453 lines
63 KiB
XML
453 lines
63 KiB
XML
<document id="DFE307261DF191D21990065110A5123D" ID-CLB-Dataset="54828" ID-DOI="10.5733/afin.054.0101" ID-GBIF-Dataset="753510a6-cffa-4637-a1bd-e9ae7d5d1660" ID-ISSN="2305-2562" ID-Zenodo-Dep="7670100" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1676639734503" checkinUser="felipe" docAuthor="Griffiths, O. L. & Herbert, D. G." docDate="2013" docId="03BC1F3CE509D93107B2FBD145B1B426" docLanguage="en" docName="AfrInvertebr.54.1.1-48.pdf" docOrigin="African Invertebrates 54 (1)" docStyle="DocumentStyle:0201127D758FBEB889983D376DC13FCB.4:AfrInvertebr.2008-2015.journal_article.1cover" docStyleId="0201127D758FBEB889983D376DC13FCB" docStyleName="AfrInvertebr.2008-2015.journal_article.1cover" docStyleVersion="4" docTitle="Kalidos maryannae Griffiths & Herbert 2013, sp. n." docType="treatment" docUuid="661CEFA0-FFA5-4FC6-BB4A-6ACE9524992F" docUuidSource="ZooBank" docVersion="8" lastPageNumber="38" masterDocId="FF856744E529D917064DFF964621B743" masterDocTitle="New species of land snails (Mollusca: Gastropoda) from two isolated karst formations in central western Madagascar: Tsingy Beanka and Antsingimavo, with additional notes on other regional endemics" masterLastPageNumber="48" masterPageNumber="1" pageNumber="32" updateTime="1699475339093" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:titleInfo id="44AF1948BF28D8704691986BDD0D299D">
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<mods:title id="5382DD1A39B129C13B955FCDF085E573">New species of land snails (Mollusca: Gastropoda) from two isolated karst formations in central western Madagascar: Tsingy Beanka and Antsingimavo, with additional notes on other regional endemics</mods:title>
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<mods:name id="6C0E032B81342006034C74D0737157E6" type="personal">
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<mods:role id="ECDCE13A004573C7D8D13CCB6337C2E4">
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<mods:roleTerm id="CA5CD14400FBFD54F90C2826CD7D8658">Author</mods:roleTerm>
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<mods:namePart id="1C7238B39911046C5D5EB0CC404D915B">Griffiths, O. L.</mods:namePart>
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<mods:affiliation id="E0D694083B8822C5A150696417152086">Research associate of the Australian Museum, Sydney, Australia and Biodiversity Conservation Madagascar (BCM) Ltd, Rivière des Anguilles, Mauritius</mods:affiliation>
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<mods:nameIdentifier id="6940FEDDD3A865D5B86EF8BEB384928B" type="email">owen@bcm.intnet.mu</mods:nameIdentifier>
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<mods:roleTerm id="E994C66784584E41F975214280D9F5E9">Author</mods:roleTerm>
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<mods:namePart id="0B2A6C570CA54F51CB3971192D6AA1DC">Herbert, D. G.</mods:namePart>
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<mods:affiliation id="A3631852D42F7608809A73F4918E7283">KwaZulu-Natal Museum, P. Bag 9070, Pietermaritzburg, 3200 South Africa and School of Life Sciences, University of KwaZulu-Natal, Pietermaritzburg, 3206 South Africa</mods:affiliation>
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<mods:nameIdentifier id="84C302CEAAA3846B9E5056E370888581" type="email">dherbert@nmsa.org.za</mods:nameIdentifier>
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<mods:typeOfResource id="60C3CE6AC010E69025FA23CE01B34292">text</mods:typeOfResource>
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<mods:title id="A739722D9F51D81FFA66578197E5EA42">African Invertebrates</mods:title>
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<mods:part id="2505858DF1F3E21038A2841B333EF0C4">
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<mods:date id="46ECD71566533BE811B6648B754163E4">2013</mods:date>
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<mods:number id="BBC88D1F2D21F2545E0F8E761891E8E4">2013-02-13</mods:number>
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<mods:number id="99B189C9D0E238EBDE044CA97A132A7C">54</mods:number>
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<treatment id="03BC1F3CE509D93107B2FBD145B1B426" ID-DOI="http://doi.org/10.5281/zenodo.7670273" ID-GBIF-Taxon="209341539" ID-Zenodo-Dep="7670273" LSID="urn:lsid:zoobank.org:act:661CEFA0-FFA5-4FC6-BB4A-6ACE9524992F" httpUri="http://treatment.plazi.org/id/03BC1F3CE509D93107B2FBD145B1B426" lastPageId="38" lastPageNumber="38" pageId="32" pageNumber="32">
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<subSubSection id="C30FFDA1E509D93707B2FBD1450DB321" box="[511,812,1095,1122]" pageId="32" pageNumber="32" type="nomenclature">
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<paragraph id="8BAAAE2AE509D93707B2FBD1450DB321" blockId="32.[511,812,1095,1122]" box="[511,812,1095,1122]" pageId="32" pageNumber="32">
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<heading id="D0E21946E509D93707B2FBD1450DB321" box="[511,812,1095,1122]" centered="true" fontSize="11" level="2" pageId="32" pageNumber="32" reason="2">
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<taxonomicName id="4C15D5A9E509D93707B2FBD144C8B321" ID-CoL="3QYYB" authority="Griffiths & Herbert, 2013" authorityName="Griffiths & Herbert" authorityYear="2013" box="[511,745,1095,1122]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="32" pageNumber="32" phylum="Mollusca" rank="species" species="maryannae" status="sp. nov.">
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<emphasis id="B9617238E509D93707B2FBD144C8B321" bold="true" box="[511,745,1095,1122]" pageId="32" pageNumber="32">Kalidos maryannae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A252CF43E509D93704BDFBDE450DB321" box="[752,812,1096,1122]" pageId="32" pageNumber="32" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C30FFDA1E509D9310475FBF945B1B426" lastPageId="38" lastPageNumber="38" pageId="32" pageNumber="32" type="description">
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<paragraph id="8BAAAE2AE509D9370475FBF944D2B3CA" blockId="32.[568,755,1135,1161]" box="[568,755,1135,1161]" pageId="32" pageNumber="32">
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<figureCitation id="132EB2AFE509D9370475FBF944ACB3CA" box="[568,653,1135,1161]" captionStart="Fig" captionStartId="33.[143,175,1082,1103]" captionTargetBox="[145,1101,214,1060]" captionTargetId="figure-155@33.[143,1103,204,1068]" captionTargetPageId="33" captionText="Fig.23. Kalidos maryannae sp.n.:(A–C) holotype, max.diameter 27.3 mm,height17.7 mm,AMSC.474168; (D–J) paratypes, NMSA L8470/T2904: (D) specimen with bands of intermediate strength, max. diameter 29.3 mm; (E) specimen with broad brown bands and low spire, max.diameter 27.0 mm; (F) specimen spire of intermediate height, max. diameter 28.3 mm; (G, H) fresh subadult with unfaded colour pattern, max. diameter 24.8 mm; (I) large specimen with additional ⅛ whorl of growth, max. diameter 29.3 mm; (J) juvenile showing angular periphery, max. diameter 15.2 mm; (K) holotype, portion of last adult whorl showing sculpture of growth-lines and spiral threads." figureDoi="http://doi.org/10.5281/zenodo.7670157" httpUri="https://zenodo.org/record/7670157/files/figure.png" pageId="32" pageNumber="32">Figs 23</figureCitation>
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,
