treatments-xml/data/2D/3C/87/2D3C87B2C438642CFF1CFCF0FD4CF92A.xml

<document id="03B79E27EA4D895D933FFCD05F7A88DA" ID-CLB-Dataset="265951" ID-DOI="10.11646/zootaxa.5343.1.5" ID-GBIF-Dataset="b4f97f21-3643-41f6-8b55-cbbc32c1a46a" ID-ISSN="1175-5326" ID-Zenodo-Dep="8324403" ID-ZooBank="8379B120-3178-4321-963F-EC50AE1BA283" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1694069880281" checkinUser="plazi" docAuthor="Tu, Yuezheng, Zheng, Yuchen &amp; Liu, Xingyue" docDate="2023" docId="2D3C87B2C438642CFF1CFCF0FD4CF92A" docLanguage="en" docName="zootaxa.5343.1.5.pdf" docOrigin="Zootaxa 5343 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5343.1.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sialis kouwenkaii Tu &amp; Zheng &amp; Liu 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="87" masterDocId="D105FFCAC4396428FF8BFFACFFE4FFCE" masterDocTitle="A new species of the alderfly genus Sialis Latreille, 1802 (Megaloptera: Sialidae) from southeastern China" masterLastPageNumber="90" masterPageNumber="83" pageNumber="84" updateTime="1694071510902" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4C430D72715688CEF6F0AFA7D18239B5">A new species of the alderfly genus Sialis Latreille, 1802 (Megaloptera: Sialidae) from southeastern China</mods:title>
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<mods:namePart id="E95FC14D3156F5FA755AFD6C95DF9A7B">Tu, Yuezheng</mods:namePart>
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<mods:affiliation id="9D284E1F9408324CEC265674802A5C1E">Department of Entomology, China Agricultural University, Beijing 100193, China &amp; tuyuezheng @ 126. com; https: // orcid. org / 0000 - 0001 - 5825 - 3391</mods:affiliation>
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<mods:namePart id="DEC1684C8266FCE879BEC1F4724FA898">Zheng, Yuchen</mods:namePart>
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<mods:affiliation id="C6E7AA58C810ED53AD41270F8D4B9639">Department of Entomology, China Agricultural University, Beijing 100193, China &amp; zhengyuchenantlion @ gmail. com; https: // orcid. org / 0000 - 0003 - 2397 - 2008</mods:affiliation>
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<mods:namePart id="FA8553309AB0644528F1F468AAF5DFD8">Liu, Xingyue</mods:namePart>
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<subSubSection id="ED8F652FC4386429FF1CFCF0FE36FCB9" box="[151,466,860,887]" pageId="1" pageNumber="84" type="nomenclature">
<paragraph id="A52A36A4C4386429FF1CFCF0FE36FCB9" blockId="1.[151,466,860,923]" box="[151,466,860,887]" pageId="1" pageNumber="84">
<heading id="FE6281C8C4386429FF1CFCF0FE36FCB9" bold="true" box="[151,466,860,887]" fontSize="11" level="1" pageId="1" pageNumber="84" reason="1">
<emphasis id="97E1EAB6C4386429FF1CFCF0FE36FCB9" bold="true" box="[151,466,860,887]" pageId="1" pageNumber="84">
<taxonomicName id="62954D27C4386429FF1CFCF0FE89FCB9" authority="Tu &amp; Zheng &amp; Liu, 2023" authorityName="Tu &amp; Zheng &amp; Liu" authorityYear="2023" box="[151,365,860,887]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="1" pageNumber="84" phylum="Arthropoda" rank="species" species="kouwenkaii" status="sp. nov.">
<emphasis id="97E1EAB6C4386429FF1CFCF0FE89FCB9" bold="true" box="[151,365,860,887]" italics="true" pageId="1" pageNumber="84">Sialis kouwenkaii</emphasis>
</taxonomicName>
<taxonomicNameLabel id="8CD257CDC4386429FEFEFCF1FE36FCB9" box="[373,466,861,887]" pageId="1" pageNumber="84" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="ED8F652FC4386429FF1CFC2CFECAFC54" box="[151,302,896,923]" pageId="1" pageNumber="84" type="description">
<paragraph id="A52A36A4C4386429FF1CFC2CFECAFC54" blockId="1.[151,466,860,923]" box="[151,302,896,923]" pageId="1" pageNumber="84">
(
<figureCitation id="3DAE2A21C4386429FF14FC2CFF09FC55" box="[159,237,896,923]" captionStart="FIGURE 1" captionStartId="2.[151,250,1675,1699]" captionTargetBox="[160,1428,187,1641]" captionTargetId="figure-22@2.[151,1436,179,1649]" captionTargetPageId="2" captionText="FIGURE 1. Adult habitus of Sialis kouwenkaii sp. nov. A. male holotype; B. female paratype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8324405" httpUri="https://zenodo.org/record/8324405/files/figure.png" pageId="1" pageNumber="84">Figs. 1</figureCitation>
, 
<figureCitation id="3DAE2A21C4386429FF70FC2DFEEFFC55" box="[251,267,897,923]" captionStart="FIGURE 2" captionStartId="3.[151,250,988,1012]" captionTargetBox="[220,1368,185,961]" captionTargetId="figure-19@3.[216,1372,181,965]" captionTargetPageId="3" captionText="FIGURE 2. Genitalia of Sialis kouwenkaii sp. nov. A. male, dorsal view; B. male, ventral view; C. male sternum 9, ventral view; D. male, lateral view; E. male, caudal view; F. female, ventral view. e = ectoproct; gp8 = gonapophyses 8; gst9, 11 = gonostylus 9, 11; gx8, 9, 11 = gonocoxites 8, 9, 11; S7, 9 = sternum 7, 9; T9 = tergum 9. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8324407" httpUri="https://zenodo.org/record/8324407/files/figure.png" pageId="1" pageNumber="84">2</figureCitation>
, 
<figureCitation id="3DAE2A21C4386429FE9CFC2CFEC2FC54" box="[279,294,896,922]" captionStart="FIGURE 6" captionStartId="6.[151,250,1523,1547]" captionTargetBox="[158,1430,237,1499]" captionTargetId="figure-41@6.[158,1430,237,1499]" captionTargetPageId="6" captionText="FIGURE 6. Distribution map of S. kouwenkaii sp. nov. and its closely related species (hollow circle depicts type locality)." figureDoi="http://doi.org/10.5281/zenodo.8324419" httpUri="https://zenodo.org/record/8324419/files/figure.png" pageId="1" pageNumber="84">6</figureCitation>
)
</paragraph>
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<subSubSection id="ED8F652FC4386429FF1CFC64FB76FBBC" pageId="1" pageNumber="84" type="diagnosis">
<paragraph id="A52A36A4C4386429FF1CFC64FB76FBBC" blockId="1.[151,1437,968,1859]" pageId="1" pageNumber="84">
<emphasis id="97E1EAB6C4386429FF1CFC64FEF6FC2C" bold="true" box="[151,274,968,994]" pageId="1" pageNumber="84">Diagnosis.</emphasis>
Head entirely black; male compound eyes prominent; forewing with RP 4-branched; male sternum 9 broadly lingulate, slightly longer than wide, almost as long as tergum 9 plus ectoprocts, distinctly elevated laterally, forming a V-shaped incision posteriorly in ventral view; gonocoxite 9 with ventral portion medially produced; fused gonocoxites 11 proximally with a pair of narrow pterygoid lobes, distally with a pair of digitiform processes (= gonostyli 11), each of which bears a broad semicircular ventral lobe and is hook-like at tip.
</paragraph>
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<subSubSection id="ED8F652FC4386429FF4CFBD0FCDEF88D" pageId="1" pageNumber="84" type="description">
<paragraph id="A52A36A4C4386429FF4CFBD0FB1AFB59" blockId="1.[151,1437,968,1859]" box="[199,1278,1148,1175]" pageId="1" pageNumber="84">
<emphasis id="97E1EAB6C4386429FF4CFBD0FEBDFB58" bold="true" box="[199,345,1148,1174]" pageId="1" pageNumber="84">Description.</emphasis>
Male. Body length 10.0 mm; forewing length 12.0 mm, hindwing length 10.0 mm.
</paragraph>
<paragraph id="A52A36A4C4386429FF4CFB0CFBD6FB10" blockId="1.[151,1437,968,1859]" pageId="1" pageNumber="84">Head entirely black, immaculate. Vertex medially with a pair of longitudinal raised scars. Compound eyes black, moderately prominent. Antennae black, with black setae. Mouthparts black.</paragraph>
<paragraph id="A52A36A4C4386429FF4CFB44FBA8FA79" blockId="1.[151,1437,968,1859]" pageId="1" pageNumber="84">
Thorax black. Legs black, with dense blackish setae. Wings dark blackish brown. Forewing nearly three times as long as wide; costal area with 11 crossveins; sc-r present; 3 or 4 crossveins between RA and RP; RP 4-branched; MA bifurcate; MP proximally fused with CuA, distally separated into a simple anterior branch and a bifurcate posterior branch; CuA bifurcate; CuP simple; 1A and 3A simple; 2A bifurcate. Hind wing slightly shorter than forewing; costal area with 9 crossveins; 3 crossveins between RA and RP; RP 3 or 4-branched; MA bifurcate; both MP1 and MP2 bifurcate; CuA bifurcate; CuP simple; 1A, 2A and 3A simple (
<figureCitation id="3DAE2A21C4386429FC72FA30FBDEFA79" box="[1017,1082,1436,1463]" captionStart="FIGURE 1" captionStartId="2.[151,250,1675,1699]" captionTargetBox="[160,1428,187,1641]" captionTargetId="figure-22@2.[151,1436,179,1649]" captionTargetPageId="2" captionText="FIGURE 1. Adult habitus of Sialis kouwenkaii sp. nov. A. male holotype; B. female paratype. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.8324405" httpUri="https://zenodo.org/record/8324405/files/figure.png" pageId="1" pageNumber="84">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="A52A36A4C4386429FF4CFA6CFB1AF97D" blockId="1.[151,1437,968,1859]" pageId="1" pageNumber="84">
Abdomen black. Tergum 9 subtrapezoid with slightly arcuate lateral margins, anterior and posterior margins with shallow trapezoid incision (
<figureCitation id="3DAE2A21C4386429FD98FA48FD94FA31" box="[531,624,1508,1535]" captionStart="FIGURE 2" captionStartId="3.[151,250,988,1012]" captionTargetBox="[220,1368,185,961]" captionTargetId="figure-19@3.[216,1372,181,965]" captionTargetPageId="3" captionText="FIGURE 2. Genitalia of Sialis kouwenkaii sp. nov. A. male, dorsal view; B. male, ventral view; C. male sternum 9, ventral view; D. male, lateral view; E. male, caudal view; F. female, ventral view. e = ectoproct; gp8 = gonapophyses 8; gst9, 11 = gonostylus 9, 11; gx8, 9, 11 = gonocoxites 8, 9, 11; S7, 9 = sternum 7, 9; T9 = tergum 9. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8324407" httpUri="https://zenodo.org/record/8324407/files/figure.png" pageId="1" pageNumber="84">Fig. 2A</figureCitation>
). Sternum 9 broadly lingulate, nearly as long as wide, almost as long as tergum 9 plus ectoprocts, distinctly elevated laterally, forming a V-shaped incision posteriorly in ventral view (
<figureCitation id="3DAE2A21C4386429FF14F980FEC7F989" box="[159,291,1580,1607]" captionStart="FIGURE 2" captionStartId="3.[151,250,988,1012]" captionTargetBox="[220,1368,185,961]" captionTargetId="figure-19@3.[216,1372,181,965]" captionTargetPageId="3" captionText="FIGURE 2. Genitalia of Sialis kouwenkaii sp. nov. A. male, dorsal view; B. male, ventral view; C. male sternum 9, ventral view; D. male, lateral view; E. male, caudal view; F. female, ventral view. e = ectoproct; gp8 = gonapophyses 8; gst9, 11 = gonostylus 9, 11; gx8, 9, 11 = gonocoxites 8, 9, 11; S7, 9 = sternum 7, 9; T9 = tergum 9. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8324407" httpUri="https://zenodo.org/record/8324407/files/figure.png" pageId="1" pageNumber="84">Figs. 2B, C</figureCitation>
). Gonocoxites 9 broad, in lateral view subtrapezoid, slightly prominent posteroventrad, in caudal view distinctly produced medially on ventral portion (
<figureCitation id="3DAE2A21C4386429FD32F9FCFCDAF9A5" box="[697,830,1616,1643]" captionStart="FIGURE 2" captionStartId="3.[151,250,988,1012]" captionTargetBox="[220,1368,185,961]" captionTargetId="figure-19@3.[216,1372,181,965]" captionTargetPageId="3" captionText="FIGURE 2. Genitalia of Sialis kouwenkaii sp. nov. A. male, dorsal view; B. male, ventral view; C. male sternum 9, ventral view; D. male, lateral view; E. male, caudal view; F. female, ventral view. e = ectoproct; gp8 = gonapophyses 8; gst9, 11 = gonostylus 9, 11; gx8, 9, 11 = gonocoxites 8, 9, 11; S7, 9 = sternum 7, 9; T9 = tergum 9. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8324407" httpUri="https://zenodo.org/record/8324407/files/figure.png" pageId="1" pageNumber="84">Figs. 2D, E</figureCitation>
). Ectoprocts short, fused as a ring around anus. Fused gonocoxites 11 proximally with a pair of narrow pterygoid lobes, distally with a pair of digitiform processes (= gonostyli 11), each of which bears a broad semicircular ventral lobe and is hook-like at tip (
<figureCitation id="3DAE2A21C4386429FB11F934FB15F97D" box="[1178,1265,1688,1715]" captionStart="FIGURE 2" captionStartId="3.[151,250,988,1012]" captionTargetBox="[220,1368,185,961]" captionTargetId="figure-19@3.[216,1372,181,965]" captionTargetPageId="3" captionText="FIGURE 2. Genitalia of Sialis kouwenkaii sp. nov. A. male, dorsal view; B. male, ventral view; C. male sternum 9, ventral view; D. male, lateral view; E. male, caudal view; F. female, ventral view. e = ectoproct; gp8 = gonapophyses 8; gst9, 11 = gonostylus 9, 11; gx8, 9, 11 = gonocoxites 8, 9, 11; S7, 9 = sternum 7, 9; T9 = tergum 9. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8324407" httpUri="https://zenodo.org/record/8324407/files/figure.png" pageId="1" pageNumber="84">Fig. 2E</figureCitation>
).
</paragraph>
<paragraph id="A52A36A4C4386429FF4CF910FCDEF88D" blockId="1.[151,1437,968,1859]" pageId="1" pageNumber="84">
Female. Body length 11.0 mm; forewing length 13.0 mm, hindwing length 12.0 mm. Similar to male in general appearance. Sternum 7 subquadrate in ventral view, slightly longer than wide. Fused gonocoxites 8 broad in ventral view, nearly twice as wide as long. Gonapophyses 8 present as a subtrapezoid membranous lobe with anterior corners produced as a pair of tongue-shaped lobes (
<figureCitation id="3DAE2A21C4386429FD5CF884FCC9F88D" box="[727,813,1832,1859]" captionStart="FIGURE 2" captionStartId="3.[151,250,988,1012]" captionTargetBox="[220,1368,185,961]" captionTargetId="figure-19@3.[216,1372,181,965]" captionTargetPageId="3" captionText="FIGURE 2. Genitalia of Sialis kouwenkaii sp. nov. A. male, dorsal view; B. male, ventral view; C. male sternum 9, ventral view; D. male, lateral view; E. male, caudal view; F. female, ventral view. e = ectoproct; gp8 = gonapophyses 8; gst9, 11 = gonostylus 9, 11; gx8, 9, 11 = gonocoxites 8, 9, 11; S7, 9 = sternum 7, 9; T9 = tergum 9. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8324407" httpUri="https://zenodo.org/record/8324407/files/figure.png" pageId="1" pageNumber="84">Fig. 2F</figureCitation>
).
</paragraph>
</subSubSection>
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<paragraph id="A52A36A4C43B642AFF1CF927FACDF96D" blockId="2.[151,1321,1675,1700]" box="[151,1321,1675,1700]" pageId="2" pageNumber="85">
<emphasis id="97E1EAB6C43B642AFF1CF927FEF7F96D" bold="true" box="[151,275,1675,1699]" pageId="2" pageNumber="85">FIGURE 1.</emphasis>
Adult habitus of 
<taxonomicName id="62954D27C43B642AFE48F920FD91F96A" authority="Tu &amp; Zheng &amp; Liu, 2023" authorityName="Tu &amp; Zheng &amp; Liu" authorityYear="2023" box="[451,629,1676,1700]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="2" pageNumber="85" phylum="Arthropoda" rank="species" species="kouwenkaii" status="sp. nov.">
<emphasis id="97E1EAB6C43B642AFE48F920FD91F96A" box="[451,629,1676,1700]" italics="true" pageId="2" pageNumber="85">Sialis kouwenkaii</emphasis>
</taxonomicName>
<emphasis id="97E1EAB6C43B642AFDF0F920FD28F96A" bold="true" box="[635,716,1676,1700]" pageId="2" pageNumber="85">
<taxonomicNameLabel id="8CD257CDC43B642AFDF0F920FD28F96A" box="[635,716,1676,1700]" pageId="2" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. male holotype; B. female paratype. Scale bar = 1.0 mm.
</paragraph>
</caption>
<subSubSection id="ED8F652FC43B642AFF4CF95AFA62F896" pageId="2" pageNumber="85" type="materials_examined">
<paragraph id="A52A36A4C43B642AFF4CF95AFA62F896" blockId="2.[151,1436,1781,1880]" pageId="2" pageNumber="85">
<emphasis id="97E1EAB6C43B642AFF4CF95AFE97F8C1" bold="true" box="[199,371,1781,1808]" pageId="2" pageNumber="85">Type material.</emphasis>
<materialsCitation id="15FD3CF9C43B642AFEF3F95AFC20F8FA" collectingDate="2022-06-11" collectionCode="CAU" country="China" county="Hangzhou" elevation="1480" latitude="30.333334" location="Mt. Western Tianmu" longLatPrecision="1222" longitude="119.416664" municipality="Lin-an" pageId="2" pageNumber="85" specimenCount="1" stateProvince="Zhejiang" typeStatus="holotype">
<typeStatus id="7A2E8806C43B642AFEF3F95AFE04F8DE" box="[376,480,1782,1808]" pageId="2" pageNumber="85">Holotype</typeStatus>
J, 
<collectingCountry id="DD827634C43B642AFD8DF95AFDA8F8DE" box="[518,588,1782,1808]" name="China" pageId="2" pageNumber="85">China</collectingCountry>
, 
<collectingRegion id="6751F846C43B642AFDDEF95AFD58F8DE" box="[597,700,1782,1808]" country="China" name="Zhejiang" pageId="2" pageNumber="85">Zhejiang</collectingRegion>
, 
<collectingCounty id="4C4B4E28C43B642AFD4EF95AFCDFF8DE" box="[709,827,1782,1808]" pageId="2" pageNumber="85">Hangzhou</collectingCounty>
, 
<collectingMunicipality id="454EACDEC43B642AFCCFF95AFC76F8DE" box="[836,914,1782,1808]" pageId="2" pageNumber="85">Lin-an</collectingMunicipality>
, 
<location id="A04A607FC43B642AFC10F95AFB98F8DE" LSID="urn:lsid:plazi:treatment:2D3C87B2C438642CFF1CFCF0FD4CF92A:A04A607FC43B642AFC10F95AFB98F8DE" box="[923,1148,1782,1808]" country="China" county="Hangzhou" latitude="30.333334" longLatPrecision="1222" longitude="119.416664" municipality="Lin-an" name="Mt. Western Tianmu" pageId="2" pageNumber="85" stateProvince="Zhejiang">Mt. Western Tianmu</location>
[
<emphasis id="97E1EAB6C43B642AFB00F95BFB1FF8C0" bold="true" box="[1163,1275,1783,1806]" pageId="2" pageNumber="85">Wƙϐψ</emphasis>
], 
<location id="A04A607FC43B642AFA9BF95AFA78F8DE" LSID="urn:lsid:plazi:treatment:2D3C87B2C438642CFF1CFCF0FD4CF92A:A04A607FC43B642AFA9BF95AFA78F8DE" box="[1296,1436,1782,1808]" country="China" county="Hangzhou" latitude="30.333334" longLatPrecision="1222" longitude="119.416664" municipality="Lin-an" name="Xianrending" pageId="2" pageNumber="85" stateProvince="Zhejiang">Xianrending</location>
[
<emphasis id="97E1EAB6C43B642AFF2AF8B7FF11F8FC" bold="true" box="[161,245,1819,1842]" pageId="2" pageNumber="85">ŇKffl</emphasis>
], [
<geoCoordinate id="C0A15063C43B642AFE9AF8B6FE92F8FA" box="[273,374,1818,1844]" degrees="30" direction="north" minutes="20" orientation="latitude" pageId="2" pageNumber="85" precision="925" value="30.333334">30°20’N</geoCoordinate>
, 
<geoCoordinate id="C0A15063C43B642AFE0AF8B6FE14F8FB" box="[385,496,1818,1845]" degrees="119" direction="east" minutes="25" orientation="longitude" pageId="2" pageNumber="85" precision="925" value="119.416664">119°25’E</geoCoordinate>
], 
<quantity id="626D9B41C43B642AFD8AF8B6FDB2F8FA" box="[513,598,1818,1844]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="2" pageNumber="85" unit="m" value="1480.0">
<elevation id="2EB8D197C43B642AFD8AF8B6FDB2F8FA" box="[513,598,1818,1844]" metricMagnitude="3" metricUnit="m" metricValue="1.48" pageId="2" pageNumber="85" unit="m" value="1480.0">1480 m</elevation>
</quantity>
, 
<date id="D12B1064C43B642AFDEBF8B6FD3DF8FA" box="[608,729,1818,1844]" pageId="2" pageNumber="85" value="2022-06-11">
<collectingDate id="C16FE98CC43B642AFDEBF8B6FD3DF8FA" box="[608,729,1818,1844]" pageId="2" pageNumber="85" value="2022-06-11">11.vi.2022</collectingDate>
</date>
, Wenkai Kou (
