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<document ID-DOI="10.11646/zootaxa.5231.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7575308" ID-ZooBank="EBD53535-397E-4894-B0A4-9AC3CC38DF3C" checkinTime="1674811458118" checkinUser="plazi" docAuthor="Lelej, Arkady S. &amp; Williams, Kevin A." docDate="2023" docId="03CF87A9313BB516FF17FC83F24C32BE" docLanguage="en" docName="zootaxa.5231.2.1.pdf" docOrigin="Zootaxa 5231 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5231.2.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Corytilla Arnold 1956" docType="treatment" docVersion="3" lastPageNumber="109" masterDocId="FFF6FFD13133B51EFF80FF97F345374A" masterDocTitle="Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae)" masterLastPageNumber="140" masterPageNumber="101" pageNumber="109" updateTime="1674840100621" updateUser="valdenar" zenodo-license-document="CLOSED">
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<mods:title>Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Lelej, Arkady S.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0001-7501-0981</mods:nameIdentifier>
<mods:affiliation>Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far East Branch of the Russian Academy of Sciences, Vladivostok, 690022, Russia. lelej @ biosoil. ru; https: // orcid. org / 0000 - 0001 - 7501 - 0981</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Williams, Kevin A.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-6073-1070</mods:nameIdentifier>
<mods:affiliation>Plant Pest Diagnostic Center, California Department of Food &amp; Agriculture, 3294 Meadowview Road, Sacramento, CA 95832, USA. Kevin. Williams @ cdfa. ca. gov; https: // orcid. org / 0000 - 0002 - 6073 - 1070</mods:affiliation>
<mods:nameIdentifier type="email">illiams@cdfa.ca.gov</mods:nameIdentifier>
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<mods:title>Zootaxa</mods:title>
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<mods:part>
<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-01-27</mods:number>
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<mods:detail type="volume">
<mods:number>5231</mods:number>
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<mods:number>2</mods:number>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<emphasis bold="true" box="[151,455,788,815]" pageId="8" pageNumber="109">
3.
<taxonomicName authority="Arnold, 1956" authorityName="Arnold" authorityYear="1956" box="[181,455,788,815]" class="Insecta" family="Mutillidae" genus="Corytilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[181,287,788,815]" italics="true" pageId="8" pageNumber="109">Corytilla</emphasis>
<bibRefCitation author="Arnold, G." box="[292,455,788,814]" pageId="8" pageNumber="109" pagination="52 - 77" refId="ref25000" refString="Arnold, G. (1956) New species of African Hymenoptera. No. 12. Occasional Papers of the National Museum of Southern Rhodesia, 3 (21 B), 52 - 77." type="journal article" year="1956">Arnold, 1956</bibRefCitation>
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</emphasis>
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</paragraph>
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<paragraph blockId="8.[151,455,788,851]" box="[151,267,824,851]" pageId="8" pageNumber="109">
(
<figureCitation box="[159,259,824,851]" captionStart="FIGURES 16" captionStartId="9.[152,270,1647,1671]" captionTargetBox="[151,1436,342,1619]" captionTargetId="figure-105@9.[151,1436,342,1621]" captionTargetPageId="9" captionText="FIGURES 16. 1. Andreimyrme long Lelej, 1995, ♁, holotype, lateral habitus; 23. A. rong Williams in Okayasu et al., 2021, ♀, holotype, habitus (2. Dorsal view, 3. Lateral view); 4a, 4b. Antennotilla phoebe (Péringuey, 1899), ♁: (4a. Lateral view, 4b. Dorsal view); 56. Corytilla pallidipes Arnold, 1956, ♁ (5. Head, lateral view, 6. Lateral habitus). (Photographs by V. Loktionov (1), D. Brothers (4b), and K. Williams (2, 3, 4a, 5, 6))." figureDoi="http://doi.org/10.5281/zenodo.7575310" httpUri="https://zenodo.org/record/7575310/files/figure.png" pageId="8" pageNumber="109">Figs 56</figureCitation>
)
</paragraph>
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<subSubSection pageId="8" pageNumber="109" type="reference_group">
<paragraph blockId="8.[151,1437,893,950]" pageId="8" pageNumber="109">
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<treatmentCitation author="Arnold, G." box="[151,421,893,917]" page="63" pageId="8" pageNumber="109" year="1956">
<taxonomicName authority="Arnold, 1956: 63" authorityName="Arnold" authorityPageNumber="63" authorityYear="1956" box="[151,421,893,917]" class="Insecta" genus="Corytilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis box="[151,243,893,917]" italics="true" pageId="8" pageNumber="109">Corytilla</emphasis>
<bibRefCitation author="Arnold, G." box="[247,421,893,917]" pageId="8" pageNumber="109" pagination="52 - 77" refId="ref25000" refString="Arnold, G. (1956) New species of African Hymenoptera. No. 12. Occasional Papers of the National Museum of Southern Rhodesia, 3 (21 B), 52 - 77." type="journal article" year="1956">Arnold, 1956: 63</bibRefCitation>
</taxonomicName>
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, figs 12, 12a, 12b, ♁; 1962: 846, figs 2, 2a, ♁,
<specimenCount box="[893,913,893,916]" pageId="8" pageNumber="109" type="female"></specimenCount>
;
<treatmentCitation author="Lelej, A. S. &amp; Brothers, D. J." box="[924,1192,893,917]" page="17" pageId="8" pageNumber="109" year="2008">
<bibRefCitation author="Lelej, A. S. &amp; Brothers, D. J." box="[924,1192,893,917]" pageId="8" pageNumber="109" pagination="1 - 79" refId="ref27054" refString="Lelej, A. S. &amp; Brothers, D. J. (2008) The genus-group names of Mutillidae (Hymenoptera) and their type species, with a new genus, new name, new synonymies, new combinations and lectotypifications. Zootaxa, 1889 (1), 1 - 79. https: // doi. org / 10.11646 / zootaxa. 1889.1.1" type="journal article" year="2008">Lelej &amp; Brothers 2008: 17</bibRefCitation>
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;
<treatmentCitation author="Brothers, D. J. &amp; Lelej, A. S." page="95" pageId="8" pageNumber="109" year="2017">
<bibRefCitation author="Brothers, D. J. &amp; Lelej, A. S." pageId="8" pageNumber="109" pagination="1 - 97" refId="ref25501" refString="Brothers, D. J. &amp; Lelej, A. S. (2017) Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses. Journal of Hymenoptera Research, 60, 1 - 97. https: // doi. org / 10.3897 / jhr. 60.20091" type="journal article" year="2017">Brothers &amp; Lelej 2017: 95</bibRefCitation>
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, ♁,
<specimenCount box="[269,289,924,947]" pageId="8" pageNumber="109" type="female"></specimenCount>
;
<treatmentCitation author="Pagliano, G. &amp; Brothers, D. J. &amp; Cambra, R. &amp; Lelej, A. S. &amp; Lo Cascio, P. &amp; Matteini Palmerini, M. &amp; Scaramozzino, P. L. &amp; Williams, K. A. &amp; Romano, M." box="[301,558,923,948]" page="169" pageId="8" pageNumber="109" year="2020">
<bibRefCitation author="Pagliano, G. &amp; Brothers, D. J. &amp; Cambra, R. &amp; Lelej, A. S. &amp; Lo Cascio, P. &amp; Matteini Palmerini, M. &amp; Scaramozzino, P. L. &amp; Williams, K. A. &amp; Romano, M." box="[301,558,923,948]" pageId="8" pageNumber="109" pagination="5 - 425" refId="ref28648" refString="Pagliano, G., Brothers, D. J., Cambra, R., Lelej, A. S., Lo Cascio, P., Matteini Palmerini, M., Scaramozzino, P. L., Williams, K. A. &amp; Romano, M. (2020 [&quot; 2018 &quot;]) Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species. Bollettino del Museo Regionale di Scienze Naturali di Torin o, 36 (1 - 2), 5 - 425." type="journal article" year="2020">
Pagliano
<emphasis box="[396,448,924,948]" italics="true" pageId="8" pageNumber="109">et al.</emphasis>
2020: 169
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.
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</paragraph>
</subSubSection>
<subSubSection box="[151,994,993,1022]" pageId="8" pageNumber="109" type="type_taxon">
<paragraph blockId="8.[151,1437,993,1524]" box="[151,994,993,1022]" pageId="8" pageNumber="109">
<emphasis bold="true" box="[151,299,993,1020]" pageId="8" pageNumber="109">
<typeStatus box="[151,210,994,1020]" pageId="8" pageNumber="109">Type</typeStatus>
species
</emphasis>
<taxonomicName authority="Arnold, 1956" authorityName="Arnold" authorityYear="1956" box="[306,683,994,1021]" class="Insecta" genus="Corytilla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="pallidipes">
<emphasis box="[306,524,994,1020]" italics="true" pageId="8" pageNumber="109">Corytilla pallidipes</emphasis>
<bibRefCitation author="Arnold, G." box="[530,683,994,1021]" pageId="8" pageNumber="109" pagination="52 - 77" refId="ref25000" refString="Arnold, G. (1956) New species of African Hymenoptera. No. 12. Occasional Papers of the National Museum of Southern Rhodesia, 3 (21 B), 52 - 77." type="journal article" year="1956">Arnold, 1956</bibRefCitation>
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, ♁, by original designation.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="109" type="diagnosis">
<paragraph blockId="8.[151,1437,993,1524]" pageId="8" pageNumber="109">
<emphasis bold="true" box="[199,314,1029,1055]" pageId="8" pageNumber="109">Diagnosis</emphasis>
. MALE. Mandible acuminated apically, beneath with large subbasal tooth. F2 a little shorter than F3. Vertex with raised subtriangular carinated platform between posterior ocelli. Mesonotum posterolaterally with full length notauli and short parapdis, hind angles produced into short blunt teeth. T2 with lateral felt lines, S2 without any traces of lateral felt lines. Forewings with very large radial and three radiomedial cells. FEMALE. Mandibles falcate and edentate with acute apex. F2 1.25 × F1 and equal in length F3. Mesosoma little wider behind than in front, one-fifth longer than wide in front. Scutellar scale lacking. T2 with oblong spot of pale golden setae, apical margin and also that of T3 with narrow fringe of same setae. Pygidial plate marginate, the basal two-thirds closely and longitudinally striate, the apical third smooth.
</paragraph>
<paragraph blockId="8.[151,1437,993,1524]" pageId="8" pageNumber="109">
<emphasis bold="true" box="[199,517,1317,1344]" pageId="8" pageNumber="109">Diversity and distribution.</emphasis>
Four species, three based on males only and one from both sexes, are recognized in the Afrotropical
<collectingRegion box="[365,450,1354,1380]" country="Iraq" pageId="8" pageNumber="109">Region</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="109" type="discussion">
<paragraph blockId="8.[151,1437,993,1524]" pageId="8" pageNumber="109">
<emphasis bold="true" box="[199,308,1390,1416]" pageId="8" pageNumber="109">Remarks</emphasis>
. Each of the four species in this genus is known from males, but only one species is known from the female as well. There are no available keys to separate all the species. The males are relatively abundant in Malaise trap samples from southern Africa (KAW, pers. obs.), but we have never seen a female that fits
<bibRefCitation author="Arnold, G." box="[1254,1436,1462,1488]" pageId="8" pageNumber="109" pagination="844 - 855" refId="ref25037" refString="Arnold, G. (1962) New species of African Hymenoptera. No. 16. Occasional Papers of the National Museum of Southern Rhodesia, 3 (26 B), 844 - 855." type="journal article" year="1962">Arnolds (1962)</bibRefCitation>
diagnosis.
</paragraph>
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</treatment>
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