146 lines
16 KiB
XML
146 lines
16 KiB
XML
<document id="8A4903A2F9E59791E7FF81D0CF45EBA5" ID-CLB-Dataset="24540" ID-DOI="10.11646/zootaxa.4708.1.1" ID-GBIF-Dataset="653d580d-e1c7-499c-8604-4b5a7b1920ef" ID-ISSN="1175-5326" ID-Zenodo-Dep="3587181" ID-ZooBank="09F24F0B-5090-4089-A525-8E5150C91EF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1576825058981" checkinUser="plazi" docAuthor="Perkins, Philip D." docDate="2019" docId="03A587E7A869FFEE0CBA12A5FB1D972E" docLanguage="en" docName="zootaxa.4708.1.1.pdf" docOrigin="Zootaxa 4708 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Hydrochus bituberculatus Perkins 2019, new species" docType="treatment" docVersion="8" lastPageNumber="11" masterDocId="FF9CFF9FA863FFE40C2D1244FFE19255" masterDocTitle="Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)" masterLastPageNumber="59" masterPageNumber="1" pageNumber="11" updateTime="1698775642300" updateUser="ExternalLinkService">
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<mods:mods id="9E18F1B3F526F188C217A176088C74C5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="875DF9F92862ED1475DF6FEC5CD1008E">
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<mods:title id="6AE3A366EE7D5AB17C15234FADA14AB2">Taxonomy of Venezuelan water beetles in the genus Hydrochus Leach, 1817, and an analysis of male genitalia morphology (Coleoptera: Hydrochidae)</mods:title>
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<mods:name id="076F4F1D526898B6C781FB1CD21D86B2" type="personal">
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<mods:namePart id="A15049AA7966C3FA1E47983441596121">Perkins, Philip D.</mods:namePart>
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<mods:typeOfResource id="90631B4796508D9A1AEFC5C601862F67">text</mods:typeOfResource>
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<mods:titleInfo id="162DCDDC7C52EE0831872B71D2D2D658">
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<mods:title id="195C35E869B2071116D3514F7FF0030D">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="773488CBF956C4F0A920D2B0B63A5348">
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<mods:date id="2DA6927E555FAB7D00FEB328FB4890F4">2019</mods:date>
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<mods:detail id="F6D1CE1CA05274480A578EEF0E9719B4" type="pubDate">
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<mods:number id="3B953068CD905066FEAFA240DAF4BC40">2019-12-12</mods:number>
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<mods:detail id="D16AB7A61170DAFE62D7097445C56350" type="volume">
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<mods:number id="133EDC846D6C59BDFBAB00AAFD539B55">4708</mods:number>
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<mods:detail id="468CE8353FF8AA99626F163DBE5AA68A" type="issue">
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<mods:number id="86EEE4034B03087B7EE4B780885B0112">1</mods:number>
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<mods:classification id="1859F6BBC20E9BC577C86612DD44526F">journal article</mods:classification>
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<mods:identifier id="039755C4236747F10F6F7753B6EE7D9A" type="CLB-Dataset">24540</mods:identifier>
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<mods:identifier id="546FE6F86B9C8AFB65E580581E982518" type="DOI">10.11646/zootaxa.4708.1.1</mods:identifier>
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<mods:identifier id="59D6645FA87B38E5CF698E6F10D71D87" type="GBIF-Dataset">653d580d-e1c7-499c-8604-4b5a7b1920ef</mods:identifier>
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<mods:identifier id="075ED3494BDBDB8F4D34BC38D5C88AD7" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03A587E7A869FFEE0CBA12A5FB1D972E" ID-DOI="http://doi.org/10.5281/zenodo.5941779" ID-GBIF-Taxon="160798407" ID-Zenodo-Dep="5941779" LSID="urn:lsid:plazi:treatment:03A587E7A869FFEE0CBA12A5FB1D972E" httpUri="http://treatment.plazi.org/id/03A587E7A869FFEE0CBA12A5FB1D972E" lastPageNumber="11" pageId="10" pageNumber="11">
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<subSubSection id="C316657AA869FFEE0CBA12A5FD8B92AE" box="[151,618,225,252]" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="8BB336F1A869FFEE0CBA12A5FD8B92AE" blockId="10.[151,618,225,288]" box="[151,618,225,252]" pageId="10" pageNumber="11">
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<heading id="D0FB819DA869FFEE0CBA12A5FD8B92AE" bold="true" box="[151,618,225,252]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
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<emphasis id="B978EAE3A869FFEE0CBA12A5FD8B92AE" bold="true" box="[151,618,225,252]" pageId="10" pageNumber="11">
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<taxonomicName id="4C0C4D72A869FFEE0CBA12A5FE3392A9" ID-CoL="8Y4PG" authority="Perkins, 2019" authorityName="Perkins" authorityYear="2019" box="[151,466,225,252]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bituberculatus" status="sp. nov.">
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<emphasis id="B978EAE3A869FFEE0CBA12A5FE3392A9" bold="true" box="[151,466,225,252]" italics="true" pageId="10" pageNumber="11">Hydrochus bituberculatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A24B5798A869FFEE0DF712A6FD8B92AE" box="[474,618,225,252]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C316657AA869FFEE0CBA1341FF389375" box="[151,217,261,288]" pageId="10" pageNumber="11" type="description">
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<paragraph id="8BB336F1A869FFEE0CBA1341FF389375" blockId="10.[151,618,225,288]" box="[151,217,261,288]" pageId="10" pageNumber="11">
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<figureCitation id="13372A74A869FFEE0CBA1341FF389375" box="[151,217,261,288]" captionStart="FIGURE 7" captionStartId="21.[151,250,1716,1741]" captionTargetBox="[186,1413,181,1668]" captionTargetId="figure@21.[151,1435,181,1691]" captionTargetPageId="21" captionText="FIGURE 7. Hydrochus bituberculatus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587195" httpUri="https://zenodo.org/record/3587195/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C316657AA869FFEE0CBA130AFE94904E" pageId="10" pageNumber="11" type="materials_examined">
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<paragraph id="8BB336F1A869FFEE0CBA130AFBCE93DE" blockId="10.[151,1437,333,1404]" pageId="10" pageNumber="11">
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<emphasis id="B978EAE3A869FFEE0CBA130AFEA6933D" bold="true" box="[151,327,333,360]" pageId="10" pageNumber="11">Type Material.</emphasis>
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<materialsCitation id="3B643CACA869FFEE0D60130AFBCE93DE" ID-GBIF-Occurrence="2521263718" collectingDate="1989-01-25" collectionCode="NMNH" country="Venezuela" location="Rio Horero" municipality="Puerto Ayacucho" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Amazonas" typeStatus="holotype">
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<emphasis id="B978EAE3A869FFEE0D60130AFDF4933D" bold="true" box="[333,533,333,360]" pageId="10" pageNumber="11">
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<typeStatus id="54B78853A869FFEE0D60130AFE5B933D" box="[333,442,334,360]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
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(male):
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</emphasis>
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“
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<collectingCountry id="F31B7661A869FFEE0E081309FD7E9332" box="[549,671,333,359]" name="Venezuela" pageId="10" pageNumber="11">Venezuela</collectingCountry>
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: T.F.
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<collectingRegion id="49C8F813A869FFEE0EFA130AFCB1933D" box="[727,848,334,360]" country="Venezuela" name="Amazonas" pageId="10" pageNumber="11">Amazonas</collectingRegion>
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,
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<collectingMunicipality id="6BD7AC8BA869FFEE0F76130AFBF69332" box="[859,1047,333,360]" pageId="10" pageNumber="11">Puerto Ayacucho</collectingMunicipality>
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(66kmN),
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<location id="8ED3602AA869FFEE08BC1309FAF3933D" LSID="urn:lsid:plazi:treatment:03A587E7A869FFEE0CBA12A5FB1D972E:8ED3602AA869FFEE08BC1309FAF3933D" box="[1169,1298,333,360]" country="Venezuela" municipality="Puerto Ayacucho" name="Rio Horero" pageId="10" pageNumber="11" stateProvince="Amazonas">Rio Horero</location>
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,
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<date id="FFB21031A869FFEE09311309FA79933D" box="[1308,1432,333,360]" pageId="10" pageNumber="11" value="1989-01-25">
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<collectingDate id="EFF6E9D9A869FFEE09311309FA79933D" box="[1308,1432,333,360]" pageId="10" pageNumber="11" value="1989-01-25">25Jan1989</collectingDate>
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</date>
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, netted, stream edge // Collected by PJSpangler, RAFaitoute & CBBarr” (
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<collectionCode id="ED1DAE34A869FFEE0FEA1335FBC293DE" box="[967,1059,369,395]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="10" pageNumber="11" type="Museum">NMNH</collectionCode>
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).
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</materialsCitation>
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</paragraph>
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<paragraph id="8BB336F1A869FFEE0CEA13D1FE94904E" blockId="10.[151,1437,333,1404]" pageId="10" pageNumber="11">
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<emphasis id="B978EAE3A869FFEE0CEA13D1FE3393E5" bold="true" box="[199,466,405,432]" pageId="10" pageNumber="11">Differential Diagnosis.</emphasis>
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Among Venezuelan species, recognized by the combination of moderate size (ca.
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<quantity id="4CF49B14A869FFEE094613D1FF269381" metricMagnitude="-3" metricUnit="m" metricValue="3.72" pageId="10" pageNumber="11" unit="mm" value="3.72">3.72 mm</quantity>
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), the shallow pronotal depressions, the large elytral punctures, the large elytral tubercles, and the male genitalia (
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<figureCitation id="13372A74A869FFEE0C8D1399FF0793AD" box="[160,230,477,504]" captionStart="FIGURE 7" captionStartId="21.[151,250,1716,1741]" captionTargetBox="[186,1413,181,1668]" captionTargetId="figure@21.[151,1435,181,1691]" captionTargetPageId="21" captionText="FIGURE 7. Hydrochus bituberculatus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587195" httpUri="https://zenodo.org/record/3587195/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
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; described below). This species has a generalized
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<taxonomicName id="4C0C4D72A869FFEE0F051399FC4393A2" authorityName="Leach" authorityYear="1817" box="[808,930,477,503]" class="Insecta" family="Hydrochidae" genus="Hydrochus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
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<emphasis id="B978EAE3A869FFEE0F051399FC4393A2" box="[808,930,477,503]" italics="true" pageId="10" pageNumber="11">Hydrochus</emphasis>
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</taxonomicName>
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habitus; reliable determinations will require dissection of males.
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</paragraph>
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</subSubSection>
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<subSubSection id="C316657AA869FFEE0CEA1061FBE99702" pageId="10" pageNumber="11" type="description">
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<paragraph id="8BB336F1A869FFEE0CEA1061FE5D90FE" blockId="10.[151,1437,333,1404]" pageId="10" pageNumber="11">
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<emphasis id="B978EAE3A869FFEE0CEA1061FEB09015" bold="true" box="[199,337,549,576]" pageId="10" pageNumber="11">Description</emphasis>
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. Habitus as illustrated (
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<figureCitation id="13372A74A869FFEE0E701061FD439015" box="[605,674,549,576]" captionStart="FIGURE 7" captionStartId="21.[151,250,1716,1741]" captionTargetBox="[186,1413,181,1668]" captionTargetId="figure@21.[151,1435,181,1691]" captionTargetPageId="21" captionText="FIGURE 7. Hydrochus bituberculatus n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587195" httpUri="https://zenodo.org/record/3587195/files/figure.png" pageId="10" pageNumber="11">Fig. 7</figureCitation>
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). Size:
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<typeStatus id="54B78853A869FFEE0EDA1061FCB8906A" box="[759,857,549,575]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 3.72/1.28; head width 0.90; pronotum l/w 0.82/0.82; PA 0.74; PB 0.72; elytra 2.36/1.28. Body size moderately large. Dorsum grey to silver grey, with slight to moderately strong iridescence, elytra with black spots. Legs brown, with tibiofemoral joints darker.
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</paragraph>
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<paragraph id="8BB336F1A869FFEE0CEA10F1FCE29142" blockId="10.[151,1437,333,1404]" pageId="10" pageNumber="11">
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Punctation on elytra ca. 2–
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<date id="FFB21031A869FFEE0DD810F1FDEE9085" box="[501,527,693,720]" pageId="10" pageNumber="11">5x</date>
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that of pronotum. Elytra interstriae ca. 1–1.5x strial puncture diameter. Interstria 9
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<superScript id="7C799BB9A869FFEE0C88109CFF5390B3" attach="left" box="[165,178,728,742]" fontSize="6" pageId="10" pageNumber="11">th</superScript>
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more convex than others, overhanging 10
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<superScript id="7C799BB9A869FFEE0EB6109CFD4990B3" attach="left" box="[667,680,728,742]" fontSize="6" pageId="10" pageNumber="11">th</superScript>
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interstria. Callus on 5
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<superScript id="7C799BB9A869FFEE0F86109CFC5990B3" attach="left" box="[939,952,728,742]" fontSize="6" pageId="10" pageNumber="11">th</superScript>
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at usual area, very large and prominently raised; 4
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<superScript id="7C799BB9A869FFEE0CDA10B8FEE5915F" attach="left" box="[247,260,764,778]" fontSize="6" pageId="10" pageNumber="11">th</superScript>
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interstriae raised in area anterior to 5
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<superScript id="7C799BB9A869FFEE0E8A10B8FD55915F" attach="left" box="[679,692,764,778]" fontSize="6" pageId="10" pageNumber="11">th</superScript>
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callus.
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</paragraph>
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<paragraph id="8BB336F1A869FFEE0CEA1166FD9191D6" blockId="10.[151,1437,333,1404]" pageId="10" pageNumber="11">Pronotum length and width equal, widest at about anterior 1/4, narrowed at base, sides weakly sinuate, smooth; anterior margin arcuate or slightly bisinuate; punctures small and rather sparse; depressions shallow, more densely punctate than reliefs between depressions.</paragraph>
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<paragraph id="8BB336F1A869FFEE0CEA11C9FB6091BA" blockId="10.[151,1437,333,1404]" pageId="10" pageNumber="11">
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Elytra with spaces between strial punctures ca. 0.5–
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<date id="FFB21031A869FFEE0F3811C9FCCC91FD" box="[789,813,909,936]" pageId="10" pageNumber="11">1x</date>
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their diameter; Punctures becoming very large at summit of posterior declivity; suture rather prominently raised at summit of posterior declivity; apices rather sharply conjointly rounded in dorsal view, in lateral view outer margin with moderate angulation.
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</paragraph>
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<paragraph id="8BB336F1A869FFEE0CEA11BDFE619662" blockId="10.[151,1437,333,1404]" pageId="10" pageNumber="11">Ventral characters: Mentum moderately coarsely punctate, with shallow median fovea. Submentum with two very shallow foveae.</paragraph>
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<paragraph id="8BB336F1A869FFEE0CEA1605FD3296BE" blockId="10.[151,1437,333,1404]" pageId="10" pageNumber="11">
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Male genitalia (
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<figureCitation id="13372A74A869FFEE0D531605FE33960E" box="[382,466,1089,1115]" captionStart="FIGURE 17" captionStartId="34.[151,250,1794,1819]" captionTargetBox="[200,1384,203,1755]" captionTargetId="figure@34.[183,1403,181,1768]" captionTargetPageId="34" captionText="FIGURE 17. Hydrochus basilaris n. sp., habitus and male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.3587217" httpUri="https://zenodo.org/record/3587217/files/figure.png" pageId="10" pageNumber="11">Fig. 17</figureCitation>
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) general characters: paramere tips sub-sagittate, distal ends pointed, lateral margins arcuate, plsa very small, angulate, medial margins of tips weakly arcuate and well separated from lateral margins of aedeagus, most of tips extending beyond aedeagus, in lateral view tips narrow, sharply pointed, arcuate toward dorsad; aedeagus moderately wide at widest point, margins arcuate at midlength, lateral margins weakly sclerotized; basal piece with rim of orifice strongly sclerotized.
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</paragraph>
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<paragraph id="8BB336F1A869FFEE0CEA16B1FF289766" blockId="10.[151,1437,333,1404]" pageId="10" pageNumber="11">Dorsal surface: adtl located slightly distal to midlength of aedeagus; adbl large; pdmm slightly overlapping alm.</paragraph>
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<paragraph id="8BB336F1A869FFEE0CEA1779FBE99702" blockId="10.[151,1437,333,1404]" box="[199,1032,1341,1368]" pageId="10" pageNumber="11">Ventral surface: avtl subbasal; pvmm approximating alm, not overlapping.</paragraph>
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</subSubSection>
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<subSubSection id="C316657AA869FFEE0CEA1726FB1D972E" box="[199,1276,1376,1404]" pageId="10" pageNumber="11" type="etymology">
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<paragraph id="8BB336F1A869FFEE0CEA1726FB1D972E" blockId="10.[151,1437,333,1404]" box="[199,1276,1376,1404]" pageId="10" pageNumber="11">
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<emphasis id="B978EAE3A869FFEE0CEA1726FEAF9729" bold="true" box="[199,334,1378,1404]" pageId="10" pageNumber="11">Etymology.</emphasis>
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Named in reference to the large tubercles of the elytra, present on the 5
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<superScript id="7C799BB9A869FFEE085D1724FB9C973B" attach="left" box="[1136,1149,1376,1390]" fontSize="6" pageId="10" pageNumber="11">th</superScript>
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interstriae.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |