450 lines
66 KiB
XML
450 lines
66 KiB
XML
<document id="626EC8EF1DCA0D35222F36E3D4169654" ID-DOI="10.11646/zootaxa.3872.3.3" ID-GBIF-Dataset="457f0311-e092-4e32-9a42-0e0b83a8dac5" ID-ISSN="1175-5326" ID-Zenodo-Dep="228599" ID-ZooBank="3B4F0F36-2469-4504-9021-AD7C9263E379" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460927101523" checkinUser="plazi" docAuthor="Janák, Jiří & Bordoni, Arnaldo" docDate="2014" docId="03A51E7737573875FF5EBFA6FD85FC26" docLanguage="en" docName="zt03872p274.pdf" docOrigin="Zootaxa 3872 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Achmonia Bordoni 2004" docType="treatment" docVersion="8" lastPageNumber="262" masterDocId="FF9C660F37553870FFC9BE36FFEAFFE1" masterDocTitle="Revision of the genus Achmonia of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="274" masterPageNumber="257" pageNumber="259" updateTime="1698362774903" updateUser="plazi">
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<mods:titleInfo id="714BC140E739E2732839E8F33A7C0AC7">
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<mods:title id="89B721882C0D1259CD3DE50CD37098BB">Revision of the genus Achmonia of Africa south of the Sahara (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="0F4C79A4534C6CA049726745810E07DD">Janák, Jiří</mods:namePart>
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<mods:namePart id="D2A94E653C71DDE3A6FF39EC994BB4F6">Bordoni, Arnaldo</mods:namePart>
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<mods:title id="B8E4679B81AA30948AC789BDCCD8AF52">Zootaxa</mods:title>
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<mods:part id="8803038AB8C5785FA29A21A5A4D0F264">
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<mods:date id="2916B1BDE71B201DF27334AB39AC0115">2014</mods:date>
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<mods:number id="48A2A6250FCCA916954D755DBCA2E35E">3872</mods:number>
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<treatment id="03A51E7737573875FF5EBFA6FD85FC26" ID-DOI="http://doi.org/10.5281/zenodo.6142453" ID-GBIF-Taxon="119589606" ID-Zenodo-Dep="6142453" LSID="urn:lsid:plazi:treatment:03A51E7737573875FF5EBFA6FD85FC26" httpUri="http://treatment.plazi.org/id/03A51E7737573875FF5EBFA6FD85FC26" lastPageId="5" lastPageNumber="262" pageId="2" pageNumber="259">
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<subSubSection id="C316FCEA37573872FF5EBFA6FE26FE4A" box="[151,460,400,427]" pageId="2" pageNumber="259" type="nomenclature">
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<paragraph id="8BB3AF6137573872FF5EBFA6FE26FE4A" blockId="2.[151,460,400,427]" box="[151,460,400,427]" pageId="2" pageNumber="259">
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<heading id="D0FB180D37573872FF5EBFA6FE26FE4A" bold="true" box="[151,460,400,427]" fontSize="11" level="1" pageId="2" pageNumber="259" reason="1">
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<taxonomicName id="4C0CD4E237573872FF5EBFA6FE26FE4A" ID-CoL="8XJ92" authority="Bordoni, 2004" authorityName="Bordoni" authorityYear="2004" box="[151,460,400,427]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="259" phylum="Arthropoda" rank="genus">
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<emphasis id="B978737337573872FF5EBFA6FE26FE4A" bold="true" box="[151,460,400,427]" pageId="2" pageNumber="259">
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<emphasis id="B978737337573872FF5EBFA6FEFEFE4B" bold="true" box="[151,276,400,426]" italics="true" pageId="2" pageNumber="259">Achmonia</emphasis>
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Bordoni, 2004
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C316FCEA37573872FF5EBFEEFCE0FD74" pageId="2" pageNumber="259" type="reference_group">
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<paragraph id="8BB3AF6137573872FF5EBFEEFD5CFE0F" blockId="2.[151,1251,472,555]" box="[151,694,472,494]" pageId="2" pageNumber="259">
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<treatmentCitationGroup id="AB1C884F37573872FF5EBFEEFD5CFE0F" box="[151,694,472,494]" pageId="2" pageNumber="259">
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<taxonomicName id="4C0CD4E237573872FF5EBFEEFE21FE0F" ID-CoL="8XJ92" authority="Bordoni, 2004a: 57" authorityName="Bordoni" authorityPageNumber="57" authorityYear="2004" box="[151,459,472,494]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="259" phylum="Arthropoda" rank="genus">
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<emphasis id="B978737337573872FF5EBFEEFF14FE0F" box="[151,254,472,494]" italics="true" pageId="2" pageNumber="259">Achmonia</emphasis>
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<treatmentCitation id="0AAD897037573872FECDBFEEFE21FE0F" author="Bordoni" box="[260,459,472,494]" page="57" pageId="2" pageNumber="259" year="2004">
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<bibRefCitation id="EF9DD29037573872FECDBFEEFE4FFE0F" author="Bordoni" box="[260,421,472,494]" pageId="2" pageNumber="259" refString="Bordoni, A. (2004 a) Nuovi dati per la conoscenza degli Xantholinini della Regione Orientale. II. Un nuovo genere di Giava (Coleoptera, Staphylinidae). Fragmenta Entomologica, Roma, 36 (1), 57 - 62." type="journal article" year="2004" yearSuffix="a">Bordoni, 2004a</bibRefCitation>
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: 57
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</treatmentCitation>
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</taxonomicName>
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; 2012: 107; 2013: 218.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BB3AF6137573872FF5EBFC0FD83FDCA" blockId="2.[151,1251,472,555]" pageId="2" pageNumber="259">
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<treatmentCitationGroup id="AB1C884F37573872FF5EBFC0FD83FDCA" pageId="2" pageNumber="259">
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<taxonomicName id="4C0CD4E237573872FF5EBFC0FE45FDED" ID-CoL="8XKS8" authority="Bordoni, 2004b: 84" authorityName="Bordoni" authorityPageNumber="84" authorityYear="2004" box="[151,431,502,524]" class="Insecta" family="Staphylinidae" genus="Daolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="259" phylum="Arthropoda" rank="genus">
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<emphasis id="B978737337573872FF5EBFC0FF0AFDED" box="[151,224,502,524]" italics="true" pageId="2" pageNumber="259">Daolus</emphasis>
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<treatmentCitation id="0AAD897037573872FF2FBFC0FE45FDED" author="Bordoni" box="[230,431,502,524]" page="84" pageId="2" pageNumber="259" year="2004">
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<bibRefCitation id="EF9DD29037573872FF2FBFC0FE63FDED" author="Bordoni" box="[230,393,502,524]" pageId="2" pageNumber="259" refString="Bordoni, A. (2004 b) Nuovi dati per la conoscenza degli Xantholinini della Regione Orientale. V. Daolus hromadkai gen. n., sp. n. del Nepal e Thyreocephalus perakensis sp. n. della Malesia (Coleoptera Staphylinidae). Entomologica, Bari, 38, 83 - 89." type="journal article" year="2004" yearSuffix="b">Bordoni, 2004b</bibRefCitation>
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: 84
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</treatmentCitation>
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</taxonomicName>
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; 2008: 60; 2009: 1867; 2010: 57; 2012: 107; 2013: 212 (synonym of
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<taxonomicName id="4C0CD4E237573872FBA4BFC0FB39FDED" ID-CoL="8XJ92" box="[1133,1235,502,524]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="259" phylum="Arthropoda" rank="genus">
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<emphasis id="B978737337573872FBA4BFC0FB39FDED" box="[1133,1235,502,524]" italics="true" pageId="2" pageNumber="259">Achmonia</emphasis>
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</taxonomicName>
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).
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<emphasis id="B978737337573872FF5EBC23FECFFDCA" bold="true" box="[151,293,533,555]" pageId="2" pageNumber="259">
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<typeStatus id="54B711C337573872FF5EBC23FF26FDCA" box="[151,204,533,555]" pageId="2" pageNumber="259">Type</typeStatus>
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species:
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</emphasis>
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<taxonomicName id="4C0CD4E237573872FEE2BC23FD83FDCA" ID-CoL="9872" authority="Bordoni, 2004" authorityName="Bordoni" authorityYear="2004" box="[299,617,533,555]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="259" phylum="Arthropoda" rank="species" species="doloc">
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<emphasis id="B978737337573872FEE2BC23FE3AFDCA" box="[299,464,533,555]" italics="true" pageId="2" pageNumber="259">Achmonia doloc</emphasis>
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Bordoni, 2004
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</paragraph>
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<paragraph id="8BB3AF6137573872FF5EBC61FCE0FD74" blockId="2.[151,1437,599,1993]" pageId="2" pageNumber="259">
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The original description of the genus was based only on the
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<typeStatus id="54B711C337573872FC4DBC6EFC5EFD91" box="[900,948,600,624]" pageId="2" pageNumber="259">type</typeStatus>
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species (
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<bibRefCitation id="EF9DD29037573872FBE1BC61FB37FD91" author="Bordoni" box="[1064,1245,599,624]" pageId="2" pageNumber="259" refString="Bordoni, A. (2004 a) Nuovi dati per la conoscenza degli Xantholinini della Regione Orientale. II. Un nuovo genere di Giava (Coleoptera, Staphylinidae). Fragmenta Entomologica, Roma, 36 (1), 57 - 62." type="journal article" year="2004" yearSuffix="a">Bordoni 2004a</bibRefCitation>
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). The following redescription is extended for all hitherto known species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C316FCEA37573872FF0EBCA9FAB3F911" pageId="2" pageNumber="259" type="description">
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<paragraph id="8BB3AF6137573872FF0EBCA9FBEEFD59" blockId="2.[151,1437,599,1993]" box="[199,1028,671,696]" pageId="2" pageNumber="259">
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<emphasis id="B978737337573872FF0EBCA9FE9EFD59" bold="true" box="[199,372,671,696]" pageId="2" pageNumber="259">Redescription.</emphasis>
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Form elongate, stout, of medium to large size (
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<quantity id="4CF4028437573872FC4EBC96FC12FD59" box="[903,1016,672,696]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" metricValueMax="2.0" metricValueMin="0.9" pageId="2" pageNumber="259" unit="mm" value="14.5" valueMax="20.0" valueMin="9.0">9–20 mm</quantity>
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).
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</paragraph>
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<paragraph id="8BB3AF6137573872FF0EBCF2FB94FB6C" blockId="2.[151,1437,599,1993]" pageId="2" pageNumber="259">
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Head scarcely punctate, with or without coalescent grooves; frontal grooves short or moderately long, reaching at most to the middle of eyes, more or less convergent posteromedially, ocular grooves very superficial. Anterior margin of frons between antennal insertion extended into short and very wide, apically truncate process, limited on each side by rounded emargination, and convex or slightly impressed dorsally. Eyes small, temples considerably longer than length of eyes as seen above, evenly rounded, posterior angles rounded or acute. Antennae geniculate, moderately short, antennal insertions separated from each other by distance about equal to distance separating each insertion from anteromedian margin of eye; first segment long, thickened towards apex, equal in length to at least four following segments combined; second segment shorter than third, outer segments more or less transverse. Labrum dark, completely chitinised, short, transverse, more or less distinctly quadrilobate (
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<figureCitation id="1337B3E437573872FB5CBDD2FAF4FC1D" box="[1173,1310,996,1021]" captionStart="FIGURES 13 – 24. 13, 14, 21, 22" captionStartId="6.[151,264,1465,1487]" captionTargetBox="[163,1431,203,1425]" captionTargetId="figure@6.[151,1436,193,1444]" captionTargetPageId="6" captionText="FIGURES 13 – 24. 13, 14, 21, 22, Achmonia doloc (Bord.), HT from Java; 15, A. eppelsheimi (Bernh. & Schub.) from Nepal; 16, A. shibatai (Bord.), HT from Taiwan; 17, A. nigra (Bord.), HT from China; 18, 23, 24, A. amabilis (Boh.) from South Africa; 19, A. simulator sp. nov., PT from Mposhi; 20, A. carinata sp. nov., HT. 13, head and pronotum; 14 – 20, labrum; 21, 23, maxillary palpus; 22, 24, labial palpus. 13, 14, 21, 22 from Bordoni (2004 a), 15 from Bordoni (2004 b), 16 from Bordoni (2010), 17 from Bordoni (2009). Scale 0.1 mm; 13, 15 and 17 without scale." httpUri="https://zenodo.org/record/228601/files/figure.png" pageId="2" pageNumber="259">Figs. 14–20</figureCitation>
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), with long and strong apical setae. Mandible strong, lateral furrow absent or reduced to vague impression at base. Maxillary palpi moderately long, segment 3 shorter than segment 2, segment 4 longer and somewhat narrower than segment 3, subacute apically (
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<figureCitation id="1337B3E437573872FE4EBA79FDFBFB89" box="[391,529,1103,1128]" captionStart="FIGURES 13 – 24. 13, 14, 21, 22" captionStartId="6.[151,264,1465,1487]" captionTargetBox="[163,1431,203,1425]" captionTargetId="figure@6.[151,1436,193,1444]" captionTargetPageId="6" captionText="FIGURES 13 – 24. 13, 14, 21, 22, Achmonia doloc (Bord.), HT from Java; 15, A. eppelsheimi (Bernh. & Schub.) from Nepal; 16, A. shibatai (Bord.), HT from Taiwan; 17, A. nigra (Bord.), HT from China; 18, 23, 24, A. amabilis (Boh.) from South Africa; 19, A. simulator sp. nov., PT from Mposhi; 20, A. carinata sp. nov., HT. 13, head and pronotum; 14 – 20, labrum; 21, 23, maxillary palpus; 22, 24, labial palpus. 13, 14, 21, 22 from Bordoni (2004 a), 15 from Bordoni (2004 b), 16 from Bordoni (2010), 17 from Bordoni (2009). Scale 0.1 mm; 13, 15 and 17 without scale." httpUri="https://zenodo.org/record/228601/files/figure.png" pageId="2" pageNumber="259">Figs. 21, 23</figureCitation>
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). Labial palpi moderately long, last segment at least as long as segment 2 (
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<figureCitation id="1337B3E437573872FAADBA79FF0BFB6D" captionStart="FIGURES 13 – 24. 13, 14, 21, 22" captionStartId="6.[151,264,1465,1487]" captionTargetBox="[163,1431,203,1425]" captionTargetId="figure@6.[151,1436,193,1444]" captionTargetPageId="6" captionText="FIGURES 13 – 24. 13, 14, 21, 22, Achmonia doloc (Bord.), HT from Java; 15, A. eppelsheimi (Bernh. & Schub.) from Nepal; 16, A. shibatai (Bord.), HT from Taiwan; 17, A. nigra (Bord.), HT from China; 18, 23, 24, A. amabilis (Boh.) from South Africa; 19, A. simulator sp. nov., PT from Mposhi; 20, A. carinata sp. nov., HT. 13, head and pronotum; 14 – 20, labrum; 21, 23, maxillary palpus; 22, 24, labial palpus. 13, 14, 21, 22 from Bordoni (2004 a), 15 from Bordoni (2004 b), 16 from Bordoni (2010), 17 from Bordoni (2009). Scale 0.1 mm; 13, 15 and 17 without scale." httpUri="https://zenodo.org/record/228601/files/figure.png" pageId="2" pageNumber="259">Figs. 22, 24</figureCitation>
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). Mentum short, transverse, pentagonal. Gula very short, gular sutures contiguous.
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</paragraph>
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<paragraph id="8BB3AF6137573872FF0EBAAFFDA2FAA1" blockId="2.[151,1437,599,1993]" pageId="2" pageNumber="259">
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Pronotum without dorsal rows of punctures (
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<figureCitation id="1337B3E437573872FD1ABAA1FCA9FB51" box="[723,835,1175,1200]" captionStart="FIGURES 3 – 12. 3, 8, 12" captionStartId="4.[151,264,1753,1775]" captionTargetBox="[154,1422,213,1729]" captionTargetId="figure@4.[151,1435,193,1732]" captionTargetPageId="4" captionText="FIGURES 3 – 12. 3, 8, 12, Achmonia carinata sp. nov., HT male; 4, 7, 9, A. simulator sp. nov., PT male, 4, 9 from Mposhi, 7 from Joko; 5, 10, A. amabilis (Boheman), female from Ithala; 6, 11, A. congoensis (Bernh.), male from P. N. Garamba. 3 – 6, habitus; 7, lateral part of pronotum, ventral; 8 – 11, head; 12, pronotum and elytra. SL, superior line; IL, inferior line; EL, elytral length; p, postocular puncture. Scales 1 mm (3 – 6) and 0.5 mm (7 – 12)." httpUri="https://zenodo.org/record/228600/files/figure.png" pageId="2" pageNumber="259">Figs. 3–6</figureCitation>
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,
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<figureCitation id="1337B3E437573872FC9BBAAEFC9AFB51" box="[850,880,1176,1200]" captionStart="FIGURES 13 – 24. 13, 14, 21, 22" captionStartId="6.[151,264,1465,1487]" captionTargetBox="[163,1431,203,1425]" captionTargetId="figure@6.[151,1436,193,1444]" captionTargetPageId="6" captionText="FIGURES 13 – 24. 13, 14, 21, 22, Achmonia doloc (Bord.), HT from Java; 15, A. eppelsheimi (Bernh. & Schub.) from Nepal; 16, A. shibatai (Bord.), HT from Taiwan; 17, A. nigra (Bord.), HT from China; 18, 23, 24, A. amabilis (Boh.) from South Africa; 19, A. simulator sp. nov., PT from Mposhi; 20, A. carinata sp. nov., HT. 13, head and pronotum; 14 – 20, labrum; 21, 23, maxillary palpus; 22, 24, labial palpus. 13, 14, 21, 22 from Bordoni (2004 a), 15 from Bordoni (2004 b), 16 from Bordoni (2010), 17 from Bordoni (2009). Scale 0.1 mm; 13, 15 and 17 without scale." httpUri="https://zenodo.org/record/228601/files/figure.png" pageId="2" pageNumber="259">13</figureCitation>
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); large lateral puncture situated far from lateral margin; superior line of pronotal hypomeron not turning downwards until close to anterior angle and not joining inferior line (
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<figureCitation id="1337B3E437573872FEE3BAE9FE93FB19" box="[298,377,1247,1272]" captionStart="FIGURES 3 – 12. 3, 8, 12" captionStartId="4.[151,264,1753,1775]" captionTargetBox="[154,1422,213,1729]" captionTargetId="figure@4.[151,1435,193,1732]" captionTargetPageId="4" captionText="FIGURES 3 – 12. 3, 8, 12, Achmonia carinata sp. nov., HT male; 4, 7, 9, A. simulator sp. nov., PT male, 4, 9 from Mposhi, 7 from Joko; 5, 10, A. amabilis (Boheman), female from Ithala; 6, 11, A. congoensis (Bernh.), male from P. N. Garamba. 3 – 6, habitus; 7, lateral part of pronotum, ventral; 8 – 11, head; 12, pronotum and elytra. SL, superior line; IL, inferior line; EL, elytral length; p, postocular puncture. Scales 1 mm (3 – 6) and 0.5 mm (7 – 12)." httpUri="https://zenodo.org/record/228600/files/figure.png" pageId="2" pageNumber="259">Figs. 7</figureCitation>
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,
|
||
<figureCitation id="1337B3E437573872FE4CBAE9FE4FFB19" box="[389,421,1247,1272]" captionStart="FIGURES 25 – 37. 25 – 31" captionStartId="7.[151,264,1760,1782]" captionTargetBox="[166,1428,208,1734]" captionTargetId="figure@7.[151,1436,193,1739]" captionTargetPageId="7" captionText="FIGURES 25 – 37. 25 – 31, Achmonia knirschi sp. nov., HT male; 32 – 34, A. carinata sp. nov., 32, PT female, 33, 34, HT male; 35, A. flavomarginata (Bernh.), HT male; 36, A. kapangana sp. nov., HT male; 37, A. simulator sp. nov., HT male. 25, 32, 35 – 37, head; 26, pronotum dorsal; 27, aedeagus; 28, median lobe of aedeagus; 29, paramere; 30, 33, male tergite 10; 31, 34, male sternite 9. Scales 1 mm (25 = 26; 32 = 35 – 37), 0.5 mm (27 = 30, 31, 33, 34) and 0.1 mm (28 = 29)." httpUri="https://zenodo.org/record/228602/files/figure.png" pageId="2" pageNumber="259">26</figureCitation>
|
||
). Prosternum elevated medially, with or without short median carina posteriorly, intercoxal process minute and short, triangular; epimera present. Mesosternum very short, transverse, widely separating middle coxae. Metasternum very long.
|
||
</paragraph>
|
||
<paragraph id="8BB3AF6137573872FF0EBB7AFE6AF9F9" blockId="2.[151,1437,599,1993]" pageId="2" pageNumber="259">
|
||
Elytra overlapping at suture, variably punctured. Legs moderately long; protarsus simple (not dilated) in either sex, first four segments gradually becoming shorter, last segment about as long as three preceding segments combined; protibia with numerous strong spines on outer margin; mesotibia strongly spinose, with apical and subapical ctenidium; first segment of mesotarsus and metatarsus about equally long as second, last segment about as long as three preceding segments combined; metatibia (
|
||
<figureCitation id="1337B3E437573872FC84BBEAFC49FA14" box="[845,931,1500,1525]" captionStart="FIGURES 54 – 68. 54 – 63" captionStartId="11.[151,264,1777,1799]" captionTargetBox="[171,1414,219,1741]" captionTargetId="figure@11.[151,1436,193,1756]" captionTargetPageId="11" captionText="FIGURES 54 – 68. 54 – 63, Achmonia simulator sp. nov., 54 – 57, 63, PT male from Joko, 58 – 61, HT male, 62, PT from Bolama; 64 – 67, A. amabilis (Boheman), 64, 66, 67, LT male, 65, female from Ithala; 68, A. congoensis (Bernh.), male from Malo. 54, aedeagus lateral; 55, 58, 61, aedeagus dorsal; 56, 59, paramere; 57, 60, median lobe of aedeagus; 62, sternite 9 and sclerites of tergite 10; 63, left metatibia and metatarsus; 64, 68, head; 66, sternite 9; 67, tergite 10; 65, lateral part of head. Scales 1 mm (64 = 65, 68), 0.5 mm (54 = 55, 58, 61, 62; 63; 66 = 67) and 0.1 mm (56 = 57, 59, 60)." httpUri="https://zenodo.org/record/228604/files/figure.png" pageId="2" pageNumber="259">Fig. 63</figureCitation>
|
||
) spinose on outer margin, with apical and subapical ctenidium.
|
||
</paragraph>
|
||
<paragraph id="8BB3AF6137573872FF0EB812FD33F981" blockId="2.[151,1437,599,1993]" pageId="2" pageNumber="259">
|
||
Abdomen (
|
||
<figureCitation id="1337B3E437573872FE86B812FE2EF9DC" box="[335,452,1572,1597]" captionStart="FIGURES 3 – 12. 3, 8, 12" captionStartId="4.[151,264,1753,1775]" captionTargetBox="[154,1422,213,1729]" captionTargetId="figure@4.[151,1435,193,1732]" captionTargetPageId="4" captionText="FIGURES 3 – 12. 3, 8, 12, Achmonia carinata sp. nov., HT male; 4, 7, 9, A. simulator sp. nov., PT male, 4, 9 from Mposhi, 7 from Joko; 5, 10, A. amabilis (Boheman), female from Ithala; 6, 11, A. congoensis (Bernh.), male from P. N. Garamba. 3 – 6, habitus; 7, lateral part of pronotum, ventral; 8 – 11, head; 12, pronotum and elytra. SL, superior line; IL, inferior line; EL, elytral length; p, postocular puncture. Scales 1 mm (3 – 6) and 0.5 mm (7 – 12)." httpUri="https://zenodo.org/record/228600/files/figure.png" pageId="2" pageNumber="259">Figs. 3–6</figureCitation>
|
||
) ‘Staphylininae-shaped’ with fourth segment broadest. Tergite 7 with complete membranous palisade fringe at its posterior margin.
|
||
</paragraph>
|
||
<paragraph id="8BB3AF6137573872FF0EB85AFAB3F911" blockId="2.[151,1437,599,1993]" pageId="2" pageNumber="259">
|
||
<collectingCountry id="F31BEFF137573872FF0EB85AFEEEF964" box="[199,260,1644,1669]" name="Malta" pageId="2" pageNumber="259">Male</collectingCountry>
|
||
. Tergite and sternite 8 of male not modified. Tergite 10 of male genital segment rather narrowly exposed between sclerites of tergite 9, strongly narrowed proximally, sclerites of tergite 9 therefore contiguous mediobasally. Sternite 9 of male symmetrical, located centrally (
|
||
<figureCitation id="1337B3E437573872FCA4B882FC2BF92C" box="[877,961,1716,1741]" captionStart="FIGURES 54 – 68. 54 – 63" captionStartId="11.[151,264,1777,1799]" captionTargetBox="[171,1414,219,1741]" captionTargetId="figure@11.[151,1436,193,1756]" captionTargetPageId="11" captionText="FIGURES 54 – 68. 54 – 63, Achmonia simulator sp. nov., 54 – 57, 63, PT male from Joko, 58 – 61, HT male, 62, PT from Bolama; 64 – 67, A. amabilis (Boheman), 64, 66, 67, LT male, 65, female from Ithala; 68, A. congoensis (Bernh.), male from Malo. 54, aedeagus lateral; 55, 58, 61, aedeagus dorsal; 56, 59, paramere; 57, 60, median lobe of aedeagus; 62, sternite 9 and sclerites of tergite 10; 63, left metatibia and metatarsus; 64, 68, head; 66, sternite 9; 67, tergite 10; 65, lateral part of head. Scales 1 mm (64 = 65, 68), 0.5 mm (54 = 55, 58, 61, 62; 63; 66 = 67) and 0.1 mm (56 = 57, 59, 60)." httpUri="https://zenodo.org/record/228604/files/figure.png" pageId="2" pageNumber="259">Fig. 62</figureCitation>
|
||
). Aedeagus with voluminous basal bulbus (
|
||
<figureCitation id="1337B3E437573872FF56B8E1FF19F911" box="[159,243,1751,1776]" captionStart="FIGURES 38 – 53. 38 – 41" captionStartId="9.[151,264,1728,1750]" captionTargetBox="[162,1428,208,1702]" captionTargetId="figure@9.[151,1436,193,1707]" captionTargetPageId="9" captionText="FIGURES 38 – 53. 38 – 41, Achmonia carinata sp. nov., HT male; 42 – 44, A. kapangana sp. nov., HT male; 45 – 50, A. flavomarginata (Bernh.), LT male; 51 – 53, A. simulator sp. nov., HT male. 38, 45, aedeagus lateral; 39, 42, 46, aedeagus dorsal; 40, 43, 47, paramere; 41, 44, 48, median lobe of aedeagus; 49, 52, male sternite 9; 50, 53, male tergite 10; 51, pronotum dorsal. B, basal bulbus; ML, median lobe. Scales 0.5 mm (38 = 39; 42 = 45, 46, 49, 50, 52, 53; 51) and 0.1 mm (40 = 41; 47 = 43, 44, 48)." httpUri="https://zenodo.org/record/228603/files/figure.png" pageId="2" pageNumber="259">Fig. 39</figureCitation>
|
||
), distinct median lobe (
|
||
<figureCitation id="1337B3E437573872FE30B8E1FDA1F911" box="[505,587,1751,1776]" captionStart="FIGURES 38 – 53. 38 – 41" captionStartId="9.[151,264,1728,1750]" captionTargetBox="[162,1428,208,1702]" captionTargetId="figure@9.[151,1436,193,1707]" captionTargetPageId="9" captionText="FIGURES 38 – 53. 38 – 41, Achmonia carinata sp. nov., HT male; 42 – 44, A. kapangana sp. nov., HT male; 45 – 50, A. flavomarginata (Bernh.), LT male; 51 – 53, A. simulator sp. nov., HT male. 38, 45, aedeagus lateral; 39, 42, 46, aedeagus dorsal; 40, 43, 47, paramere; 41, 44, 48, median lobe of aedeagus; 49, 52, male sternite 9; 50, 53, male tergite 10; 51, pronotum dorsal. B, basal bulbus; ML, median lobe. Scales 0.5 mm (38 = 39; 42 = 45, 46, 49, 50, 52, 53; 51) and 0.1 mm (40 = 41; 47 = 43, 44, 48)." httpUri="https://zenodo.org/record/228603/files/figure.png" pageId="2" pageNumber="259">Fig. 39</figureCitation>
|
||
) and symmetrical parameres (
|
||
<figureCitation id="1337B3E437573872FC54B8E1FC11F911" box="[925,1019,1751,1776]" captionStart="FIGURES 25 – 37. 25 – 31" captionStartId="7.[151,264,1760,1782]" captionTargetBox="[166,1428,208,1734]" captionTargetId="figure@7.[151,1436,193,1739]" captionTargetPageId="7" captionText="FIGURES 25 – 37. 25 – 31, Achmonia knirschi sp. nov., HT male; 32 – 34, A. carinata sp. nov., 32, PT female, 33, 34, HT male; 35, A. flavomarginata (Bernh.), HT male; 36, A. kapangana sp. nov., HT male; 37, A. simulator sp. nov., HT male. 25, 32, 35 – 37, head; 26, pronotum dorsal; 27, aedeagus; 28, median lobe of aedeagus; 29, paramere; 30, 33, male tergite 10; 31, 34, male sternite 9. Scales 1 mm (25 = 26; 32 = 35 – 37), 0.5 mm (27 = 30, 31, 33, 34) and 0.1 mm (28 = 29)." httpUri="https://zenodo.org/record/228602/files/figure.png" pageId="2" pageNumber="259">Figs. 27</figureCitation>
|
||
,
|
||
<figureCitation id="1337B3E437573872FBCEB8EEFB93F911" box="[1031,1145,1751,1776]" captionStart="FIGURES 38 – 53. 38 – 41" captionStartId="9.[151,264,1728,1750]" captionTargetBox="[162,1428,208,1702]" captionTargetId="figure@9.[151,1436,193,1707]" captionTargetPageId="9" captionText="FIGURES 38 – 53. 38 – 41, Achmonia carinata sp. nov., HT male; 42 – 44, A. kapangana sp. nov., HT male; 45 – 50, A. flavomarginata (Bernh.), LT male; 51 – 53, A. simulator sp. nov., HT male. 38, 45, aedeagus lateral; 39, 42, 46, aedeagus dorsal; 40, 43, 47, paramere; 41, 44, 48, median lobe of aedeagus; 49, 52, male sternite 9; 50, 53, male tergite 10; 51, pronotum dorsal. B, basal bulbus; ML, median lobe. Scales 0.5 mm (38 = 39; 42 = 45, 46, 49, 50, 52, 53; 51) and 0.1 mm (40 = 41; 47 = 43, 44, 48)." httpUri="https://zenodo.org/record/228603/files/figure.png" pageId="2" pageNumber="259">39, 42, 46</figureCitation>
|
||
,
|
||
<figureCitation id="1337B3E437573872FB4DB8E1FB1DF911" box="[1156,1271,1751,1776]" captionStart="FIGURES 54 – 68. 54 – 63" captionStartId="11.[151,264,1777,1799]" captionTargetBox="[171,1414,219,1741]" captionTargetId="figure@11.[151,1436,193,1756]" captionTargetPageId="11" captionText="FIGURES 54 – 68. 54 – 63, Achmonia simulator sp. nov., 54 – 57, 63, PT male from Joko, 58 – 61, HT male, 62, PT from Bolama; 64 – 67, A. amabilis (Boheman), 64, 66, 67, LT male, 65, female from Ithala; 68, A. congoensis (Bernh.), male from Malo. 54, aedeagus lateral; 55, 58, 61, aedeagus dorsal; 56, 59, paramere; 57, 60, median lobe of aedeagus; 62, sternite 9 and sclerites of tergite 10; 63, left metatibia and metatarsus; 64, 68, head; 66, sternite 9; 67, tergite 10; 65, lateral part of head. Scales 1 mm (64 = 65, 68), 0.5 mm (54 = 55, 58, 61, 62; 63; 66 = 67) and 0.1 mm (56 = 57, 59, 60)." httpUri="https://zenodo.org/record/228604/files/figure.png" pageId="2" pageNumber="259">55, 58, 61</figureCitation>
|
||
,
|
||
<figureCitation id="1337B3E437573872FACAB8E1FAA6F911" box="[1283,1356,1751,1776]" captionStart="FIGURES 69 – 74. 69 – 70, A" captionStartId="14.[151,264,1567,1589]" captionTargetBox="[159,1421,205,1535]" captionTargetId="figure@14.[151,1436,195,1545]" captionTargetPageId="14" captionText="FIGURES 69 – 74. 69 – 70, A. amabilis (Boheman), LT male; 71 – 74, A. congoensis (Bernh.), male from Malo. 69 – 70, aedeagus dorsal; 70, 72, paramere; 73, sternite 9; 74, tergite 10. Scale 0.5 mm (69 = 71, 73, 74; 70 = 72)." httpUri="https://zenodo.org/record/228605/files/figure.png" pageId="2" pageNumber="259">69, 71</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C316FCEA37573875FF0EB8CAFB62FDDC" lastPageId="5" lastPageNumber="262" pageId="2" pageNumber="259" type="discussion">
|
||
<paragraph id="8BB3AF6137573872FF0EB8CAFF04F828" blockId="2.[151,1437,599,1993]" pageId="2" pageNumber="259">
|
||
<emphasis id="B978737337573872FF0EB8CAFEA6F8F4" bold="true" box="[199,332,1788,1813]" pageId="2" pageNumber="259">Discussion.</emphasis>
|
||
Some characters such as the shape of the labrum and of the maxillary and labial palpi are variable within our genus concept and can be specific for different species groups. However, the labrum in
|
||
<taxonomicName id="4C0CD4E237573872FAC7B929FA6AF8D9" box="[1294,1408,1823,1848]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="259" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337573872FAC7B929FA6AF8D9" box="[1294,1408,1823,1848]" italics="true" pageId="2" pageNumber="259">Achmonia</emphasis>
|
||
</taxonomicName>
|
||
is always dark, completely chitinised and more or less distinctly quadrilobate (
|
||
<figureCitation id="1337B3E437573872FC24B972FB9DF8BD" box="[1005,1143,1860,1885]" captionStart="FIGURES 13 – 24. 13, 14, 21, 22" captionStartId="6.[151,264,1465,1487]" captionTargetBox="[163,1431,203,1425]" captionTargetId="figure@6.[151,1436,193,1444]" captionTargetPageId="6" captionText="FIGURES 13 – 24. 13, 14, 21, 22, Achmonia doloc (Bord.), HT from Java; 15, A. eppelsheimi (Bernh. & Schub.) from Nepal; 16, A. shibatai (Bord.), HT from Taiwan; 17, A. nigra (Bord.), HT from China; 18, 23, 24, A. amabilis (Boh.) from South Africa; 19, A. simulator sp. nov., PT from Mposhi; 20, A. carinata sp. nov., HT. 13, head and pronotum; 14 – 20, labrum; 21, 23, maxillary palpus; 22, 24, labial palpus. 13, 14, 21, 22 from Bordoni (2004 a), 15 from Bordoni (2004 b), 16 from Bordoni (2010), 17 from Bordoni (2009). Scale 0.1 mm; 13, 15 and 17 without scale." httpUri="https://zenodo.org/record/228601/files/figure.png" pageId="2" pageNumber="259">Figs. 14–20</figureCitation>
|
||
). The maxillary and labial palpi are more or less always flattened but the relative width and length of the last two segments are variable among species. A similar situation was observed in other genera of Xanthonini (e.g.,
|
||
<bibRefCitation id="EF9DD29037573872FB94B9BAFAEEF845" author="Marquez" box="[1117,1284,1932,1956]" pageId="2" pageNumber="259" refString="Marquez, J. (2010) Revision of the genus Renda Blackwelder, 1952 (Coleoptera: Staphylinidae: Xantholinini). Zootaxa, 2686, 1 - 61." type="journal article" year="2010">Márquez 2010</bibRefCitation>
|
||
:
|
||
<figureCitation id="1337B3E437573872FAD9B9BAFF08F828" captionStart="FIGURES 38 – 53. 38 – 41" captionStartId="9.[151,264,1728,1750]" captionTargetBox="[162,1428,208,1702]" captionTargetId="figure@9.[151,1436,193,1707]" captionTargetPageId="9" captionText="FIGURES 38 – 53. 38 – 41, Achmonia carinata sp. nov., HT male; 42 – 44, A. kapangana sp. nov., HT male; 45 – 50, A. flavomarginata (Bernh.), LT male; 51 – 53, A. simulator sp. nov., HT male. 38, 45, aedeagus lateral; 39, 42, 46, aedeagus dorsal; 40, 43, 47, paramere; 41, 44, 48, median lobe of aedeagus; 49, 52, male sternite 9; 50, 53, male tergite 10; 51, pronotum dorsal. B, basal bulbus; ML, median lobe. Scales 0.5 mm (38 = 39; 42 = 45, 46, 49, 50, 52, 53; 51) and 0.1 mm (40 = 41; 47 = 43, 44, 48)." httpUri="https://zenodo.org/record/228603/files/figure.png" pageId="2" pageNumber="259">Figs. 38–40, 44–46</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF73FFE937563873FF5EBA11FCECFBDC" box="[151,774,1063,1085]" pageId="3" pageNumber="260">
|
||
<paragraph id="8BB3AF6137563873FF5EBA11FCECFBDC" blockId="3.[151,774,1063,1085]" box="[151,774,1063,1085]" pageId="3" pageNumber="260">
|
||
<emphasis id="B978737337563873FF5EBA11FEE7FBDC" bold="true" box="[151,269,1063,1085]" pageId="3" pageNumber="260">FIGURE 1</emphasis>
|
||
. Distribution of
|
||
<taxonomicName id="4C0CD4E237563873FE7CBA11FD57FBDC" box="[437,701,1063,1085]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="260" phylum="Arthropoda" rank="species" species="flavomarginata">
|
||
<emphasis id="B978737337563873FE7CBA11FD57FBDC" box="[437,701,1063,1085]" italics="true" pageId="3" pageNumber="260">Achmonia flavomarginata</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF73FFE937563873FF5EB9FBFD28F802" box="[151,706,1997,2019]" pageId="3" pageNumber="260">
|
||
<paragraph id="8BB3AF6137563873FF5EB9FBFD28F802" blockId="3.[151,706,1997,2019]" box="[151,706,1997,2019]" pageId="3" pageNumber="260">
|
||
<emphasis id="B978737337563873FF5EB9FBFEE6F802" bold="true" box="[151,268,1997,2019]" pageId="3" pageNumber="260">FIGURE 2</emphasis>
|
||
. Distribution of
|
||
<taxonomicName id="4C0CD4E237563873FE7CB9FBFD93F802" box="[437,633,1997,2019]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="260" phylum="Arthropoda" rank="species" species="amabilis">
|
||
<emphasis id="B978737337563873FE7CB9FBFD93F802" box="[437,633,1997,2019]" italics="true" pageId="3" pageNumber="260">Achmonia amabilis</emphasis>
|
||
</taxonomicName>
|
||
group.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF73FFE937513874FF5EB8EFFCB3F8AA" httpUri="https://zenodo.org/record/228600/files/figure.png" pageId="4" pageNumber="261" targetBox="[154,1422,213,1729]" targetPageId="4">
|
||
<paragraph id="8BB3AF6137513874FF5EB8EFFCB3F8AA" blockId="4.[151,1436,1753,1867]" pageId="4" pageNumber="261">
|
||
<emphasis id="B978737337513874FF5EB8EFFEABF90E" bold="true" box="[151,321,1753,1775]" pageId="4" pageNumber="261">FIGURES 3–12</emphasis>
|
||
. 3, 8, 12,
|
||
<taxonomicName id="4C0CD4E237513874FE60B8EFFD86F90E" box="[425,620,1753,1775]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="carinata" status="sp. nov.">
|
||
<emphasis id="B978737337513874FE60B8EFFD86F90E" box="[425,620,1753,1775]" italics="true" pageId="4" pageNumber="261">Achmonia carinata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24BCE0837513874FDBAB8EFFD2EF90E" box="[627,708,1753,1775]" pageId="4" pageNumber="261" rank="species">
|
||
<emphasis id="B978737337513874FDBAB8EFFD2EF90E" bold="true" box="[627,708,1753,1775]" pageId="4" pageNumber="261">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, HT male; 4, 7, 9,
|
||
<taxonomicName id="4C0CD4E237513874FC41B8ECFBECF90E" box="[904,1030,1753,1775]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="simulator" status="sp. nov.">
|
||
<emphasis id="B978737337513874FC41B8ECFBECF90E" box="[904,1030,1753,1775]" italics="true" pageId="4" pageNumber="261">A. simulator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A24BCE0837513874FBC4B8EFFBB4F90E" box="[1037,1118,1753,1775]" pageId="4" pageNumber="261" rank="species">
|
||
<emphasis id="B978737337513874FBC4B8EFFBB4F90E" bold="true" box="[1037,1118,1753,1775]" pageId="4" pageNumber="261">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, PT male, 4, 9 from Mposhi, 7 from Joko; 5, 10,
|
||
<taxonomicName id="4C0CD4E237513874FE98B8CFFDA8F8EF" authority="Boheman" authorityName="Boheman" box="[337,578,1784,1806]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="amabilis">
|
||
<emphasis id="B978737337513874FE98B8CFFE2CF8EF" box="[337,454,1784,1806]" italics="true" pageId="4" pageNumber="261">A. amabilis</emphasis>
|
||
(Boheman)
|
||
</taxonomicName>
|
||
, female from Ithala; 6, 11,
|
||
<taxonomicName id="4C0CD4E237513874FC97B8CFFBA1F8EF" authority="Bernh." authorityName="Bernh." box="[862,1099,1784,1806]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="261" phylum="Arthropoda" rank="species" species="congoensis">
|
||
<emphasis id="B978737337513874FC97B8CFFC06F8EF" box="[862,1004,1785,1806]" italics="true" pageId="4" pageNumber="261">A. congoensis</emphasis>
|
||
(Bernh.)
|
||
</taxonomicName>
|
||
, male from P. N. Garamba. 3–6, habitus; 7, lateral part of pronotum, ventral; 8–11, head; 12, pronotum and elytra. SL, superior line; IL, inferior line; EL, elytral length; p, postocular puncture. Scales 1mm (3–6) and 0.5 mm (7–12).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BB3AF6137513875FF0EB94EFB62FDDC" blockId="4.[151,1436,1912,2009]" lastBlockId="5.[151,1437,151,968]" lastPageId="5" lastPageNumber="262" pageId="4" pageNumber="261">
|
||
<emphasis id="B978737337513874FF0EB94EFE3BF870" bold="true" box="[199,465,1912,1937]" pageId="4" pageNumber="261">Differential diagnosis.</emphasis>
|
||
This genus differs from several other genera of
|
||
<taxonomicName id="4C0CD4E237513874FBC1B94EFB7FF870" box="[1032,1173,1912,1937]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="261" phylum="Arthropoda" rank="tribe" tribe="Xantholinini">Xantholinini</taxonomicName>
|
||
in Africa south of the Sahara by the absence of dorsal rows of punctures on the pronotum. Most characters in
|
||
<taxonomicName id="4C0CD4E237513874FBB3B9AAFB01F854" box="[1146,1259,1948,1973]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="261" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337513874FBB3B9AAFB01F854" box="[1146,1259,1948,1973]" italics="true" pageId="4" pageNumber="261">Achmonia</emphasis>
|
||
</taxonomicName>
|
||
are identical to those in
|
||
<taxonomicName id="4C0CD4E237513874FEC9B9F6FE58F838" box="[256,434,1984,2009]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="261" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337513874FEC9B9F6FE58F838" box="[256,434,1984,2009]" italics="true" pageId="4" pageNumber="261">Thyreocephalus</emphasis>
|
||
</taxonomicName>
|
||
(for redescription see
|
||
<bibRefCitation id="EF9DD29037513874FD0BB9F6FCA2F839" author="Janak" box="[706,840,1984,2009]" pageId="4" pageNumber="261" refString="Janak, J. (2010) On the genus Thyreocephalus from Africa south of Sahara with description of four new species (Coleoptera: Staphylinidae: Xantholinini). Studies and Reports, Taxonomical Series, 6, 129 - 147." type="journal article" year="2010">Janák 2010</bibRefCitation>
|
||
) but
|
||
<taxonomicName id="4C0CD4E237513874FC42B9F6FC17F838" box="[907,1021,1984,2009]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="261" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337513874FC42B9F6FC17F838" box="[907,1021,1984,2009]" italics="true" pageId="4" pageNumber="261">Achmonia</emphasis>
|
||
</taxonomicName>
|
||
can be easily distinguished by the superior line of the pronotal hypomeron not joining the inferior line.
|
||
<taxonomicName id="4C0CD4E237503875FC6BBEA1FBFEFF51" box="[930,1044,151,176]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FC6BBEA1FBFEFF51" box="[930,1044,151,176]" italics="true" pageId="5" pageNumber="262">Achmonia</emphasis>
|
||
</taxonomicName>
|
||
shares the previous character state with
|
||
<taxonomicName id="4C0CD4E237503875FF1ABE8AFEDAFF34" box="[211,304,188,213]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FF1ABE8AFEDAFF34" box="[211,304,188,213]" italics="true" pageId="5" pageNumber="262">Eulissus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54B711C337503875FE8ABE8AFE9FFF35" box="[323,373,188,212]" pageId="5" pageNumber="262">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C0CD4E237503875FE13BE8AFC49FF35" authority="Mannerheim, 1830" authorityName="Mannerheim" authorityYear="1830" box="[474,931,188,213]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="chalybaeus">
|
||
<emphasis id="B978737337503875FE13BE8AFD2AFF34" box="[474,704,188,213]" italics="true" pageId="5" pageNumber="262">Eulissus chalybaeus</emphasis>
|
||
<bibRefCitation id="EF9DD29037503875FD02BE8AFC49FF35" author="Mannerheim" box="[715,931,188,213]" pageId="5" pageNumber="262" refString="Mannerheim, C. G. von (1830) Precis d'un nouvel arrangement de la famille des brachelytres de l'ordre des insectes coleopteres. St. Petersbourg, 87 pp." type="book" year="1830">Mannerheim, 1830</bibRefCitation>
|
||
</taxonomicName>
|
||
described from
|
||
<collectingCountry id="F31BEFF137503875FBADBE8AFB40FF34" box="[1124,1194,188,213]" name="Brazil" pageId="5" pageNumber="262">Brazil</collectingCountry>
|
||
) and
|
||
<taxonomicName id="4C0CD4E237503875FB26BE8AFAB2FF34" box="[1263,1368,188,213]" class="Insecta" family="Staphylinidae" genus="Agerodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FB26BE8AFAB2FF34" box="[1263,1368,188,213]" italics="true" pageId="5" pageNumber="262">Agerodes</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<typeStatus id="54B711C337503875FAA3BE8AFA76FF35" box="[1386,1436,188,212]" pageId="5" pageNumber="262">type</typeStatus>
|
||
species
|
||
<taxonomicName id="4C0CD4E237503875FF39BEE9FD47FF19" authority="Motschulsky, 1858" authorityName="Motschulsky" authorityYear="1858" box="[240,685,223,248]" class="Insecta" family="Staphylinidae" genus="Agerodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="coeruleus">
|
||
<emphasis id="B978737337503875FF39BEE9FE24FF19" box="[240,462,223,248]" italics="true" pageId="5" pageNumber="262">Agerodes coeruleus</emphasis>
|
||
<bibRefCitation id="EF9DD29037503875FE1FBEE9FD47FF19" author="Motschulsky" box="[470,685,223,248]" pageId="5" pageNumber="262" refString="Motschulsky, V. (1858) Enumeration des nouvelles especes de Coleopteres rapportes de ses voyages. Bulletin de la Societe Imperiale des Naturalistes de Moscou, 31 (3), 204 - 264." type="journal article" year="1858">Motschulsky, 1858</bibRefCitation>
|
||
</taxonomicName>
|
||
described from
|
||
<collectingCountry id="F31BEFF137503875FCA1BEE9FC30FF19" box="[872,986,223,248]" name="Colombia" pageId="5" pageNumber="262">Colombia</collectingCountry>
|
||
; a female
|
||
<typeStatus id="54B711C337503875FB99BED6FB6AFF19" box="[1104,1152,224,248]" pageId="5" pageNumber="262">type</typeStatus>
|
||
specimen of this species from Motschulsky’s collection was examined by Janák) (for
|
||
<taxonomicName id="4C0CD4E237503875FC95BF32FBA8FEFC" box="[860,1090,260,285]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="chalybaeus">
|
||
<emphasis id="B978737337503875FC95BF32FBA8FEFC" box="[860,1090,260,285]" italics="true" pageId="5" pageNumber="262">Eulissus chalybaeus</emphasis>
|
||
</taxonomicName>
|
||
see
|
||
<bibRefCitation id="EF9DD29037503875FBB2BF32FA72FEFD" author="Marquez" box="[1147,1432,260,285]" pageId="5" pageNumber="262" refString="Marquez, J. & Asiain, J. (2002) Eulissus chalybaeus Mannerheim, 1830. Dugesiana, 9, 67 - 68." type="journal article" year="2002">Márquez & Asiain 2002</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EF9DD29037503875FF5EBF11FE7DFEA1" author="Almeida" box="[151,407,295,320]" pageId="5" pageNumber="262" refString="Almeida, M. L. & Mise, K. M. (2009) Diagnosis and key of the main families and species of South American Coleoptera of forensic importance. Revista Brasileira de Entomologia, 53, 227 - 244. http: // dx. doi. org / 10.1590 / s 0085 - 56262009000200006" type="journal article" year="2009">Almeida & Mise 2009</bibRefCitation>
|
||
(
|
||
<figureCitation id="1337B3E437503875FE61BF11FE16FEA1" box="[424,508,295,320]" captionStart="FIGURES 13 – 24. 13, 14, 21, 22" captionStartId="6.[151,264,1465,1487]" captionTargetBox="[163,1431,203,1425]" captionTargetId="figure@6.[151,1436,193,1444]" captionTargetPageId="6" captionText="FIGURES 13 – 24. 13, 14, 21, 22, Achmonia doloc (Bord.), HT from Java; 15, A. eppelsheimi (Bernh. & Schub.) from Nepal; 16, A. shibatai (Bord.), HT from Taiwan; 17, A. nigra (Bord.), HT from China; 18, 23, 24, A. amabilis (Boh.) from South Africa; 19, A. simulator sp. nov., PT from Mposhi; 20, A. carinata sp. nov., HT. 13, head and pronotum; 14 – 20, labrum; 21, 23, maxillary palpus; 22, 24, labial palpus. 13, 14, 21, 22 from Bordoni (2004 a), 15 from Bordoni (2004 b), 16 from Bordoni (2010), 17 from Bordoni (2009). Scale 0.1 mm; 13, 15 and 17 without scale." httpUri="https://zenodo.org/record/228601/files/figure.png" pageId="5" pageNumber="262">Fig. 16</figureCitation>
|
||
),
|
||
<bibRefCitation id="EF9DD29037503875FDDABF11FD07FEA1" author="Aballay" box="[531,749,295,320]" pageId="5" pageNumber="262" refString="Aballay, F. H., Chani-Posse, M. R., Ayon, M. R., Maldonado, M. B. & Centeno, N. D. (2014) An illustrated key to and diagnoses of the species of Staphylinidae (Coleoptera) associated with decaying carcasses in Argentina. Zootaxa, 3860, 101 - 124. http: // dx. doi. org / 10.11646 / zootaxa. 3860.2.1" type="journal article" year="2014">Aballay et al. 2014</bibRefCitation>
|
||
(
|
||
<figureCitation id="1337B3E437503875FD37BF11FCB8FEA1" box="[766,850,295,320]" captionStart="FIGURES 25 – 37. 25 – 31" captionStartId="7.[151,264,1760,1782]" captionTargetBox="[166,1428,208,1734]" captionTargetId="figure@7.[151,1436,193,1739]" captionTargetPageId="7" captionText="FIGURES 25 – 37. 25 – 31, Achmonia knirschi sp. nov., HT male; 32 – 34, A. carinata sp. nov., 32, PT female, 33, 34, HT male; 35, A. flavomarginata (Bernh.), HT male; 36, A. kapangana sp. nov., HT male; 37, A. simulator sp. nov., HT male. 25, 32, 35 – 37, head; 26, pronotum dorsal; 27, aedeagus; 28, median lobe of aedeagus; 29, paramere; 30, 33, male tergite 10; 31, 34, male sternite 9. Scales 1 mm (25 = 26; 32 = 35 – 37), 0.5 mm (27 = 30, 31, 33, 34) and 0.1 mm (28 = 29)." httpUri="https://zenodo.org/record/228602/files/figure.png" pageId="5" pageNumber="262">Fig. 33</figureCitation>
|
||
)).
|
||
<taxonomicName id="4C0CD4E237503875FCBBBF11FC0EFEA1" box="[882,996,295,320]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FCBBBF11FC0EFEA1" box="[882,996,295,320]" italics="true" pageId="5" pageNumber="262">Achmonia</emphasis>
|
||
</taxonomicName>
|
||
markedly differs from
|
||
<taxonomicName id="4C0CD4E237503875FB26BF11FAA6FEA1" box="[1263,1356,295,320]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FB26BF11FAA6FEA1" box="[1263,1356,295,320]" italics="true" pageId="5" pageNumber="262">Eulissus</emphasis>
|
||
</taxonomicName>
|
||
by the dark, strongly chitinised and more or less quadrilobate labrum. The labrum in
|
||
<taxonomicName id="4C0CD4E237503875FBD4BF7AFB90FE84" box="[1053,1146,332,357]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FBD4BF7AFB90FE84" box="[1053,1146,332,357]" italics="true" pageId="5" pageNumber="262">Eulissus</emphasis>
|
||
</taxonomicName>
|
||
is bilobate with only the middle part strongly chitinised and the rest weak and semi-transparent (for details see
|
||
<bibRefCitation id="EF9DD29037503875FB45BF59FAE1FE69" author="Steel" box="[1164,1291,367,392]" pageId="5" pageNumber="262" refString="Steel, W. O. (1938) On the xantholinid genera Thyreocephalus Guer. and Eulissus Mannh. (Col. Staphylinidae). The Entomologist's Monthly Magazine, 74, 55 - 56." type="journal article" year="1938">Steel 1938</bibRefCitation>
|
||
: 55,
|
||
<figureCitation id="1337B3E437503875FA8EBF59FA61FE69" box="[1351,1419,367,392]" captionStart="FIGURE 1" captionStartId="3.[151,250,1063,1085]" captionText="FIGURE 1. Distribution of Achmonia flavomarginata group." pageId="5" pageNumber="262">Fig. 1</figureCitation>
|
||
).
|
||
<taxonomicName id="4C0CD4E237503875FF5EBFA2FEE3FE4C" box="[151,265,404,429]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FF5EBFA2FEE3FE4C" box="[151,265,404,429]" italics="true" pageId="5" pageNumber="262">Achmonia</emphasis>
|
||
</taxonomicName>
|
||
(and
|
||
<taxonomicName id="4C0CD4E237503875FE82BFA2FE42FE4C" box="[331,424,404,429]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FE82BFA2FE42FE4C" box="[331,424,404,429]" italics="true" pageId="5" pageNumber="262">Eulissus</emphasis>
|
||
</taxonomicName>
|
||
) differs from
|
||
<taxonomicName id="4C0CD4E237503875FD80BFA2FD58FE4C" box="[585,690,404,429]" class="Insecta" family="Staphylinidae" genus="Agerodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FD80BFA2FD58FE4C" box="[585,690,404,429]" italics="true" pageId="5" pageNumber="262">Agerodes</emphasis>
|
||
</taxonomicName>
|
||
by the presence of both an apical and subapical ctenidium on the metatibia. In
|
||
<taxonomicName id="4C0CD4E237503875FEE7BF81FE7DFE31" box="[302,407,439,464]" class="Insecta" family="Staphylinidae" genus="Agerodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FEE7BF81FE7DFE31" box="[302,407,439,464]" italics="true" pageId="5" pageNumber="262">Agerodes</emphasis>
|
||
</taxonomicName>
|
||
the subapical ctenidium is absent (observed in several undetermined Neotropical species). We consider
|
||
<taxonomicName id="4C0CD4E237503875FEEEBFEAFE73FE14" box="[295,409,476,501]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FEEEBFEAFE73FE14" box="[295,409,476,501]" italics="true" pageId="5" pageNumber="262">Achmonia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C0CD4E237503875FE07BFEAFDC1FE14" box="[462,555,476,501]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FE07BFEAFDC1FE14" box="[462,555,476,501]" italics="true" pageId="5" pageNumber="262">Eulissus</emphasis>
|
||
</taxonomicName>
|
||
as sister groups, differing mainly by the shape of the labrum, but probably also by differences in the shape and structure of the aedeagus, which may be revealed by a revision of
|
||
<taxonomicName id="4C0CD4E237503875FAFEBFC9FA7EFDF9" box="[1335,1428,511,536]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FAFEBFC9FA7EFDF9" box="[1335,1428,511,536]" italics="true" pageId="5" pageNumber="262">Eulissus</emphasis>
|
||
</taxonomicName>
|
||
. According to our opinion, the genus
|
||
<taxonomicName id="4C0CD4E237503875FDFBBC12FD65FDDC" box="[562,655,548,573]" class="Insecta" family="Staphylinidae" genus="Eulissus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FDFBBC12FD65FDDC" box="[562,655,548,573]" italics="true" pageId="5" pageNumber="262">Eulissus</emphasis>
|
||
</taxonomicName>
|
||
is only distributed in the Neotropical region.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C316FCEA37503875FF0EBC71FD85FC26" pageId="5" pageNumber="262" type="distribution">
|
||
<paragraph id="8BB3AF6137503875FF0EBC71FEA6FCD9" blockId="5.[151,1437,151,968]" pageId="5" pageNumber="262">
|
||
<emphasis id="B978737337503875FF0EBC71FE8AFD81" bold="true" box="[199,352,583,608]" pageId="5" pageNumber="262">Distribution.</emphasis>
|
||
<taxonomicName id="4C0CD4E237503875FEA3BC71FE36FD81" box="[362,476,583,608]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FEA3BC71FE36FD81" box="[362,476,583,608]" italics="true" pageId="5" pageNumber="262">Achmonia</emphasis>
|
||
</taxonomicName>
|
||
was previously confused with
|
||
<taxonomicName id="4C0CD4E237503875FC81BC71FC10FD81" box="[840,1018,583,608]" class="Insecta" family="Staphylinidae" genus="Thyreocephalus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FC81BC71FC10FD81" box="[840,1018,583,608]" italics="true" pageId="5" pageNumber="262">Thyreocephalus</emphasis>
|
||
</taxonomicName>
|
||
and in some regions they co-occur.
|
||
<taxonomicName id="4C0CD4E237503875FF5EBC5AFEE3FD64" box="[151,265,620,645]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B978737337503875FF5EBC5AFEE3FD64" box="[151,265,620,645]" italics="true" pageId="5" pageNumber="262">Achmonia</emphasis>
|
||
</taxonomicName>
|
||
has been hitherto represented by the following eight species occurring in the Oriental and Australian regions:
|
||
<taxonomicName id="4C0CD4E237503875FF31BCA6FDA5FD49" authority="Bernhauer, 1915" authorityName="Bernhauer" authorityYear="1915" box="[248,591,655,680]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="sondaica">
|
||
<emphasis id="B978737337503875FF31BCA6FE96FD49" box="[248,380,655,680]" italics="true" pageId="5" pageNumber="262">A. sondaica</emphasis>
|
||
(Bernhauer, 1915)
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F31BEFF137503875FD97BCB9FD23FD49" box="[606,713,655,680]" name="Malaysia" pageId="5" pageNumber="262">Malaysia</collectingCountry>
|
||
, Java) (=
|
||
<taxonomicName id="4C0CD4E237503875FCFEBCA6FBA8FD49" authority="Bordoni, 2004" authorityName="Bordoni" authorityYear="2004" box="[823,1090,655,680]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="doloc">
|
||
<emphasis id="B978737337503875FCFEBCA6FC7EFD49" box="[823,916,655,680]" italics="true" pageId="5" pageNumber="262">A. doloc</emphasis>
|
||
Bordoni, 2004
|
||
</taxonomicName>
|
||
);
|
||
<taxonomicName id="4C0CD4E237503875FB9CBCA6FEA7FD2D" authority="Bernhauer & Schubert, 1914" authorityName="Bernhauer & Schubert" authorityYear="1914" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="eppelsheimi">
|
||
<emphasis id="B978737337503875FB9CBCA6FB13FD49" box="[1109,1273,655,680]" italics="true" pageId="5" pageNumber="262">A. eppelsheimi</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9DD29037503875FAC0BCB9FEAEFD2D" author="Bernhauer" pageId="5" pageNumber="262" refString="Bernhauer, M. & Schubert, K. (1914) Staphylinidae IV. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus, 5 (57). Junk, Berlin, pp. 289 - 408." type="journal article" year="1914">Bernhauer & Schubert, 1914</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(NE
|
||
<collectingCountry id="F31BEFF137503875FE46BC82FE21FD2C" box="[399,459,692,717]" name="India" pageId="5" pageNumber="262">India</collectingCountry>
|
||
,
|
||
<collectingCountry id="F31BEFF137503875FE13BC82FDCAFD2C" box="[474,544,692,717]" name="Nepal" pageId="5" pageNumber="262">Nepal</collectingCountry>
|
||
, Sikkim,
|
||
<collectingCountry id="F31BEFF137503875FD59BC82FD09FD2C" box="[656,739,692,717]" name="Bhutan" pageId="5" pageNumber="262">Bhutan</collectingCountry>
|
||
) (=
|
||
<taxonomicName id="4C0CD4E237503875FCDEBC82FB8FFD2D" authority="Bordoni, 2004" authorityName="Bordoni" authorityYear="2004" box="[791,1125,692,717]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="hromadkai">
|
||
<emphasis id="B978737337503875FCDEBC82FC5BFD2C" box="[791,945,692,717]" italics="true" pageId="5" pageNumber="262">A. hromadkai</emphasis>
|
||
Bordoni, 2004
|
||
</taxonomicName>
|
||
);
|
||
<taxonomicName id="4C0CD4E237503875FBB2BC82FA76FD2C" authority="Fauvel, 1895" authorityName="Fauvel" authorityYear="1895" box="[1147,1436,692,717]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="gestroi">
|
||
<emphasis id="B978737337503875FBB2BC82FB03FD2D" box="[1147,1257,692,716]" italics="true" pageId="5" pageNumber="262">A. gestroi</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9DD29037503875FB35BC82FA7EFD2C" author="Fauvel" box="[1276,1428,692,717]" pageId="5" pageNumber="262" refString="Fauvel, A. (1895) Staphylinides nouveaux de l'Inde et de la Malaisie. Revue d'Entomologie, 14, 180 - 286." type="journal article" year="1895">Fauvel, 1895</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F31BEFF137503875FF57BCEEFF05FD11" box="[158,239,728,752]" name="Myanmar" pageId="5" pageNumber="262">Burma</collectingCountry>
|
||
,
|
||
<collectingCountry id="F31BEFF137503875FF32BCE1FE8EFD11" box="[251,356,727,752]" name="Thailand" pageId="5" pageNumber="262">Thailand</collectingCountry>
|
||
);
|
||
<taxonomicName id="4C0CD4E237503875FEBEBCEEFCDAFD11" authority="Bordoni, 2012" authorityName="Bordoni" authorityYear="2012" box="[375,816,727,752]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="solodovnikovi">
|
||
<emphasis id="B978737337503875FEBEBCEEFDD9FD11" box="[375,563,727,752]" italics="true" pageId="5" pageNumber="262">A. solodovnikovi</emphasis>
|
||
<bibRefCitation id="EF9DD29037503875FDF3BCE1FD36FD11" author="Bordoni" box="[570,732,727,752]" pageId="5" pageNumber="262" refString="Bordoni, A. (2012) New data on the Xantholinini of the Oriental Region 31. New species and new records from Laos of the Zoological Museum of Copenhagen. Quaderno di Studi e Notizie di Storia Naturale della Romagna, 36, 105 - 113." type="journal article" year="2012">Bordoni, 2012</bibRefCitation>
|
||
(
|
||
<collectingCountry id="F31BEFF137503875FD25BCEEFCC2FD11" box="[748,808,728,752]" name="Laos" pageId="5" pageNumber="262">Laos</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C0CD4E237503875FCF5BCEEFBA9FD11" authority="Bordoni, 2009" authorityName="Bordoni" authorityYear="2009" box="[828,1091,727,752]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="nigra">
|
||
<emphasis id="B978737337503875FCF5BCEEFC73FD11" box="[828,921,728,752]" italics="true" pageId="5" pageNumber="262">A. nigra</emphasis>
|
||
<bibRefCitation id="EF9DD29037503875FC69BCE1FBA9FD11" author="Bordoni" box="[928,1091,727,752]" pageId="5" pageNumber="262" refString="Bordoni, A. (2009) New data on Xantholinini from China. XVII. Daolus niger nov. sp. from Zhejiang (Coleoptera: Staphylinidae, Xantholinini). Linzer Biologische Beitrage, 41 (2), 1867 - 1870." type="journal article" year="2009">Bordoni, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
(
|
||
<collectingCountry id="F31BEFF137503875FB9BBCE1FB70FD11" box="[1106,1178,727,752]" name="China" pageId="5" pageNumber="262">China</collectingCountry>
|
||
: Zhejiang);
|
||
<taxonomicName id="4C0CD4E237503875FAEBBCEEFE5CFCF4" authority="Bordoni, 2010" authorityName="Bordoni" authorityYear="2010" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="shibatai">
|
||
<emphasis id="B978737337503875FAEBBCEEFA76FD11" box="[1314,1436,727,752]" italics="true" pageId="5" pageNumber="262">A. shibatai</emphasis>
|
||
<bibRefCitation id="EF9DD29037503875FF5EBCCAFEAAFCF5" author="Bordoni" box="[151,320,764,789]" pageId="5" pageNumber="262" refString="Bordoni, A. (2010) Description of the first Thyreocephalus and Daolus from Taiwan (Insecta: Coleoptera: Staphylinidae). Annalen des Naturhistorisches Museums in Wien, B, 111, 55 - 59." type="journal article" year="2010">Bordoni, 2010</bibRefCitation>
|
||
(
|
||
<collectingCountry id="F31BEFF137503875FE9CBCCAFE44FCF4" box="[341,430,764,789]" name="Taiwan" pageId="5" pageNumber="262">Taiwan</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C0CD4E237503875FE01BCCAFC67FCF4" authority="Bordoni, 2005" authorityName="Bordoni" authorityYear="2005" box="[456,909,764,789]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="wogwog">
|
||
<emphasis id="B978737337503875FE01BCCAFDA1FCF5" box="[456,587,764,788]" italics="true" pageId="5" pageNumber="262">A. wogwog</emphasis>
|
||
<bibRefCitation id="EF9DD29037503875FD93BCCAFCE8FCF4" author="Bordoni" box="[602,770,764,789]" pageId="5" pageNumber="262" refString="Bordoni, A. (2005) Revision of the Xantholinini of Australia (Coleoptera: Staphylinidae). In: Daccordi, M. & Giachino, P. M. (Eds.), Results of the Zoological Missions to Australia of the Regional Museum of Natural Sciences of Turin, Italy. II. Monografie del Museo regionale di Scienze naturali, Torino, 42, 435 - 614." type="journal article" year="2005">Bordoni, 2005</bibRefCitation>
|
||
(
|
||
<collectingCountry id="F31BEFF137503875FCD1BCCAFC6FFCF4" box="[792,901,764,789]" name="Australia" pageId="5" pageNumber="262">Australia</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C0CD4E237503875FC56BCCAFA76FCF4" authority="(Erichson, 1839) (Australia)" authorityName="Erichson" authorityYear="1839" baseAuthorityName="Erichson" baseAuthorityYear="1839" box="[927,1436,764,789]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="cyanoptera">
|
||
<emphasis id="B978737337503875FC56BCCAFBA9FCF5" box="[927,1091,764,788]" italics="true" pageId="5" pageNumber="262">A. cyanoptera</emphasis>
|
||
(
|
||
<bibRefCitation id="EF9DD29037503875FB90BCCAFAE6FCF5" author="Erichson" box="[1113,1292,764,789]" pageId="5" pageNumber="262" refString="Erichson, W. F. (1839) Genera et species Staphylinorum insectorum coleopterorum familiae (1). F. H. Morin, Berlin, 400 pp. http: // dx. doi. org / 10.5962 / bhl. title. 59644" type="book" year="1839">Erichson, 1839</bibRefCitation>
|
||
) (
|
||
<collectingCountry id="F31BEFF137503875FAE3BCCAFA7EFCF4" box="[1322,1428,764,789]" name="Australia" pageId="5" pageNumber="262">Australia</collectingCountry>
|
||
)
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="EF9DD29037503875FF57BD29FEAAFCD9" author="Bordoni" box="[158,320,799,824]" pageId="5" pageNumber="262" refString="Bordoni, A. (2012) New data on the Xantholinini of the Oriental Region 31. New species and new records from Laos of the Zoological Museum of Copenhagen. Quaderno di Studi e Notizie di Storia Naturale della Romagna, 36, 105 - 113." type="journal article" year="2012">Bordoni 2012</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BB3AF6137503875FF0EBD73FD85FC26" blockId="5.[151,1437,151,968]" pageId="5" pageNumber="262">
|
||
In Africa the genus can be divided into two distinct species groups, the first composed of species with setiferous postocular punctures on the head (
|
||
<taxonomicName id="4C0CD4E237503875FD51BD51FCAEFC61" box="[664,836,871,896]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="flavomarginata">
|
||
<emphasis id="B978737337503875FD51BD51FCAEFC61" box="[664,836,871,896]" italics="true" pageId="5" pageNumber="262">flavomarginata</emphasis>
|
||
</taxonomicName>
|
||
group) and the second composed of species lacking postocular punctures (
|
||
<taxonomicName id="4C0CD4E237503875FE5DBDBAFE1FFC44" box="[404,501,908,933]" class="Insecta" family="Staphylinidae" genus="Achmonia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="262" phylum="Arthropoda" rank="species" species="amabilis">
|
||
<emphasis id="B978737337503875FE5DBDBAFE1FFC44" box="[404,501,908,933]" italics="true" pageId="5" pageNumber="262">amabilis</emphasis>
|
||
</taxonomicName>
|
||
group). Both these groups can be distinguished from non-Afrotropical species of the genus by the non-carinate prosternum.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |