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<document id="F96F69F717EEFD4D2408BFCC33A7CAF9" ID-CLB-Dataset="22824" ID-DOI="10.11646/zootaxa.4764.1.1" ID-GBIF-Dataset="55f6b110-0a37-45fe-be1c-34e9a035fe0e" ID-ISSN="1175-5334" ID-Zenodo-Dep="3764923" ID-ZooBank="urn:lsid:zoobank.org:pub:53C5289A-156C-4AF1-B84A-73099A332C05" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1587733789776" checkinUser="carolina" docAuthor="Van Ofwegen, Leen P." docDate="2020" docId="03BCB7172314FFEAFF3580D9FE1AFD45" docLanguage="en" docName="zootaxa.4764.1.1.pdf.imf" docOrigin="Zootaxa 4764 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Litophyton brewerensis Van 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="59" masterDocId="FF85CF6F2321FFD0FFA2856EFFC1FFB1" masterDocTitle="The genus Litophyton Forskål, 1775 (Octocorallia: Alcyonacea: Nephtheidae) from Australia" masterLastPageNumber="131" masterPageNumber="1" pageNumber="54" updateTime="1698837010817" updateUser="ExternalLinkService">
<mods:mods id="6C4F69BB720AA970AC6B4C436B7423B1" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0CF9544B7E909150B4E65C3C300D5B12">
<mods:title id="7261D7F58F5C28A880AE8842924FF205">The genus Litophyton Forskål, 1775 (Octocorallia: Alcyonacea: Nephtheidae) from Australia</mods:title>
</mods:titleInfo>
<mods:name id="EA7D295BC8F0BC178C7D3025D8730EBF" type="personal">
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<mods:namePart id="BB7D87F763273D99970921F59666FAE7">Van Ofwegen, Leen P.</mods:namePart>
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<mods:typeOfResource id="94DC1DA95D9129BF2E00D4574712D3BB">text</mods:typeOfResource>
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<mods:title id="401ECCB196D3D9EE9F4A0B0DD1D156CD">Zootaxa</mods:title>
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<mods:part id="47E0E0245BF58DDB6BD1402FDEE94FB6">
<mods:date id="B8360E07757A0F043072AD45C156DA4A">2020</mods:date>
<mods:detail id="861C257118836602CC1ADADE233BEE7D" type="pubDate">
<mods:number id="7468724B9BFF59C71881B390BE93F8A0">2020-04-16</mods:number>
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<mods:number id="4ACD1F4C1754884E92DE67471255FA32">4764</mods:number>
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<mods:classification id="03AEBCF9C71CAE37E93BC471BFE48154">journal article</mods:classification>
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<mods:identifier id="57F2E783E34CD37C995FB6885350F0D4" type="DOI">10.11646/zootaxa.4764.1.1</mods:identifier>
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<treatment id="03BCB7172314FFEAFF3580D9FE1AFD45" ID-DOI="http://doi.org/10.5281/zenodo.3803538" ID-GBIF-Taxon="163737717" ID-Zenodo-Dep="3803538" LSID="urn:lsid:plazi:treatment:03BCB7172314FFEAFF3580D9FE1AFD45" httpUri="http://treatment.plazi.org/id/03BCB7172314FFEAFF3580D9FE1AFD45" lastPageId="58" lastPageNumber="59" pageId="53" pageNumber="54">
<subSubSection id="C30F558A2314FFE5FF3580D9FDCFFA60" box="[151,526,1463,1489]" pageId="53" pageNumber="54" type="nomenclature">
<paragraph id="8BAA06012314FFE5FF3580D9FDCFFA60" blockId="53.[151,526,1463,1525]" box="[151,526,1463,1489]" pageId="53" pageNumber="54">
<heading id="D0E2B16D2314FFE5FF3580D9FDCFFA60" box="[151,526,1463,1489]" pageId="53" pageNumber="54">
<taxonomicName id="4C157D822314FFE5FF3580D9FE6BFA60" ID-CoL="866MH" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[151,426,1463,1489]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="53" pageNumber="54" phylum="Cnidaria" rank="species" species="brewerensis" status="sp. nov.">
<emphasis id="B961DA132314FFE5FF3580D9FE6BFA60" box="[151,426,1463,1489]" italics="true" pageId="53" pageNumber="54">Litophyton brewerensis</emphasis>
</taxonomicName>
<emphasis id="B961DA132314FFE5FE1380D9FDCFFA60" bold="true" box="[433,526,1463,1489]" pageId="53" pageNumber="54">
<taxonomicNameLabel id="A25267682314FFE5FE1380D9FDCFFA60" box="[433,526,1463,1489]" pageId="53" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30F558A2314FFE5FF3580B5FE4EFA44" box="[151,399,1499,1525]" pageId="53" pageNumber="54" type="description">
<paragraph id="8BAA06012314FFE5FF3580B5FE4EFA44" blockId="53.[151,526,1463,1525]" box="[151,399,1499,1525]" pageId="53" pageNumber="54">
(
<figureCitation id="132E1A842314FFE5FF3D80B5FEC7FA44" box="[159,262,1499,1525]" captionStart="FIGURE 41" captionStartId="54.[151,250,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@54.[178,1409,181,1919]" captionTargetPageId="54" captionText="FIGURE 41. Litophyton specimens: a-b, L. brewerensis n. sp., a, holotype NTM C4050, b, paratype NTM C2248; c-d, L. legiopolypum (Verseveldt &amp; Alderslade, 1982), c, NTM C713, d, NTM C714; e, L. oxleyensis n. sp., holotype NTM C4669; all natural size." figureDoi="http://doi.org/10.5281/zenodo.3765011" httpUri="https://zenodo.org/record/3765011/files/figure.png" pageId="53" pageNumber="54">Figs. 41a</figureCitation>
,
<figureCitation id="132E1A842314FFE5FEB380B5FEFAFA44" box="[273,315,1499,1525]" captionStart="FIGURE 42" captionStartId="55.[151,250,1910,1935]" captionTargetBox="[187,1390,185,1880]" captionTargetId="figure@55.[178,1409,181,1884]" captionTargetPageId="55" captionText="FIGURE 42. Lateral views polyp armature of Litophyton species: a, L. brewerensis n. sp., holotype NTM C4050; b, L. legiopolypum (Verseveldt &amp; Alderslade, 1982) holotype AMG 14806; c, L. oxleyensis n. sp., holotype NTM C4669; d, L. simplex n. sp., holotype NTM C3949; e, L. snakeensis n. sp., holotype NTM C10439; f, L. squamatum n. sp., holotype NTM C 4136; g-h, L. compactum (Verseveldt, 1966) holotype RMNH 2236 (after Verseveldt; magnification was not indicated by him)." figureDoi="http://doi.org/10.5281/zenodo.3765013" httpUri="https://zenodo.org/record/3765013/files/figure.png" pageId="53" pageNumber="54">42a</figureCitation>