<collectionCode id="C384AE61C43B642AFCF0F8B6FC58F8FA" box="[891,956,1818,1844]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="2" pageNumber="85">CAU</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="15FD3CF9C43B642AFC46F8B6FA66F896" collectingDate="2011-06-12" collectingDateMax="2011-06-16" collectingDateMin="2011-06-12" collectionCode="CAU" collectorName="Meicai Wei &amp; Gengyun Niu" country="China" county="Hangzhou" elevation="1100" latitude="30.333334" location="Mt. Western Tianmu" longLatPrecision="1222" longitude="119.433334" municipality="Lin-an" pageId="2" pageNumber="85" specimenCount="1" specimenCount-female="1" stateProvince="Zhejiang" typeStatus="paratype">
<typeStatus id="7A2E8806C43B642AFC46F8B6FBCBF8FA" box="[973,1071,1818,1844]" pageId="2" pageNumber="85">Paratype</typeStatus>
<specimenCount id="B393FD2DC43B642AFBBFF8B6FBB3F8FA" box="[1076,1111,1818,1844]" pageId="2" pageNumber="85" type="female">1♀</specimenCount>
, 
<collectingCountry id="DD827634C43B642AFBE8F8B6FB4DF8FA" box="[1123,1193,1818,1844]" name="China" pageId="2" pageNumber="85">China</collectingCountry>
, 
<collectingRegion id="6751F846C43B642AFB39F8B6FAFDF8FA" box="[1202,1305,1818,1844]" country="China" name="Zhejiang" pageId="2" pageNumber="85">Zhejiang</collectingRegion>
, 
<collectingCounty id="4C4B4E28C43B642AFAA9F8B6FA7CF8FA" box="[1314,1432,1818,1844]" pageId="2" pageNumber="85">Hangzhou</collectingCounty>
, 
<collectingMunicipality id="454EACDEC43B642AFF1CF892FF01F896" box="[151,229,1854,1880]" pageId="2" pageNumber="85">Lin-an</collectingMunicipality>
, 
<location id="A04A607FC43B642AFF7BF892FE3DF896" LSID="urn:lsid:plazi:treatment:2D3C87B2C438642CFF1CFCF0FD4CF92A:A04A607FC43B642AFF7BF892FE3DF896" box="[240,473,1854,1880]" country="China" county="Hangzhou" latitude="30.333334" longLatPrecision="1222" longitude="119.433334" municipality="Lin-an" name="Mt. Western Tianmu" pageId="2" pageNumber="85" stateProvince="Zhejiang">Mt. Western Tianmu</location>
, [
<geoCoordinate id="C0A15063C43B642AFE60F892FDB4F896" box="[491,592,1854,1880]" degrees="30" direction="north" minutes="20" orientation="latitude" pageId="2" pageNumber="85" precision="925" value="30.333334">30°20’N</geoCoordinate>
, 
<geoCoordinate id="C0A15063C43B642AFDD7F892FD2FF896" box="[604,715,1854,1880]" degrees="119" direction="east" minutes="26" orientation="longitude" pageId="2" pageNumber="85" precision="925" value="119.433334">119°26’E</geoCoordinate>
], 
<quantity id="626D9B41C43B642AFD55F892FCD0F896" box="[734,820,1854,1880]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="85" unit="m" value="1100.0">
<elevation id="2EB8D197C43B642AFD55F892FCD0F896" box="[734,820,1854,1880]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="85" unit="m" value="1100.0">1100 m</elevation>
</quantity>
, 
<date id="D12B1064C43B642AFCB4F892FC07F896" box="[831,995,1854,1880]" pageId="2" pageNumber="85" value="2011-06-12" valueMax="2011-06-16" valueMin="2011-06-12">
<collectingDate id="C16FE98CC43B642AFCB4F892FC07F896" box="[831,995,1854,1880]" pageId="2" pageNumber="85" value="2011-06-12" valueMax="2011-06-16" valueMin="2011-06-12">12–16.vi.2011</collectingDate>
</date>
, 
<collectorName id="08605372C43B642AFC65F892FB8AF896" box="[1006,1134,1854,1880]" pageId="2" pageNumber="85">Meicai Wei</collectorName>
&amp; 
<collectorName id="08605372C43B642AFB19F892FACEF896" box="[1170,1322,1854,1880]" pageId="2" pageNumber="85">Gengyun Niu</collectorName>
(
<collectionCode id="C384AE61C43B642AFAB2F892FA9EF896" box="[1337,1402,1854,1880]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33074" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="2" pageNumber="85">CAU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="F1EA662CC43A642BFF1CFC70FB0CFBF2" ID-DOI="http://doi.org/10.5281/zenodo.8324407" ID-Zenodo-Dep="8324407" httpUri="https://zenodo.org/record/8324407/files/figure.png" pageId="3" pageNumber="86" startId="3.[151,250,988,1012]" targetBox="[220,1368,185,961]" targetPageId="3" targetType="figure">
<paragraph id="A52A36A4C43A642BFF1CFC70FB0CFBF2" blockId="3.[151,1436,988,1085]" pageId="3" pageNumber="86">
<emphasis id="97E1EAB6C43A642BFF1CFC70FEF1FC3A" bold="true" box="[151,277,988,1012]" pageId="3" pageNumber="86">FIGURE 2.</emphasis>
Genitalia of 
<taxonomicName id="62954D27C43A642BFE14FC71FDB6FC3B" authority="Tu &amp; Zheng &amp; Liu, 2023" authorityName="Tu &amp; Zheng &amp; Liu" authorityYear="2023" box="[415,594,989,1013]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="kouwenkaii" status="sp. nov.">
<emphasis id="97E1EAB6C43A642BFE14FC71FDB6FC3B" box="[415,594,989,1013]" italics="true" pageId="3" pageNumber="86">Sialis kouwenkaii</emphasis>
</taxonomicName>
<emphasis id="97E1EAB6C43A642BFDD0FC71FD49FC3B" bold="true" box="[603,685,989,1013]" pageId="3" pageNumber="86">
<taxonomicNameLabel id="8CD257CDC43A642BFDD0FC71FD49FC3B" box="[603,685,989,1013]" pageId="3" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
A. male, dorsal view; B. male, ventral view; C. male sternum 9, ventral view; D. male, lateral view; E. male, caudal view; F. female, ventral view. e = ectoproct; gp8 = gonapophyses 8; gst9, 11 = gonostylus 9, 11; gx8, 9, 11 = gonocoxites 8, 9, 11; S7, 9 = sternum 7, 9; T9 = tergum 9. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="F1EA662CC43A642BFF1CF83AFD05F838" ID-DOI="http://doi.org/10.5281/zenodo.8324409" ID-Zenodo-Dep="8324409" httpUri="https://zenodo.org/record/8324409/files/figure.png" pageId="3" pageNumber="86" startId="3.[151,250,1942,1966]" targetBox="[218,1368,1112,1915]" targetPageId="3" targetType="figure">
<paragraph id="A52A36A4C43A642BFF1CF83AFD05F838" blockId="3.[151,1437,1942,2038]" pageId="3" pageNumber="86">
<emphasis id="97E1EAB6C43A642BFF1CF83AFEF0F860" bold="true" box="[151,276,1942,1966]" pageId="3" pageNumber="86">FIGURE 3.</emphasis>
Male genitalia of closely related species of 
<taxonomicName id="62954D27C43A642BFD51F83AFC69F860" authority="Tu &amp; Zheng &amp; Liu, 2023" authorityName="Tu &amp; Zheng &amp; Liu" authorityYear="2023" box="[730,909,1942,1966]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="kouwenkaii" status="sp. nov.">
<emphasis id="97E1EAB6C43A642BFD51F83AFC69F860" box="[730,909,1942,1966]" italics="true" pageId="3" pageNumber="86">Sialis kouwenkaii</emphasis>
</taxonomicName>
<emphasis id="97E1EAB6C43A642BFC1EF83AFC02F860" bold="true" box="[917,998,1942,1966]" pageId="3" pageNumber="86">
<taxonomicNameLabel id="8CD257CDC43A642BFC1EF83AFC02F860" box="[917,998,1942,1966]" pageId="3" pageNumber="86" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, ventral view. A. 
<taxonomicName id="62954D27C43A642BFB15F83BFECCF81C" authority="Liu, Hayashi &amp; Yang, 2015" authorityName="Liu, Hayashi &amp; Yang" authorityYear="2015" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="97E1EAB6C43A642BFB15F83BFAF5F860" box="[1182,1297,1942,1966]" italics="true" pageId="3" pageNumber="86">S. australis</emphasis>
Liu, Hayashi &amp; Yang, 2015
</taxonomicName>
; B. 
<taxonomicName id="62954D27C43A642BFEDBF817FE58F81C" authorityName="Okamoto" authorityYear="1910" box="[336,444,1979,2002]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="jezoensi">
<emphasis id="97E1EAB6C43A642BFEDBF817FE58F81C" box="[336,444,1979,2002]" italics="true" pageId="3" pageNumber="86">S. jezoensi</emphasis>
</taxonomicName>
s 
<bibRefCitation id="C1044B55C43A642BFE47F816FD89F81C" author="Okamoto, H." box="[460,621,1978,2002]" pageId="3" pageNumber="86" pagination="255 - 263" refId="ref3057" refString="Okamoto, H. (1910) Die Sialiden Japans. Wiener Entomologische Zeitung, 29, 255 - 263. https: // doi. org / 10.5962 / bhl. part. 23335" type="journal article" year="1910">Okamoto, 1910</bibRefCitation>
; C. 
<taxonomicName id="62954D27C43A642BFD1EF817FBEEF81C" authority="Liu, Hayashi &amp; Yang, 2015" authorityName="Liu, Hayashi &amp; Yang" authorityYear="2015" box="[661,1034,1978,2003]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="jiyuni">
<emphasis id="97E1EAB6C43A642BFD1EF817FD0DF81C" box="[661,745,1979,2002]" italics="true" pageId="3" pageNumber="86">S. jiyuni</emphasis>
Liu, Hayashi &amp; Yang, 2015
</taxonomicName>
; D. 
<taxonomicName id="62954D27C43A642BFBB8F817FA86F81C" authority="Okamoto, 1910" authorityName="Okamoto" authorityYear="1910" box="[1075,1378,1978,2002]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="kumejimae">
<emphasis id="97E1EAB6C43A642BFBB8F817FB5FF81C" box="[1075,1211,1978,2002]" italics="true" pageId="3" pageNumber="86">S. kumejimae</emphasis>
<bibRefCitation id="C1044B55C43A642BFB4AF816FA86F81C" author="Okamoto, H." box="[1217,1378,1978,2002]" pageId="3" pageNumber="86" pagination="255 - 263" refId="ref3057" refString="Okamoto, H. (1910) Die Sialiden Japans. Wiener Entomologische Zeitung, 29, 255 - 263. https: // doi. org / 10.5962 / bhl. part. 23335" type="journal article" year="1910">Okamoto, 1910</bibRefCitation>
</taxonomicName>
; E. 
<taxonomicName id="62954D27C43A642BFA02F817FDEFF838" authority="(Hayashi &amp; Suda, 1995)" baseAuthorityName="Hayashi &amp; Suda" baseAuthorityYear="1995" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="3" pageNumber="86" phylum="Arthropoda" rank="species" species="kuwayamai">
<emphasis id="97E1EAB6C43A642BFA02F817FEEEF838" italics="true" pageId="3" pageNumber="86">S. kuwayamai</emphasis>
(
<bibRefCitation id="C1044B55C43A642BFE93F872FDE6F838" author="Hayashi, F. &amp; Suda, S." box="[280,514,2014,2038]" pageId="3" pageNumber="86" pagination="1 - 15" refId="ref2531" refString="Hayashi, F. &amp; Suda, S. (1995) Sialidae (Megaloptera) from Japan. Aquatic Insects, 17, 1 - 15. https: // doi. org / 10.1080 / 01650429509361564" type="journal article" year="1995">Hayashi &amp; Suda, 1995</bibRefCitation>
)
</taxonomicName>
. Scale bar = 0.5 mm.
</paragraph>
</caption>
<caption id="F1EA662CC43D642CFF1CFBEFFEC3FB6C" ID-DOI="http://doi.org/10.5281/zenodo.8324413" ID-Zenodo-Dep="8324413" httpUri="https://zenodo.org/record/8324413/files/figure.png" pageId="4" pageNumber="87" startId="4.[151,250,1091,1115]" targetBox="[224,1363,182,1060]" targetPageId="4" targetType="figure">
<paragraph id="A52A36A4C43D642CFF1CFBEFFEC3FB6C" blockId="4.[151,1437,1091,1187]" pageId="4" pageNumber="87">
<emphasis id="97E1EAB6C43D642CFF1CFBEFFEF1FB95" bold="true" box="[151,277,1091,1115]" pageId="4" pageNumber="87">FIGURE 4.</emphasis>
Male genitalia of 
<taxonomicName id="62954D27C43D642CFE5EFBEFFDA6FB94" box="[469,578,1091,1115]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="97E1EAB6C43D642CFE5EFBEFFDE9FB95" box="[469,525,1091,1115]" italics="true" pageId="4" pageNumber="87">Sialis</emphasis>
spp.
</taxonomicName>
, lateral view. A. 
<taxonomicName id="62954D27C43D642CFD79FBEFFB6AFB95" authority="Liu, Hayashi &amp; Yang, 2015" authorityName="Liu, Hayashi &amp; Yang" authorityYear="2015" box="[754,1166,1091,1115]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="97E1EAB6C43D642CFD79FBEFFC82FB95" box="[754,870,1091,1115]" italics="true" pageId="4" pageNumber="87">S. australis</emphasis>
Liu, Hayashi &amp; Yang, 2015
</taxonomicName>
; B. 
<taxonomicName id="62954D27C43D642CFB32FBEFFAC2FB94" authorityName="Okamoto" authorityYear="1910" box="[1209,1318,1091,1114]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="jezoensi">
<emphasis id="97E1EAB6C43D642CFB32FBEFFAC2FB94" box="[1209,1318,1091,1114]" italics="true" pageId="4" pageNumber="87">S. jezoensi</emphasis>
</taxonomicName>
s 
<bibRefCitation id="C1044B55C43D642CFAB3FBEFFF29FBB1" author="Okamoto, H." pageId="4" pageNumber="87" pagination="255 - 263" refId="ref3057" refString="Okamoto, H. (1910) Die Sialiden Japans. Wiener Entomologische Zeitung, 29, 255 - 263. https: // doi. org / 10.5962 / bhl. part. 23335" type="journal article" year="1910">Okamoto, 1910</bibRefCitation>
; C. 
<taxonomicName id="62954D27C43D642CFF78FBCBFDBBFBB1" authority="Liu, Hayashi &amp; Yang, 2015" authorityName="Liu, Hayashi &amp; Yang" authorityYear="2015" box="[243,607,1127,1151]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="jiyuni">
<emphasis id="97E1EAB6C43D642CFF78FBCBFEA1FBB0" box="[243,325,1127,1150]" italics="true" pageId="4" pageNumber="87">S. jiyuni</emphasis>
Liu, Hayashi &amp; Yang, 2015
</taxonomicName>
; D. 
<taxonomicName id="62954D27C43D642CFD0DFBCBFC55FBB1" authority="Okamoto, 1910" authorityName="Okamoto" authorityYear="1910" box="[646,945,1127,1151]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="kumejimae">
<emphasis id="97E1EAB6C43D642CFD0DFBCBFCE8FBB1" box="[646,780,1127,1151]" italics="true" pageId="4" pageNumber="87">S. kumejimae</emphasis>
<bibRefCitation id="C1044B55C43D642CFC9AFBCBFC55FBB1" author="Okamoto, H." box="[785,945,1127,1151]" pageId="4" pageNumber="87" pagination="255 - 263" refId="ref3057" refString="Okamoto, H. (1910) Die Sialiden Japans. Wiener Entomologische Zeitung, 29, 255 - 263. https: // doi. org / 10.5962 / bhl. part. 23335" type="journal article" year="1910">Okamoto, 1910</bibRefCitation>
</taxonomicName>
; E. 
<taxonomicName id="62954D27C43D642CFC5DFBCBFAB9FBB1" authority="(Hayashi &amp; Suda, 1995)" baseAuthorityName="Hayashi &amp; Suda" baseAuthorityYear="1995" box="[982,1373,1127,1151]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="kuwayamai">
<emphasis id="97E1EAB6C43D642CFC5DFBCBFB85FBB1" box="[982,1121,1127,1151]" italics="true" pageId="4" pageNumber="87">S. kuwayamai</emphasis>
(
<bibRefCitation id="C1044B55C43D642CFBE6FBCBFAB1FBB1" author="Hayashi, F. &amp; Suda, S." box="[1133,1365,1127,1151]" pageId="4" pageNumber="87" pagination="1 - 15" refId="ref2531" refString="Hayashi, F. &amp; Suda, S. (1995) Sialidae (Megaloptera) from Japan. Aquatic Insects, 17, 1 - 15. https: // doi. org / 10.1080 / 01650429509361564" type="journal article" year="1995">Hayashi &amp; Suda, 1995</bibRefCitation>
)
</taxonomicName>
. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="ED8F652FC43D642CFF4CFB59FEB1FAFD" pageId="4" pageNumber="87" type="etymology">
<paragraph id="A52A36A4C43D642CFF4CFB59FEB1FAFD" blockId="4.[151,1436,1269,1764]" pageId="4" pageNumber="87">
<emphasis id="97E1EAB6C43D642CFF4CFB59FEAAFAC1" bold="true" box="[199,334,1269,1295]" pageId="4" pageNumber="87">Etymology.</emphasis>
The new species is dedicated to Mr. Wenkai Kou [
<emphasis id="97E1EAB6C43D642CFC1DFB5AFC0EFAC3" bold="true" box="[918,1002,1270,1293]" pageId="4" pageNumber="87">ĖẊffi</emphasis>
], who collected the male specimen of this new species.
</paragraph>
</subSubSection>
<subSubSection id="ED8F652FC43D642CFF4CFA91FDCBFA99" box="[199,559,1341,1367]" pageId="4" pageNumber="87" type="distribution">
<paragraph id="A52A36A4C43D642CFF4CFA91FDCBFA99" blockId="4.[151,1436,1269,1764]" box="[199,559,1341,1367]" pageId="4" pageNumber="87">
<emphasis id="97E1EAB6C43D642CFF4CFA91FE85FA99" bold="true" box="[199,353,1341,1367]" pageId="4" pageNumber="87">Distribution.</emphasis>
<collectingCountry id="DD827634C43D642CFEECFA91FE4EFA99" box="[359,426,1341,1367]" name="China" pageId="4" pageNumber="87">China</collectingCountry>
(
<collectingRegion id="6751F846C43D642CFE32FA91FDC0FA99" box="[441,548,1341,1367]" country="China" name="Zhejiang" pageId="4" pageNumber="87">Zhejiang</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="ED8F652FC43D642CFF4CFACDFD4CF92A" pageId="4" pageNumber="87" type="discussion">
<paragraph id="A52A36A4C43D642CFF4CFACDFD4CF92A" blockId="4.[151,1436,1269,1764]" pageId="4" pageNumber="87">
<emphasis id="97E1EAB6C43D642CFF4CFACDFEDFFAB5" bold="true" box="[199,315,1377,1403]" pageId="4" pageNumber="87">Remarks.</emphasis>
The new species appears to be similar to 
<taxonomicName id="62954D27C43D642CFCA1FACDFB13FAB2" authority="Liu, Hayashi &amp; Yang, 2015" authorityName="Liu, Hayashi &amp; Yang" authorityYear="2015" box="[810,1271,1377,1404]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="97E1EAB6C43D642CFCA1FACDFC4FFAB5" box="[810,939,1377,1403]" italics="true" pageId="4" pageNumber="87">S. australis</emphasis>
Liu, Hayashi &amp; Yang, 2015
</taxonomicName>
, 
<taxonomicName id="62954D27C43D642CFA8EFACDFE7EFA6E" authority="Liu, Hayashi &amp; Yang, 2015" authorityName="Liu, Hayashi &amp; Yang" authorityYear="2015" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="jiyuni">
<emphasis id="97E1EAB6C43D642CFA8EFACDFA81FAB5" box="[1285,1381,1377,1403]" italics="true" pageId="4" pageNumber="87">S. jiyuni</emphasis>
Liu, Hayashi &amp; Yang, 2015
</taxonomicName>
, 
<taxonomicName id="62954D27C43D642CFE2DFA29FD04FA51" authority="Okamoto, 1910" authorityName="Okamoto" authorityYear="1910" box="[422,736,1413,1439]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="jezoensis">
<emphasis id="97E1EAB6C43D642CFE2DFA29FDC3FA51" box="[422,551,1413,1439]" italics="true" pageId="4" pageNumber="87">S. jezoensis</emphasis>
<bibRefCitation id="C1044B55C43D642CFDA4FA29FD04FA51" author="Okamoto, H." box="[559,736,1413,1439]" pageId="4" pageNumber="87" pagination="255 - 263" refId="ref3057" refString="Okamoto, H. (1910) Die Sialiden Japans. Wiener Entomologische Zeitung, 29, 255 - 263. https: // doi. org / 10.5962 / bhl. part. 23335" type="journal article" year="1910">Okamoto, 1910</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="62954D27C43D642CFD67FA29FBDFFA51" authority="Okamoto, 1910" authorityName="Okamoto" authorityYear="1910" box="[748,1083,1413,1439]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="kumejimae">
<emphasis id="97E1EAB6C43D642CFD67FA29FC65FA51" box="[748,897,1413,1439]" italics="true" pageId="4" pageNumber="87">S. kumejimae</emphasis>
<bibRefCitation id="C1044B55C43D642CFC02FA29FBDFFA51" author="Okamoto, H." box="[905,1083,1413,1439]" pageId="4" pageNumber="87" pagination="255 - 263" refId="ref3057" refString="Okamoto, H. (1910) Die Sialiden Japans. Wiener Entomologische Zeitung, 29, 255 - 263. https: // doi. org / 10.5962 / bhl. part. 23335" type="journal article" year="1910">Okamoto, 1910</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="62954D27C43D642CFBFDFA29FEFAFA0A" authority="(Hayashi &amp; Suda, 1995)" baseAuthorityName="Hayashi &amp; Suda" baseAuthorityYear="1995" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="kuwayamai">
<emphasis id="97E1EAB6C43D642CFBFDFA29FAF7FA51" box="[1142,1299,1413,1439]" italics="true" pageId="4" pageNumber="87">S. kuwayamai</emphasis>
(
<bibRefCitation id="C1044B55C43D642CFAAAFA29FEF2FA0A" author="Hayashi, F. &amp; Suda, S." pageId="4" pageNumber="87" pagination="1 - 15" refId="ref2531" refString="Hayashi, F. &amp; Suda, S. (1995) Sialidae (Megaloptera) from Japan. Aquatic Insects, 17, 1 - 15. https: // doi. org / 10.1080 / 01650429509361564" type="journal article" year="1995">Hayashi &amp; Suda, 1995</bibRefCitation>
)
</taxonomicName>
by having the diagnostic characters of the formerly recognized 
<taxonomicName id="62954D27C43D642CFC6DFA05FB66FA0D" authorityName="Kuwayama" authorityYear="1962" box="[998,1154,1449,1475]" class="Insecta" family="Sialidae" genus="Nipponosialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="97E1EAB6C43D642CFC6DFA05FB66FA0D" box="[998,1154,1449,1475]" italics="true" pageId="4" pageNumber="87">Nipponosialis</emphasis>
</taxonomicName>
, now considered a junior synonym of 
<taxonomicName id="62954D27C43D642CFEACFA61FDDFFA26" authority="(Liu et al. 2015 a)" baseAuthorityName="Liu" baseAuthorityYear="2015" box="[295,571,1485,1512]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="genus">
<emphasis id="97E1EAB6C43D642CFEACFA61FE81FA29" box="[295,357,1485,1511]" italics="true" pageId="4" pageNumber="87">Sialis</emphasis>
(
<bibRefCitation id="C1044B55C43D642CFEFDFA61FDD7FA26" author="Liu, X. Y. &amp; Hayashi, F. &amp; Yang, D." box="[374,563,1485,1512]" pageId="4" pageNumber="87" pagination="18 - 49" refId="ref2722" refString="Liu, X. Y., Hayashi, F. &amp; Yang, D. (2015 a) Phylogeny of the family Sialidae (Insecta: Megaloptera) inferred from morphological data, with implications for generic classification and historical biogeography. Cladistics, 31, 18 - 49. https: // doi. org / 10.1111 / cla. 12071" type="journal article" year="2015">
Liu 
<emphasis id="97E1EAB6C43D642CFE2CFA62FE3EFA29" box="[423,474,1485,1511]" italics="true" pageId="4" pageNumber="87">et al</emphasis>
. 2015a
</bibRefCitation>
)
</taxonomicName>
. The new species is similar to other related species by external appearance, however it may be distinguished from others by the broadly lingulate male sternum 9 with elevated lateral edges, the male gonocoxites 9 with ventral portion produced medially, and the shape of the male gonoxoxites and gonostyli 11 (
<figureCitation id="3DAE2A21C43D642CFF48F995FE8BF99D" box="[195,367,1593,1619]" captionStart="FIGURE 2" captionStartId="3.[151,250,988,1012]" captionTargetBox="[220,1368,185,961]" captionTargetId="figure-19@3.[216,1372,181,965]" captionTargetPageId="3" captionText="FIGURE 2. Genitalia of Sialis kouwenkaii sp. nov. A. male, dorsal view; B. male, ventral view; C. male sternum 9, ventral view; D. male, lateral view; E. male, caudal view; F. female, ventral view. e = ectoproct; gp8 = gonapophyses 8; gst9, 11 = gonostylus 9, 11; gx8, 9, 11 = gonocoxites 8, 9, 11; S7, 9 = sternum 7, 9; T9 = tergum 9. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8324407" httpUri="https://zenodo.org/record/8324407/files/figure.png" pageId="4" pageNumber="87">Figs. 2B, C, D</figureCitation>
). In 
<taxonomicName id="62954D27C43D642CFE2FF995FDC7F99D" authorityName="Liu, Hayashi &amp; Yang" authorityYear="2015" box="[420,547,1593,1619]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="97E1EAB6C43D642CFE2FF995FDC7F99D" box="[420,547,1593,1619]" italics="true" pageId="4" pageNumber="87">S. australis</emphasis>
</taxonomicName>
, 
<taxonomicName id="62954D27C43D642CFDB8F995FD75F99D" authorityName="Liu, Hayashi &amp; Yang" authorityYear="2015" box="[563,657,1593,1619]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="jiyuni">
<emphasis id="97E1EAB6C43D642CFDB8F995FD75F99D" box="[563,657,1593,1619]" italics="true" pageId="4" pageNumber="87">S. jiyuni</emphasis>
</taxonomicName>
, 
<taxonomicName id="62954D27C43D642CFD2BF995FCC7F99D" authorityName="Okamoto" authorityYear="1910" box="[672,803,1593,1619]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="jezoensis">
<emphasis id="97E1EAB6C43D642CFD2BF995FCC7F99D" box="[672,803,1593,1619]" italics="true" pageId="4" pageNumber="87">S. jezoensis</emphasis>
</taxonomicName>
, 
<taxonomicName id="62954D27C43D642CFCB9F995FC2DF99D" authorityName="Okamoto" authorityYear="1910" box="[818,969,1593,1619]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="kumejimae">
<emphasis id="97E1EAB6C43D642CFCB9F995FC2DF99D" box="[818,969,1593,1619]" italics="true" pageId="4" pageNumber="87">S. kumejimae</emphasis>
</taxonomicName>
, and 
<taxonomicName id="62954D27C43D642CFB81F995FB43F99D" baseAuthorityName="Hayashi &amp; Suda" baseAuthorityYear="1995" box="[1034,1191,1593,1619]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="kuwayamai">
<emphasis id="97E1EAB6C43D642CFB81F995FB43F99D" box="[1034,1191,1593,1619]" italics="true" pageId="4" pageNumber="87">S. kuwayamai</emphasis>
</taxonomicName>
, the male sternums 9 are short, nearly semicircular (in 
<taxonomicName id="62954D27C43D642CFD9AF9F1FD77F9B9" authorityName="Okamoto" authorityYear="1910" box="[529,659,1629,1655]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="jezoensis">
<emphasis id="97E1EAB6C43D642CFD9AF9F1FD77F9B9" box="[529,659,1629,1655]" italics="true" pageId="4" pageNumber="87">S. jezoensis</emphasis>
</taxonomicName>
and 
<taxonomicName id="62954D27C43D642CFD46F9F1FCCEF9B9" authorityName="Liu, Hayashi &amp; Yang" authorityYear="2015" box="[717,810,1629,1655]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="jiyuni">
<emphasis id="97E1EAB6C43D642CFD46F9F1FCCEF9B9" box="[717,810,1629,1655]" italics="true" pageId="4" pageNumber="87">S. jiyuni</emphasis>
</taxonomicName>
, 
