treatments-xml/data/03/BC/87/03BC87D0FFA9FFE6FF6EF83FF838D6B2.xml
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<document id="B43A0DB0C87E9CBA7760629248A2FEBE" ID-DOI="10.11646/zootaxa.4095.1.1" ID-GBIF-Dataset="f1972764-7444-46ba-97b1-517daec66224" ID-ISSN="1175-5326" ID-Zenodo-Dep="261389" ID-ZooBank="690FFEBE-F71B-4EFD-865A-944D81A12897" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459240875275" checkinUser="plazi" docAuthor="P. R. Pugh" docDate="2016" docId="03BC87D0FFA9FFE6FF6EF83FF838D6B2" docLanguage="en" docName="zootaxa_4095_1.pdf.imf" docOrigin="Zootaxa 4095 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Cardianecta parchelion Pugh, 2016, sp. nov." docType="treatment" docVersion="17" lastPageNumber="58" masterDocId="FF85FFA8FF99FFDFFFF9FFACF93CD442" masterDocTitle="A synopsis of the Family Cordagalmatidae fam. nov. (Cnidaria, Siphonophora, Physonectae)" masterLastPageNumber="64" masterPageNumber="1" pageNumber="49" updateTime="1698186612131" updateUser="plazi">
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<mods:title id="53956B631B4172D3B99D4E740870BC47">A synopsis of the Family Cordagalmatidae fam. nov. (Cnidaria, Siphonophora, Physonectae)</mods:title>
</mods:titleInfo>
<mods:name id="01E8CF64015F75542A4331F43FFA0C76" type="personal">
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<mods:namePart id="4BFA0BC3ECB1BA369576007A5B28B813">P. R. Pugh</mods:namePart>
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<mods:title id="56AA0E0DC05FC079BDBB3C2E6AD628E2">Zootaxa</mods:title>
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<mods:part id="166D6AD445C89D75AA78DE77DD1B0367">
<mods:date id="96ACD835B201ABDC8C5C310DCEB33602">2016</mods:date>
<mods:detail id="60C4502078921EDF32483816C73B829C" type="volume">
<mods:number id="D3621C49C8E9EB4D114A64EE568E82E5">4095</mods:number>
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<mods:identifier id="D6160FE61B26671AE12255902CCE0302" type="DOI">10.11646/zootaxa.4095.1.1</mods:identifier>
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<treatment id="03BC87D0FFA9FFE6FF6EF83FF838D6B2" ID-DOI="http://doi.org/10.5281/zenodo.6087827" ID-GBIF-Taxon="119319697" ID-Zenodo-Dep="6087827" LSID="urn:lsid:plazi:treatment:03BC87D0FFA9FFE6FF6EF83FF838D6B2" httpUri="http://treatment.plazi.org/id/03BC87D0FFA9FFE6FF6EF83FF838D6B2" lastPageId="57" lastPageNumber="58" pageId="48" pageNumber="49">
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<paragraph id="8BAA36C6FFA9FFEFFF6EF83FFB25D3EC" blockId="48.[151,537,1939,1966]" box="[151,537,1939,1966]" pageId="48" pageNumber="49">
<heading id="D0E281AAFFA9FFEFFF6EF83FFB25D3EC" bold="true" box="[151,537,1939,1966]" fontSize="11" level="1" pageId="48" pageNumber="49" reason="1">
<emphasis id="B961EAD4FFA9FFEFFF6EF83FFB25D3EC" bold="true" box="[151,537,1939,1966]" pageId="48" pageNumber="49">
<taxonomicName id="4C154D45FFA9FFEFFF6EF83FF888D3EF" box="[151,436,1939,1965]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="48" pageNumber="49" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFA9FFEFFF6EF83FF888D3EF" bold="true" box="[151,436,1939,1965]" italics="true" pageId="48" pageNumber="49">Cardianecta parchelion</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257AFFFA9FFEFFE45F838FB25D3EC" box="[444,537,1940,1966]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30F654DFFA9FFEEFF6EF876F9EED655" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49" type="diagnosis">
<paragraph id="8BAA36C6FFA9FFEEFF6EF876F9EED655" blockId="48.[151,1436,2010,2035]" lastBlockId="49.[151,1437,151,789]" lastPageId="49" lastPageNumber="50" pageId="48" pageNumber="49">
<emphasis id="B961EAD4FFA9FFEFFF6EF876F82DD3B1" bold="true" box="[151,273,2010,2035]" pageId="48" pageNumber="49">Diagnosis.</emphasis>
Most mature nectophores relatively large and flimsy, with typical heart-shaped appearance. Younger ones with distinct notch on lateral wings. Long ascending and descending mantle canals of approximately equal length. No pigmentation. Bracts in enantiomorphic pairs; broad, leaf-like, tapering toward the proximal end; lower surface flat, upper domed. Indistinct longitudinal ridge from distal, median point to about half the length of the bract. Distal point, in young bracts, bearing nematocysts. Bracteal canal in contact with lower surface of the bract for about one-third its length; distal part, inflected into the mesogloea; relatively long in youngest bracts but gradually shortens to a short process in mature bracts. Tentilla of unique form, with involucrum forming two lateral flaps, and inflated terminal process bearing two asymmetrically placed horns, one terminal the other sub-terminal. At some stage in its development, the cnidoband spirally coiled and containing only one kind of nematocyst, probably mastigophores. Desmonemes (?) present in the terminal process, particularly in the horns. Proximally attached palpons possess a palpacle although often lost before the palpon becomes fully developed in the preserved state.
</paragraph>
</subSubSection>
<subSubSection id="C30F654DFFA8FFEEFF3EFD88FD48D756" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="8BAA36C6FFA8FFEEFF3EFD88F813D622" blockId="49.[151,1437,151,789]" pageId="49" pageNumber="50">
<emphasis id="B961EAD4FFA8FFEEFF3EFD88F88DD67F" bold="true" box="[199,433,548,573]" pageId="49" pageNumber="50">Material examined.</emphasis>
<specimenCount id="9D13FD4FFFA8FFEEFE41FD88FB7ED67F" box="[440,578,548,573]" pageId="49" pageNumber="50" type="generic">5 specimens</specimenCount>
collected by the Johnson-Sea-Link (
<collectionCode id="ED04AE03FFA8FFEEFC26FD88FD31D67E" box="[991,1037,548,572]" collectionName="JSL" pageId="49" pageNumber="50">JSL</collectionCode>
) submersibles, all in the vicinity of
<collectingCountry id="F3027656FFA8FFEEFF6EFDEBF817D622" box="[151,299,583,608]" name="Bahamas" pageId="49" pageNumber="50">the Bahamas</collectingCountry>
:
</paragraph>
<paragraph id="8BAA36C6FFA8FFEEFF6EFDC0FD48D6C6" blockId="49.[151,1437,151,789]" box="[151,1140,620,645]" pageId="49" pageNumber="50">
<materialsCitation id="3B7D3C9BFFA8FFEEFF6EFDC0FD48D6C6" ID-GBIF-Occurrence="1262419317" box="[151,1140,620,645]" collectingCountry="Bahamas" collectingDate="1984-10-28" elevation="0" latitude="25.908333" location="JSL II Dive 981" longLatPrecision="124" longitude="-77.29667" pageId="49" pageNumber="50" specimenCount="1">
<location id="8ECA601DFFA8FFEEFF6EFDC0F876D6C6" LSID="urn:lsid:plazi:treatment:03BC87D0FFA9FFE6FF6EF83FF838D6B2:8ECA601DFFA8FFEEFF6EFDC0F876D6C6" box="[151,330,620,645]" latitude="25.908333" longLatPrecision="124" longitude="-77.29667" name="JSL II Dive 981" pageId="49" pageNumber="50">JSL II Dive 981</location>
<geoCoordinate id="EE215001FFA8FFEEFE78FDC0F8C5D6C7" box="[385,505,620,645]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="92" value="25.908333">25°54.5N</geoCoordinate>
,
<geoCoordinate id="EE215001FFA8FFEEFDFCFDC0FBBCD6C6" box="[517,640,620,644]" direction="west" orientation="longitude" pageId="49" pageNumber="50" precision="92" value="-77.29667">77°17.8W</geoCoordinate>
<date id="FFAB1006FFA8FFEEFD22FDC0FA56D6C6" box="[731,874,620,644]" pageId="49" pageNumber="50" value="1984-10-28">
<collectingDate id="EFEFE9EEFFA8FFEEFD22FDC0FA56D6C6" box="[731,874,620,644]" pageId="49" pageNumber="50" value="1984-10-28">28-Oct-1984</collectingDate>
</date>
<elevation id="0038D1F5FFA8FFEEFC10FDC0FD48D6C6" box="[1001,1140,620,645]" pageId="49" pageNumber="50">
depth
<quantity id="4CED9B23FFA8FFEEFBD7FDC0FD48D6C6" box="[1070,1140,620,645]" metricMagnitude="2" metricUnit="m" metricValue="6.4" pageId="49" pageNumber="50" unit="m" value="640.0">640 m</quantity>
</elevation>
</materialsCitation>
</paragraph>
<paragraph id="8BAA36C6FFA8FFEEFF6EFD3CFD48D6E5" blockId="49.[151,1437,151,789]" box="[151,1140,655,680]" pageId="49" pageNumber="50">
<materialsCitation id="3B7D3C9BFFA8FFEEFF6EFD3CFD48D6E5" ID-GBIF-Occurrence="1262419323" box="[151,1140,655,680]" collectingCountry="Bahamas" collectingDate="1984-10-31" elevation="0" latitude="25.366667" location="JSL II Dive 998" longLatPrecision="124" longitude="-77.86" pageId="49" pageNumber="50" specimenCount="1">
<location id="8ECA601DFFA8FFEEFF6EFD3CF876D6EA" LSID="urn:lsid:plazi:treatment:03BC87D0FFA9FFE6FF6EF83FF838D6B2:8ECA601DFFA8FFEEFF6EFD3CF876D6EA" box="[151,330,655,680]" latitude="25.366667" longLatPrecision="124" longitude="-77.86" name="JSL II Dive 998" pageId="49" pageNumber="50">JSL II Dive 998</location>
<geoCoordinate id="EE215001FFA8FFEEFE78FD23F8C5D6EA" box="[385,505,655,680]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="92" value="25.366667">25°22.0N</geoCoordinate>
,
<geoCoordinate id="EE215001FFA8FFEEFDFCFD23FBBCD6EA" box="[517,640,655,680]" direction="west" orientation="longitude" pageId="49" pageNumber="50" precision="92" value="-77.86">77°51.6W</geoCoordinate>
<date id="FFAB1006FFA8FFEEFD22FD3CFA56D6EA" box="[731,874,656,680]" pageId="49" pageNumber="50" value="1984-10-31">
<collectingDate id="EFEFE9EEFFA8FFEEFD22FD3CFA56D6EA" box="[731,874,656,680]" pageId="49" pageNumber="50" value="1984-10-31">31-Oct-1984</collectingDate>
</date>
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depth
<quantity id="4CED9B23FFA8FFEEFBD7FD23FD48D6E5" box="[1070,1140,655,680]" metricMagnitude="2" metricUnit="m" metricValue="6.42" pageId="49" pageNumber="50" unit="m" value="642.0">642 m</quantity>
</elevation>
</materialsCitation>
</paragraph>
<paragraph id="8BAA36C6FFA8FFEEFF6EFD18FD48D68E" blockId="49.[151,1437,151,789]" box="[151,1140,692,717]" pageId="49" pageNumber="50">
<materialsCitation id="3B7D3C9BFFA8FFEEFF6EFD18FD48D68E" ID-GBIF-Occurrence="1262419328" box="[151,1140,692,717]" collectingCountry="Bahamas" collectingDate="1989-11-03" elevation="0" latitude="26.246666" location="JSL I Dive 2633" longLatPrecision="124" longitude="-77.72833" pageId="49" pageNumber="50" specimenCount="1">
<location id="8ECA601DFFA8FFEEFF6EFD18F873D68F" LSID="urn:lsid:plazi:treatment:03BC87D0FFA9FFE6FF6EF83FF838D6B2:8ECA601DFFA8FFEEFF6EFD18F873D68F" box="[151,335,692,717]" latitude="26.246666" longLatPrecision="124" longitude="-77.72833" name="JSL I Dive 2633" pageId="49" pageNumber="50">JSL I Dive 2633</location>
<geoCoordinate id="EE215001FFA8FFEEFE78FD18F8C5D68F" box="[385,505,692,717]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="92" value="26.246666">26°14.8N</geoCoordinate>
,
<geoCoordinate id="EE215001FFA8FFEEFDFCFD18FBBCD68E" box="[517,640,692,716]" direction="west" orientation="longitude" pageId="49" pageNumber="50" precision="92" value="-77.72833">77°43.7W</geoCoordinate>
<date id="FFAB1006FFA8FFEEFD22FD18FA58D68E" box="[731,868,692,716]" pageId="49" pageNumber="50" value="1989-11-03">
<collectingDate id="EFEFE9EEFFA8FFEEFD22FD18FA58D68E" box="[731,868,692,716]" pageId="49" pageNumber="50" value="1989-11-03">3-Nov-1989</collectingDate>
</date>
<elevation id="0038D1F5FFA8FFEEFC10FD18FD48D68E" box="[1001,1140,692,717]" pageId="49" pageNumber="50">
depth
<quantity id="4CED9B23FFA8FFEEFBD7FD18FD48D68E" box="[1070,1140,692,716]" metricMagnitude="2" metricUnit="m" metricValue="8.84" pageId="49" pageNumber="50" unit="m" value="884.0">884 m</quantity>
</elevation>
</materialsCitation>
</paragraph>
<paragraph id="8BAA36C6FFA8FFEEFF6EFD74FD48D6AD" blockId="49.[151,1437,151,789]" box="[151,1140,727,752]" pageId="49" pageNumber="50">
<materialsCitation id="3B7D3C9BFFA8FFEEFF6EFD74FD48D6AD" ID-GBIF-Occurrence="1262419336" box="[151,1140,727,752]" collectingCountry="Bahamas" collectingDate="1989-11-08" elevation="0" latitude="26.038334" location="JSL I Dive 2643" longLatPrecision="124" longitude="-77.53333" pageId="49" pageNumber="50" specimenCount="1">
<location id="8ECA601DFFA8FFEEFF6EFD74F873D6B2" LSID="urn:lsid:plazi:treatment:03BC87D0FFA9FFE6FF6EF83FF838D6B2:8ECA601DFFA8FFEEFF6EFD74F873D6B2" box="[151,335,727,752]" latitude="26.038334" longLatPrecision="124" longitude="-77.53333" name="JSL I Dive 2643" pageId="49" pageNumber="50">JSL I Dive 2643</location>
<geoCoordinate id="EE215001FFA8FFEEFE78FD7BF8C5D6B2" box="[385,505,727,752]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="92" value="26.038334">26°02.3N</geoCoordinate>
,
<geoCoordinate id="EE215001FFA8FFEEFDFCFD74FBBCD6B2" box="[517,640,728,752]" direction="west" orientation="longitude" pageId="49" pageNumber="50" precision="92" value="-77.53333">77°32.0W</geoCoordinate>
<date id="FFAB1006FFA8FFEEFD22FD74FA58D6B2" box="[731,868,728,752]" pageId="49" pageNumber="50" value="1989-11-08">
<collectingDate id="EFEFE9EEFFA8FFEEFD22FD74FA58D6B2" box="[731,868,728,752]" pageId="49" pageNumber="50" value="1989-11-08">8-Nov-1989</collectingDate>
</date>
<elevation id="0038D1F5FFA8FFEEFC10FD7BFD48D6AD" box="[1001,1140,727,752]" pageId="49" pageNumber="50">
depth
<quantity id="4CED9B23FFA8FFEEFBD7FD7BFD48D6AD" box="[1070,1140,727,752]" metricMagnitude="2" metricUnit="m" metricValue="8.65" pageId="49" pageNumber="50" unit="m" value="865.0">865 m</quantity>
</elevation>
</materialsCitation>
</paragraph>
<paragraph id="8BAA36C6FFA8FFEEFF6EFD50FD48D756" blockId="49.[151,1437,151,789]" box="[151,1140,764,789]" pageId="49" pageNumber="50">
<materialsCitation id="3B7D3C9BFFA8FFEEFF6EFD50FD48D756" ID-GBIF-Occurrence="1262419334" box="[151,1140,764,789]" collectingCountry="Bahamas" collectingDate="1989-11-13" elevation="0" latitude="26.066668" location="JSL I Dive 2650" longLatPrecision="124" longitude="-77.558334" pageId="49" pageNumber="50" specimenCount="1">
<location id="8ECA601DFFA8FFEEFF6EFD50F873D757" LSID="urn:lsid:plazi:treatment:03BC87D0FFA9FFE6FF6EF83FF838D6B2:8ECA601DFFA8FFEEFF6EFD50F873D757" box="[151,335,764,789]" latitude="26.066668" longLatPrecision="124" longitude="-77.558334" name="JSL I Dive 2650" pageId="49" pageNumber="50">JSL I Dive 2650</location>
<geoCoordinate id="EE215001FFA8FFEEFE78FD50F8C5D757" box="[385,505,764,789]" direction="north" orientation="latitude" pageId="49" pageNumber="50" precision="92" value="26.066668">26°04.0N</geoCoordinate>
,
<geoCoordinate id="EE215001FFA8FFEEFDFCFD50FBBCD757" box="[517,640,764,789]" direction="west" orientation="longitude" pageId="49" pageNumber="50" precision="92" value="-77.558334">77°33.5W</geoCoordinate>
<collectingDate id="EFEFE9EEFFA8FFEEFD22FD50FA4ED756" box="[731,882,764,788]" pageId="49" pageNumber="50" value="1989-11-13">
<date id="FFAB1006FFA8FFEEFD22FD50FA4ED756" box="[731,882,764,788]" pageId="49" pageNumber="50" value="1989-11-13">13-Nov-1989</date>
</collectingDate>
<elevation id="0038D1F5FFA8FFEEFC10FD50FD48D756" box="[1001,1140,764,789]" pageId="49" pageNumber="50">
depth
<quantity id="4CED9B23FFA8FFEEFBD7FD50FD48D756" box="[1070,1140,764,789]" metricMagnitude="2" metricUnit="m" metricValue="7.57" pageId="49" pageNumber="50" unit="m" value="757.0">757 m</quantity>
</elevation>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30F654DFFA8FFEEFF3EFCE8FB39D052" pageId="49" pageNumber="50" type="materials_examined">
<paragraph id="8BAA36C6FFA8FFEEFF3EFCE8FB39D052" blockId="49.[151,1436,836,1112]" pageId="49" pageNumber="50">
<typeStatus id="54AE8864FFA8FFEEFF3EFCE8F812D71F" box="[199,302,836,861]" pageId="49" pageNumber="50" type="holotype">
<emphasis id="B961EAD4FFA8FFEEFF3EFCE8F812D71F" box="[199,302,836,861]" italics="true" pageId="49" pageNumber="50">Holotype</emphasis>
</typeStatus>
: The specimen collected during the Johnson-Sea-Link I Dive 2650 has been designated the
<typeStatus id="54AE8864FFA8FFEEFACAFCE8FCA4D71F" box="[1331,1432,836,861]" pageId="49" pageNumber="50" type="holotype">holotype</typeStatus>
, and will be deposited at the
<collectingCountry id="F3027656FFA8FFEEFE28FCCBFB5AD7C2" box="[465,614,871,896]" name="United States of America" pageId="49" pageNumber="50">United States</collectingCountry>
National Museum (Smithsonian Institution), Washington, DC.
<typeStatus id="54AE8864FFA8FFEEFF3EFC21F812D7E6" box="[199,302,909,932]" pageId="49" pageNumber="50" type="paratype">
<emphasis id="B961EAD4FFA8FFEEFF3EFC21F812D7E6" box="[199,302,909,932]" italics="true" pageId="49" pageNumber="50">Paratype</emphasis>
</typeStatus>
: The specimen collected during the Johnson-Sea-Link I Dive 2633 has been designated the
<typeStatus id="54AE8864FFA8FFEEFACCFC21FCA4D7E7" box="[1333,1432,909,933]" pageId="49" pageNumber="50" type="paratype">paratype</typeStatus>
, and will also be deposited at the
<collectingCountry id="F3027656FFA8FFEEFDFCFC03FBA6D78A" box="[517,666,943,968]" name="United States of America" pageId="49" pageNumber="50">United States</collectingCountry>
National Museum (Smithsonian Institution), Washington, DC. The remaining specimens will be placed in the collections of Dr Casey Dunn, at Brown University, Providence, Rhode
<collectingCountry id="F3027656FFA8FFEEFE8AFC5BF886D052" box="[371,442,1015,1040]" name="Iceland" pageId="49" pageNumber="50">Island</collectingCountry>
,
<collectingCountry id="F3027656FFA8FFEEFE3CFC54FB3DD052" box="[453,513,1016,1040]" name="United States of America" pageId="49" pageNumber="50">USA</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F654DFFA8FFE8FF3EFBB0FBBDD3B7" lastPageId="55" lastPageNumber="56" pageId="49" pageNumber="50" type="description">
<paragraph id="8BAA36C6FFA8FFEEFF3EFBB0FCACD01A" blockId="49.[151,1436,836,1112]" pageId="49" pageNumber="50">
<emphasis id="B961EAD4FFA8FFEEFF3EFBB0F865D077" bold="true" box="[199,345,1052,1077]" pageId="49" pageNumber="50">Description.</emphasis>
<emphasis id="B961EAD4FFA8FFEEFE9BFBB0FB28D077" box="[354,532,1052,1077]" italics="true" pageId="49" pageNumber="50">Pneumatophore</emphasis>
: The preserved pneumatophores were quite distorted, measuring c.
<quantity id="4CED9B23FFA8FFEEFADDFBB0FC41D076" box="[1316,1405,1052,1076]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="49" pageNumber="50" unit="mm" value="2.2">2.2 mm</quantity>
in length and
<quantity id="4CED9B23FFA8FFEEFEEAFB93F84BD015" box="[275,375,1087,1112]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="49" pageNumber="50" unit="mm" value="1.25">1.25 mm</quantity>
in diameter, but otherwise were quite featureless, without any obvious signs of pigmentation.
</paragraph>
<caption id="DF6A664EFFA8FFEEFF6EF807F9F7D3A2" httpUri="https://zenodo.org/record/261446/files/figure.png" pageId="49" pageNumber="50" targetBox="[277,1305,1167,1925]" targetPageId="49">
<paragraph id="8BAA36C6FFA8FFEEFF6EF807F9F7D3A2" blockId="49.[151,1435,1963,2016]" pageId="49" pageNumber="50">
<emphasis id="B961EAD4FFA8FFEEFF6EF807F81DD383" bold="true" box="[151,289,1963,1985]" pageId="49" pageNumber="50">FIGURE 58.</emphasis>
<taxonomicName id="4C154D45FFA8FFEEFED3F807FB21D383" box="[298,541,1963,1985]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="49" pageNumber="50" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFA8FFEEFED3F807FB21D383" box="[298,541,1963,1985]" italics="true" pageId="49" pageNumber="50">Cardianecta parchelion</emphasis>
</taxonomicName>
<emphasis id="B961EAD4FFA8FFEEFDDCF807FB4BD383" bold="true" box="[549,631,1963,1985]" pageId="49" pageNumber="50">
<taxonomicNameLabel id="A25257AFFFA8FFEEFDDCF807FB4BD383" box="[549,631,1963,1985]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Ostial view of large, flimsy nectophore from paratype specimen. Scale bar 1 mm..
</paragraph>
</caption>
<paragraph id="8BAA36C6FFABFFEDFF3EFF35FD8ED4ED" blockId="50.[151,1437,151,501]" box="[199,1202,151,176]" pageId="50" pageNumber="51">
<emphasis id="B961EAD4FFABFFEDFF3EFF35F87ED4F2" box="[199,322,153,176]" italics="true" pageId="50" pageNumber="51">Nectosome</emphasis>
: The nectophores were clearly budded from the ventral side of the nectosome.
</paragraph>
<paragraph id="8BAA36C6FFABFFEDFF3EFF10FD28D5B7" blockId="50.[151,1437,151,501]" pageId="50" pageNumber="51">
<emphasis id="B961EAD4FFABFFEDFF3EFF10F875D497" box="[199,329,188,213]" italics="true" pageId="50" pageNumber="51">Nectophore</emphasis>
: The preserved nectophores were heart-shaped measuring up to
<quantity id="4CED9B23FFABFFEDFBD8FF10FDBAD496" box="[1057,1158,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="8.75" pageId="50" pageNumber="51" unit="mm" value="8.75">8.75 mm</quantity>
in length and
<quantity id="4CED9B23FFABFFEDFAD1FF10FC43D496" box="[1320,1407,188,213]" metricMagnitude="-3" metricUnit="m" metricValue="9.6" pageId="50" pageNumber="51" unit="mm" value="9.6">9.6 mm</quantity>
in width, without any obvious pigmentation. The largest nectophores were very flimsy and generally distorted in preservation (
<figureCitation id="132E2A43FFABFFEDFECBFEA8F899D55F" box="[306,421,260,285]" captionStart="FIGURE 58" captionStartId="49.[151,250,1963,1985]" captionTargetBox="[277,1305,1167,1925]" captionTargetId="figure@49.[277,1306,1167,1925]" captionTargetPageId="49" captionText="FIGURE 58. Cardianecta parchelion sp. nov. Ostial view of large, flimsy nectophore from paratype specimen. Scale bar 1 mm .." httpUri="https://zenodo.org/record/261446/files/figure.png" pageId="50" pageNumber="51">Figure 58</figureCitation>
), although those of the
<typeStatus id="54AE8864FFABFFEDFD40FEA8FA27D55F" box="[697,795,260,285]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
specimen were better preserved. In ostial or axial view they were heart-shaped (
<figureCitation id="132E2A43FFABFFEDFE51FE8BFB24D502" box="[424,536,295,320]" captionStart="FIGURE 59" captionStartId="50.[151,250,1092,1114]" captionTargetBox="[190,1393,597,1063]" captionTargetId="figure@50.[184,1402,591,1067]" captionTargetPageId="50" captionText="FIGURE 59. Cardianecta parchelion sp. nov. A. Ostial and B. axial views of larger nectophore from holotype specimen. amc, dmc: ascending and descending mantle canal, respectively; larc, lrc, orc and urc: lateral, lower, ostial ring and upper radial canals, respectively; llr and ulr: lower and upper lateral ridges respectively; ns: nectosac; pedc: pedicular canal; tec: tract of ectodermal cells. Scale bar 2 mm." httpUri="https://zenodo.org/record/261447/files/figure.png" pageId="50" pageNumber="51">Figure 59</figureCitation>
), with a squarish ostium measuring c.
<quantity id="4CED9B23FFABFFEDFC3AFE84FD3FD57D" box="[963,1027,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="50" pageNumber="51" unit="mm" value="2.0">2 mm</quantity>
across. The nectosac, in lateral view (
<figureCitation id="132E2A43FFABFFEDFF67FEE0F87BD527" box="[158,327,332,357]" captionStart-0="FIGURE 60" captionStart-1="FIGURE 61" captionStart-2="FIGURE 62" captionStartId-0="50.[151,250,1763,1785]" captionStartId-1="51.[151,250,1496,1518]" captionStartId-2="52.[151,250,991,1013]" captionTargetBox-0="[219,1367,1292,1733]" captionTargetBox-1="[361,1225,517,1470]" captionTargetBox-2="[359,1232,350,954]" captionTargetId-0="figure@50.[215,1371,1289,1738]" captionTargetId-1="figure@51.[355,1231,514,1474]" captionTargetId-2="figure@52.[355,1239,342,957]" captionTargetPageId-0="50" captionTargetPageId-1="51" captionTargetPageId-2="52" captionText-0="FIGURE 60. Cardianecta parchelion sp. nov. A. ostial and B. lateral views of larger nectophore; C. lateral view of young nectophore, both from holotype specimen. n: notch in lateral wings. Scale bars A and B 2 mm, C 1 mm." captionText-1="FIGURE 61. Cardianecta parchelion sp. nov. Lateral (A), ostial (B), and ostio basal (C) views of younger nectophore from JSL I Dive 2643 specimen before preservation. See Figures 59 and 60 for legend. Scale bar 1 mm." captionText-2="FIGURE 62. Upper views of bracts of Cardianecta parchelion sp. nov. A. mature, and B. developing bracts from the holotype (scale bar 1 mm). The grey part of the bracteal canal is within the mesogloea. C. Very young bracts from the JSL I 2633 specimen. Proximal end at bottom. Scale bars 0. 5 mm." httpUri-0="https://zenodo.org/record/261448/files/figure.png" httpUri-1="https://zenodo.org/record/261449/files/figure.png" httpUri-2="https://zenodo.org/record/261450/files/figure.png" pageId="50" pageNumber="51">Figures 6062</figureCitation>
), was L-shaped, but deeply divided in the mid-line. The ascending and descending mantle canals were of approximately equal length. The pedicular canal passed straight to the nectosac where it gave rise to all four radial canals. The upper and lower canals were straight; the laterals were broadly looped and connected with the ostial ring canal at the upper corners of the squarish ostium, where a distinct tract of ectodermal cells extended obliquely up the sides of the nectophore (
<figureCitation id="132E2A43FFABFFEDFD9CFE70FBE9D5B7" box="[613,725,476,501]" captionStart="FIGURE 59" captionStartId="50.[151,250,1092,1114]" captionTargetBox="[190,1393,597,1063]" captionTargetId="figure@50.[184,1402,591,1067]" captionTargetPageId="50" captionText="FIGURE 59. Cardianecta parchelion sp. nov. A. Ostial and B. axial views of larger nectophore from holotype specimen. amc, dmc: ascending and descending mantle canal, respectively; larc, lrc, orc and urc: lateral, lower, ostial ring and upper radial canals, respectively; llr and ulr: lower and upper lateral ridges respectively; ns: nectosac; pedc: pedicular canal; tec: tract of ectodermal cells. Scale bar 2 mm." httpUri="https://zenodo.org/record/261447/files/figure.png" pageId="50" pageNumber="51">Figure 59</figureCitation>
,
<figureCitation id="132E2A43FFABFFEDFD18FE70FBC1D5B7" box="[737,765,476,501]" captionStart="FIGURE 61" captionStartId="51.[151,250,1496,1518]" captionTargetBox="[361,1225,517,1470]" captionTargetId="figure@51.[355,1231,514,1474]" captionTargetPageId="51" captionText="FIGURE 61. Cardianecta parchelion sp. nov. Lateral (A), ostial (B), and ostio basal (C) views of younger nectophore from JSL I Dive 2643 specimen before preservation. See Figures 59 and 60 for legend. Scale bar 1 mm." httpUri="https://zenodo.org/record/261449/files/figure.png" pageId="50" pageNumber="51">61</figureCitation>
<emphasis id="B961EAD4FFABFFEDFCFDFE70FA1AD5B7" bold="true" box="[772,806,476,501]" pageId="50" pageNumber="51">tec</emphasis>
) for a short distance.
</paragraph>
<caption id="DF6A664EFFABFFEDFF6EFBE8F8D2D0F4" httpUri="https://zenodo.org/record/261447/files/figure.png" pageId="50" pageNumber="51" targetBox="[190,1393,597,1063]" targetPageId="50">
<paragraph id="8BAA36C6FFABFFEDFF6EFBE8F8D2D0F4" blockId="50.[151,1436,1092,1206]" pageId="50" pageNumber="51">
<emphasis id="B961EAD4FFABFFEDFF6EFBE8F823D018" bold="true" box="[151,287,1092,1114]" pageId="50" pageNumber="51">FIGURE 59.</emphasis>
<taxonomicName id="4C154D45FFABFFEDFEDCFBE8FB2AD018" box="[293,534,1092,1114]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="50" pageNumber="51" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFABFFEDFEDCFBE8FB2AD018" box="[293,534,1092,1114]" italics="true" pageId="50" pageNumber="51">Cardianecta parchelion</emphasis>
</taxonomicName>
<emphasis id="B961EAD4FFABFFEDFDE5FBE8FBB8D018" bold="true" box="[540,644,1092,1114]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="A25257AFFFABFFEDFDE5FBE8FB50D018" box="[540,620,1092,1114]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
. Ostial and
<emphasis id="B961EAD4FFABFFEDFD07FBE8FA33D018" bold="true" box="[766,783,1092,1114]" pageId="50" pageNumber="51">B</emphasis>
. axial views of larger nectophore from holotype specimen.
<emphasis id="B961EAD4FFABFFEDFA90FBE8F9F9D03B" bold="true" pageId="50" pageNumber="51">amc, dmc</emphasis>
: ascending and descending mantle canal, respectively;
<emphasis id="B961EAD4FFABFFEDFCFBFBCFFAB4D03B" bold="true" box="[770,904,1123,1145]" pageId="50" pageNumber="51">larc, lrc, orc</emphasis>
and
<emphasis id="B961EAD4FFABFFEDFC45FBCFFADCD03B" bold="true" box="[956,992,1123,1145]" pageId="50" pageNumber="51">urc</emphasis>
: lateral, lower, ostial ring and upper radial canals, respectively;
<emphasis id="B961EAD4FFABFFEDFE94FB2EF8BAD0DA" bold="true" box="[365,390,1154,1176]" pageId="50" pageNumber="51">llr</emphasis>
and
<emphasis id="B961EAD4FFABFFEDFE43FB2EF8E6D0DA" bold="true" box="[442,474,1154,1176]" pageId="50" pageNumber="51">ulr</emphasis>
: lower and upper lateral ridges respectively;
<emphasis id="B961EAD4FFABFFEDFC54FB2EFAF9D0DA" bold="true" box="[941,965,1154,1176]" pageId="50" pageNumber="51">ns</emphasis>
: nectosac;
<emphasis id="B961EAD4FFABFFEDFBC0FB2EFD50D0DA" bold="true" box="[1081,1132,1154,1176]" pageId="50" pageNumber="51">pedc</emphasis>
: pedicular canal;
<emphasis id="B961EAD4FFABFFEDFADDFB2EFC7FD0DA" bold="true" box="[1316,1347,1154,1176]" pageId="50" pageNumber="51">tec</emphasis>
: tract of ectodermal cells. Scale bar 2 mm.
</paragraph>
</caption>
<caption id="DF6A664EFFABFFEDFF6EF94FFD8AD355" httpUri="https://zenodo.org/record/261448/files/figure.png" pageId="50" pageNumber="51" targetBox="[219,1367,1292,1733]" targetPageId="50">
<paragraph id="8BAA36C6FFABFFEDFF6EF94FFD8AD355" blockId="50.[151,1435,1763,1815]" pageId="50" pageNumber="51">
<emphasis id="B961EAD4FFABFFEDFF6EF94FF81ED2BB" bold="true" box="[151,290,1763,1785]" pageId="50" pageNumber="51">FIGURE 60.</emphasis>
<taxonomicName id="4C154D45FFABFFEDFED3F94FFB21D2BB" box="[298,541,1763,1785]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="50" pageNumber="51" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFABFFEDFED3F94FFB21D2BB" box="[298,541,1763,1785]" italics="true" pageId="50" pageNumber="51">Cardianecta parchelion</emphasis>
</taxonomicName>
<emphasis id="B961EAD4FFABFFEDFDDFF94FFBA8D2BB" bold="true" box="[550,660,1763,1785]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="A25257AFFFABFFEDFDDFF94FFB44D2BB" box="[550,632,1763,1785]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
. ostial and
<emphasis id="B961EAD4FFABFFEDFCF6F94FFA1CD2BB" bold="true" box="[783,800,1763,1785]" pageId="50" pageNumber="51">B</emphasis>
. lateral views of larger nectophore;
<emphasis id="B961EAD4FFABFFEDFB65F94FFD92D2BB" bold="true" box="[1180,1198,1763,1785]" pageId="50" pageNumber="51">C</emphasis>
. lateral view of young nectophore, both from holotype specimen.
<emphasis id="B961EAD4FFABFFEDFDB3F8ADFB64D355" bold="true" box="[586,600,1793,1815]" pageId="50" pageNumber="51">n</emphasis>
: notch in lateral wings. Scale bars
<emphasis id="B961EAD4FFABFFEDFC43F8ADFAF0D355" bold="true" box="[954,972,1793,1815]" pageId="50" pageNumber="51">A</emphasis>
and
<emphasis id="B961EAD4FFABFFEDFC07F8ADFD33D355" bold="true" box="[1022,1039,1793,1815]" pageId="50" pageNumber="51">B</emphasis>
2 mm,
<emphasis id="B961EAD4FFABFFEDFBA5F8ADFD52D355" bold="true" box="[1116,1134,1793,1815]" pageId="50" pageNumber="51">C</emphasis>
1 mm.
</paragraph>
</caption>
<paragraph id="8BAA36C6FFABFFECFF3EF893F9DFD527" blockId="50.[151,1436,1855,2024]" lastBlockId="51.[151,1436,151,429]" lastPageId="51" lastPageNumber="52" pageId="50" pageNumber="51">
One feature of the younger nectophores was that there was a distinct notch in each lateral wing (
<figureCitation id="132E2A43FFABFFEDFAF0F893FCB8D31A" box="[1289,1412,1855,1880]" captionStart="FIGURE 60" captionStartId="50.[151,250,1763,1785]" captionTargetBox="[219,1367,1292,1733]" captionTargetId="figure@50.[215,1371,1289,1738]" captionTargetPageId="50" captionText="FIGURE 60. Cardianecta parchelion sp. nov. A. ostial and B. lateral views of larger nectophore; C. lateral view of young nectophore, both from holotype specimen. n: notch in lateral wings. Scale bars A and B 2 mm, C 1 mm." httpUri="https://zenodo.org/record/261448/files/figure.png" pageId="50" pageNumber="51">Figures 60</figureCitation>
C, 61A
<emphasis id="B961EAD4FFABFFEDFF29F8CFF9E3D33E" bold="true" box="[208,223,1891,1916]" pageId="50" pageNumber="51">n</emphasis>
), but the more mature the nectophore the more indistinct this notch became. Another was the presence of two pairs of lateral ridges, although they were often difficult to discern (Figure
<emphasis id="B961EAD4FFABFFEDFBDAF82BFD12D3E2" bold="true" box="[1059,1070,1927,1952]" pageId="50" pageNumber="51">s</emphasis>
5961). The upper pair (
<figureCitation id="132E2A43FFABFFEDFAA8F82BF988D386" captionStart="FIGURE 59" captionStartId="50.[151,250,1092,1114]" captionTargetBox="[190,1393,597,1063]" captionTargetId="figure@50.[184,1402,591,1067]" captionTargetPageId="50" captionText="FIGURE 59. Cardianecta parchelion sp. nov. A. Ostial and B. axial views of larger nectophore from holotype specimen. amc, dmc: ascending and descending mantle canal, respectively; larc, lrc, orc and urc: lateral, lower, ostial ring and upper radial canals, respectively; llr and ulr: lower and upper lateral ridges respectively; ns: nectosac; pedc: pedicular canal; tec: tract of ectodermal cells. Scale bar 2 mm." httpUri="https://zenodo.org/record/261447/files/figure.png" pageId="50" pageNumber="51">Figure 59</figureCitation>
A, 61A
<emphasis id="B961EAD4FFABFFEDFEF5F807F80CD386" bold="true" box="[268,304,1963,1988]" pageId="50" pageNumber="51">ulr</emphasis>
) stretched upwards on either side from the upper side of the squarish ostium, approximately half way between the points of insertion of the upper and lateral radial canals, toward the upper end of the nectophore, but petered out approximately on a level with the apex of the nectosac. They were more indistinct than the lower lateral ridges and, because of damage to the ostial region it was uncertain whether they actually connected with the ostium itself. The pair of lower lateral ridges (
<figureCitation id="132E2A43FFAAFFECFDBFFF73FBFCD4BA" box="[582,704,223,248]" captionStart="FIGURE 59" captionStartId="50.[151,250,1092,1114]" captionTargetBox="[190,1393,597,1063]" captionTargetId="figure@50.[184,1402,591,1067]" captionTargetPageId="50" captionText="FIGURE 59. Cardianecta parchelion sp. nov. A. Ostial and B. axial views of larger nectophore from holotype specimen. amc, dmc: ascending and descending mantle canal, respectively; larc, lrc, orc and urc: lateral, lower, ostial ring and upper radial canals, respectively; llr and ulr: lower and upper lateral ridges respectively; ns: nectosac; pedc: pedicular canal; tec: tract of ectodermal cells. Scale bar 2 mm." httpUri="https://zenodo.org/record/261447/files/figure.png" pageId="51" pageNumber="52">Figures 59</figureCitation>
A, 61A
<emphasis id="B961EAD4FFAAFFECFCE1FF73FA08D4BA" bold="true" box="[792,820,223,248]" pageId="51" pageNumber="52">llr</emphasis>
) was usually more distinct and often was paralleled by a furrow on their upper side, although this might have been a preservation artefact. They arose from the sides of the ostium just below the point of insertion of the lateral radial canals and followed a similar course to the upper lateral ridges.
</paragraph>
<paragraph id="8BAA36C6FFAAFFECFF3EFEC3FAD1D5EF" blockId="51.[151,1436,151,429]" pageId="51" pageNumber="52">
<emphasis id="B961EAD4FFAAFFECFF3EFEC3F87CD5CA" box="[199,320,367,392]" italics="true" pageId="51" pageNumber="52">Siphosome</emphasis>
: The siphosomal stem, if present, was always greatly fragmented such that it was not possible to glean any information regarding the organisation of the individual cormidia.
</paragraph>
<caption id="DF6A664EFFAAFFECFF6EFA74FD47D24F" httpUri="https://zenodo.org/record/261449/files/figure.png" pageId="51" pageNumber="52" targetBox="[361,1225,517,1470]" targetPageId="51">
<paragraph id="8BAA36C6FFAAFFECFF6EFA74FD47D24F" blockId="51.[151,1436,1496,1549]" pageId="51" pageNumber="52">
<emphasis id="B961EAD4FFAAFFECFF6EFA74F81CD1AC" bold="true" box="[151,288,1496,1518]" pageId="51" pageNumber="52">FIGURE 61.</emphasis>
<taxonomicName id="4C154D45FFAAFFECFED1FA74FB26D1AC" box="[296,538,1496,1518]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="51" pageNumber="52" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFAAFFECFED1FA74FB26D1AC" box="[296,538,1496,1518]" italics="true" pageId="51" pageNumber="52">Cardianecta parchelion</emphasis>
</taxonomicName>
<emphasis id="B961EAD4FFAAFFECFDD8FA74FB4FD1AC" bold="true" box="[545,627,1496,1518]" pageId="51" pageNumber="52">
<taxonomicNameLabel id="A25257AFFFAAFFECFDD8FA74FB4FD1AC" box="[545,627,1496,1518]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Lateral (
<emphasis id="B961EAD4FFAAFFECFD2AFA74FBD9D1AC" bold="true" box="[723,741,1496,1518]" pageId="51" pageNumber="52">A</emphasis>
), ostial (
<emphasis id="B961EAD4FFAAFFECFCBBFA74FA6FD1AC" bold="true" box="[834,851,1496,1518]" pageId="51" pageNumber="52">B</emphasis>
), and ostiobasal (
<emphasis id="B961EAD4FFAAFFECFBEEFA74FD15D1AC" bold="true" box="[1047,1065,1496,1518]" pageId="51" pageNumber="52">C</emphasis>
) views of younger nectophore from JSL I Dive 2643 specimen before preservation. See Figures 59 and 60 for legend. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="8BAA36C6FFAAFFEBFF3EF997FDE6D4BA" blockId="51.[151,1437,1593,2014]" lastBlockId="52.[151,1437,151,248]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
<emphasis id="B961EAD4FFAAFFECFF3EF997F839D210" box="[199,261,1595,1618]" italics="true" pageId="51" pageNumber="52">Bract</emphasis>
: The bracts (
<figureCitation id="132E2A43FFAAFFECFE61F995FB01D210" box="[408,573,1593,1618]" captionStart-0="FIGURE 62" captionStart-1="FIGURE 63" captionStartId-0="52.[151,250,991,1013]" captionStartId-1="52.[151,250,1694,1716]" captionTargetBox-0="[359,1232,350,954]" captionTargetBox-1="[340,1246,1157,1670]" captionTargetId-0="figure@52.[355,1239,342,957]" captionTargetId-1="figure@52.[340,1247,1157,1670]" captionTargetPageId-0="52" captionTargetPageId-1="52" captionText-0="FIGURE 62. Upper views of bracts of Cardianecta parchelion sp. nov. A. mature, and B. developing bracts from the holotype (scale bar 1 mm). The grey part of the bracteal canal is within the mesogloea. C. Very young bracts from the JSL I 2633 specimen. Proximal end at bottom. Scale bars 0. 5 mm." captionText-1="FIGURE 63. Cardianecta parchelion sp. nov. Photographs of upper side of bracts from paratype specimen before preservation. Proximal end at bottom. Scale bar 1 mm." httpUri-0="https://zenodo.org/record/261450/files/figure.png" httpUri-1="https://zenodo.org/record/261451/files/figure.png" pageId="51" pageNumber="52">Figures 6263</figureCitation>
) were of a single
<typeStatus id="54AE8864FFAAFFECFCFFF996FA0AD210" box="[774,822,1594,1618]" pageId="51" pageNumber="52">type</typeStatus>
and occurred in enantiomorphic pairs. They measured up to
<quantity id="4CED9B23FFAAFFECFF2CF9F1F82AD237" box="[213,278,1629,1654]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="51" pageNumber="52" unit="mm" value="6.0">6 mm</quantity>
in length and
<quantity id="4CED9B23FFAAFFECFE4AF9F1FB35D237" box="[435,521,1629,1654]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="51" pageNumber="52" unit="mm" value="5.2">5.2 mm</quantity>
in greatest width. They were broadest in their distal half, rapidly tapering down to a small, ventral distal point; with a flat lower side and a domed upper one. In the younger bracts this point was occupied by an accumulation of c. 20 nematocysts, measuring
<date id="FFAB1006FFAAFFECFC73F90AFAD6D2FC" box="[906,1002,1702,1726]" pageId="51" pageNumber="52" value="1932-10-18">32 x 18</date>
µm. However, in older bracts these nematocysts had usually been abraded. In the preserved specimens there was an indistinct ridge that ran from the distal point of the bract to about half its length. However, in the bracts photographed before preservation (
<figureCitation id="132E2A43FFAAFFECFAA8F941F988D368" captionStart="FIGURE 63" captionStartId="52.[151,250,1694,1716]" captionTargetBox="[340,1246,1157,1670]" captionTargetId="figure@52.[340,1247,1157,1670]" captionTargetPageId="52" captionText="FIGURE 63. Cardianecta parchelion sp. nov. Photographs of upper side of bracts from paratype specimen before preservation. Proximal end at bottom. Scale bar 1 mm." httpUri="https://zenodo.org/record/261451/files/figure.png" pageId="51" pageNumber="52">Figure 63</figureCitation>
) this ridge can clearly been seen. In the youngest bracts (
<figureCitation id="132E2A43FFAAFFECFCC1F8BDFA9BD368" box="[824,935,1809,1834]" captionStart="FIGURE 62" captionStartId="52.[151,250,991,1013]" captionTargetBox="[359,1232,350,954]" captionTargetId="figure@52.[355,1239,342,957]" captionTargetPageId="52" captionText="FIGURE 62. Upper views of bracts of Cardianecta parchelion sp. nov. A. mature, and B. developing bracts from the holotype (scale bar 1 mm). The grey part of the bracteal canal is within the mesogloea. C. Very young bracts from the JSL I 2633 specimen. Proximal end at bottom. Scale bars 0. 5 mm." httpUri="https://zenodo.org/record/261450/files/figure.png" pageId="51" pageNumber="52">Figure 62</figureCitation>
) the ridge was quite broad and included two or three rows of distinctive ectodermal cells. The bract tapered down, proximally, more rapidly on the inner side than the outer to form a prominent protuberance at the proximal end. The length of the bracteal canal varied according to the age of the bract, but it was never seen to extend the whole length of the bract. In the youngest ones the canal stretched to about three-quarters the length of the bract, and the part enclosed within the mesogloea was twice as long as the part that ran along the lower surface. The former part was also much broader, and included a broad canal.
<collectingCountry id="F3027656FFADFFEBFED3FF34F875D4F2" box="[298,329,152,176]" name="American Samoa" pageId="52" pageNumber="53">As</collectingCountry>
the bract developed the internal portion of the canal decreased in length, while the proximal part lying against the lower surface increased. Eventually, in the largest bracts the proximal part of the bracteal canal extended to about one-third their length, and the part inflected into the mesogloea was very short.
</paragraph>
<caption id="DF6A664EFFADFFEBFF6EFC73FBFAD070" httpUri="https://zenodo.org/record/261450/files/figure.png" pageId="52" pageNumber="53" targetBox="[359,1232,350,954]" targetPageId="52">
<paragraph id="8BAA36C6FFADFFEBFF6EFC73FBFAD070" blockId="52.[151,1436,991,1074]" pageId="52" pageNumber="53">
<emphasis id="B961EAD4FFADFFEBFF6EFC73F823D7B7" bold="true" box="[151,287,991,1013]" pageId="52" pageNumber="53">FIGURE 62.</emphasis>
Upper views of bracts of
<taxonomicName id="4C154D45FFADFFEBFDDCFC73FA2AD7B7" box="[549,790,991,1013]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="52" pageNumber="53" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFADFFEBFDDCFC73FA2AD7B7" box="[549,790,991,1013]" italics="true" pageId="52" pageNumber="53">Cardianecta parchelion</emphasis>
</taxonomicName>
<emphasis id="B961EAD4FFADFFEBFCE5FC73FAB9D7B7" bold="true" box="[796,901,991,1013]" pageId="52" pageNumber="53">
<taxonomicNameLabel id="A25257AFFFADFFEBFCE5FC73FA51D7B7" box="[796,877,991,1013]" pageId="52" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
. mature, and
<emphasis id="B961EAD4FFADFFEBFBE9FC73FD1DD7B7" bold="true" box="[1040,1057,991,1013]" pageId="52" pageNumber="53">B</emphasis>
. developing bracts from the holotype (scale bar 1 mm). The grey part of the bracteal canal is within the mesogloea.
<emphasis id="B961EAD4FFADFFEBFC23FC52FAD0D056" bold="true" box="[986,1004,1022,1044]" pageId="52" pageNumber="53">C</emphasis>
. Very young bracts from the JSL I 2633 specimen. Proximal end at bottom. Scale bars 0. 5 mm.
</paragraph>
</caption>
<caption id="DF6A664EFFADFFEBFF6EF932FBFDD291" httpUri="https://zenodo.org/record/261451/files/figure.png" pageId="52" pageNumber="53" targetBox="[340,1246,1157,1670]" targetPageId="52">
<paragraph id="8BAA36C6FFADFFEBFF6EF932FBFDD291" blockId="52.[151,1436,1694,1747]" pageId="52" pageNumber="53">
<emphasis id="B961EAD4FFADFFEBFF6EF932F814D2F6" bold="true" box="[151,296,1694,1716]" pageId="52" pageNumber="53">FIGURE 63.</emphasis>
<taxonomicName id="4C154D45FFADFFEBFECFF932FB0CD2F6" box="[310,560,1694,1716]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="52" pageNumber="53" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFADFFEBFECFF932FB0CD2F6" box="[310,560,1694,1716]" italics="true" pageId="52" pageNumber="53">Cardianecta parchelion</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257AFFFADFFEBFDC7F932FBABD2F6" box="[574,663,1694,1716]" pageId="52" pageNumber="53" rank="species">
<emphasis id="B961EAD4FFADFFEBFDC7F932FBABD2F6" bold="true" box="[574,663,1694,1716]" pageId="52" pageNumber="53">sp. nov.</emphasis>
</taxonomicNameLabel>
Photographs of upper side of bracts from paratype specimen before preservation. Proximal end at bottom. Scale bar 1 mm.
</paragraph>
</caption>
<paragraph id="8BAA36C6FFADFFEBFF3EF8ACF83CD38F" blockId="52.[151,1436,1792,1997]" pageId="52" pageNumber="53">
<emphasis id="B961EAD4FFADFFEBFF3EF8ACF8D1D35B" box="[199,493,1792,1817]" italics="true" pageId="52" pageNumber="53">Gastrozooid and tentacle</emphasis>
: The gastrozooids (
<figureCitation id="132E2A43FFADFFEBFD27F8ACFA6FD35B" box="[734,851,1792,1817]" captionStart="FIGURE 64" captionStartId="53.[151,250,912,934]" captionTargetBox="[339,1245,185,887]" captionTargetId="figure@53.[339,1247,185,887]" captionTargetPageId="53" captionText="FIGURE 64. Cardianecta parchelion sp. nov. Gastrozooids from JSL II 998 - D 4. Proximal at bottom, distal at top. Scale bars 1 mm." httpUri="https://zenodo.org/record/261452/files/figure.png" pageId="52" pageNumber="53">Figure 64</figureCitation>
) typically were very variable in size and shape, particularly with regard to the stomach section. The largest measured up to
<quantity id="4CED9B23FFADFFEBFC15F888FD12D37E" box="[1004,1070,1828,1852]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="52" pageNumber="53" unit="mm" value="9.0">9 mm</quantity>
in length and
<quantity id="4CED9B23FFADFFEBFB28F888FC1CD37E" box="[1233,1312,1828,1852]" metricMagnitude="-2" metricUnit="m" metricValue="2.3" pageId="52" pageNumber="53" unit="mm" value="23.0">23 mm</quantity>
in greatest width. There was a prominent, cylindrical basigaster that measured about
<quantity id="4CED9B23FFADFFEBFC09F8E4FD75D322" box="[1008,1097,1864,1888]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="52" pageNumber="53" unit="mm" value="2.3">2.3 mm</quantity>
in length and slightly less in width, which was separated from the main stomach region by a distinct constriction. There was no obvious distal proboscis region, but the mouth and much of the stomach could open widely, revealing the hepatic stripes (
<figureCitation id="132E2A43FFADFFEBFAA8F83CF98FD38F" captionStart="FIGURE 64" captionStartId="53.[151,250,912,934]" captionTargetBox="[339,1245,185,887]" captionTargetId="figure@53.[339,1247,185,887]" captionTargetPageId="53" captionText="FIGURE 64. Cardianecta parchelion sp. nov. Gastrozooids from JSL II 998 - D 4. Proximal at bottom, distal at top. Scale bars 1 mm." httpUri="https://zenodo.org/record/261452/files/figure.png" pageId="52" pageNumber="53">Figure 64</figureCitation>
right).
</paragraph>
<caption id="DF6A664EFFACFFEAFF6EFC3CF9FAD786" httpUri="https://zenodo.org/record/261452/files/figure.png" pageId="53" pageNumber="54" targetBox="[339,1245,185,887]" targetPageId="53">
<paragraph id="8BAA36C6FFACFFEAFF6EFC3CF9FAD786" blockId="53.[151,1435,912,964]" pageId="53" pageNumber="54">
<emphasis id="B961EAD4FFACFFEAFF6EFC3CF823D7E4" bold="true" box="[151,287,912,934]" pageId="53" pageNumber="54">FIGURE 64.</emphasis>
<taxonomicName id="4C154D45FFACFFEAFEDCFC3CFB29D7E4" box="[293,533,912,934]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="53" pageNumber="54" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFACFFEAFEDCFC3CFB29D7E4" box="[293,533,912,934]" italics="true" pageId="53" pageNumber="54">Cardianecta parchelion</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257AFFFACFFEAFDE2FC3CFB57D7E4" box="[539,619,912,934]" pageId="53" pageNumber="54" rank="species">
<emphasis id="B961EAD4FFACFFEAFDE2FC3CFB57D7E4" bold="true" box="[539,619,912,934]" pageId="53" pageNumber="54">sp. nov.</emphasis>
</taxonomicNameLabel>
Gastrozooids from JSL II 998-D4. Proximal at bottom, distal at top. Scale bars 1 mm.
</paragraph>
</caption>
<paragraph id="8BAA36C6FFACFFEAFF3EFC5DFB6AD130" blockId="53.[151,1437,1009,2006]" pageId="53" pageNumber="54">
<emphasis id="B961EAD4FFACFFEAFF3EFC5DF827D048" box="[199,283,1009,1034]" italics="true" pageId="53" pageNumber="54">Tentilla</emphasis>
: The tentilla first appeared as short, thickened, undifferentiated tubes at the proximal end of the tentacle (
<figureCitation id="132E2A43FFACFFEAFF04FBB9F852D06C" box="[253,366,1045,1070]" captionStart="FIGURE 64" captionStartId="53.[151,250,912,934]" captionTargetBox="[339,1245,185,887]" captionTargetId="figure@53.[339,1247,185,887]" captionTargetPageId="53" captionText="FIGURE 64. Cardianecta parchelion sp. nov. Gastrozooids from JSL II 998 - D 4. Proximal at bottom, distal at top. Scale bars 1 mm." httpUri="https://zenodo.org/record/261452/files/figure.png" pageId="53" pageNumber="54">Figure 64</figureCitation>
). In the preserved specimens the distal end of this tube was often curved to one side, and it was part that next began to elongate and to twist itself into 23 spirals to form the rudimentary cnidoband, which at this stage did not appear to be covered by a membrane. At the same time the involucrum, between the pedicle and the cnidoband began to differentiate. It formed a horseshoe shaped swelling with the two loose ends closely apposed. Simultaneously the terminal process began to form as a flattened irregular swelling at the distal end of the cnidoband, with two short, broad distal protuberances (
<figureCitation id="132E2A43FFACFFEAFD04FB65FA50D0A0" box="[765,876,1225,1250]" captionStart="FIGURE 65" captionStartId="54.[151,250,1652,1674]" captionTargetBox="[371,1217,236,1622]" captionTargetId="figure@54.[364,1224,231,1628]" captionTargetPageId="54" captionText="FIGURE 65. Cardianecta parchelion sp. nov. Stages in development of the tentilla from the type specimen. cn. cnidoband; h. horns; i. involucrum; p. pedicle; tp. terminal process. Scale bars: A B 0.25 mm; C E 0.5 mm; F H 1.0 mm." httpUri="https://zenodo.org/record/261453/files/figure.png" pageId="53" pageNumber="54">Figure 65</figureCitation>
A). The terminal process continued to enlarge into an almost spherical, relatively thick-walled swelling, bearing two, asymmetrically placed &quot;horns&quot;; one terminal and the other sub-terminal (
<figureCitation id="132E2A43FFACFFEAFE5DFABDFB2BD168" box="[420,535,1297,1322]" captionStart="FIGURE 65" captionStartId="54.[151,250,1652,1674]" captionTargetBox="[371,1217,236,1622]" captionTargetId="figure@54.[364,1224,231,1628]" captionTargetPageId="54" captionText="FIGURE 65. Cardianecta parchelion sp. nov. Stages in development of the tentilla from the type specimen. cn. cnidoband; h. horns; i. involucrum; p. pedicle; tp. terminal process. Scale bars: A B 0.25 mm; C E 0.5 mm; F H 1.0 mm." httpUri="https://zenodo.org/record/261453/files/figure.png" pageId="53" pageNumber="54">Figure 65</figureCitation>
B, C). Up to this stage the pedicle remained short and broad, with a relatively wide central canal that at first was straight, but later began to twist into coils (
<figureCitation id="132E2A43FFACFFEAFBF6FA99FDBDD10C" box="[1039,1153,1333,1358]" captionStart="FIGURE 65" captionStartId="54.[151,250,1652,1674]" captionTargetBox="[371,1217,236,1622]" captionTargetId="figure@54.[364,1224,231,1628]" captionTargetPageId="54" captionText="FIGURE 65. Cardianecta parchelion sp. nov. Stages in development of the tentilla from the type specimen. cn. cnidoband; h. horns; i. involucrum; p. pedicle; tp. terminal process. Scale bars: A B 0.25 mm; C E 0.5 mm; F H 1.0 mm." httpUri="https://zenodo.org/record/261453/files/figure.png" pageId="53" pageNumber="54">Figure 65</figureCitation>
C), while the involucrum showed no further sign of development.
</paragraph>
<paragraph id="8BAA36C6FFACFFEAFF3EFAD2FD23D22C" blockId="53.[151,1437,1009,2006]" pageId="53" pageNumber="54">
At the next stage of development the pedicle began to elongate considerably, becoming thinner walled, and with its central canal straightening out again, except at its distal end where, in the preserved state at least, it remained coiled. At the same time the cnidoband began to increase in length by the addition of further regular coils (
<figureCitation id="132E2A43FFACFFEAFF67FA45F826D240" box="[158,282,1513,1538]" captionStart="FIGURE 65" captionStartId="54.[151,250,1652,1674]" captionTargetBox="[371,1217,236,1622]" captionTargetId="figure@54.[364,1224,231,1628]" captionTargetPageId="54" captionText="FIGURE 65. Cardianecta parchelion sp. nov. Stages in development of the tentilla from the type specimen. cn. cnidoband; h. horns; i. involucrum; p. pedicle; tp. terminal process. Scale bars: A B 0.25 mm; C E 0.5 mm; F H 1.0 mm." httpUri="https://zenodo.org/record/261453/files/figure.png" pageId="53" pageNumber="54">Figures 65</figureCitation>
D; 66A, B, C) up to a maximum of six to seven.
<collectingCountry id="F3027656FFACFFEAFCBAFA45FA5ED243" box="[835,866,1513,1537]" name="American Samoa" pageId="53" pageNumber="54">As</collectingCountry>
the cnidoband reached its maximum development the involucrum began to develop further, by the lengthening of the free ends of the horseshoe and their gradual spreading apart (
<figureCitation id="132E2A43FFACFFEAFEAFF99DF8EED208" box="[342,466,1585,1610]" captionStart="FIGURE 65" captionStartId="54.[151,250,1652,1674]" captionTargetBox="[371,1217,236,1622]" captionTargetId="figure@54.[364,1224,231,1628]" captionTargetPageId="54" captionText="FIGURE 65. Cardianecta parchelion sp. nov. Stages in development of the tentilla from the type specimen. cn. cnidoband; h. horns; i. involucrum; p. pedicle; tp. terminal process. Scale bars: A B 0.25 mm; C E 0.5 mm; F H 1.0 mm." httpUri="https://zenodo.org/record/261453/files/figure.png" pageId="53" pageNumber="54">Figures 65</figureCitation>
E, F; 66D). Eventually these formed long lateral flaps that extended out sideways or curved distally to cover the cnidoband on two sides (
<figureCitation id="132E2A43FFACFFEAFD1CF9F9FA62D22C" box="[741,862,1621,1646]" captionStart="FIGURE 65" captionStartId="54.[151,250,1652,1674]" captionTargetBox="[371,1217,236,1622]" captionTargetId="figure@54.[364,1224,231,1628]" captionTargetPageId="54" captionText="FIGURE 65. Cardianecta parchelion sp. nov. Stages in development of the tentilla from the type specimen. cn. cnidoband; h. horns; i. involucrum; p. pedicle; tp. terminal process. Scale bars: A B 0.25 mm; C E 0.5 mm; F H 1.0 mm." httpUri="https://zenodo.org/record/261453/files/figure.png" pageId="53" pageNumber="54">Figures 65</figureCitation>
EH; 66E,F; 67).
</paragraph>
<paragraph id="8BAA36C6FFACFFEAFF3EF9D5F8A3D394" blockId="53.[151,1437,1009,2006]" pageId="53" pageNumber="54">
During this stage an external membrane began to cover the cnidoband. At first it was tight against the cnidoband, but later began to expand so as to leave a gap between. This was the ultimate stage found in most of the specimens, but in the
<typeStatus id="54AE8864FFACFFEAFE70F96DF8D7D298" box="[393,491,1729,1754]" pageId="53" pageNumber="54" type="holotype">holotype</typeStatus>
three tentilla were found that showed further development (
<figureCitation id="132E2A43FFACFFEAFB70F96DFDCBD298" box="[1161,1271,1729,1754]" captionStart="FIGURE 65" captionStartId="54.[151,250,1652,1674]" captionTargetBox="[371,1217,236,1622]" captionTargetId="figure@54.[364,1224,231,1628]" captionTargetPageId="54" captionText="FIGURE 65. Cardianecta parchelion sp. nov. Stages in development of the tentilla from the type specimen. cn. cnidoband; h. horns; i. involucrum; p. pedicle; tp. terminal process. Scale bars: A B 0.25 mm; C E 0.5 mm; F H 1.0 mm." httpUri="https://zenodo.org/record/261453/files/figure.png" pageId="53" pageNumber="54">Figure 65</figureCitation>
G,H). On these the terminal process expanded in size and its walls became thinner and indistinct from the membrane covering the cnidoband. Thus a thin-walled spherical cavity was formed that enclosed, usually in its proximal half, the cnidoband, and still retained the terminal and sub-terminal &quot;horns&quot;. On one tentillum (
<figureCitation id="132E2A43FFACFFEAFB90F881FDE4D304" box="[1129,1240,1837,1862]" captionStart="FIGURE 65" captionStartId="54.[151,250,1652,1674]" captionTargetBox="[371,1217,236,1622]" captionTargetId="figure@54.[364,1224,231,1628]" captionTargetPageId="54" captionText="FIGURE 65. Cardianecta parchelion sp. nov. Stages in development of the tentilla from the type specimen. cn. cnidoband; h. horns; i. involucrum; p. pedicle; tp. terminal process. Scale bars: A B 0.25 mm; C E 0.5 mm; F H 1.0 mm." httpUri="https://zenodo.org/record/261453/files/figure.png" pageId="53" pageNumber="54">Figure 65</figureCitation>
H) the cnidoband lost its regular coiling and formed a series of irregular loops. Whether this was an artefact caused by the breaking away of the proximal part of the cnidoband from the pedicle, is not clear, although other, less developed tentilla (e.g.
<figureCitation id="132E2A43FFACFFEAFF28F835F87CD3F0" box="[209,320,1945,1970]" captionStart="FIGURE 66" captionStartId="55.[151,250,1413,1435]" captionTargetBox="[322,1266,211,1376]" captionTargetId="figure@55.[302,1285,195,1389]" captionTargetPageId="55" captionText="FIGURE 66. Cardianecta parchelion sp. nov. Stages in development of tentillum. JSLII 998 - D 4 specimen. Scale bars: A C 0.25 mm; D F 0.5 mm." httpUri="https://zenodo.org/record/261454/files/figure.png" pageId="53" pageNumber="54">Figure 66</figureCitation>
E) also showed a less regularly coiled cnidoband. No clear evidence for the presence of elastic or angle bands was found.
</paragraph>
<caption id="DF6A664EFFAFFFE9FF6EF9D8FDDDD2EB" httpUri="https://zenodo.org/record/261453/files/figure.png" pageId="54" pageNumber="55" targetBox="[371,1217,236,1622]" targetPageId="54">
<paragraph id="8BAA36C6FFAFFFE9FF6EF9D8FDDDD2EB" blockId="54.[151,1435,1652,1705]" pageId="54" pageNumber="55">
<emphasis id="B961EAD4FFAFFFE9FF6EF9D8F81CD2C8" bold="true" box="[151,288,1652,1674]" pageId="54" pageNumber="55">FIGURE 65.</emphasis>
<taxonomicName id="4C154D45FFAFFFE9FEDFF9D8FB2BD2C8" box="[294,535,1652,1674]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="54" pageNumber="55" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFAFFFE9FEDFF9D8FB2BD2C8" box="[294,535,1652,1674]" italics="true" pageId="54" pageNumber="55">Cardianecta parchelion</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257AFFFAFFFE9FDE4F9D8FB51D2C8" box="[541,621,1652,1674]" pageId="54" pageNumber="55" rank="species">
<emphasis id="B961EAD4FFAFFFE9FDE4F9D8FB51D2C8" bold="true" box="[541,621,1652,1674]" pageId="54" pageNumber="55">sp. nov.</emphasis>
</taxonomicNameLabel>
Stages in development of the tentilla from the type specimen.
<emphasis id="B961EAD4FFAFFFE9FB13F9D8FC3FD2C8" bold="true" box="[1258,1283,1652,1674]" pageId="54" pageNumber="55">cn</emphasis>
. cnidoband;
<emphasis id="B961EAD4FFAFFFE9FA7EF9D8FCA9D2C8" bold="true" box="[1415,1429,1652,1674]" pageId="54" pageNumber="55">h</emphasis>
. horns;
<emphasis id="B961EAD4FFAFFFE9FF25F93FF9DFD2EB" bold="true" box="[220,227,1683,1705]" pageId="54" pageNumber="55">i</emphasis>
. involucrum;
<emphasis id="B961EAD4FFAFFFE9FE8BF93FF8BCD2EB" bold="true" box="[370,384,1683,1705]" pageId="54" pageNumber="55">p</emphasis>
. pedicle;
<emphasis id="B961EAD4FFAFFFE9FE1AF93FFB3DD2EB" bold="true" box="[483,513,1683,1705]" pageId="54" pageNumber="55">tp.</emphasis>
terminal process. Scale bars: AB 0.25 mm; CE 0.5 mm; FH 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BAA36C6FFAFFFE9FF3EF956F8D8D33D" blockId="54.[151,1437,1786,1919]" pageId="54" pageNumber="55">
Only two
<typeStatus id="54AE8864FFAFFFE9FEC3F957F849D351" box="[314,373,1787,1811]" pageId="54" pageNumber="55">types</typeStatus>
of nematocysts were found on the tentilla. Small rounded nematocysts, c. 14 µm in diameter where found on the terminal process and were mainly concentrated in the &quot;horns&quot;. These appeared to be desmonemes. The cnidoband bore only elongate nematocysts, c. 42 µm long and 15µm at their widest, which possibly were mastigophores.
</paragraph>
<caption id="DF6A664EFFAEFFE8FF6EFA29F8BAD1F8" httpUri="https://zenodo.org/record/261454/files/figure.png" pageId="55" pageNumber="56" targetBox="[322,1266,211,1376]" targetPageId="55">
<paragraph id="8BAA36C6FFAEFFE8FF6EFA29F8BAD1F8" blockId="55.[151,1436,1413,1466]" pageId="55" pageNumber="56">
<emphasis id="B961EAD4FFAEFFE8FF6EFA29F81DD1D9" bold="true" box="[151,289,1413,1435]" pageId="55" pageNumber="56">FIGURE 66.</emphasis>
<taxonomicName id="4C154D45FFAEFFE8FED1FA29FB26D1D9" box="[296,538,1413,1435]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="55" pageNumber="56" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFAEFFE8FED1FA29FB26D1D9" box="[296,538,1413,1435]" italics="true" pageId="55" pageNumber="56">Cardianecta parchelion</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257AFFFAEFFE8FDDBFA29FB48D1D9" box="[546,628,1413,1435]" pageId="55" pageNumber="56" rank="species">
<emphasis id="B961EAD4FFAEFFE8FDDBFA29FB48D1D9" bold="true" box="[546,628,1413,1435]" pageId="55" pageNumber="56">sp. nov.</emphasis>
</taxonomicNameLabel>
Stages in development of tentillum. JSLII 998-D4 specimen. Scale bars: AC 0.25 mm; DF 0.5 mm.
</paragraph>
</caption>
<paragraph id="8BAA36C6FFAEFFE8FF3EFA48F8F6D297" blockId="55.[151,1437,1508,2037]" pageId="55" pageNumber="56">
<emphasis id="B961EAD4FFAEFFE8FF3EFA48F824D1BF" box="[199,280,1508,1533]" italics="true" pageId="55" pageNumber="56">Palpon</emphasis>
: The palpons (
<figureCitation id="132E2A43FFAEFFE8FE3CFA48FB09D1BF" box="[453,565,1508,1533]" captionStart="FIGURE 68" captionStartId="56.[151,250,1723,1745]" captionTargetBox="[307,1284,1221,1699]" captionTargetId="figure@56.[302,1284,1221,1699]" captionTargetPageId="56" captionText="FIGURE 68. Cardianecta parchelion sp. nov. A. Palpons from JSL II 998 Specimen. Scale bar 1 mm. B. female (left) and male (right) gonophores from the paratype specimen. Scale bar 0.5 mm. Proximal top—distal bottom." httpUri="https://zenodo.org/record/261456/files/figure.png" pageId="55" pageNumber="56">Figure 68</figureCitation>
A) measured up to c.
<quantity id="4CED9B23FFAEFFE8FCC8FA48FAB5D1BE" box="[817,905,1508,1532]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="55" pageNumber="56" unit="mm" value="3.3">3.3 mm</quantity>
in length; the larger ones tending to be ovate, while the smaller ones were cylindrical or club-shaped. All possessed a small rounded proboscis region at their distal ends. Almost all of the largest palpons did not possess a palpacle, but it is clear that the latter must have been detached as such a structure was still attached to many of the younger ones (
<figureCitation id="132E2A43FFAEFFE8FBD7F9E3FD9ED22A" box="[1070,1186,1615,1640]" captionStart="FIGURE 68" captionStartId="56.[151,250,1723,1745]" captionTargetBox="[307,1284,1221,1699]" captionTargetId="figure@56.[302,1284,1221,1699]" captionTargetPageId="56" captionText="FIGURE 68. Cardianecta parchelion sp. nov. A. Palpons from JSL II 998 Specimen. Scale bar 1 mm. B. female (left) and male (right) gonophores from the paratype specimen. Scale bar 0.5 mm. Proximal top—distal bottom." httpUri="https://zenodo.org/record/261456/files/figure.png" pageId="55" pageNumber="56">Figure 68</figureCitation>
A). The palpacle was annulated, with quite long segments between each annulation and appeared to bear a few, very small, ovate nematocysts, but these were not examined in detail. The palpon itself was thin-walled and featureless, and was devoid of any nematocysts.
</paragraph>
<paragraph id="8BAA36C6FFAEFFE8FF3EF973FBBDD3B7" blockId="55.[151,1437,1508,2037]" pageId="55" pageNumber="56">
<emphasis id="B961EAD4FFAEFFE8FF3EF973F86CD2BA" box="[199,336,1759,1784]" italics="true" pageId="55" pageNumber="56">Gonophores</emphasis>
: Gonophores of both sexes were present (
<figureCitation id="132E2A43FFAEFFE8FCC3F973FA84D2BA" box="[826,952,1759,1784]" captionStart="FIGURE 68" captionStartId="56.[151,250,1723,1745]" captionTargetBox="[307,1284,1221,1699]" captionTargetId="figure@56.[302,1284,1221,1699]" captionTargetPageId="56" captionText="FIGURE 68. Cardianecta parchelion sp. nov. A. Palpons from JSL II 998 Specimen. Scale bar 1 mm. B. female (left) and male (right) gonophores from the paratype specimen. Scale bar 0.5 mm. Proximal top—distal bottom." httpUri="https://zenodo.org/record/261456/files/figure.png" pageId="55" pageNumber="56">Figures 68</figureCitation>
B). The youngest gonophores consisted of spherical processes borne on short narrow pedicles, with the sexual products occupying the whole of the subumbrella cavity, and only a narrow distal opening.
<collectingCountry id="F3027656FFAEFFE8FD04F884FA20D302" box="[765,796,1832,1856]" name="American Samoa" pageId="55" pageNumber="56">As</collectingCountry>
they grew the gonophores became more elongate with a wide pedicle, through which the gastrovascular cavity passed, occupying approximately a third of its length, and the subumbrella cavity extending distal to the sexual products, and was widely open. In its lining were four straight lateral canals. Further development saw a re-shortening of the pedicle, which still remained wide, and an enlargement of the subumbrella cavity; although the overall length of the gonophore only increased slightly. No nematocysts were noted on the gonophores.
</paragraph>
</subSubSection>
<caption id="DF6A664EFFA1FFE7FF6EFBF0F873D0D3" httpUri="https://zenodo.org/record/261455/files/figure.png" pageId="56" pageNumber="57" targetBox="[348,1234,195,1091]" targetPageId="56">
<paragraph id="8BAA36C6FFA1FFE7FF6EFBF0F873D0D3" blockId="56.[151,1437,1116,1169]" pageId="56" pageNumber="57">
<emphasis id="B961EAD4FFA1FFE7FF6EFBF0F81ED030" bold="true" box="[151,290,1116,1138]" pageId="56" pageNumber="57">FIGURE 67.</emphasis>
<taxonomicName id="4C154D45FFA1FFE7FED5FBF0FB23D030" box="[300,543,1116,1138]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="56" pageNumber="57" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFA1FFE7FED5FBF0FB23D030" box="[300,543,1116,1138]" italics="true" pageId="56" pageNumber="57">Cardianecta parchelion</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257AFFFA1FFE7FDD0FBF0FB40D030" box="[553,636,1116,1138]" pageId="56" pageNumber="57" rank="species">
<emphasis id="B961EAD4FFA1FFE7FDD0FBF0FB40D030" bold="true" box="[553,636,1116,1138]" pageId="56" pageNumber="57">sp. nov.</emphasis>
</taxonomicNameLabel>
Tentillum from paratype specimen before preservation. Proximal at bottom. Scale bar 0.5 mm.
</paragraph>
</caption>
<caption id="DF6A664EFFA1FFE7FF6EF917FD99D2B2" httpUri="https://zenodo.org/record/261456/files/figure.png" pageId="56" pageNumber="57" targetBox="[307,1284,1221,1699]" targetPageId="56">
<paragraph id="8BAA36C6FFA1FFE7FF6EF917FD99D2B2" blockId="56.[151,1435,1723,1776]" pageId="56" pageNumber="57">
<emphasis id="B961EAD4FFA1FFE7FF6EF917F81DD293" bold="true" box="[151,289,1723,1745]" pageId="56" pageNumber="57">FIGURE 68.</emphasis>
<taxonomicName id="4C154D45FFA1FFE7FED3F917FB21D293" box="[298,541,1723,1745]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="56" pageNumber="57" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFA1FFE7FED3F917FB21D293" box="[298,541,1723,1745]" italics="true" pageId="56" pageNumber="57">Cardianecta parchelion</emphasis>
</taxonomicName>
<emphasis id="B961EAD4FFA1FFE7FDDDF917FBAED293" bold="true" box="[548,658,1723,1745]" pageId="56" pageNumber="57">
<taxonomicNameLabel id="A25257AFFFA1FFE7FDDDF917FB4BD293" box="[548,631,1723,1745]" pageId="56" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
A
</emphasis>
. Palpons from JSL II 998 Specimen. Scale bar 1 mm.
<emphasis id="B961EAD4FFA1FFE7FB29F917FDDDD293" bold="true" box="[1232,1249,1723,1745]" pageId="56" pageNumber="57">B</emphasis>
. female (left) and male (right) gonophores from the paratype specimen. Scale bar 0.5 mm. Proximal top—distal bottom.
</paragraph>
</caption>
<subSubSection id="C30F654DFFA1FFE7FF3EF8B0F8F1D31B" pageId="56" pageNumber="57" type="distribution">
<paragraph id="8BAA36C6FFA1FFE7FF3EF8B0F8F1D31B" blockId="56.[151,1436,1820,2026]" pageId="56" pageNumber="57">
<emphasis id="B961EAD4FFA1FFE7FF3EF8B0F85CD377" bold="true" box="[199,352,1820,1845]" pageId="56" pageNumber="57">Distribution.</emphasis>
All five specimens were collected in the &quot;Tongue of the Ocean&quot; in
<collectingCountry id="F3027656FFA1FFE7FB72F8B0FC14D377" box="[1163,1320,1820,1845]" name="Bahamas" pageId="56" pageNumber="57">The Bahamas</collectingCountry>
at depths ranging from
<quantity id="4CED9B23FFA1FFE7FEC8F8EDF8F4D31B" box="[305,456,1857,1882]" metricMagnitude="2" metricUnit="m" metricValue="7.62" metricValueMax="8.84" metricValueMin="6.4" pageId="56" pageNumber="57" unit="m" value="762.0" valueMax="884.0" valueMin="640.0">
640 to
<specimenCount id="9D13FD4FFFA1FFE7FE87F8EDF8F4D31B" box="[382,456,1857,1881]" pageId="56" pageNumber="57" type="male">884 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C30F654DFFA1FFE6FF3EF8C8F897D6C7" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57" type="discussion">
<paragraph id="8BAA36C6FFA1FFE6FF3EF8C8FDABD5EF" blockId="56.[151,1436,1820,2026]" lastBlockId="57.[151,1437,151,752]" lastPageId="57" lastPageNumber="58" pageId="56" pageNumber="57">
<emphasis id="B961EAD4FFA1FFE7FF3EF8C8F807D33F" bold="true" box="[199,315,1892,1917]" pageId="56" pageNumber="57">Remarks.</emphasis>
<collectingCountry id="F3027656FFA1FFE7FEBBF8C9F85DD33F" box="[322,353,1893,1917]" name="American Samoa" pageId="56" pageNumber="57">As</collectingCountry>
noted above, it is far from certain that the genus
<taxonomicName id="4C154D45FFA1FFE7FC74F8C8FD24D33F" box="[909,1048,1892,1917]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="56" pageNumber="59" phylum="Cnidaria" rank="genus" status="gen. nov.">
<emphasis id="B961EAD4FFA1FFE7FC74F8C8FD24D33F" box="[909,1048,1892,1917]" italics="true" pageId="56" pageNumber="57">Cardianecta</emphasis>
</taxonomicName>
actually belongs within the newly established family Cordagalmatidae fam. nov. The only character that
<taxonomicName id="4C154D45FFA1FFE7FC35F826FD55D3E0" box="[972,1129,1929,1954]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="56" pageNumber="57" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFA1FFE7FC35F826FD55D3E0" box="[972,1129,1929,1954]" italics="true" pageId="56" pageNumber="57">C. parchelion</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257AFFFA1FFE7FB8DF825FDEDD3E0" box="[1140,1233,1929,1954]" pageId="56" pageNumber="57" rank="species">
<emphasis id="B961EAD4FFA1FFE7FB8DF825FDEDD3E0" bold="true" box="[1140,1233,1929,1954]" pageId="56" pageNumber="57">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C154D45FFA1FFE7FAF6F825FCA0D3E0" box="[1295,1436,1929,1954]" class="Hydrozoa" family="Agalmatidae" genus="Cordagalma" kingdom="Animalia" order="Siphonophorae" pageId="56" pageNumber="57" phylum="Cnidaria" rank="genus">
<emphasis id="B961EAD4FFA1FFE7FAF6F825FCA0D3E0" box="[1295,1436,1929,1954]" italics="true" pageId="56" pageNumber="57">Cordagalma</emphasis>
</taxonomicName>
species have in common is that the nectophores are heart-shaped. Although the bracts of the former are more foliaceous and do not possess a distal facet, there are fundamental differences between the other two main characters, the palpons and in particular the tentilla and their cnidome, that suggest that the genera may not be that closely related. All
<taxonomicName id="4C154D45FFA0FFE6FE8AFF10FB3CD497" box="[371,512,188,213]" class="Hydrozoa" family="Agalmatidae" genus="Cordagalma" kingdom="Animalia" order="Siphonophorae" pageId="57" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="B961EAD4FFA0FFE6FE8AFF10FB3CD497" box="[371,512,188,213]" italics="true" pageId="57" pageNumber="58">Cordagalma</emphasis>
</taxonomicName>
species, as far as we know, have palpons that do not develop a palpon, while such is clearly present with, a least, the younger palpons of
<taxonomicName id="4C154D45FFA0FFE6FCD6FF4CFAF1D4BA" box="[815,973,223,248]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="57" pageNumber="58" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFA0FFE6FCD6FF4CFAF1D4BA" box="[815,973,223,248]" italics="true" pageId="57" pageNumber="58">C. parchelion</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257AFFFA0FFE6FC23FF73FD05D4BA" box="[986,1081,223,248]" pageId="57" pageNumber="58" rank="species">
<emphasis id="B961EAD4FFA0FFE6FC23FF73FD05D4BA" bold="true" box="[986,1081,223,248]" pageId="57" pageNumber="58">sp. nov.</emphasis>
</taxonomicNameLabel>
Nonetheless, the fundamental differences between the simple, larval-type tentilla of
<taxonomicName id="4C154D45FFA0FFE6FCF8FEA8FAB2D55F" box="[769,910,260,285]" class="Hydrozoa" family="Agalmatidae" genus="Cordagalma" kingdom="Animalia" order="Siphonophorae" pageId="57" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="B961EAD4FFA0FFE6FCF8FEA8FAB2D55F" box="[769,910,260,285]" italics="true" pageId="57" pageNumber="58">Cordagalma</emphasis>
</taxonomicName>
species, and the complex involucrate form of those of
<taxonomicName id="4C154D45FFA0FFE6FF05FE84F8A4D502" box="[252,408,295,320]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="57" pageNumber="58" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFA0FFE6FF05FE84F8A4D502" box="[252,408,295,320]" italics="true" pageId="57" pageNumber="58">C. parchelion</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257AFFFA0FFE6FE5BFE8BF8C2D502" box="[418,510,295,320]" pageId="57" pageNumber="58" rank="species">
<emphasis id="B961EAD4FFA0FFE6FE5BFE8BF8C2D502" bold="true" box="[418,510,295,320]" pageId="57" pageNumber="58">sp. nov.</emphasis>
</taxonomicNameLabel>
is striking. This also applies to their cnidome, with
<taxonomicName id="4C154D45FFA0FFE6FBA6FE8BFDD0D502" box="[1119,1260,295,320]" class="Hydrozoa" family="Agalmatidae" genus="Cordagalma" kingdom="Animalia" order="Siphonophorae" pageId="57" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="B961EAD4FFA0FFE6FBA6FE8BFDD0D502" box="[1119,1260,295,320]" italics="true" pageId="57" pageNumber="58">Cordagalma</emphasis>
</taxonomicName>
species having three
<typeStatus id="54AE8864FFA0FFE6FF21FEE1F82FD527" box="[216,275,333,357]" pageId="57" pageNumber="58">types</typeStatus>
of nematocysts, including an incomplete circle of usually five stenoteles, in the cnidoband, while
<taxonomicName id="4C154D45FFA0FFE6FA7BFEE1F833D5CA" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="57" pageNumber="58" phylum="Cnidaria" rank="species" species="parchelion">
<emphasis id="B961EAD4FFA0FFE6FA7BFEE1F833D5CA" italics="true" pageId="57" pageNumber="58">C. parchelion</emphasis>
</taxonomicName>
sp nov. has just one. Meanwhile, that species possesses what appear to be desmonemes in its terminal process, with two asymmetric horns, while such a process is absent in
<taxonomicName id="4C154D45FFA0FFE6FC52FE38FD04D5EF" box="[939,1080,404,429]" class="Hydrozoa" family="Agalmatidae" genus="Cordagalma" kingdom="Animalia" order="Siphonophorae" pageId="57" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="B961EAD4FFA0FFE6FC52FE38FD04D5EF" box="[939,1080,404,429]" italics="true" pageId="57" pageNumber="58">Cordagalma</emphasis>
</taxonomicName>
species.
</paragraph>
<paragraph id="8BAA36C6FFA0FFE6FF3EFE1BF897D6C7" blockId="57.[151,1437,151,752]" pageId="57" pageNumber="58">
Thus it is clear that the species
<taxonomicName id="4C154D45FFA0FFE6FDDEFE1BFBA3D592" box="[551,671,439,464]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="57" pageNumber="58" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFA0FFE6FDDEFE1BFBA3D592" box="[551,671,439,464]" italics="true" pageId="57" pageNumber="58">parchelion</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257AFFFA0FFE6FD5EFE1BFA3CD592" box="[679,768,439,464]" pageId="57" pageNumber="58" rank="species">
<emphasis id="B961EAD4FFA0FFE6FD5EFE1BFA3CD592" bold="true" box="[679,768,439,464]" pageId="57" pageNumber="58">sp. nov.</emphasis>
</taxonomicNameLabel>
should be placed in a separate genus from the
<taxonomicName id="4C154D45FFA0FFE6FAF6FE1BFCA0D592" box="[1295,1436,439,464]" class="Hydrozoa" family="Agalmatidae" genus="Cordagalma" kingdom="Animalia" order="Siphonophorae" pageId="57" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="B961EAD4FFA0FFE6FAF6FE1BFCA0D592" box="[1295,1436,439,464]" italics="true" pageId="57" pageNumber="58">Cordagalma</emphasis>
</taxonomicName>
species but, at present, we cannot be certain whether it actually belongs within the same family. However, the two genera are both monoecious and the nectophores are developed on the ventral side of the nectosome. Molecular genetic data are now available for some
<taxonomicName id="4C154D45FFA0FFE6FD91FD88FBC9D67F" box="[616,757,548,573]" class="Hydrozoa" family="Agalmatidae" genus="Cordagalma" kingdom="Animalia" order="Siphonophorae" pageId="57" pageNumber="58" phylum="Cnidaria" rank="genus">
<emphasis id="B961EAD4FFA0FFE6FD91FD88FBC9D67F" box="[616,757,548,573]" italics="true" pageId="57" pageNumber="58">Cordagalma</emphasis>
</taxonomicName>
species, other than
<taxonomicName id="4C154D45FFA0FFE6FC20FD89FD4CD67F" box="[985,1136,548,573]" class="Hydrozoa" family="Agalmatidae" genus="Cordagalma" kingdom="Animalia" order="Siphonophorae" pageId="57" pageNumber="58" phylum="Cnidaria" rank="species" species="ordinatum">
<emphasis id="B961EAD4FFA0FFE6FC20FD89FD4CD67F" box="[985,1136,548,573]" italics="true" pageId="57" pageNumber="58">C. ordinatum</emphasis>
</taxonomicName>
(Steve Haddock, personal communication), but until such can be gathered for
<taxonomicName id="4C154D45FFA0FFE6FD19FDEBFAD7D622" box="[736,1003,583,608]" class="Hydrozoa" family="Agalmatidae" genus="Cardianecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siphonophorae" pageId="57" pageNumber="58" phylum="Cnidaria" rank="species" species="parchelion" status="sp. nov.">
<emphasis id="B961EAD4FFA0FFE6FD19FDEBFAD7D622" box="[736,1003,583,608]" italics="true" pageId="57" pageNumber="58">Cardianecta parchelion</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25257AFFFA0FFE6FC0AFDEBFD72D622" box="[1011,1102,583,608]" pageId="57" pageNumber="58" rank="species">
<emphasis id="B961EAD4FFA0FFE6FC0AFDEBFD72D622" bold="true" box="[1011,1102,583,608]" pageId="57" pageNumber="58">sp. nov.</emphasis>
</taxonomicNameLabel>
one can only speculate on its systematic relationships.
</paragraph>
</subSubSection>
<subSubSection id="C30F654DFFA0FFE6FF3EFD23F838D6B2" pageId="57" pageNumber="58" type="etymology">
<paragraph id="8BAA36C6FFA0FFE6FF3EFD23F838D6B2" blockId="57.[151,1437,151,752]" pageId="57" pageNumber="58">
<emphasis id="B961EAD4FFA0FFE6FF3EFD23F872D6EA" bold="true" box="[199,334,655,680]" pageId="57" pageNumber="58">Etymology.</emphasis>
The generic name refers to the heart-shaped nectophore. The specific name is derived from the Greek &quot;like a chela&quot;, and refers to the supposed chela-like appearance of the &quot;horns&quot; on the terminal process of the tentillum.
</paragraph>
</subSubSection>
</treatment>
</document>