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<document id="38E65FF641FBBF3D7690A413DC04DF93" ID-CLB-Dataset="8756" ID-DOI="10.5852/ejt.2021.731.1209" ID-GBIF-Dataset="3cf2c6ad-4274-4301-9cd0-e8295eace8cc" ID-ISSN="2118-9773" ID-Zenodo-Dep="4452219" ID-ZooBank="070B1EBB-976A-4EBB-B37D-6CB4A10D09FF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1611143662020" checkinUser="plazi" docAuthor="Vargas, Héctor A." docDate="2021" docId="03A587BDBB265E61617763DB9EA060CD" docLanguage="en" docName="ejt-731_117-134.pdf" docOrigin="European Journal of Taxonomy 731" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Helioandesia tarregai Vargas 2021, gen. et sp. nov." docType="treatment" docUuid="935E51EF-4505-454E-98DA-1FA3156E7ED7" docUuidSource="ZooBank" docVersion="8" lastPageNumber="130" masterDocId="FF9CFFC5BB2D5E6C634C625F9D00641F" masterDocTitle="Systematics of Helioandesia tarregai gen. et sp. nov. (Lepidoptera: Yponomeutoidea: Heliodinidae) from the Andes of Northern Chile" masterLastPageNumber="134" masterPageNumber="117" pageNumber="128" updateTime="1698924879965" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B3E73B62537977641BD9FE9F6696AB63">Systematics of Helioandesia tarregai gen. et sp. nov. (Lepidoptera: Yponomeutoidea: Heliodinidae) from the Andes of Northern Chile</mods:title>
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<mods:namePart id="0AC8295EED441CFB242B8D785919EDE9">Vargas, Héctor A.</mods:namePart>
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<mods:affiliation id="5C150E031651E3F0D4D055129EE924ED">Departamento de Recursos Ambientales, Facultad de Ciencias Agronómicas, Universidad de Tarapacá, Arica, Chile.</mods:affiliation>
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<mods:title id="06FC6BC33F33C244F7E053754D57535F">European Journal of Taxonomy</mods:title>
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<subSubSection id="C3166520BB265E67617763DB9EF96580" box="[571,1017,388,415]" pageId="11" pageNumber="128" type="nomenclature">
<paragraph id="8BB336ABBB265E67617763DB9EF96580" blockId="11.[371,1217,388,485]" box="[571,1017,388,415]" pageId="11" pageNumber="128">
<heading id="D0FB81C7BB265E67617763DB9EF96580" box="[571,1017,388,415]" centered="true" fontSize="11" level="2" pageId="11" pageNumber="128" reason="2">
<taxonomicName id="4C0C4D28BB265E67617763DB9E426581" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[571,834,388,415]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="tarregai" status="gen. et sp. nov.">
<emphasis id="B978EAB9BB265E67617763DB9E426581" bold="true" box="[571,834,388,415]" italics="true" pageId="11" pageNumber="128">Helioandesia tarregai</emphasis>
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<taxonomicNameLabel id="A24B57C2BB265E67600563DA9EF96580" box="[841,1017,389,415]" pageId="11" pageNumber="128" rank="species">gen. et sp. nov.</taxonomicNameLabel>
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<paragraph id="8BB336ABBB265E6761EA63949E8E65FA" blockId="11.[371,1217,388,485]" box="[678,910,459,485]" pageId="11" pageNumber="128">
<figureCitation id="13372A2EBB265E6761EA63949E0E65FA" box="[678,782,459,485]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStart-3="Fig" captionStartId-0="7.[189,232,1928,1954]" captionStartId-1="8.[188,231,1963,1989]" captionStartId-2="9.[189,232,1857,1883]" captionStartId-3="10.[191,234,1927,1953]" captionTargetBox-0="[189,1398,757,1889]" captionTargetBox-1="[190,1397,1272,1927]" captionTargetBox-2="[327,1261,319,1818]" captionTargetBox-3="[189,1398,559,1861]" captionTargetId-0="figure-163@7.[189,1398,757,1889]" captionTargetId-1="figure-364@8.[190,1397,1272,1927]" captionTargetId-2="figure-11@9.[326,1261,319,1818]" captionTargetId-3="figure-92@10.[189,1398,559,1861]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionTargetPageId-3="10" captionText-0="Fig. 3. Helioandesia tarregai gen. et sp. nov., ♂, holotype (IDEA-LEPI-2020-015). A. Dorsal view. B. Head, lateral. C. Basal flagellomeres of right antenna. D. Three bulging metallic spots on the basal half of forewing. E. Hair pencil from base of subcosta, ventral. Scale bar: 1 mm." captionText-1="Fig. 4. Helioandesia tarregai gen. et sp. nov., ♀, paratype (IDEA-LEPI-2020-024), wing venation showing frenulum with two bristles of unequal length. Scale bar: 1 mm." captionText-2="Fig. 5. Male genitalia and eighth segment of Helioandesia tarregai gen. et sp. nov., ♂, paratype (IDEA- LEPI-2020-021). A. Male genitalia, ventral view, phallus removed. B. Phallus, lateral view. C. Tergum and pleural lobes of eighth segment. D. Socius, closed arrow in A. E. Small spine-like cornuti on the vesica, open arrow in A. F. Medial part of the cornutus. G. Coremata between pleural lobes of segment VIII and genitalia. Scale bars: 0.1 mm." captionText-3="Fig. 6. Female genitalia of Helioandesia tarregai gen. et sp. nov., ♀, paratype (IDEA-LEPI-2020-024) A. Female genitalia, ventral view. B. Papillae analis. C. Ornamentation of the proximal part of the signum. D. Ornamentation of the distal part of the signum. Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4452227" figureDoi-1="http://doi.org/10.5281/zenodo.4452229" figureDoi-2="http://doi.org/10.5281/zenodo.4452231" figureDoi-3="http://doi.org/10.5281/zenodo.4452233" httpUri-0="https://zenodo.org/record/4452227/files/figure.png" httpUri-1="https://zenodo.org/record/4452229/files/figure.png" httpUri-2="https://zenodo.org/record/4452231/files/figure.png" httpUri-3="https://zenodo.org/record/4452233/files/figure.png" pageId="11" pageNumber="128">Figs 36</figureCitation>
, Table
<tableCitation id="C68E0310BB265E67602E63949E6C65FA" box="[866,876,459,485]" captionStart="Table 1" captionStartId="3.[189,258,265,291]" captionTargetBox="[200,1381,372,1967]" captionTargetPageId="3" captionText="Table 1. Character states coded for Helioandesia tarregai gen. et sp. nov. included in the data matrix of Hsu &amp; Powell (2004) for the cladistic analysis." httpUri="http://table.plazi.org/id/DF736623BB2E5E6F63F163569FE46559" pageId="11" pageNumber="128" tableUuid="DF736623BB2E5E6F63F163569FE46559">1</tableCitation>
<tableCitation id="C68E0310BB265E67603263949E8E65FA" box="[894,910,459,485]" captionStart="Table 2" captionStartId="4.[189,258,265,291]" captionTargetBox="[200,1381,334,838]" captionTargetPageId="4" captionText="Table 2. DNA barcode sequences used in the molecular analyses." httpUri="http://table.plazi.org/id/DF736623BB295E6863F163569EBD653C" pageId="11" pageNumber="128" tableUuid="DF736623BB295E6863F163569EBD653C">2</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3166520BB265E6763F160539C5B664A" box="[189,347,524,597]" pageId="11" pageNumber="128" type="diagnosis">
<paragraph id="8BB336ABBB265E6763F160539C366639" blockId="11.[189,310,524,550]" box="[189,310,524,550]" pageId="11" pageNumber="128">
<heading id="D0FB81C7BB265E6763F160539C366639" bold="true" box="[189,310,524,550]" fontSize="11" level="3" pageId="11" pageNumber="128" reason="3">
<emphasis id="B978EAB9BB265E6763F160539C366639" bold="true" box="[189,310,524,550]" pageId="11" pageNumber="128">Diagnosis</emphasis>
</heading>
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<paragraph id="8BB336ABBB265E6763F160649C5B664A" blockId="11.[189,347,571,597]" box="[189,347,571,597]" pageId="11" pageNumber="128">As for genus.</paragraph>
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<caption id="DF736623BB265E6763F1653B9E1A63F6" ID-DOI="http://doi.org/10.5281/zenodo.4452235" ID-Zenodo-Dep="4452235" httpUri="https://zenodo.org/record/4452235/files/figure.png" pageId="11" pageNumber="128" startId="11.[189,232,1892,1918]" targetBox="[189,1398,704,1852]" targetPageId="11">
<paragraph id="8BB336ABBB265E6763F1653B9E1A63F6" blockId="11.[187,1399,1892,2025]" pageId="11" pageNumber="128">
<emphasis id="B978EAB9BB265E6763F1653B9C086360" bold="true" box="[189,264,1892,1919]" pageId="11" pageNumber="128">Fig. 7.</emphasis>
Habitat and host plant of
<taxonomicName id="4C0C4D28BB265E676165653A9E2A6360" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[553,810,1893,1919]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="11" pageNumber="128" phylum="Arthropoda" rank="species" species="tarregai" status="gen. et sp. nov.">
<emphasis id="B978EAB9BB265E676165653A9E2A6360" box="[553,810,1893,1919]" italics="true" pageId="11" pageNumber="128">Helioandesia tarregai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB265E67607D653A9EDB6360" box="[817,987,1893,1919]" pageId="11" pageNumber="128" rank="species">gen. et sp. nov.</taxonomicNameLabel>
<emphasis id="B978EAB9BB265E676091653B9EF26361" bold="true" box="[989,1010,1892,1918]" pageId="11" pageNumber="128">A</emphasis>
. The type locality near Zapahuira Village at about 3400 m elevation on the western slopes of the Andes of the Parinacota Province, northern Chile.
<emphasis id="B978EAB9BB265E67624265F49C2263DA" bold="true" box="[270,290,1963,1989]" pageId="11" pageNumber="128">B</emphasis>
. The host plant
<taxonomicName id="4C0C4D28BB265E6762A765F4994763D9" authority="(Reiche) Heimerl" authorityName="Heimerl" authorityYear="1932" baseAuthorityName="Reiche" box="[491,1095,1963,1990]" class="Magnoliopsida" family="Nyctaginaceae" genus="Mirabilis" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="acuta">
<emphasis id="B978EAB9BB265E6762A765F49FA263DA" box="[491,674,1963,1989]" italics="true" pageId="11" pageNumber="128">Mirabilis acuta</emphasis>
(Reiche) Heimerl (Nyctaginaceae)
</taxonomicName>
.
<emphasis id="B978EAB9BB265E67671A65F4996963DA" bold="true" box="[1110,1129,1963,1989]" pageId="11" pageNumber="128">C</emphasis>
. Detail of the damage pattern on a leaf of the host.
<emphasis id="B978EAB9BB265E67614165919F2263F7" bold="true" box="[525,546,1998,2024]" pageId="11" pageNumber="128">D</emphasis>
. Flower of
<taxonomicName id="4C0C4D28BB265E6761E765909E1363F7" authorityName="Heimerl" authorityYear="1932" baseAuthorityName="Reiche" box="[683,787,1999,2024]" class="Magnoliopsida" family="Nyctaginaceae" genus="Mirabilis" kingdom="Plantae" order="Caryophyllales" pageId="11" pageNumber="128" phylum="Tracheophyta" rank="species" species="acuta">
<emphasis id="B978EAB9BB265E6761E765909E1363F7" box="[683,787,1999,2024]" italics="true" pageId="11" pageNumber="128">M. acuta</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="C3166520BB215E6063F163569CB36587" pageId="12" pageNumber="129" type="etymology">
<paragraph id="8BB336ABBB215E6063F163569C44653C" blockId="12.[189,324,265,291]" box="[189,324,265,291]" pageId="12" pageNumber="129">
<heading id="D0FB81C7BB215E6063F163569C44653C" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="12" pageNumber="129" reason="3">
<emphasis id="B978EAB9BB215E6063F163569C44653C" bold="true" box="[189,324,265,291]" pageId="12" pageNumber="129">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BB336ABBB215E6063F163689CB36587" blockId="12.[187,1398,311,408]" pageId="12" pageNumber="129">
<taxonomicName id="4C0C4D28BB215E6063F163689CBE654E" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[189,446,311,337]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="tarregai" status="gen. et sp. nov.">
<emphasis id="B978EAB9BB215E6063F163689CBE654E" box="[189,446,311,337]" italics="true" pageId="12" pageNumber="129">Helioandesia tarregai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB215E60628F63679F6D654D" box="[451,621,312,338]" pageId="12" pageNumber="129" rank="species">gen. et sp. nov.</taxonomicNameLabel>
is named in memory of the eminent Spanish guitarist and composer Francisco Tárrega, for his huge contribution to the repertory of the classical guitar, especially for the wonderful ʻMarietaʼ.
</paragraph>
</subSubSection>
<subSubSection id="C3166520BB215E6063F163E09ED2677C" pageId="12" pageNumber="129" type="materials_examined">
<paragraph id="8BB336ABBB215E6063F163E09CAA65C6" blockId="12.[189,426,447,473]" box="[189,426,447,473]" pageId="12" pageNumber="129">
<heading id="D0FB81C7BB215E6063F163E09CAA65C6" bold="true" box="[189,426,447,473]" fontSize="11" level="3" pageId="12" pageNumber="129" reason="3">
<emphasis id="B978EAB9BB215E6063F163E09CAA65C6" bold="true" box="[189,426,447,473]" pageId="12" pageNumber="129">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BB336ABBB215E6063A663B19C5C6617" blockId="12.[186,1399,494,626]" box="[234,348,494,520]" pageId="12" pageNumber="129">
<emphasis id="B978EAB9BB215E6063A663B19C5C6617" bold="true" box="[234,348,494,520]" pageId="12" pageNumber="129">
<typeStatus id="54B78809BB215E6063A663B19C5C6617" box="[234,348,494,520]" pageId="12" pageNumber="129" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BB336ABBB215E6063F1604E980A666D" blockId="12.[186,1399,494,626]" pageId="12" pageNumber="129">
<materialsCitation id="3B643CF6BB215E6063F1604E980A666D" ID-GBIF-Occurrence="3019938302" collectingDate="2019-03" collectingDateMax="2019-04" collectingDateMin="2019-03" collectorName="H. A. Vargas" country="Chile" elevation="3400" latitude="-18.329168" location="Parinacota Province" longLatPrecision="21" longitude="-69.58222" municipality="Zapahuira" pageId="12" pageNumber="129" specimenCode="IDEA-LEPI-2020-015" specimenCount="1" typeStatus="holotype">
<collectingCountry id="F31B763BBB215E6063F1604E9C136634" box="[189,275,529,555]" name="Chile" pageId="12" pageNumber="129">CHILE</collectingCountry>
• ♂;
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,
<collectingMunicipality id="6BD7ACD1BB215E606171604E9FB86634" box="[573,696,529,555]" pageId="12" pageNumber="129">Zapahuira</collectingMunicipality>
;
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,
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;
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<quantity id="4CF49B4EBB215E6060B4604E994E6634" box="[1016,1102,529,555]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="129" unit="m" value="3400.0">3400 m</quantity>
a.s.l.
</elevation>
; emerged
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<collectingDate id="EFF6E983BB215E6067B0604E98726634" box="[1276,1394,529,555]" pageId="12" pageNumber="129" value="2019-04">Apr. 2019</collectingDate>
</date>
;
<collectorName id="26F9537DBB215E6063F6606B9C486651" box="[186,328,564,590]" pageId="12" pageNumber="129">H.A. Vargas</collectorName>
leg.; ex-larva,
<taxonomicName id="4C0C4D28BB215E6062BB606B9E826651" authority="(Reiche) Heimerl" authorityName="Heimerl" authorityYear="1932" baseAuthorityName="Reiche" box="[503,898,564,591]" class="Magnoliopsida" family="Nyctaginaceae" genus="Mirabilis" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="129" phylum="Tracheophyta" rank="species" species="acuta">
<emphasis id="B978EAB9BB215E6062BB606B9FAB6651" box="[503,683,564,590]" italics="true" pageId="12" pageNumber="129">Mirabilis acuta</emphasis>
(Reiche) Heimerl
</taxonomicName>
; coll.
<date id="FFB2106BBB215E60608A606B99406651" box="[966,1088,564,590]" pageId="12" pageNumber="129" value="2019-03">
<collectingDate id="EFF6E983BB215E60608A606B99406651" box="[966,1088,564,590]" pageId="12" pageNumber="129" value="2019-03">Mar. 2019</collectingDate>
</date>
; “
<typeStatus id="54B78809BB215E606719606B99C56651" box="[1109,1221,564,590]" pageId="12" pageNumber="129" type="holotype">Holotype</typeStatus>
/
<taxonomicName id="4C0C4D28BB215E606795606B98756651" authorityName="Vargas" authorityYear="2021" box="[1241,1397,564,590]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="B978EAB9BB215E606795606B98756651" box="[1241,1397,564,590]" italics="true" pageId="12" pageNumber="129">Helioandesia</emphasis>
</taxonomicName>
/
<taxonomicName id="4C0C4D28BB215E60638060079C2B666E" authorityName="Vargas" authorityYear="2021" box="[204,299,600,625]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="tarregai">
<emphasis id="B978EAB9BB215E60638060079C2B666E" box="[204,299,600,625]" italics="true" pageId="12" pageNumber="129">tarregai</emphasis>
</taxonomicName>
/ Vargas” [red handwritten label]; genitalia slide HAV-1379;
<specimenCode id="DBAA9ED0BB215E6060B660089805666D" box="[1018,1285,599,626]" pageId="12" pageNumber="129">IDEA-LEPI-2020-015</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BB336ABBB215E6063A660C69C6766AC" blockId="12.[189,1399,665,867]" box="[234,359,665,691]" pageId="12" pageNumber="129">
<emphasis id="B978EAB9BB215E6063A660C69C6766AC" bold="true" box="[234,359,665,691]" pageId="12" pageNumber="129">
<typeStatus id="54B78809BB215E6063A660C69C6766AC" box="[234,359,665,691]" pageId="12" pageNumber="129" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8BB336ABBB215E6063F160E39ED2677C" blockId="12.[189,1399,665,867]" pageId="12" pageNumber="129">
<materialsCitation id="3B643CF6BB215E6063F160E39E1966E6" ID-GBIF-Occurrence="3019938303" collectingDate="2019-03" collectingDateMax="2019-04" collectingDateMin="2019-03" collectionCode="IDEA" collectorName="H. A. Vargas" country="Chile" elevation="3400" latitude="-18.329168" location="CHILE" longLatPrecision="21" longitude="-69.58222" municipality="Zapahuira" pageId="12" pageNumber="129" specimenCode="IDEA-LEPI-2020-016, IDEA- LEPI-2020-020" specimenCount="5" specimenCount-female="2" specimenCount-male="3" typeStatus="paratype">
<collectingCountry id="F31B763BBB215E6063F160E39C1366C9" box="[189,275,700,726]" name="Chile" pageId="12" pageNumber="129">CHILE</collectingCountry>
<specimenCount id="9D0AFD22BB215E60627160E39C8966C9" box="[317,393,700,726]" pageId="12" pageNumber="129" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D0AFD22BB215E6062D260E39CEA66C9" box="[414,490,700,726]" pageId="12" pageNumber="129" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype;
<specimenCode id="DBAA9ED0BB215E6060A560E499F166CA" box="[1001,1265,699,725]" pageId="12" pageNumber="129">IDEA-LEPI-2020-016</specimenCode>
to
<specimenCode id="DBAA9ED0BB215E60666B60E39C7966E6" pageId="12" pageNumber="129">IDEA- LEPI-2020-020</specimenCode>
; genitalia slide HAV-1374 female
</materialsCitation>
<materialsCitation id="3B643CF6BB215E60607060809ECE677C" ID-GBIF-Occurrence="3019938301" collectingDate="2018-04" collectingDateMax="2018-05" collectingDateMin="2018-04" collectionCode="IDEA" collectorName="H. A. Vargas" country="Chile" elevation="3400" latitude="-18.329168" location="Zapahuira" longLatPrecision="21" longitude="-69.58222" municipality="Zapahuira" pageId="12" pageNumber="129" specimenCode="IDEA-LEPI-2020-021, IDEA-LEPI-2020-024" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Parinacota Province" typeStatus="paratype">
<specimenCount id="9D0AFD22BB215E60607060809E8566E6" box="[828,901,735,761]" pageId="12" pageNumber="129" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="9D0AFD22BB215E6060DB60809EDF66E6" box="[919,991,735,761]" pageId="12" pageNumber="129" type="female">2 ♀♀</specimenCount>
;
<collectingRegion id="49C8F849BB215E6060BF608099E666E6" box="[1011,1254,735,761]" pageId="12" pageNumber="129">Parinacota Province</collectingRegion>
,
<location id="8ED36070BB215E6067BB6080987266E6" LSID="urn:lsid:plazi:treatment:03A587BDBB265E61617763DB9EA060CD:8ED36070BB215E6067BB6080987266E6" box="[1271,1394,735,761]" country="Chile" latitude="-18.329168" longLatPrecision="21" longitude="-69.58222" municipality="Zapahuira" name="Zapahuira" pageId="12" pageNumber="129" stateProvince="Parinacota Province">Zapahuira</location>
;
<geoCoordinate id="EE38506CBB215E6063F1615D9C526703" box="[189,338,768,797]" degrees="18" direction="south" minutes="19" orientation="latitude" pageId="12" pageNumber="129" precision="15" seconds="45" value="-18.329168">18°1945″ S</geoCoordinate>
;
<geoCoordinate id="EE38506CBB215E60622E615D9F016702" box="[354,513,768,797]" degrees="69" direction="west" minutes="34" orientation="longitude" pageId="12" pageNumber="129" precision="15" seconds="56" value="-69.58222">69°3456″ W</geoCoordinate>
;
<elevation id="0021D198BB215E60615D615D9FB26703" box="[529,690,770,797]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="129" unit="m" value="3400.0">
<quantity id="4CF49B4EBB215E60615D615D9F6E6702" box="[529,622,770,797]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="129" unit="m" value="3400.0">3400 m</quantity>
a.s.l.
</elevation>
; emerged
<date id="FFB2106BBB215E60607E615C9EB16703" box="[818,945,770,797]" pageId="12" pageNumber="129" value="2018-05">
<collectingDate id="EFF6E983BB215E60607E615C9EB16703" box="[818,945,770,797]" pageId="12" pageNumber="129" value="2018-05">May 2018</collectingDate>
</date>
;
<collectorName id="26F9537DBB215E60608D615D99546702" box="[961,1108,770,797]" pageId="12" pageNumber="129">H.A. Vargas</collectorName>
leg.; ex-larva
<taxonomicName id="4C0C4D28BB215E606645615D9CE06720" authority="(Reiche) Heimerl" authorityName="Heimerl" authorityYear="1932" baseAuthorityName="Reiche" class="Magnoliopsida" family="Nyctaginaceae" genus="Mirabilis" kingdom="Plantae" order="Caryophyllales" pageId="12" pageNumber="129" phylum="Tracheophyta" rank="species" species="acuta">
<emphasis id="B978EAB9BB215E606645615D9DFE6720" italics="true" pageId="12" pageNumber="129">Mirabilis acuta</emphasis>
(Reiche) Heimerl
</taxonomicName>
; coll.
<date id="FFB2106BBB215E606163617A9FAC6720" box="[559,684,805,831]" pageId="12" pageNumber="129" value="2018-04">
<collectingDate id="EFF6E983BB215E606163617A9FAC6720" box="[559,684,805,831]" pageId="12" pageNumber="129" value="2018-04">Apr. 2018</collectingDate>
</date>
;
<specimenCode id="DBAA9ED0BB215E6061F1617A9EC66720" box="[701,966,805,831]" pageId="12" pageNumber="129">IDEA-LEPI-2020-021</specimenCode>
to
<specimenCode id="DBAA9ED0BB215E6060B9617A98006720" box="[1013,1280,805,831]" pageId="12" pageNumber="129">IDEA-LEPI-2020-024</specimenCode>
; genitalia slides HAV-1371 male, HAV-1372 female, HAV-1373 male;
<collectionCode id="ED1DAE6EBB215E6060C861169ECE677C" box="[900,974,841,867]" collectionName="Chile, Arica, Universidad de Tarapaca, Instituto de Agronomia" country="Chile" httpUri="http://grbio.org/cool/t1d4-ec1s" name="Instituto de Agronomia" pageId="12" pageNumber="129">IDEA</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3166520BB215E6163F161D59FAC6779" lastPageId="13" lastPageNumber="130" pageId="12" pageNumber="129" type="description">
<paragraph id="8BB336ABBB215E6063F161D59C4E67BB" blockId="12.[189,334,906,932]" box="[189,334,906,932]" pageId="12" pageNumber="129">
<heading id="D0FB81C7BB215E6063F161D59C4E67BB" bold="true" box="[189,334,906,932]" fontSize="11" level="3" pageId="12" pageNumber="129" reason="3">
<emphasis id="B978EAB9BB215E6063F161D59C4E67BB" bold="true" box="[189,334,906,932]" pageId="12" pageNumber="129">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BB336ABBB215E6063A661E69C2A67CC" blockId="12.[189,694,953,1014]" box="[234,298,953,979]" pageId="12" pageNumber="129">
<emphasis id="B978EAB9BB215E6063A661E69C2A67CC" bold="true" box="[234,298,953,979]" pageId="12" pageNumber="129">Male</emphasis>
</paragraph>
<paragraph id="8BB336ABBB215E6063F161839FB667E9" blockId="12.[189,694,953,1014]" box="[189,694,987,1014]" pageId="12" pageNumber="129">
Forewing length
<quantity id="4CF49B4EBB215E6062C961849F1667E9" box="[389,534,987,1014]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" metricValueMax="6.1" metricValueMin="5.5" pageId="12" pageNumber="129" unit="mm" value="5.8" valueMax="6.1" valueMin="5.5">5.56.1 mm</quantity>
. (
<figureCitation id="13372A2EBB215E60616661839F6E67E9" box="[554,622,988,1014]" captionStart="Fig" captionStartId="7.[189,232,1928,1954]" captionTargetBox="[189,1398,757,1889]" captionTargetId="figure-163@7.[189,1398,757,1889]" captionTargetPageId="7" captionText="Fig. 3. Helioandesia tarregai gen. et sp. nov., ♂, holotype (IDEA-LEPI-2020-015). A. Dorsal view. B. Head, lateral. C. Basal flagellomeres of right antenna. D. Three bulging metallic spots on the basal half of forewing. E. Hair pencil from base of subcosta, ventral. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452227" httpUri="https://zenodo.org/record/4452227/files/figure.png" pageId="12" pageNumber="129">Fig. 3</figureCitation>
AE).
</paragraph>
<paragraph id="8BB336ABBB215E6063F166419C2860BE" blockId="12.[188,1398,1053,1185]" pageId="12" pageNumber="129">HEAD. Vertex and frons silvery metallic. Pale orange yellow scales behind compound eyes. Antenna with alternate rings of creamy white and brownish gray on basal two third, distal third brownish gray. Labial palpus creamy white on two basal segments; brownish gray scales with creamy white margin on third segment.</paragraph>
<caption id="DF736623BB215E6063F16541983163F6" ID-DOI="http://doi.org/10.5281/zenodo.4452237" ID-Zenodo-Dep="4452237" httpUri="https://zenodo.org/record/4452237/files/figure.png" pageId="12" pageNumber="129" startId="12.[189,232,1822,1848]" targetBox="[189,1398,1242,1784]" targetPageId="12">
<paragraph id="8BB336ABBB215E6063F16541983163F6" blockId="12.[189,1399,1822,2025]" pageId="12" pageNumber="129">
<emphasis id="B978EAB9BB215E6063F165419C0C6327" bold="true" box="[189,268,1822,1848]" pageId="12" pageNumber="129">Fig. 8.</emphasis>
Geographic distribution of three Neotropical
<taxonomicName id="4C0C4D28BB215E60607F65419ECE6327" authorityName="Heinemann" authorityYear="1877" box="[819,974,1822,1848]" class="Insecta" family="Heliodinidae" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="family">Heliodinidae</taxonomicName>
. Red rectangle on left indicates the location in South America of the area enlarged on right. Circle = type locality of
<taxonomicName id="4C0C4D28BB215E606738651E98776344" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[1140,1399,1857,1883]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="tarregai" status="gen. et sp. nov.">
<emphasis id="B978EAB9BB215E606738651E98776344" box="[1140,1399,1857,1883]" italics="true" pageId="12" pageNumber="129">Helioandesia tarregai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB215E6063F1653A9C756360" box="[189,373,1893,1919]" pageId="12" pageNumber="129" rank="species">gen. et sp. nov.</taxonomicNameLabel>
in northern Chile. Triangles = distribution of
<taxonomicName id="4C0C4D28BB215E6060E0653A98446360" authority="(Meyrick, 1917)" baseAuthorityName="Meyrick" baseAuthorityYear="1917" box="[940,1348,1893,1919]" class="Insecta" family="Heliodinidae" genus="Aetole" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="demarcha">
<emphasis id="B978EAB9BB215E6060E0653A99756360" box="[940,1141,1893,1919]" italics="true" pageId="12" pageNumber="129">Aetole demarcha</emphasis>
(
<bibRefCitation id="EF9D4B5ABB215E6067CB653A983B6360" author="Meyrick E." box="[1159,1339,1893,1919]" pageId="12" pageNumber="129" pagination="61 - 66" refId="ref7150" refString="Meyrick E. 1917. Heliodinidae. Exotic Microlepidoptera 2: 61 - 66." type="journal article" year="1917">Meyrick, 1917</bibRefCitation>
)
</taxonomicName>
, the geographically nearest heliodinid species to
<taxonomicName id="4C0C4D28BB215E60619E65D69E5863BD" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[722,856,1929,1954]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="tarregai" status="gen. et sp. nov.">
<emphasis id="B978EAB9BB215E60619E65D69E5863BD" box="[722,856,1929,1954]" italics="true" pageId="12" pageNumber="129">H. tarregai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB215E60602E65D7991963BD" box="[866,1049,1928,1954]" pageId="12" pageNumber="129" rank="species">gen. et sp. nov.</taxonomicNameLabel>
on the western slopes of the Andes, in Peru. Square = type locality of
<taxonomicName id="4C0C4D28BB215E60618765F499C163DA" authority="(Meyrick, 1932)" baseAuthorityName="Meyrick" baseAuthorityYear="1932" box="[715,1217,1963,1989]" class="Insecta" family="Heliodinidae" genus="Lithariapteryx" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="loriculata">
<emphasis id="B978EAB9BB215E60618765F49EF363DA" box="[715,1011,1963,1989]" italics="true" pageId="12" pageNumber="129">Lithariapteryx loriculata</emphasis>
(Meyrick, 1932)
</taxonomicName>
on the eastern slopes of the Andes of Bolivia, the morphologically nearest species to
<taxonomicName id="4C0C4D28BB215E6060BB6590997A63F7" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[1015,1146,1999,2024]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="12" pageNumber="129" phylum="Arthropoda" rank="species" species="tarregai" status="gen. et sp. nov.">
<emphasis id="B978EAB9BB215E6060BB6590997A63F7" box="[1015,1146,1999,2024]" italics="true" pageId="12" pageNumber="129">H. tarregai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB215E6067CD6590983163F6" box="[1153,1329,1999,2025]" pageId="12" pageNumber="129" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BB336ABBB205E6163F163569E8E6606" blockId="13.[186,1398,265,538]" pageId="13" pageNumber="130">THORAX. Variable proportion of silvery metallic and brownish gray with creamy white margin scales dorsally, brownish gray scales with wider creamy white margin latero-ventrally. Legs mainly concolorous with latero-ventral side of thorax. Forewing mostly covered by brownish gray with creamy white margin scales; five metallic silvery bulging spots with black proximal margin, three near costal margin, size progressively decreasing from base to apex, two near anal margin; small metallic silvery spot near tegula, slightly differentiated metallic silvery diagonal stripe from about middle of distal margin of discal cell to near costal margin; a few brownish orange scattered scales mainly on distal half; fringe brownish gray. Hindwing brownish gray; fringe concolorous.</paragraph>
<paragraph id="8BB336ABBB205E6163F1601E9FA06661" blockId="13.[187,1398,577,639]" pageId="13" pageNumber="130">ABDOMEN. Mainly brownish gray dorsally, lighter near posterior margin of each segment; scales brownish gray with creamy white margin ventrally.</paragraph>
<paragraph id="8BB336ABBB205E6163F160F49FAC66DF" blockId="13.[189,684,677,704]" box="[189,684,677,704]" pageId="13" pageNumber="130">
GENITALIA (
<figureCitation id="13372A2EBB205E61627660FA9C8266DF" box="[314,386,677,704]" captionStart="Fig" captionStartId="9.[189,232,1857,1883]" captionTargetBox="[327,1261,319,1818]" captionTargetId="figure-11@9.[326,1261,319,1818]" captionTargetPageId="9" captionText="Fig. 5. Male genitalia and eighth segment of Helioandesia tarregai gen. et sp. nov., ♂, paratype (IDEA- LEPI-2020-021). A. Male genitalia, ventral view, phallus removed. B. Phallus, lateral view. C. Tergum and pleural lobes of eighth segment. D. Socius, closed arrow in A. E. Small spine-like cornuti on the vesica, open arrow in A. F. Medial part of the cornutus. G. Coremata between pleural lobes of segment VIII and genitalia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452231" httpUri="https://zenodo.org/record/4452231/files/figure.png" pageId="13" pageNumber="130">Fig. 5</figureCitation>
). As described for genus.
</paragraph>
<paragraph id="8BB336ABBB205E6163A660B89C7A673B" blockId="13.[189,378,743,805]" pageId="13" pageNumber="130">
<emphasis id="B978EAB9BB205E6163A660B89C45671E" bold="true" box="[234,325,743,769]" pageId="13" pageNumber="130">Female</emphasis>
Similar to male.
</paragraph>
<paragraph id="8BB336ABBB205E6163F1610E9FAC6779" blockId="13.[189,684,843,870]" box="[189,684,843,870]" pageId="13" pageNumber="130">
GENITALIA (
<figureCitation id="13372A2EBB205E61627661149C826779" box="[314,386,843,870]" captionStart="Fig" captionStartId="10.[191,234,1927,1953]" captionTargetBox="[189,1398,559,1861]" captionTargetId="figure-92@10.[189,1398,559,1861]" captionTargetPageId="10" captionText="Fig. 6. Female genitalia of Helioandesia tarregai gen. et sp. nov., ♀, paratype (IDEA-LEPI-2020-024) A. Female genitalia, ventral view. B. Papillae analis. C. Ornamentation of the proximal part of the signum. D. Ornamentation of the distal part of the signum. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.4452233" httpUri="https://zenodo.org/record/4452233/files/figure.png" pageId="13" pageNumber="130">Fig. 6</figureCitation>
). As described for genus.
</paragraph>
</subSubSection>
<subSubSection id="C3166520BB205E6163F161D29CA46020" pageId="13" pageNumber="130" type="biology_ecology">
<paragraph id="8BB336ABBB205E6163F161D29C4067B8" blockId="13.[189,320,909,935]" box="[189,320,909,935]" pageId="13" pageNumber="130">
<heading id="D0FB81C7BB205E6163F161D29C4067B8" bold="true" box="[189,320,909,935]" fontSize="11" level="3" pageId="13" pageNumber="130" reason="3">
<emphasis id="B978EAB9BB205E6163F161D29C4067B8" bold="true" box="[189,320,909,935]" pageId="13" pageNumber="130">Host plant</emphasis>
</heading>
</paragraph>
<paragraph id="8BB336ABBB205E6163F161E49CA46020" blockId="13.[189,1399,955,1087]" pageId="13" pageNumber="130">
The only known host plant is the native
<taxonomicName id="4C0C4D28BB205E6161E961E4992567C9" authorityName="Heimerl" authorityYear="1932" baseAuthorityName="Reiche" box="[677,1061,955,982]" class="Magnoliopsida" family="Nyctaginaceae" genus="Mirabilis" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="130" phylum="Tracheophyta" rank="species" species="acuta">
<emphasis id="B978EAB9BB205E6161E961E49E5D67CA" box="[677,861,955,981]" italics="true" pageId="13" pageNumber="130">Mirabilis acuta</emphasis>
(Nyctaginaceae)
</taxonomicName>
(
<figureCitation id="13372A2EBB205E61677A61E4998067C9" box="[1078,1152,955,982]" captionStart="Fig" captionStartId="11.[189,232,1892,1918]" captionTargetBox="[189,1398,704,1852]" captionTargetId="figure-49@11.[189,1398,704,1852]" captionTargetPageId="11" captionText="Fig. 7. Habitat and host plant ofHelioandesia tarregai gen. et sp. nov.A. The type locality near Zapahuira Village at about 3400 m elevation on the western slopes of theAndes of the Parinacota Province, northern Chile. B. The host plant Mirabilis acuta (Reiche) Heimerl (Nyctaginaceae). C. Detail of the damage pattern on a leaf of the host. D. Flower of M. acuta." figureDoi="http://doi.org/10.5281/zenodo.4452235" httpUri="https://zenodo.org/record/4452235/files/figure.png" pageId="13" pageNumber="130">Fig. 7</figureCitation>
), a Chilean endemic (
<bibRefCitation id="EF9D4B5ABB205E61638B61809CC867E6" author="Rodriguez R. &amp; Marticorena C. &amp; Alarcon D. &amp; Baeza C. &amp; Cavieres L. &amp; Finot V. L. &amp; Fuentes N. &amp; Kiessling A. &amp; Mihoc M. &amp; Pauchard A. &amp; Ruiz E. &amp; Sanchez P. &amp; Marticorena A." box="[199,456,991,1017]" pageId="13" pageNumber="130" pagination="1 - 430" refId="ref7763" refString="Rodriguez R., Marticorena C., Alarcon D., Baeza C., Cavieres L., Finot V. L., Fuentes N., Kiessling A., Mihoc M., Pauchard A., Ruiz E., Sanchez P. &amp; Marticorena A. 2018. Catalogo de las plantas vasculares de Chile. Gayana Botanica 75: 1 - 430." type="journal article" year="2018">
Rodríguez
<emphasis id="B978EAB9BB205E61620461809C8267E6" box="[328,386,991,1017]" italics="true" pageId="13" pageNumber="130">et al.</emphasis>
2018
</bibRefCitation>
). In the study zone
<taxonomicName id="4C0C4D28BB205E6161EB61809E0E67E7" authorityName="Heimerl" authorityYear="1932" baseAuthorityName="Reiche" box="[679,782,991,1016]" class="Magnoliopsida" family="Nyctaginaceae" genus="Mirabilis" kingdom="Plantae" order="Caryophyllales" pageId="13" pageNumber="130" phylum="Tracheophyta" rank="species" species="acuta">
<emphasis id="B978EAB9BB205E6161EB61809E0E67E7" box="[679,782,991,1016]" italics="true" pageId="13" pageNumber="130">M. acuta</emphasis>
</taxonomicName>
sprout and flower after the summer rains. The larvae of
<taxonomicName id="4C0C4D28BB205E616390665C9C5E6003" authority="Vargas, 2021" authorityName="Vargas" authorityYear="2021" box="[220,350,1027,1052]" class="Insecta" family="Heliodinidae" genus="Helioandesia" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="130" phylum="Arthropoda" rank="species" species="tarregai" status="gen. et sp. nov.">
<emphasis id="B978EAB9BB205E616390665C9C5E6003" box="[220,350,1027,1052]" italics="true" pageId="13" pageNumber="130">H. tarregai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24B57C2BB205E616229665D9F146003" box="[357,532,1026,1052]" pageId="13" pageNumber="130" rank="species">gen. et sp. nov.</taxonomicNameLabel>
feed externally as leaf skeletonizers, remaining partially covered by a fine silk layer (
<figureCitation id="13372A2EBB205E616275667A9C966020" box="[313,406,1061,1087]" captionStart="Fig" captionStartId="11.[189,232,1892,1918]" captionTargetBox="[189,1398,704,1852]" captionTargetId="figure-49@11.[189,1398,704,1852]" captionTargetPageId="11" captionText="Fig. 7. Habitat and host plant ofHelioandesia tarregai gen. et sp. nov.A. The type locality near Zapahuira Village at about 3400 m elevation on the western slopes of theAndes of the Parinacota Province, northern Chile. B. The host plant Mirabilis acuta (Reiche) Heimerl (Nyctaginaceae). C. Detail of the damage pattern on a leaf of the host. D. Flower of M. acuta." figureDoi="http://doi.org/10.5281/zenodo.4452235" httpUri="https://zenodo.org/record/4452235/files/figure.png" pageId="13" pageNumber="130">Fig. 7C</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3166520BB205E6163F166399EA060CD" pageId="13" pageNumber="130" type="distribution">
<paragraph id="8BB336ABBB205E6163F166399C56609F" blockId="13.[189,342,1126,1152]" box="[189,342,1126,1152]" pageId="13" pageNumber="130">
<heading id="D0FB81C7BB205E6163F166399C56609F" bold="true" box="[189,342,1126,1152]" fontSize="11" level="3" pageId="13" pageNumber="130" reason="3">
<emphasis id="B978EAB9BB205E6163F166399C56609F" bold="true" box="[189,342,1126,1152]" pageId="13" pageNumber="130">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BB336ABBB205E6163F166C99EA060CD" blockId="13.[189,1399,1173,1235]" pageId="13" pageNumber="130">
Known only from the
<typeStatus id="54B78809BB205E61629E66C99F0560AF" box="[466,517,1174,1200]" pageId="13" pageNumber="130">type</typeStatus>
locality (
<figureCitation id="13372A2EBB205E61613066CA9FC760B0" box="[636,711,1173,1199]" captionStart="Fig" captionStartId="12.[189,232,1822,1848]" captionTargetBox="[189,1398,1242,1784]" captionTargetId="figure-327@12.[189,1398,1242,1784]" captionTargetPageId="12" captionText="Fig. 8. Geographic distribution of three Neotropical Heliodinidae. Red rectangle on left indicates the location in South America of the area enlarged on right. Circle = type locality of Helioandesia tarregai gen. et sp. nov. in northern Chile. Triangles = distribution of Aetole demarcha (Meyrick, 1917), the geographically nearest heliodinid species to H. tarregai gen. et sp. nov. on the western slopes of the Andes, in Peru. Square = type locality of Lithariapteryx loriculata (Meyrick, 1932) on the eastern slopes of the Andes of Bolivia, the morphologically nearest species to H. tarregai gen. et sp. nov." figureDoi="http://doi.org/10.5281/zenodo.4452237" httpUri="https://zenodo.org/record/4452237/files/figure.png" pageId="13" pageNumber="130">Fig. 8</figureCitation>
), near Zapahuira, Parinacota Province, at about
<quantity id="4CF49B4EBB205E61665566CA987760AF" box="[1305,1399,1173,1200]" metricMagnitude="3" metricUnit="m" metricValue="3.4" pageId="13" pageNumber="130" unit="m" value="3400.0">3400 m</quantity>
elevation on the western slopes of the Andes of northern
<collectingCountry id="F31B763BBB205E61601566E79E9C60CD" box="[857,924,1208,1234]" name="Chile" pageId="13" pageNumber="130">Chile</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>