,
<figureCitation id="132E1A842314FFE5FEE480B5FE46FA44" box="[326,391,1499,1525]" captionStart-0="FIGURE 43" captionStart-1="FIGURE 44" captionStartId-0="56.[151,250,1945,1970]" captionStartId-1="57.[151,250,1910,1935]" captionTargetBox-0="[178,1409,181,1919]" captionTargetBox-1="[190,1397,181,1884]" captionTargetId-0="figure@56.[178,1409,181,1919]" captionTargetId-1="figure@57.[190,1397,181,1884]" captionTargetPageId-0="56" captionTargetPageId-1="57" captionText-0="FIGURE 43. Litophyton brewerensis n. sp., holotype NTM C4050; a, abaxial point spindles; b, lateral, adaxial and tentacle scales; c, supporting bundle spindle; d, sclerites of surface layer top of stalk; e, sclerites of surface layer base of stalk; f-g, spindles of interior base of stalk; h, tuberculation of interior base stalk spindle. Scale at a also applies to c-e, g." captionText-1="FIGURE 44. Litophyton brewerensis n. sp., paratype NTM C2939; a, abaxial point spindles; b, lateral, adaxial and tentacle scales; c, polyp stalk rods; d, supporting bundle spindle; e-f, sclerites of surface layer top of stalk; g, sclerites of surface layer base of stalk; h-i, spindles of interior base of stalk; j, tuberculation of interior base stalk spindle. Scale at a also applies to d, f, g, i; scale at b also applies to e." figureDoi-0="http://doi.org/10.5281/zenodo.3765015" figureDoi-1="http://doi.org/10.5281/zenodo.3765017" httpUri-0="https://zenodo.org/record/3765015/files/figure.png" httpUri-1="https://zenodo.org/record/3765017/files/figure.png" pageId="53" pageNumber="54">43-44</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C30F558A2314FFE5FF35834CFE04F940" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="8BAA06012314FFE5FF35834CFE04F940" blockId="53.[151,1437,1570,2030]" pageId="53" pageNumber="54">
<emphasis id="B961DA132314FFE5FF35834CFEB3F98C" bold="true" box="[151,370,1570,1597]" pageId="53" pageNumber="54">Material examined</emphasis>
.
<materialsCitation id="3B7D0C5C2314FFE5FEDC834DFE71F9D0" ID-GBIF-Occurrence="2608549227" collectingDate="1981-02-07" collectionCode="NTM" collectorName="Z. Dinesen" elevation="8" location="John Brewer" municipality="Great Barrier Reef" pageId="53" pageNumber="54" specimenCode="NTM C4050" specimenCount="1" stateProvince="Queensland" typeStatus="holotype">
<typeStatus id="54AEB8A32314FFE5FEDC834DFE25F98C" box="[382,484,1571,1597]" pageId="53" pageNumber="54" type="holotype">Holotype</typeStatus>
<specimenCode id="DBB3AE7A2314FFE5FE4B834AFDB9F98C" box="[489,632,1570,1598]" pageId="53" pageNumber="54">NTM C4050</specimenCode>
,
<collectingRegion id="49D1C8E32314FFE5FD23834DFCC4F98C" box="[641,773,1571,1597]" country="Australia" name="Queensland" pageId="53" pageNumber="54">Queensland</collectingRegion>
,
<collectingMunicipality id="6BCE9C7B2314FFE5FCAC834DFC19F98C" box="[782,984,1571,1597]" pageId="53" pageNumber="54">Great Barrier Reef</collectingMunicipality>
,
<location id="8ECA50DA2314FFE5FC43834DFBABF98F" LSID="urn:lsid:plazi:treatment:03BCB7172314FFEAFF3580D9FE1AFD45:8ECA50DA2314FFE5FC43834DFBABF98F" box="[993,1130,1571,1598]" municipality="Great Barrier Reef" name="John Brewer" pageId="53" pageNumber="54" stateProvince="Queensland">John Brewer</location>
reef, depth
<quantity id="4CEDABE42314FFE5FB4A834CFAF6F98F" box="[1256,1335,1570,1598]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="53" pageNumber="54" unit="m" value="7.5" valueMax="10.0" valueMin="5.0">
<elevation id="0038E1322314FFE5FB4A834CFAF6F98F" box="[1256,1335,1570,1598]" metricMagnitude="0" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="53" pageNumber="54" unit="m" value="7.5" valueMax="10.0" valueMin="5.0">5-10 m</elevation>
</quantity>
,
<date id="FFAB20C12314FFE5FAE2834AFF39F9D0" pageId="53" pageNumber="54" value="1981-02-07">
<collectingDate id="EFEFD9292314FFE5FAE2834AFF39F9D0" pageId="53" pageNumber="54" value="1981-02-07">7 February 1981</collectingDate>
</date>
, coll.
<collectorName id="26E063D72314FFE5FE948326FE71F9D0" box="[310,432,1607,1634]" pageId="53" pageNumber="54">Z. Dinesen</collectorName>
</materialsCitation>
;
<typeStatus id="54AEB8A32314FFE5FE1B8326FDE4F9D3" box="[441,549,1608,1634]" pageId="53" pageNumber="54" type="paratype">paratypes</typeStatus>
: NTM C02236-NTM C2238,
<collectingRegion id="49D1C8E32314FFE5FCCD8329FC37F9D0" box="[879,1014,1607,1633]" country="Australia" name="Queensland" pageId="53" pageNumber="54">Queensland</collectingRegion>
, Briggs reef, Bommie, surface,
<date id="FFAB20C12314FFE5FAF18329FF13F934" pageId="53" pageNumber="54" value="1978-03">March 1978</date>
, coll. P. Alderslade; NTM C2248,
<collectingRegion id="49D1C8E32314FFE5FDC78305FD28F934" box="[613,745,1643,1669]" country="Australia" name="Queensland" pageId="53" pageNumber="54">Queensland</collectingRegion>
, Briggs-Sudbury reef,
<date id="FFAB20C12314FFE5FC4B8305FBB5F934" box="[1001,1140,1643,1669]" pageId="53" pageNumber="54" value="1978-03">March 1978</date>
, coll. P. Alderslade; NTM C2260-NTM C2261,
<collectingRegion id="49D1C8E32314FFE5FE2B83E1FDD1F918" box="[393,528,1679,1705]" country="Australia" name="Queensland" pageId="53" pageNumber="54">Queensland</collectingRegion>
, Masthead reef,
<date id="FFAB20C12314FFE5FD6A83E1FCBDF918" box="[712,892,1679,1705]" pageId="53" pageNumber="54" value="1974-08-31">31 August 1974</date>
, coll. P. Alderslade;? NTM C2939,
<collectingRegion id="49D1C8E32314FFE5FAB383E1FA59F918" box="[1297,1432,1679,1705]" country="Australia" name="Queensland" pageId="53" pageNumber="54">Queensland</collectingRegion>
, Orpheus island, Pioneer Bay, depth
<quantity id="4CEDABE42314FFE5FD9783DCFD56F97F" box="[565,663,1714,1742]" metricMagnitude="1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="53" pageNumber="54" unit="m" value="12.5" valueMax="15.0" valueMin="10.0">10-15 m</quantity>
, coll. J. Coll; NTM C10003, cf.
<emphasis id="B961DA132314FFE5FBB183DDFB84F97C" box="[1043,1093,1715,1741]" italics="true" pageId="53" pageNumber="54">alba</emphasis>
;
<collectingRegion id="49D1C8E32314FFE5FBF483DDFB1CF97C" box="[1110,1245,1715,1741]" country="Australia" name="Queensland" pageId="53" pageNumber="54">Queensland</collectingRegion>
, Orpheus island, Pioneer Bay, coll. P. Alino.
</paragraph>
</subSubSection>
<subSubSection id="C30F558A2314FFEAFF658394FAA4FED9" lastPageId="58" lastPageNumber="59" pageId="53" pageNumber="54" type="description">
<paragraph id="8BAA06012314FFE5FF658394FC00F888" blockId="53.[151,1437,1570,2030]" pageId="53" pageNumber="54">
<emphasis id="B961DA132314FFE5FF658394FE98F8A4" bold="true" box="[199,345,1786,1813]" pageId="53" pageNumber="54">Description.</emphasis>
The
<typeStatus id="54AEB8A32314FFE5FE308395FE35F8A4" box="[402,500,1787,1813]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
NTM C4050 is
<quantity id="4CEDABE42314FFE5FD0E8395FD22F8A7" box="[684,739,1787,1814]" metricMagnitude="-2" metricUnit="m" metricValue="8.0" pageId="53" pageNumber="54" unit="cm" value="8.0">8 cm</quantity>
high and wide, the stalk is
<quantity id="4CEDABE42314FFE5FBB58394FBA2F8A7" box="[1047,1123,1786,1814]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="53" pageNumber="54" unit="cm" value="1.5">1.5 cm</quantity>
high (
<figureCitation id="132E1A842314FFE5FB098395FACDF8A4" box="[1195,1292,1787,1813]" captionStart="FIGURE 41" captionStartId="54.[151,250,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@54.[178,1409,181,1919]" captionTargetPageId="54" captionText="FIGURE 41. Litophyton specimens: a-b, L. brewerensis n. sp., a, holotype NTM C4050, b, paratype NTM C2248; c-d, L. legiopolypum (Verseveldt &amp; Alderslade, 1982), c, NTM C713, d, NTM C714; e, L. oxleyensis n. sp., holotype NTM C4669; all natural size." figureDoi="http://doi.org/10.5281/zenodo.3765011" httpUri="https://zenodo.org/record/3765011/files/figure.png" pageId="53" pageNumber="54">Fig. 41a</figureCitation>
). The polyps are up to
<quantity id="4CEDABE42314FFE5FEA28271FEA5F88B" box="[256,356,1823,1850]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="53" pageNumber="54" unit="mm" value="0.8">0.80 mm</quantity>
wide and
<quantity id="4CEDABE42314FFE5FE7A8271FDFDF88B" box="[472,572,1823,1850]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="53" pageNumber="54" unit="mm" value="0.9">0.90 mm</quantity>
high, and grouped in long catkins.
</paragraph>
<paragraph id="8BAA06012314FFE5FF65822DFD73F85F" blockId="53.[151,1437,1570,2030]" pageId="53" pageNumber="54">
<emphasis id="B961DA132314FFE5FF65822DFED5F8EC" bold="true" box="[199,276,1859,1885]" pageId="53" pageNumber="54">Polyps</emphasis>
. Without clear arrangement of sclerites (
<figureCitation id="132E1A842314FFE5FD47822DFC86F8EC" box="[741,839,1859,1885]" captionStart="FIGURE 42" captionStartId="55.[151,250,1910,1935]" captionTargetBox="[187,1390,185,1880]" captionTargetId="figure@55.[178,1409,181,1884]" captionTargetPageId="55" captionText="FIGURE 42. Lateral views polyp armature of Litophyton species: a, L. brewerensis n. sp., holotype NTM C4050; b, L. legiopolypum (Verseveldt &amp; Alderslade, 1982) holotype AMG 14806; c, L. oxleyensis n. sp., holotype NTM C4669; d, L. simplex n. sp., holotype NTM C3949; e, L. snakeensis n. sp., holotype NTM C10439; f, L. squamatum n. sp., holotype NTM C 4136; g-h, L. compactum (Verseveldt, 1966) holotype RMNH 2236 (after Verseveldt; magnification was not indicated by him)." figureDoi="http://doi.org/10.5281/zenodo.3765013" httpUri="https://zenodo.org/record/3765013/files/figure.png" pageId="53" pageNumber="54">Fig. 42a</figureCitation>
). Abaxial point sclerites are spindles with simple tubercles, up to
<quantity id="4CEDABE42314FFE5FE8C8209FE4EF833" box="[302,399,1895,1922]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="53" pageNumber="54" unit="mm" value="0.4">0.40 mm</quantity>
long (
<figureCitation id="132E1A842314FFE5FE768209FDF0F830" box="[468,561,1895,1921]" captionStart="FIGURE 43" captionStartId="56.[151,250,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@56.[178,1409,181,1919]" captionTargetPageId="56" captionText="FIGURE 43. Litophyton brewerensis n. sp., holotype NTM C4050; a, abaxial point spindles; b, lateral, adaxial and tentacle scales; c, supporting bundle spindle; d, sclerites of surface layer top of stalk; e, sclerites of surface layer base of stalk; f-g, spindles of interior base of stalk; h, tuberculation of interior base stalk spindle. Scale at a also applies to c-e, g." figureDoi="http://doi.org/10.5281/zenodo.3765015" httpUri="https://zenodo.org/record/3765015/files/figure.png" pageId="53" pageNumber="54">Fig. 43a</figureCitation>
), laterally and adaxially scales and rods are present, which are also present in the tentacles and polyp stalk (
<figureCitation id="132E1A842314FFE5FE1482E5FDD7F817" box="[438,534,1931,1958]" captionStart="FIGURE 43" captionStartId="56.[151,250,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@56.[178,1409,181,1919]" captionTargetPageId="56" captionText="FIGURE 43. Litophyton brewerensis n. sp., holotype NTM C4050; a, abaxial point spindles; b, lateral, adaxial and tentacle scales; c, supporting bundle spindle; d, sclerites of surface layer top of stalk; e, sclerites of surface layer base of stalk; f-g, spindles of interior base of stalk; h, tuberculation of interior base stalk spindle. Scale at a also applies to c-e, g." figureDoi="http://doi.org/10.5281/zenodo.3765015" httpUri="https://zenodo.org/record/3765015/files/figure.png" pageId="53" pageNumber="54">Fig. 43b</figureCitation>
). Supporting bundle without projecting spindles or one projecting for a maximum distance of
<quantity id="4CEDABE42314FFE5FEB782C1FEB8F87B" box="[277,377,1967,1994]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="53" pageNumber="54" unit="mm" value="0.1">0.10 mm</quantity>
. Spindles of the supporting bundle up to
<quantity id="4CEDABE42314FFE5FC9E82C1FC5CF87B" box="[828,925,1967,1994]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" pageId="53" pageNumber="54" unit="mm" value="1.4">1.40 mm</quantity>
long, with simple tubercles (
<figureCitation id="132E1A842314FFE5FB7882C1FAF9F878" box="[1242,1336,1967,1993]" captionStart="FIGURE 43" captionStartId="56.[151,250,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@56.[178,1409,181,1919]" captionTargetPageId="56" captionText="FIGURE 43. Litophyton brewerensis n. sp., holotype NTM C4050; a, abaxial point spindles; b, lateral, adaxial and tentacle scales; c, supporting bundle spindle; d, sclerites of surface layer top of stalk; e, sclerites of surface layer base of stalk; f-g, spindles of interior base of stalk; h, tuberculation of interior base stalk spindle. Scale at a also applies to c-e, g." figureDoi="http://doi.org/10.5281/zenodo.3765015" httpUri="https://zenodo.org/record/3765015/files/figure.png" pageId="53" pageNumber="54">Fig. 43c</figureCitation>
). Several polyp sclerites have an optically black inner core.
</paragraph>
<caption id="DF6A56892317FFE6FF3582F7FED3F84B" ID-DOI="http://doi.org/10.5281/zenodo.3765011" ID-Zenodo-Dep="3765011" httpUri="https://zenodo.org/record/3765011/files/figure.png" pageId="54" pageNumber="55" startId="54.[151,250,1945,1970]" targetBox="[178,1409,181,1919]" targetPageId="54">
<paragraph id="8BAA06012317FFE6FF3582F7FED3F84B" blockId="54.[151,1436,1945,2042]" pageId="54" pageNumber="55">
<emphasis id="B961DA132317FFE6FF3582F7FEDDF803" bold="true" box="[151,284,1945,1970]" pageId="54" pageNumber="55">FIGURE 41</emphasis>
.
<taxonomicName id="4C157D822317FFE6FE8E82F4FE5BF803" authorityName="Forskal" authorityYear="1775" box="[300,410,1946,1970]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="54" pageNumber="55" phylum="Cnidaria" rank="genus">
<emphasis id="B961DA132317FFE6FE8E82F4FE5BF803" box="[300,410,1946,1970]" italics="true" pageId="54" pageNumber="55">Litophyton</emphasis>
</taxonomicName>
specimens: a-b,
<taxonomicName id="4C157D822317FFE6FDEE82F4FD23F803" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[588,738,1946,1970]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="54" pageNumber="55" phylum="Cnidaria" rank="species" species="brewerensis" status="sp. nov.">
<emphasis id="B961DA132317FFE6FDEE82F4FD9BF800" box="[588,602,1946,1969]" italics="true" pageId="54" pageNumber="55">L</emphasis>
.
<emphasis id="B961DA132317FFE6FDCB82F4FD23F803" box="[617,738,1946,1970]" italics="true" pageId="54" pageNumber="55">brewerensis</emphasis>
</taxonomicName>
<emphasis id="B961DA132317FFE6FD4982F4FCECF803" bold="true" box="[747,813,1946,1970]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="A25267682317FFE6FD4982F4FCE9F803" box="[747,808,1946,1970]" pageId="54" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
a, holotype NTM C4050, b, paratype NTM C2248; c-d,
<taxonomicName id="4C157D822317FFE6FA2A82F4FDAAF867" authority="(Verseveldt &amp; Alderslade, 1982)" authorityName="Great Barrier Reef" baseAuthorityName="Verseveldt &amp; Alderslade" baseAuthorityYear="1982" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="54" pageNumber="55" phylum="Cnidaria" rank="species" species="legiopolypum">
<emphasis id="B961DA132317FFE6FA2A82F4FA57F800" box="[1416,1430,1946,1969]" italics="true" pageId="54" pageNumber="55">L</emphasis>
.
<emphasis id="B961DA132317FFE6FF3582D0FEE0F867" box="[151,289,1982,2006]" italics="true" pageId="54" pageNumber="55">legiopolypum</emphasis>
(
<bibRefCitation id="EF847BF02317FFE6FE8D82D3FDA5F867" author="Verseveldt &amp; Alderslade" box="[303,612,1981,2006]" firstAuthor="Verseveldt" pageId="54" pageNumber="55" pagination="619 - 647" refId="ref42721" refString="Verseveldt, J. &amp; Alderslade, P. (1982) Descriptions of types and other Alcyonacean material (Coelenterata: Octocorallia) in the Australian Museum, Sydney. Records of the Australian Museum, 34, 619 - 647. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.245" type="journal article" year="1982">Verseveldt &amp; Alderslade, 1982</bibRefCitation>
)
</taxonomicName>
, c, NTM C713, d, NTM C714; e,
<taxonomicName id="4C157D822317FFE6FC6082D0FB82F867" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[962,1091,1982,2006]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="54" pageNumber="55" phylum="Cnidaria" rank="species" species="oxleyensis" status="sp. nov.">
<emphasis id="B961DA132317FFE6FC6082D0FC11F864" box="[962,976,1982,2005]" italics="true" pageId="54" pageNumber="55">L</emphasis>
.