<figureCitation id="3DAE2A21C43D642CFCB2F9F1FC59F9B9" box="[825,957,1629,1655]" captionStart="FIGURE 3" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[218,1368,1112,1915]" captionTargetId="figure-92@3.[215,1372,1109,1919]" captionTargetPageId="3" captionText="FIGURE 3. Male genitalia of closely related species of Sialis kouwenkaii sp. nov., ventral view. A. S. australis Liu, Hayashi &amp; Yang, 2015; B. S. jezoensis Okamoto, 1910; C. S. jiyuni Liu, Hayashi &amp; Yang, 2015; D. S. kumejimae Okamoto, 1910; E. S. kuwayamai (Hayashi &amp; Suda, 1995). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8324409" httpUri="https://zenodo.org/record/8324409/files/figure.png" pageId="4" pageNumber="87">Figs. 3B, C</figureCitation>
), or subquadrilateral with shallow to deep incisions on the posterior margins (in 
<taxonomicName id="62954D27C43D642CFDDEF92DFD33F955" authorityName="Liu, Hayashi &amp; Yang" authorityYear="2015" box="[597,727,1665,1691]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="australis">
<emphasis id="97E1EAB6C43D642CFDDEF92DFD33F955" box="[597,727,1665,1691]" italics="true" pageId="4" pageNumber="87">S. australis</emphasis>
</taxonomicName>
, 
<taxonomicName id="62954D27C43D642CFD62F92DFC65F955" authorityName="Okamoto" authorityYear="1910" box="[745,897,1665,1691]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="kumejimae">
<emphasis id="97E1EAB6C43D642CFD62F92DFC65F955" box="[745,897,1665,1691]" italics="true" pageId="4" pageNumber="87">S. kumejimae</emphasis>
</taxonomicName>
and 
<taxonomicName id="62954D27C43D642CFC34F92DFBBBF955" baseAuthorityName="Hayashi &amp; Suda" baseAuthorityYear="1995" box="[959,1119,1665,1691]" class="Insecta" family="Sialidae" genus="Sialis" kingdom="Animalia" order="Megaloptera" pageId="4" pageNumber="87" phylum="Arthropoda" rank="species" species="kuwayamai">
<emphasis id="97E1EAB6C43D642CFC34F92DFBBBF955" box="[959,1119,1665,1691]" italics="true" pageId="4" pageNumber="87">S. kuwayamai</emphasis>
</taxonomicName>
, 
<figureCitation id="3DAE2A21C43D642CFBFBF92DFAC4F955" box="[1136,1312,1665,1692]" captionStart="FIGURE 3" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[218,1368,1112,1915]" captionTargetId="figure-92@3.[215,1372,1109,1919]" captionTargetPageId="3" captionText="FIGURE 3. Male genitalia of closely related species of Sialis kouwenkaii sp. nov., ventral view. A. S. australis Liu, Hayashi &amp; Yang, 2015; B. S. jezoensis Okamoto, 1910; C. S. jiyuni Liu, Hayashi &amp; Yang, 2015; D. S. kumejimae Okamoto, 1910; E. S. kuwayamai (Hayashi &amp; Suda, 1995). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8324409" httpUri="https://zenodo.org/record/8324409/files/figure.png" pageId="4" pageNumber="87">Figs. 3A, D, E</figureCitation>
); the male gonocoxites 9 have ventral portion not produced medially (
<figureCitation id="3DAE2A21C43D642CFCB3F909FC6CF971" box="[824,904,1701,1727]" captionStart="FIGURE 3" captionStartId="3.[151,250,1942,1966]" captionTargetBox="[218,1368,1112,1915]" captionTargetId="figure-92@3.[215,1372,1109,1919]" captionTargetPageId="3" captionText="FIGURE 3. Male genitalia of closely related species of Sialis kouwenkaii sp. nov., ventral view. A. S. australis Liu, Hayashi &amp; Yang, 2015; B. S. jezoensis Okamoto, 1910; C. S. jiyuni Liu, Hayashi &amp; Yang, 2015; D. S. kumejimae Okamoto, 1910; E. S. kuwayamai (Hayashi &amp; Suda, 1995). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8324409" httpUri="https://zenodo.org/record/8324409/files/figure.png" pageId="4" pageNumber="87">Figs. 3</figureCitation>
, 
<figureCitation id="3DAE2A21C43D642CFC1DF909FC42F90E" box="[918,934,1701,1728]" captionStart="FIGURE 5" captionStartId="5.[151,250,1142,1166]" captionTargetBox="[222,1369,191,1118]" captionTargetId="figure-19@5.[215,1372,181,1118]" captionTargetPageId="5" captionText="FIGURE 5. Male genitalia of Sialis spp., caudal view. A. S. australis Liu, Hayashi &amp; Yang, 2015; B. S. jezoensis Okamoto, 1910; C. S. jiyuni Liu, Hayashi &amp; Yang, 2015; D. S. kumejimae Okamoto, 1910; E. S. kuwayamai (Hayashi &amp; Suda, 1995). Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.8324417" httpUri="https://zenodo.org/record/8324417/files/figure.png" pageId="4" pageNumber="87">5</figureCitation>
); the male gonocoxites and gonostyli 11 are also different from the new species (
<figureCitation id="3DAE2A21C43D642CFDA4F965FD7FF92A" box="[559,667,1737,1764]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="3.[151,250,1942,1966]" captionStartId-1="4.[151,250,1091,1115]" captionStartId-2="5.[151,250,1142,1166]" captionTargetBox-0="[218,1368,1112,1915]" captionTargetBox-1="[224,1363,182,1060]" captionTargetBox-2="[222,1369,191,1118]" captionTargetId-0="figure-92@3.[215,1372,1109,1919]" captionTargetId-1="figure-22@4.[215,1372,181,1066]" captionTargetId-2="figure-19@5.[215,1372,181,1118]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 3. Male genitalia of closely related species of Sialis kouwenkaii sp. nov., ventral view. A. S. australis Liu, Hayashi &amp; Yang, 2015; B. S. jezoensis Okamoto, 1910; C. S. jiyuni Liu, Hayashi &amp; Yang, 2015; D. S. kumejimae Okamoto, 1910; E. S. kuwayamai (Hayashi &amp; Suda, 1995). Scale bar = 0.5 mm." captionText-1="FIGURE 4. Male genitalia of Sialis spp., lateral view. A. S. australis Liu, Hayashi &amp; Yang, 2015; B. S. jezoensis Okamoto, 1910; C. S. jiyuni Liu, Hayashi &amp; Yang, 2015; D. S. kumejimae Okamoto, 1910; E. S. kuwayamai (Hayashi &amp; Suda, 1995). Scale bar = 0.5 mm." captionText-2="FIGURE 5. Male genitalia of Sialis spp., caudal view. A. S. australis Liu, Hayashi &amp; Yang, 2015; B. S. jezoensis Okamoto, 1910; C. S. jiyuni Liu, Hayashi &amp; Yang, 2015; D. S. kumejimae Okamoto, 1910; E. S. kuwayamai (Hayashi &amp; Suda, 1995). Scale bar = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8324409" figureDoi-1="http://doi.org/10.5281/zenodo.8324413" figureDoi-2="http://doi.org/10.5281/zenodo.8324417" httpUri-0="https://zenodo.org/record/8324409/files/figure.png" httpUri-1="https://zenodo.org/record/8324413/files/figure.png" httpUri-2="https://zenodo.org/record/8324417/files/figure.png" pageId="4" pageNumber="87">Figs. 3–5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>