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<figureCitation id="132EB2AFE509D93704D4FBF94496B3CA" box="[665,695,1135,1161]" captionStart="Fig" captionStartId="34.[181,214,1510,1531]" captionTargetBox="[187,1137,895,1481]" captionTargetId="figure-244@34.[181,1141,870,1494]" captionTargetPageId="34" captionText="Fig. 24. Kalidos maryannae sp. n., genitalia, paratype, NMSA L8470/T2904: (A) entire reproductive tract; (B) penis opened longitudinally to show well-developed penial verge extending from epiphallus; (C) allospermatophore from bursa copulatrix, head end missing.Abbreviations:ag – albumen gland, bc – bursa copulatrix, bcd – bursa copulatrix duct, ep – epiphallus, f – flagellum, ga – genital atrium, og – oviducal gland, p – penis, ps – penis sheath, pv – penial verge, prm – penial retractor muscle, sc – spermatophore capsule, sod – sperm-oviduct, st – spermatophore tail, v – vagina, vd – vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7670159" httpUri="https://zenodo.org/record/7670159/files/figure.png" pageId="32" pageNumber="32">24</figureCitation>
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,
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<figureCitation id="132EB2AFE509D937048EFBF944D2B3CA" box="[707,755,1135,1161]" captionStart="Fig" captionStartId="36.[181,214,1462,1483]" captionTargetBox="[190,1134,203,1445]" captionTargetId="figure-127@36.[189,1134,203,1445]" captionTargetPageId="36" captionText="Fig. 25. Living animals from Tsingy Beanka: (A) Acroptychia bathiei Fischer-Piette & Bedoucha, 1965, st’n 13/06, shell diameter 23.2 mm, NMSA L7315; (B) Tropidophora humbug sp. n., st’n 12/06, shell diameter 28.3 mm, paratype,NMSA L7204/T2984; (C) Tropidophora secunda Fischer-Piette & Bedoucha, 1965, st’n 03/06, shell diameter approx.19.0 mm; (D) Clavator griffithsjonesi Emberton, 1999, st’n 12/06, shell height 94.8 mm, NMSA L7192; (E) Ampelita milloti Fischer-Piette,1952,st’n 07/09, shell diameter 19.2 mm, NMSA L8502; (F) Ampelita namerokoensis Fischer-Piette, 1952, st’n 12/06, shell diameter 20.3 mm, NMSA L8442; (G) Kalidos griffithshauchleri Emberton, 2002, st’n 12/06, shell diameter approx. 60 mm; (H) Kalidos maryannae sp. n., st’n 12/06, shell diameter 22.5 mm, paratype, NMSA L7193/T2944." figureDoi="http://doi.org/10.5281/zenodo.7670161" httpUri="https://zenodo.org/record/7670161/files/figure.png" pageId="32" pageNumber="32">25H</figureCitation>
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</paragraph>
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<paragraph id="8BAAAE2AE509D93706F8FB0C4584B3F7" blockId="32.[181,1142,1178,1676]" box="[181,933,1178,1204]" pageId="32" pageNumber="32">Etymology: Named for Mary-Ann Stanley, wife of Owen Griffiths.</paragraph>
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<paragraph id="8BAAAE2AE509D93706F8FB544499B27F" blockId="32.[181,1142,1178,1676]" pageId="32" pageNumber="32">Diagnosis: Shell globose-lenticular, periphery rounded, aperture weakly descendant, umbilicus narrow; sculpture of fine, uneven spiral lines crossed by growth-lines; lustreless, pale coffee-brown, with a bold white spiral band just above periphery, bordered above and below by thinner, dark brown spiral lines.</paragraph>
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<paragraph id="8BAAAE2AE509D93706F8FADC4760B227" blockId="32.[181,1142,1178,1676]" box="[181,321,1354,1380]" pageId="32" pageNumber="32">Description:</paragraph>
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<paragraph id="8BAAAE2AE509D93606F8FAE34727B288" blockId="32.[181,1142,1178,1676]" lastBlockId="33.[144,1105,1297,1676]" lastPageId="33" lastPageNumber="33" pageId="32" pageNumber="32">
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<emphasis id="B9617238E509D93706F8FAE346CCB2CE" box="[181,237,1397,1421]" italics="true" pageId="32" pageNumber="32">Shell</emphasis>
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: Medium sized, globose-lenticular with a relatively elevated spire (H:D=0.57– 0.67); periphery at mid-whorl, rounded in adults, somewhat angled in juveniles; suture indented but not strongly so, situated at whorl periphery, occasionally slightly descendant prior to aperture; umbilicus narrow. Protoconch of approx 1¼ whorls, smooth except for numerous, close-set, microscopic, spiral threads; junction with teleoconch often indistinct. Teleoconch of a further 3½–4 whorls; initially lustreless and smooth, save for weak growth-lines, but with fine, irregular spiral sculpture developing with growth; last adult whorl with surface cut by somewhat stronger, rather uneven spiral lines, crossed by irregular growth-lines, creating low, weakly and unevenly pustular spiral ridges (
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<figureCitation id="132EB2AFE508D93606AEFA87476BB268" box="[227,330,1297,1323]" captionStart="Fig" captionStartId="33.[143,175,1082,1103]" captionTargetBox="[145,1101,214,1060]" captionTargetId="figure-155@33.[143,1103,204,1068]" captionTargetPageId="33" captionText="Fig.23. Kalidos maryannae sp.n.:(A–C) holotype, max.diameter 27.3 mm,height17.7 mm,AMSC.474168; (D–J) paratypes, NMSA L8470/T2904: (D) specimen with bands of intermediate strength, max. diameter 29.3 mm; (E) specimen with broad brown bands and low spire, max.diameter 27.0 mm; (F) specimen spire of intermediate height, max. diameter 28.3 mm; (G, H) fresh subadult with unfaded colour pattern, max. diameter 24.8 mm; (I) large specimen with additional ⅛ whorl of growth, max. diameter 29.3 mm; (J) juvenile showing angular periphery, max. diameter 15.2 mm; (K) holotype, portion of last adult whorl showing sculpture of growth-lines and spiral threads." figureDoi="http://doi.org/10.5281/zenodo.7670157" httpUri="https://zenodo.org/record/7670157/files/figure.png" pageId="33" pageNumber="33">Fig. 23K</figureCitation>
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), these more distinct toward periphery, but evanescing on base which is more glossy and sculptured only with close-set, microscopic, incised spiral threads, rendered irregularly undulant by growth-lines. Aperture ovate-reniform, peristome strongly oblique to vertical axis of shell, interrupted by base of penultimate whorl; outer lip simple and thin; upper part of columella lip reflected and partially covering umbilicus.
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</paragraph>
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<caption id="DF6AFEA2E508D93606C2FBAC4584B39C" ID-DOI="http://doi.org/10.5281/zenodo.7670157" ID-Zenodo-Dep="7670157" httpUri="https://zenodo.org/record/7670157/files/figure.png" pageId="33" pageNumber="33" startId="33.[143,175,1082,1103]" targetBox="[145,1101,214,1060]" targetPageId="33">
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<paragraph id="8BAAAE2AE508D93606C2FBAC4584B39C" blockId="33.[143,1105,1082,1247]" pageId="33" pageNumber="33">
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Fig. 23.
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<taxonomicName id="4C15D5A9E508D936069BFBAA47A2B30C" authority="Griffiths & Herbert, 2013" authorityName="Griffiths & Herbert" authorityYear="2013" box="[214,387,1084,1103]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="33" pageNumber="33" phylum="Mollusca" rank="species" species="maryannae" status="sp. nov.">
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<emphasis id="B9617238E508D936069BFBAA47A2B30C" box="[214,387,1084,1103]" italics="true" pageId="33" pageNumber="33">Kalidos maryannae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A252CF43E508D93607CAFBAC4799B30C" box="[391,440,1082,1103]" pageId="33" pageNumber="33" rank="species">sp. n.</taxonomicNameLabel>
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: (A–C) holotype, max.diameter 27.3 mm, height 17.7 mm,AMS C.474168; (D–J) paratypes, NMSA L8470/T2904: (D) specimen with bands of intermediate strength, max. diameter 29.3 mm; (E) specimen with broad brown bands and low spire, max. diameter 27.0 mm; (F) specimen spire of intermediate height, max. diameter 28.3 mm; (G, H) fresh subadult with unfaded colour pattern, max. diameter 24.8 mm; (I) large specimen with additional ⅛ whorl of growth, max. diameter 29.3 mm; (J) juvenile showing angular periphery, max. diameter 15.2 mm; (K) holotype, portion of last adult whorl showing sculpture of growth-lines and spiral threads.
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</paragraph>
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</caption>
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<paragraph id="8BAAAE2AE508D93506E7FA474247B665" blockId="33.[144,1105,1297,1676]" lastBlockId="34.[181,1142,203,822]" lastPageId="34" lastPageNumber="34" pageId="33" pageNumber="33">
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Shell strikingly coloured, with a bold white spiral band (1.0–2.0 mm wide) just above periphery, bordered above and below by thinner, dark brown spiral lines; white band and upper brown line visible on penultimate whorl, but generally absent on spire whorls; remainder of shell rather unevenly washed with buff to pale coffee-brown in fresh specimens, fading to dirty white with time and loss of periostracum. Colour bands often ceasing a short distance before lip edge, and clearly visible inside aperture. In the largest specimens, an additional ⅛ whorl is added to the mature lip (
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<figureCitation id="132EB2AFE50BD935058DFF5D423FB7A6" box="[960,1054,203,229]" captionStart="Fig" captionStartId="33.[143,175,1082,1103]" captionTargetBox="[145,1101,214,1060]" captionTargetId="figure-155@33.[143,1103,204,1068]" captionTargetPageId="33" captionText="Fig.23. Kalidos maryannae sp.n.:(A–C) holotype, max.diameter 27.3 mm,height17.7 mm,AMSC.474168; (D–J) paratypes, NMSA L8470/T2904: (D) specimen with bands of intermediate strength, max. diameter 29.3 mm; (E) specimen with broad brown bands and low spire, max.diameter 27.0 mm; (F) specimen spire of intermediate height, max. diameter 28.3 mm; (G, H) fresh subadult with unfaded colour pattern, max. diameter 24.8 mm; (I) large specimen with additional ⅛ whorl of growth, max. diameter 29.3 mm; (J) juvenile showing angular periphery, max. diameter 15.2 mm; (K) holotype, portion of last adult whorl showing sculpture of growth-lines and spiral threads." figureDoi="http://doi.org/10.5281/zenodo.7670157" httpUri="https://zenodo.org/record/7670157/files/figure.png" pageId="34" pageNumber="34">Fig. 23I</figureCitation>
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). These may represent individuals that survive through the dry season after maturity has been reached and begin another phase of growth when the rains return the following year.
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</paragraph>
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<paragraph id="8BAAAE2AE50BD935069DFEBD44A7B625" blockId="34.[181,1142,203,822]" pageId="34" pageNumber="34">
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Dimensions:
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<typeStatus id="54AE1088E50BD9350728FEBD47F1B606" box="[357,464,299,325]" pageId="34" pageNumber="34" type="holotype">Holotype</typeStatus>
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, max. diameter
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<quantity id="4CED03CFE50BD93504CFFEBD44C9B605" box="[642,744,299,326]" metricMagnitude="-2" metricUnit="m" metricValue="2.73" pageId="34" pageNumber="34" unit="mm" value="27.3">27.3 mm</quantity>
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, height
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<quantity id="4CED03CFE50BD935050CFEBD4589B605" box="[833,936,299,326]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" pageId="34" pageNumber="34" unit="mm" value="17.7">17.7 mm</quantity>
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; largest specimen, max. diameter
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<quantity id="4CED03CFE50BD9350710FEDD47E2B625" box="[349,451,331,358]" metricMagnitude="-2" metricUnit="m" metricValue="2.94" pageId="34" pageNumber="34" unit="mm" value="29.4">29.4 mm</quantity>
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, height
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<quantity id="4CED03CFE50BD9350456FEDD44A3B625" box="[539,642,331,358]" metricMagnitude="-2" metricUnit="m" metricValue="1.88" pageId="34" pageNumber="34" unit="mm" value="18.8">18.8 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BAAAE2AE50BD93506F8FEE04419B54E" blockId="34.[181,1142,203,822]" pageId="34" pageNumber="34">
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<emphasis id="B9617238E50BD93506F8FEE04756B6CD" box="[181,375,374,398]" italics="true" pageId="34" pageNumber="34">External features</emphasis>
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(
|
||
<figureCitation id="132EB2AFE50BD93507CAFEE547D0B6CE" box="[391,497,371,397]" captionStart="Fig" captionStartId="36.[181,214,1462,1483]" captionTargetBox="[190,1134,203,1445]" captionTargetId="figure-127@36.[189,1134,203,1445]" captionTargetPageId="36" captionText="Fig. 25. Living animals from Tsingy Beanka: (A) Acroptychia bathiei Fischer-Piette & Bedoucha, 1965, st’n 13/06, shell diameter 23.2 mm, NMSA L7315; (B) Tropidophora humbug sp. n., st’n 12/06, shell diameter 28.3 mm, paratype,NMSA L7204/T2984; (C) Tropidophora secunda Fischer-Piette & Bedoucha, 1965, st’n 03/06, shell diameter approx.19.0 mm; (D) Clavator griffithsjonesi Emberton, 1999, st’n 12/06, shell height 94.8 mm, NMSA L7192; (E) Ampelita milloti Fischer-Piette,1952,st’n 07/09, shell diameter 19.2 mm, NMSA L8502; (F) Ampelita namerokoensis Fischer-Piette, 1952, st’n 12/06, shell diameter 20.3 mm, NMSA L8442; (G) Kalidos griffithshauchleri Emberton, 2002, st’n 12/06, shell diameter approx. 60 mm; (H) Kalidos maryannae sp. n., st’n 12/06, shell diameter 22.5 mm, paratype, NMSA L7193/T2944." figureDoi="http://doi.org/10.5281/zenodo.7670161" httpUri="https://zenodo.org/record/7670161/files/figure.png" pageId="34" pageNumber="34">Fig. 25H</figureCitation>
|
||
): Head-foot mostly pale, but somewhat darker anteriorly, with yellowish white tubercles contrasting with darker, brownish skin grooves; lateral pedal grooves and peripodial groove distinct; caudal pit and surmounting horn well developed; optic tentacles darker greyish brown; mantle lobes pale with numerous yellowish white pigment granules.
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE50BD93406F8FD884519B506" blockId="34.[181,1142,203,822]" lastBlockId="35.[143,1105,203,1675]" lastPageId="35" lastPageNumber="35" pageId="34" pageNumber="34">
|
||
<emphasis id="B9617238E50BD93506F8FD884744B575" box="[181,357,542,566]" italics="true" pageId="34" pageNumber="34">Distal genitalia</emphasis>
|
||
(
|
||
<figureCitation id="132EB2AFE50BD9350739FD8D47E9B576" box="[372,456,539,565]" captionStart="Fig" captionStartId="34.[181,214,1510,1531]" captionTargetBox="[187,1137,895,1481]" captionTargetId="figure-244@34.[181,1141,870,1494]" captionTargetPageId="34" captionText="Fig. 24. Kalidos maryannae sp. n., genitalia, paratype, NMSA L8470/T2904: (A) entire reproductive tract; (B) penis opened longitudinally to show well-developed penial verge extending from epiphallus; (C) allospermatophore from bursa copulatrix, head end missing.Abbreviations:ag – albumen gland, bc – bursa copulatrix, bcd – bursa copulatrix duct, ep – epiphallus, f – flagellum, ga – genital atrium, og – oviducal gland, p – penis, ps – penis sheath, pv – penial verge, prm – penial retractor muscle, sc – spermatophore capsule, sod – sperm-oviduct, st – spermatophore tail, v – vagina, vd – vas deferens." figureDoi="http://doi.org/10.5281/zenodo.7670159" httpUri="https://zenodo.org/record/7670159/files/figure.png" pageId="34" pageNumber="34">Fig. 24</figureCitation>
|
||
): Penis of moderate length (approx.
|
||
<quantity id="4CED03CFE50BD935052DFD8D45E9B575" box="[864,968,539,566]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="34" pageNumber="34" unit="mm" value="12.5">12.5 mm</quantity>
|
||
), apical portion swollen and with a deep indentation; basal ¾ encased in a thin sheath; apical bulb of penis containing a well-developed verge; verge much contracted but possessing a deep longitudinal groove confluent with opening of epiphallus; lower portion of penis with 5 or 6 broad, low folds with a microscopically velvety texture; epiphallus cylindrical, with a sharp bend at insertion of penial retractor muscle and thus divided into proximal and distal limbs; proximal limb slightly shorter than distal one (distal limb might also be considered part of penis rather than epiphallus); a short flagellum, comprising just over one tight loop, arises at junction of vas deferens and epiphallus; flagellum and vas deferens wrapped in connective tissue, but flagellum not connected to penis by muscle bands; lumen of flagellum with a rod-like process extending from its blind end; vas deferens passes along vagina and free oviduct to its origin at base of sperm-oviduct; retractor muscle of right optic tentacle passes to right of penis, i.e. between penis and vagina; bursa copulatrix large, narrowing apically and with a short broad duct; wall of duct thick and muscular, lined internally with longitudinal folds, wall of bursa much thinner; bursa with one moderately fresh allospermatophore and the remains of at least one more; free oviduct swollen prior to its junction with vagina (peri-vaginal or oviducal gland); sperm-oviduct well developed, twisted and with numerous superficial folds. Spermatophore with well-developed capsule (head and body) and a looped tail piece corresponding with shape of flagellum; tail piece with prominent flange-like spiral ridge; a small ‘filling nipple’ present at level of vas deferens.
|
||
</paragraph>
|
||
<caption id="DF6AFEA2E50BD93506F8FA704768B1C8" ID-DOI="http://doi.org/10.5281/zenodo.7670159" ID-Zenodo-Dep="7670159" httpUri="https://zenodo.org/record/7670159/files/figure.png" pageId="34" pageNumber="34" startId="34.[181,214,1510,1531]" targetBox="[187,1137,895,1481]" targetPageId="34">
|
||
<paragraph id="8BAAAE2AE50BD93506F8FA704768B1C8" blockId="34.[181,1142,1510,1675]" pageId="34" pageNumber="34">
|
||
Fig. 24.
|
||
<taxonomicName id="4C15D5A9E50BD935074DFA7E4793B2B8" authority="Griffiths & Herbert, 2013" authorityName="Griffiths & Herbert" authorityYear="2013" box="[256,434,1512,1531]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="34" pageNumber="34" phylum="Mollusca" rank="species" species="maryannae" status="sp. nov.">
|
||
<emphasis id="B9617238E50BD935074DFA7E4793B2B8" box="[256,434,1512,1531]" italics="true" pageId="34" pageNumber="34">Kalidos maryannae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A252CF43E50BD93507FAFA7047C8B2B8" box="[439,489,1510,1531]" pageId="34" pageNumber="34" rank="species">sp. n.</taxonomicNameLabel>
|
||
, genitalia, paratype, NMSA L8470/T2904: (A) entire reproductive tract; (B) penis opened longitudinally to show well-developed penial verge extending from epiphallus; (C) allospermatophore from bursa copulatrix, head end missing.Abbreviations: ag – albumen gland, bc – bursa copulatrix, bcd – bursa copulatrix duct, ep – epiphallus, f – flagellum, ga – genital atrium, og – oviducal gland, p – penis, ps – penis sheath, pv – penial verge, prm – penial retractor muscle, sc – spermatophore capsule, sod – sperm-oviduct, st – spermatophore tail, v – vagina, vd – vas deferens.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAAAE2AE50AD93406DDFDC44768B459" blockId="35.[143,1105,203,1675]" pageId="35" pageNumber="35">
|
||
<materialsCitation id="3B7DA477E50AD93406DDFDC4426DB5D4" collectingDate="2010-02" collectionCode="AMS" collectorName="R. Randalana" country="Madagascar" elevation="345" latitude="-18.06325" location="Tana - Maintirano" longLatPrecision="1" longitude="44.52861" pageId="35" pageNumber="35" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54AE1088E50AD93406DDFDC446C9B524" box="[144,232,594,615]" pageId="35" pageNumber="35" type="holotype">Holotype</typeStatus>
|
||
:
|
||
<collectingCountry id="F302EEBAE50AD93406A2FDC447B5B524" box="[239,404,594,615]" name="Madagascar" pageId="35" pageNumber="35">MADAGASCAR</collectingCountry>
|
||
:
|
||
<collectorName id="26E0CBFCE50AD93407D6FDC4444AB524" box="[411,619,594,615]" pageId="35" pageNumber="35">Central W Madagascar</collectorName>
|
||
,
|
||
<emphasis id="B9617238E50AD934043FFDC244A6B524" box="[626,647,596,615]" italics="true" pageId="35" pageNumber="35">ca</emphasis>
|
||
<locationDeviation id="9A18B708E50AD93404C0FDC4457BB524" box="[653,858,594,615]" location="Maintirano" pageId="35" pageNumber="35">
|
||
<quantity id="4CED03CFE50AD93404C0FDC444E4B524" box="[653,709,594,615]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="35" pageNumber="35" unit="km" value="60.0">60 km</quantity>
|
||
E of Maintirano
|
||
</locationDeviation>
|
||
, S part of
|
||
<taxonomicName id="4C15D5A9E50AD93405FEFDC4421AB524" box="[947,1083,594,615]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
|
||
, S side of
|
||
<location id="8ECAF8F1E50AD934069DFDFC474DB53C" LSID="urn:lsid:plazi:treatment:03BC1F3CE509D93107B2FBD145B1B426:8ECAF8F1E50AD934069DFDFC474DB53C" box="[208,364,618,639]" country="Madagascar" latitude="-18.06325" longLatPrecision="1" longitude="44.52861" name="Tana - Maintirano" pageId="35" pageNumber="35">Tana–Maintirano</location>
|
||
road, above N bank of
|
||
<collectorName id="26E0CBFCE50AD9340476FDFC44F9B53C" box="[571,728,618,639]" pageId="35" pageNumber="35">Kimanambolo R.</collectorName>
|
||
, tall moist forest growing on limestone on south-facing slopes,
|
||
<geoCoordinate id="EE21C8EDE50AD934070EFD14478DB5D4" box="[323,428,642,663]" degrees="18.06325" direction="south" orientation="latitude" pageId="35" pageNumber="35" precision="1" value="-18.06325">18.06325°S</geoCoordinate>
|
||
<geoCoordinate id="EE21C8EDE50AD93407FDFD14443DB5D4" box="[432,540,642,663]" degrees="44.52861" direction="east" orientation="longitude" pageId="35" pageNumber="35" precision="1" value="44.52861">44.52861°E</geoCoordinate>
|
||
,
|
||
<emphasis id="B9617238E50AD9340469FD124418B5D4" box="[548,569,644,663]" italics="true" pageId="35" pageNumber="35">ca</emphasis>
|
||
<quantity id="4CED03CFE50AD9340470FD144457B5D4" box="[573,630,642,663]" metricMagnitude="2" metricUnit="m" metricValue="3.45" pageId="35" pageNumber="35" unit="m" value="345.0">
|
||
<elevation id="00384919E50AD9340470FD144457B5D4" box="[573,630,642,663]" metricMagnitude="2" metricUnit="m" metricValue="3.45" pageId="35" pageNumber="35" unit="m" value="345.0">345 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="FFAB88EAE50AD9340430FD14449FB5D4" box="[637,702,642,663]" pageId="35" pageNumber="35" value="2010-02">
|
||
<collectingDate id="EFEF7102E50AD9340430FD14449FB5D4" box="[637,702,642,663]" pageId="35" pageNumber="35" value="2010-02">ii.2010</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="26E0CBFCE50AD9340488FD14451CB5D4" box="[709,829,642,663]" pageId="35" pageNumber="35">R. Randalana</collectorName>
|
||
, st’n R01/10 (
|
||
<collectionCode id="ED0436EFE50AD93405FBFD1445C9B5D4" box="[950,1000,642,663]" pageId="35" pageNumber="35">AMS</collectionCode>
|
||
C.474168)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B7DA477E50AD93406DDFD33426DB591" collectingDate="2010-02" collectionCode="NMSA" collectorName="R. Randalana" country="Madagascar" elevation="345" latitude="-18.06325" location="Tana - Maintirano" longLatPrecision="1" longitude="44.52861" pageId="35" pageNumber="35" specimenCode="L8470, L7193, L8471" specimenCount="14" typeStatus="paratype">
|
||
<typeStatus id="54AE1088E50AD93406DDFD3346CDB5F9" box="[144,236,677,698]" pageId="35" pageNumber="35" type="paratype">Paratypes</typeStatus>
|
||
: Same data as holotype (
|
||
<collectionCode id="ED0436EFE50AD9340798FD334436B5F9" box="[469,535,677,698]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="35" pageNumber="35" type="Museum">NMSA</collectionCode>
|
||
<specimenCode id="DBB30651E50AD9340453FD33447BB5F9" box="[542,602,677,698]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="35" pageNumber="35" type="Museum">L8470</specimenCode>
|
||
/T2904,
|
||
<specimenCount id="9D1365A3E50AD93404E5FD334509B5F9" box="[680,808,677,698]" pageId="35" pageNumber="35" type="generic">10 specimens</specimenCount>
|
||
, body of one in alcohol); 12/06 (
|
||
<collectionCode id="ED0436EFE50AD93406DBFD2B46F9B591" box="[150,216,701,722]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="35" pageNumber="35" type="Museum">NMSA</collectionCode>
|
||
<specimenCode id="DBB30651E50AD9340691FD2B4739B591" box="[220,280,701,722]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="35" pageNumber="35" type="Museum">L7193</specimenCode>
|
||
/T2944,
|
||
<specimenCount id="9D1365A3E50AD934072EFD2B47F5B591" box="[355,468,701,722]" pageId="35" pageNumber="35" type="generic">3 specimens</specimenCount>
|
||
, body of one in alcohol); st’n 8/10 (
|
||
<collectionCode id="ED0436EFE50AD9340542FD2B4570B591" box="[783,849,701,722]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="35" pageNumber="35" type="Museum">NMSA</collectionCode>
|
||
<specimenCode id="DBB30651E50AD9340518FD2B45B0B591" box="[853,913,701,722]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="35" pageNumber="35" type="Museum">L8471</specimenCode>
|
||
/T2943,
|
||
<specimenCount id="9D1365A3E50AD9340591FD2B4267B591" box="[988,1094,701,722]" pageId="35" pageNumber="35" type="generic">1 specimen</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B7DA477E50AD93406DDFD43426DB5A9" box="[144,1100,725,746]" collectionCode="NMSA, AMS" country="Madagascar" pageId="35" pageNumber="35" specimenCode="L8475" specimenCount="13" typeStatus="paratype">
|
||
st’n R01/09 (
|
||
<collectionCode id="ED0436EFE50AD934074CFD434763B5A9" box="[257,322,725,746]" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="35" pageNumber="35" type="Museum">NMSA</collectionCode>
|
||
<specimenCode id="DBB30651E50AD934070BFD4347A0B5A9" box="[326,385,725,746]" collectionCode="NMSA" country="South Africa" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34985" lsid="urn:lsid:biocol.org:col:34985" name="KwaZulu-Natal Museum" pageId="35" pageNumber="35" type="Museum">L8475</specimenCode>
|
||
/T2912,
|
||
<specimenCount id="9D1365A3E50AD9340787FD434410B5A9" box="[458,561,725,746]" pageId="35" pageNumber="35" type="generic">1 specimen</specimenCount>
|
||
, body in alcohol); st’n 08/09 (
|
||
<collectionCode id="ED0436EFE50AD9340579FD434547B5A9" box="[820,870,725,746]" pageId="35" pageNumber="35">AMS</collectionCode>
|
||
C.469578,
|
||
<specimenCount id="9D1365A3E50AD9340586FD434267B5A9" box="[971,1094,725,746]" pageId="35" pageNumber="35" type="generic">12 specimens</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="3B7DA477E50AD93406DDFD7B4765B459" collectionCode="MNHN, TMAM" country="Madagascar" pageId="35" pageNumber="35" specimenCode="NHMUK 20120015, T166" specimenCount="12" typeStatus="paratype">
|
||
st’n 03/10 (
|
||
<collectionCode id="ED0436EFE50AD93406BAFD7B471FB441" box="[247,318,749,770]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="35">MNHN</collectionCode>
|
||
IM-2010-20072,
|
||
<specimenCount id="9D1365A3E50AD93407ADFD7B4473B441" box="[480,594,749,770]" pageId="35" pageNumber="35" type="generic">3 specimens</specimenCount>
|
||
;
|
||
<specimenCode id="DBB30651E50AD9340416FD7B4535B441" box="[603,788,749,770]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="35" pageNumber="35" type="Museum">NHMUK 20120015</specimenCode>
|
||
,
|
||
<specimenCount id="9D1365A3E50AD9340550FD7B45B0B441" box="[797,913,749,770]" pageId="35" pageNumber="35" type="generic">3 specimens</specimenCount>
|
||
); st’n 07/09 (
|
||
<collectionCode id="ED0436EFE50AD934024AFD7B426EB441" box="[1031,1103,749,770]" pageId="35" pageNumber="35">TMAM</collectionCode>
|
||
<specimenCode id="DBB30651E50AD93406DDFC9346E0B459" box="[144,193,773,794]" collectionCode="TMAM" pageId="35" pageNumber="35">T166</specimenCode>
|
||
,
|
||
<specimenCount id="9D1365A3E50AD9340686FC93471EB459" box="[203,319,773,794]" pageId="35" pageNumber="35" type="generic">6 specimens</specimenCount>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE50AD93406DDFCB3459CB411" blockId="35.[143,1105,203,1675]" pageId="35" pageNumber="35">
|
||
Additional locality data:
|
||
<emphasis id="B9617238E50AD934073BFCB147D2B479" box="[374,499,807,826]" italics="true" pageId="35" pageNumber="35">Antsingimavo</emphasis>
|
||
: st’n 04/06 (subfossil).
|
||
<taxonomicName id="4C15D5A9E50AD934049CFCB14577B479" box="[721,854,807,826]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">
|
||
<emphasis id="B9617238E50AD934049CFCB14577B479" box="[721,854,807,826]" italics="true" pageId="35" pageNumber="35">Tsingy Beanka</emphasis>
|
||
</taxonomicName>
|
||
: st’ns 12/06, 14/06, 15/06, 18/06, 02/09, 06/09, 07/09, 08/09, 09/09, 11/09, 01/10, 04/10, 06/10, 07/10, 08/10, 09/10.
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE50AD93406DDFCF747EAB4F8" blockId="35.[143,1105,203,1675]" pageId="35" pageNumber="35">
|
||
Distribution: Evidently now confined to the
|
||
<taxonomicName id="4C15D5A9E50AD93404DCFCF7451FB438" box="[657,830,865,891]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
|
||
, where it is moderately common; not recorded from the Tsingy de Bemaraha, but a single subfossil specimen was found at Antsingimavo.
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE50AD93406DDFC5F47ACB300" blockId="35.[143,1105,203,1675]" pageId="35" pageNumber="35">
|
||
Habitat: Fresh dead shells are found most commonly amongst limestone rocks, predominantly in the taller moister forests of the southern part of
|
||
<taxonomicName id="4C15D5A9E50AD9340502FC7F45DAB340" box="[847,1019,1001,1027]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
|
||
. Living specimens are usually found in leaf-litter.Appears to aestivate sealed to limestone rocks or deep in tsingy slots.
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE50AD93406DDFBC74489B228" blockId="35.[143,1105,203,1675]" pageId="35" pageNumber="35">
|
||
Remarks: In its moderate size and pattern of peripheral banding (two dark brown spiral lines separated by a broader whitish band) this species resembles
|
||
<taxonomicName id="4C15D5A9E50AD9340511FBE546F2B3E8" authority="(Robson, 1914)" baseAuthorityName="Robson" baseAuthorityYear="1914" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="bathensis">
|
||
<emphasis id="B9617238E50AD9340511FBE545C4B3C8" box="[860,997,1139,1163]" italics="true" pageId="35" pageNumber="35">K. bathensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EF84D3DBE50AD93405BCFBE746EAB3E8" author="ROBSON, G. C." pageId="35" pageNumber="35" pagination="375 - 389" refId="ref18819" refString="ROBSON, G. C. 1914. On a collection of land and freshwater Gastropoda from Madagascar, with descriptions of new genera and new species. Zoological Journal of the Linnean Society 32: 375 - 389." type="journal article" year="1914">Robson, 1914</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C15D5A9E50AD9340690FB0547D2B3E8" authority="Robson, 1914" authorityName="Robson" authorityYear="1914" box="[221,499,1169,1195]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="bournei">
|
||
<emphasis id="B9617238E50AD9340690FB054772B3E8" box="[221,339,1171,1195]" italics="true" pageId="35" pageNumber="35">K. bournei</emphasis>
|
||
<bibRefCitation id="EF84D3DBE50AD934071AFB0747D2B3E8" author="ROBSON, G. C." box="[343,499,1169,1195]" pageId="35" pageNumber="35" pagination="375 - 389" refId="ref18819" refString="ROBSON, G. C. 1914. On a collection of land and freshwater Gastropoda from Madagascar, with descriptions of new genera and new species. Zoological Journal of the Linnean Society 32: 375 - 389." type="journal article" year="1914">Robson, 1914</bibRefCitation>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C15D5A9E50AD93407B0FB05451AB3E8" authority="(Angas, 1877)" baseAuthorityName="Angas" baseAuthorityYear="1877" box="[509,827,1169,1195]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="ekongensis">
|
||
<emphasis id="B9617238E50AD93407B0FB0544B7B3E8" box="[509,662,1171,1195]" italics="true" pageId="35" pageNumber="35">K. ekongensis</emphasis>
|
||
(Angas, 1877)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C15D5A9E50AD9340508FB05426AB3E8" authority="(Odhner, 1919)" baseAuthorityName="Odhner" baseAuthorityYear="1919" box="[837,1099,1169,1195]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="hova">
|
||
<emphasis id="B9617238E50AD9340508FB0545BBB3E8" box="[837,922,1171,1195]" italics="true" pageId="35" pageNumber="35">K. hova</emphasis>
|
||
(Odhner, 1919)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C15D5A9E50AD93406F0FB254432B388" authority="Emberton, 2007" authorityName="Emberton" authorityYear="2007" box="[189,531,1201,1227]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="namorokae">
|
||
<emphasis id="B9617238E50AD93406F0FB254778B388" box="[189,345,1203,1227]" italics="true" pageId="35" pageNumber="35">K. namorokae</emphasis>
|
||
Emberton, 2007
|
||
</taxonomicName>
|
||
. In
|
||
<taxonomicName id="4C15D5A9E50AD9340475FB25448FB388" authorityName="Robson" authorityYear="1914" box="[568,686,1203,1227]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="bournei">
|
||
<emphasis id="B9617238E50AD9340475FB25448FB388" box="[568,686,1203,1227]" italics="true" pageId="35" pageNumber="35">K. bournei</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C15D5A9E50AD93404ACFB25455CB388" authorityName="Emberton" authorityYear="2007" box="[737,893,1203,1227]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="namorokae">
|
||
<emphasis id="B9617238E50AD93404ACFB25455CB388" box="[737,893,1203,1227]" italics="true" pageId="35" pageNumber="35">K. namorokae</emphasis>
|
||
</taxonomicName>
|
||
, however, the shell is smaller and the upper brown spiral line is situated further from the whorl periphery. In contrast,
|
||
<taxonomicName id="4C15D5A9E50AD93406BBFB6547A1B248" baseAuthorityName="Robson" baseAuthorityYear="1914" box="[246,384,1267,1291]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="bathensis">
|
||
<emphasis id="B9617238E50AD93406BBFB6547A1B248" box="[246,384,1267,1291]" italics="true" pageId="35" pageNumber="35">K. bathensis</emphasis>
|
||
</taxonomicName>
|
||
is larger than the present species, paler in colour and the median spiral band between the brown spiral lines is not distinctly paler than the remaining shell. In addition, it has a third brown spiral line below the suture and the supra-peripheral spiral line is clearly present on the spire whorls.
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE50AD93406E7FAE74412B1C8" blockId="35.[143,1105,203,1675]" pageId="35" pageNumber="35">
|
||
Undoubtedly, the most similar species in terms of size and colour pattern are
|
||
<taxonomicName id="4C15D5A9E50AD93405BEFAE546F4B2E8" baseAuthorityName="Angas" baseAuthorityYear="1877" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="ekongensis">
|
||
<emphasis id="B9617238E50AD93405BEFAE546F4B2E8" italics="true" pageId="35" pageNumber="35">K. ekongensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C15D5A9E50AD934074AFA05477DB2E8" baseAuthorityName="Odhner" baseAuthorityYear="1919" box="[263,348,1427,1451]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="hova">
|
||
<emphasis id="B9617238E50AD934074AFA05477DB2E8" box="[263,348,1427,1451]" italics="true" pageId="35" pageNumber="35">K. hova</emphasis>
|
||
</taxonomicName>
|
||
, both of which, as presently conceived, are poorly delineated species. In his description of
|
||
<taxonomicName id="4C15D5A9E50AD9340727FA254445B288" authority="Odhner (1919)" authorityName="Odhner" authorityYear="1919" box="[362,612,1457,1483]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="hova">
|
||
<emphasis id="B9617238E50AD9340727FA25479EB288" box="[362,447,1459,1483]" italics="true" pageId="35" pageNumber="35">K. hova</emphasis>
|
||
Odhner (1919)
|
||
</taxonomicName>
|
||
noted a fawn peripheral band on an otherwise brownish shell, but he made no mention of this band being bordered above and below by dark brown spiral lines, neither are these clearly evident in his figure nor in the type material (
|
||
<figureCitation id="132EB2AFE50AD93406B1F987475CB168" box="[252,381,1553,1579]" captionStart="Fig" captionStartId="37.[144,176,1510,1531]" captionTargetBox="[157,1095,235,1476]" captionTargetId="figure-89@37.[147,1108,223,1487]" captionTargetPageId="37" captionText="Fig.26.Speciesillustrated for comparative purposes:(A–C) Tropidophora moulinsii, holotype of Cyclostoma moulinsii Grateloup, 1840, ‘Madagascar’, max. diameter 33.1 mm, NHMUK 1907.11.22.12; (D–F) Tropidophora thesauri Fischer-Piette, 1949, Orangea, Diego Suarez [= Antsiranana] district, max. diameter 38.0 mm, NMSA L8438; (G–I) Tropidophora reticulata, one of three syntypes of Cyclostoma reticulatum (Adams & Reeve, 1850), ‘Madagascar’, max.diameter 15.1 mm, NHMUK 1874.12.11.231; (J–L) Kalidos hova, syntype of Nanina hova Odhner, 1919, Catsèpe, Madagascar, max. diameter 24.4 mm (SMNH)." figureDoi="http://doi.org/10.5281/zenodo.7670163" httpUri="https://zenodo.org/record/7670163/files/figure.png" pageId="35" pageNumber="35">Fig. 26J–L</figureCitation>
|
||
,
|
||
<typeStatus id="54AE1088E50AD93407C4F98447C2B16F" box="[393,483,1554,1580]" pageId="35" pageNumber="35" type="syntype">syntype</typeStatus>
|
||
, SMNH). He also stated that the shell was smooth and glossy and that the spiral microsculpture was more distinct on the base. This is not the case in
|
||
<taxonomicName id="4C15D5A9E50AD93406AEF9C547A3B128" authorityName="Griffiths & Herbert" authorityYear="2013" box="[227,386,1619,1643]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="35" pageNumber="35" phylum="Mollusca" rank="species" species="maryannae">
|
||
<emphasis id="B9617238E50AD93406AEF9C547A3B128" box="[227,386,1619,1643]" italics="true" pageId="35" pageNumber="35">K. maryannae</emphasis>
|
||
</taxonomicName>
|
||
in which the shell is lustreless and the spiral sculpture strongest above the periphery of the last whorl.
|
||
</paragraph>
|
||
<caption id="DF6AFEA2E50DD93306F8FA204455B1C8" ID-DOI="http://doi.org/10.5281/zenodo.7670161" ID-Zenodo-Dep="7670161" httpUri="https://zenodo.org/record/7670161/files/figure.png" pageId="36" pageNumber="36" startId="36.[181,214,1462,1483]" targetBox="[190,1134,203,1445]" targetPageId="36">
|
||
<paragraph id="8BAAAE2AE50DD93306F8FA204455B1C8" blockId="36.[181,1142,1462,1675]" pageId="36" pageNumber="36">
|
||
Fig. 25. Living animals from
|
||
<taxonomicName id="4C15D5A9E50DD933078AFA204475B288" box="[455,596,1462,1483]" class="Gastropoda" family="Cyclophoridae" genus="Tsingy" kingdom="Animalia" order="Architaenioglossa" pageId="36" pageNumber="35" phylum="Mollusca" rank="species" species="beanka">Tsingy Beanka</taxonomicName>
|
||
: (A)
|
||
<taxonomicName id="4C15D5A9E50DD93304CEFA2E4253B288" authority="Fischer-Piette & Bedoucha, 1965" authorityName=": Fischer-Piette & Bedoucha" authorityYear="1965" box="[643,1138,1462,1483]" class="Gastropoda" family="Cyclophoridae" genus="Acroptychia" kingdom="Animalia" order="Architaenioglossa" pageId="36" pageNumber="36" phylum="Mollusca" rank="species" species="bathiei">
|
||
<emphasis id="B9617238E50DD93304CEFA2E4516B288" box="[643,823,1464,1483]" italics="true" pageId="36" pageNumber="36">Acroptychia bathiei</emphasis>
|
||
Fischer-Piette & Bedoucha, 1965
|
||
</taxonomicName>
|
||
, st’n 13/06, shell diameter 23.2 mm, NMSA L7315; (B)
|
||
<taxonomicName id="4C15D5A9E50DD93304B3FA4645EDB2A0" authority="Griffiths & Herbert, 2013" authorityName="Griffiths & Herbert" authorityYear="2013" box="[766,972,1488,1507]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="36" pageNumber="36" phylum="Mollusca" rank="species" species="humbug" status="sp. nov.">
|
||
<emphasis id="B9617238E50DD93304B3FA4645EDB2A0" box="[766,972,1488,1507]" italics="true" pageId="36" pageNumber="36">Tropidophora humbug</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A252CF43E50DD9330599FA584229B2A0" box="[980,1032,1486,1507]" pageId="36" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
|
||
, st’n 12/06, shell diameter 28.3 mm, paratype, NMSA L7204/T2984; (C)
|
||
<taxonomicName id="4C15D5A9E50DD9330559FA7E47A8B150" authority="Fischer-Piette & Bedoucha, 1965" authorityName=": Fischer-Piette & Bedoucha" authorityYear="1965" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="36" pageNumber="36" phylum="Mollusca" rank="species" species="secunda">
|
||
<emphasis id="B9617238E50DD9330559FA7E45FEB2B8" box="[788,991,1512,1531]" italics="true" pageId="36" pageNumber="36">Tropidophora secunda</emphasis>
|
||
Fischer-Piette & Bedoucha, 1965
|
||
</taxonomicName>
|
||
, st’n 03/06, shell diameter approx. 19.0 mm; (D)
|
||
<taxonomicName id="4C15D5A9E50DD933050DF9964705B168" authority="Emberton, 1999" authorityName=": Emberton" authorityYear="1999" class="Gastropoda" family="Clavatoridae" genus="Clavator" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="36" phylum="Mollusca" rank="species" species="griffithsjonesi">
|
||
<emphasis id="B9617238E50DD933050DF9964233B150" box="[832,1042,1534,1555]" italics="true" pageId="36" pageNumber="36">Clavator griffithsjonesi</emphasis>
|
||
Emberton, 1999
|
||
</taxonomicName>
|
||
, st’n 12/06, shell height 94.8 mm, NMSA L7192; (E)
|
||
<taxonomicName id="4C15D5A9E50DD933054DF98E426AB168" authority="Fischer-Piette, 1952" authorityName=": Fischer-Piette" authorityYear="1952" box="[768,1099,1558,1579]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="36" phylum="Mollusca" rank="species" species="milloti">
|
||
<emphasis id="B9617238E50DD933054DF98E45B0B168" box="[768,913,1560,1579]" italics="true" pageId="36" pageNumber="36">Ampelita milloti</emphasis>
|
||
Fischer-Piette, 1952
|
||
</taxonomicName>
|
||
, st’n 07/09, shell diameter 19.2 mm, NMSA L8502; (F)
|
||
<taxonomicName id="4C15D5A9E50DD933049CF9A64253B100" authority="Fischer-Piette, 1952" authorityName=": Fischer-Piette" authorityYear="1952" box="[721,1138,1582,1603]" class="Gastropoda" family="Acavidae" genus="Ampelita" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="36" phylum="Mollusca" rank="species" species="namerokoensis">
|
||
<emphasis id="B9617238E50DD933049CF9A64590B100" box="[721,945,1584,1603]" italics="true" pageId="36" pageNumber="36">Ampelita namerokoensis</emphasis>
|
||
Fischer-Piette, 1952
|
||
</taxonomicName>
|
||
, st’n 12/06, shell diameter 20.3 mm, NMSA L8442; (G)
|
||
<taxonomicName id="4C15D5A9E50DD93304A2F9DE4253B118" authority="Emberton, 2002" authorityName=": Emberton" authorityYear="2002" box="[751,1138,1606,1627]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="36" phylum="Mollusca" rank="species" species="griffithshauchleri">
|
||
<emphasis id="B9617238E50DD93304A2F9DE45F9B118" box="[751,984,1606,1627]" italics="true" pageId="36" pageNumber="36">Kalidos griffithshauchleri</emphasis>
|
||
Emberton, 2002
|
||
</taxonomicName>
|
||
, st’n 12/06, shell diameter approx. 60 mm; (H)
|
||
<taxonomicName id="4C15D5A9E50DD93304D1F9F6456FB130" authority="Griffiths & Herbert, 2013" authorityName="Griffiths & Herbert" authorityYear="2013" box="[668,846,1632,1651]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="36" pageNumber="36" phylum="Mollusca" rank="species" species="maryannae" status="sp. nov.">
|
||
<emphasis id="B9617238E50DD93304D1F9F6456FB130" box="[668,846,1632,1651]" italics="true" pageId="36" pageNumber="36">Kalidos maryannae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A252CF43E50DD9330519F9C845A7B130" box="[852,902,1630,1651]" pageId="36" pageNumber="36" rank="species">sp. n.</taxonomicNameLabel>
|
||
, st’n 12/06, shell diameter 22.5 mm, paratype, NMSA L7193/T2944.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF6AFEA2E50CD93206DDFA704424B1C8" ID-DOI="http://doi.org/10.5281/zenodo.7670163" ID-Zenodo-Dep="7670163" httpUri="https://zenodo.org/record/7670163/files/figure.png" pageId="37" pageNumber="37" startId="37.[144,176,1510,1531]" targetBox="[157,1095,235,1476]" targetPageId="37">
|
||
<paragraph id="8BAAAE2AE50CD93206DDFA704424B1C8" blockId="37.[144,1104,1510,1675]" pageId="37" pageNumber="37">
|
||
Fig. 26. Species illustrated for comparative purposes: (A–C)
|
||
<emphasis id="B9617238E50CD93204EFFA7E455BB2B8" box="[674,890,1512,1531]" italics="true" pageId="37" pageNumber="37">
|
||
<taxonomicName id="4C15D5A9E50CD93204EFFA7E4556B2B8" baseAuthorityName="Grateloup" baseAuthorityYear="1840" box="[674,887,1512,1531]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="37" phylum="Mollusca" rank="species" species="moulinsii">Tropidophora moulinsii</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
holotype of
|
||
<taxonomicName id="4C15D5A9E50CD93205ABFA7E47EFB150" authority="Grateloup, 1840" authorityName="Grateloup" authorityYear="1840" class="Gastropoda" family="Pomatiidae" genus="Cyclostoma" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="37" phylum="Mollusca" rank="species" species="moulinsii">
|
||
<emphasis id="B9617238E50CD93205ABFA7E4706B150" italics="true" pageId="37" pageNumber="37">Cyclostoma moulinsii</emphasis>
|
||
Grateloup, 1840
|
||
</taxonomicName>
|
||
, ‘Madagascar’, max. diameter 33.1 mm, NHMUK 1907.11.22.12; (D–F)
|
||
<taxonomicName id="4C15D5A9E50CD9320743F98E44B9B168" authority="Fischer-Piette, 1949" authorityName="Fischer-Piette" authorityYear="1949" box="[270,664,1558,1579]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="37" phylum="Mollusca" rank="species" species="thesauri">
|
||
<emphasis id="B9617238E50CD9320743F98E47FAB168" box="[270,475,1560,1579]" italics="true" pageId="37" pageNumber="37">Tropidophora thesauri</emphasis>
|
||
Fischer-Piette, 1949
|
||
</taxonomicName>
|
||
, Orangea, Diego Suarez [= Antsiranana] district, max. diameter 38.0 mm, NMSA L8438; (G–I)
|
||
<taxonomicName id="4C15D5A9E50CD93204C8F9A64543B100" baseAuthorityName="Adams & Reeve" baseAuthorityYear="1850" box="[645,866,1584,1603]" class="Gastropoda" family="Pomatiidae" genus="Tropidophora" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="37" phylum="Mollusca" rank="species" species="reticulata">
|
||
<emphasis id="B9617238E50CD93204C8F9A64543B100" box="[645,866,1584,1603]" italics="true" pageId="37" pageNumber="37">Tropidophora reticulata</emphasis>
|
||
</taxonomicName>
|
||
, one of three syntypes of
|
||
<taxonomicName id="4C15D5A9E50CD932069DF9DE44ABB118" authority="(Adams & Reeve, 1850)" baseAuthorityName="Adams & Reeve" baseAuthorityYear="1850" box="[208,650,1606,1627]" class="Gastropoda" family="Pomatiidae" genus="Cyclostoma" kingdom="Animalia" order="Littorinimorpha" pageId="37" pageNumber="37" phylum="Mollusca" rank="species" species="reticulatum">
|
||
<emphasis id="B9617238E50CD932069DF9DE4789B118" box="[208,424,1608,1627]" italics="true" pageId="37" pageNumber="37">Cyclostoma reticulatum</emphasis>
|
||
(Adams & Reeve, 1850)
|
||
</taxonomicName>
|
||
, ‘Madagascar’, max. diameter 15.1 mm, NHMUK 1874.12.11.231; (J–L)
|
||
<taxonomicName id="4C15D5A9E50CD93207ECF9F64438B130" authorityName="Odhner" authorityYear="1919" box="[417,537,1632,1651]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="37" phylum="Mollusca" rank="species" species="hova">
|
||
<emphasis id="B9617238E50CD93207ECF9F64438B130" box="[417,537,1632,1651]" italics="true" pageId="37" pageNumber="37">Kalidos hova</emphasis>
|
||
</taxonomicName>
|
||
, syntype of
|
||
<taxonomicName id="4C15D5A9E50CD93204C4F9F645A0B130" authority="Odhner, 1919" authorityName="Odhner" authorityYear="1919" box="[649,897,1630,1651]" class="Gastropoda" family="Ariophantidae" genus="Nanina" kingdom="Animalia" order="Stylommatophora" pageId="37" pageNumber="37" phylum="Mollusca" rank="species" species="hova">
|
||
<emphasis id="B9617238E50CD93204C4F9F644DFB130" box="[649,766,1632,1651]" italics="true" pageId="37" pageNumber="37">Nanina hova</emphasis>
|
||
Odhner, 1919
|
||
</taxonomicName>
|
||
, Catsèpe, Madagascar, max. diameter 24.4 mm (SMNH).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAAAE2AE50FD931069DFF58476FB546" blockId="38.[181,1142,203,869]" pageId="38" pageNumber="38">
|
||
<taxonomicName id="4C15D5A9E50FD931069DFF58474CB7A5" baseAuthorityName="Angas" baseAuthorityYear="1877" box="[208,365,206,230]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="38" phylum="Mollusca" rank="species" species="ekongensis">
|
||
<emphasis id="B9617238E50FD931069DFF58474CB7A5" box="[208,365,206,230]" italics="true" pageId="38" pageNumber="38">K. ekongensis</emphasis>
|
||
</taxonomicName>
|
||
is more difficult to assess since the type material is now missing. A
|
||
<typeStatus id="54AE1088E50FD93106F8FF7A4734B645" box="[181,277,236,262]" pageId="38" pageNumber="38" type="paratype">paratype</typeStatus>
|
||
reportedly in the NHMUK (Fischer-Piette
|
||
<emphasis id="B9617238E50FD931054CFF784515B645" box="[769,820,238,262]" italics="true" pageId="38" pageNumber="38">et al</emphasis>
|
||
. 1966) is not in fact present there (Ablett pers. comm.
|
||
<date id="FFAB88EAE50FD9310790FE9A4474B666" box="[477,597,267,294]" pageId="38" pageNumber="38" value="2011-11">Nov. 2011</date>
|
||
). The original description is not detailed, but the figure shows a shell closely resembling the present material. However, Fischer-Piette
|
||
<emphasis id="B9617238E50FD93106F8FED846C7B625" box="[181,230,334,358]" italics="true" pageId="38" pageNumber="38">et al</emphasis>
|
||
. (1966) noted, after examination of the NHMUK
|
||
<typeStatus id="54AE1088E50FD9310559FEDA4556B625" box="[788,887,332,358]" pageId="38" pageNumber="38" type="paratype">paratype</typeStatus>
|
||
, that the shell was thin and very glossy. Again this is not a description that would apply to the present material. Fischer-Piette
|
||
<emphasis id="B9617238E50FD9310715FE1847ABB6E5" box="[344,394,398,422]" italics="true" pageId="38" pageNumber="38">et al</emphasis>
|
||
. (1966, 1994) interpreted
|
||
<taxonomicName id="4C15D5A9E50FD93104FEFE18456EB6E5" baseAuthorityName="Angas" baseAuthorityYear="1877" box="[691,847,398,422]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="38" phylum="Mollusca" rank="species" species="ekongensis">
|
||
<emphasis id="B9617238E50FD93104FEFE18456EB6E5" box="[691,847,398,422]" italics="true" pageId="38" pageNumber="38">K. ekongensis</emphasis>
|
||
</taxonomicName>
|
||
very loosely, including in it a variety of different forms from widely disjunct localities with widely differing habitats. Almost certainly these are not conspecific and their true identity requires further investigation.
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE50FD931069DFD9A47FAB5E6" blockId="38.[181,1142,203,869]" pageId="38" pageNumber="38">
|
||
In addition to having relatively coarse spiral sculpture on the last adult whorl, rendering it lustreless,
|
||
<taxonomicName id="4C15D5A9E50FD93107C6FDB84409B505" authorityName="Griffiths & Herbert" authorityYear="2013" box="[395,552,558,582]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="38" phylum="Mollusca" rank="species" species="maryannae">
|
||
<emphasis id="B9617238E50FD93107C6FDB84409B505" box="[395,552,558,582]" italics="true" pageId="38" pageNumber="38">K. maryannae</emphasis>
|
||
</taxonomicName>
|
||
also differs from
|
||
<taxonomicName id="4C15D5A9E50FD93104A5FDB845A3B505" baseAuthorityName="Angas" baseAuthorityYear="1877" box="[744,898,558,582]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="38" phylum="Mollusca" rank="species" species="ekongensis">
|
||
<emphasis id="B9617238E50FD93104A5FDB845A3B505" box="[744,898,558,582]" italics="true" pageId="38" pageNumber="38">K. ekongensis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C15D5A9E50FD93105F9FDB84228B505" authorityName="Odhner" authorityYear="1919" box="[948,1033,558,582]" class="Gastropoda" family="Ariophantidae" genus="Nanina" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="38" phylum="Mollusca" rank="species" species="hova">
|
||
<emphasis id="B9617238E50FD93105F9FDB84228B505" box="[948,1033,558,582]" italics="true" pageId="38" pageNumber="38">K. hova</emphasis>
|
||
</taxonomicName>
|
||
in having more rapidly expanding whorls. Another distinctive feature of
|
||
<taxonomicName id="4C15D5A9E50FD93105C8FDD84207B525" authorityName="Griffiths & Herbert" authorityYear="2013" box="[901,1062,590,614]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="38" phylum="Mollusca" rank="species" species="maryannae">
|
||
<emphasis id="B9617238E50FD93105C8FDD84207B525" box="[901,1062,590,614]" italics="true" pageId="38" pageNumber="38">K. maryannae</emphasis>
|
||
</taxonomicName>
|
||
is that the colour bands are evident only on the penultimate and final whorls, the spire whorls being of a uniform colour.
|
||
</paragraph>
|
||
<paragraph id="8BAAAE2AE50FD931069DFD3D45B1B426" blockId="38.[181,1142,203,869]" pageId="38" pageNumber="38">
|
||
The genital anatomy of
|
||
<taxonomicName id="4C15D5A9E50FD9310790FD38445CB585" authorityName="Griffiths & Herbert" authorityYear="2013" box="[477,637,686,710]" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="38" phylum="Mollusca" rank="species" species="maryannae">
|
||
<emphasis id="B9617238E50FD9310790FD38445CB585" box="[477,637,686,710]" italics="true" pageId="38" pageNumber="38">K. maryannae</emphasis>
|
||
</taxonomicName>
|
||
conforms to the common pattern evinced by
|
||
<taxonomicName id="4C15D5A9E50FD93106F8FD584742B5A6" box="[181,355,715,742]" pageId="38" pageNumber="38">
|
||
<emphasis id="B9617238E50FD93106F8FD58472DB5A5" box="[181,268,718,742]" italics="true" pageId="38" pageNumber="38">Kalidos</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(Fischer-Piette
|
||
<emphasis id="B9617238E50FD931045FFD584460B5A5" box="[530,577,718,742]" italics="true" pageId="38" pageNumber="38">et al</emphasis>
|
||
. 1975; Schileyko 2002) and is fully consistent with referral of the species to this genus. In this regard it is perhaps most similar to
|
||
<taxonomicName id="4C15D5A9E50FD9310253FD784790B466" authority="(Ancey, 1902)" baseAuthorityName="Ancey" baseAuthorityYear="1902" class="Gastropoda" family="Helicarionidae" genus="Kalidos" kingdom="Animalia" order="Stylommatophora" pageId="38" pageNumber="38" phylum="Mollusca" rank="species" species="oleatus">
|
||
<emphasis id="B9617238E50FD9310253FD784727B465" italics="true" pageId="38" pageNumber="38">Kalidos oleatus</emphasis>
|
||
(Ancey, 1902)
|
||
</taxonomicName>
|
||
as figured by Schileyko (2002), but in the present species the flagellum is shorter and more tightly coiled and it is not connected to the penis by muscle bands, and the bursa copulatrix duct is shorter and much broader.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |