treatments-xml/data/03/A5/87/03A587FA7828A030FCFBBB6D31555375.xml

<document id="080C413047DEF7733E5C303D3BCBEB0A" ID-DOI="10.5281/zenodo.5460634" ID-ISSN="2345-7600" ID-Zenodo-Dep="5460634" ID-ZooBank="1E7214C5-24E8-4860-BF13-203FD93A2C19" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630239335276" checkinUser="valdenar" docAuthor="Pholyotha, Arthit, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2018" docId="03A587FA7828A030FCFBBB6D31555375" docLanguage="en" docName="66rbz763-781.pdf" docOrigin="Raffles Bulletin of Zoology 66" docStyle="DocumentStyle:F2D34FF25DBF9998BB8DF6F24A5B097A.5:RafflesBZool.2014-2018.journal_article" docStyleId="F2D34FF25DBF9998BB8DF6F24A5B097A" docStyleName="RafflesBZool.2014-2018.journal_article" docStyleVersion="5" docTitle="Macrochlamys tanymentula Pholyotha &amp; Sutcharit &amp; Panha 2018, new species" docType="treatment" docUuid="4849420C-A790-4CB0-9BC2-61AC66C8E595" docUuidSource="ZooBank" docVersion="3" lastPageNumber="774" masterDocId="FF9CFF827821A03BFFC7BF1034285476" masterDocTitle="The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae)" masterLastPageNumber="781" masterPageNumber="763" pageNumber="772" updateTime="1699195389588" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="339ED083ACB60E870703F3732A8E0A4F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6CB80BFA6B457E6FB64ED3F1DF79B89C">
<mods:title id="97174109E9800744BDAFBE4B95E522C3">The land snail genus Macrochlamys Gray, 1847 from Thailand, with descriptions of five new species (Pulmonata: Ariophantidae)</mods:title>
</mods:titleInfo>
<mods:name id="B28C2E930625937877BE13BD518A1A5B" type="personal">
<mods:role id="6D1351C45C62E8FC56FFE5102625FEC5">
<mods:roleTerm id="C32BCC2FE2FECB18A4979ED61FD35FE1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F825635C68481585BA5A384BD81CA916">Pholyotha, Arthit</mods:namePart>
</mods:name>
<mods:name id="ED702E3C8B5E2780F4BA0E6A96C9958F" type="personal">
<mods:role id="0507E1399A964BD9D27E750ED8F8A8D6">
<mods:roleTerm id="48924DCA2E09A948D952B0303C1C691E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="83E396CB6C96AC83DF94778C15C1A2ED">Sutcharit, Chirasak</mods:namePart>
</mods:name>
<mods:name id="5ED4CEFBA8DFCBD34CEF5F487453900C" type="personal">
<mods:role id="47B5335F6B0FFA38BC9F8070E04855C8">
<mods:roleTerm id="2CD0F7770B6E1E502D808450F851F0FC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="98B6A572B913678A9536C603EE8D5084">Panha, Somsak</mods:namePart>
</mods:name>
<mods:typeOfResource id="C24E68E327396A693D1795CBFFD410E9">text</mods:typeOfResource>
<mods:relatedItem id="86E7368E230282CAC89BCA053994067F" type="host">
<mods:titleInfo id="11BBA85F02B6198E2688B1C145E38549">
<mods:title id="29AA90A336C01101F71D2D5C80B02045">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="BB24BC0243FEC9E822DC29A84C3A6C0F">
<mods:date id="396F15EC1620E91976D8FA7876442AE6">2018</mods:date>
<mods:detail id="DFD76760E83043F21C911995829D2A93" type="pubDate">
<mods:number id="B0753453F8812025ACC19F87DAF606B0">2018-12-03</mods:number>
</mods:detail>
<mods:detail id="4B88D011DC8E41AC6DB663A73728F45B" type="volume">
<mods:number id="5FA19D5D90A217CB070A11640C54B64B">66</mods:number>
</mods:detail>
<mods:extent id="8013933A4E41B88095D9A8BE748FDACF" unit="page">
<mods:start id="A4688B92CBA63FD395D6D04FCBF46F3B">763</mods:start>
<mods:end id="70D09523C5E196C184C51774335866C3">781</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="ACCC7E769E539BDF7F49C76743FE0CC4">journal article</mods:classification>
<mods:identifier id="FA7D13C463A625D257162A5A82FCFCFA" type="DOI">10.5281/zenodo.5460634</mods:identifier>
<mods:identifier id="F66E15FAD5795E41CBE03FE98E334634" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="64B4C55ED15181EDA1A6AE74900C8E74" type="Zenodo-Dep">5460634</mods:identifier>
<mods:identifier id="F5B4F6731E9B28F7482C61314261498F" type="ZooBank">1E7214C5-24E8-4860-BF13-203FD93A2C19</mods:identifier>
</mods:mods>
<treatment id="03A587FA7828A030FCFBBB6D31555375" LSID="urn:lsid:zoobank.org:act:4849420C-A790-4CB0-9BC2-61AC66C8E595" httpUri="http://treatment.plazi.org/id/03A587FA7828A030FCFBBB6D31555375" lastPageId="11" lastPageNumber="774" pageId="9" pageNumber="772">
<subSubSection id="C31665677828A032FCFBBB6D30BC50C3" pageId="9" pageNumber="772" type="nomenclature">
<paragraph id="8BB336EC7828A032FCFBBB6D30BC50C3" blockId="9.[828,1436,1149,1237]" pageId="9" pageNumber="772">
<heading id="D0FB81807828A032FCFBBB6D30BC50C3" bold="true" centered="true" fontSize="10" level="2" pageId="9" pageNumber="772" reason="2">
<emphasis id="B978EAFE7828A032FCFBBB6D30BC50C3" bold="true" pageId="9" pageNumber="772">
<taxonomicName id="4C0C4D6F7828A032FCFBBB6D314850E3" ID-CoL="3X446" authority="Pholyotha &amp; Panha" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[828,1376,1149,1173]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="772" phylum="Mollusca" rank="species" species="tanymentula" status="sp. nov.">
<emphasis id="B978EAFE7828A032FCFBBB6D305A50E3" bold="true" box="[828,1138,1149,1173]" italics="true" pageId="9" pageNumber="772">Macrochlamys tanymentula</emphasis>
Pholyotha &amp; Panha
</taxonomicName>
, 
<taxonomicNameLabel id="A24B57857828A032FAA9BB6D30BC50C3" pageId="9" pageNumber="772" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31665677828A032FC7FBBAD310950A3" box="[952,1313,1213,1237]" pageId="9" pageNumber="772" type="description">
<paragraph id="8BB336EC7828A032FC7FBBAD310950A3" blockId="9.[828,1436,1149,1237]" box="[952,1313,1213,1237]" pageId="9" pageNumber="772">
(
<figureCitation id="13372A697828A032FC07BBAD303650A3" box="[960,1054,1213,1237]" captionStart="Fig" captionStartId="2.[135,170,1546,1568]" captionTargetBox="[136,1474,192,1529]" captionTargetId="figure-7@2.[136,1474,189,1531]" captionTargetPageId="2" captionText="Fig. 2. Living snails of Macrochlamys spp. from Thailand. A, Macrochlamys aurantia new species, paratype (CUMZ 7103), Thong Pha Phum, Kanchanaburi, shell width about 15 mm; B, M. coleus new species, paratype (CUMZ 7106), Sai Yok, Kanchanaburi, shell width about 15 mm; C,M. caverna new species, paratype (CUMZ 7109), Tha Wung, Lopburi, shell width about 17 mm; D, M. lemma new species, paratype (CUMZ 7116), Ban Mi, Lopburi, shell width about 15 mm; E, M. tanymentula new species, paratype (CUMZ 7119), Thong Pha Phum, Kanchanaburi, shell width about 10 mm; F, M. kelantanensis (CUMZ 7125), Lam Kaen, Phang-Nga, shell width about 20 mm." figureDoi="http://doi.org/10.5281/zenodo.5460640" httpUri="https://zenodo.org/record/5460640/files/figure.png" pageId="9" pageNumber="772">Figs. 2E</figureCitation>
, 
<figureCitation id="13372A697828A032FBECBBAD304650A3" box="[1067,1134,1213,1237]" captionStart="Fig" captionStartId="11.[113,148,1249,1271]" captionTargetBox="[113,1451,189,1234]" captionTargetId="figure-9@11.[113,1451,189,1234]" captionTargetPageId="11" captionText="Fig. 7. Shells of Macrochlamys lemma new species (A, B), and M. tanymentula new species (C, D). A, Macrochlamys lemma holotype (CUMZ 7114); B, paratype (CUMZ 7115); C, M. tanymentula holotype (CUMZ 7117); D, paratype (CUMZ 7118)." figureDoi="http://doi.org/10.5281/zenodo.5460652" httpUri="https://zenodo.org/record/5460652/files/figure.png" pageId="9" pageNumber="772">7C, D</figureCitation>
, 
<figureCitation id="13372A697828A032FBBBBBAD30E850A3" box="[1148,1216,1213,1237]" captionStart="Fig" captionStartId="14.[136,171,1853,1875]" captionTargetBox="[136,1474,285,1838]" captionTargetId="figure-7@14.[136,1474,285,1838]" captionTargetPageId="14" captionText="Fig. 9. Genitalia of Macrochlamys tanymentula new species (A, B), and M. kelantanensis (C, D). A, B, Macrochlamys tanymentula paratype (CUMZ 7119): A, whole reproductive system; B, internal structure of penis with penial verge. C, D, Macrochlamys kelantanensis (CUMZ 7125): C, whole reproductive system; D, internal structure of penis with penial verge. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Arrow indicates position of the penial verge." figureDoi="http://doi.org/10.5281/zenodo.5460656" httpUri="https://zenodo.org/record/5460656/files/figure.png" pageId="9" pageNumber="772">9A, B</figureCitation>
, 
<figureCitation id="13372A697828A032FB0ABBAD313150A3" box="[1229,1305,1213,1237]" captionStart="Fig" captionStartId="15.[113,148,1222,1244]" captionTargetBox="[113,1452,189,1207]" captionTargetId="figure-347@15.[113,1452,189,1207]" captionTargetPageId="15" captionText="Fig. 10. SEM images of the radula of Macrochlamys lemma new species (A–C) (paratype; CUMZ 7116), M. tanymentula new species (D–F) (paratype; CUMZ 7119), and M. kelantanensis Möllendorff, 1902 (G–I) (CUMZ 7125). Central tooth (indicated by ‘C’) with lateral teeth (A, D, G); lateral teeth with tricuspid marginal teeth transition (B, E, H); bicuspid marginal teeth (C, F, I)." figureDoi="http://doi.org/10.5281/zenodo.5460660" httpUri="https://zenodo.org/record/5460660/files/figure.png" pageId="9" pageNumber="772">10D–F</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31665677828A032FCEABBED30FB5263" pageId="9" pageNumber="772" type="materials_examined">
<paragraph id="8BB336EC7828A032FCEABBED30FB5263" blockId="9.[811,1452,1277,1557]" pageId="9" pageNumber="772">
<emphasis id="B978EAFE7828A032FCEABBED30205163" bold="true" box="[813,1032,1277,1301]" pageId="9" pageNumber="772">Material examined.</emphasis>
<materialsCitation id="3B643CB17828A032FBCBBBED37B451C3" collectionCode="CUMZ" country="Thailand" county="Thong Pha Phum District" latitude="14.737139" location="Khao Lok" longLatPrecision="1" longitude="98.64186" municipality="Tha Kha-nun Village" pageId="9" pageNumber="772" specimenCode="CUMZ 7117" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="holotype">
<typeStatus id="54B7884E7828A032FBCBBBED30465163" box="[1036,1134,1277,1301]" pageId="9" pageNumber="772" type="holotype">Holotype</typeStatus>
(
<figureCitation id="13372A697828A032FBBBBBED30E65163" box="[1148,1230,1277,1301]" captionStart="Fig" captionStartId="11.[113,148,1249,1271]" captionTargetBox="[113,1451,189,1234]" captionTargetId="figure-9@11.[113,1451,189,1234]" captionTargetPageId="11" captionText="Fig. 7. Shells of Macrochlamys lemma new species (A, B), and M. tanymentula new species (C, D). A, Macrochlamys lemma holotype (CUMZ 7114); B, paratype (CUMZ 7115); C, M. tanymentula holotype (CUMZ 7117); D, paratype (CUMZ 7118)." figureDoi="http://doi.org/10.5281/zenodo.5460652" httpUri="https://zenodo.org/record/5460652/files/figure.png" pageId="9" pageNumber="772">Fig. 7C</figureCitation>
; shell height 
<quantity id="4CF49B097828A032FA92BBED318F5163" box="[1365,1447,1277,1301]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="9" pageNumber="772" unit="mm" value="5.4">5.4 mm</quantity>
, shell width 
<quantity id="4CF49B097828A032FC62BA0D302D5143" box="[933,1029,1309,1333]" metricMagnitude="-2" metricUnit="m" metricValue="1.09" pageId="9" pageNumber="772" unit="mm" value="10.9">10.9 mm</quantity>
, aperture height 
<quantity id="4CF49B097828A032FB73BA0D312F5143" box="[1204,1287,1309,1333]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="9" pageNumber="772" unit="mm" value="4.2">4.2 mm</quantity>
, aperture width 
<quantity id="4CF49B097828A032FCECBA2D37AA5123" box="[811,898,1341,1365]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="9" pageNumber="772" unit="mm" value="5.3">5.3 mm</quantity>
, 5½ whorls) (
<specimenCode id="DBAA9E977828A032FBDCBA2D30845123" box="[1051,1196,1341,1365]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="9" pageNumber="772">CUMZ 7117</specimenCode>
), limestone outcrops at 
<location id="8ED360377828A032FCEABA4D37B35103" LSID="urn:lsid:plazi:treatment:03A587FA7828A030FCFBBB6D31555375:8ED360377828A032FCEABA4D37B35103" box="[813,923,1373,1397]" country="Thailand" county="Thong Pha Phum District" latitude="14.737139" longLatPrecision="1" longitude="98.64186" municipality="Tha Kha-nun Village" name="Khao Lok" pageId="9" pageNumber="772" stateProvince="Kanchanaburi">Khao Lok</location>
, 
<collectingMunicipality id="6BD7AC967828A032FC62BA4D30A25103" box="[933,1162,1373,1397]" pageId="9" pageNumber="772">Tha Kha-nun Village</collectingMunicipality>
, 
<collectingCounty id="62D24E607828A032FB52BA4D31805103" box="[1173,1448,1373,1397]" pageId="9" pageNumber="772">Thong Pha Phum District</collectingCounty>
, 
<collectingRegion id="49C8F80E7828A032FCEABA6D306951E3" box="[813,1089,1405,1429]" country="Thailand" name="Kanchanaburi" pageId="9" pageNumber="772">Kanchanaburi Province</collectingRegion>
, 
<collectingCountry id="F31B767C7828A032FB97BA6D309151E3" box="[1104,1209,1405,1429]" name="Thailand" pageId="9" pageNumber="772">Thailand</collectingCountry>
, 
<geoCoordinate id="EE38502B7828A032FB0EBA6D315C51E3" box="[1225,1396,1404,1429]" degrees="14" direction="north" minutes="44" orientation="latitude" pageId="9" pageNumber="772" precision="1" seconds="13.7" value="14.737139">14° 44′13.7″N</geoCoordinate>
, 
<geoCoordinate id="EE38502B7828A032FA43BA6D37B451C3" degrees="98" direction="east" minutes="38" orientation="longitude" pageId="9" pageNumber="772" precision="1" seconds="30.7" value="98.64186">98° 38′30.7″E</geoCoordinate>
</materialsCitation>
. 
<materialsCitation id="3B643CB17828A032FC6DBA8E30E75263" collectionCode="CUMZ" country="Thailand" county="Thong Pha Phum District" latitude="14.737139" location="Khao Lok" longLatPrecision="1" longitude="98.64186" municipality="Tha Kha-nun Village" pageId="9" pageNumber="772" specimenCode="CUMZ 7118, CUMZ 7119" specimenCount="1" stateProvince="Kanchanaburi" typeStatus="paratype">
<typeStatus id="54B7884E7828A032FC6DBA8E303F51C0" box="[938,1047,1438,1462]" pageId="9" pageNumber="772" type="paratype">Paratypes</typeStatus>
: one shell (
<figureCitation id="13372A697828A032FB5EBA8D30DB51C3" box="[1177,1267,1437,1461]" captionStart="Fig" captionStartId="11.[113,148,1249,1271]" captionTargetBox="[113,1451,189,1234]" captionTargetId="figure-9@11.[113,1451,189,1234]" captionTargetPageId="11" captionText="Fig. 7. Shells of Macrochlamys lemma new species (A, B), and M. tanymentula new species (C, D). A, Macrochlamys lemma holotype (CUMZ 7114); B, paratype (CUMZ 7115); C, M. tanymentula holotype (CUMZ 7117); D, paratype (CUMZ 7118)." figureDoi="http://doi.org/10.5281/zenodo.5460652" httpUri="https://zenodo.org/record/5460652/files/figure.png" pageId="9" pageNumber="772">Fig. 7D</figureCitation>
; shell height 
<quantity id="4CF49B097828A032FA4CBA8D377151A3" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="772" unit="mm" value="5.5">5.5 mm</quantity>
, shell width 
<quantity id="4CF49B097828A032FC26BAAD306A51A3" box="[993,1090,1469,1493]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="9" pageNumber="772" unit="mm" value="11.1">11.1 mm</quantity>
, aperture height 4.0 mm, aperture width 
<quantity id="4CF49B097828A032FCB5BACD37E05183" box="[882,968,1501,1525]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="9" pageNumber="772" unit="mm" value="5.2">5.2 mm</quantity>
, 5¾ whorls) (
<specimenCode id="DBAA9E977828A032FBA6BACD30D95183" box="[1121,1265,1501,1525]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="9" pageNumber="772">CUMZ 7118</specimenCode>
), 
<specimenCount id="9D0AFD657828A032FAC1BACD31845183" box="[1286,1452,1501,1525]" pageId="9" pageNumber="772" type="generic">nine specimens</specimenCount>
(
<specimenCode id="DBAA9E977828A032FCF1BAED37EE5263" box="[822,966,1533,1557]" collectionCode="CUMZ" country="0" httpUri="http://grbio.org/cool/da58-p0xp" name="Cameroon University, Museum of Zoology" pageId="9" pageNumber="772">CUMZ 7119</specimenCode>
), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C31665677828A032FCEAB92D37DF52A0" pageId="9" pageNumber="772" type="etymology">
<paragraph id="8BB336EC7828A032FCEAB92D37DF52A0" blockId="9.[813,1452,1597,1750]" pageId="9" pageNumber="772">
<emphasis id="B978EAFE7828A032FCEAB92D379D5223" bold="true" box="[813,949,1597,1621]" pageId="9" pageNumber="772">Etymology.</emphasis>
The specific name “ 
<taxonomicName id="4C0C4D6F7828A032FB69B92D31115223" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[1198,1337,1597,1621]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="772" phylum="Mollusca" rank="species" species="tanymentula">
<emphasis id="B978EAFE7828A032FB69B92D31115223" box="[1198,1337,1597,1621]" italics="true" pageId="9" pageNumber="772">tanymentula</emphasis>
</taxonomicName>
” is made up of the Greek word “ 
<emphasis id="B978EAFE7828A032FBFDB94E30415203" box="[1082,1129,1630,1653]" italics="true" pageId="9" pageNumber="772">tany</emphasis>
” meaning “long or slender” and the Latin “ 
<emphasis id="B978EAFE7828A032FC1DB96D301A52E3" box="[986,1074,1661,1685]" italics="true" pageId="9" pageNumber="772">mentula</emphasis>
” meaning “male organ including epiphallus” to refer to the very long and slender epiphallus of the male organ.
</paragraph>
</subSubSection>
<subSubSection id="C31665677828A031FCEAB9ED305F5288" lastPageId="10" lastPageNumber="773" pageId="9" pageNumber="772" type="description">
<paragraph id="8BB336EC7828A031FCEAB9ED35185108" blockId="9.[812,1454,1789,2101]" lastBlockId="10.[136,775,1350,1406]" lastPageId="10" lastPageNumber="773" pageId="9" pageNumber="772">
<emphasis id="B978EAFE7828A032FCEAB9ED37905363" bold="true" box="[813,952,1789,1813]" pageId="9" pageNumber="772">Description.</emphasis>
Shell medium-sized (
<figureCitation id="13372A697828A032FB60B9ED31085363" box="[1191,1312,1789,1813]" captionStart="Fig" captionStartId="11.[113,148,1249,1271]" captionTargetBox="[113,1451,189,1234]" captionTargetId="figure-9@11.[113,1451,189,1234]" captionTargetPageId="11" captionText="Fig. 7. Shells of Macrochlamys lemma new species (A, B), and M. tanymentula new species (C, D). A, Macrochlamys lemma holotype (CUMZ 7114); B, paratype (CUMZ 7115); C, M. tanymentula holotype (CUMZ 7117); D, paratype (CUMZ 7118)." figureDoi="http://doi.org/10.5281/zenodo.5460652" httpUri="https://zenodo.org/record/5460652/files/figure.png" pageId="9" pageNumber="772">Fig. 7C, D</figureCitation>
; shell height up to 
<quantity id="4CF49B097828A032FCADB80D37965343" box="[874,958,1821,1845]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="772" unit="mm" value="5.5">5.5 mm</quantity>
; shell width up to 
<quantity id="4CF49B097828A032FBBBB80D30F65343" box="[1148,1246,1821,1845]" metricMagnitude="-2" metricUnit="m" metricValue="1.1099999999999999" pageId="9" pageNumber="772" unit="mm" value="11.1">11.1 mm</quantity>
), dextral, globosely depressed, rather thin, shiny, translucent and light brown in colour. Shell surface smooth with thin growth lines. Embryonic shell smooth, with about 2½ whorls. Whorls 5–6, regularly increasing, with weakly impressed suture. Spire low conical to a little convex. Apex slightly raised with spire angle of about 151–154°. Last whorl large with rounded periphery. Aperture slightly large, ovately lunate in shape, obliquely open with simple lip. Columellar margin gradually reflected near umbilicus. Umbilicus narrowly open and very small.
</paragraph>
<caption id="DF736664782BA031FF4FBBAF308C5178" ID-DOI="http://doi.org/10.5281/zenodo.5460650" ID-Zenodo-Dep="5460650" httpUri="https://zenodo.org/record/5460650/files/figure.png" pageId="10" pageNumber="773" startId="10.[136,171,1215,1237]" targetBox="[136,1474,189,1200]" targetPageId="10">
<paragraph id="8BB336EC782BA031FF4FBBAF308C5178" blockId="10.[136,1477,1215,1295]" pageId="10" pageNumber="773">
Fig. 6. SEM images of the radula of 
<taxonomicName id="4C0C4D6F782BA031FE30BBAF36C850A3" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[503,736,1215,1237]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="773" phylum="Mollusca" rank="species" species="aurantia" status="sp. nov.">
<emphasis id="B978EAFE782BA031FE30BBAF36C850A3" box="[503,736,1215,1237]" italics="true" pageId="10" pageNumber="773">Macrochlamys aurantia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782BA031FD2FBBD0377450A3" box="[744,860,1215,1237]" pageId="10" pageNumber="773" rank="species">new species</taxonomicNameLabel>
(A–C) (paratype; CUMZ 7103), 
<taxonomicName id="4C0C4D6F782BA031FB61BBD0312C50A2" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[1190,1284,1215,1237]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="773" phylum="Mollusca" rank="species" species="coleus" status="sp. nov.">
<emphasis id="B978EAFE782BA031FB61BBD0312C50A2" box="[1190,1284,1215,1237]" italics="true" pageId="10" pageNumber="773">M. coleus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782BA031FACABBD031A950A3" box="[1293,1409,1215,1237]" pageId="10" pageNumber="773" rank="species">new species</taxonomicNameLabel>
(D–F) (paratype; CUMZ 7106), and 
<taxonomicName id="4C0C4D6F782BA031FE68BBCD36365084" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[431,542,1245,1266]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="773" phylum="Mollusca" rank="species" species="caverna" status="sp. nov.">
<emphasis id="B978EAFE782BA031FE68BBCD36365084" box="[431,542,1245,1266]" italics="true" pageId="10" pageNumber="773">M. caverna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782BA031FDEFBBCC36B55084" box="[552,669,1244,1266]" pageId="10" pageNumber="773" rank="species">new species</taxonomicNameLabel>
(G–I) (paratype; CUMZ 7109). Central tooth (indicated by ‘C’) with lateral teeth (A, D, G); lateral teeth with tricuspid marginal teeth transition (B, E, H); bicuspid marginal teeth (C, F, I).
</paragraph>
</caption>
<paragraph id="8BB336EC782BA031FF4FBAB6359152C8" blockId="10.[135,777,1446,1726]" pageId="10" pageNumber="773">
<emphasis id="B978EAFE782BA031FF4FBAB6351B51C8" box="[136,307,1446,1470]" italics="true" pageId="10" pageNumber="773">Genital organs</emphasis>
. Atrium (at) very short. Penis (p) short, enlarged cylindrical in shape, and about two times greater than epiphallus diameter. Epiphallus (e) small, a slender tube about five times longer than penis. Epiphallic caecum (ec) coiled about one circle and located at middle of epiphallus. Penial retractor muscle (prm) thin. Flagellum (fl) very long, slender, and about half of epiphallic length. Vas deference (vd) a thin tube connected between free oviduct and distal end of epiphallus (
<figureCitation id="13372A69782BA031FE93B9B6358452C8" box="[340,428,1702,1726]" captionStart="Fig" captionStartId="14.[136,171,1853,1875]" captionTargetBox="[136,1474,285,1838]" captionTargetId="figure-7@14.[136,1474,285,1838]" captionTargetPageId="14" captionText="Fig. 9. Genitalia of Macrochlamys tanymentula new species (A, B), and M. kelantanensis (C, D). A, B, Macrochlamys tanymentula paratype (CUMZ 7119): A, whole reproductive system; B, internal structure of penis with penial verge. C, D, Macrochlamys kelantanensis (CUMZ 7125): C, whole reproductive system; D, internal structure of penis with penial verge. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Arrow indicates position of the penial verge." figureDoi="http://doi.org/10.5281/zenodo.5460656" httpUri="https://zenodo.org/record/5460656/files/figure.png" pageId="10" pageNumber="773">Fig. 9A</figureCitation>
).
</paragraph>
<paragraph id="8BB336EC782BA031FF4FB9F636965348" blockId="10.[135,776,1766,1854]" pageId="10" pageNumber="773">
Internal wall of penis with small wrinkled longitudinal penial pilasters (pp). Penial verge (pv) slightly small, cylindricalshaped and situated at distal end of penis (
<figureCitation id="13372A69782BA031FD9CB83636995348" box="[603,689,1830,1854]" captionStart="Fig" captionStartId="14.[136,171,1853,1875]" captionTargetBox="[136,1474,285,1838]" captionTargetId="figure-7@14.[136,1474,285,1838]" captionTargetPageId="14" captionText="Fig. 9. Genitalia of Macrochlamys tanymentula new species (A, B), and M. kelantanensis (C, D). A, B, Macrochlamys tanymentula paratype (CUMZ 7119): A, whole reproductive system; B, internal structure of penis with penial verge. C, D, Macrochlamys kelantanensis (CUMZ 7125): C, whole reproductive system; D, internal structure of penis with penial verge. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Arrow indicates position of the penial verge." figureDoi="http://doi.org/10.5281/zenodo.5460656" httpUri="https://zenodo.org/record/5460656/files/figure.png" pageId="10" pageNumber="773">Fig. 9B</figureCitation>
).
</paragraph>
<paragraph id="8BB336EC782BA031FF4FB87636BD5C48" blockId="10.[134,775,1894,2110]" pageId="10" pageNumber="773">
Vagina (v) very short. Dart apparatus (da) large and cylindrical in shape, situated at proximal end of vagina. Gametolytic sac (gs) long and bulbous. Gametolytic duct (gd) long and cylindrical in shape. Free oviduct (fo) a small tube, approximately same length of penis, with thick brownish tissue near proximal end. Oviduct (ov) large lobules, with prostate gland (pg) running alongside (
<figureCitation id="13372A69782BA031FDF6B73636A05C48" box="[561,648,2086,2110]" captionStart="Fig" captionStartId="14.[136,171,1853,1875]" captionTargetBox="[136,1474,285,1838]" captionTargetId="figure-7@14.[136,1474,285,1838]" captionTargetPageId="14" captionText="Fig. 9. Genitalia of Macrochlamys tanymentula new species (A, B), and M. kelantanensis (C, D). A, B, Macrochlamys tanymentula paratype (CUMZ 7119): A, whole reproductive system; B, internal structure of penis with penial verge. C, D, Macrochlamys kelantanensis (CUMZ 7125): C, whole reproductive system; D, internal structure of penis with penial verge. Abbreviations: at, atrium; da, dart apparatus; drm, dart retractor muscle; e, epiphallus; ec, epiphallic caecum; fl, flagellum; fo, free oviduct; gd, gametolytic duct; gs, gametolytic sac; hf, head filament; ov, oviduct; p, penis; pc, penial caecum; pg, prostate gland; pp, penial pilaster; prm, penial retractor muscle; pv, penial verge; ss, sperm sac; tf, tail filament; v, vagina and vd, vas deferens. Arrow indicates position of the penial verge." figureDoi="http://doi.org/10.5281/zenodo.5460656" httpUri="https://zenodo.org/record/5460656/files/figure.png" pageId="10" pageNumber="773">Fig. 9A</figureCitation>
).
</paragraph>
<paragraph id="8BB336EC782BA031FC84BA56300F5188" blockId="10.[835,1475,1350,1534]" pageId="10" pageNumber="773">
<emphasis id="B978EAFE782BA031FC84BA5637B85128" box="[835,912,1350,1374]" italics="true" pageId="10" pageNumber="773">Radula</emphasis>
. Each row consists of about 99 teeth with formula (50-(14-12)-1-(12-13)-48). Central tooth symmetrical tricuspid with long and slender mesocone. Lateral teeth asymmetrical tricuspid with deeply indented endocone and ectocone. Bicuspid marginal teeth start from tooth number 12–14 (
<figureCitation id="13372A69782BA031FC5EBAF630335188" box="[921,1051,1510,1534]" captionStart="Fig" captionStartId="15.[113,148,1222,1244]" captionTargetBox="[113,1452,189,1207]" captionTargetId="figure-347@15.[113,1452,189,1207]" captionTargetPageId="15" captionText="Fig. 10. SEM images of the radula of Macrochlamys lemma new species (A–C) (paratype; CUMZ 7116), M. tanymentula new species (D–F) (paratype; CUMZ 7119), and M. kelantanensis Möllendorff, 1902 (G–I) (CUMZ 7125). Central tooth (indicated by ‘C’) with lateral teeth (A, D, G); lateral teeth with tricuspid marginal teeth transition (B, E, H); bicuspid marginal teeth (C, F, I)." figureDoi="http://doi.org/10.5281/zenodo.5460660" httpUri="https://zenodo.org/record/5460660/files/figure.png" pageId="10" pageNumber="773">Fig. 10D–F</figureCitation>
).
</paragraph>
<paragraph id="8BB336EC782BA031FC84B936305F5288" blockId="10.[834,1476,1574,1790]" pageId="10" pageNumber="773">
<emphasis id="B978EAFE782BA031FC84B93637D05248" box="[835,1016,1574,1598]" italics="true" pageId="10" pageNumber="773">External features</emphasis>
. Animal with blackish grey reticulated skin, becoming pale yellow near foot sole and darker grey at eye stalks. Caudal foss (cf) large. Caudal horn (ch) raised with blackish grey colouration. Mantle edge well developed and blackish grey (
<figureCitation id="13372A69782BA031FC23B9B6301052C8" box="[996,1080,1702,1726]" captionStart="Fig" captionStartId="2.[135,170,1546,1568]" captionTargetBox="[136,1474,192,1529]" captionTargetId="figure-7@2.[136,1474,189,1531]" captionTargetPageId="2" captionText="Fig. 2. Living snails of Macrochlamys spp. from Thailand. A, Macrochlamys aurantia new species, paratype (CUMZ 7103), Thong Pha Phum, Kanchanaburi, shell width about 15 mm; B, M. coleus new species, paratype (CUMZ 7106), Sai Yok, Kanchanaburi, shell width about 15 mm; C,M. caverna new species, paratype (CUMZ 7109), Tha Wung, Lopburi, shell width about 17 mm; D, M. lemma new species, paratype (CUMZ 7116), Ban Mi, Lopburi, shell width about 15 mm; E, M. tanymentula new species, paratype (CUMZ 7119), Thong Pha Phum, Kanchanaburi, shell width about 10 mm; F, M. kelantanensis (CUMZ 7125), Lam Kaen, Phang-Nga, shell width about 20 mm." figureDoi="http://doi.org/10.5281/zenodo.5460640" httpUri="https://zenodo.org/record/5460640/files/figure.png" pageId="10" pageNumber="773">Fig. 2E</figureCitation>
). Shell and dorsal lobes morphology similar to 
<taxonomicName id="4C0C4D6F782BA031FC6BB9D7300352A8" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[940,1067,1735,1758]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="773" phylum="Mollusca" rank="species" species="aurantia" status="sp. nov.">
<emphasis id="B978EAFE782BA031FC6BB9D7300352A8" box="[940,1067,1735,1758]" italics="true" pageId="10" pageNumber="773">M. aurantia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782BA031FBF7B9D6309A52A8" box="[1072,1202,1734,1758]" pageId="10" pageNumber="773" rank="species">new species</taxonomicNameLabel>
. Snails secrete yellowishgreen slime when disturbed.
</paragraph>
</subSubSection>
<subSubSection id="C3166567782BA031FC84B83637DD53C8" pageId="10" pageNumber="773" type="distribution">
<paragraph id="8BB336EC782BA031FC84B83637DD53C8" blockId="10.[834,1475,1830,1982]" pageId="10" pageNumber="773">
<emphasis id="B978EAFE782BA031FC84B83630485348" bold="true" box="[835,1120,1830,1854]" pageId="10" pageNumber="773">Distribution and habitat.</emphasis>
<taxonomicName id="4C0C4D6F782BA031FBA1B83631A75348" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[1126,1423,1830,1854]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="773" phylum="Mollusca" rank="species" species="tanymentula" status="sp. nov.">
<emphasis id="B978EAFE782BA031FBA1B83631A75348" box="[1126,1423,1830,1854]" italics="true" pageId="10" pageNumber="773">Macrochlamys tanymentula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782BA031FA51B83637B85328" pageId="10" pageNumber="773" rank="species">new species</taxonomicNameLabel>
has a narrow distribution and is currently known only from the 
<typeStatus id="54B7884E782BA031FC62B87637FB5308" box="[933,979,1894,1918]" pageId="10" pageNumber="773">type</typeStatus>
locality with a low population density. Snails were found among leaf litter on the ground or crawling on limestone walls.
</paragraph>
</subSubSection>
<subSubSection id="C3166567782BA030FC84B8F631555375" lastPageId="11" lastPageNumber="774" pageId="10" pageNumber="773" type="discussion">
<paragraph id="8BB336EC782BA031FC84B8F631EA5C48" blockId="10.[835,1474,2022,2110]" pageId="10" pageNumber="773">
<emphasis id="B978EAFE782BA031FC84B8F6379A5388" bold="true" box="[835,946,2022,2046]" pageId="10" pageNumber="773">Remarks.</emphasis>
<taxonomicName id="4C0C4D6F782BA031FC7CB8F630CE5388" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[955,1254,2022,2046]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="773" phylum="Mollusca" rank="species" species="tanymentula" status="sp. nov.">
<emphasis id="B978EAFE782BA031FC7CB8F630CE5388" box="[955,1254,2022,2046]" italics="true" pageId="10" pageNumber="773">Macrochlamys tanymentula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782BA031FB28B8F6315A5388" box="[1263,1394,2022,2046]" pageId="10" pageNumber="773" rank="species">new species</taxonomicNameLabel>
can be distinguished by its globosely depressed shell, with wide shallow suture and a very narrow umbilicus. The genitalia
</paragraph>
<paragraph id="8BB336EC782AA030FE22BF6435AD5195" blockId="11.[485,1080,116,140]" lastBlockId="11.[113,752,1355,1507]" pageId="11" pageNumber="774">
Pholyotha et al.: 
<taxonomicName id="4C0C4D6F782AA030FD67BF6437BC54FA" box="[672,916,116,140]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="undetermined">Macrochlamys species</taxonomicName>
from 
<collectingCountry id="F31B767C782AA030FC1EBF64301054FA" box="[985,1080,116,140]" name="Thailand" pageId="11" pageNumber="774">Thailand</collectingCountry>
have a very long and slender epiphallus and flagellum, coiled epiphallic caecum of about one circle, very short vagina, and a large dart apparatus located at the vagina base opposite the penis. Its radula morphology is very similar to 
<taxonomicName id="4C0C4D6F782AA030FFB5BADC34DC5195" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[114,244,1484,1507]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="aurantia" status="sp. nov.">
<emphasis id="B978EAFE782AA030FFB5BADC34DC5195" box="[114,244,1484,1507]" italics="true" pageId="11" pageNumber="774">M. aurantia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782AA030FF3CBADC35A95195" box="[251,385,1483,1507]" pageId="11" pageNumber="774" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
<caption id="DF736664782AA030FFB6BBF130CB5162" ID-DOI="http://doi.org/10.5281/zenodo.5460652" ID-Zenodo-Dep="5460652" httpUri="https://zenodo.org/record/5460652/files/figure.png" pageId="11" pageNumber="774" startId="11.[113,148,1249,1271]" targetBox="[113,1451,189,1234]" targetPageId="11">
<paragraph id="8BB336EC782AA030FFB6BBF130CB5162" blockId="11.[113,1451,1249,1300]" pageId="11" pageNumber="774">
Fig. 7. Shells of 
<taxonomicName id="4C0C4D6F782AA030FEDFBBF135C65080" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[280,494,1249,1270]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="lemma" status="sp. nov.">
<emphasis id="B978EAFE782AA030FEDFBBF135C65080" box="[280,494,1249,1270]" italics="true" pageId="11" pageNumber="774">Macrochlamys lemma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782AA030FE30BBF236445081" box="[503,620,1249,1271]" pageId="11" pageNumber="774" rank="species">new species</taxonomicNameLabel>
(A, B), and 
<taxonomicName id="4C0C4D6F782AA030FD2ABBF237AE5080" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[749,902,1249,1271]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="tanymentula" status="sp. nov.">
<emphasis id="B978EAFE782AA030FD2ABBF237AE5080" box="[749,902,1249,1271]" italics="true" pageId="11" pageNumber="774">M. tanymentula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782AA030FC4ABBF2302B5081" box="[909,1027,1249,1271]" pageId="11" pageNumber="774" rank="species">new species</taxonomicNameLabel>
(C, D). A, 
<taxonomicName id="4C0C4D6F782AA030FBBFBBF131675080" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[1144,1359,1249,1270]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="lemma">
<emphasis id="B978EAFE782AA030FBBFBBF131675080" box="[1144,1359,1249,1270]" italics="true" pageId="11" pageNumber="774">Macrochlamys lemma</emphasis>
</taxonomicName>
holotype (CUMZ 7114); B, paratype (CUMZ 7115); C, 
<taxonomicName id="4C0C4D6F782AA030FD87BBEF36F05165" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[576,728,1278,1300]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="tanymentula">
<emphasis id="B978EAFE782AA030FD87BBEF36F05165" box="[576,728,1278,1300]" italics="true" pageId="11" pageNumber="774">M. tanymentula</emphasis>
</taxonomicName>
holotype (CUMZ 7117); D, paratype (CUMZ 7118).
</paragraph>
</caption>
<paragraph id="8BB336EC782AA030FFB6B91B35E65315" blockId="11.[112,754,1547,1891]" pageId="11" pageNumber="774">
This new species can be clearly distinguished from the other four new species proposed herein by its smaller shell with weakly impressed suture, narrow umbilicus, and very long epiphallus and flagellum, short vagina, and no penial caecum. The other four new species have relatively larger shells (
<tableCitation id="C68E0357782AA030FF7CB9BB352352B5" box="[187,267,1707,1731]" captionStart="Table 1" captionStartId="12.[136,191,189,211]" captionText="Table 1. Comparison of shell and genital system of Macrochlamys species in Thailand." pageId="11" pageNumber="774">Table 1</tableCitation>
), shorter epiphallus and flagellum, and with a short penial caecum. Furthermore, 
<taxonomicName id="4C0C4D6F782AA030FE2EB9DC36415295" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[489,617,1740,1763]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="aurantia" status="sp. nov.">
<emphasis id="B978EAFE782AA030FE2EB9DC36415295" box="[489,617,1740,1763]" italics="true" pageId="11" pageNumber="774">M. aurantia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782AA030FDB7B9DC36D85295" box="[624,752,1739,1763]" pageId="11" pageNumber="774" rank="species">new species</taxonomicNameLabel>
and 
<taxonomicName id="4C0C4D6F782AA030FF5AB9FC352C5375" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[157,260,1771,1795]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="coleus" status="sp. nov.">
<emphasis id="B978EAFE782AA030FF5AB9FC352C5375" box="[157,260,1771,1795]" italics="true" pageId="11" pageNumber="774">M. coleus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782AA030FECDB9FC35A15375" box="[266,393,1771,1795]" pageId="11" pageNumber="774" rank="species">new species</taxonomicNameLabel>
have a channel-shaped suture and orange to pale orange body colour, while 
<taxonomicName id="4C0C4D6F782AA030FDF8B81C36935355" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[575,699,1804,1827]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="caverna" status="sp. nov.">
<emphasis id="B978EAFE782AA030FDF8B81C36935355" box="[575,699,1804,1827]" italics="true" pageId="11" pageNumber="774">M. caverna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782AA030FD03B81C34965335" pageId="11" pageNumber="774" rank="species">new species</taxonomicNameLabel>
and 
<taxonomicName id="4C0C4D6F782AA030FF30B83C354C5335" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[247,356,1835,1859]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="lemma" status="sp. nov.">
<emphasis id="B978EAFE782AA030FF30B83C354C5335" box="[247,356,1835,1859]" italics="true" pageId="11" pageNumber="774">M. lemma</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782AA030FEAAB83C35C75335" box="[365,495,1835,1859]" pageId="11" pageNumber="774" rank="species">new species</taxonomicNameLabel>
have oblique trapezoid penial pilasters inside the penis.
</paragraph>
<paragraph id="8BB336EC782AA030FFB6B89B31555375" blockId="11.[113,753,1931,2115]" lastBlockId="11.[811,1453,1355,1795]" pageId="11" pageNumber="774">
<taxonomicName id="4C0C4D6F782AA030FFB6B89B358D53D5" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[113,421,1931,1955]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="tanymentula" status="sp. nov.">
<emphasis id="B978EAFE782AA030FFB6B89B358D53D5" box="[113,421,1931,1955]" italics="true" pageId="11" pageNumber="774">Macrochlamys tanymentula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782AA030FE77B89C361F53D5" box="[432,567,1931,1955]" pageId="11" pageNumber="774" rank="species">new species</taxonomicNameLabel>
differs from 
<taxonomicName id="4C0C4D6F782AA030FD14B89C355653B5" authority="(Benson, 1859)" baseAuthorityName="Benson" baseAuthorityYear="1859" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="petasus">
<emphasis id="B978EAFE782AA030FD14B89C34EC53B5" italics="true" pageId="11" pageNumber="774">M. petasus</emphasis>
(
<bibRefCitation id="EF9D4B1D782AA030FF10B8BB355E53B5" author="Benson WH" box="[215,374,1963,1987]" pageId="11" pageNumber="774" pagination="387 - 393" refId="ref12352" refString="Benson WH (1859) New Helicidae collected by W. Theobald, Esq., jun., in Burmah and the Khasia Hills, and described by W. H. Benson. Annals and Magazine of Natural History, Series 3, Volume 3: 387 - 393." type="journal article" year="1859">Benson, 1859</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="4C0C4D6F782AA030FE7CB8BC36D853B5" authority="(Benson, 1863)" baseAuthorityName="Benson" baseAuthorityYear="1863" box="[443,752,1963,1987]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="aspides">
<emphasis id="B978EAFE782AA030FE7CB8BC361F53B5" box="[443,567,1963,1987]" italics="true" pageId="11" pageNumber="774">M. aspides</emphasis>
(
<bibRefCitation id="EF9D4B1D782AA030FD8DB8BB36C053B5" author="Benson WH" box="[586,744,1963,1987]" pageId="11" pageNumber="774" pagination="318 - 323" refId="ref12436" refString="Benson WH (1863) Characters of new land-shells from the Andaman Islands, Burmah, and Ceylon, and of the animal of Sophina. Annals and Magazine of Natural History, Series 3, Volume 11: 318 - 323." type="journal article" year="1863">Benson, 1863</bibRefCitation>
)
</taxonomicName>
from 
<collectingCountry id="F31B767C782AA030FF72B8DC350C5392" box="[181,292,1996,2020]" name="Myanmar" pageId="11" pageNumber="774">Myanmar</collectingCountry>
by its rounded last whorl and simple apertural lip. In comparison, 
<taxonomicName id="4C0C4D6F782AA030FE72B8FC36045C75" baseAuthorityName="Benson" baseAuthorityYear="1859" box="[437,556,2028,2051]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="petasus">
<emphasis id="B978EAFE782AA030FE72B8FC36045C75" box="[437,556,2028,2051]" italics="true" pageId="11" pageNumber="774">M. petasus</emphasis>
</taxonomicName>
has a rounded to slightly subangulated last whorl, and the inside edge of the peristome is slightly expanded, while 
<taxonomicName id="4C0C4D6F782AA030FDFBB73C36995C35" baseAuthorityName="Benson" baseAuthorityYear="1863" box="[572,689,2091,2115]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="aspides">
<emphasis id="B978EAFE782AA030FDFBB73C36995C35" box="[572,689,2091,2115]" italics="true" pageId="11" pageNumber="774">M. aspides</emphasis>
</taxonomicName>
has a strongly thickened basal margin at the edge of the peristome (
<bibRefCitation id="EF9D4B1D782AA030FCF4BA7B309651F5" author="Blanford WT &amp; Godwin-Austen HH" box="[819,1214,1387,1411]" pageId="11" pageNumber="774" pagination="1 - 311" refId="ref12680" refString="Blanford WT &amp; Godwin-Austen HH (1908) Mollusca: Testacellidae and Zonitidae. In: Bingham CT (ed.), The Fauna of British India including Ceylon and Burma. Taylor and Francis, London. Pp. 1 - 311." type="book chapter" year="1908">Blanford &amp; Godwin-Austen, 1908</bibRefCitation>
). Compared with 
<taxonomicName id="4C0C4D6F782AA030FA49BA7C303B51D5" authority="Blanford, 1905" authorityName="Blanford" authorityYear="1905" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="notha">
<emphasis id="B978EAFE782AA030FA49BA7C318C51F5" box="[1422,1444,1388,1411]" italics="true" pageId="11" pageNumber="774">M</emphasis>
. 
<emphasis id="B978EAFE782AA030FCEABA9B374251D5" box="[813,874,1419,1443]" italics="true" pageId="11" pageNumber="774">notha</emphasis>
<bibRefCitation id="EF9D4B1D782AA030FCB6BA9B303B51D5" author="Blanford WT" box="[881,1043,1419,1443]" pageId="11" pageNumber="774" pagination="441 - 447" refId="ref12631" refString="Blanford WT (1905) Descriptions of Indian and Burmese land-shells referred to the genera Macrochlamys, Bensonia, Taphrospira, (gen. nov.), Microcystina, Euplecta, and Polita. Proceedings of the Zoological Society of London, 2: 441 - 447." type="journal article" year="1905">Blanford, 1905</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="F31B767C782AA030FB91BA9C309651D2" box="[1110,1214,1420,1444]" name="Myanmar" pageId="11" pageNumber="774">Myanmar</collectingCountry>
and 
<taxonomicName id="4C0C4D6F782AA030FB33BA9C315951D5" authorityName="Mollendorff" authorityYear="1902" box="[1268,1393,1419,1443]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="brunnea">
<emphasis id="B978EAFE782AA030FB33BA9C315951D5" box="[1268,1393,1419,1443]" italics="true" pageId="11" pageNumber="774">M. brunnea</emphasis>
</taxonomicName>
from 
<collectingCountry id="F31B767C782AA030FCEABABB37A751B5" box="[813,911,1451,1475]" name="Thailand" pageId="11" pageNumber="774">Thailand</collectingCountry>
, 
<taxonomicName id="4C0C4D6F782AA030FC5BBABC306F51B5" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[924,1095,1451,1475]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="tanymentula" status="sp. nov.">
<emphasis id="B978EAFE782AA030FC5BBABC306F51B5" box="[924,1095,1451,1475]" italics="true" pageId="11" pageNumber="774">M. tanymentula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782AA030FB97BABC30FC51B5" box="[1104,1236,1451,1475]" pageId="11" pageNumber="774" rank="species">new species</taxonomicNameLabel>
has a smooth shell surface, relatively large and thin shell, with a slightly wider umbilicus. 
<taxonomicName id="4C0C4D6F782AA030FC63BAFB30AF5275" authorityName="Blanford" authorityYear="1905" box="[932,1159,1515,1539]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="notha">
<emphasis id="B978EAFE782AA030FC63BAFB30AF5275" box="[932,1159,1515,1539]" italics="true" pageId="11" pageNumber="774">Macrochlamys notha</emphasis>
</taxonomicName>
has a dull shell with radial sculpture while 
<taxonomicName id="4C0C4D6F782AA030FC14B91C30665255" authorityName="Mollendorff" authorityYear="1902" box="[979,1102,1547,1571]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="brunnea">
<emphasis id="B978EAFE782AA030FC14B91C30665255" box="[979,1102,1547,1571]" italics="true" pageId="11" pageNumber="774">M. brunnea</emphasis>
</taxonomicName>
has a relatively smaller shell and narrower umbilicus. 
<taxonomicName id="4C0C4D6F782AA030FBD1B93B374B5215" authority="Stoliczka, 1873" authorityName="Stoliczka" authorityYear="1873" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="stephoides">
<emphasis id="B978EAFE782AA030FBD1B93B311D5235" box="[1046,1333,1579,1603]" italics="true" pageId="11" pageNumber="774">Macrochlamys stephoides</emphasis>
<bibRefCitation id="EF9D4B1D782AA030FAF8B93B374B5215" author="Stoliczka F" pageId="11" pageNumber="774" pagination="11 - 38" refId="ref14126" refString="Stoliczka F (1873) On the land shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea. Journal of the Asiatic Society of Bengal, 42: 11 - 38." type="journal article" year="1873">Stoliczka, 1873</bibRefCitation>
</taxonomicName>
from 
<collectingCountry id="F31B767C782AA030FC69B95B303A5215" box="[942,1042,1611,1635]" name="Malaysia" pageId="11" pageNumber="774">Malaysia</collectingCountry>
has a higher spire compared to the depressed spire of 
<taxonomicName id="4C0C4D6F782AA030FC33B97C30B452F5" authority="Pholyotha &amp; Sutcharit &amp; Panha, 2018" authorityName="Pholyotha &amp; Sutcharit &amp; Panha" authorityYear="2018" box="[1012,1180,1643,1667]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="tanymentula" status="sp. nov.">
<emphasis id="B978EAFE782AA030FC33B97C30B452F5" box="[1012,1180,1643,1667]" italics="true" pageId="11" pageNumber="774">M. tanymentula</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B5785782AA030FB64B97C310E52F5" box="[1187,1318,1643,1667]" pageId="11" pageNumber="774" rank="species">new species</taxonomicNameLabel>
. In addition, 
<bibRefCitation id="EF9D4B1D782AA030FCEAB99B37C952D5" author="Stoliczka F" box="[813,993,1675,1699]" pageId="11" pageNumber="774" pagination="11 - 38" refId="ref14126" refString="Stoliczka F (1873) On the land shells of Penang Island, with descriptions of the animals and anatomical notes; part second, Helicacea. Journal of the Asiatic Society of Bengal, 42: 11 - 38." type="journal article" year="1873">Stoliczka (1873)</bibRefCitation>
mentioned that 
<taxonomicName id="4C0C4D6F782AA030FB52B99C310452D5" authorityName="Stoliczka" authorityYear="1873" box="[1173,1324,1675,1699]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="stephoides">
<emphasis id="B978EAFE782AA030FB52B99C308352D5" box="[1173,1195,1676,1699]" italics="true" pageId="11" pageNumber="774">M</emphasis>
. 
<emphasis id="B978EAFE782AA030FB7CB99B310452D5" box="[1211,1324,1675,1699]" italics="true" pageId="11" pageNumber="774">stephoides</emphasis>
</taxonomicName>
has similar genitalia to 
<taxonomicName id="4C0C4D6F782AA030FC69B9BC318052B5" authority="Benson sensu Godwin-Austen, 1883" authorityName="Godwin-Austen" authorityYear="1883" box="[942,1448,1707,1731]" class="Gastropoda" family="Ariophantidae" genus="Macrochlamys" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="774" phylum="Mollusca" rank="species" species="indica">
<emphasis id="B978EAFE782AA030FC69B9BC303D52B5" box="[942,1045,1707,1731]" italics="true" pageId="11" pageNumber="774">M. indica</emphasis>
Benson sensu 
<bibRefCitation id="EF9D4B1D782AA030FB70B9BB318052B5" author="Godwin-Austen HH" box="[1207,1448,1707,1731]" pageId="11" pageNumber="774" pagination="47 - 238" refId="ref12935" refString="Godwin-Austen HH (1883 - 1907) Land and Freshwater Mollusca of India, Including South Arabia, Baluchistan, Afghanistan, Kashmir, Nepal, Burmah, Pegu, Tenasserim, Malay Peninsula, Ceylon, and other Islands of the Indian Ocean. Taylor and Francis, London, Volume 1 (1883 - 1887): 67 - 206, pls. 13 - 62; Volume 2 (1898 - 1907): 47 - 238, pls. 70 - 117." type="journal article" year="1883">Godwin-Austen, 1883</bibRefCitation>
</taxonomicName>
. In contrast, this new species has a very long epiphallus and flagellum (see 
<bibRefCitation id="EF9D4B1D782AA030FC09B9FB31585375" author="Blanford WT &amp; Godwin-Austen HH" box="[974,1392,1771,1795]" pageId="11" pageNumber="774" pagination="1 - 311" refId="ref12680" refString="Blanford WT &amp; Godwin-Austen HH (1908) Mollusca: Testacellidae and Zonitidae. In: Bingham CT (ed.), The Fauna of British India including Ceylon and Burma. Taylor and Francis, London. Pp. 1 - 311." type="book chapter" year="1908">Blanford &amp; Godwin-Austen, 1908: 95</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>