, stout, only anterior projections of radial and interradial plates free; posterior paired processes of radial plates divided into several pieces. Body wall ossicles tables with a squarish to oval disc usually perforated by four large central and four smaller peripheral holes, the latter sometimes reduced or absent; spire of two pillars joined at apex and terminating in few blunt teeth.</paragraph>
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</subSubSection>
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<subSubSection id="3F8FAAD04B53FFAABDEEFA3EFE82F9A3" pageId="1" pageNumber="584" type="materials_examined">
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||
<paragraph id="772AF95B4B53FFAABDEEFA3EFB8FFA53" blockId="1.[151,1436,1261,1466]" box="[199,1059,1441,1466]" pageId="1" pageNumber="584">
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||
<emphasis id="45E125494B53FFAABDEEFA3EFEF7FA50" bold="true" box="[199,347,1441,1466]" pageId="1" pageNumber="584">
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||
<typeStatus id="A82E47F94B53FFAABDEEFA3EFEAEFA50" box="[199,258,1442,1466]" pageId="1" pageNumber="584">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="B09582D84B53FFAABC43FA3DFCACFA53" authority="Pearson, 1903" authorityName="Pearson" authorityYear="1903" box="[362,768,1441,1465]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="584" phylum="Echinodermata" rank="species" species="herdmani">
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<emphasis id="45E125494B53FFAABC43FA3DFDF6FA53" box="[362,602,1441,1465]" italics="true" pageId="1" pageNumber="584">Havelockia herdmani</emphasis>
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Pearson, 1903
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</taxonomicName>
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(by original designation).
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</paragraph>
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<paragraph id="772AF95B4B53FFAABDBEF991FE82F9A3" blockId="1.[151,433,1549,1609]" pageId="1" pageNumber="584">
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<emphasis id="45E125494B53FFAABDBEF991FE1DF9CF" bold="true" box="[151,433,1549,1574]" pageId="1" pageNumber="584">
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<taxonomicName id="B09582D84B53FFAABDBEF991FEFCF9CC" box="[151,336,1549,1574]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="584" phylum="Echinodermata" rank="species" species="pegi" status="sp. nov.">
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<emphasis id="45E125494B53FFAABDBEF991FEFCF9CC" bold="true" box="[151,336,1549,1574]" italics="true" pageId="1" pageNumber="584">Havelockia pegi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="5ED298324B53FFAABC71F991FE1DF9CF" box="[344,433,1549,1573]" pageId="1" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(
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<figureCitation id="EFAEE5DE4B53FFAABDB7F9ADFE89F9A3" box="[158,293,1585,1609]" captionStart-0="FIGURE 1. A" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3. A" captionStartId-0="2.[151,250,2015,2038]" captionStartId-1="3.[151,250,984,1007]" captionStartId-2="3.[151,250,1929,1952]" captionTargetBox-0="[155,1432,1047,2004]" captionTargetBox-1="[155,1432,193,962]" captionTargetBox-2="[155,1432,1117,1907]" captionTargetId-0="figure@2.[155,1433,1047,2005]" captionTargetId-1="figure@3.[155,1432,193,963]" captionTargetId-2="figure@3.[155,1433,1117,1907]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="3" captionText-0="FIGURE 1. A. Holotype of Havelockia pegi sp. nov., MACN-In 39019, scale 2 cm; B. Calcareous ring." captionText-1="FIGURE 2. Ossicles of Havelockia pegi sp. nov. A. Tables from body wall; B. Quadrilocular tables from tube feet; C. endplate; D. Rosettes from introvert and tentacles. Scale 50 µm." captionText-2="FIGURE 3. A. table from body wall near anus; B. Quadrilocular table from tube foot. Scale 20 µm." httpUri-0="https://zenodo.org/record/215564/files/figure.png" httpUri-1="https://zenodo.org/record/215565/files/figure.png" httpUri-2="https://zenodo.org/record/215566/files/figure.png" pageId="1" pageNumber="584">Figures 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="3F8FAAD04B53FFAABDBEF9E5FF58F933" pageId="1" pageNumber="584" type="diagnosis">
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||
<paragraph id="772AF95B4B53FFAABDBEF9E5FF58F933" blockId="1.[151,1437,1657,2006]" pageId="1" pageNumber="584">
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||
<emphasis id="45E125494B53FFAABDBEF9E5FEBDF978" bold="true" box="[151,273,1657,1682]" pageId="1" pageNumber="584">Diagnosis.</emphasis>
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Tentacles purple in colour; body wall ossicles scarce, only few detected, especially from the anal region and base of tube feet, comprising tables with a 2-pillared, often regressed spire; tentacles and introvert with rosettes.
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</paragraph>
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</subSubSection>
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<subSubSection id="3F8FAAD04B53FFAABDEEF979FD94F8CB" pageId="1" pageNumber="584" type="etymology">
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||
<paragraph id="772AF95B4B53FFAABDEEF979FD94F8CB" blockId="1.[151,1437,1657,2006]" pageId="1" pageNumber="584">
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<emphasis id="45E125494B53FFAABDEEF979FEE2F917" bold="true" box="[199,334,1765,1789]" pageId="1" pageNumber="584">Etymology.</emphasis>
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The specific name
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<taxonomicName id="B09582D84B53FFAABF0EF97AFDFBF917" box="[551,599,1766,1789]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="1" pageNumber="584" phylum="Echinodermata" rank="species" species="pegi">
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<emphasis id="45E125494B53FFAABF0EF97AFDFBF917" box="[551,599,1766,1789]" italics="true" pageId="1" pageNumber="584">pegi</emphasis>
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</taxonomicName>
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is used to honour the Martinez-Ferreyra family to whom this name is quite significant. It is a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="3F8FAAD04B53FFA9BDEEF8B1FB2DFF3E" lastPageId="2" lastPageNumber="585" pageId="1" pageNumber="584" type="materials_examined">
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||
<paragraph id="772AF95B4B53FFAABDEEF8B1FA31F83F" blockId="1.[151,1437,1657,2006]" pageId="1" pageNumber="584">
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||
<emphasis id="45E125494B53FFAABDEEF8B1FE1CF8AC" bold="true" box="[199,432,1837,1862]" pageId="1" pageNumber="584">Material examined.</emphasis>
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<emphasis id="45E125494B53FFAABC91F8B1FDB3F8AF" box="[440,543,1837,1861]" italics="true" pageId="1" pageNumber="584">
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||
<typeStatus id="A82E47F94B53FFAABC91F8B1FDB3F8AF" box="[440,543,1837,1861]" pageId="1" pageNumber="584" type="holotype">Holotype</typeStatus>
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</emphasis>
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:
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<quantity id="B06D54BE4B53FFAABF04F8B1FDD1F8AF" box="[557,637,1837,1861]" metricMagnitude="-2" metricUnit="m" metricValue="8.6" pageId="1" pageNumber="584" unit="mm" value="86.0">86 mm</quantity>
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length, one male individual and 3 slides with ossicles, Villarino, Golfo San José, Chubut Province (
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<date id="032BDF9B4B53FFAABCF8F8CDFDE9F883" box="[465,581,1873,1897]" pageId="1" pageNumber="584" value="2006-09">Sep. 2006</date>
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), (
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||
<geoCoordinate id="12A19F9C4B53FFAABF48F8CDFD10F883" box="[609,700,1873,1897]" direction="south" orientation="latitude" pageId="1" pageNumber="584" precision="925" value="-42.416668">42°25’S</geoCoordinate>
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||
–
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||
<geoCoordinate id="12A19F9C4B53FFAABFE4F8CDFC9AF883" box="[717,822,1873,1897]" direction="west" orientation="longitude" pageId="1" pageNumber="584" precision="925" value="-64.51667">64°31’W</geoCoordinate>
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||
),
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||
<specimenCount id="619332D24B53FFAABE63F8CDFC25F883" box="[842,905,1873,1897]" pageId="1" pageNumber="584" type="generic">
|
||
<quantity id="B06D54BE4B53FFAABE63F8CDFC29F883" box="[842,901,1873,1897]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="584" unit="m" value="10.0">10 m</quantity>
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||
.
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||
</specimenCount>
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||
MACN-In 39019.
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||
<emphasis id="45E125494B53FFAAB977F8CEFB7CF883" box="[1118,1232,1874,1897]" italics="true" pageId="1" pageNumber="584">
|
||
<typeStatus id="A82E47F94B53FFAAB977F8CEFB7CF883" box="[1118,1232,1874,1897]" pageId="1" pageNumber="584" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
:
|
||
<quantity id="B06D54BE4B53FFAAB9F7F8CDFAFFF883" box="[1246,1363,1873,1897]" metricMagnitude="-2" metricUnit="m" metricValue="3.27" pageId="1" pageNumber="584" unit="mm" value="32.7">32,70 mm</quantity>
|
||
,
|
||
<quantity id="B06D54BE4B53FFAAB874F8CDFF69F867" metricMagnitude="-2" metricUnit="m" metricValue="5.595" pageId="1" pageNumber="584" unit="mm" value="55.95">55,95 mm</quantity>
|
||
, Buenos Aires Province (
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||
<date id="032BDF9B4B53FFAABCC8F8E9FDC5F867" box="[481,617,1909,1933]" pageId="1" pageNumber="584" value="2009-09-09">9 Sep. 2009</date>
|
||
), (
|
||
<geoCoordinate id="12A19F9C4B53FFAABFACF8E9FD72F867" box="[645,734,1909,1933]" direction="south" orientation="latitude" pageId="1" pageNumber="584" precision="925" value="-38.25">38°15’S</geoCoordinate>
|
||
–
|
||
<geoCoordinate id="12A19F9C4B53FFAABFC6F8E9FCF6F867" box="[751,858,1909,1933]" direction="west" orientation="longitude" pageId="1" pageNumber="584" precision="925" value="-57.25">57°15’W</geoCoordinate>
|
||
),
|
||
<specimenCount id="619332D24B53FFAABE47F8E9FC03F867" box="[878,943,1909,1933]" pageId="1" pageNumber="584" type="generic">
|
||
<quantity id="B06D54BE4B53FFAABE47F8E9FC07F867" box="[878,939,1909,1933]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="1" pageNumber="584" unit="m" value="48.0">48 m</quantity>
|
||
.
|
||
</specimenCount>
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||
, MACN-In 39021 (2
|
||
<typeStatus id="A82E47F94B53FFAAB988F8E9FABEF867" box="[1185,1298,1909,1933]" pageId="1" pageNumber="584" type="paratype">paratypes</typeStatus>
|
||
);
|
||
<quantity id="B06D54BE4B53FFAAB80DF8E9FA3BF867" box="[1316,1431,1909,1933]" metricMagnitude="-2" metricUnit="m" metricValue="3.5549999999999997" pageId="1" pageNumber="584" unit="mm" value="35.55">35,55 mm</quantity>
|
||
, Villarino, Golfo San José, Chubut Province (
|
||
<geoCoordinate id="12A19F9C4B53FFAABFBFF805FD5EF85B" box="[662,754,1945,1969]" direction="south" orientation="latitude" pageId="1" pageNumber="584" precision="925" value="-42.416668">42°25’S</geoCoordinate>
|
||
–
|
||
<geoCoordinate id="12A19F9C4B53FFAABE2AF805FCC7F85B" box="[771,875,1945,1969]" direction="west" orientation="longitude" pageId="1" pageNumber="584" precision="925" value="-64.51667">64°31’W</geoCoordinate>
|
||
),
|
||
<quantity id="B06D54BE4B53FFAABEA8F805FC1AF85B" box="[897,950,1945,1969]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="584" unit="m" value="10.0">10m</quantity>
|
||
, MACN-In 39020 (1
|
||
<typeStatus id="A82E47F94B53FFAAB986F805FAB9F85B" box="[1199,1301,1945,1969]" pageId="1" pageNumber="584" type="paratype">paratype</typeStatus>
|
||
);
|
||
<quantity id="B06D54BE4B53FFAAB800F805FA30F85B" box="[1321,1436,1945,1969]" metricMagnitude="-2" metricUnit="m" metricValue="3.043" pageId="1" pageNumber="584" unit="mm" value="30.43">30,43 mm</quantity>
|
||
length, one individual and 1 slide with ossicles; El sótano, Golfo San Matías, Río Negro Province,
|
||
<collectingCountry id="0F82B9CB4B53FFAAB9CEF821FAFAF83F" box="[1255,1366,1981,2005]" name="Argentina" pageId="1" pageNumber="584">Argentina</collectingCountry>
|
||
(Nov.
|
||
</paragraph>
|
||
<paragraph id="772AF95B4B50FFA9BDBEFF0BFB2DFF3E" blockId="2.[151,1436,151,1004]" pageId="2" pageNumber="585">
|
||
2011), (
|
||
<geoCoordinate id="12A19F9C4B50FFA9BDC6FF0BFEFDFF45" box="[239,337,151,175]" direction="south" orientation="latitude" pageId="2" pageNumber="585" precision="925" value="-41.0">41º00’ S</geoCoordinate>
|
||
–
|
||
<geoCoordinate id="12A19F9C4B50FFA9BC48FF0BFE7CFF45" box="[353,464,151,175]" direction="west" orientation="longitude" pageId="2" pageNumber="585" precision="925" value="-65.13333">65º08’ W</geoCoordinate>
|
||
),
|
||
<quantity id="B06D54BE4B50FFA9BCCFFF0BFD8EFF45" box="[486,546,151,175]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="585" unit="m" value="15.0">15 m</quantity>
|
||
, ICML-UNAM 5.195.0 (1
|
||
<typeStatus id="A82E47F94B50FFA9BE70FF0BFC13FF45" box="[857,959,151,175]" pageId="2" pageNumber="585" type="paratype">paratype</typeStatus>
|
||
);
|
||
<quantity id="B06D54BE4B50FFA9BEFAFF0BFB8AFF45" box="[979,1062,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="585" unit="mm" value="40.0">40 mm</quantity>
|
||
, one individual; San Matías, Río Negro Province (
|
||
<date id="032BDF9B4B50FFA9BC7EFF20FE5FFF3E" box="[343,499,188,212]" pageId="2" pageNumber="585" value="2009-11-16">16 Nov. 2009</date>
|
||
), (
|
||
<geoCoordinate id="12A19F9C4B50FFA9BF26FF20FDC0FF3E" box="[527,620,188,212]" direction="south" orientation="latitude" pageId="2" pageNumber="585" precision="925" value="-41.333332">41º 20’S</geoCoordinate>
|
||
–
|
||
<geoCoordinate id="12A19F9C4B50FFA9BF55FF20FD40FF3E" box="[636,748,188,212]" direction="west" orientation="longitude" pageId="2" pageNumber="585" precision="925" value="-62.983334">62º 59’W</geoCoordinate>
|
||
),
|
||
<quantity id="B06D54BE4B50FFA9BE28FF20FC90FF3E" box="[769,828,188,212]" metricMagnitude="1" metricUnit="m" metricValue="3.4" pageId="2" pageNumber="585" unit="m" value="34.0">34 m</quantity>
|
||
, CNP-INV 415 (1
|
||
<typeStatus id="A82E47F94B50FFA9B926FF20FBD9FF3E" box="[1039,1141,188,212]" pageId="2" pageNumber="585" type="paratype">paratype</typeStatus>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3F8FAAD04B50FFA9BDEEFF43FD27FC95" pageId="2" pageNumber="585" type="description">
|
||
<paragraph id="772AF95B4B50FFA9BDEEFF43FBB4FDB5" blockId="2.[151,1436,151,1004]" pageId="2" pageNumber="585">
|
||
<emphasis id="45E125494B50FFA9BDEEFF43FEF5FF12" bold="true" box="[199,345,223,248]" pageId="2" pageNumber="585">Description.</emphasis>
|
||
All specimens partially eviscerated; length up to
|
||
<quantity id="B06D54BE4B50FFA9BEB4FF43FC5EFF1D" box="[925,1010,223,247]" metricMagnitude="-2" metricUnit="m" metricValue="8.6" pageId="2" pageNumber="585" unit="mm" value="86.0">86 mm</quantity>
|
||
. Body form U-shaped to cylindrical. Colour in life and in alcohol, brown; tentacles purple in life, turning to light violet or brown in alcohol. Tube feet scattered, covering entire body with no indication of any regular arrangement (
|
||
<figureCitation id="EFAEE5DE4B50FFA9B935FEBBFBF3FED5" box="[1052,1119,295,319]" captionStart="FIGURE 1. A" captionStartId="2.[151,250,2015,2038]" captionTargetBox="[155,1432,1047,2004]" captionTargetId="figure@2.[155,1433,1047,2005]" captionTargetPageId="2" captionText="FIGURE 1. A. Holotype of Havelockia pegi sp. nov., MACN-In 39019, scale 2 cm; B. Calcareous ring." httpUri="https://zenodo.org/record/215564/files/figure.png" pageId="2" pageNumber="585">Fig. 1</figureCitation>
|
||
). Larger tube feet around
|
||
<quantity id="B06D54BE4B50FFA9B8A7FEBBFF6FFE8E" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="585" unit="mm" value="2.0">2 mm</quantity>
|
||
in length, suckers well developed. Tentacles 10, bushy, ventral two reduced to quarter the size of large tentacles. Anal teeth present. Calcareous ring short-tubular with plates compact, fused for most of their length; posterior paired processes of radial plates of moderate length, divided into 3–4 pieces of calcite. Polian vesicle single, saccular, free, on left side of mesentery; stone canal long, straight, on right side of mesentery, madreporite lenticular. Gonad of
|
||
<typeStatus id="A82E47F94B50FFA9BCA9FE40FE4EFE1E" box="[384,482,476,500]" pageId="2" pageNumber="585" type="holotype">holotype</typeStatus>
|
||
mature, in two tufts, attached to the middle of body, each with numerous undivided branches, filling greater part of coelom. Respiratory trees, each with two branches, extending along the total length of body, the right tree slightly longer. Longitudinal muscle bands well developed. Retractor muscles attach to the radial plates. Cloaca elongate, about quarter to one fifth the total length of body.
|
||
</paragraph>
|
||
<paragraph id="772AF95B4B50FFA9BDEEFDF0FED6FCB6" blockId="2.[151,1436,151,1004]" pageId="2" pageNumber="585">
|
||
Body wall ossicles rare, only a few detected, especially in the anal region and bases of tube feet, comprising tables with an oblong or irregular, often regressive disc (65–160 µm) with 4–8 holes and a low, 2-pillared spire, pillars sometimes fused (
|
||
<figureCitation id="EFAEE5DE4B50FFA9BC9AFD28FE55FD26" box="[435,505,692,716]" captionStart="FIGURE 2" captionStartId="3.[151,250,984,1007]" captionTargetBox="[155,1432,193,962]" captionTargetId="figure@3.[155,1432,193,963]" captionTargetPageId="3" captionText="FIGURE 2. Ossicles of Havelockia pegi sp. nov. A. Tables from body wall; B. Quadrilocular tables from tube feet; C. endplate; D. Rosettes from introvert and tentacles. Scale 50 µm." httpUri="https://zenodo.org/record/215565/files/figure.png" pageId="2" pageNumber="585">Fig. 2</figureCitation>
|
||
a, 3a). Tube feet ossicles as tables with a quadrilocular, straight or slightly curved disc (115–172 µm), with four central holes and often an additional hole at each end; spire 2-pillared, ending in several blunt teeth or spire regressive or absent (
|
||
<figureCitation id="EFAEE5DE4B50FFA9BFE3FD60FCBCFCFE" box="[714,784,764,788]" captionStart="FIGURE 2" captionStartId="3.[151,250,984,1007]" captionTargetBox="[155,1432,193,962]" captionTargetId="figure@3.[155,1432,193,963]" captionTargetPageId="3" captionText="FIGURE 2. Ossicles of Havelockia pegi sp. nov. A. Tables from body wall; B. Quadrilocular tables from tube feet; C. endplate; D. Rosettes from introvert and tentacles. Scale 50 µm." httpUri="https://zenodo.org/record/215565/files/figure.png" pageId="2" pageNumber="585">Fig. 2</figureCitation>
|
||
b, 3b), end-plate circular, ca. 225 µm, with small central holes and larger ones outside these in no regular arrangement (
|
||
<figureCitation id="EFAEE5DE4B50FFA9BE7BFC83FC3AFCDD" box="[850,918,799,823]" captionStart="FIGURE 2" captionStartId="3.[151,250,984,1007]" captionTargetBox="[155,1432,193,962]" captionTargetId="figure@3.[155,1432,193,963]" captionTargetPageId="3" captionText="FIGURE 2. Ossicles of Havelockia pegi sp. nov. A. Tables from body wall; B. Quadrilocular tables from tube feet; C. endplate; D. Rosettes from introvert and tentacles. Scale 50 µm." httpUri="https://zenodo.org/record/215565/files/figure.png" pageId="2" pageNumber="585">Fig. 2</figureCitation>
|
||
c). Introvert and tentacles with closed rosettes, 30–62 µm (
|
||
<figureCitation id="EFAEE5DE4B50FFA9BC30FCD8FEF0FCB6" box="[281,348,836,860]" captionStart="FIGURE 2" captionStartId="3.[151,250,984,1007]" captionTargetBox="[155,1432,193,962]" captionTargetId="figure@3.[155,1432,193,963]" captionTargetPageId="3" captionText="FIGURE 2. Ossicles of Havelockia pegi sp. nov. A. Tables from body wall; B. Quadrilocular tables from tube feet; C. endplate; D. Rosettes from introvert and tentacles. Scale 50 µm." httpUri="https://zenodo.org/record/215565/files/figure.png" pageId="2" pageNumber="585">Fig. 2</figureCitation>
|
||
d).
|
||
</paragraph>
|
||
<paragraph id="772AF95B4B50FFA9BDEEFCFBFD27FC95" blockId="2.[151,1436,151,1004]" box="[199,651,871,896]" pageId="2" pageNumber="585">
|
||
<emphasis id="45E125494B50FFA9BDEEFCFBFE8FFC6A" bold="true" box="[199,291,871,896]" pageId="2" pageNumber="585">Habitat</emphasis>
|
||
: sand, broken shells and gravel.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="3F8FAAD04B50FFA9BDEEFC17FE24FC06" pageId="2" pageNumber="585" type="distribution">
|
||
<paragraph id="772AF95B4B50FFA9BDEEFC17FD03FC2D" blockId="2.[151,1436,151,1004]" pageId="2" pageNumber="585">
|
||
<emphasis id="45E125494B50FFA9BDEEFC17FEF5FC4E" bold="true" box="[199,345,907,932]" pageId="2" pageNumber="585">Distribution</emphasis>
|
||
: South Atlantic Ocean, from Buenos Aires Province, Mar del Plata (
|
||
<geoCoordinate id="12A19F9C4B50FFA9B9B7FC10FB5AFC4E" box="[1182,1270,908,932]" direction="south" orientation="latitude" pageId="2" pageNumber="585" precision="925" value="-38.25">38°15’S</geoCoordinate>
|
||
–
|
||
<geoCoordinate id="12A19F9C4B50FFA9B82EFC10FADCFC4E" box="[1287,1392,908,932]" direction="west" orientation="longitude" pageId="2" pageNumber="585" precision="925" value="-57.25">57°15’W</geoCoordinate>
|
||
) to Chubut Province, Villarino (
|
||
<geoCoordinate id="12A19F9C4B50FFA9BCFCFC33FD83FC2D" box="[469,559,943,967]" direction="south" orientation="latitude" pageId="2" pageNumber="585" precision="925" value="-42.416668">42°25’S</geoCoordinate>
|
||
–
|
||
<geoCoordinate id="12A19F9C4B50FFA9BF69FC33FD04FC2D" box="[576,680,943,967]" direction="west" orientation="longitude" pageId="2" pageNumber="585" precision="925" value="-64.51667">64°31’W</geoCoordinate>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="772AF95B4B50FFA9BDEEFC48FE24FC06" blockId="2.[151,1436,151,1004]" box="[199,392,980,1004]" pageId="2" pageNumber="585">
|
||
<emphasis id="45E125494B50FFA9BDEEFC48FEBCFC06" bold="true" box="[199,272,980,1004]" pageId="2" pageNumber="585">Depth</emphasis>
|
||
:
|
||
<quantity id="B06D54BE4B50FFA9BC37FC48FE28FC06" box="[286,388,980,1004]" metricMagnitude="1" metricUnit="m" metricValue="2.9" metricValueMax="4.8" metricValueMin="1.0" pageId="2" pageNumber="585" unit="m" value="29.0" valueMax="48.0" valueMin="10.0">
|
||
10–
|
||
<specimenCount id="619332D24B50FFA9BC61FC48FE28FC06" box="[328,388,980,1004]" pageId="2" pageNumber="585" type="generic">48 m</specimenCount>
|
||
</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="23EAA9D34B50FFA9BDBEF843FB15F81C" box="[151,1209,2015,2038]" httpUri="https://zenodo.org/record/215564/files/figure.png" pageId="2" pageNumber="585" targetBox="[155,1432,1047,2004]" targetPageId="2">
|
||
<paragraph id="772AF95B4B50FFA9BDBEF843FB15F81C" blockId="2.[151,1209,2015,2038]" box="[151,1209,2015,2038]" pageId="2" pageNumber="585">
|
||
<emphasis id="45E125494B50FFA9BDBEF843FEBFF81F" bold="true" box="[151,275,2015,2038]" pageId="2" pageNumber="585">FIGURE 1.</emphasis>
|
||
A. Holotype of
|
||
<taxonomicName id="B09582D84B50FFA9BC91F87CFDF1F81C" box="[440,605,2016,2038]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="2" pageNumber="585" phylum="Echinodermata" rank="species" species="pegi" status="sp. nov.">
|
||
<emphasis id="45E125494B50FFA9BC91F87CFDF1F81C" box="[440,605,2016,2038]" italics="true" pageId="2" pageNumber="585">Havelockia pegi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="45E125494B50FFA9BF4AF87CFD18F81F" bold="true" box="[611,692,2016,2037]" pageId="2" pageNumber="585">
|
||
<taxonomicNameLabel id="5ED298324B50FFA9BF4AF87CFD18F81F" box="[611,692,2016,2037]" pageId="2" pageNumber="585" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, MACN-In 39019, scale 2 cm; B. Calcareous ring.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="23EAA9D34B51FFA8BDBEFC44FD50FBE7" httpUri="https://zenodo.org/record/215565/files/figure.png" pageId="3" pageNumber="586" targetBox="[155,1432,193,962]" targetPageId="3">
|
||
<paragraph id="772AF95B4B51FFA8BDBEFC44FD50FBE7" blockId="3.[151,1436,984,1037]" pageId="3" pageNumber="586">
|
||
<emphasis id="45E125494B51FFA8BDBEFC44FEB8FC07" bold="true" box="[151,276,984,1007]" pageId="3" pageNumber="586">FIGURE 2.</emphasis>
|
||
Ossicles of
|
||
<taxonomicName id="B09582D84B51FFA8BCBCFC44FD90FC04" box="[405,572,984,1006]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="3" pageNumber="586" phylum="Echinodermata" rank="species" species="pegi" status="sp. nov.">
|
||
<emphasis id="45E125494B51FFA8BCBCFC44FD90FC04" box="[405,572,984,1006]" italics="true" pageId="3" pageNumber="586">Havelockia pegi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="45E125494B51FFA8BF6DFC45FD3AFC04" bold="true" box="[580,662,985,1006]" pageId="3" pageNumber="586">
|
||
<taxonomicNameLabel id="5ED298324B51FFA8BF6DFC45FD3AFC04" box="[580,662,985,1006]" pageId="3" pageNumber="586" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A. Tables from body wall; B. Quadrilocular tables from tube feet; C. endplate; D. Rosettes from introvert and tentacles. Scale 50 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="23EAA9D34B51FFA8BDBEF815FB20F875" box="[151,1164,1929,1952]" httpUri="https://zenodo.org/record/215566/files/figure.png" pageId="3" pageNumber="586" targetBox="[155,1432,1117,1907]" targetPageId="3">
|
||
<paragraph id="772AF95B4B51FFA8BDBEF815FB20F875" blockId="3.[151,1164,1929,1952]" box="[151,1164,1929,1952]" pageId="3" pageNumber="586">
|
||
<emphasis id="45E125494B51FFA8BDBEF815FEBEF875" bold="true" box="[151,274,1929,1952]" pageId="3" pageNumber="586">FIGURE 3.</emphasis>
|
||
A. table from body wall near anus; B. Quadrilocular table from tube foot. Scale 20 µm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="3F8FAAD04B56FFAFBDEEFF04FCD4FB96" pageId="4" pageNumber="587" type="discussion">
|
||
<paragraph id="772AF95B4B56FFAFBDEEFF04FBEAFF1D" blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="587">
|
||
<emphasis id="45E125494B56FFAFBDEEFF04FE98FF5A" bold="true" box="[199,308,152,176]" pageId="4" pageNumber="587">Remarks</emphasis>
|
||
. In all individuals examined, the tentacles, calcareous ring and the first part of the gut are partially eviscerated. The dark purple colour of the tentacles, in living, and brown or light-violet in preserved specimens, is quite distinctive in comparison with that of some other Argentinian dendrochirotids.
|
||
</paragraph>
|
||
<paragraph id="772AF95B4B56FFAFBDEEFE98FC9AFD4D" blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="587">
|
||
The family
|
||
<taxonomicName id="B09582D84B56FFAFBC7CFE98FDBCFEF6" box="[341,528,260,284]" class="Holothuroidea" family="Sclerodactylidae" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="family">Sclerodactylidae</taxonomicName>
|
||
(sensu Pawson & Fell 1965) comprises over 80 species with a world-wide distribution, but it is surprising that besides
|
||
<taxonomicName id="B09582D84B56FFAFBF9AFEB4FBF8FED5" authority="Ludwig, 1887" authorityName="Ludwig" authorityYear="1887" box="[691,1108,295,320]" class="Holothuroidea" family="Sclerodactylidae" genus="Pseudothyone" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="species" species="belli">
|
||
<emphasis id="45E125494B56FFAFBF9AFEB4FC23FEAA" box="[691,911,296,320]" italics="true" pageId="4" pageNumber="587">Pseudothyone belli</emphasis>
|
||
(Ludwig, 1887)
|
||
</taxonomicName>
|
||
and the recently described
|
||
<taxonomicName id="B09582D84B56FFAFBDBEFED1FD2EFE8E" authority="Martins & Souto, 2012" authorityName="Martins & Souto" authorityYear="2012" box="[151,642,332,356]" class="Holothuroidea" family="Sclerodactylidae" genus="Coronatum" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="species" species="baiensis">
|
||
<emphasis id="45E125494B56FFAFBDBEFED1FED4FE8E" box="[151,376,332,356]" italics="true" pageId="4" pageNumber="587">Coronatum baiensis</emphasis>
|
||
Martins & Souto, 2012
|
||
</taxonomicName>
|
||
, no other true sclerodactylid has been recorded from the south-western Atlantic Ocean so far. In comparison with most species of the genus,
|
||
<taxonomicName id="B09582D84B56FFAFBEB0FEECFBE3FE62" box="[921,1103,368,392]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="species" species="pegi" status="sp. nov.">
|
||
<emphasis id="45E125494B56FFAFBEB0FEECFBE3FE62" box="[921,1103,368,392]" italics="true" pageId="4" pageNumber="587">Havelockia pegi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="45E125494B56FFAFB97CFEECFB02FE62" bold="true" box="[1109,1198,368,392]" pageId="4" pageNumber="587">
|
||
<taxonomicNameLabel id="5ED298324B56FFAFB97CFEECFB02FE62" box="[1109,1198,368,392]" pageId="4" pageNumber="587" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has scarce body wall ossicles, a circular end-plate, and rosettes in both the introvert and tentacles. These features in combination make it distinctive. There are only a couple of
|
||
<taxonomicName id="B09582D84B56FFAFBF49FE24FD4CFE3A" box="[608,736,440,464]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="45E125494B56FFAFBF49FE24FD4CFE3A" box="[608,736,440,464]" italics="true" pageId="4" pageNumber="587">Havelockia</emphasis>
|
||
</taxonomicName>
|
||
species with scarce or no body wall deposits. These are
|
||
<taxonomicName id="B09582D84B56FFAFB8A8FE25FE0BFE1E" authority="Sluiter, 1901" authorityName="Sluiter" authorityYear="1901" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="species" species="discolor">
|
||
<emphasis id="45E125494B56FFAFB8A8FE25FF5FFE1E" italics="true" pageId="4" pageNumber="587">H. discolor</emphasis>
|
||
(Sluiter, 1901)
|
||
</taxonomicName>
|
||
with no body wall deposits and
|
||
<taxonomicName id="B09582D84B56FFAFBE15FE41FB36FE1E" authority="Sluiter, 1901" authorityName="Sluiter" authorityYear="1901" box="[828,1178,476,500]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="species" species="conciliatrix">
|
||
<emphasis id="45E125494B56FFAFBE15FE41FC49FE1E" box="[828,997,476,500]" italics="true" pageId="4" pageNumber="587">H. conciliatrix</emphasis>
|
||
(Sluiter, 1901)
|
||
</taxonomicName>
|
||
with scarce deposits, restricted to the podia. Both are warm-water West Pacific species and differ from the new species in many features, e.g.
|
||
<taxonomicName id="B09582D84B56FFAFBDEFFDB9FEC5FDD6" box="[198,361,548,572]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="species" species="conciliatrix">
|
||
<emphasis id="45E125494B56FFAFBDEFFDB9FEC5FDD6" box="[198,361,548,572]" italics="true" pageId="4" pageNumber="587">H. conciliatrix</emphasis>
|
||
</taxonomicName>
|
||
has conical papillae and naked dorsal interradii and
|
||
<taxonomicName id="B09582D84B56FFAFBE91FDB9FB9AFDD6" box="[952,1078,548,572]" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="species" species="discolor">
|
||
<emphasis id="45E125494B56FFAFBE91FDB9FB9AFDD6" box="[952,1078,548,572]" italics="true" pageId="4" pageNumber="587">H. discolor</emphasis>
|
||
</taxonomicName>
|
||
has rods in the tentacles and no deposits in the body wall and tube feet. In comparison with the other sclerodactylids (sensu Panning, 1949, non Pawson & Fell, 1965) from South
|
||
<collectingCountry id="0F82B9CB4B56FFAFBF01FDF0FD27FD6E" box="[552,651,620,644]" name="United States of America" pageId="4" pageNumber="587">America</collectingCountry>
|
||
,
|
||
<taxonomicName id="B09582D84B56FFAFBFB0FDF1FCB4FD6E" box="[665,792,620,644]" class="Holothuroidea" family="Sclerodactylidae" genus="Coronatum" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="species" species="baiensis">
|
||
<emphasis id="45E125494B56FFAFBFB0FDF1FCB4FD6E" box="[665,792,620,644]" italics="true" pageId="4" pageNumber="587">C. baiensis</emphasis>
|
||
</taxonomicName>
|
||
has tables with oval, knobbed disc while
|
||
<taxonomicName id="B09582D84B56FFAFB9D6FDF0FF65FD42" class="Holothuroidea" family="Sclerodactylidae" genus="Pseudothyone" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="species" species="belli">
|
||
<emphasis id="45E125494B56FFAFB9D6FDF0FF65FD42" italics="true" pageId="4" pageNumber="587">Pseudothyone belli</emphasis>
|
||
</taxonomicName>
|
||
has knobbed buttons instead of tables in the body wall.
|
||
</paragraph>
|
||
<paragraph id="772AF95B4B56FFAFBDEEFD28FB84FC06" blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="587">
|
||
The new species might be mistaken to represent a
|
||
<emphasis id="45E125494B56FFAFBE37FD29FC0CFD26" box="[798,928,693,716]" italics="true" pageId="4" pageNumber="587">
|
||
<taxonomicName id="B09582D84B56FFAFBE37FD29FC30FD26" box="[798,924,693,716]" class="Holothuroidea" family="Phyllophoridae" genus="Pentamera" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="genus">Pentamera</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
notably
|
||
<emphasis id="45E125494B56FFAFB920FD29FB0CFD26" box="[1033,1184,692,716]" italics="true" pageId="4" pageNumber="587">
|
||
<taxonomicName id="B09582D84B56FFAFB920FD29FB30FD26" box="[1033,1180,692,716]" class="Holothuroidea" family="Phyllophoridae" genus="Pentamera" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="species" species="chiloensis">P. chiloensis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
which also occurs in Argentinian waters. However, the genus
|
||
<taxonomicName id="B09582D84B56FFAFBF4DFD45FD73FD1A" box="[612,735,729,752]" class="Holothuroidea" family="Phyllophoridae" genus="Pentamera" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="genus">
|
||
<emphasis id="45E125494B56FFAFBF4DFD45FD73FD1A" box="[612,735,729,752]" italics="true" pageId="4" pageNumber="587">Pentamera</emphasis>
|
||
</taxonomicName>
|
||
is well characterized by the ambulacral restriction of the tube feet, a different
|
||
<typeStatus id="A82E47F94B56FFAFBC78FD61FE2DFCFF" box="[337,385,765,789]" pageId="4" pageNumber="587">type</typeStatus>
|
||
of calcareous ring and well developed body wall ossicles, often accompanied by plates.
|
||
<taxonomicName id="B09582D84B56FFAFB8AEFD61FEABFCD2" class="Holothuroidea" family="Phyllophoridae" genus="Pentamera" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="species" species="chiloensis">
|
||
<emphasis id="45E125494B56FFAFB8AEFD61FEABFCD2" italics="true" pageId="4" pageNumber="587">P. chiloensis</emphasis>
|
||
</taxonomicName>
|
||
in particular, although lacking plate-like ossicles, has well-developed table deposits with numerous holes, table spires with 1–4 pillars and stellate end-plates (Pawson 1969).
|
||
<taxonomicName id="B09582D84B56FFAFBEC8FCD8FB44FCB6" box="[993,1256,836,860]" class="Holothuroidea" family="Sclerodactylidae" genus="Thandarum" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="species" species="hernandezi">
|
||
<emphasis id="45E125494B56FFAFBEC8FCD8FB44FCB6" box="[993,1256,836,860]" italics="true" pageId="4" pageNumber="587">Thandarum hernandezi</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="B09582D84B56FFAFB8A8FCD9FF64FC6A" class="Holothuroidea" family="Sclerodactylidae" genus="Havelockia" kingdom="Animalia" order="Dendrochirotida" pageId="4" pageNumber="587" phylum="Echinodermata" rank="species" species="pegi">
|
||
<emphasis id="45E125494B56FFAFB8A8FCD9FF64FC6A" italics="true" pageId="4" pageNumber="587">H. pegi</emphasis>
|
||
</taxonomicName>
|
||
in the ambulacral restriction of the tube feet and the presence of 4-pillared tables in the body wall. However, both species have a northern distribution, but restricted to the Rio de la Plata and are not found further north as the low salinity prevent their influx into the northern waters. Perhaps the same factor intervenes to prevent Brazilian species extending southwards (Tommasi et al. 1988b, Martinez & Brogger 2012).
|
||
</paragraph>
|
||
<paragraph id="772AF95B4B56FFAFBDEEFC64FCD4FB96" blockId="4.[151,1437,151,1149]" pageId="4" pageNumber="587">
|
||
More collections along the long Argentinian coastline will undoubtedly reveal more dendrochirotids as this order is notorious for its distribution in temperate waters of all the world’s oceans. In fact, the temperate waters of most continents or subcontinents of the southern hemisphere, including
|
||
<collectingCountry id="0F82B9CB4B56FFAFBECDFBDCFB2DFBBD" box="[996,1153,1087,1111]" name="New Zealand" pageId="4" pageNumber="587">New Zealand</collectingCountry>
|
||
, are well endowed with endemic dendrochirotids—so
|
||
<collectingCountry id="0F82B9CB4B56FFAFBCC1FBF8FDFBFB96" box="[488,599,1124,1148]" name="Argentina" pageId="4" pageNumber="587">Argentina</collectingCountry>
|
||
may not be an exception.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |