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<document id="D52FE62257BD1D8D39712B4C0EB2B9CF" ID-CLB-Dataset="7531" ID-DOI="10.5852/ejt.2021.742.1291" ID-GBIF-Dataset="62f9dc57-bade-4fbc-8af7-f92ff107593c" ID-ISSN="2118-9773" ID-Zenodo-Dep="4655404" ID-ZooBank="23424EC3-8440-4C72-AD9E-C1EC3087F7CA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617262825393" checkinUser="plazi" docAuthor="De Oliveira, Luiz Fernando M. &amp; Brescovit, Antonio Domingos" docDate="2021" docId="03CF87ADFF8C6608FD069756FE3FFD3E" docLanguage="en" docName="ejt-742_1-77.pdf" docOrigin="European Journal of Taxonomy 742" docSource="http://dx.doi.org/10.5852/ejt.2021.742.1291" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Tafana ruizi De Oliveira &amp; Brescovit 2021, sp. nov." docType="treatment" docUuid="DAF38D70-D874-40E0-9C78-A041F4B36B88" docUuidSource="ZooBank" docVersion="8" lastPageNumber="39" masterDocId="FFF6FFD5FFA8662EFF99965EFF99FFCF" masterDocTitle="Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species" masterLastPageNumber="77" masterPageNumber="1" pageNumber="37" updateTime="1698935806814" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="AAF81338E5FB0572501D64BFE106C024">Taxonomic revision and cladistic analysis of ghost spiders of the genus Tafana Simon, 1903 (Araneae: Dionycha, Anyphaenidae), with the descriptions of twelve new species</mods:title>
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<mods:namePart id="961FAB1602E729F3F87C7A0EBD574033">De Oliveira, Luiz Fernando M.</mods:namePart>
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<mods:affiliation id="D00834EF674F42E677BED91F83631CF9">Laboratório de Coleções Zoológicas, Instituto Butantan - Av. Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil. &amp; Instituto de Biociências, Universidade de São Paulo, Rua do Matão, 321, 05508 - 090, São Paulo, SP, Brazil.</mods:affiliation>
<mods:nameIdentifier id="A05291B57E811DA353776AD265E21566" type="email">luiz.moura.lfm@gmail.com</mods:nameIdentifier>
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<mods:namePart id="0C6554443BD25F19C3E685052DA6CBF7">Brescovit, Antonio Domingos</mods:namePart>
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<mods:affiliation id="3718BC1E76BEEBABD5EE7960647ACBF6">Laboratório de Coleções Zoológicas, Instituto Butantan - Av. Vital Brazil, 1500, 05503 - 900, São Paulo, SP, Brazil.</mods:affiliation>
<mods:nameIdentifier id="D65DC40110E15E0B3314FA17E4937D30" type="email">antonio.brescovit@butantan.gov.br</mods:nameIdentifier>
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<mods:title id="D0A7AE9F42583013B1E3C708FA61A459">European Journal of Taxonomy</mods:title>
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<mods:date id="8F0491241ACD6F98366E9DF1703062F8">2021</mods:date>
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<treatment id="03CF87ADFF8C6608FD069756FE3FFD3E" ID-DOI="http://doi.org/10.5281/zenodo.4658753" ID-GBIF-Taxon="180784548" ID-Zenodo-Dep="4658753" LSID="urn:lsid:zoobank.org:act:DAF38D70-D874-40E0-9C78-A041F4B36B88" httpUri="http://treatment.plazi.org/id/03CF87ADFF8C6608FD069756FE3FFD3E" lastPageId="38" lastPageNumber="39" pageId="36" pageNumber="37">
<subSubSection id="C37C6530FF8C660AFD069756FC0CFEEC" box="[671,917,264,291]" pageId="36" pageNumber="37" type="nomenclature">
<paragraph id="8BD936BBFF8C660AFD069756FC0CFEEC" blockId="36.[366,1222,264,361]" box="[671,917,264,291]" pageId="36" pageNumber="37">
<heading id="D09181D7FF8C660AFD069756FC0CFEEC" box="[671,917,264,291]" centered="true" fontSize="11" level="2" pageId="36" pageNumber="37" reason="2">
<taxonomicName id="4C664D38FF8C660AFD069756FCAAFEEC" ID-CoL="98Z2Y" authority="De Oliveira &amp; Brescovit, 2021" authorityName="De Oliveira &amp; Brescovit" authorityYear="2021" box="[671,819,264,291]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="ruizi" status="sp. nov.">
<emphasis id="B912EAA9FF8C660AFD069756FCAAFEEC" bold="true" box="[671,819,264,291]" italics="true" pageId="36" pageNumber="37">Tafana ruizi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157D2FF8C660AFCA39757FC0CFEEC" box="[826,917,265,291]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
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<subSubSection id="C37C6530FF8C660AFEF79772FCE2FEA6" pageId="36" pageNumber="37" type="description">
<paragraph id="8BD936BBFF8C660AFEF79772FB5FFE89" blockId="36.[366,1222,264,361]" box="[366,1222,300,326]" pageId="36" pageNumber="37">
<uri id="FFF73AB9FF8C660AFEF79772FB5FFE89" box="[366,1222,300,326]" pageId="36" pageNumber="37">
urn:lsid:zoobank.org:act:
<uuid id="FFC00C6EFF8C660AFD169772FB5FFE89" box="[655,1222,300,326]" pageId="36" pageNumber="37">DAF38D70-D874-40E0-9C78-A041F4B36B88</uuid>
</uri>
</paragraph>
<paragraph id="8BD936BBFF8C660AFD219711FCE2FEA6" blockId="36.[366,1222,264,361]" box="[696,891,335,361]" pageId="36" pageNumber="37">
<figureCitation id="135D2A3EFF8C660AFD219711FC8BFEA6" box="[696,786,335,361]" captionStart="Fig" captionStartId="37.[189,232,1785,1811]" captionTargetBox="[189,1399,458,1743]" captionTargetId="figure-195@37.[189,1399,458,1743]" captionTargetPageId="37" captionText="Fig. 21. Tafana ruizi sp. nov. (A, CE =holotype, ♂, IBSP 258183; B, EF =paratype, ♀, IBSP 258183). A. Habitus, dorsal view.B. Habitus, dorsal view.C. Left palp, ventral view (arrow indicating the base of the embolus). D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A = atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; MA =median apophysis; RTA=retrolateral tibial apophysis; S1=primary spermathecae; S2 = secondary spermathecae; T =tegulum. Scale bars: AB=3.5 mm; C=0.5 mm; DG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655467" httpUri="https://zenodo.org/record/4655467/files/figure.png" pageId="36" pageNumber="37">Figs 21</figureCitation>
,
<figureCitation id="135D2A3EFF8C660AFC879711FCC8FEA6" box="[798,849,335,361]" captionStart="Fig" captionStartId="57.[187,230,1824,1850]" captionTargetBox="[189,1398,495,1783]" captionTargetId="figure-161@57.[189,1398,495,1783]" captionTargetPageId="57" captionText="Fig. 33. Female genitalia, digested. A. Tafana huatanay sp. nov. (MUSM). B. Tafana ruizi sp. nov. (IBSP 258183). C. Tafana silhavyi (Caporiacco, 1955) (MNHN). D. Tafana maracay sp. nov. (MNHN 11314). E. Tafana arawak sp. nov. (MNHN 11320). F. Tafana pittieri sp. nov. (AMNH). G. Tafana oliviae sp. nov. (MCZ). H. Tafana orinoco sp. nov. (MIZA 07062). Abbreviations: CD =copulatory ducts; FD =fertilization ducts; H =hood; S1 =primary spermathecae; S2 =secondary spermathecae. Scale bars: AH =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655511" httpUri="https://zenodo.org/record/4655511/files/figure.png" pageId="36" pageNumber="37">33B</figureCitation>
,
<figureCitation id="135D2A3EFF8C660AFCC79711FCE2FEA6" box="[862,891,335,361]" captionStart="Fig" captionStartId="58.[189,232,1997,2023]" captionTargetBox="[217,1370,264,1952]" captionTargetId="figure-27@58.[217,1370,264,1953]" captionTargetPageId="58" captionText="Fig. 34. Distribution map of species of Tafana." figureDoi="http://doi.org/10.5281/zenodo.4655515" httpUri="https://zenodo.org/record/4655515/files/figure.png" pageId="36" pageNumber="37">34</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37C6530FF8C660AFF2497CEFD2AFDA9" pageId="36" pageNumber="37" type="diagnosis">
<paragraph id="8BD936BBFF8C660AFF2497CEFEAFFE65" blockId="36.[189,310,400,426]" box="[189,310,400,426]" pageId="36" pageNumber="37">
<heading id="D09181D7FF8C660AFF2497CEFEAFFE65" bold="true" box="[189,310,400,426]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B912EAA9FF8C660AFF2497CEFEAFFE65" bold="true" box="[189,310,400,426]" pageId="36" pageNumber="37">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BD936BBFF8C660AFF2497E1FD2AFDA9" blockId="36.[188,1399,447,614]" pageId="36" pageNumber="37">
Males of
<taxonomicName id="4C664D38FF8C660AFEB397E1FE2EFE16" authority="De Oliveira &amp; Brescovit, 2021" authorityName="De Oliveira &amp; Brescovit" authorityYear="2021" box="[298,439,447,473]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="ruizi" status="sp. nov.">
<emphasis id="B912EAA9FF8C660AFEB397E1FE2EFE16" box="[298,439,447,473]" italics="true" pageId="36" pageNumber="37">Tafana ruizi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157D2FF8C660AFE27979EFD81FE15" box="[446,536,448,474]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
resemble those of
<taxonomicName id="4C664D38FF8C660AFD6A979EFC1DFE16" authorityName="AMNH" baseAuthorityName="L. Koch" baseAuthorityYear="1866" box="[755,900,448,473]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="straminea">
<emphasis id="B912EAA9FF8C660AFD6A979EFC1DFE16" box="[755,900,448,473]" italics="true" pageId="36" pageNumber="37">T. straminea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C664D38FF8C660AFC25979EFBE8FE16" authority="De Oliveira &amp; Brescovit, 2021" authorityName="De Oliveira &amp; Brescovit" authorityYear="2021" box="[956,1137,447,473]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="kunturmarqa" status="sp. nov.">
<emphasis id="B912EAA9FF8C660AFC25979EFBE8FE16" box="[956,1137,447,473]" italics="true" pageId="36" pageNumber="37">T. kunturmarqa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157D2FF8C660AFBE1979EFB4BFE15" box="[1144,1234,448,474]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
by having the base of the embolus large and inclined, but differ by having the ventral tegular process large and apex of tegulum not rounded (
<figureCitation id="135D2A3EFF8C660AFE469458FDD3FDEF" box="[479,586,518,544]" captionStart="Fig" captionStartId="37.[189,232,1785,1811]" captionTargetBox="[189,1399,458,1743]" captionTargetId="figure-195@37.[189,1399,458,1743]" captionTargetPageId="37" captionText="Fig. 21. Tafana ruizi sp. nov. (A, CE =holotype, ♂, IBSP 258183; B, EF =paratype, ♀, IBSP 258183). A. Habitus, dorsal view.B. Habitus, dorsal view.C. Left palp, ventral view (arrow indicating the base of the embolus). D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A = atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; MA =median apophysis; RTA=retrolateral tibial apophysis; S1=primary spermathecae; S2 = secondary spermathecae; T =tegulum. Scale bars: AB=3.5 mm; C=0.5 mm; DG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655467" httpUri="https://zenodo.org/record/4655467/files/figure.png" pageId="36" pageNumber="37">Fig. 21C</figureCitation>
). Females resemble those of
<taxonomicName id="4C664D38FF8C660AFC009459FBB3FDEF" authorityName="AMNH" baseAuthorityName="L. Koch" baseAuthorityYear="1866" box="[921,1066,519,544]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="straminea">
<emphasis id="B912EAA9FF8C660AFC009459FBB3FDEF" box="[921,1066,519,544]" italics="true" pageId="36" pageNumber="37">T. straminea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C664D38FF8C660AFBFB9459FA8FFDEF" authority="De Oliveira &amp; Brescovit, 2021" authorityName="De Oliveira &amp; Brescovit" authorityYear="2021" box="[1122,1302,518,544]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="kunturmarqa" status="sp. nov.">
<emphasis id="B912EAA9FF8C660AFBFB9459FA8FFDEF" box="[1122,1302,518,544]" italics="true" pageId="36" pageNumber="37">T. kunturmarqa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157D2FF8C660AFA849458FAEEFDEF" box="[1309,1399,518,544]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
by having the atrial septum, but differ by having the wide atrium, showing the copulation openings and hood long and circular (
<figureCitation id="135D2A3EFF8C660AFE4C9412FDFCFDA9" box="[469,613,588,614]" captionStart="Fig" captionStartId="37.[189,232,1785,1811]" captionTargetBox="[189,1399,458,1743]" captionTargetId="figure-195@37.[189,1399,458,1743]" captionTargetPageId="37" captionText="Fig. 21. Tafana ruizi sp. nov. (A, CE =holotype, ♂, IBSP 258183; B, EF =paratype, ♀, IBSP 258183). A. Habitus, dorsal view.B. Habitus, dorsal view.C. Left palp, ventral view (arrow indicating the base of the embolus). D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A = atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; MA =median apophysis; RTA=retrolateral tibial apophysis; S1=primary spermathecae; S2 = secondary spermathecae; T =tegulum. Scale bars: AB=3.5 mm; C=0.5 mm; DG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655467" httpUri="https://zenodo.org/record/4655467/files/figure.png" pageId="36" pageNumber="37">Figs 21FG</figureCitation>
,
<figureCitation id="135D2A3EFF8C660AFDE89412FD3FFDA9" box="[625,678,588,614]" captionStart="Fig" captionStartId="57.[187,230,1824,1850]" captionTargetBox="[189,1398,495,1783]" captionTargetId="figure-161@57.[189,1398,495,1783]" captionTargetPageId="57" captionText="Fig. 33. Female genitalia, digested. A. Tafana huatanay sp. nov. (MUSM). B. Tafana ruizi sp. nov. (IBSP 258183). C. Tafana silhavyi (Caporiacco, 1955) (MNHN). D. Tafana maracay sp. nov. (MNHN 11314). E. Tafana arawak sp. nov. (MNHN 11320). F. Tafana pittieri sp. nov. (AMNH). G. Tafana oliviae sp. nov. (MCZ). H. Tafana orinoco sp. nov. (MIZA 07062). Abbreviations: CD =copulatory ducts; FD =fertilization ducts; H =hood; S1 =primary spermathecae; S2 =secondary spermathecae. Scale bars: AH =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655511" httpUri="https://zenodo.org/record/4655511/files/figure.png" pageId="36" pageNumber="37">33B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37C6530FF8C660AFF2494D0FD39FD36" pageId="36" pageNumber="37" type="etymology">
<paragraph id="8BD936BBFF8C660AFF2494D0FEDDFD67" blockId="36.[189,324,654,680]" box="[189,324,654,680]" pageId="36" pageNumber="37">
<heading id="D09181D7FF8C660AFF2494D0FEDDFD67" bold="true" box="[189,324,654,680]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B912EAA9FF8C660AFF2494D0FEDDFD67" bold="true" box="[189,324,654,680]" pageId="36" pageNumber="37">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BD936BBFF8C660AFF2494E2FD39FD36" blockId="36.[189,1398,700,762]" pageId="36" pageNumber="37">The specific name is a patronym in honor of the Cuban arachnologist Alexandre Sánchez-Ruiz for his contributions to the taxonomy of spiders.</paragraph>
</subSubSection>
<subSubSection id="C37C6530FF8C660AFF24957EFB9FFBDA" pageId="36" pageNumber="37" type="materials_examined">
<paragraph id="8BD936BBFF8C660AFF24957EFE33FCF5" blockId="36.[189,426,800,826]" box="[189,426,800,826]" pageId="36" pageNumber="37">
<heading id="D09181D7FF8C660AFF24957EFE33FCF5" bold="true" box="[189,426,800,826]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B912EAA9FF8C660AFF24957EFE33FCF5" bold="true" box="[189,426,800,826]" pageId="36" pageNumber="37">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BD936BBFF8C660AFF739511FDDFFC7F" blockId="36.[189,1398,847,944]" pageId="36" pageNumber="37">
<materialsCitation id="3B0E3CE6FF8C660AFF739511FDDFFC7F" ID-GBIF-Occurrence="3066856326" collectingDate="2003-10-10" collectionCode="IBSP" collectorName="A. Sanchez-Ruiz" country="Colombia" latitude="5.3838887" location="Pensilvania" longLatPrecision="21" longitude="-75.16111" municipality="Pensilvania" pageId="36" pageNumber="37" specimenCode="IBSP 258183" specimenCount="1" specimenCount-male="1" stateProvince="Departamento de Caldas" typeStatus="holotype">
<emphasis id="B912EAA9FF8C660AFF739511FEC5FCA6" bold="true" box="[234,348,847,873]" pageId="36" pageNumber="37">
<typeStatus id="54DD8819FF8C660AFF739511FEC5FCA6" box="[234,348,847,873]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F371762BFF8C660AFF24952DFEC0FC42" box="[189,345,883,909]" name="Colombia" pageId="36" pageNumber="37">COLOMBIA</collectingCountry>
<specimenCount id="9D60FD32FF8C660AFEF6952DFE04FC42" box="[367,413,883,909]" pageId="36" pageNumber="37" type="male">1 ♂</specimenCount>
;
<collectingRegion id="49A2F859FF8C660AFE30952DFD53FC43" box="[425,714,882,909]" country="Colombia" name="Caldas" pageId="36" pageNumber="37">Departamento de Caldas</collectingRegion>
,
<collectingMunicipality id="6BBDACC1FF8C660AFD4C952CFCF9FC43" box="[725,864,882,908]" pageId="36" pageNumber="37">Pensilvania</collectingMunicipality>
; [
<geoCoordinate id="EE52507CFF8C660AFCEB952FFC62FC42" box="[882,1019,881,909]" degrees="5" direction="north" minutes="23" orientation="latitude" pageId="36" pageNumber="37" precision="15" seconds="02" value="5.3838887">5°2302″ N</geoCoordinate>
,
<geoCoordinate id="EE52507CFF8C660AFB9F952FFB3AFC42" box="[1030,1187,881,909]" degrees="75" direction="west" minutes="09" orientation="longitude" pageId="36" pageNumber="37" precision="15" seconds="40" value="-75.16111">75°0940″ W</geoCoordinate>
];
<date id="FFD8107BFF8C660AFB2F952DFAC9FC42" box="[1206,1360,883,909]" pageId="36" pageNumber="37" value="2003-10-10">
<collectingDate id="EF9CE993FF8C660AFB2F952DFAC9FC42" box="[1206,1360,883,909]" pageId="36" pageNumber="37" value="2003-10-10">10 Oct. 2003</collectingDate>
</date>
;
<collectorName id="2693536DFF8C660AFAC3952DFEF9FC7F" pageId="36" pageNumber="37">A. Sánchez-Ruiz</collectorName>
leg.;
<specimenCode id="DBC09EC0FF8C660AFE3B95C8FDDBFC7F" box="[418,578,917,944]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="36" pageNumber="37">IBSP 258183</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BD936BBFF8C660AFF739589FB9FFBDA" blockId="36.[189,1030,983,1045]" pageId="36" pageNumber="37">
<materialsCitation id="3B0E3CE6FF8C660AFF739589FB9AFBDA" ID-GBIF-Occurrence="3066856344" collectingDate="2003-10-10" collectionCode="IBSP" collectorName="A. Sanchez-Ruiz" country="Colombia" latitude="5.3838887" location="COLOMBIA" longLatPrecision="21" longitude="-75.16111" pageId="36" pageNumber="37" specimenCode="IBSP 258183" specimenCount="1" specimenCount-female="1" stateProvince="Caldas" typeStatus="paratype">
<emphasis id="B912EAA9FF8C660AFF739589FEC5FC3E" bold="true" box="[234,348,983,1009]" pageId="36" pageNumber="37">
<typeStatus id="54DD8819FF8C660AFF739589FEC5FC3E" box="[234,348,983,1009]" pageId="36" pageNumber="37" type="paratype">Paratype</typeStatus>
</emphasis>
<collectingCountry id="F371762BFF8C660AFF2495A4FEC0FBDB" box="[189,345,1018,1044]" name="Colombia" pageId="36" pageNumber="37">COLOMBIA</collectingCountry>
<specimenCount id="9D60FD32FF8C660AFEE895A4FE07FBDA" box="[369,414,1018,1045]" pageId="36" pageNumber="37" type="female">1 ♀</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBC09EC0FF8C660AFCFA95A4FB9AFBDA" box="[867,1027,1018,1045]" collectionCode="IBSP" country="Brazil" httpUri="http://grbio.org/cool/23qr-v2gj" name="Instituto Biologico de Sao Paulo" pageId="36" pageNumber="37">IBSP 258183</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37C6530FF8C6608FF249265FAEEFD49" lastPageId="38" lastPageNumber="39" pageId="36" pageNumber="37" type="description">
<paragraph id="8BD936BBFF8C660AFF249265FED7FB9A" blockId="36.[189,334,1083,1109]" box="[189,334,1083,1109]" pageId="36" pageNumber="37">
<heading id="D09181D7FF8C660AFF249265FED7FB9A" bold="true" box="[189,334,1083,1109]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B912EAA9FF8C660AFF249265FED7FB9A" bold="true" box="[189,334,1083,1109]" pageId="36" pageNumber="37">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BD936BBFF8C660AFF739234FDCFFB4A" blockId="36.[189,1401,1130,1262]" box="[234,598,1130,1157]" pageId="36" pageNumber="37">
<emphasis id="B912EAA9FF8C660AFF739234FEB3FB4B" bold="true" box="[234,298,1130,1156]" pageId="36" pageNumber="37">Male</emphasis>
(
<typeStatus id="54DD8819FF8C660AFEA09234FE3DFB4B" box="[313,420,1130,1156]" pageId="36" pageNumber="37" type="holotype">holotype</typeStatus>
, IBSP 258183)
</paragraph>
<paragraph id="8BD936BBFF8C660AFF2492D0FEDBFB21" blockId="36.[189,1401,1130,1262]" pageId="36" pageNumber="37">
COLOR. Carapace reddish brown (
<figureCitation id="135D2A3EFF8C660AFDD792D3FD27FB67" box="[590,702,1165,1192]" captionStart="Fig" captionStartId="37.[189,232,1785,1811]" captionTargetBox="[189,1399,458,1743]" captionTargetId="figure-195@37.[189,1399,458,1743]" captionTargetPageId="37" captionText="Fig. 21. Tafana ruizi sp. nov. (A, CE =holotype, ♂, IBSP 258183; B, EF =paratype, ♀, IBSP 258183). A. Habitus, dorsal view.B. Habitus, dorsal view.C. Left palp, ventral view (arrow indicating the base of the embolus). D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A = atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; MA =median apophysis; RTA=retrolateral tibial apophysis; S1=primary spermathecae; S2 = secondary spermathecae; T =tegulum. Scale bars: AB=3.5 mm; C=0.5 mm; DG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655467" httpUri="https://zenodo.org/record/4655467/files/figure.png" pageId="36" pageNumber="37">Fig. 21A</figureCitation>
). Labium, sternum and endites yellow. Chelicerae reddish brown. Legs IIV yellow with spots, darker at tarsi. Abdomen dark with spots yellow in dorsal view (
<figureCitation id="135D2A3EFF8C660AFF51928AFEACFB21" box="[200,309,1236,1262]" captionStart="Fig" captionStartId="37.[189,232,1785,1811]" captionTargetBox="[189,1399,458,1743]" captionTargetId="figure-195@37.[189,1399,458,1743]" captionTargetPageId="37" captionText="Fig. 21. Tafana ruizi sp. nov. (A, CE =holotype, ♂, IBSP 258183; B, EF =paratype, ♀, IBSP 258183). A. Habitus, dorsal view.B. Habitus, dorsal view.C. Left palp, ventral view (arrow indicating the base of the embolus). D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A = atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; MA =median apophysis; RTA=retrolateral tibial apophysis; S1=primary spermathecae; S2 = secondary spermathecae; T =tegulum. Scale bars: AB=3.5 mm; C=0.5 mm; DG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655467" httpUri="https://zenodo.org/record/4655467/files/figure.png" pageId="36" pageNumber="37">Fig. 21A</figureCitation>
).
</paragraph>
<paragraph id="8BD936BBFF8C660AFF249348FE4DFA56" blockId="36.[189,1399,1301,1433]" pageId="36" pageNumber="37">MEASUREMENTS. Total length 6.9, carapace length 3.2, width 2.5. Clypeus height 0.1. Eye diameters and interdistances: AME 0.1, ALE 0.14, PME 0.14, PLE 0.14; AMEAME 0.08, AMEALE 0.1, PME PME 0.16, PMEPLE 0.16, ALEPLE 0.04. Chelicerae 2.0 long, with four promarginal teeth and five retromarginal denticles.</paragraph>
<paragraph id="8BD936BBFF8C660AFF24939FFB4EFA31" blockId="36.[189,1398,1472,1534]" pageId="36" pageNumber="37">LEG MEASUREMENTS. I—femur 2.7 /patella 1.2/ tibia 2.7/ metatarsus 2.3 / tarsus 1.1/ total 10.0; II— 2.5/ 1.1/ 2.5 /2.0 / 1.0/ 9.1; III—2.0 / 0.9/ 1.4/ 1.7/ 0.6/ 6.6; IV—2.4 / 1.1/ 2.3/ 2.7 /0.8 / 9.3.</paragraph>
<paragraph id="8BD936BBFF8C660AFF249078FBC2F9AD" blockId="36.[189,1399,1573,1635]" pageId="36" pageNumber="37">LEG SPINATION. I—tibia v1-2-1-1-1, metatarsus v2-0-0; II—tibia v0-2-1; III—tibia v1-2-2, p0-0-1, r0-1-1, metatarsus v2-0-2, p0-1-1, r0-1-1; IV—tibia v1-2-2, r1-1-0, metatarsus v2-0-2.</paragraph>
<paragraph id="8BD936BBFF8C660AFF2490D4FF79F8C1" blockId="36.[189,1400,1674,1806]" pageId="36" pageNumber="37">
PALP. Retrolateral tibial apophysis long and curved; cymbium with retrolateral projection; apophysis at the base of the embolus short and furrowed; embolus long; ventral tegular process wide and near the median apophysis; sperm duct with the secondary looping almost reaching apex of tegulum (
<figureCitation id="135D2A3EFF8C660AFA9B908EFF4AF8C1" captionStart="Fig" captionStartId="37.[189,232,1785,1811]" captionTargetBox="[189,1399,458,1743]" captionTargetId="figure-195@37.[189,1399,458,1743]" captionTargetPageId="37" captionText="Fig. 21. Tafana ruizi sp. nov. (A, CE =holotype, ♂, IBSP 258183; B, EF =paratype, ♀, IBSP 258183). A. Habitus, dorsal view.B. Habitus, dorsal view.C. Left palp, ventral view (arrow indicating the base of the embolus). D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A = atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; MA =median apophysis; RTA=retrolateral tibial apophysis; S1=primary spermathecae; S2 = secondary spermathecae; T =tegulum. Scale bars: AB=3.5 mm; C=0.5 mm; DG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655467" httpUri="https://zenodo.org/record/4655467/files/figure.png" pageId="36" pageNumber="37">Fig. 21C E</figureCitation>
).
</paragraph>
<paragraph id="8BD936BBFF8C660AFF24916BFAEEF880" blockId="36.[189,1399,1845,1871]" box="[189,1399,1845,1871]" pageId="36" pageNumber="37">ABDOMEN. Length 3.6, epigastric furrow 0.9 from tracheal spiracle, spiracle 1.1 from base of spinnerets.</paragraph>
<paragraph id="8BD936BBFF8C660AFF739128FDE9F85E" blockId="36.[189,1398,1910,2007]" box="[234,624,1910,1937]" pageId="36" pageNumber="37">
<emphasis id="B912EAA9FF8C660AFF739128FEDCF85F" bold="true" box="[234,325,1910,1936]" pageId="36" pageNumber="37">Female</emphasis>
(
<typeStatus id="54DD8819FF8C660AFECC9128FE26F85F" box="[341,447,1910,1936]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
, IBSP 258183)
</paragraph>
<paragraph id="8BD936BBFF8C660AFF2491C7FB19F818" blockId="36.[189,1398,1910,2007]" pageId="36" pageNumber="37">
COLOR. Coloration as in males of this species, except for the reddish carapace and chelicerae reddish brown and robust (
<figureCitation id="135D2A3EFF8C660AFE0491E2FD91F818" box="[413,520,1980,2007]" captionStart="Fig" captionStartId="37.[189,232,1785,1811]" captionTargetBox="[189,1399,458,1743]" captionTargetId="figure-195@37.[189,1399,458,1743]" captionTargetPageId="37" captionText="Fig. 21. Tafana ruizi sp. nov. (A, CE =holotype, ♂, IBSP 258183; B, EF =paratype, ♀, IBSP 258183). A. Habitus, dorsal view.B. Habitus, dorsal view.C. Left palp, ventral view (arrow indicating the base of the embolus). D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A = atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; MA =median apophysis; RTA=retrolateral tibial apophysis; S1=primary spermathecae; S2 = secondary spermathecae; T =tegulum. Scale bars: AB=3.5 mm; C=0.5 mm; DG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655467" httpUri="https://zenodo.org/record/4655467/files/figure.png" pageId="36" pageNumber="37">Fig. 21B</figureCitation>
). Abdomen gray with spots in dorsal view (
<figureCitation id="135D2A3EFF8C660AFB9E91E2FBEBF818" box="[1031,1138,1980,2007]" captionStart="Fig" captionStartId="37.[189,232,1785,1811]" captionTargetBox="[189,1399,458,1743]" captionTargetId="figure-195@37.[189,1399,458,1743]" captionTargetPageId="37" captionText="Fig. 21. Tafana ruizi sp. nov. (A, CE =holotype, ♂, IBSP 258183; B, EF =paratype, ♀, IBSP 258183). A. Habitus, dorsal view.B. Habitus, dorsal view.C. Left palp, ventral view (arrow indicating the base of the embolus). D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A = atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; MA =median apophysis; RTA=retrolateral tibial apophysis; S1=primary spermathecae; S2 = secondary spermathecae; T =tegulum. Scale bars: AB=3.5 mm; C=0.5 mm; DG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655467" httpUri="https://zenodo.org/record/4655467/files/figure.png" pageId="36" pageNumber="37">Fig. 21B</figureCitation>
).
</paragraph>
<paragraph id="8BD936BBFF8D660BFF249757FE64FE43" blockId="37.[189,1398,265,396]" pageId="37" pageNumber="38">MEASUREMENTS. Total length 7.4, carapace length 3.0, width 2.4. Clypeus height 0.10. Eye diameters and interdistances: AME 0.12, ALE 0.14, PME 0.14, PLE 0.14; AMEAME 0.06, AMEALE 0.06, PMEPME 0.2, PMEPLE 0.12, ALEPLE 0.04. Chelicerae 1.2 long, with five promarginal teeth and six retromarginal denticles.</paragraph>
<caption id="DF196633FF8D660BFF2490A7FB0EF828" ID-DOI="http://doi.org/10.5281/zenodo.4655467" ID-Zenodo-Dep="4655467" httpUri="https://zenodo.org/record/4655467/files/figure.png" pageId="37" pageNumber="38" startId="37.[189,232,1785,1811]" targetBox="[189,1399,458,1743]" targetPageId="37">
<paragraph id="8BD936BBFF8D660BFF2490A7FB0EF828" blockId="37.[185,1399,1785,2023]" pageId="37" pageNumber="38">
<emphasis id="B912EAA9FF8D660BFF2490A7FE81F8DC" bold="true" box="[189,280,1785,1811]" pageId="37" pageNumber="38">Fig. 21.</emphasis>
<taxonomicName id="4C664D38FF8D660BFE8790A7FE32F8DC" authority="De Oliveira &amp; Brescovit, 2021" authorityName="De Oliveira &amp; Brescovit" authorityYear="2021" box="[286,427,1785,1811]" class="Arachnida" family="Anyphaenidae" genus="Tafana" kingdom="Animalia" order="Araneae" pageId="37" pageNumber="38" phylum="Arthropoda" rank="species" species="ruizi" status="sp. nov.">
<emphasis id="B912EAA9FF8D660BFE8790A7FE32F8DC" box="[286,427,1785,1811]" italics="true" pageId="37" pageNumber="38">Tafana ruizi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22157D2FF8D660BFE2890A4FD93F8DB" box="[433,522,1786,1812]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
(A, CE =holotype, ♂, IBSP 258183; B, EF =paratype, ♀, IBSP 258183).
<emphasis id="B912EAA9FF8D660BFF249142FF4BF8F9" bold="true" box="[189,210,1820,1846]" pageId="37" pageNumber="38">A</emphasis>
. Habitus, dorsal view.
<emphasis id="B912EAA9FF8D660BFE569143FE7AF8F8" bold="true" box="[463,483,1821,1847]" pageId="37" pageNumber="38">B</emphasis>
. Habitus, dorsal view.
<emphasis id="B912EAA9FF8D660BFD7B9142FD6EF8F9" bold="true" box="[738,759,1820,1846]" pageId="37" pageNumber="38">C</emphasis>
. Left palp, ventral view (arrow indicating the base of the embolus).
<emphasis id="B912EAA9FF8D660BFEAE911EFED5F895" bold="true" box="[311,332,1856,1882]" pageId="37" pageNumber="38">D</emphasis>
. Left palp, retrolateral view.
<emphasis id="B912EAA9FF8D660BFD00911EFD34F895" bold="true" box="[665,685,1856,1882]" pageId="37" pageNumber="38">E</emphasis>
. Left palp, dorsal view.
<emphasis id="B912EAA9FF8D660BFC5B911EFC4DF895" bold="true" box="[962,980,1856,1882]" pageId="37" pageNumber="38">F</emphasis>
. Epigynum, ventral view.
<emphasis id="B912EAA9FF8D660BFA9D9161FA82F896" bold="true" box="[1284,1307,1855,1881]" pageId="37" pageNumber="38">G</emphasis>
. Vulva, dorsal view. Abbreviations: A = atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; MA =median apophysis; RTA=retrolateral tibial apophysis; S1=primary spermathecae; S2 = secondary spermathecae; T =tegulum. Scale bars: AB=3.5 mm; C=0.5 mm; DG =0.25 mm.
</paragraph>
</caption>
<paragraph id="8BD936BBFF8E6608FF249757FB6DFE8B" blockId="38.[189,1398,265,325]" pageId="38" pageNumber="39">LEG MEASUREMENTS. I—femur 2.1 /patella 1.1/ tibia 1.9/ metatarsus 1.5/ tarsus 0.8 /total 7.4; II— 2.0/ 1.1/ 1.8 /1.5 / 0.8/ 7.2; III—1.52 /0.9 / 1.1/ 1.4/ 0.5/ 5.42; IV—2.0 / 1.0 /2.0 / 2.2/ 0.6/ 7.8.</paragraph>
<paragraph id="8BD936BBFF8E6608FF249734FD1BFE6B" blockId="38.[189,1398,361,421]" pageId="38" pageNumber="39">LEG SPINATION. I—tibia v2-2-0, metatarsus v2-0-0; III—metatarsus v2-0-2, p1-0-1, r0-0-1; IV—tibia r0- 1-0, metatarsus v2-0-2, p0-1-1, r0-1-1.</paragraph>
<paragraph id="8BD936BBFF8E6608FF249794FAD1FD88" blockId="38.[188,1399,457,583]" pageId="38" pageNumber="39">
EPIGYNUM. Hood circular; atrial septum present; epigynal plate covered by cuticle in the posterior region; lateral lobes extremely curved in anterior region. Internally, copulatory ducts sinuous and laminar; primary spermathecae large and oval; secondary spermathecae short and rounded in the anterior region on the copulatory ducts; fertilization ducts long over the primary spermathecae (
<figureCitation id="135D2A3EFF8E6608FBF39472FB63FD88" box="[1130,1274,556,583]" captionStart="Fig" captionStartId="37.[189,232,1785,1811]" captionTargetBox="[189,1399,458,1743]" captionTargetId="figure-195@37.[189,1399,458,1743]" captionTargetPageId="37" captionText="Fig. 21. Tafana ruizi sp. nov. (A, CE =holotype, ♂, IBSP 258183; B, EF =paratype, ♀, IBSP 258183). A. Habitus, dorsal view.B. Habitus, dorsal view.C. Left palp, ventral view (arrow indicating the base of the embolus). D. Left palp, retrolateral view. E. Left palp, dorsal view. F. Epigynum, ventral view. G. Vulva, dorsal view. Abbreviations: A = atrium; ABE= apophysis of base of the embolus; CD =copulatory ducts; CO =copulatory opening; E =embolus; FD =fertilization ducts; H =hood; LL= lateral lobes; MA =median apophysis; RTA=retrolateral tibial apophysis; S1=primary spermathecae; S2 = secondary spermathecae; T =tegulum. Scale bars: AB=3.5 mm; C=0.5 mm; DG =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655467" httpUri="https://zenodo.org/record/4655467/files/figure.png" pageId="38" pageNumber="39">Figs 21FG</figureCitation>
,
<figureCitation id="135D2A3EFF8E6608FA9F9473FAA2FD88" box="[1286,1339,557,583]" captionStart="Fig" captionStartId="57.[187,230,1824,1850]" captionTargetBox="[189,1398,495,1783]" captionTargetId="figure-161@57.[189,1398,495,1783]" captionTargetPageId="57" captionText="Fig. 33. Female genitalia, digested. A. Tafana huatanay sp. nov. (MUSM). B. Tafana ruizi sp. nov. (IBSP 258183). C. Tafana silhavyi (Caporiacco, 1955) (MNHN). D. Tafana maracay sp. nov. (MNHN 11314). E. Tafana arawak sp. nov. (MNHN 11320). F. Tafana pittieri sp. nov. (AMNH). G. Tafana oliviae sp. nov. (MCZ). H. Tafana orinoco sp. nov. (MIZA 07062). Abbreviations: CD =copulatory ducts; FD =fertilization ducts; H =hood; S1 =primary spermathecae; S2 =secondary spermathecae. Scale bars: AH =0.25 mm." figureDoi="http://doi.org/10.5281/zenodo.4655511" httpUri="https://zenodo.org/record/4655511/files/figure.png" pageId="38" pageNumber="39">33B</figureCitation>
).
</paragraph>
<paragraph id="8BD936BBFF8E6608FF249432FAEEFD49" blockId="38.[189,1399,620,646]" box="[189,1399,620,646]" pageId="38" pageNumber="39">ABDOMEN. Length 4.1, epigastric furrow 0.98 from tracheal spiracle, spiracle 1.9 from base of spinnerets.</paragraph>
</subSubSection>
<subSubSection id="C37C6530FF8E6608FF2494F5FE3FFD3E" box="[189,422,683,753]" pageId="38" pageNumber="39" type="distribution">
<paragraph id="8BD936BBFF8E6608FF2494F5FECFFD0A" blockId="38.[189,422,683,753]" box="[189,342,683,709]" pageId="38" pageNumber="39">
<heading id="D09181D7FF8E6608FF2494F5FECFFD0A" bold="true" box="[189,342,683,709]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="B912EAA9FF8E6608FF2494F5FECFFD0A" bold="true" box="[189,342,683,709]" pageId="38" pageNumber="39">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BD936BBFF8E6608FF249489FE3FFD3E" blockId="38.[189,422,683,753]" box="[189,422,727,753]" pageId="38" pageNumber="39">
<collectingCountry id="F371762BFF8E6608FF249489FEA8FD3E" box="[189,305,727,753]" name="Colombia" pageId="38" pageNumber="39">Colombia</collectingCountry>
(
<figureCitation id="135D2A3EFF8E6608FED89489FE00FD3E" box="[321,409,727,753]" captionStart="Fig" captionStartId="58.[189,232,1997,2023]" captionTargetBox="[217,1370,264,1952]" captionTargetId="figure-27@58.[217,1370,264,1953]" captionTargetPageId="58" captionText="Fig. 34. Distribution map of species of Tafana." figureDoi="http://doi.org/10.5281/zenodo.4655515" httpUri="https://zenodo.org/record/4655515/files/figure.png" pageId="38" pageNumber="39">Fig. 34</figureCitation>
).
</paragraph>
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