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<document id="AC680409B3E5179CFCD3C12F0678E45F" ID-DOI="http://dx.doi.org/10.3897/zookeys.689.12021" ID-GBIF-Dataset="907713bd-c7f2-4f8e-a75e-b7f9c958865f" ID-PMC="PMC5672588" ID-Pensoft-Pub="1313-2970-689-1" ID-PubMed="29200920" ID-ZooBank="2F40BF4AD35F4CC697D5976EC201E652" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-689-1" ModsDocOrigin="ZooKeys 689" ModsDocTitle="A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)" checkinTime="1502744367222" checkinUser="pensoft" docAuthor="Lackner, Tomas &amp; Leschen, Richard A. B." docDate="2017" docId="5A66DE90C8206714228BBC441025C0FE" docLanguage="en" docName="ZooKeys 689: 1-263" docOrigin="ZooKeys 689" docSource="http://dx.doi.org/10.3897/zookeys.689.12021" docTitle="Hypocaccus (Hypocaccus) brasiliensis Paykull 1811" docType="treatment" docVersion="6" lastPageNumber="35" masterDocId="D302FFB6FFA66345FFBAFF89311EFFDE" masterDocTitle="A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)" masterLastPageNumber="263" masterPageNumber="1" pageNumber="33" updateTime="1701316739810" updateUser="plazi">
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<mods:title id="2C2C460AA9340BDF7F7CCA9FBA6B10B9">A monograph of the Australopacific Saprininae (Coleoptera, Histeridae)</mods:title>
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<mods:namePart id="66A36850EBEE72FB69E94164287F628D">Lackner, Tomas</mods:namePart>
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<treatment id="5A66DE90C8206714228BBC441025C0FE" ID-GBIF-Taxon="132648721" LSID="urn:lsid:plazi:treatment:5A66DE90C8206714228BBC441025C0FE" httpUri="http://treatment.plazi.org/id/5A66DE90C8206714228BBC441025C0FE" lastPageId="34" lastPageNumber="35" pageId="32" pageNumber="33">
<subSubSection id="4FBB2F54593E9FDE19C2CC8E5E1D7FD1" pageId="32" pageNumber="33" type="nomenclature">
<paragraph id="730F50E63767A12A970BF05D4ADABFB7" pageId="32" pageNumber="33">
<taxonomicName id="C6003ED07B77793AAC4967B2D2120B31" ID-CoL="7VHRV" authority="Paykull, 1811" authorityName="Paykull" authorityYear="1811" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Hypocaccus) brasiliensis" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="brasiliensis" subGenus="Hypocaccus">
<pageBreakToken id="1425F9FD4B509DBC4272C5F8A0012BB9" pageId="32" pageNumber="33" start="start">Hypocaccus</pageBreakToken>
(Hypocaccus) brasiliensis (Paykull, 1811)
</taxonomicName>
Figs 144, 145-153, 154-160, 755, 756, 760
</paragraph>
</subSubSection>
<subSubSection id="A6DE20F545810866B9FAFDBE7CF90604" pageId="32" pageNumber="33" type="reference_group">
<paragraph id="4784606C9AD0945700C90B16365F9D91" pageId="32" pageNumber="33">
<taxonomicName id="1387CFDFD3227AE728C91A7F12661D39" class="Insecta" family="Histeridae" genus="Hister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hister brasiliensis" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="brasiliensis">Hister brasiliensis</taxonomicName>
Paykull, 1811: 66.
</paragraph>
<paragraph id="0DEBC2C79CA2335B20872611F254F19C" pageId="32" pageNumber="33">
<taxonomicName id="C741D0605F00C5443C82B5D9E0DAA624" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus apricarius" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="apricarius">Saprinus apricarius</taxonomicName>
Erichson, 1834: 194 - Synonymized by Bickhardt (1910): 225.
</paragraph>
<paragraph id="A4D1560BB7FCDED91CE306F82637A60A" pageId="32" pageNumber="33">
<taxonomicName id="B504F5114C5FF0B04304E1DD37B24C70" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus bistrigifrons" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="bistrigifrons">Saprinus bistrigifrons</taxonomicName>
Marseul, 1855: 729 - Synonymized by
<bibRefCitation id="D9F81D75860BB9B68F64095F17A1C690" author="Mazur, S" journalOrPublisher="Entomologische Blaetter" pageId="217" pageNumber="218" pagination="28 - 32" title="Description of a new Pholioxenus from Morocco with additional histerid notes (Col., Histeridae)." volume="83" year="1987">Mazur (1987)</bibRefCitation>
: 32.
</paragraph>
<paragraph id="1931D1D92461383A63B6F0DCDDDFA92E" pageId="32" pageNumber="33">
<taxonomicName id="76617C56910BA25319F807B9928CE911" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus dentipes" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="dentipes">Saprinus dentipes</taxonomicName>
Marseul, 1855: 728 - Synonymized by
<bibRefCitation id="D448A32C0C311401742B4AFBD8DF8C48" author="Mazur, S" journalOrPublisher="Entomologische Blaetter" pageId="217" pageNumber="218" pagination="28 - 32" title="Description of a new Pholioxenus from Morocco with additional histerid notes (Col., Histeridae)." volume="83" year="1987">Mazur (1987)</bibRefCitation>
: 32.
</paragraph>
<paragraph id="9B24B4CC98F85D721A41EAB6C02F8FAB" pageId="32" pageNumber="33">
<taxonomicName id="E4292DBEFE7D72CD8818816C2B692C3E" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus piscarius" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="piscarius">Saprinus piscarius</taxonomicName>
Blackburn, 1903: 108 - Synonymized by
<bibRefCitation id="A045A40063293EC8AAE159826BC65CCB" author="Mazur, S" journalOrPublisher="Entomologische Blaetter" pageId="217" pageNumber="218" pagination="28 - 32" title="Description of a new Pholioxenus from Morocco with additional histerid notes (Col., Histeridae)." volume="83" year="1987">Mazur (1987)</bibRefCitation>
: 32.
</paragraph>
</subSubSection>
<subSubSection id="8E83A50D52BA2976A5B627146CBBDA1B" pageId="32" pageNumber="33" type="type locality">
<paragraph id="580971524B1A4C7B0319EAF88BCEE386" pageId="32" pageNumber="33">Type locality.</paragraph>
<paragraph id="8F67C57D4D80D7935AFB83D56DA0A4F2" pageId="32" pageNumber="33">Brazil.</paragraph>
</subSubSection>
<subSubSection id="BDDC59FBB4BF0BD50ED3B6FA9BBBA4FB" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="type material">
<paragraph id="B9069FF2626D6696A5611EBDC6F09554" pageId="32" pageNumber="33">Type material examined.</paragraph>
<paragraph id="F4E7EB3ACAB0858ACB46A25698D440E8" pageId="32" pageNumber="33">
<taxonomicName id="B35E525FCFB29D2FF98DE03F256590CE" class="Insecta" family="Histeridae" genus="Hister" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hister brasiliensis" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="brasiliensis">Hister brasiliensis</taxonomicName>
Paykull, 1811: Lectotype, ♀, designated by Dahlgren in 1968, together with extracted genitalia glued to a rectangular mounting card, right foreleg missing, right elytron missing, right metatibia broken off, glued next to the specimen, with the following labels:
<normalizedToken id="D28EF19BCB50152952A1FC2D145B7D3A" originalValue="“24”">&quot;24&quot;</normalizedToken>
(pencil-written); followed by: &quot;Uppsala Univ. Zool. Mus. / Gyllenhals saml. TYP nr. / 1153&quot; (red label, printed); followed by: &quot;LECTOTYP / HYPOCACCUS / BRASILIENSIS / PAYK. G. DAHL- / GREN 27.II.1968&quot; (written in black ink) (UUZM).
</paragraph>
<caption id="9FDA4546AD4614F59CC82A23A9B9622A" pageId="32" pageNumber="33">
<paragraph id="CE9B98131397472700D2DFAA5E891420" pageId="32" pageNumber="33">
Figures 145-153. 145
<taxonomicName id="1CCDB2F5D915CB2BE2BE2E579F79AFC0" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Hypocaccus) brasiliensis" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="brasiliensis" subGenus="Hypocaccus">Hypocaccus (Hypocaccus) brasiliensis</taxonomicName>
(Paykull, 1811) head, dorsal view 146 prosternum 147 mesoventrite 148 lateral disc of metaventrite + metepisternum 149 protibia, dorsal view 150 ditto, ventral view 151 propygidium + pygidium 152 metatibia, dorsal view 153 ditto, ventral view.
</paragraph>
</caption>
<paragraph id="9D1391536D47C3500B91CDAB35EB1932" pageId="32" pageNumber="33">
<taxonomicName id="487877972873808E25955D78E7A8BA19" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus apricarius" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="apricarius">Saprinus apricarius</taxonomicName>
Erichson, 1834: Lectotype, present designation: most likely a ♀, pinned, left hind leg missing, with the following labels: &quot;Aegypt / xxxxiv Er.(?)&quot; (dark green label, written); followed by:
<normalizedToken id="2EB4582701C876EC0B90E96E849B1725" originalValue="“49233”">&quot;49233&quot;</normalizedToken>
(printed); followed by: &quot;Hist. -Coll. (
<taxonomicName id="2C2919B78C45676581EAE309E68E0FF1" lsidName="" order="Coleoptera" pageId="32" pageNumber="33" rank="order">Coleoptera</taxonomicName>
) / Nr. 49233 /
<taxonomicName id="17A0C0033F79321A437B489742E1EFDC" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus apricarius" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="apricarius">Saprinus apricarius</taxonomicName>
Er. x / Aegypt., Ehrenberg / Zool. Mus. Berlin&quot; (black-framed label, printed); followed by: &quot;
<taxonomicName id="BBBE1C76A6E4BDB1948B59AA0CC18889" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/
<taxonomicName id="9CDC8225553D8662F448B79EDBA0286B" lsidName="apricarius" pageId="32" pageNumber="33" rank="species" species="apricarius">apricarius</taxonomicName>
/ Erichson, 1834 / LECTOTYPE 2014 / des. T. Lackner&quot; (red label, written) (ZMHUB). Paralectotypes, present designation: 1 spec., possibly a ♀, pinned, with the following label: &quot;Hist. -Coll. (
<taxonomicName id="EE4F2A771BE349C2496800A03D7D9BEF" lsidName="" order="Coleoptera" pageId="32" pageNumber="33" rank="order">Coleoptera</taxonomicName>
) / Nr. 49233 /
<taxonomicName id="D6CB44C54F67C306A64B23FF3ED9D3FE" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus apricarius" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="apricarius">Saprinus apricarius</taxonomicName>
Er. x / Aegypt., Ehrenberg / Zool. Mus. Berlin&quot; (black-framed label, printed); followed by: &quot;
<taxonomicName id="B3730B3CA2CCCD6E4401102536C98653" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/
<taxonomicName id="0C9A41595F57686C73DADCB5BCCC010B" lsidName="apricarius" pageId="32" pageNumber="33" rank="species" species="apricarius">apricarius</taxonomicName>
/ Erichson, 1834 / PARALECTOTYPE 2014 / des. T. Lackner&quot; (red label, written) (ZMHUB). 1 spec., possibly a ♀, pinned, with the following labels:
<normalizedToken id="C507213ED30DE934AE57019CE341A738" originalValue="“49233”">&quot;49233&quot;</normalizedToken>
(printed); followed by: &quot;Hist. -Coll. (
<taxonomicName id="D88B484D811D2D1B0F1B65A04C518C42" lsidName="" order="Coleoptera" pageId="32" pageNumber="33" rank="order">Coleoptera</taxonomicName>
) / Nr. 49233 /
<taxonomicName id="22554057E90DDA89E4A9DAED86C42D93" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus apricarius" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="apricarius">Saprinus apricarius</taxonomicName>
Er. x / Sicil, Dahl / Zool. Mus. Berlin&quot; (black-framed label, printed); followed by: &quot;
<taxonomicName id="7BA44132C350D3FD1E5C062996A19516" lsidName="apricarius" pageId="32" pageNumber="33" rank="species" species="apricarius">apricarius</taxonomicName>
/ Er. / Sicil. Dahl.&quot; (black-framed label, written); followed by: &quot;
<taxonomicName id="0355FE020C70E167F77CC3D2D81C2F87" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/
<taxonomicName id="3E90FAD98487D0DBCAD151746A63EC55" lsidName="apricarius" pageId="32" pageNumber="33" rank="species" species="apricarius">apricarius</taxonomicName>
/ Erichson, 1834 / PARALECTOTYPE 2014 / des. T. Lackner&quot; (red label, written) (ZMHUB). 1 spec., possibly a ♀, pinned, with the following labels:
<normalizedToken id="F261C18B15A847B93F5AF7F8A1EE57F6" originalValue="“47”">&quot;47&quot;</normalizedToken>
; followed by: &quot;Hist. -Coll. (
<taxonomicName id="CF7F1ECFE72855FC3E130FD722AA8196" lsidName="" order="Coleoptera" pageId="32" pageNumber="33" rank="order">Coleoptera</taxonomicName>
) / Nr. 49233 /
<taxonomicName id="BD89C4E1E8646B56D05DB9FA9445936B" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus apricarius" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="apricarius">Saprinus apricarius</taxonomicName>
Er. x / Sicil, Dahl / Zool. Mus. Berlin&quot; (black-framed label, printed); followed by: &quot;
<taxonomicName id="204D6CD56293A4839F7DE730A9ED5734" lsidName="apricarius" pageId="32" pageNumber="33" rank="species" species="apricarius">apricarius</taxonomicName>
/ Er. / Sicil. Dahl.&quot; (black-framed label, written); followed by: &quot;
<taxonomicName id="EEBCAA0AA8FC92320031B1A2E34CC259" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/
<taxonomicName id="2100A6BC8004D1F6F3CCBF35C1807B48" lsidName="apricarius" pageId="32" pageNumber="33" rank="species" species="apricarius">apricarius</taxonomicName>
/ Erichson, 1834 / PARALECTOTYPE 2014 / des. T. Lackner&quot; (red label, written) (ZMHUB). 1 spec., pinned, with the following labels:
<normalizedToken id="954059D09889FECBDEBC24E6A1E9A9E4" originalValue="“54”">&quot;54&quot;</normalizedToken>
; followed by: &quot;Hist. -Coll. (
<taxonomicName id="3DD5035838B5355FF627A76F36A8893B" lsidName="" order="Coleoptera" pageId="32" pageNumber="33" rank="order">Coleoptera</taxonomicName>
) / Nr. 49233 /
<taxonomicName id="FAF0DFE5E6FF08CC82910B207FD18EB9" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus apricarius" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="apricarius">Saprinus apricarius</taxonomicName>
Er. x / Sicil, Dahl / Zool. Mus. Berlin&quot; (black-framed label, printed); followed by: &quot;
<taxonomicName id="95B48C8E436AD955043B89E7ECA89F5C" lsidName="apricarius" pageId="32" pageNumber="33" rank="species" species="apricarius">apricarius</taxonomicName>
/ Er. / Sicil. Dahl.&quot; (black-framed label, written); followed by: &quot;
<taxonomicName id="6FA887DAEAD674566C6508EE657CAA0B" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/
<taxonomicName id="A1F54D1B00A0FD96B902D5C69EB7B6B8" lsidName="apricarius" pageId="32" pageNumber="33" rank="species" species="apricarius">apricarius</taxonomicName>
/ Erichson, 1834 / PARALECTOTYPE 2014 / des. T. Lackner&quot; (red label, written) (ZMHUB). 2 specs., pinned, with the following labels: &quot;Hist. -Coll. (
<taxonomicName id="00FC1C7F012479050329D9AA6C501FC0" lsidName="" order="Coleoptera" pageId="32" pageNumber="33" rank="order">Coleoptera</taxonomicName>
) / Nr. 49233 /
<taxonomicName id="FE179FBA9E501B8275CA1C73EA351251" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus apricarius" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="apricarius">Saprinus apricarius</taxonomicName>
Er. x / Sicil, Dahl / Zool. Mus. Berlin&quot; (black-framed label, printed); followed by: &quot;
<taxonomicName id="472AF48C1760294BABE518FF8B418EC1" lsidName="apricarius" pageId="32" pageNumber="33" rank="species" species="apricarius">apricarius</taxonomicName>
/ Er. / Sicil. Dahl.&quot; (black-framed label, written); followed by: &quot;
<taxonomicName id="3B19FCF7906B7B87BA08C88D7CD56993" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/
<taxonomicName id="30A4E46C0A00EE8FE7E03BCFB8166B37" lsidName="apricarius" pageId="32" pageNumber="33" rank="species" species="apricarius">apricarius</taxonomicName>
/ Erichson, 1834 / PARALECTOTYPE 2014 / des. T. Lackner&quot; (red label, written) (ZMHUB). This species has been described from both Sicily and Egypt, among the type series we designate a single female specimen from Egypt as lectotype to fix the taxonomic identity of the species.
</paragraph>
<paragraph id="664AC39A4B0AE582724B070F0D1E0790" pageId="32" pageNumber="33">
<taxonomicName id="7FA854EF327758855F606F61732E28EF" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus bistrigifrons" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="bistrigifrons">Saprinus bistrigifrons</taxonomicName>
Marseul, 1855: Lectotype, present designation: ♂, pinned, right mesotarsus missing, with following labels: tiny pink label, followed by: &quot;161 /
<taxonomicName id="822F5AD16010819E972A6421EC102654" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/
<taxonomicName id="83E046B8275AD9F9F9139C48CC92D6A8" lsidName="bistrigifrons" pageId="32" pageNumber="33" rank="species" species="bistrigifrons">bistrigifrons</taxonomicName>
/ m. / Mexique / illegible text&quot; (green, round label, written); followed by: &quot;MUSEUM PARIS / COLL. / DE MARSEUL 1890&quot; (green, printed label); followed by:
<normalizedToken id="2F8B814A8F4FD38C04C151189E54AADF" originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
(red-printed label); followed by: &quot;
<taxonomicName id="3C53617B94B6FE463ABE5101E8936FA4" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/
<taxonomicName id="47C53475BC93E385C38B1C03242081BA" lsidName="bistrigifrons" pageId="32" pageNumber="33" rank="species" species="bistrigifrons">bistrigifrons</taxonomicName>
/ Marseul, 1855 / LECTOTYPE 2014 / des. T. Lackner&quot; (red label, written) (MNHN). This species has been described from unknown number of specimens and the lectotype designation fixes the species identity.
</paragraph>
<paragraph id="9F90F3B6666AB6A6CC770F75825379CA" pageId="33" pageNumber="34">
<taxonomicName id="35730EB26E9952816CB234F095DAAFC9" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus dentipes" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="dentipes">
<pageBreakToken id="273EC6DED4CF27B4430E82DDAD48BC8C" pageId="33" pageNumber="34" start="start">Saprinus</pageBreakToken>
dentipes
</taxonomicName>
Marseul, 1855: Lectotype, present designation: probably a ♂, pinned, left metatarsus and right hind leg missing, with the following labels: &quot;160 /
<taxonomicName id="C20316D1E753F30C2CF9A78315B7F0E4" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/
<taxonomicName id="38254DD93AE734D57EF575B4D7E181BA" lsidName="dentipes" pageId="33" pageNumber="34" rank="species" species="dentipes">dentipes</taxonomicName>
/ illegible text / Mexique / illegible text&quot; (round, green label, written); followed by: &quot;MUSEUM PARIS / COLL. / DE MARSEUL 1890&quot; (green, printed label); followed by:
<normalizedToken id="DBFFE461B6D48D83DFB440F6E368040F" originalValue="“TYPE”">&quot;TYPE&quot;</normalizedToken>
(red-printed label); followed by: &quot;
<taxonomicName id="4739483D6DFCCA9ED754C20E3E599ED3" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus dentipes" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="dentipes">Saprinus dentipes</taxonomicName>
/ Marseul, 1855 / LECTOTYPE 2014 / des. T. Lackner&quot; (red label, written) (MNHN). This species has been described from unknown number of specimens and the lectotype designation fixes the species identity.
</paragraph>
<paragraph id="557D92B0C0991C4A828358F24406A778" pageId="33" pageNumber="34">
<taxonomicName id="86D6B8407DF0809F6B74696F624FBAA5" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus piscarius" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="piscarius">Saprinus piscarius</taxonomicName>
Blackburn, 1903: Cotype, sex unidentified, with printed label: &quot;Australia /
<normalizedToken id="29103D5630EEDEFA1E6FBD2D47BD2618" originalValue="Blackb’s">Blackb's</normalizedToken>
Coll&quot; (printed), followed by &quot;
<taxonomicName id="90DBDFBFD9D36900C7147C77FEF6BD55" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Hypocaccus</taxonomicName>
/
<taxonomicName id="3BD2BAC5018A97804D7D0F9E0C3A8E91" lsidName="vernulus" pageId="33" pageNumber="34" rank="species" species="vernulus">vernulus</taxonomicName>
Blackb&quot; (written), followed by &quot;
<taxonomicName id="202BC636D732EF4B740FB4E5915A8E42" lsidName="piscarius" pageId="33" pageNumber="34" rank="species" species="piscarius">piscarius</taxonomicName>
Bl. Cot. /
<taxonomicName id="BA970AEFC7180DC49D23A27E47242B95" lsidName="vernulus" pageId="33" pageNumber="34" rank="species" species="vernulus">vernulus</taxonomicName>
placed with Cotype /
<taxonomicName id="EF7784DA0CA93E17C243825C8C900AFA" lsidName="piscarius" pageId="33" pageNumber="34" rank="species" species="piscarius">piscarius</taxonomicName>
in the Bl. Coll&quot; (written), followed by &quot; SAMA Database / No. 25-029592&quot; (printed); 1 spec., &quot;
<taxonomicName id="0E405A2449F1B18CC1CF9DABAC7C36BA" lsidName="Hypo. piscarius" pageId="33" pageNumber="34" rank="species" species="piscarius">Hypo. piscarius</taxonomicName>
/ co-type&quot; (written), followed by &quot; SAMA Database / No. 25-029591 (printed)&quot;; 1 spec., &quot;
<taxonomicName id="74096A2D5564F1251A4A8F8566F7E4BC" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="genus">Saprinus</taxonomicName>
/
<taxonomicName id="9CCA12E04D283F884CB37B5F4D516E96" lsidName="piscarius" pageId="33" pageNumber="34" rank="species" species="piscarius">piscarius</taxonomicName>
/ Cotype&quot; (written), followed by &quot; SAMA Database / No. 25-029590&quot; (written) (SAMA). This species was described from 'near
<normalizedToken id="DB46F3665B23CF29C739BBE535DF1A9B" originalValue="Adelaide’">Adelaide'</normalizedToken>
, as well as 'Also from Victoria (Frankston, Mr. Kershaw)' (Blackburn, 1903: 108). Because of the ambiguous label data (see above) as well as no locality indication of the cotype we decided not to designate a lectotype of this species.
</paragraph>
<caption id="E11F802BC955F20418EE4E7F4DD55010" pageId="33" pageNumber="34">
<paragraph id="769B6A6C35CE539D9FF22E0306244F1A" pageId="33" pageNumber="34">
Figures 154-160. 154
<taxonomicName id="D97FA73F2F07911A3B1767F774905396" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Hypocaccus) brasiliensis" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="brasiliensis" subGenus="Hypocaccus">Hypocaccus (Hypocaccus) brasiliensis</taxonomicName>
(Paykull, 1811) male terminalia: 8th sternite + 8th tergite, ventral view 155 ditto, dorsal view 156 ditto, lateral view 157 male terminalia: 9th + 10th tergites, dorsal view; spiculum gastrale, ventral view 158 male terminalia: 9th + 10th tergites; spiculum gastrale, lateral view 159 male terminalia: aedeagus, dorsal view 160 ditto, lateral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="ACE657B8856C00A3372AE0571B6EB1C2" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="B9E99663D4711DA5255CFD7BE9912FC2" pageId="33" pageNumber="34">Additional material examined.</paragraph>
<paragraph id="D82935CB795F0113783DEA5903FC36C7" pageId="33" pageNumber="34">NEW CALEDONIA. 1 ♂ &amp; 4 specs., Isle Atire, 1.ix.1982, T. Lovegrove (ex dead white-capped noddy) (AMNZ).</paragraph>
<paragraph id="39A0B234DA98A5E3EB97F8BCBEC12123" pageId="34" pageNumber="35">
<pageBreakToken id="0EB715683C58EEBB28D317CB1F1CC3B9" pageId="34" pageNumber="35" start="start">New</pageBreakToken>
Guinea. New Britain: 12 specs., Ralum, F. Dahl S. (ZMHUB; 2 specs. in coll TLAN).
</paragraph>
<paragraph id="85E0F0C0C3184B25972D0DF4E57A2F31" pageId="34" pageNumber="35">
AUSTRALIA. New South Wales: 2 specs., Tomakin, 35.49S 150.11E, 25.xii.1988, W. Dressler (ex dead shark head) (ANIC); 1 spec., Colo River, Near Colo, 8.xii.1979, D.P. Came (on river sands) (ANIC); 2 specs., Sydney, K.K. Spence (ANIC); 1 spec., Sydney, Lea (SAMA). South Australia: 4 specs., Adelaide, Griffith (SAMA); 1 spec., Marino Rook: Spelhore, 4.-5.i.1978, G.
<normalizedToken id="DF47568F6B3D7DF684D7F749099D16C4" originalValue="Szelényi">Szelenyi</normalizedToken>
leg. (HMNH). Western Australia: 2 specs., Geraldton (SAMA).
</paragraph>
</subSubSection>
<subSubSection id="2C9F313909875304D5C449617BB76BF7" pageId="34" pageNumber="35" type="remarks">
<paragraph id="D2E956837B3B71552091EFEA186E69F3" pageId="34" pageNumber="35">Remarks.</paragraph>
<paragraph id="51EED619E4F1F72B21421793A8789D13" pageId="34" pageNumber="35">
<taxonomicName id="D747B360EB822FFAECFC50834F8CFF8B" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus dentipes" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="dentipes">Saprinus dentipes</taxonomicName>
has been synonymized with
<taxonomicName id="D109BA83E5C23B2B4F326D2424507C9D" lsidName="H. (H.) brasiliensis" pageId="34" pageNumber="35" rank="species" species="brasiliensis" subGenus="H.">H. (H.) brasiliensis</taxonomicName>
by
<bibRefCitation id="DA4A7489EAFEEBA4332818687B9F5EA1" author="Mazur, S" journalOrPublisher="Entomologische Blaetter" pageId="217" pageNumber="218" pagination="28 - 32" title="Description of a new Pholioxenus from Morocco with additional histerid notes (Col., Histeridae)." volume="83" year="1987">Mazur (1987)</bibRefCitation>
. However, upon inspecting its type specimen we conclude that it is not synonymous with
<taxonomicName id="3E55A9D5A116529C5C8848D8F2B636E2" lsidName="H. (H.) brasiliensis" pageId="34" pageNumber="35" rank="species" species="brasiliensis" subGenus="H.">H. (H.) brasiliensis</taxonomicName>
, but with the species
<taxonomicName id="B045B3F5BF82EAC7E4A31762B0EEB026" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Baeckmanniolus) gaudens" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="gaudens" subGenus="Baeckmanniolus">Hypocaccus (Baeckmanniolus) gaudens</taxonomicName>
(J.L. LeConte, 1851) instead. Thus
<taxonomicName id="18B47F72FAB5E38475F6060B38122363" class="Insecta" family="Histeridae" genus="Saprinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Saprinus dentipes" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="dentipes">Saprinus dentipes</taxonomicName>
Marseul, 1855 =
<taxonomicName id="EB087BF5192BB3923C775B8AA4280796" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (Baeckmanniolus) gaudens" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="gaudens" subGenus="Baeckmanniolus">Hypocaccus (Baeckmanniolus) gaudens</taxonomicName>
(J.L. LeConte, 1851) syn. n.
<taxonomicName id="8348D17CB3C91F8E8642E085E7899A6C" lsidName="H. (H.) brasiliensis" pageId="34" pageNumber="35" rank="species" species="brasiliensis" subGenus="H.">H. (H.) brasiliensis</taxonomicName>
is a widespread and variable species with variable dorsal punctation. The taxonomic status of both
<taxonomicName id="420F72669D4C3EA2784682500AC1B5EC" lsidName="H. (H.) brasiliensis" pageId="34" pageNumber="35" rank="species" species="brasiliensis" subGenus="H.">H. (H.) brasiliensis</taxonomicName>
and the following
<taxonomicName id="8D6BDADE590BF71BC7047ACD7D24C66E" lsidName="H. (H.) sinae" pageId="34" pageNumber="35" rank="species" species="sinae" subGenus="H.">H. (H.) sinae</taxonomicName>
requires further clarification, especially with respect to their rather similar genitalia and minimal differences in dorsal sculpture.
<taxonomicName id="34ECE3FEC1C606B1E68AFA12B3EEF584" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (H.) brasiliensis" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="brasiliensis" subGenus="H.">Hypocaccus (H.) brasiliensis</taxonomicName>
is above included in the key to the Australopacific species of
<taxonomicName id="5DE32A0A79E08CD13EAE14AB628FBC38" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">Hypocaccus</taxonomicName>
that separates it from the related
<taxonomicName id="0E1C77B61C98945237E3042C4FAB2468" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (H.) sinae" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="sinae" subGenus="H.">Hypocaccus (H.) sinae</taxonomicName>
. We chose not to fully re-describe
<taxonomicName id="017D59FA1C767A5452C89D96082E0279" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus (H.) brasiliensis" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="brasiliensis" subGenus="H.">Hypocaccus (H.) brasiliensis</taxonomicName>
here, leaving its re-description to the revision of the genus
<taxonomicName id="AEB9B63BC94B1591B0DC9A31E7C85BB1" class="Insecta" family="Histeridae" genus="Hypocaccus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Hypocaccus" order="Coleoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">Hypocaccus</taxonomicName>
. For the sake of the better species recognition, however, we decided to depict it here, including its male terminalia.
</paragraph>
</subSubSection>
</treatment>
</document>
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