<emphasis id="B961DA132317FFE6FC7C82D0FB82F867" box="[990,1091,1982,2006]" italics="true" pageId="54" pageNumber="55">oxleyensis</emphasis>
</taxonomicName>
<emphasis id="B961DA132317FFE6FBE882D0FB4BF867" bold="true" box="[1098,1162,1982,2006]" pageId="54" pageNumber="55">
<taxonomicNameLabel id="A25267682317FFE6FBE882D0FB44F867" box="[1098,1157,1982,2006]" pageId="54" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
holotype NTM C4669; all natural size.
</paragraph>
</caption>
<caption id="DF6A56892316FFE7FF358218FACFF84A" ID-DOI="http://doi.org/10.5281/zenodo.3765013" ID-Zenodo-Dep="3765013" httpUri="https://zenodo.org/record/3765013/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,250,1910,1935]" targetBox="[187,1390,185,1880]" targetPageId="55">
<paragraph id="8BAA06012316FFE7FF358218FACFF84A" blockId="55.[151,1436,1910,2043]" pageId="55" pageNumber="56">
<emphasis id="B961DA132316FFE7FF358218FEDBF83E" bold="true" box="[151,282,1910,1935]" pageId="55" pageNumber="56">FIGURE 42</emphasis>
. Lateral views polyp armature of
<taxonomicName id="4C157D822316FFE7FDD88219FD29F83E" authorityName="Forskal" authorityYear="1775" box="[634,744,1911,1935]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="55" pageNumber="56" phylum="Cnidaria" rank="genus">
<emphasis id="B961DA132316FFE7FDD88219FD29F83E" box="[634,744,1911,1935]" italics="true" pageId="55" pageNumber="56">Litophyton</emphasis>
</taxonomicName>
species: a,
<taxonomicName id="4C157D822316FFE7FCC28216FC37F83E" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[864,1014,1911,1935]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="55" pageNumber="56" phylum="Cnidaria" rank="species" species="brewerensis" status="sp. nov.">
<emphasis id="B961DA132316FFE7FCC28216FCAFF83E" box="[864,878,1912,1935]" italics="true" pageId="55" pageNumber="56">L</emphasis>
.
<emphasis id="B961DA132316FFE7FCDF8219FC37F83E" box="[893,1014,1911,1935]" italics="true" pageId="55" pageNumber="56">brewerensis</emphasis>
</taxonomicName>
<emphasis id="B961DA132316FFE7FC5F8219FBFCF83E" bold="true" box="[1021,1085,1911,1935]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="A25267682316FFE7FC5F8219FBF9F83E" box="[1021,1080,1911,1935]" pageId="55" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
holotype NTM C4050; b,
<taxonomicName id="4C157D822316FFE7FAF08216FD82F802" authority="(Verseveldt &amp; Alderslade, 1982)" authorityName="Great Barrier Reef" baseAuthorityName="Verseveldt &amp; Alderslade" baseAuthorityYear="1982" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="55" pageNumber="56" phylum="Cnidaria" rank="species" species="legiopolypum">
<emphasis id="B961DA132316FFE7FAF08216FAA1F83E" box="[1362,1376,1912,1935]" italics="true" pageId="55" pageNumber="56">L</emphasis>
.
<emphasis id="B961DA132316FFE7FACC8219FF3AF802" italics="true" pageId="55" pageNumber="56">legiopolypum</emphasis>
(
<bibRefCitation id="EF847BF02316FFE7FEAB82F4FDFDF802" author="Verseveldt &amp; Alderslade" box="[265,572,1946,1971]" firstAuthor="Verseveldt" pageId="55" pageNumber="56" pagination="619 - 647" refId="ref42721" refString="Verseveldt, J. &amp; Alderslade, P. (1982) Descriptions of types and other Alcyonacean material (Coelenterata: Octocorallia) in the Australian Museum, Sydney. Records of the Australian Museum, 34, 619 - 647. https: // doi. org / 10.3853 / j. 0067 - 1975.34.1982.245" type="journal article" year="1982">Verseveldt &amp; Alderslade, 1982</bibRefCitation>
)
</taxonomicName>
holotype AMG 14806; c,
<taxonomicName id="4C157D822316FFE7FCEB82F2FC08F802" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[841,969,1947,1971]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="55" pageNumber="56" phylum="Cnidaria" rank="species" species="oxleyensis" status="sp. nov.">
<emphasis id="B961DA132316FFE7FCEB82F2FC96F802" box="[841,855,1948,1971]" italics="true" pageId="55" pageNumber="56">L</emphasis>
.
<emphasis id="B961DA132316FFE7FCC682F5FC08F802" box="[868,969,1947,1971]" italics="true" pageId="55" pageNumber="56">oxleyensis</emphasis>
</taxonomicName>
<emphasis id="B961DA132316FFE7FC7282F5FBD1F802" bold="true" box="[976,1040,1947,1971]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="A25267682316FFE7FC7282F5FBCAF802" box="[976,1035,1947,1971]" pageId="55" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
holotype NTM C4669; d,
<taxonomicName id="4C157D822316FFE7FAB982F2FA43F802" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[1307,1410,1947,1971]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="55" pageNumber="56" phylum="Cnidaria" rank="species" species="simplex" status="sp. nov.">
<emphasis id="B961DA132316FFE7FAB982F2FAE8F802" box="[1307,1321,1948,1971]" italics="true" pageId="55" pageNumber="56">L</emphasis>
.
<emphasis id="B961DA132316FFE7FA9482F5FA43F802" box="[1334,1410,1947,1971]" italics="true" pageId="55" pageNumber="56">simplex</emphasis>
</taxonomicName>
<emphasis id="B961DA132316FFE7FA2A82F5FF7AF866" bold="true" pageId="55" pageNumber="56">
<taxonomicNameLabel id="A25267682316FFE7FA2A82F5FF77F866" pageId="55" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
holotype NTM C3949; e,
<taxonomicName id="4C157D822316FFE7FE6582AEFD8CF866" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[455,589,1983,2007]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="55" pageNumber="56" phylum="Cnidaria" rank="species" species="snakeensis" status="sp. nov.">
<emphasis id="B961DA132316FFE7FE6582AEFD8CF866" box="[455,589,1983,2007]" italics="true" pageId="55" pageNumber="56">L. snakeensis</emphasis>
</taxonomicName>
<emphasis id="B961DA132316FFE7FDF182D1FD53F866" bold="true" box="[595,658,1983,2007]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="A25267682316FFE7FDF182D1FD4CF866" box="[595,653,1983,2007]" pageId="55" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
holotype NTM C10439; f,
<taxonomicName id="4C157D822316FFE7FC0582AEFBF7F866" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[935,1078,1984,2007]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="55" pageNumber="56" phylum="Cnidaria" rank="species" species="squamatum" status="sp. nov.">
<emphasis id="B961DA132316FFE7FC0582AEFBF7F866" box="[935,1078,1984,2007]" italics="true" pageId="55" pageNumber="56">L. squamatum</emphasis>
</taxonomicName>
<emphasis id="B961DA132316FFE7FB9E82D1FBBBF866" bold="true" box="[1084,1146,1983,2007]" pageId="55" pageNumber="56">
<taxonomicNameLabel id="A25267682316FFE7FB9E82D1FBB7F866" box="[1084,1142,1983,2007]" pageId="55" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
holotype NTM C 4136; g-h,
<taxonomicName id="4C157D822316FFE7FF35828AFE1EF84A" authority="(Verseveldt, 1966)" baseAuthorityName="Verseveldt" baseAuthorityYear="1966" box="[151,479,2018,2043]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="55" pageNumber="56" phylum="Cnidaria" rank="species" species="compactum">
<emphasis id="B961DA132316FFE7FF35828AFEE2F84A" box="[151,291,2020,2043]" italics="true" pageId="55" pageNumber="56">L. compactum</emphasis>
(
<bibRefCitation id="EF847BF02316FFE7FE92828CFE19F84A" author="Verseveldt" box="[304,472,2018,2043]" firstAuthor="Verseveldt" pageId="55" pageNumber="56" refId="ref42688" refString="Verseveldt, J. (1966) Octocorallia from the Malay Archipelago (Part II). Biological Results of the Snellius Expedition XXII. Zoologische Verhandelingen Leiden, 80,1 - 107." type="book" year="1966">Verseveldt, 1966</bibRefCitation>
)
</taxonomicName>
holotype RMNH 2236 (after Verseveldt; magnification was not indicated by him).
</paragraph>
</caption>
<caption id="DF6A56892319FFE8FF3582F7FBBCF84B" ID-DOI="http://doi.org/10.5281/zenodo.3765015" ID-Zenodo-Dep="3765015" httpUri="https://zenodo.org/record/3765015/files/figure.png" pageId="56" pageNumber="57" startId="56.[151,250,1945,1970]" targetBox="[178,1409,181,1919]" targetPageId="56">
<paragraph id="8BAA06012319FFE8FF3582F7FBBCF84B" blockId="56.[151,1437,1945,2042]" pageId="56" pageNumber="57">
<emphasis id="B961DA132319FFE8FF3582F7FEDDF803" bold="true" box="[151,284,1945,1970]" pageId="56" pageNumber="57">FIGURE 43</emphasis>
.
<taxonomicName id="4C157D822319FFE8FE8E82F4FDDDF803" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[300,540,1946,1970]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="56" pageNumber="57" phylum="Cnidaria" rank="species" species="brewerensis" status="sp. nov.">
<emphasis id="B961DA132319FFE8FE8E82F4FDDDF803" box="[300,540,1946,1970]" italics="true" pageId="56" pageNumber="57">Litophyton brewerensis</emphasis>
</taxonomicName>
<emphasis id="B961DA132319FFE8FD8682F4FDA7F803" bold="true" box="[548,614,1946,1970]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="A25267682319FFE8FD8682F4FDA0F803" box="[548,609,1946,1970]" pageId="56" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
holotype NTM C4050; a, abaxial point spindles; b, lateral, adaxial and tentacle scales; c, supporting bundle spindle; d, sclerites of surface layer top of stalk; e, sclerites of surface layer base of stalk; f-g, spindles of interior base of stalk; h, tuberculation of interior base stalk spindle. Scale at a also applies to c-e, g.
</paragraph>
</caption>
<caption id="DF6A56892318FFE9FF358218FECCF84B" ID-DOI="http://doi.org/10.5281/zenodo.3765017" ID-Zenodo-Dep="3765017" httpUri="https://zenodo.org/record/3765017/files/figure.png" pageId="57" pageNumber="58" startId="57.[151,250,1910,1935]" targetBox="[190,1397,181,1884]" targetPageId="57">
<paragraph id="8BAA06012318FFE9FF358218FECCF84B" blockId="57.[151,1437,1910,2043]" pageId="57" pageNumber="58">
<emphasis id="B961DA132318FFE9FF358218FED9F83E" bold="true" box="[151,280,1910,1935]" pageId="57" pageNumber="58">FIGURE 44</emphasis>
.
<taxonomicName id="4C157D822318FFE9FE868219FDCEF83E" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[292,527,1911,1935]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="57" pageNumber="58" phylum="Cnidaria" rank="species" species="brewerensis" status="sp. nov.">
<emphasis id="B961DA132318FFE9FE868219FDCEF83E" box="[292,527,1911,1935]" italics="true" pageId="57" pageNumber="58">Litophyton brewerensis</emphasis>
</taxonomicName>
<emphasis id="B961DA132318FFE9FDB68219FD93F83E" bold="true" box="[532,594,1911,1935]" pageId="57" pageNumber="58">
<taxonomicNameLabel id="A25267682318FFE9FDB68219FD8CF83E" box="[532,589,1911,1935]" pageId="57" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
,
</emphasis>
paratype NTM C2939; a, abaxial point spindles; b, lateral, adaxial and tentacle scales; c, polyp stalk rods; d, supporting bundle spindle; e-f, sclerites of surface layer top of stalk; g, sclerites of surface layer base of stalk; h-i, spindles of interior base of stalk; j, tuberculation of interior base stalk spindle. Scale at a also applies to d, f, g, i; scale at b also applies to e.
</paragraph>
</caption>
<paragraph id="8BAA0601231BFFEAFF6585F6FEB5FF69" blockId="58.[151,1437,152,756]" pageId="58" pageNumber="59">
<emphasis id="B961DA13231BFFEAFF6585F6FE0DFF02" bold="true" box="[199,460,152,179]" pageId="58" pageNumber="59">Surface layer top stalk</emphasis>
. Spindles and unilaterally spinose spindles, up to
<quantity id="4CEDABE4231BFFEAFC4485F7FB86FF02" box="[998,1095,153,179]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="58" pageNumber="59" unit="mm" value="0.6">0.60 mm</quantity>
long, with simple and complex tubercles (
<figureCitation id="132E1A84231BFFEAFEAA85D3FEA9FF69" box="[264,360,189,216]" captionStart="FIGURE 43" captionStartId="56.[151,250,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@56.[178,1409,181,1919]" captionTargetPageId="56" captionText="FIGURE 43. Litophyton brewerensis n. sp., holotype NTM C4050; a, abaxial point spindles; b, lateral, adaxial and tentacle scales; c, supporting bundle spindle; d, sclerites of surface layer top of stalk; e, sclerites of surface layer base of stalk; f-g, spindles of interior base of stalk; h, tuberculation of interior base stalk spindle. Scale at a also applies to c-e, g." figureDoi="http://doi.org/10.5281/zenodo.3765015" httpUri="https://zenodo.org/record/3765015/files/figure.png" pageId="58" pageNumber="59">Fig. 43d</figureCitation>
).
</paragraph>
<paragraph id="8BAA0601231BFFEAFF65858EFE10FEF2" blockId="58.[151,1437,152,756]" pageId="58" pageNumber="59">
<emphasis id="B961DA13231BFFEAFF65858EFE25FF4A" bold="true" box="[199,484,224,251]" pageId="58" pageNumber="59">Surface layer base stalk</emphasis>
. Capstans and derivatives of capstans,
<quantity id="4CEDABE4231BFFEAFC02858EFBFCFF4A" box="[928,1085,224,251]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="58" pageNumber="59" unit="mm" value="0.1" valueMax="0.15" valueMin="0.05">0.05-0.15 mm</quantity>
long. Spindles and unilaterally spinose spindles are also present, up to
<quantity id="4CEDABE4231BFFEAFDE3846BFD63FEAE" box="[577,674,261,287]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="58" pageNumber="59" unit="mm" value="0.7">0.70 mm</quantity>
long (
<figureCitation id="132E1A84231BFFEAFD4A846BFC84FEAE" box="[744,837,261,287]" captionStart="FIGURE 43" captionStartId="56.[151,250,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@56.[178,1409,181,1919]" captionTargetPageId="56" captionText="FIGURE 43. Litophyton brewerensis n. sp., holotype NTM C4050; a, abaxial point spindles; b, lateral, adaxial and tentacle scales; c, supporting bundle spindle; d, sclerites of surface layer top of stalk; e, sclerites of surface layer base of stalk; f-g, spindles of interior base of stalk; h, tuberculation of interior base stalk spindle. Scale at a also applies to c-e, g." figureDoi="http://doi.org/10.5281/zenodo.3765015" httpUri="https://zenodo.org/record/3765015/files/figure.png" pageId="58" pageNumber="59">Fig. 43e</figureCitation>
). The smaller sclerites have simple tubercles, the larger ones also complex tubercles.
</paragraph>
<paragraph id="8BAA0601231BFFEAFF658422FAA4FED9" blockId="58.[151,1437,152,756]" box="[199,1381,332,360]" pageId="58" pageNumber="59">
<emphasis id="B961DA13231BFFEAFF658422FEA3FED6" bold="true" box="[199,354,332,359]" pageId="58" pageNumber="59">Interior stalk</emphasis>
. Spindles, up to
<quantity id="4CEDABE4231BFFEAFDBA8423FDB8FED6" box="[536,633,333,359]" metricMagnitude="-3" metricUnit="m" metricValue="1.2" pageId="58" pageNumber="59" unit="mm" value="1.2">1.20 mm</quantity>
long, with pointed ends (
<figureCitation id="132E1A84231BFFEAFC2F8423FC1AFED9" box="[909,987,333,360]" captionStart="FIGURE 43" captionStartId="56.[151,250,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@56.[178,1409,181,1919]" captionTargetPageId="56" captionText="FIGURE 43. Litophyton brewerensis n. sp., holotype NTM C4050; a, abaxial point spindles; b, lateral, adaxial and tentacle scales; c, supporting bundle spindle; d, sclerites of surface layer top of stalk; e, sclerites of surface layer base of stalk; f-g, spindles of interior base of stalk; h, tuberculation of interior base stalk spindle. Scale at a also applies to c-e, g." figureDoi="http://doi.org/10.5281/zenodo.3765015" httpUri="https://zenodo.org/record/3765015/files/figure.png" pageId="58" pageNumber="59">Fig. 43</figureCitation>
f-g) and simple tubercles (
<figureCitation id="132E1A84231BFFEAFB5B8423FA99FED9" box="[1273,1368,333,360]" captionStart="FIGURE 43" captionStartId="56.[151,250,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@56.[178,1409,181,1919]" captionTargetPageId="56" captionText="FIGURE 43. Litophyton brewerensis n. sp., holotype NTM C4050; a, abaxial point spindles; b, lateral, adaxial and tentacle scales; c, supporting bundle spindle; d, sclerites of surface layer top of stalk; e, sclerites of surface layer base of stalk; f-g, spindles of interior base of stalk; h, tuberculation of interior base stalk spindle. Scale at a also applies to c-e, g." figureDoi="http://doi.org/10.5281/zenodo.3765015" httpUri="https://zenodo.org/record/3765015/files/figure.png" pageId="58" pageNumber="59">Fig. 43h</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30F558A231BFFEAFF65841FFD55FE3A" box="[199,660,369,395]" pageId="58" pageNumber="59" type="etymology">
<paragraph id="8BAA0601231BFFEAFF65841FFD55FE3A" blockId="58.[151,1437,152,756]" box="[199,660,369,395]" pageId="58" pageNumber="59">
<emphasis id="B961DA13231BFFEAFF65841FFE8AFE3A" bold="true" box="[199,331,369,395]" pageId="58" pageNumber="59">Etymology.</emphasis>
Named after the
<typeStatus id="54AEB8A3231BFFEAFDA5841FFDF7FE3A" box="[519,566,369,395]" pageId="58" pageNumber="59">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C30F558A231BFFEAFF6584FAFB6EFE1E" box="[199,1199,404,431]" pageId="58" pageNumber="59" type="distribution">
<paragraph id="8BAA0601231BFFEAFF6584FAFB6EFE1E" blockId="58.[151,1437,152,756]" box="[199,1199,404,431]" pageId="58" pageNumber="59">
<emphasis id="B961DA13231BFFEAFF6584FAFE94FE1E" bold="true" box="[199,341,404,431]" pageId="58" pageNumber="59">Distribution</emphasis>
. Great Barrier Reef: John Brewer reef, Briggs-Sudbury reef,? Orpheus island.
</paragraph>
</subSubSection>
<subSubSection id="C30F558A231BFFEAFF6584D7FE1AFD45" pageId="58" pageNumber="59" type="discussion">
<paragraph id="8BAA0601231BFFEAFF6584D7FE9AFE46" blockId="58.[151,1437,152,756]" pageId="58" pageNumber="59">
<emphasis id="B961DA13231BFFEAFF6584D7FEF8FE62" bold="true" box="[199,313,441,467]" pageId="58" pageNumber="59">Remarks.</emphasis>
NTM C2248 differs from the other material in colony shape (
<figureCitation id="132E1A84231BFFEAFC7D84D7FBFCFE65" box="[991,1085,441,468]" captionStart="FIGURE 41" captionStartId="54.[151,250,1945,1970]" captionTargetBox="[178,1409,181,1919]" captionTargetId="figure@54.[178,1409,181,1919]" captionTargetPageId="54" captionText="FIGURE 41. Litophyton specimens: a-b, L. brewerensis n. sp., a, holotype NTM C4050, b, paratype NTM C2248; c-d, L. legiopolypum (Verseveldt &amp; Alderslade, 1982), c, NTM C713, d, NTM C714; e, L. oxleyensis n. sp., holotype NTM C4669; all natural size." figureDoi="http://doi.org/10.5281/zenodo.3765011" httpUri="https://zenodo.org/record/3765011/files/figure.png" pageId="58" pageNumber="59">Fig. 41b</figureCitation>
), probably the colony was dried out at some stage.
</paragraph>
<paragraph id="8BAA0601231BFFEAFF65876FFC7EFD8E" blockId="58.[151,1437,152,756]" pageId="58" pageNumber="59">
The species mostly resembles
<taxonomicName id="4C157D82231BFFEAFDB1876CFD64FDAA" authority="Van, 2020" authorityName="Van" authorityYear="2020" box="[531,677,513,539]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="58" pageNumber="59" phylum="Cnidaria" rank="species" species="snakeensis" status="sp. nov.">
<emphasis id="B961DA13231BFFEAFDB1876CFDE3FDAA" box="[531,546,514,539]" italics="true" pageId="58" pageNumber="59">L</emphasis>
.
<emphasis id="B961DA13231BFFEAFD92876FFD64FDAA" box="[560,677,513,539]" italics="true" pageId="58" pageNumber="59">snakeensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2526768231BFFEAFD09876FFCC0FDAA" box="[683,769,513,539]" pageId="58" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
, but differs in having more slender interior stalk spindles and many capstans and derivatives of capstans in the surface layer of the stalk.
</paragraph>
<paragraph id="8BAA0601231BFFEAFF658727FC6FFDD2" blockId="58.[151,1437,152,756]" box="[199,942,585,611]" pageId="58" pageNumber="59">NTM C2236-NTM C2238 were examined using temporary slides.</paragraph>
<paragraph id="8BAA0601231BFFEAFF658703FE1AFD45" blockId="58.[151,1437,152,756]" pageId="58" pageNumber="59">
NTM C2939 and NTM C10003 are included in this species, but with some doubts. They differ from the other specimens in having numerous rods in polyp stalk (
<figureCitation id="132E1A84231BFFEAFD6B87FFFCE5FD1A" box="[713,804,657,683]" captionStart="FIGURE 44" captionStartId="57.[151,250,1910,1935]" captionTargetBox="[190,1397,181,1884]" captionTargetId="figure@57.[190,1397,181,1884]" captionTargetPageId="57" captionText="FIGURE 44. Litophyton brewerensis n. sp., paratype NTM C2939; a, abaxial point spindles; b, lateral, adaxial and tentacle scales; c, polyp stalk rods; d, supporting bundle spindle; e-f, sclerites of surface layer top of stalk; g, sclerites of surface layer base of stalk; h-i, spindles of interior base of stalk; j, tuberculation of interior base stalk spindle. Scale at a also applies to d, f, g, i; scale at b also applies to e." figureDoi="http://doi.org/10.5281/zenodo.3765017" httpUri="https://zenodo.org/record/3765017/files/figure.png" pageId="58" pageNumber="59">Fig. 44c</figureCitation>
) instead of scales. In this aspect these specimens resemble
<taxonomicName id="4C157D82231BFFEAFF3587D8FE8BFD7E" authorityName="Great Barrier Reef" baseAuthorityName="Verseveldt &amp; Alderslade" baseAuthorityYear="1982" box="[151,330,693,719]" class="Anthozoa" family="Nephtheidae" genus="Litophyton" kingdom="Animalia" order="Alcyonacea" pageId="58" pageNumber="59" phylum="Cnidaria" rank="species" species="legiopolypum">
<emphasis id="B961DA13231BFFEAFF3587D8FF67FD7E" box="[151,166,694,719]" italics="true" pageId="58" pageNumber="59">L</emphasis>
.
<emphasis id="B961DA13231BFFEAFF1787DBFE8BFD7E" box="[181,330,693,719]" italics="true" pageId="58" pageNumber="59">legiopolypum</emphasis>
</taxonomicName>
(see below). However, they lack the short branched spindles with rough tubercles in the base of the stalk of the colony (
<figureCitation id="132E1A84231BFFEAFECD87B7FE0EFD45" box="[367,463,729,756]" captionStart="FIGURE 44" captionStartId="57.[151,250,1910,1935]" captionTargetBox="[190,1397,181,1884]" captionTargetId="figure@57.[190,1397,181,1884]" captionTargetPageId="57" captionText="FIGURE 44. Litophyton brewerensis n. sp., paratype NTM C2939; a, abaxial point spindles; b, lateral, adaxial and tentacle scales; c, polyp stalk rods; d, supporting bundle spindle; e-f, sclerites of surface layer top of stalk; g, sclerites of surface layer base of stalk; h-i, spindles of interior base of stalk; j, tuberculation of interior base stalk spindle. Scale at a also applies to d, f, g, i; scale at b also applies to e." figureDoi="http://doi.org/10.5281/zenodo.3765017" httpUri="https://zenodo.org/record/3765017/files/figure.png" pageId="58" pageNumber="59">Fig. 44h</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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