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<document id="1FE93D5A6D0CE7A952DF9ECCEA03F502" ID-CLB-Dataset="26038" ID-DOI="10.11646/zootaxa.4653.1.1" ID-GBIF-Dataset="2429a921-404f-482c-8f90-741676225442" ID-ISSN="1175-5326" ID-Zenodo-Dep="3366542" ID-ZooBank="FCDFD71B-D7AD-480D-A8CA-18297FD8D5EC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1565679017730" checkinUser="plazi" docAuthor="Meshram, Naresh M." docDate="2019" docId="CD4987FE4A64E015FF0CFB6CFB53F8A5" docLanguage="en" docName="zootaxa.4653.1.1.pdf" docOrigin="Zootaxa 4653 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Calodia deergha Meshram 2019, sp.nov." docType="treatment" docVersion="18" lastPageNumber="9" masterDocId="3170FF864A6CE01DFF9BFF9CFFA0FFEE" masterDocTitle="Leafhopper tribe Coelidiini (Hemiptera: Cicadellidae: Coelidiinae) of the Indian subcontinent" masterLastPageNumber="91" masterPageNumber="1" pageNumber="9" updateTime="1698688541026" updateUser="plazi">
<mods:mods id="579DFCD97C19FB4043B7F2286A28D225" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7C01B120088DDA7F56714C2557626D12">
<mods:title id="7E8D194A1AABBAB59A7C971AADCC0F39">Leafhopper tribe Coelidiini (Hemiptera: Cicadellidae: Coelidiinae) of the Indian subcontinent</mods:title>
</mods:titleInfo>
<mods:name id="518E899E8FBE2E5727AD7BCE14717F14" type="personal">
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<mods:namePart id="6DC332ADA7F9EA23DD9216B8A8685351">Meshram, Naresh M.</mods:namePart>
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<mods:date id="45E87D5549A632B4A4A389EB9772AAFE">2019</mods:date>
<mods:detail id="6531E35EFC8EDC3126CBB73CDE52941B" type="pubDate">
<mods:number id="0CFECFE250691D56BC14695B265F36AF">2019-08-12</mods:number>
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<mods:number id="56D03B6838F9A75B88A0BCC84DE2BE2B">4653</mods:number>
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<mods:classification id="6268BB4DB7B0DE6DC7C652CF524EC0B6">journal article</mods:classification>
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<mods:identifier id="6033F674ED3AF1986A798E2443491D9A" type="DOI">10.11646/zootaxa.4653.1.1</mods:identifier>
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<treatment id="CD4987FE4A64E015FF0CFB6CFB53F8A5" ID-DOI="http://doi.org/10.5281/zenodo.5191045" ID-GBIF-Taxon="159125981" ID-Zenodo-Dep="5191045" LSID="urn:lsid:plazi:treatment:CD4987FE4A64E015FF0CFB6CFB53F8A5" httpUri="http://treatment.plazi.org/id/CD4987FE4A64E015FF0CFB6CFB53F8A5" lastPageNumber="9" pageId="8" pageNumber="9">
<subSubSection id="0DFA65634A64E015FF0CFB6CFE1CFAE5" box="[151,444,1264,1291]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="455F36E84A64E015FF0CFB6CFE1CFAE5" blockId="8.[151,463,1264,1327]" box="[151,444,1264,1291]" pageId="8" pageNumber="9">
<heading id="1E1781844A64E015FF0CFB6CFE1CFAE5" bold="true" box="[151,444,1264,1291]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis id="7794EAFA4A64E015FF0CFB6CFE1CFAE5" bold="true" box="[151,444,1264,1291]" pageId="8" pageNumber="9">
<taxonomicName id="82E04D6B4A64E015FF0CFB6CFEFFFAE5" ID-CoL="8JQQK" authority="Meshram, 2019" authorityName="Meshram" authorityYear="2019" box="[151,351,1264,1291]" class="Insecta" family="Cicadellidae" genus="Calodia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="deergha" status="sp. nov.">
<emphasis id="7794EAFA4A64E015FF0CFB6CFEFFFAE5" bold="true" box="[151,351,1264,1291]" italics="true" pageId="8" pageNumber="9">Calodia deergha</emphasis>
</taxonomicName>
<taxonomicNameLabel id="6CA757814A64E015FEFDFB6DFE1CFAE5" box="[358,444,1265,1291]" pageId="8" pageNumber="9" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="0DFA65634A64E015FF0CFA89FB67F8ED" pageId="8" pageNumber="9" type="description">
<paragraph id="455F36E84A64E015FF0CFA89FE6FFAC1" blockId="8.[151,463,1264,1327]" box="[151,463,1300,1327]" pageId="8" pageNumber="9">
<figureCitation id="DDDB2A6D4A64E015FF0CFA89FF59FAC1" box="[151,249,1301,1327]" captionStart="FIGURES 1" captionStartId="36.[151,264,1896,1921]" captionTargetBox="[177,1410,169,1875]" captionTargetId="figure@36.[159,1428,166,1875]" captionTargetPageId="36" captionText="FIGURES 1 A–O. Habitus of Coelidiini species. A. Calodia deergha sp. nov., male; B. C. keralica sp. nov., male; C &amp; D. C. kodikanelensis Nielson, male and female; E. C. kumari sp. nov., male; F &amp; G. C. neofusca sp. nov., male and female; H &amp; I. C. ostenta (Distant), male and female; J. C. periyari sp. nov., male; K. C. subcrista Nielson, male; L. C. tridenta sp. nov., male; M. Glaberana acuta sp. nov., male; N &amp; O. G. khasiensis (Rao), male and female." httpUri="https://zenodo.org/record/3366546/files/figure.png" pageId="8" pageNumber="9">Figs.1 A</figureCitation>
,
<figureCitation id="DDDB2A6D4A64E015FE9FFA89FE8FFAC1" box="[260,303,1301,1327]" captionStart="FIGURES 4" captionStartId="39.[151,264,1897,1922]" captionTargetBox="[188,1409,181,1867]" captionTargetId="figure@39.[164,1423,166,1884]" captionTargetPageId="39" captionText="FIGURES 4 A–O. Habitus of Coelidiini species, lateral view. A. Calodia deergha sp. nov., male; B. C. keralica sp. nov., male; C &amp; D. C. kodikanelensis Nielson, male and female; E. C. kumari sp. nov., male; F &amp; G. C. neofusca sp. nov., male and female; H &amp; I. C. ostenta (Distant), male and female; J. C. periyari sp. nov., male; K. C. subcrista Nielson, male; L. C. tridenta sp. nov., male; M. Glaberana acuta sp. nov., male; N &amp; O. G. khasiensis (Rao), male and female." httpUri="https://zenodo.org/record/3366552/files/figure.png" pageId="8" pageNumber="9">4 A</figureCitation>
,
<figureCitation id="DDDB2A6D4A64E015FEA1FA89FEC4FAC1" box="[314,356,1301,1327]" captionStart="FIGURES 7" captionStartId="42.[151,264,1892,1917]" captionTargetBox="[175,1425,310,1744]" captionTargetId="figure@42.[151,1435,264,1750]" captionTargetPageId="42" captionText="FIGURES 7 A–P. Faces of Coelidiini species. A. Calodia deergha sp. nov., male; B. C. keralica sp. nov., male; C &amp; D. C. kodikanelensis Nielson, male and female; E. C. kumari sp. nov., male; F &amp; G. C. neofusca sp. nov., male and female; H &amp; I. C. ostenta (Distant), male and female; J. C. periyari sp. nov., male; K. C. subcrista Nielson, male; L. C. tridenta sp. nov., male; M. Glaberana acuta sp. nov., Male; N &amp; O. G. khasiensis (Rao), male and female. P. Glaberana purva sp. nov., male." httpUri="https://zenodo.org/record/3366558/files/figure.png" pageId="8" pageNumber="9">7 A</figureCitation>
,
<figureCitation id="DDDB2A6D4A64E015FEF4FA88FE2BFAC1" box="[367,395,1300,1327]" captionStart="FIGURES 15" captionStartId="50.[151,264,1968,1993]" captionTargetBox="[183,1406,166,1916]" captionTargetId="figure@50.[163,1425,166,1948]" captionTargetPageId="50" captionText="FIGURES 15 A–H. Calodia deergha sp. nov., male genitalia. A &amp; B. Aedeagus, dorsal view; C &amp; D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view." httpUri="https://zenodo.org/record/3366574/files/figure.png" pageId="8" pageNumber="9">15</figureCitation>
A–H.
</paragraph>
<paragraph id="455F36E84A64E015FF0CFAC1FB55FA75" blockId="8.[151,1437,1373,1867]" pageId="8" pageNumber="9">
<emphasis id="7794EAFA4A64E015FF0CFAC1FF71FA99" box="[151,209,1373,1399]" italics="true" pageId="8" pageNumber="9">
<collectingRegion id="8724F80A4A64E015FF0CFAC1FF71FA99" box="[151,209,1373,1399]" country="Maldives" name="Maale" pageId="8" pageNumber="9">Male</collectingRegion>
</emphasis>
. Head ochraceous, ocelli black; eyes brown; pronotum and mesonotum black with posterior margin paler; forewings ochraceous; thoracic sternites and pleurites largely black with broad ochraceous margins.
</paragraph>
<paragraph id="455F36E84A64E015FF5CFA39FB29F9A1" blockId="8.[151,1437,1373,1867]" pageId="8" pageNumber="9">
<emphasis id="7794EAFA4A64E015FF5CFA39FED5FA51" box="[199,373,1445,1471]" italics="true" pageId="8" pageNumber="9">
<collectingRegion id="8724F80A4A64E015FF5CFA39FEA1FA51" box="[199,257,1445,1471]" country="Maldives" name="Maale" pageId="8" pageNumber="9">Male</collectingRegion>
genitalia.
</emphasis>
Pygofer with digitate caudoventral process, without caudodorsal process, small lobe on caudodorsal margin (
<figureCitation id="DDDB2A6D4A64E015FEFBFA55FE72FA0D" box="[352,466,1480,1507]" captionStart="FIGURES 15" captionStartId="50.[151,264,1968,1993]" captionTargetBox="[183,1406,166,1916]" captionTargetId="figure@50.[163,1425,166,1948]" captionTargetPageId="50" captionText="FIGURES 15 A–H. Calodia deergha sp. nov., male genitalia. A &amp; B. Aedeagus, dorsal view; C &amp; D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view." httpUri="https://zenodo.org/record/3366574/files/figure.png" pageId="8" pageNumber="9">Fig. 15 H</figureCitation>
). Subgenital plate of uniform width, slightly broader medially and basally, with short setae (15 E). Style short, broad basally (
<figureCitation id="DDDB2A6D4A64E015FDC7FA71FD67F9E9" box="[604,711,1516,1543]" captionStart="FIGURES 15" captionStartId="50.[151,264,1968,1993]" captionTargetBox="[183,1406,166,1916]" captionTargetId="figure@50.[163,1425,166,1948]" captionTargetPageId="50" captionText="FIGURES 15 A–H. Calodia deergha sp. nov., male genitalia. A &amp; B. Aedeagus, dorsal view; C &amp; D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view." httpUri="https://zenodo.org/record/3366574/files/figure.png" pageId="8" pageNumber="9">Fig. 15 F</figureCitation>
). Connective as long as wide at base (
<figureCitation id="DDDB2A6D4A64E015FBEDFA71FB47F9E9" box="[1142,1255,1516,1543]" captionStart="FIGURES 15" captionStartId="50.[151,264,1968,1993]" captionTargetBox="[183,1406,166,1916]" captionTargetId="figure@50.[163,1425,166,1948]" captionTargetPageId="50" captionText="FIGURES 15 A–H. Calodia deergha sp. nov., male genitalia. A &amp; B. Aedeagus, dorsal view; C &amp; D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view." httpUri="https://zenodo.org/record/3366574/files/figure.png" pageId="8" pageNumber="9">Fig. 15 G</figureCitation>
). Aedeagus narrowed at apical half in lateral view, with two subapical, glabrous spine-like processes, distal process
<date id="315E10284A64E015FB60F98DFAB7F9C5" box="[1275,1303,1553,1579]" pageId="8" pageNumber="9">2x</date>
longer than the proximal process, gonopore arising distad of base of proximal process (
<figureCitation id="DDDB2A6D4A64E015FC7AF9A9FB9CF9A1" box="[993,1084,1588,1615]" captionStart="FIGURES 15" captionStartId="50.[151,264,1968,1993]" captionTargetBox="[183,1406,166,1916]" captionTargetId="figure@50.[163,1425,166,1948]" captionTargetPageId="50" captionText="FIGURES 15 A–H. Calodia deergha sp. nov., male genitalia. A &amp; B. Aedeagus, dorsal view; C &amp; D. Aedeagus, lateral view; E. Subgenital plate, dorsal view; F. Style, dorsal view; G. Connective, dorsal view; H. Pygofer, lateral view." httpUri="https://zenodo.org/record/3366574/files/figure.png" pageId="8" pageNumber="9">Figs. 15</figureCitation>
A–D).
</paragraph>
<paragraph id="455F36E84A64E015FF5CF9C6FEACF979" blockId="8.[151,1437,1373,1867]" pageId="8" pageNumber="9">
<emphasis id="7794EAFA4A64E015FF5CF9C6FED1F99D" box="[199,369,1626,1651]" italics="true" pageId="8" pageNumber="9">Measurements.</emphasis>
<collectingRegion id="8724F80A4A64E015FEE2F9C5FE13F99D" box="[377,435,1625,1651]" country="Maldives" name="Maale" pageId="8" pageNumber="9">Male</collectingRegion>
<quantity id="82189B0D4A64E015FE21F9C4FDB2F99D" box="[442,530,1624,1651]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="8" pageNumber="9" unit="mm" value="6.5">6.5 mm</quantity>
long,
<quantity id="82189B0D4A64E015FDC1F9C5FD11F99D" box="[602,689,1625,1651]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="8" pageNumber="9" unit="mm" value="1.9">1.9 mm</quantity>
wide across eyes and
<quantity id="82189B0D4A64E015FC2AF9C5FBA8F99D" box="[945,1032,1625,1651]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="8" pageNumber="9" unit="mm" value="2.1">2.1 mm</quantity>
wide across poserolateral angles of pronotum.
</paragraph>
<paragraph id="455F36E84A64E015FF5CF93DFF5CF931" blockId="8.[151,1437,1373,1867]" pageId="8" pageNumber="9">
<emphasis id="7794EAFA4A64E015FF5CF93DFE09F955" box="[199,425,1697,1723]" italics="true" pageId="8" pageNumber="9">Material examined.</emphasis>
<materialsCitation id="F5883CB54A64E015FE2DF93DFF5CF931" ID-GBIF-Occurrence="2332229189" collectingDate="1978-07-27" collectionCode="UASB" collectorName="S. Viraktamath Coll." country="India" location="Courtallam" pageId="8" pageNumber="9" specimenCount="1" specimenCount-male="1" stateProvince="Tamil Nadu" typeStatus="holotype">
<typeStatus id="9A5B884A4A64E015FE2DF93DFDEAF955" box="[438,586,1697,1723]" pageId="8" pageNumber="9" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="53E6FD614A64E015FDCCF93DFDCEF955" box="[599,622,1697,1723]" pageId="8" pageNumber="9" type="male">♂</specimenCount>
,
<collectingCountry id="3DF776784A64E015FD1BF93DFD74F955" box="[640,724,1697,1723]" name="India" pageId="8" pageNumber="9">INDIA</collectingCountry>
:
<collectingRegion id="8724F80A4A64E015FD7FF93DFCD3F955" box="[740,883,1697,1723]" country="India" name="Tamil Nadu" pageId="8" pageNumber="9">Tamil Nadu</collectingRegion>
:
<location id="403F60334A64E015FC1FF93DFBA5F955" LSID="urn:lsid:plazi:treatment:CD4987FE4A64E015FF0CFB6CFB53F8A5:403F60334A64E015FC1FF93DFBA5F955" box="[900,1029,1697,1723]" country="India" name="Courtallam" pageId="8" pageNumber="9" stateProvince="Tamil Nadu">Courtallam</location>
,
<date id="315E10284A64E015FB8DF93DFB38F955" box="[1046,1176,1697,1723]" pageId="8" pageNumber="9" value="1978-07-27">
<collectingDate id="211AE9C04A64E015FB8DF93DFB38F955" box="[1046,1176,1697,1723]" pageId="8" pageNumber="9" value="1978-07-27">27.vii.1978</collectingDate>
</date>
,
<collectorName id="E815533E4A64E015FB31F93DFA3DF955" box="[1194,1437,1697,1723]" pageId="8" pageNumber="9">S. Viraktamath Coll.</collectorName>
(
<collectionCode id="23F1AE2D4A64E015FF04F959FF50F931" box="[159,240,1733,1759]" country="India" httpUri="http://grbio.org/cool/yr7i-jnjm" name="University of Agricultural Sciences" pageId="8" pageNumber="9">UASB</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="455F36E84A64E015FF5CF975FB67F8ED" blockId="8.[151,1437,1373,1867]" box="[199,1223,1769,1795]" pageId="8" pageNumber="9">
<emphasis id="7794EAFA4A64E015FF5CF975FE9FF8ED" box="[199,319,1769,1795]" italics="true" pageId="8" pageNumber="9">Etymology</emphasis>
. The species name is Samskrit meaning long for processes of the aedeagal shaft.
</paragraph>
</subSubSection>
<subSubSection id="0DFA65634A64E015FF5CF891FB53F8A5" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="455F36E84A64E015FF5CF891FB53F8A5" blockId="8.[151,1437,1373,1867]" pageId="8" pageNumber="9">
<emphasis id="7794EAFA4A64E015FF5CF891FE9BF8C9" bold="true" box="[199,315,1805,1831]" pageId="8" pageNumber="9">Remarks.</emphasis>
<taxonomicName id="82E04D6B4A64E015FEDEF891FDA3F8C9" authority="Meshram, 2019" authorityName="Meshram" authorityYear="2019" box="[325,515,1805,1831]" class="Insecta" family="Cicadellidae" genus="Calodia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="deergha" status="sp. nov.">
<emphasis id="7794EAFA4A64E015FEDEF891FDA3F8C9" box="[325,515,1805,1831]" italics="true" pageId="8" pageNumber="9">Calodia deergha</emphasis>
</taxonomicName>
<emphasis id="7794EAFA4A64E015FD96F891FDC9F8C9" bold="true" box="[525,617,1805,1831]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="6CA757814A64E015FD96F891FDC9F8C9" box="[525,617,1805,1831]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles
<taxonomicName id="82E04D6B4A64E015FD77F891FCC0F8C9" baseAuthorityName="Distant" box="[748,864,1805,1831]" class="Insecta" family="Cicadellidae" genus="Calodia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ostenta">
<emphasis id="7794EAFA4A64E015FD77F891FCC0F8C9" box="[748,864,1805,1831]" italics="true" pageId="8" pageNumber="9">C. ostenta</emphasis>
</taxonomicName>
in general habitus but differs in having the distal process of the aedeagus
<date id="315E10284A64E015FE3CF8ADFE63F8A5" box="[423,451,1841,1867]" pageId="8" pageNumber="9">2x</date>
longer than the proximal process. They are equal in length in
<taxonomicName id="82E04D6B4A64E015FBE0F8ADFB4CF8A5" baseAuthorityName="Distant" box="[1147,1260,1841,1867]" class="Insecta" family="Cicadellidae" genus="Calodia" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ostenta">
<emphasis id="7794EAFA4A64E015FBE0F8ADFB4CF8A5" box="[1147,1260,1841,1867]" italics="true" pageId="8" pageNumber="9">C. ostenta</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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