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<document id="EC32AEF5C99224812341C44B71825861" ID-CLB-Dataset="28156" ID-DOI="10.11646/zootaxa.4563.1.3" ID-GBIF-Dataset="65ff8808-27b5-43df-af44-fc56857e8906" ID-ISSN="1175-5326" ID-Zenodo-Dep="2600975" ID-ZooBank="7DF1DFC3-1B49-43AD-A4AC-2DA9BAC1E6C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553171079484" checkinUser="plazi" docAuthor="Chen, Weicai, Liao, Xiaowen, Zhou, Shichu &amp; Mo, Yunming" docDate="2019" docId="03A50C1C6824835EFF53FF66FD84FE49" docLanguage="en" docName="zootaxa.4563.1.3.pdf" docOrigin="Zootaxa 4563 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leptobrachella shangsiensis Chen &amp; Liao &amp; Zhou &amp; Mo 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="79" masterDocId="FF9C746468218352FFC4FFF1FFDFFFE4" masterDocTitle="A new species of Leptobrachella (Anura: Megophryidae) from southern Guangxi, China" masterLastPageNumber="82" masterPageNumber="67" pageNumber="72" updateTime="1698702976614" updateUser="ExternalLinkService">
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<mods:title id="45D2CBBFB66AE5C4A5E292AD786FCC3D">A new species of Leptobrachella (Anura: Megophryidae) from southern Guangxi, China</mods:title>
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<mods:namePart id="B7503853DEE6CCA16A614CFF0E3AB90D">Chen, Weicai</mods:namePart>
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<mods:namePart id="5DF4D7C52A7C5F2DDDA701F8F447DCB1">Liao, Xiaowen</mods:namePart>
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<mods:namePart id="C2BBF3D621ADB3F688FDF994C1DA0BC1">Zhou, Shichu</mods:namePart>
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<mods:namePart id="5B670FB07EA9FEAAAFE02115DB04249C">Mo, Yunming</mods:namePart>
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<treatment id="03A50C1C6824835EFF53FF66FD84FE49" ID-DOI="http://doi.org/10.5281/zenodo.5933270" ID-GBIF-Taxon="154474446" ID-Zenodo-Dep="5933270" LSID="urn:lsid:plazi:treatment:03A50C1C6824835EFF53FF66FD84FE49" httpUri="http://treatment.plazi.org/id/03A50C1C6824835EFF53FF66FD84FE49" lastPageId="12" lastPageNumber="79" pageId="5" pageNumber="72">
<subSubSection id="C316EE8168248357FF53FF66FD8DFF56" box="[151,594,151,178]" pageId="5" pageNumber="72" type="nomenclature">
<paragraph id="8BB3BD0A68248357FF53FF66FD8DFF56" blockId="5.[151,594,151,211]" box="[151,594,151,178]" pageId="5" pageNumber="72">
<heading id="D0FB0A6668248357FF53FF66FD8DFF56" bold="true" box="[151,594,151,178]" fontSize="11" level="1" pageId="5" pageNumber="72" reason="1">
<emphasis id="B978611868248357FF53FF66FD8DFF56" bold="true" box="[151,594,151,178]" pageId="5" pageNumber="72">
<taxonomicName id="4C0CC68968248357FF53FF66FE31FF55" ID-CoL="6PG45" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[151,494,151,177]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B978611868248357FF53FF66FE31FF55" bold="true" box="[151,494,151,177]" italics="true" pageId="5" pageNumber="72">Leptobrachella shangsiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24BDC6368248357FE31FF69FD8DFF56" box="[501,594,152,178]" pageId="5" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C316EE8168248357FF53FF4BFF20FF37" box="[151,255,186,211]" pageId="5" pageNumber="72" type="description">
<paragraph id="8BB3BD0A68248357FF53FF4BFF20FF37" blockId="5.[151,594,151,211]" box="[151,255,186,211]" pageId="5" pageNumber="72">
<figureCitation id="1337A18F68248357FF53FF4BFF20FF37" box="[151,255,186,211]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="6.[151,250,1627,1649]" captionStartId-1="7.[151,250,725,747]" captionTargetBox-0="[167,1400,196,1584]" captionTargetBox-1="[151,1436,193,704]" captionTargetId-0="figure@6.[167,1400,196,1584]" captionTargetId-1="figure@7.[151,1436,193,704]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURE 2. Holotype of Leptobrachella shangsiensis sp. nov. (NHMG1704002): (A) Dorsal view, (B) dorsolateral view, (C) flank and lateral view, (D) ventral view, (E) volar view of the left hand, and (F) thenar view of the left foot." captionText-1="FIGURE 3. Leptobrachella shangsiensis sp. nov. in habitat (Photo taken in situ)." httpUri-0="https://zenodo.org/record/2600979/files/figure.png" httpUri-1="https://zenodo.org/record/2600983/files/figure.png" pageId="5" pageNumber="72">Figs. 23</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C316EE8168248357FF53FEF3FE26FDF2" pageId="5" pageNumber="72" type="materials_examined">
<paragraph id="8BB3BD0A68248357FF53FEF3FE47FE87" blockId="5.[151,1437,258,1975]" pageId="5" pageNumber="72">
<materialsCitation id="3B64B75768248357FF53FEF3FE47FE87" ID-GBIF-Occurrence="2013950969" collectingDate="2017-04-26" collectionCode="NHMG" collectorName="Shiwandashan National NR" country="China" elevation="500" latitude="22.455" location="Guangxi" longLatPrecision="75" longitude="107.048" pageId="5" pageNumber="72" specimenCode="NHMG1704002" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<emphasis id="B978611868248357FF53FEF3FEDBFEFF" bold="true" box="[151,260,258,283]" pageId="5" pageNumber="72">
<typeStatus id="54B703A868248357FF53FEF3FEDBFEFF" box="[151,260,258,283]" pageId="5" pageNumber="72" type="holotype">Holotype</typeStatus>
</emphasis>
.
<specimenCode id="DBAA157168248357FED7FEF2FE12FEFF" box="[275,461,259,283]" collectionCode="NHMG" country="Sweden" httpUri="http://grbio.org/cool/1jid-0yf9" name="Goteborgs Naturhistoriska Museet" pageId="5" pageNumber="72">NHMG1704002</specimenCode>
, adult male (
<figureCitation id="1337A18F68248357FDA5FEF3FD7BFEFF" box="[609,676,258,283]" captionStart="FIGURE 2" captionStartId="6.[151,250,1627,1649]" captionTargetBox="[167,1400,196,1584]" captionTargetId="figure@6.[167,1400,196,1584]" captionTargetPageId="6" captionText="FIGURE 2. Holotype of Leptobrachella shangsiensis sp. nov. (NHMG1704002): (A) Dorsal view, (B) dorsolateral view, (C) flank and lateral view, (D) ventral view, (E) volar view of the left hand, and (F) thenar view of the left foot." httpUri="https://zenodo.org/record/2600979/files/figure.png" pageId="5" pageNumber="72">Fig. 2</figureCitation>
AF), collected at
<collectorName id="26F9D8DC68248357FC45FEF3FB62FEFE" box="[897,1213,258,283]" pageId="5" pageNumber="72">Shiwandashan National NR</collectorName>
,
<collectingRegion id="49C873E868248357FB0DFEF3FAF5FEFF" box="[1225,1322,258,283]" country="China" name="Guangxi" pageId="5" pageNumber="72">Guangxi</collectingRegion>
Province,
<collectingCountry id="F31BFD9A68248357FF53FED7FF05FEDB" box="[151,218,294,319]" name="China" pageId="5" pageNumber="72">China</collectingCountry>
(
<geoCoordinate id="EE38DBCD68248357FF2EFED7FEBEFEDA" box="[234,353,294,319]" direction="north" orientation="latitude" pageId="5" pageNumber="72" precision="55" value="22.455">22.455° N</geoCoordinate>
,
<geoCoordinate id="EE38DBCD68248357FEAAFED6FE30FEDA" box="[366,495,295,319]" direction="east" orientation="longitude" pageId="5" pageNumber="72" precision="55" value="107.048">107.048° E</geoCoordinate>
; alt.
<quantity id="4CF410EF68248357FDECFED7FDAAFEDB" box="[552,629,294,319]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="72" unit="m" value="500.0">
<elevation id="00215A3968248357FDECFED7FDAAFEDB" box="[552,629,294,319]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="72" unit="m" value="500.0">500 m</elevation>
</quantity>
). Collected on
<date id="FFB29BCA68248357FCE6FED7FC1CFEDB" box="[802,963,294,319]" pageId="5" pageNumber="72" value="2017-04-26">
<collectingDate id="EFF6622268248357FCE6FED7FC1CFEDB" box="[802,963,294,319]" pageId="5" pageNumber="72" value="2017-04-26">26 April 2017</collectingDate>
</date>
by Weicai Chen, Yunming Mo, Xiaowen Liao and Shichu Zhou.
</materialsCitation>
</paragraph>
<paragraph id="8BB3BD0A68248357FF03FE9FFE26FDF2" blockId="5.[151,1437,258,1975]" pageId="5" pageNumber="72">
<materialsCitation id="3B64B75768248357FF03FE9FFD97FE63" ID-GBIF-Occurrence="2013950968" box="[199,584,366,391]" collectionCode="NHMG" pageId="5" pageNumber="72" specimenCode="NHMG1401024-031" specimenCount="1" typeStatus="paratype">
<emphasis id="B978611868248357FF03FE9FFEE0FE63" bold="true" box="[199,319,366,391]" pageId="5" pageNumber="72">
<typeStatus id="54B703A868248357FF03FE9FFEE0FE63" box="[199,319,366,391]" pageId="5" pageNumber="72" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<specimenCode id="DBAA157168248357FE96FE9EFD9BFE63" box="[338,580,367,391]" collectionCode="NHMG" country="Sweden" httpUri="http://grbio.org/cool/1jid-0yf9" name="Goteborgs Naturhistoriska Museet" pageId="5" pageNumber="72">NHMG1401024031</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B64B75768248357FD90FE9EFC99FE63" ID-GBIF-Occurrence="2013950965" box="[596,838,367,391]" collectionCode="NHMG" pageId="5" pageNumber="72" specimenCode="NHMG1704003-004" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBAA157168248357FD90FE9EFC99FE63" box="[596,838,367,391]" collectionCode="NHMG" country="Sweden" httpUri="http://grbio.org/cool/1jid-0yf9" name="Goteborgs Naturhistoriska Museet" pageId="5" pageNumber="72">NHMG1704003004</specimenCode>
</materialsCitation>
, ten adult males,
<materialsCitation id="3B64B75768248357FBDAFE9EFACFFE63" ID-GBIF-Occurrence="2013950963" box="[1054,1296,367,391]" collectionCode="NHMG" pageId="5" pageNumber="72" specimenCode="NHMG1401032-034" specimenCount="1" typeStatus="paratype">
<specimenCode id="DBAA157168248357FBDAFE9EFACFFE63" box="[1054,1296,367,391]" collectionCode="NHMG" country="Sweden" httpUri="http://grbio.org/cool/1jid-0yf9" name="Goteborgs Naturhistoriska Museet" pageId="5" pageNumber="72">NHMG1401032034</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B64B75768248357FAE4FE9FFA47FE4E" ID-GBIF-Occurrence="2013950966" collectingDate="2014-01-03" collectionCode="NHMG" collectorName="Shiwandashan National NR" country="China" elevation="500" latitude="22.455" location="Guangxi" longLatPrecision="75" longitude="107.048" pageId="5" pageNumber="72" specimenCount="1" stateProvince="Guangxi" typeStatus="paratype">
three adult females, collected at same locality as holotype on
<date id="FFB29BCA68248357FCE9FE62FC19FE4F" box="[813,966,403,427]" pageId="5" pageNumber="72" value="2014-01-03">
<collectingDate id="EFF6622268248357FCE9FE62FC19FE4F" box="[813,966,403,427]" pageId="5" pageNumber="72" value="2014-01-03">3 Jan. 2014</collectingDate>
</date>
by Weicai Chen and Yunming Mo
</materialsCitation>
.
<materialsCitation id="3B64B75768248357FF53FE47FEB2FE2B" ID-GBIF-Occurrence="2013950971" box="[151,365,438,463]" collectionCode="NHMG" country="China" pageId="5" pageNumber="72" specimenCode="NHMG201604001" specimenCount="1" stateProvince="Guangxi" typeStatus="paratype">
<specimenCode id="DBAA157168248357FF53FE47FEB2FE2B" box="[151,365,438,463]" collectionCode="NHMG" country="Sweden" httpUri="http://grbio.org/cool/1jid-0yf9" name="Goteborgs Naturhistoriska Museet" pageId="5" pageNumber="72">NHMG201604001</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B64B75768248357FEBDFE46FEECFE16" ID-GBIF-Occurrence="2013950964" collectingDate="2016-04-25" collectionCode="NHMG" collectorName="Shiwandashan National NR" country="China" elevation="500" latitude="22.455" location="Guangxi" longLatPrecision="75" longitude="107.048" pageId="5" pageNumber="72" specimenCount="1" stateProvince="Guangxi" typeStatus="paratype">
one adult female, collected at same locality as holotype on
<date id="FFB29BCA68248357FBD0FE47FB6DFE2B" box="[1044,1202,438,463]" pageId="5" pageNumber="72" value="2016-04-25">
<collectingDate id="EFF6622268248357FBD0FE47FB6DFE2B" box="[1044,1202,438,463]" pageId="5" pageNumber="72" value="2016-04-25">25 April 2016</collectingDate>
</date>
by Weicai Chen and Yunming Mo
</materialsCitation>
.
<materialsCitation id="3B64B75768248357FE86FE2BFE22FE17" ID-GBIF-Occurrence="2013950970" box="[322,509,474,499]" collectionCode="NHMG" country="China" pageId="5" pageNumber="72" specimenCode="NHMG1704005" specimenCount="1" stateProvince="Guangxi" typeStatus="paratype">
<specimenCode id="DBAA157168248357FE86FE2BFE22FE17" box="[322,509,474,499]" collectionCode="NHMG" country="Sweden" httpUri="http://grbio.org/cool/1jid-0yf9" name="Goteborgs Naturhistoriska Museet" pageId="5" pageNumber="72">NHMG1704005</specimenCode>
</materialsCitation>
,
<materialsCitation id="3B64B75768248357FDC8FE2AFE2AFDF2" ID-GBIF-Occurrence="2013950967" collectingDate="2017-04-26" collectionCode="NHMG" collectorName="Shiwandashan National NR" country="China" elevation="500" latitude="22.455" location="Guangxi" longLatPrecision="75" longitude="107.048" pageId="5" pageNumber="72" specimenCount="1" stateProvince="Guangxi" typeStatus="paratype">
one adult female, collected at same locality as holotype on
<date id="FFB29BCA68248357FB0BFE2BFAAAFE17" box="[1231,1397,474,499]" pageId="5" pageNumber="72" value="2017-04-26">
<collectingDate id="EFF6622268248357FB0BFE2BFAAAFE17" box="[1231,1397,474,499]" pageId="5" pageNumber="72" value="2017-04-26">26 April 2017</collectingDate>
</date>
by Weicai Chen and Yunming Mo
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C316EE8168248357FF03FDD3FCE5FDBB" pageId="5" pageNumber="72" type="etymology">
<paragraph id="8BB3BD0A68248357FF03FDD3FCE5FDBB" blockId="5.[151,1437,258,1975]" pageId="5" pageNumber="72">
<emphasis id="B978611868248357FF03FDD3FE99FDDF" bold="true" box="[199,326,546,571]" pageId="5" pageNumber="72">Etymology</emphasis>
. The specific epithet is in reference to the
<typeStatus id="54B703A868248357FC81FDD2FCAAFDDF" box="[837,885,547,571]" pageId="5" pageNumber="72">type</typeStatus>
locality of Shangsi County,
<collectingRegion id="49C873E868248357FB02FDD3FAF8FDDF" box="[1222,1319,546,571]" country="China" name="Guangxi" pageId="5" pageNumber="72">Guangxi</collectingRegion>
Province,
<collectingCountry id="F31BFD9A68248357FF53FDB7FF01FDBB" box="[151,222,582,607]" name="China" pageId="5" pageNumber="72">China</collectingCountry>
. Suggested English name is Shangsi Leaf Litter Toad.
</paragraph>
</subSubSection>
<subSubSection id="C316EE8168248357FF03FD9BFD30FC9A" pageId="5" pageNumber="72" type="diagnosis">
<paragraph id="8BB3BD0A68248357FF03FD9BFD30FC9A" blockId="5.[151,1437,258,1975]" pageId="5" pageNumber="72">
<emphasis id="B978611868248357FF03FD9BFEE5FD67" bold="true" box="[199,314,618,643]" pageId="5" pageNumber="72">Diagnosis</emphasis>
. The specimens were assigned to the genus
<taxonomicName id="4C0CC68968248357FCEEFD9AFC0AFD67" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[810,981,619,643]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="72" phylum="Chordata" rank="genus">
<emphasis id="B978611868248357FCEEFD9AFC0AFD67" box="[810,981,619,643]" italics="true" pageId="5" pageNumber="72">Leptobrachella</emphasis>
</taxonomicName>
on the basis of the following characters: small body size, having an elevated inner metacarpal tubercle, having macro-glands on body (including supraaxillary, femoral and ventrolateral glands), lacking vomerine teeth and having small tubercles on eyelids (Dubois 1980;
<bibRefCitation id="EF9DC0FB68248357FF24FD27FE1DFD0B" author="Lathrop, A. &amp; Murphy, R. W. &amp; Orlov, N. &amp; Ho, C. T." box="[224,450,726,751]" pageId="5" pageNumber="72" pagination="253 - 267" refId="ref9702" refString="Lathrop, A., Murphy, R. W., Orlov, N. &amp; Ho, C. T. (1998) Two new species of Leptolalax (Anura: Megophryidae) from northern Vietnam. Amphibia-Reptilia, 19, 253 - 267. https: // doi. org / 10.1163 / 156853898 X 00160" type="journal article" year="1998">
Lathrop
<emphasis id="B978611868248357FE87FD26FEA1FD0A" box="[323,382,726,750]" italics="true" pageId="5" pageNumber="72">et al.</emphasis>
1998
</bibRefCitation>
; Delorme
<emphasis id="B978611868248357FDF8FD26FDA8FD0A" box="[572,631,726,750]" italics="true" pageId="5" pageNumber="72">et al.</emphasis>
2006).
<taxonomicName id="4C0CC68968248357FD17FD27FBCAFD0A" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[723,1045,726,750]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="5" pageNumber="72" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B978611868248357FD17FD27FBCAFD0A" box="[723,1045,726,750]" italics="true" pageId="5" pageNumber="72">Leptobrachella shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B978611868248357FBDBFD27FBA3FD0B" bold="true" box="[1055,1148,726,751]" pageId="5" pageNumber="72">
<taxonomicNameLabel id="A24BDC6368248357FBDBFD27FBA3FD0B" box="[1055,1148,726,751]" pageId="5" pageNumber="72" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished from its congeners by a combination of (1) SVL
<quantity id="4CF410EF68248357FD98FD0AFD20FCF6" box="[604,767,763,787]" metricMagnitude="-2" metricUnit="m" metricValue="2.715" metricValueMax="2.94" metricValueMin="2.4899999999999998" pageId="5" pageNumber="72" unit="mm" value="27.15" valueMax="29.4" valueMin="24.9">24.929.4 mm</quantity>
in males (n = 11) and
<quantity id="4CF410EF68248357FC3FFD0BFB40FCF6" box="[1019,1183,762,787]" metricMagnitude="-2" metricUnit="m" metricValue="3.335" metricValueMax="3.59" metricValueMin="3.08" pageId="5" pageNumber="72" unit="mm" value="33.35" valueMax="35.9" valueMin="30.8">30.835.9 mm</quantity>
in females (n = 5), (2) black supratympanic fold with dark reddish rim, (3) ventrolateral glands distinct, forming a broken line, (4) ventral surface yellowish creamy-white with marble texture, (5) iris copper in the upper and silver in the lower fifth and (6) toe webbing basal and lateral fringes distinct, narrow.
</paragraph>
</subSubSection>
<subSubSection id="C316EE8168248355FF03FC7BFA52F849" lastPageId="7" lastPageNumber="74" pageId="5" pageNumber="72" type="description">
<paragraph id="8BB3BD0A68248357FF03FC7BFE18FA07" blockId="5.[151,1437,258,1975]" pageId="5" pageNumber="72">
<emphasis id="B978611868248357FF03FC7BFE3EFC47" bold="true" box="[199,481,906,931]" pageId="5" pageNumber="72">
Description of
<typeStatus id="54B703A868248357FEBFFC7BFE3EFC47" box="[379,481,906,931]" pageId="5" pageNumber="72" type="holotype">holotype</typeStatus>
</emphasis>
. Head width greater than head length (HW / HL = 1.15); snout bluntly rounded in profile and dorsal view, projecting slightly over lower jaw; nostril oval-shaped and closer to tip of snout than eye; canthus rostralis distinct; loreal region distinctly sloping, concave; pupil vertical; diameter of eye less than length of snout (ED / SNT = 0.78); tympanum distinct, rounded, diameter 55% that of eye (TD / ED = 0.55); tympanic rim elevated relative to skin of temporal region; vomerine teeth absent; pineal ocellus absent; vocal sac openings present, located laterally on floor of mouth; tongue large with shadow notch at posterior tip; supratympanic fold distinct, running from corner of eye to axillary gland (
<figureCitation id="1337A18F68248357FCD4FB93FC8AFB9F" box="[784,853,1122,1147]" captionStart="FIGURE 2" captionStartId="6.[151,250,1627,1649]" captionTargetBox="[167,1400,196,1584]" captionTargetId="figure@6.[167,1400,196,1584]" captionTargetPageId="6" captionText="FIGURE 2. Holotype of Leptobrachella shangsiensis sp. nov. (NHMG1704002): (A) Dorsal view, (B) dorsolateral view, (C) flank and lateral view, (D) ventral view, (E) volar view of the left hand, and (F) thenar view of the left foot." httpUri="https://zenodo.org/record/2600979/files/figure.png" pageId="5" pageNumber="72">Fig. 2</figureCitation>
AD).
<emphasis id="B978611868248357FC74FB93FBF9FB9F" bold="true" box="[944,1062,1122,1147]" italics="true" pageId="5" pageNumber="72">Forelimbs</emphasis>
. Forelimbs thin, slender; tips of fingers rounded, slightly swollen; relative fingers lengths I &lt;II &lt;IV &lt;III; nuptial pad absent; subarticular tubercles absent; a large, inner palmar tubercle separated from small, laterally compressed outer palmar tubercle; finger webbing and dermal fringes absent (
<figureCitation id="1337A18F68248357FD88FB3FFD6AFB03" box="[588,693,1230,1255]" captionStart="FIGURE 2" captionStartId="6.[151,250,1627,1649]" captionTargetBox="[167,1400,196,1584]" captionTargetId="figure@6.[167,1400,196,1584]" captionTargetPageId="6" captionText="FIGURE 2. Holotype of Leptobrachella shangsiensis sp. nov. (NHMG1704002): (A) Dorsal view, (B) dorsolateral view, (C) flank and lateral view, (D) ventral view, (E) volar view of the left hand, and (F) thenar view of the left foot." httpUri="https://zenodo.org/record/2600979/files/figure.png" pageId="5" pageNumber="72">Fig. 2 E</figureCitation>
).
<emphasis id="B978611868248357FD0BFB3FFC97FB03" bold="true" box="[719,840,1230,1255]" italics="true" pageId="5" pageNumber="72">Hindlimbs</emphasis>
. Hindlimbs slender; tips of toes rounded, slightly swollen, similar those of fingers; relative toe lengths I &lt;II &lt;V &lt;III &lt;IV; subarticular tubercles absent, replaced by distinct dermal ridges; larger, oval inner metatarsal tubercle pronounced, outer metatarsal tubercle absent; toe webbing basal; lateral fringes distinct, narrow (
<figureCitation id="1337A18F68248357FD72FACBFCC8FAB7" box="[694,791,1338,1363]" captionStart="FIGURE 2" captionStartId="6.[151,250,1627,1649]" captionTargetBox="[167,1400,196,1584]" captionTargetId="figure@6.[167,1400,196,1584]" captionTargetPageId="6" captionText="FIGURE 2. Holotype of Leptobrachella shangsiensis sp. nov. (NHMG1704002): (A) Dorsal view, (B) dorsolateral view, (C) flank and lateral view, (D) ventral view, (E) volar view of the left hand, and (F) thenar view of the left foot." httpUri="https://zenodo.org/record/2600979/files/figure.png" pageId="5" pageNumber="72">Fig. 2 F</figureCitation>
).
<emphasis id="B978611868248357FCE8FACBFBA0FAB7" bold="true" box="[812,1151,1338,1363]" italics="true" pageId="5" pageNumber="72">Skin texture and skin glands</emphasis>
. Skin on dorsum mostly smooth with numerous tiny tubercles; upper eyelid with small tubercles; ventral skin smooth; pectoral glands visible; femoral glands oval, approximately
<quantity id="4CF410EF68248357FD42FA72FD03FA7E" box="[646,732,1411,1435]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="72" unit="mm" value="1.1">1.1 mm</quantity>
in diameter, located on posteroventral surface of thighs, closer to knee than to vent; supra-axillary gland small, oval, approximately
<quantity id="4CF410EF68248357FC5EFA56FC2DFA5A" box="[922,1010,1447,1471]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="72" unit="mm" value="1.1">1.1 mm</quantity>
; ventrolateral glands distinct, forming a broken line (
<figureCitation id="1337A18F68248357FEFCFA3BFE65FA07" box="[312,442,1482,1507]" captionStart="FIGURE 2" captionStartId="6.[151,250,1627,1649]" captionTargetBox="[167,1400,196,1584]" captionTargetId="figure@6.[167,1400,196,1584]" captionTargetPageId="6" captionText="FIGURE 2. Holotype of Leptobrachella shangsiensis sp. nov. (NHMG1704002): (A) Dorsal view, (B) dorsolateral view, (C) flank and lateral view, (D) ventral view, (E) volar view of the left hand, and (F) thenar view of the left foot." httpUri="https://zenodo.org/record/2600979/files/figure.png" pageId="5" pageNumber="72">Fig. 2 C, D</figureCitation>
).
</paragraph>
<paragraph id="8BB3BD0A68248357FF03FA1FFE08F8C3" blockId="5.[151,1437,258,1975]" pageId="5" pageNumber="72">
<emphasis id="B978611868248357FF03FA1FFE2AF9E3" bold="true" box="[199,501,1518,1543]" pageId="5" pageNumber="72">
Color of
<typeStatus id="54B703A868248357FEFEFA1FFE7EF9E3" box="[314,417,1518,1543]" pageId="5" pageNumber="72" type="holotype">holotype</typeStatus>
in life
</emphasis>
. Dorsum reddish brown with dark brown markings between eyes and scapular region, and scattered some deep reddish tubercles concentrated on upper eyelid and scapular region; black supratympanic fold with dark reddish rim; canthus rostralis with black spots; lateral surfaces of trunk with creamy whitish spots, scattered with about 5 spots; transverse dark-brown bars present on dorsal surface of thighs, tibia, and lower arms; elbows and upper arms orange with light reddish-brown spots on tubercles; ventral surface yellowish creamy-white with marble texture; ventral surfaces of forelimbs and hindlimbs pinkish milk-white with dense whitish speckling; two distinct blackish-grey spots on edge of upper jaw located from snout to area under eye; supra-axillary gland milk-white; femoral and ventrolateral glands white. Iris copper in the upper and silver in the lower fifth (
<figureCitation id="1337A18F68248357FE82F8FFFE56F8C3" box="[326,393,1806,1831]" captionStart="FIGURE 2" captionStartId="6.[151,250,1627,1649]" captionTargetBox="[167,1400,196,1584]" captionTargetId="figure@6.[167,1400,196,1584]" captionTargetPageId="6" captionText="FIGURE 2. Holotype of Leptobrachella shangsiensis sp. nov. (NHMG1704002): (A) Dorsal view, (B) dorsolateral view, (C) flank and lateral view, (D) ventral view, (E) volar view of the left hand, and (F) thenar view of the left foot." httpUri="https://zenodo.org/record/2600979/files/figure.png" pageId="5" pageNumber="72">Fig. 2</figureCitation>
AD).
</paragraph>
<paragraph id="8BB3BD0A68248357FF03F8C3FC2AF853" blockId="5.[151,1437,258,1975]" pageId="5" pageNumber="72">
<emphasis id="B978611868248357FF03F8C3FD92F8AF" bold="true" box="[199,589,1842,1867]" pageId="5" pageNumber="72">
Color of
<typeStatus id="54B703A868248357FEEBF8C3FE49F8AF" box="[303,406,1842,1867]" pageId="5" pageNumber="72" type="holotype">holotype</typeStatus>
in preservative
</emphasis>
. In preservation, coloration fades to dark grey-brown on dorsum, forelimbs, and hindlimbs. Transverse dark-brown bars are distinct on forelimbs and hindlimbs. Ventral surface is creamywhite. Margin of throat and ventrolateral surfaces of forelimbs and hindlimbs are pale-brown. Macro-glands including supra-axillary glands, femoral and ventrolateral glands are creamy.
</paragraph>
<caption id="DF73ED8268278354FF53F9AAFB0AF974" httpUri="https://zenodo.org/record/2600979/files/figure.png" pageId="6" pageNumber="73" startId="6.[151,250,1627,1649]" targetBox="[167,1400,196,1584]" targetPageId="6">
<paragraph id="8BB3BD0A68278354FF53F9AAFB0AF974" blockId="6.[151,1436,1627,1680]" pageId="6" pageNumber="73">
<emphasis id="B978611868278354FF53F9AAFECCF995" bold="true" box="[151,275,1627,1649]" pageId="6" pageNumber="73">FIGURE 2.</emphasis>
Holotype of
<taxonomicName id="4C0CC68968278354FE58F9AAFD61F995" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[412,702,1627,1649]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="6" pageNumber="73" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B978611868278354FE58F9AAFD61F995" box="[412,702,1627,1649]" italics="true" pageId="6" pageNumber="73">Leptobrachella shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B978611868278354FD01F9AAFCC8F995" bold="true" box="[709,791,1627,1649]" pageId="6" pageNumber="73">
<taxonomicNameLabel id="A24BDC6368278354FD01F9AAFCC8F995" box="[709,791,1627,1649]" pageId="6" pageNumber="73" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(NHMG1704002): (A) Dorsal view, (B) dorsolateral view, (C) flank and lateral view, (D) ventral view, (E) volar view of the left hand, and (F) thenar view of the left foot.
</paragraph>
</caption>
<paragraph id="8BB3BD0A68278354FF03F924FC3BF8F6" blockId="6.[151,1436,1749,1913]" pageId="6" pageNumber="73">
<emphasis id="B978611868278354FF03F924FDCBF90A" bold="true" box="[199,532,1749,1774]" pageId="6" pageNumber="73">
Measurements of
<typeStatus id="54B703A868278354FE59F924FDDBF90A" box="[413,516,1749,1774]" pageId="6" pageNumber="73" type="holotype">holotype</typeStatus>
(
</emphasis>
mm): SVL 26.8, HL 8.1, HW 9.3, SNT 4.6, ED 3.8, IOD 2.8, TD 2.1, TED 1.3, IN 3.2, EN 1.8, TIB 12.9, FLL 13.4, THL 14.1, PL 12.0, ML 7.0, FEM 1.1.
</paragraph>
<paragraph id="8BB3BD0A68278354FF03F8ECFBBBF89D" blockId="6.[151,1436,1749,1913]" pageId="6" pageNumber="73">
<emphasis id="B978611868278354FF03F8ECFEEAF8D2" bold="true" box="[199,309,1821,1846]" pageId="6" pageNumber="73">Variation</emphasis>
. Measurements of the type series are shown in
<tableCitation id="C68E88B168278354FC89F8ECFC7CF8D2" box="[845,931,1821,1846]" captionStart="TABLE 2" captionStartId="7.[151,239,822,845]" captionText="TABLE 2. Measurements of Leptobrachella shangsiensis sp. nov. Abbreviations defined in Material and methods." pageId="6" pageNumber="73">Table 2</tableCitation>
. Females (mean SVL 33.3 ± 2.0, n = 5) have larger body size than males (mean SVL 27.4 ± 1.1, n = 11). In life, all specimens have similar coloration (
<figureCitation id="1337A18F68278354FA8DF8CEFA52F8B3" box="[1353,1421,1855,1880]" captionStart="FIGURE 3" captionStartId="7.[151,250,725,747]" captionTargetBox="[151,1436,193,704]" captionTargetId="figure@7.[151,1436,193,704]" captionTargetPageId="7" captionText="FIGURE 3. Leptobrachella shangsiensis sp. nov. in habitat (Photo taken in situ)." httpUri="https://zenodo.org/record/2600983/files/figure.png" pageId="6" pageNumber="73">Fig. 3</figureCitation>
), but in preservation, some specimens have slightly darker brown compared to
<typeStatus id="54B703A868278354FC3FF891FBBEF89D" box="[1019,1121,1888,1913]" pageId="6" pageNumber="73" type="holotype">holotype</typeStatus>
.
</paragraph>
<caption id="DF73ED8268268355FF53FD24FC0FFD0F" box="[151,976,725,747]" httpUri="https://zenodo.org/record/2600983/files/figure.png" pageId="7" pageNumber="74" startId="7.[151,250,725,747]" targetBox="[151,1436,193,704]" targetPageId="7">
<paragraph id="8BB3BD0A68268355FF53FD24FC0FFD0F" blockId="7.[151,976,725,747]" box="[151,976,725,747]" pageId="7" pageNumber="74">
<emphasis id="B978611868268355FF53FD24FECDFD0F" bold="true" box="[151,274,725,747]" pageId="7" pageNumber="74">FIGURE 3.</emphasis>
<taxonomicName id="4C0CC68968268355FEDDFD24FDE5FD0F" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[281,570,725,747]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="7" pageNumber="74" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B978611868268355FEDDFD24FDE5FD0F" box="[281,570,725,747]" italics="true" pageId="7" pageNumber="74">Leptobrachella shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B978611868268355FD84FD24FD4EFD0F" bold="true" box="[576,657,725,747]" pageId="7" pageNumber="74">
<taxonomicNameLabel id="A24BDC6368268355FD84FD24FD4EFD0F" box="[576,657,725,747]" pageId="7" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in habitat (Photo taken
<emphasis id="B978611868268355FC40FD27FC1DFD0F" box="[900,962,726,747]" italics="true" pageId="7" pageNumber="74">in situ</emphasis>
).
</paragraph>
</caption>
<caption id="DF73ED8268268355FF53FCC7FABDFCA9" box="[151,1378,821,845]" pageId="7" pageNumber="74">
<paragraph id="8BB3BD0A68268355FF53FCC7FABDFCA9" blockId="7.[151,1378,821,845]" box="[151,1378,821,845]" pageId="7" pageNumber="74">
<emphasis id="B978611868268355FF53FCC7FEDCFCA9" bold="true" box="[151,259,822,845]" pageId="7" pageNumber="74">TABLE 2</emphasis>
. Measurements of
<taxonomicName id="4C0CC68968268355FE0BFCC4FCDFFCA9" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[463,768,821,845]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="7" pageNumber="74" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B978611868268355FE0BFCC4FCDFFCA9" box="[463,768,821,845]" italics="true" pageId="7" pageNumber="74">Leptobrachella shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B978611868268355FCC2FCC7FC83FCA9" bold="true" box="[774,860,822,845]" pageId="7" pageNumber="74">
<taxonomicNameLabel id="A24BDC6368268355FCC2FCC7FC83FCA9" box="[774,860,822,845]" pageId="7" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Abbreviations defined in Material and methods.
</paragraph>
</caption>
<paragraph id="8BB3BD0A68268355FE65FC9BFAD7F907" pageId="7" pageNumber="74">
<table id="F90C4FAA68267CADFF5BFC98FAFDF907" box="[159,1314,873,1763]" gridcols="5" gridrows="22" pageId="7" pageNumber="74">
<tr id="353CBF4868267CADFF5BFC98FAFDFC9B" box="[159,1314,873,895]" gridrow="0" pageId="7" pageNumber="74" rowspan-0="1">
<th id="76EDD63468267CADFE65FC98FDF4FC9B" box="[417,555,873,895]" gridcol="1" gridrow="0" pageId="7" pageNumber="74">Range</th>
<th id="76EDD63468267CADFD5BFC98FCFEFC9B" box="[671,801,873,895]" gridcol="2" gridrow="0" pageId="7" pageNumber="74">Mean ± S.D.</th>
<th id="76EDD63468267CADFC66FC98FBE8FC9B" box="[930,1079,873,895]" gridcol="3" gridrow="0" pageId="7" pageNumber="74">Range</th>
<th id="76EDD63468267CADFB65FC98FAFDFC9B" box="[1185,1314,873,895]" gridcol="4" gridrow="0" pageId="7" pageNumber="74">Mean ± S.D.</th>
</tr>
<tr id="353CBF4868267CADFF5BFC62FAFDFC4D" box="[159,1314,915,937]" gridrow="1" pageId="7" pageNumber="74" rowspan-2="1" rowspan-4="1">
<th id="76EDD63468267CADFF5BFC62FF21FC4D" box="[159,254,915,937]" gridcol="0" gridrow="1" pageId="7" pageNumber="74">Sex</th>
<td id="76EDD63468267CADFE65FC62FDF4FC4D" box="[417,555,915,937]" gridcol="1" gridrow="1" pageId="7" pageNumber="74">Male (n = 11)</td>
<td id="76EDD63468267CADFC66FC62FBE8FC4D" box="[930,1079,915,937]" gridcol="3" gridrow="1" pageId="7" pageNumber="74">Female (n = 5)</td>
</tr>
<tr id="353CBF4868267CADFF5BFC4DFAFDFC36" box="[159,1314,956,978]" gridrow="2" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BFC4DFF21FC36" box="[159,254,956,978]" gridcol="0" gridrow="2" pageId="7" pageNumber="74">SVL</th>
<td id="76EDD63468267CADFE65FC4DFDF4FC36" box="[417,555,956,978]" gridcol="1" gridrow="2" pageId="7" pageNumber="74">24.929.4</td>
<td id="76EDD63468267CADFD5BFC4DFCFEFC36" box="[671,801,956,978]" gridcol="2" gridrow="2" pageId="7" pageNumber="74">27.4±1.1</td>
<td id="76EDD63468267CADFC66FC4DFBE8FC36" box="[930,1079,956,978]" gridcol="3" gridrow="2" pageId="7" pageNumber="74">30.835.9</td>
<td id="76EDD63468267CADFB65FC4DFAFDFC36" box="[1185,1314,956,978]" gridcol="4" gridrow="2" pageId="7" pageNumber="74">33.3±2.0</td>
</tr>
<tr id="353CBF4868267CADFF5BFC14FAFDFC1F" box="[159,1314,997,1019]" gridrow="3" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BFC14FF21FC1F" box="[159,254,997,1019]" gridcol="0" gridrow="3" pageId="7" pageNumber="74">HL</th>
<td id="76EDD63468267CADFE65FC14FDF4FC1F" box="[417,555,997,1019]" gridcol="1" gridrow="3" pageId="7" pageNumber="74">8.09.8</td>
<td id="76EDD63468267CADFD5BFC14FCFEFC1F" box="[671,801,997,1019]" gridcol="2" gridrow="3" pageId="7" pageNumber="74">8.8±0.6</td>
<td id="76EDD63468267CADFC66FC14FBE8FC1F" box="[930,1079,997,1019]" gridcol="3" gridrow="3" pageId="7" pageNumber="74">9.710.9</td>
<td id="76EDD63468267CADFB65FC14FAFDFC1F" box="[1185,1314,997,1019]" gridcol="4" gridrow="3" pageId="7" pageNumber="74">10.34±0.5</td>
</tr>
<tr id="353CBF4868267CADFF5BFBFFFAFDFBC0" box="[159,1314,1038,1060]" gridrow="4" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BFBFFFF21FBC0" box="[159,254,1038,1060]" gridcol="0" gridrow="4" pageId="7" pageNumber="74">HW</th>
<td id="76EDD63468267CADFE65FBFFFDF4FBC0" box="[417,555,1038,1060]" gridcol="1" gridrow="4" pageId="7" pageNumber="74">8.910.5</td>
<td id="76EDD63468267CADFD5BFBFFFCFEFBC0" box="[671,801,1038,1060]" gridcol="2" gridrow="4" pageId="7" pageNumber="74">9.6±0.5</td>
<td id="76EDD63468267CADFC66FBFFFBE8FBC0" box="[930,1079,1038,1060]" gridcol="3" gridrow="4" pageId="7" pageNumber="74">10.911.8</td>
<td id="76EDD63468267CADFB65FBFFFAFDFBC0" box="[1185,1314,1038,1060]" gridcol="4" gridrow="4" pageId="7" pageNumber="74">11.3±0.4</td>
</tr>
<tr id="353CBF4868267CADFF5BFBC9FAFDFBAA" box="[159,1314,1080,1102]" gridrow="5" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BFBC9FF21FBAA" box="[159,254,1080,1102]" gridcol="0" gridrow="5" pageId="7" pageNumber="74">SNT</th>
<td id="76EDD63468267CADFE65FBC9FDF4FBAA" box="[417,555,1080,1102]" gridcol="1" gridrow="5" pageId="7" pageNumber="74">3.84.8</td>
<td id="76EDD63468267CADFD5BFBC9FCFEFBAA" box="[671,801,1080,1102]" gridcol="2" gridrow="5" pageId="7" pageNumber="74">4.3±0.4</td>
<td id="76EDD63468267CADFC66FBC9FBE8FBAA" box="[930,1079,1080,1102]" gridcol="3" gridrow="5" pageId="7" pageNumber="74">4.55.0</td>
<td id="76EDD63468267CADFB65FBC9FAFDFBAA" box="[1185,1314,1080,1102]" gridcol="4" gridrow="5" pageId="7" pageNumber="74">4.8±0.2</td>
</tr>
<tr id="353CBF4868267CADFF5BFB90FAFDFB93" box="[159,1314,1121,1143]" gridrow="6" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BFB90FF21FB93" box="[159,254,1121,1143]" gridcol="0" gridrow="6" pageId="7" pageNumber="74">ED</th>
<td id="76EDD63468267CADFE65FB90FDF4FB93" box="[417,555,1121,1143]" gridcol="1" gridrow="6" pageId="7" pageNumber="74">3.54.2</td>
<td id="76EDD63468267CADFD5BFB90FCFEFB93" box="[671,801,1121,1143]" gridcol="2" gridrow="6" pageId="7" pageNumber="74">3.9±0.2</td>
<td id="76EDD63468267CADFC66FB90FBE8FB93" box="[930,1079,1121,1143]" gridcol="3" gridrow="6" pageId="7" pageNumber="74">4.24.9</td>
<td id="76EDD63468267CADFB65FB90FAFDFB93" box="[1185,1314,1121,1143]" gridcol="4" gridrow="6" pageId="7" pageNumber="74">4.6±0.3</td>
</tr>
<tr id="353CBF4868267CADFF5BFB7AFAFDFB45" box="[159,1314,1163,1185]" gridrow="7" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BFB7AFF21FB45" box="[159,254,1163,1185]" gridcol="0" gridrow="7" pageId="7" pageNumber="74">IOD</th>
<td id="76EDD63468267CADFE65FB7AFDF4FB45" box="[417,555,1163,1185]" gridcol="1" gridrow="7" pageId="7" pageNumber="74">2.53.8</td>
<td id="76EDD63468267CADFD5BFB7AFCFEFB45" box="[671,801,1163,1185]" gridcol="2" gridrow="7" pageId="7" pageNumber="74">3.0±0.4</td>
<td id="76EDD63468267CADFC66FB7AFBE8FB45" box="[930,1079,1163,1185]" gridcol="3" gridrow="7" pageId="7" pageNumber="74">3.13.8</td>
<td id="76EDD63468267CADFB65FB7AFAFDFB45" box="[1185,1314,1163,1185]" gridcol="4" gridrow="7" pageId="7" pageNumber="74">3.5±0.3</td>
</tr>
<tr id="353CBF4868267CADFF5BFB45FAFDFB2E" box="[159,1314,1204,1226]" gridrow="8" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BFB45FF21FB2E" box="[159,254,1204,1226]" gridcol="0" gridrow="8" pageId="7" pageNumber="74">TD</th>
<td id="76EDD63468267CADFE65FB45FDF4FB2E" box="[417,555,1204,1226]" gridcol="1" gridrow="8" pageId="7" pageNumber="74">1.62.2</td>
<td id="76EDD63468267CADFD5BFB45FCFEFB2E" box="[671,801,1204,1226]" gridcol="2" gridrow="8" pageId="7" pageNumber="74">1.9±0.2</td>
<td id="76EDD63468267CADFC66FB45FBE8FB2E" box="[930,1079,1204,1226]" gridcol="3" gridrow="8" pageId="7" pageNumber="74">2.12.4</td>
<td id="76EDD63468267CADFB65FB45FAFDFB2E" box="[1185,1314,1204,1226]" gridcol="4" gridrow="8" pageId="7" pageNumber="74">2.3±0.1</td>
</tr>
<tr id="353CBF4868267CADFF5BFB2CFAFDFB17" box="[159,1314,1245,1267]" gridrow="9" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BFB2CFF21FB17" box="[159,254,1245,1267]" gridcol="0" gridrow="9" pageId="7" pageNumber="74">TED</th>
<td id="76EDD63468267CADFE65FB2CFDF4FB17" box="[417,555,1245,1267]" gridcol="1" gridrow="9" pageId="7" pageNumber="74">1.01.5</td>
<td id="76EDD63468267CADFD5BFB2CFCFEFB17" box="[671,801,1245,1267]" gridcol="2" gridrow="9" pageId="7" pageNumber="74">1.2±0.1</td>
<td id="76EDD63468267CADFC66FB2CFBE8FB17" box="[930,1079,1245,1267]" gridcol="3" gridrow="9" pageId="7" pageNumber="74">1.31.4</td>
<td id="76EDD63468267CADFB65FB2CFAFDFB17" box="[1185,1314,1245,1267]" gridcol="4" gridrow="9" pageId="7" pageNumber="74">1.31±0.1</td>
</tr>
<tr id="353CBF4868267CADFF5BFAF7FAFDFAF8" box="[159,1314,1286,1308]" gridrow="10" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BFAF7FF21FAF8" box="[159,254,1286,1308]" gridcol="0" gridrow="10" pageId="7" pageNumber="74">IN</th>
<td id="76EDD63468267CADFE65FAF7FDF4FAF8" box="[417,555,1286,1308]" gridcol="1" gridrow="10" pageId="7" pageNumber="74">2.93.5</td>
<td id="76EDD63468267CADFD5BFAF7FCFEFAF8" box="[671,801,1286,1308]" gridcol="2" gridrow="10" pageId="7" pageNumber="74">3.2±0.1</td>
<td id="76EDD63468267CADFC66FAF7FBE8FAF8" box="[930,1079,1286,1308]" gridcol="3" gridrow="10" pageId="7" pageNumber="74">3.43.6</td>
<td id="76EDD63468267CADFB65FAF7FAFDFAF8" box="[1185,1314,1286,1308]" gridcol="4" gridrow="10" pageId="7" pageNumber="74">3.5±0.1</td>
</tr>
<tr id="353CBF4868267CADFF5BFAC1FAFDFAA2" box="[159,1314,1328,1350]" gridrow="11" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BFAC1FF21FAA2" box="[159,254,1328,1350]" gridcol="0" gridrow="11" pageId="7" pageNumber="74">EN</th>
<td id="76EDD63468267CADFE65FAC1FDF4FAA2" box="[417,555,1328,1350]" gridcol="1" gridrow="11" pageId="7" pageNumber="74">1.62.3</td>
<td id="76EDD63468267CADFD5BFAC1FCFEFAA2" box="[671,801,1328,1350]" gridcol="2" gridrow="11" pageId="7" pageNumber="74">1.9±0.3</td>
<td id="76EDD63468267CADFC66FAC1FBE8FAA2" box="[930,1079,1328,1350]" gridcol="3" gridrow="11" pageId="7" pageNumber="74">2.22.7</td>
<td id="76EDD63468267CADFB65FAC1FAFDFAA2" box="[1185,1314,1328,1350]" gridcol="4" gridrow="11" pageId="7" pageNumber="74">2.4±0.2</td>
</tr>
<tr id="353CBF4868267CADFF5BFAA8FAFDFA8B" box="[159,1314,1369,1391]" gridrow="12" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BFAA8FF21FA8B" box="[159,254,1369,1391]" gridcol="0" gridrow="12" pageId="7" pageNumber="74">TIB</th>
<td id="76EDD63468267CADFE65FAA8FDF4FA8B" box="[417,555,1369,1391]" gridcol="1" gridrow="12" pageId="7" pageNumber="74">11.913.8</td>
<td id="76EDD63468267CADFD5BFAA8FCFEFA8B" box="[671,801,1369,1391]" gridcol="2" gridrow="12" pageId="7" pageNumber="74">13.1±0.5</td>
<td id="76EDD63468267CADFC66FAA8FBE8FA8B" box="[930,1079,1369,1391]" gridcol="3" gridrow="12" pageId="7" pageNumber="74">14.316.8</td>
<td id="76EDD63468267CADFB65FAA8FAFDFA8B" box="[1185,1314,1369,1391]" gridcol="4" gridrow="12" pageId="7" pageNumber="74">15.6±0.9</td>
</tr>
<tr id="353CBF4868267CADFF5BFA72FAFDFA7D" box="[159,1314,1411,1433]" gridrow="13" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BFA72FF21FA7D" box="[159,254,1411,1433]" gridcol="0" gridrow="13" pageId="7" pageNumber="74">FLL</th>
<td id="76EDD63468267CADFE65FA72FDF4FA7D" box="[417,555,1411,1433]" gridcol="1" gridrow="13" pageId="7" pageNumber="74">12.414.6</td>
<td id="76EDD63468267CADFD5BFA72FCFEFA7D" box="[671,801,1411,1433]" gridcol="2" gridrow="13" pageId="7" pageNumber="74">13.4±0.6</td>
<td id="76EDD63468267CADFC66FA72FBE8FA7D" box="[930,1079,1411,1433]" gridcol="3" gridrow="13" pageId="7" pageNumber="74">15.116.5</td>
<td id="76EDD63468267CADFB65FA72FAFDFA7D" box="[1185,1314,1411,1433]" gridcol="4" gridrow="13" pageId="7" pageNumber="74">15.9±0.7</td>
</tr>
<tr id="353CBF4868267CADFF5BFA5DFAFDFA26" box="[159,1314,1452,1474]" gridrow="14" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BFA5DFF21FA26" box="[159,254,1452,1474]" gridcol="0" gridrow="14" pageId="7" pageNumber="74">THL</th>
<td id="76EDD63468267CADFE65FA5DFDF4FA26" box="[417,555,1452,1474]" gridcol="1" gridrow="14" pageId="7" pageNumber="74">12.314.5</td>
<td id="76EDD63468267CADFD5BFA5DFCFEFA26" box="[671,801,1452,1474]" gridcol="2" gridrow="14" pageId="7" pageNumber="74">13.7±0.6</td>
<td id="76EDD63468267CADFC66FA5DFBE8FA26" box="[930,1079,1452,1474]" gridcol="3" gridrow="14" pageId="7" pageNumber="74">14.119.2</td>
<td id="76EDD63468267CADFB65FA5DFAFDFA26" box="[1185,1314,1452,1474]" gridcol="4" gridrow="14" pageId="7" pageNumber="74">16.4±1.9</td>
</tr>
<tr id="353CBF4868267CADFF5BFA24FAFDFA0F" box="[159,1314,1493,1515]" gridrow="15" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BFA24FF21FA0F" box="[159,254,1493,1515]" gridcol="0" gridrow="15" pageId="7" pageNumber="74">ML</th>
<td id="76EDD63468267CADFE65FA24FDF4FA0F" box="[417,555,1493,1515]" gridcol="1" gridrow="15" pageId="7" pageNumber="74">6.17.9</td>
<td id="76EDD63468267CADFD5BFA24FCFEFA0F" box="[671,801,1493,1515]" gridcol="2" gridrow="15" pageId="7" pageNumber="74">6.9±0.4</td>
<td id="76EDD63468267CADFC66FA24FBE8FA0F" box="[930,1079,1493,1515]" gridcol="3" gridrow="15" pageId="7" pageNumber="74">7.78.8</td>
<td id="76EDD63468267CADFB65FA24FAFDFA0F" box="[1185,1314,1493,1515]" gridcol="4" gridrow="15" pageId="7" pageNumber="74">8.0±0.4</td>
</tr>
<tr id="353CBF4868267CADFF5BFA0FFAFDF9F0" box="[159,1314,1534,1556]" gridrow="16" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BFA0FFF21F9F0" box="[159,254,1534,1556]" gridcol="0" gridrow="16" pageId="7" pageNumber="74">PL</th>
<td id="76EDD63468267CADFE65FA0FFDF4F9F0" box="[417,555,1534,1556]" gridcol="1" gridrow="16" pageId="7" pageNumber="74">11.813.8</td>
<td id="76EDD63468267CADFD5BFA0FFCFEF9F0" box="[671,801,1534,1556]" gridcol="2" gridrow="16" pageId="7" pageNumber="74">12.8±0.6</td>
<td id="76EDD63468267CADFC66FA0FFBE8F9F0" box="[930,1079,1534,1556]" gridcol="3" gridrow="16" pageId="7" pageNumber="74">12.815.0</td>
<td id="76EDD63468267CADFB65FA0FFAFDF9F0" box="[1185,1314,1534,1556]" gridcol="4" gridrow="16" pageId="7" pageNumber="74">13.9±0.8</td>
</tr>
<tr id="353CBF4868267CADFF5BF9D9FAFDF9DA" box="[159,1314,1576,1598]" gridrow="17" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BF9D9FF21F9DA" box="[159,254,1576,1598]" gridcol="0" gridrow="17" pageId="7" pageNumber="74">FEM</th>
<td id="76EDD63468267CADFE65F9D9FDF4F9DA" box="[417,555,1576,1598]" gridcol="1" gridrow="17" pageId="7" pageNumber="74">0.81.8</td>
<td id="76EDD63468267CADFD5BF9D9FCFEF9DA" box="[671,801,1576,1598]" gridcol="2" gridrow="17" pageId="7" pageNumber="74">1.3±0.3</td>
<td id="76EDD63468267CADFC66F9D9FBE8F9DA" box="[930,1079,1576,1598]" gridcol="3" gridrow="17" pageId="7" pageNumber="74">1.31.8</td>
<td id="76EDD63468267CADFB65F9D9FAFDF9DA" box="[1185,1314,1576,1598]" gridcol="4" gridrow="17" pageId="7" pageNumber="74">1.6±0.2</td>
</tr>
<tr id="353CBF4868267CADFF5BF9A0FAFDF983" box="[159,1314,1617,1639]" gridrow="18" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BF9A0FF21F983" box="[159,254,1617,1639]" gridcol="0" gridrow="18" pageId="7" pageNumber="74">HL:HW</th>
<td id="76EDD63468267CADFE65F9A0FDF4F983" box="[417,555,1617,1639]" gridcol="1" gridrow="18" pageId="7" pageNumber="74">0.870.98</td>
<td id="76EDD63468267CADFD5BF9A0FCFEF983" box="[671,801,1617,1639]" gridcol="2" gridrow="18" pageId="7" pageNumber="74">0.92±0.03</td>
<td id="76EDD63468267CADFC66F9A0FBE8F983" box="[930,1079,1617,1639]" gridcol="3" gridrow="18" pageId="7" pageNumber="74">0.890.94</td>
<td id="76EDD63468267CADFB65F9A0FAFDF983" box="[1185,1314,1617,1639]" gridcol="4" gridrow="18" pageId="7" pageNumber="74">0.92±0.01</td>
</tr>
<tr id="353CBF4868267CADFF5BF98AFAFDF975" box="[159,1314,1659,1681]" gridrow="19" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BF98AFF21F975" box="[159,254,1659,1681]" gridcol="0" gridrow="19" pageId="7" pageNumber="74">HL:SVL</th>
<td id="76EDD63468267CADFE65F98AFDF4F975" box="[417,555,1659,1681]" gridcol="1" gridrow="19" pageId="7" pageNumber="74">0.290.34</td>
<td id="76EDD63468267CADFD5BF98AFCFEF975" box="[671,801,1659,1681]" gridcol="2" gridrow="19" pageId="7" pageNumber="74">0.32±0.01</td>
<td id="76EDD63468267CADFC66F98AFBE8F975" box="[930,1079,1659,1681]" gridcol="3" gridrow="19" pageId="7" pageNumber="74">0.290.34</td>
<td id="76EDD63468267CADFB65F98AFAFDF975" box="[1185,1314,1659,1681]" gridcol="4" gridrow="19" pageId="7" pageNumber="74">0.31±0.01</td>
</tr>
<tr id="353CBF4868267CADFF5BF955FAFDF95E" box="[159,1314,1700,1722]" gridrow="20" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BF955FF21F95E" box="[159,254,1700,1722]" gridcol="0" gridrow="20" pageId="7" pageNumber="74">TIB:SVL</th>
<td id="76EDD63468267CADFE65F955FDF4F95E" box="[417,555,1700,1722]" gridcol="1" gridrow="20" pageId="7" pageNumber="74">0.460.50</td>
<td id="76EDD63468267CADFD5BF955FCFEF95E" box="[671,801,1700,1722]" gridcol="2" gridrow="20" pageId="7" pageNumber="74">0.48±0.01</td>
<td id="76EDD63468267CADFC66F955FBE8F95E" box="[930,1079,1700,1722]" gridcol="3" gridrow="20" pageId="7" pageNumber="74">0.430.50</td>
<td id="76EDD63468267CADFB65F955FAFDF95E" box="[1185,1314,1700,1722]" gridcol="4" gridrow="20" pageId="7" pageNumber="74">0.47±0.02</td>
</tr>
<tr id="353CBF4868267CADFF5BF93CFAFDF907" box="[159,1314,1741,1763]" gridrow="21" pageId="7" pageNumber="74">
<th id="76EDD63468267CADFF5BF93CFF21F907" box="[159,254,1741,1763]" gridcol="0" gridrow="21" pageId="7" pageNumber="74">TD:SVL</th>
<td id="76EDD63468267CADFE65F93CFDF4F907" box="[417,555,1741,1763]" gridcol="1" gridrow="21" pageId="7" pageNumber="74">0.060.08</td>
<td id="76EDD63468267CADFD5BF93CFCFEF907" box="[671,801,1741,1763]" gridcol="2" gridrow="21" pageId="7" pageNumber="74">0.07±0.00</td>
<td id="76EDD63468267CADFC66F93CFBE8F907" box="[930,1079,1741,1763]" gridcol="3" gridrow="21" pageId="7" pageNumber="74">0.060.08</td>
<td id="76EDD63468267CADFB65F93CFAFDF907" box="[1185,1314,1741,1763]" gridcol="4" gridrow="21" pageId="7" pageNumber="74">0.07±0.00</td>
</tr>
</table>
</paragraph>
<paragraph id="8BB3BD0A68268355FF03F8D9FA52F849" blockId="7.[151,1437,1832,1965]" pageId="7" pageNumber="74">
<emphasis id="B978611868268355FF03F8D9FE74F8A5" bold="true" box="[199,427,1832,1857]" pageId="7" pageNumber="74">Advertisement call</emphasis>
. Call descriptions are on the basis of the calls of the
<typeStatus id="54B703A868268355FBE4F8D9FB5AF8A5" box="[1056,1157,1832,1857]" pageId="7" pageNumber="74" type="holotype">holotype</typeStatus>
, recorded at an ambient temperature of 21.5°C. Calls of
<taxonomicName id="4C0CC68968268355FDC3F8BCFD6BF880" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[519,692,1868,1892]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="7" pageNumber="74" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B978611868268355FDC3F8BCFD6BF880" box="[519,692,1868,1892]" italics="true" pageId="7" pageNumber="74">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B978611868268355FD7AF8BDFCC5F881" bold="true" box="[702,794,1868,1893]" pageId="7" pageNumber="74">
<taxonomicNameLabel id="A24BDC6368268355FD7AF8BDFCC5F881" box="[702,794,1868,1893]" pageId="7" pageNumber="74" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
consisted of a single note, with call duration 6469 ms (66.03 ± 1.15, n = 30), call repetition rate
<geoCoordinate id="EE38DBCD68268355FDBAF881FD6BF86C" box="[638,692,1904,1928]" direction="south" orientation="latitude" pageId="7" pageNumber="74" precision="5555" value="-3.4">3.4 s</geoCoordinate>
and intercall interval 184289 ms (250.53 ± 31.07, n = 30). The dominant frequency was between 5.56.5 kHz, but fundamental frequency and harmonics were not clear (
<figureCitation id="1337A18F68268355FAFFF865FA5FF849" box="[1339,1408,1940,1965]" captionStart="FIGURE 4" captionStartId="8.[199,298,1049,1071]" captionTargetBox="[218,1369,193,1028]" captionTargetId="figure@8.[218,1369,193,1028]" captionTargetPageId="8" captionText="FIGURE 4. The advertisement call of holotype of Leptobrachella shangsiensis sp. nov. Expanded about 2 s waveform with Oscillogram (top) and corresponding audiospectrogram (bottom). The ambient air temperature was 21.5°C." httpUri="https://zenodo.org/record/2600985/files/figure.png" pageId="7" pageNumber="74">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF73ED826829835AFF03FBE8FAD7FBB6" httpUri="https://zenodo.org/record/2600985/files/figure.png" pageId="8" pageNumber="75" startId="8.[199,298,1049,1071]" targetBox="[218,1369,193,1028]" targetPageId="8">
<paragraph id="8BB3BD0A6829835AFF03FBE8FAD7FBB6" blockId="8.[151,1436,1049,1106]" pageId="8" pageNumber="75">
<emphasis id="B97861186829835AFF03FBE8FE9BFBCB" bold="true" box="[199,324,1049,1071]" pageId="8" pageNumber="75">FIGURE 4.</emphasis>
The advertisement call of holotype of
<taxonomicName id="4C0CC6896829835AFD1FFBE8FC22FBCB" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[731,1021,1049,1071]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186829835AFD1FFBE8FC22FBCB" box="[731,1021,1049,1071]" italics="true" pageId="8" pageNumber="75">Leptobrachella shangsiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24BDC636829835AFBC1FBE8FB87FBCB" box="[1029,1112,1049,1071]" pageId="8" pageNumber="75" rank="species">
<emphasis id="B97861186829835AFBC1FBE8FB87FBCB" bold="true" box="[1029,1112,1049,1071]" pageId="8" pageNumber="75">sp. nov.</emphasis>
</taxonomicNameLabel>
Expanded about 2 s waveform with Oscillogram (top) and corresponding audiospectrogram (bottom). The ambient air temperature was 21.5°C.
</paragraph>
</caption>
<subSubSection id="C316EE816829835AFF03FB73FEB3FAE2" pageId="8" pageNumber="75" type="biology_ecology">
<paragraph id="8BB3BD0A6829835AFF03FB73FA42FB06" blockId="8.[151,1437,1154,2007]" pageId="8" pageNumber="75">
<emphasis id="B97861186829835AFF03FB73FE2CFB7F" bold="true" box="[199,499,1154,1179]" pageId="8" pageNumber="75">Ecology and distribution</emphasis>
. All specimens were found in an evergreen forest in Shiwandashan National NR located at an elevation of
<quantity id="4CF410EF6829835AFE73FB57FDE9FB5A" box="[439,566,1190,1215]" metricMagnitude="2" metricUnit="m" metricValue="5.0" metricValueMax="5.5" metricValueMin="4.5" pageId="8" pageNumber="75" unit="m" value="500.0" valueMax="550.0" valueMin="450.0">450550 m</quantity>
(
<figureCitation id="1337A18F6829835AFD82FB57FD7FFB5B" box="[582,672,1190,1215]" captionStart="FIGURE 5" captionStartId="11.[151,250,1947,1969]" captionTargetBox="[150,1436,375,1925]" captionTargetId="figure@11.[150,1436,375,1925]" captionTargetPageId="11" captionText="FIGURE 5. (A) Breeding habitat of Leptobrachella shangsiensis sp. nov. and (B) deforestation in Shiwandashan National Nature Reserve, Guangxi Province, China." httpUri="https://zenodo.org/record/2600987/files/figure.png" pageId="8" pageNumber="75">Fig. 5A</figureCitation>
). Males were calling on rocks within or adjacent to rocky streams at night between 19302400 h.
<taxonomicName id="4C0CC6896829835AFE1EFB3BFCC7FB06" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[474,792,1226,1250]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186829835AFE1EFB3BFCC7FB06" box="[474,792,1226,1250]" italics="true" pageId="8" pageNumber="75">Leptobrachella shangsiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24BDC636829835AFCDBFB3BFCA7FB07" box="[799,888,1226,1251]" pageId="8" pageNumber="75" rank="species">
<emphasis id="B97861186829835AFCDBFB3BFCA7FB07" bold="true" box="[799,888,1226,1251]" pageId="8" pageNumber="75">sp. nov.</emphasis>
</taxonomicNameLabel>
is only known from Shiwandashan National NR.
</paragraph>
<paragraph id="8BB3BD0A6829835AFF03FB1FFEB3FAE2" blockId="8.[151,1437,1154,2007]" box="[199,364,1262,1287]" pageId="8" pageNumber="75">
<emphasis id="B97861186829835AFF03FB1FFEBAFAE3" bold="true" box="[199,357,1262,1287]" pageId="8" pageNumber="75">Comparisons</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C316EE816829835EFF03FAE3FD84FE49" lastPageId="12" lastPageNumber="79" pageId="8" pageNumber="75" type="description">
<paragraph id="8BB3BD0A6829835BFF03FAE3FB5FFD61" blockId="8.[151,1437,1154,2007]" lastBlockId="9.[151,1437,151,2013]" lastPageId="9" lastPageNumber="76" pageId="8" pageNumber="75">
<taxonomicName id="4C0CC6896829835AFF03FAE3FDD6FACE" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[199,521,1298,1322]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186829835AFF03FAE3FDD6FACE" box="[199,521,1298,1322]" italics="true" pageId="8" pageNumber="75">Leptobrachella shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186829835AFDD7FAE3FDB0FACF" bold="true" box="[531,623,1298,1323]" pageId="8" pageNumber="75">
<taxonomicNameLabel id="A24BDC636829835AFDD7FAE3FDB0FACF" box="[531,623,1298,1323]" pageId="8" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from all other
<taxonomicName id="4C0CC6896829835AFCB7FAE3FBC1FACE" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[883,1054,1298,1322]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="genus">
<emphasis id="B97861186829835AFCB7FAE3FBC1FACE" box="[883,1054,1298,1322]" italics="true" pageId="8" pageNumber="75">Leptobrachella</emphasis>
</taxonomicName>
species based on morphological, molecular and acoustic data. Phylogenetically,
<taxonomicName id="4C0CC6896829835AFD6EFAC6FC88FAAA" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[682,855,1334,1358]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186829835AFD6EFAC6FC88FAAA" box="[682,855,1334,1358]" italics="true" pageId="8" pageNumber="75">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186829835AFC9AFAC7FC66FAAB" bold="true" box="[862,953,1334,1359]" pageId="8" pageNumber="75">
<taxonomicNameLabel id="A24BDC636829835AFC9AFAC7FC66FAAB" box="[862,953,1334,1359]" pageId="8" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="4C0CC6896829835AFC0CFAC6FBEFFAAA" box="[968,1072,1335,1358]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="aereus">
<emphasis id="B97861186829835AFC0CFAC6FBEFFAAA" box="[968,1072,1335,1358]" italics="true" pageId="8" pageNumber="75">L. aereus</emphasis>
</taxonomicName>
,
<emphasis id="B97861186829835AFBFBFAC6FD3CFA96" italics="true" pageId="8" pageNumber="75">
<taxonomicName id="4C0CC6896829835AFBFBFAC6FB1FFAAA" box="[1087,1216,1335,1358]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="minimus">L. minimus</taxonomicName>
,
<taxonomicName id="4C0CC6896829835AFB08FAC6FAA6FAAA" box="[1228,1401,1334,1358]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="nahangensis">L. nahangensis</taxonomicName>
,
<taxonomicName id="4C0CC6896829835AFA42FAC6FF1EFA96" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="nyx">L. nyx</taxonomicName>
,
<taxonomicName id="4C0CC6896829835AFF09FAAAFEA3FA96" box="[205,380,1370,1394]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="pelodytoides">L. pelodytoides</taxonomicName>
,
<taxonomicName id="4C0CC6896829835AFE4CFAAAFDD5FA96" box="[392,522,1370,1394]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="pluvialis">L. pluvialis</taxonomicName>
,
<taxonomicName id="4C0CC6896829835AFDD2FAAAFD3CFA96" box="[534,739,1371,1394]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="ventripunctatus">L. ventripunctatus</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0CC6896829835AFCD8FAAAFC34FA96" box="[796,1003,1370,1394]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="wuhuangmontis">
<emphasis id="B97861186829835AFCD8FAAAFC34FA96" box="[796,1003,1370,1394]" italics="true" pageId="8" pageNumber="75">L. wuhuangmontis</emphasis>
</taxonomicName>
form a clade. Genetically, among this clade, the smallest genetic distance is 5.9% (
<taxonomicName id="4C0CC6896829835AFD48FA8EFCE7FA72" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[652,824,1406,1430]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186829835AFD48FA8EFCE7FA72" box="[652,824,1406,1430]" italics="true" pageId="8" pageNumber="75">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186829835AFCFAFA8FFC47FA73" bold="true" box="[830,920,1406,1431]" pageId="8" pageNumber="75">
<taxonomicNameLabel id="A24BDC636829835AFCFAFA8FFC47FA73" box="[830,920,1406,1431]" pageId="8" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C0CC6896829835AFC0BFA8EFBCCFA72" box="[975,1043,1407,1430]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="nyx">
<emphasis id="B97861186829835AFC0BFA8EFBCCFA72" box="[975,1043,1407,1430]" italics="true" pageId="8" pageNumber="75">L. nyx</emphasis>
</taxonomicName>
) and the largest genetic distance is 10.3% (
<taxonomicName id="4C0CC6896829835AFF35FA52FE41FA5E" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[241,414,1442,1466]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186829835AFF35FA52FE41FA5E" box="[241,414,1442,1466]" italics="true" pageId="8" pageNumber="75">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186829835AFE63FA53FDDCFA5F" bold="true" box="[423,515,1442,1467]" pageId="8" pageNumber="75">
<taxonomicNameLabel id="A24BDC636829835AFE63FA53FDDCFA5F" box="[423,515,1442,1467]" pageId="8" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C0CC6896829835AFDF9FA52FCD5FA5E" box="[573,778,1443,1466]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="B97861186829835AFDF9FA52FCD5FA5E" box="[573,778,1443,1466]" italics="true" pageId="8" pageNumber="75">L. ventripunctatus</emphasis>
</taxonomicName>
). However,
<taxonomicName id="4C0CC6896829835AFC5DFA52FB99FA5E" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[921,1094,1442,1466]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186829835AFC5DFA52FB99FA5E" box="[921,1094,1442,1466]" italics="true" pageId="8" pageNumber="75">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186829835AFB8BFA53FB74FA5F" bold="true" box="[1103,1195,1442,1467]" pageId="8" pageNumber="75">
<taxonomicNameLabel id="A24BDC636829835AFB8BFA53FB74FA5F" box="[1103,1195,1442,1467]" pageId="8" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from the above-mentioned species by morphological characters. First,
<taxonomicName id="4C0CC6896829835AFC7BFA36FBF6FA3A" box="[959,1065,1479,1502]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="aereus">
<emphasis id="B97861186829835AFC7BFA36FBF6FA3A" box="[959,1065,1479,1502]" italics="true" pageId="8" pageNumber="75">L. aereus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C0CC6896829835AFB00FA36FAAFFA3A" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[1220,1392,1478,1502]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186829835AFB00FA36FAAFFA3A" box="[1220,1392,1478,1502]" italics="true" pageId="8" pageNumber="75">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186829835AFABEFA37FF17F9E7" bold="true" pageId="8" pageNumber="75">
<taxonomicNameLabel id="A24BDC636829835AFABEFA37FF17F9E7" pageId="8" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having a uniformly bronze iris (vs. iris copper in the upper and silver in the lower fifth), the higher frequency calls (6.27.9 kHz at 22.425.7°C in
<taxonomicName id="4C0CC6896829835AFD6BF9FEFCC8F9C2" box="[687,791,1551,1574]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="aereus">
<emphasis id="B97861186829835AFD6BF9FEFCC8F9C2" box="[687,791,1551,1574]" italics="true" pageId="8" pageNumber="75">L. aereus</emphasis>
</taxonomicName>
vs. 5.56.5 kHz at 21.5°C in the new species) and absent a distinct black supratympanic line and no black spots on flanks (vs. present a black supratympanic line and black spots on flanks).
<taxonomicName id="4C0CC6896829835AFE93F9A7FDB7F98A" box="[343,616,1622,1646]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="minimus">
<emphasis id="B97861186829835AFE93F9A7FDB7F98A" box="[343,616,1622,1646]" italics="true" pageId="8" pageNumber="75">Leptobrachella minimus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C0CC6896829835AFD38F9A6FC78F98A" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[764,935,1622,1646]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186829835AFD38F9A6FC78F98A" box="[764,935,1622,1646]" italics="true" pageId="8" pageNumber="75">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186829835AFC6BF9A7FBD6F98B" bold="true" box="[943,1033,1622,1647]" pageId="8" pageNumber="75">
<taxonomicNameLabel id="A24BDC636829835AFC6BF9A7FBD6F98B" box="[943,1033,1622,1647]" pageId="8" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having no lateral fringes on toes (vs. present narrow lateral fringes on toes) and iris dark gold above and grey below (vs. iris copper in the upper and silver in the lower fifth).
<taxonomicName id="4C0CC6896829835AFE76F96FFD2EF952" box="[434,753,1694,1718]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="nahangensis">
<emphasis id="B97861186829835AFE76F96FFD2EF952" box="[434,753,1694,1718]" italics="true" pageId="8" pageNumber="75">Leptobrachella nahangensis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C0CC6896829835AFC42F96EFBEDF952" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[902,1074,1694,1718]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186829835AFC42F96EFBEDF952" box="[902,1074,1694,1718]" italics="true" pageId="8" pageNumber="75">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186829835AFBFDF96FFB4CF953" bold="true" box="[1081,1171,1694,1719]" pageId="8" pageNumber="75">
<taxonomicNameLabel id="A24BDC636829835AFBFDF96FFB4CF953" box="[1081,1171,1694,1719]" pageId="8" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having a large body size (male SVL
<quantity id="4CF410EF6829835AFE8BF932FE6BF93E" box="[335,436,1731,1755]" metricMagnitude="-2" metricUnit="m" metricValue="4.08" pageId="8" pageNumber="75" unit="mm" value="40.8">40.8 mm</quantity>
vs. SVL
<quantity id="4CF410EF6829835AFDE4F932FD17F93F" box="[544,712,1731,1755]" metricMagnitude="-2" metricUnit="m" metricValue="2.715" metricValueMax="2.94" metricValueMin="2.4899999999999998" pageId="8" pageNumber="75" unit="mm" value="27.15" valueMax="29.4" valueMin="24.9">24.929.4 mm</quantity>
), a uniformly gold iris (vs. iris copper in the upper and silver in the lower fifth) and no lateral fringes on toes (vs. present narrow lateral fringes on toes).
<taxonomicName id="4C0CC6896829835AFBB0F917FA92F91A" box="[1140,1357,1766,1790]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="nyx">
<emphasis id="B97861186829835AFBB0F917FA92F91A" box="[1140,1357,1766,1790]" italics="true" pageId="8" pageNumber="75">Leptobrachella nyx</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C0CC6896829835AFF11F8FAFE5EF8C6" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[213,385,1802,1826]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186829835AFF11F8FAFE5EF8C6" box="[213,385,1802,1826]" italics="true" pageId="8" pageNumber="75">L. shangsiensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24BDC636829835AFE4EF8FBFE3AF8C7" box="[394,485,1802,1827]" pageId="8" pageNumber="75" rank="species">
<emphasis id="B97861186829835AFE4EF8FBFE3AF8C7" bold="true" box="[394,485,1802,1827]" pageId="8" pageNumber="75">sp. nov.</emphasis>
</taxonomicNameLabel>
by having a relatively large body size (SVL
<quantity id="4CF410EF6829835AFC23F8FBFB54F8C6" box="[999,1163,1802,1827]" metricMagnitude="-2" metricUnit="m" metricValue="2.965" metricValueMax="3.26" metricValueMin="2.67" pageId="8" pageNumber="75" unit="mm" value="29.65" valueMax="32.6" valueMin="26.7">26.732.6 mm</quantity>
in males and 37.041.0 mm in females vs. SVL
<quantity id="4CF410EF6829835AFE78F8DEFDBCF8A2" box="[444,611,1839,1863]" metricMagnitude="-2" metricUnit="m" metricValue="2.715" metricValueMax="2.94" metricValueMin="2.4899999999999998" pageId="8" pageNumber="75" unit="mm" value="27.15" valueMax="29.4" valueMin="24.9">24.929.4 mm</quantity>
in males and
<quantity id="4CF410EF6829835AFCCBF8DFFC69F8A2" box="[783,950,1838,1863]" metricMagnitude="-2" metricUnit="m" metricValue="3.335" metricValueMax="3.59" metricValueMin="3.08" pageId="8" pageNumber="75" unit="mm" value="33.35" valueMax="35.9" valueMin="30.8">30.835.9 mm</quantity>
in females); lateroventral glandular ridge poorly distinct (vs. distinct); dermal ridges under toes rather poorly distinct (vs. distinct); greyish-brown dorsum with dark regularly set rounded spots (vs. reddish brown dorsum); absent dermal fringe along toe V (vs. present); posterior part of thigh with glandular warts (vs. without glandular warts); HW / HL = 0.95 (vs. HW / HL = 1.15); ED / SNT = 1.00 (vs. ED / SNT = 0.78).
<taxonomicName id="4C0CC6896829835AFD47F84FFC1AF832" box="[643,965,1982,2006]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="pelodytoides">
<emphasis id="B97861186829835AFD47F84FFC1AF832" box="[643,965,1982,2006]" italics="true" pageId="8" pageNumber="75">Leptobrachella pelodytoides</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C0CC6896829835AFBA6F84EFAD0F832" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[1122,1295,1982,2006]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="8" pageNumber="75" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186829835AFBA6F84EFAD0F832" box="[1122,1295,1982,2006]" italics="true" pageId="8" pageNumber="75">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186829835AFADEF84FFAA9F833" bold="true" box="[1306,1398,1982,2007]" pageId="8" pageNumber="75">
<taxonomicNameLabel id="A24BDC636829835AFADEF84FFAA9F833" box="[1306,1398,1982,2007]" pageId="8" pageNumber="75" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having wide webbing on toes (vs. basal toe webbing in
<taxonomicName id="4C0CC6896828835BFCE6FF68FC10FF54" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[802,975,152,176]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFCE6FF68FC10FF54" box="[802,975,152,176]" italics="true" pageId="9" pageNumber="76">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFC1EFF66FBE9FF54" bold="true" box="[986,1078,151,176]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFC1EFF66FBE9FF54" box="[986,1078,151,176]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), poorly distinct dermal ridges under toes (vs. distinct) and present longitudinal skin folds on dorsum (vs. absent).
<taxonomicName id="4C0CC6896828835BFBFEFF4DFA93FF30" box="[1082,1356,188,212]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="pluvialis">
<emphasis id="B97861186828835BFBFEFF4DFA93FF30" box="[1082,1356,188,212]" italics="true" pageId="9" pageNumber="76">Leptobrachella pluvialis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C0CC6896828835BFF10FF10FE5FFF1C" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[212,384,224,248]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFF10FF10FE5FFF1C" box="[212,384,224,248]" italics="true" pageId="9" pageNumber="76">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFE42FF2EFE00FF1C" bold="true" box="[390,479,223,248]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFE42FF2EFE00FF1C" box="[390,479,223,248]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having a relatively small body size (male SVL
<quantity id="4CF410EF6828835BFBDDFF11FB63FF13" box="[1049,1212,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="2.18" metricValueMax="2.23" metricValueMin="2.13" pageId="9" pageNumber="76" unit="mm" value="21.8" valueMax="22.3" valueMin="21.3">21.322.3 mm</quantity>
vs.
<quantity id="4CF410EF6828835BFB2DFF11FA4FFF1C" box="[1257,1424,224,248]" metricMagnitude="-2" metricUnit="m" metricValue="2.715" metricValueMax="2.94" metricValueMin="2.4899999999999998" pageId="9" pageNumber="76" unit="mm" value="27.15" valueMax="29.4" valueMin="24.9">24.929.4 mm</quantity>
), absent dermal fringes on toes (vs. present), dirty white with dark brown marbling on venter (vs. yellowish creamywhite with marble texture) and dark golden iris (vs. iris copper in the upper and silver in the lower fifth).
<taxonomicName id="4C0CC6896828835BFF53FEBDFE27FE80" box="[151,504,332,356]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="ventripunctatus">
<emphasis id="B97861186828835BFF53FEBDFE27FE80" box="[151,504,332,356]" italics="true" pageId="9" pageNumber="76">Leptobrachella ventripunctatus</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C0CC6896828835BFD4AFEBCFCE5FE80" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[654,826,332,356]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFD4AFEBCFCE5FE80" box="[654,826,332,356]" italics="true" pageId="9" pageNumber="76">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFC86FEBDFC42FE81" bold="true" box="[834,925,332,357]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFC86FEBDFC42FE81" box="[834,925,332,357]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having a bicolored iris, copper above and grey-brown below (vs. iris copper in the upper and silver in the lower fifth), absent lateral fringes on toes (vs. present narrow lateral fringes on toes), chest and belly with dark brown spots (vs. yellowish creamy-white ventral surface with marble texture), present longitudinal skin folds on dorsum (vs. absent) and different dominant frequency (6.16.4 kHz at 15.0°C vs. 5.56.5 kHz at 21.5°C in
<taxonomicName id="4C0CC6896828835BFC6BFE2CFBBEFE10" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[943,1121,476,500]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFC6BFE2CFBBEFE10" box="[943,1121,476,500]" italics="true" pageId="9" pageNumber="76">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFBB4FE2DFB0DFE11" bold="true" box="[1136,1234,476,501]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFBB4FE2DFB0DFE11" box="[1136,1234,476,501]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="4C0CC6896828835BFB35FE2DFE97FDFC" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="wuhuangmontis">
<emphasis id="B97861186828835BFB35FE2DFE97FDFC" italics="true" pageId="9" pageNumber="76">Leptobrachella wuhuangmontis</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C0CC6896828835BFE1EFDF0FD5BFDFC" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[474,644,512,536]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFE1EFDF0FD5BFDFC" box="[474,644,512,536]" italics="true" pageId="9" pageNumber="76">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFD4FFE0EFD3BFDFC" bold="true" box="[651,740,511,536]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFD4FFE0EFD3BFDFC" box="[651,740,511,536]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having a greyish-white ventral surface mixed by tiny white and black dots (vs. yellowish creamy-white ventral surface with marble texture), distinct dark brown blotches on flanks from groin to axilla (vs. about 5 spots on flanks) and rough skin on dorsum body and limbs with skin ridges and dense conical tubercles (vs. nearly smooth dorsal skin with numerous tiny tubercles).
</paragraph>
<paragraph id="8BB3BD0A6828835BFF03FD61FE26FAD4" blockId="9.[151,1437,151,2013]" pageId="9" pageNumber="76">
Among the genus
<taxonomicName id="4C0CC6896828835BFE5AFD61FD96FD4C" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[414,585,656,680]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="genus">
<emphasis id="B97861186828835BFE5AFD61FD96FD4C" box="[414,585,656,680]" italics="true" pageId="9" pageNumber="76">Leptobrachella</emphasis>
</taxonomicName>
, fifteen species (
<emphasis id="B97861186828835BFCD7FD60FEF4FD14" italics="true" pageId="9" pageNumber="76">
<taxonomicName id="4C0CC6896828835BFCD7FD60FC7FFD4C" baseAuthorityName="Smith" baseAuthorityYear="1931" box="[787,928,656,680]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="baluensis">L. baluensis</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFC6BFD60FBB1FD4C" box="[943,1134,656,680]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="bondangensis">L. bondangensis</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFBB8FD60FAD6FD4C" baseAuthorityName="Dring" baseAuthorityYear="1983" box="[1148,1289,656,680]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="brevicrus">L. brevicrus</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFADCFD60FAA8FD4C" box="[1304,1399,656,680]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="fusca">L. fusca</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFA42FD60FF3EFD28" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="itiokai">L. itiokai</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFF29FD44FE4BFD28" box="[237,404,692,716]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="juliandringi">L. juliandringi</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFE5BFD44FDC2FD28" box="[415,541,692,716]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="lateralis">L. lateralis</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFDEDFD44FD71FD28" baseAuthorityName="Smith" baseAuthorityYear="1925" box="[553,686,692,716]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="mjobergi">L. mjobergi</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFD7EFD44FCEAFD28" baseAuthorityName="Gunther" baseAuthorityYear="1895" box="[698,821,693,716]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="natunae">L. natunae</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFC84FD44FC3AFD28" box="[832,997,692,716]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="nokrekensis">L. nokrekensis</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFC34FD44FBB2FD28" box="[1008,1133,692,716]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="palmata">L. palmata</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFBBDFD44FB04FD28" baseAuthorityName="Dring" baseAuthorityYear="1983" box="[1145,1243,693,716]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="parva">L. parva</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFB22FD44FA47FD28" box="[1254,1432,692,716]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="platycephala">L. platycephala</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFF53FD28FEF4FD14" baseAuthorityName="Dring" baseAuthorityYear="1983" box="[151,299,729,752]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="serasanae">L. serasanae</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0CC6896828835BFEAEFD28FE0BFD14" box="[362,468,728,752]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="tamdil">
<emphasis id="B97861186828835BFEAEFD28FE0BFD14" box="[362,468,728,752]" italics="true" pageId="9" pageNumber="76">L. tamdil</emphasis>
</taxonomicName>
) have no molecular data. Geographically, these species occur in allopatry with
<taxonomicName id="4C0CC6896828835BFA41FD28FEFAFCF0" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFA41FD28FEFAFCF0" italics="true" pageId="9" pageNumber="76">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFEEFFD0DFE5AFCF1" bold="true" box="[299,389,764,789]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFEEFFD0DFE5AFCF1" box="[299,389,764,789]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
<emphasis id="B97861186828835BFE48FD0DFF06FCDC" italics="true" pageId="9" pageNumber="76">
<taxonomicName id="4C0CC6896828835BFE48FD0DFD74FCF0" baseAuthorityName="Smith" baseAuthorityYear="1931" box="[396,683,764,788]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="baluensis">Leptobrachella baluensis</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFD72FD0CFCE1FCF0" baseAuthorityName="Dring" baseAuthorityYear="1983" box="[694,830,764,788]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="brevicrus">L. brevicrus</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFC8EFD0CFC6EFCF0" box="[842,945,764,788]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="itiokai">L. itiokai</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFC79FD0CFBBBFCF0" box="[957,1124,764,788]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="juliandringi">L. juliandringi</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFBABFD0CFB2CFCF0" baseAuthorityName="Smith" baseAuthorityYear="1925" box="[1135,1267,764,788]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="mjobergi">L. mjobergi</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFB3AFD0CFAA4FCF0" box="[1278,1403,764,788]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="palmata">L. palmata</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFA42FD0CFF06FCDC" baseAuthorityName="Dring" baseAuthorityYear="1983" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="parva">L. parva</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0CC6896828835BFED2FCD0FE17FCDC" box="[278,456,800,824]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="platycephala">
<emphasis id="B97861186828835BFED2FCD0FE17FCDC" box="[278,456,800,824]" italics="true" pageId="9" pageNumber="76">L. platycephala</emphasis>
</taxonomicName>
come from
<collectingCountry id="F31BFD9A6828835BFD9FFCEEFD1AFCDC" box="[603,709,799,824]" name="Malaysia" pageId="9" pageNumber="76">Malaysia</collectingCountry>
;
<emphasis id="B97861186828835BFD10FCD0FB55FCDC" box="[724,1162,800,824]" italics="true" pageId="9" pageNumber="76">
<taxonomicName id="4C0CC6896828835BFD10FCD0FC4DFCDC" box="[724,914,800,824]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="bondangensis">L. bondangensis</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFC65FCD0FBDFFCDC" box="[929,1024,800,824]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="fusca">L. fusca</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFBCBFCD0FB55FCDC" baseAuthorityName="Gunther" baseAuthorityYear="1895" box="[1039,1162,801,824]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="natunae">L. natunae</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0CC6896828835BFB0CFCD0FA84FCDC" baseAuthorityName="Dring" baseAuthorityYear="1983" box="[1224,1371,801,824]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="serasanae">
<emphasis id="B97861186828835BFB0CFCD0FA84FCDC" box="[1224,1371,801,824]" italics="true" pageId="9" pageNumber="76">L. serasanae</emphasis>
</taxonomicName>
from
<collectingCountry id="F31BFD9A6828835BFF53FCB5FED7FCB9" box="[151,264,836,861]" name="Indonesia" pageId="9" pageNumber="76">Indonesia</collectingCountry>
;
<emphasis id="B97861186828835BFED6FCB4FDE5FCB8" box="[274,570,836,860]" italics="true" pageId="9" pageNumber="76">
<taxonomicName id="4C0CC6896828835BFED6FCB4FE50FCB8" box="[274,399,836,860]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="lateralis">L. lateralis</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFE5EFCB4FDE5FCB8" box="[410,570,836,860]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="nokrekensis">L. nokrekensis</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0CC6896828835BFDABFCB4FD0CFCB8" box="[623,723,836,860]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="tamdil">
<emphasis id="B97861186828835BFDABFCB4FD0CFCB8" box="[623,723,836,860]" italics="true" pageId="9" pageNumber="76">L. tamdil</emphasis>
</taxonomicName>
from northeast
<collectingCountry id="F31BFD9A6828835BFC40FCB5FC1EFCB9" box="[900,961,836,861]" name="India" pageId="9" pageNumber="76">India</collectingCountry>
. Morphologically,
<taxonomicName id="4C0CC6896828835BFB50FCB4FAC4FCB8" baseAuthorityName="Smith" baseAuthorityYear="1931" box="[1172,1307,836,860]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="baluensis">
<emphasis id="B97861186828835BFB50FCB4FAC4FCB8" box="[1172,1307,836,860]" italics="true" pageId="9" pageNumber="76">L. baluensis</emphasis>
</taxonomicName>
(male SVL
<quantity id="4CF410EF6828835BFF53FC96FE9FFC64" box="[151,320,871,896]" metricMagnitude="-2" metricUnit="m" metricValue="1.54" metricValueMax="1.59" metricValueMin="1.49" pageId="9" pageNumber="76" unit="mm" value="15.4" valueMax="15.9" valueMin="14.9">14.915.9 mm</quantity>
),
<taxonomicName id="4C0CC6896828835BFE91FC98FDD0FC64" box="[341,527,872,896]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="bondangensis">
<emphasis id="B97861186828835BFE91FC98FDD0FC64" box="[341,527,872,896]" italics="true" pageId="9" pageNumber="76">L. bondangensis</emphasis>
</taxonomicName>
(male SVL
<quantity id="4CF410EF6828835BFD64FC99FCD4FC64" box="[672,779,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="9" pageNumber="76" unit="mm" value="17.8">17.8 mm</quantity>
),
<taxonomicName id="4C0CC6896828835BFCE4FC98FC76FC64" baseAuthorityName="Dring" baseAuthorityYear="1983" box="[800,937,872,896]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="brevicrus">
<emphasis id="B97861186828835BFCE4FC98FC76FC64" box="[800,937,872,896]" italics="true" pageId="9" pageNumber="76">L. brevicrus</emphasis>
</taxonomicName>
(male SVL
<quantity id="4CF410EF6828835BFBFDFC99FB3CFC64" box="[1081,1251,872,896]" metricMagnitude="-2" metricUnit="m" metricValue="1.745" metricValueMax="1.78" metricValueMin="1.71" pageId="9" pageNumber="76" unit="mm" value="17.45" valueMax="17.8" valueMin="17.1">17.117.8 mm</quantity>
),
<taxonomicName id="4C0CC6896828835BFB3CFC98FA8BFC64" box="[1272,1364,872,896]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="fusca">
<emphasis id="B97861186828835BFB3CFC98FA8BFC64" box="[1272,1364,872,896]" italics="true" pageId="9" pageNumber="76">L. fusca</emphasis>
</taxonomicName>
(male SVL
<quantity id="4CF410EF6828835BFF10FC7DFEE3FC40" box="[212,316,908,933]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="9" pageNumber="76" unit="mm" value="16.3">16.3 mm</quantity>
),
<taxonomicName id="4C0CC6896828835BFE94FC7CFE69FC40" box="[336,438,908,932]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="itiokai">
<emphasis id="B97861186828835BFE94FC7CFE69FC40" box="[336,438,908,932]" italics="true" pageId="9" pageNumber="76">L. itiokai</emphasis>
</taxonomicName>
(male SVL
<quantity id="4CF410EF6828835BFD87FC7DFD34FC40" box="[579,747,908,933]" metricMagnitude="-2" metricUnit="m" metricValue="1.595" metricValueMax="1.67" metricValueMin="1.52" pageId="9" pageNumber="76" unit="mm" value="15.95" valueMax="16.7" valueMin="15.2">15.216.7 mm</quantity>
),
<taxonomicName id="4C0CC6896828835BFD3BFC7CFC7CFC40" box="[767,931,908,932]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="juliandringi">
<emphasis id="B97861186828835BFD3BFC7CFC7CFC40" box="[767,931,908,932]" italics="true" pageId="9" pageNumber="76">L. juliandringi</emphasis>
</taxonomicName>
(SVL17.0
<quantity id="4CF410EF6828835BFBECFC7DFB52FC40" box="[1064,1165,908,932]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" pageId="9" pageNumber="76" unit="mm" value="17.2">17.2 mm</quantity>
in males and
<quantity id="4CF410EF6828835BFAE8FC7DFF1CFC23" metricMagnitude="-2" metricUnit="m" metricValue="1.9" metricValueMax="1.91" metricValueMin="1.89" pageId="9" pageNumber="76" unit="mm" value="19.0" valueMax="19.1" valueMin="18.9">18.919.1 mm</quantity>
in females),
<taxonomicName id="4C0CC6896828835BFE92FC40FE07FC2C" baseAuthorityName="Smith" baseAuthorityYear="1925" box="[342,472,944,968]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="mjobergi">
<emphasis id="B97861186828835BFE92FC40FE07FC2C" box="[342,472,944,968]" italics="true" pageId="9" pageNumber="76">L. mjobergi</emphasis>
</taxonomicName>
(male SVL 15.719.0 mm),
<taxonomicName id="4C0CC6896828835BFCDBFC40FC48FC2C" baseAuthorityName="Gunther" baseAuthorityYear="1895" box="[799,919,945,968]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="natunae">
<emphasis id="B97861186828835BFCDBFC40FC48FC2C" box="[799,919,945,968]" italics="true" pageId="9" pageNumber="76">L. natunae</emphasis>
</taxonomicName>
(male SVL
<quantity id="4CF410EF6828835BFBE6FC5EFB54FC2C" box="[1058,1163,943,968]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" pageId="9" pageNumber="76" unit="mm" value="17.6">17.6 mm</quantity>
),
<taxonomicName id="4C0CC6896828835BFB5AFC40FAC7FC2C" box="[1182,1304,944,968]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="palmata">
<emphasis id="B97861186828835BFB5AFC40FAC7FC2C" box="[1182,1304,944,968]" italics="true" pageId="9" pageNumber="76">L. palmata</emphasis>
</taxonomicName>
(male SVL
<quantity id="4CF410EF6828835BFF53FC25FEE1FC08" box="[151,318,980,1005]" metricMagnitude="-2" metricUnit="m" metricValue="1.56" metricValueMax="1.68" metricValueMin="1.44" pageId="9" pageNumber="76" unit="mm" value="15.6" valueMax="16.8" valueMin="14.4">14.416.8 mm</quantity>
),
<taxonomicName id="4C0CC6896828835BFE94FC24FE6FFC08" baseAuthorityName="Dring" baseAuthorityYear="1983" box="[336,432,981,1004]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="parva">
<emphasis id="B97861186828835BFE94FC24FE6FFC08" box="[336,432,981,1004]" italics="true" pageId="9" pageNumber="76">L. parva</emphasis>
</taxonomicName>
(SVL15.0
<quantity id="4CF410EF6828835BFDF0FC25FD48FC08" box="[564,663,980,1005]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="9" pageNumber="76" unit="mm" value="16.9">16.9 mm</quantity>
in males and
<quantity id="4CF410EF6828835BFCF5FC25FC4AFC08" box="[817,917,980,1004]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" pageId="9" pageNumber="76" unit="mm" value="17.8">17.8 mm</quantity>
in female) and
<taxonomicName id="4C0CC6896828835BFB80FC24FB0CFC08" baseAuthorityName="Dring" baseAuthorityYear="1983" box="[1092,1235,981,1004]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="serasanae">
<emphasis id="B97861186828835BFB80FC24FB0CFC08" box="[1092,1235,981,1004]" italics="true" pageId="9" pageNumber="76">L. serasanae</emphasis>
</taxonomicName>
(female
<quantity id="4CF410EF6828835BFAF5FC25FA4BFC08" box="[1329,1428,980,1005]" metricMagnitude="-2" metricUnit="m" metricValue="1.69" pageId="9" pageNumber="76" unit="mm" value="16.9">16.9 mm</quantity>
) have a small body size (vs. SVL
<quantity id="4CF410EF6828835BFDC1FC09FD77FBEB" box="[517,680,1016,1040]" metricMagnitude="-2" metricUnit="m" metricValue="2.715" metricValueMax="2.94" metricValueMin="2.4899999999999998" pageId="9" pageNumber="76" unit="mm" value="27.15" valueMax="29.4" valueMin="24.9">24.929.4 mm</quantity>
in males and
<quantity id="4CF410EF6828835BFC86FC06FC3BFBEB" box="[834,996,1015,1040]" metricMagnitude="-2" metricUnit="m" metricValue="3.335" metricValueMax="3.59" metricValueMin="3.08" pageId="9" pageNumber="76" unit="mm" value="33.35" valueMax="35.9" valueMin="30.8">30.835.9 mm</quantity>
in females in
<taxonomicName id="4C0CC6896828835BFB45FC08FAF3FBF4" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[1153,1324,1016,1040]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFB45FC08FAF3FBF4" box="[1153,1324,1016,1040]" italics="true" pageId="9" pageNumber="76">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFAF6FC06FA54FBF4" bold="true" box="[1330,1419,1015,1040]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFAF6FC06FA54FBF4" box="[1330,1419,1015,1040]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). Furthermore,
<emphasis id="B97861186828835BFEF2FBECFE31FBBC" italics="true" pageId="9" pageNumber="76">
<taxonomicName id="4C0CC6896828835BFEF2FBECFE1BFBD0" baseAuthorityName="Smith" baseAuthorityYear="1931" box="[310,452,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="baluensis">L. baluensis</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFE11FBECFDBCFBD0" baseAuthorityName="Dring" baseAuthorityYear="1983" box="[469,611,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="brevicrus">L. brevicrus</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFDB7FBECFCECFBD0" box="[627,819,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="bondangensis">L. bondangensis</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFC87FBECFC7CFBD0" box="[835,931,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="fusca">L. fusca</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFC77FBECFBFFFBD0" box="[947,1056,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="itiokai">L. itiokai</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFBF5FBECFB03FBD0" box="[1073,1244,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="juliandringi">L. juliandringi</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFB28FBECFAA9FBD0" baseAuthorityName="Smith" baseAuthorityYear="1925" box="[1260,1398,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="mjobergi">L. mjobergi</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFA42FBECFF2BFBBC" baseAuthorityName="Gunther" baseAuthorityYear="1895" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="natunae">L. natunae</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFEC5FBB0FE5FFBBC" box="[257,384,1088,1112]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="palmata">L. palmata</taxonomicName>
,
<taxonomicName id="4C0CC6896828835BFE49FBB0FE31FBBC" baseAuthorityName="Dring" baseAuthorityYear="1983" box="[397,494,1089,1112]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="parva">L. parva</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0CC6896828835BFDEDFBB0FD64FBBC" baseAuthorityName="Dring" baseAuthorityYear="1983" box="[553,699,1089,1112]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="serasanae">
<emphasis id="B97861186828835BFDEDFBB0FD64FBBC" box="[553,699,1089,1112]" italics="true" pageId="9" pageNumber="76">L. serasanae</emphasis>
</taxonomicName>
exhibit an arrow-like projection of digital tip (vs. no arrow-like projection of digital tip).
<taxonomicName id="4C0CC6896828835BFE72FB95FD19FB98" box="[438,710,1124,1148]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="lateralis">
<emphasis id="B97861186828835BFE72FB95FD19FB98" box="[438,710,1124,1148]" italics="true" pageId="9" pageNumber="76">Leptobrachella lateralis</emphasis>
</taxonomicName>
has no lateral fringes on toes (vs. present narrow lateral fringes on toes).
<taxonomicName id="4C0CC6896828835BFF39FB79FDEDFB44" box="[253,562,1160,1184]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="nokrekensis">
<emphasis id="B97861186828835BFF39FB79FDEDFB44" box="[253,562,1160,1184]" italics="true" pageId="9" pageNumber="76">Leptobrachella nokrekensis</emphasis>
</taxonomicName>
has reddish orange irises in the upper (vs. irises copper in the upper and silver in the lower fifth).
<taxonomicName id="4C0CC6896828835BFEB1FB5DFD19FB20" baseAuthorityName="Dehling" baseAuthorityYear="2012" box="[373,710,1196,1220]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="B97861186828835BFEB1FB5DFD19FB20" box="[373,710,1196,1220]" italics="true" pageId="9" pageNumber="76">Leptobrachella platycephalus</emphasis>
</taxonomicName>
has a large body size (male SVL
<quantity id="4CF410EF6828835BFB90FB5DFB64FB20" box="[1108,1211,1196,1221]" metricMagnitude="-2" metricUnit="m" metricValue="3.51" pageId="9" pageNumber="76" unit="mm" value="35.1">35.1 mm</quantity>
vs. SVL
<quantity id="4CF410EF6828835BFAE8FB5DFF1BFB0C" metricMagnitude="-2" metricUnit="m" metricValue="2.715" metricValueMax="2.94" metricValueMin="2.4899999999999998" pageId="9" pageNumber="76" unit="mm" value="27.15" valueMax="29.4" valueMin="24.9">24.929.4 mm</quantity>
) and orange red or reddish brown irises (vs. irises copper in the upper and silver in the lower fifth).
<taxonomicName id="4C0CC6896828835BFF53FB05FE50FAE8" box="[151,399,1268,1292]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="tamdil">
<emphasis id="B97861186828835BFF53FB05FE50FAE8" box="[151,399,1268,1292]" italics="true" pageId="9" pageNumber="76">Leptobrachella tamdil</emphasis>
</taxonomicName>
has wide lateral fringes on toes (vs. present narrow lateral fringes on toes) and wide webbing on toes (vs. basal toe webbing).
</paragraph>
<paragraph id="8BB3BD0A68288358FF03FACDFF0EFB20" blockId="9.[151,1437,151,2013]" lastBlockId="10.[151,1437,151,2013]" lastPageId="10" lastPageNumber="77" pageId="9" pageNumber="76">
<taxonomicName id="4C0CC6896828835BFF03FACDFDDAFAB0" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[199,517,1340,1364]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFF03FACDFDDAFAB0" box="[199,517,1340,1364]" italics="true" pageId="9" pageNumber="76">Leptobrachella shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFDC8FACDFDBAFAB1" bold="true" box="[524,613,1340,1365]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFDC8FACDFDBAFAB1" box="[524,613,1340,1365]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
occurs in sympatry with
<emphasis id="B97861186828835BFCBAFACCFC01FAB0" box="[894,990,1341,1364]" italics="true" pageId="9" pageNumber="76">
<taxonomicName id="4C0CC6896828835BFCBAFACCFC05FAB0" box="[894,986,1341,1364]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="sungi">L. sungi</taxonomicName>
,
</emphasis>
but
<taxonomicName id="4C0CC6896828835BFBCBFACCFBB6FAB0" box="[1039,1129,1341,1364]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="sungi">
<emphasis id="B97861186828835BFBCBFACCFBB6FAB0" box="[1039,1129,1341,1364]" italics="true" pageId="9" pageNumber="76">L. sungi</emphasis>
</taxonomicName>
has a large body size (male SVL
<quantity id="4CF410EF6828835BFF10FAAEFEA6FA93" box="[212,377,1375,1400]" metricMagnitude="-2" metricUnit="m" metricValue="5.05" metricValueMax="5.27" metricValueMin="4.83" pageId="9" pageNumber="76" unit="mm" value="50.5" valueMax="52.7" valueMin="48.3">48.352.7 mm</quantity>
vs. SVL
<quantity id="4CF410EF6828835BFE2CFA91FD4EFA9C" box="[488,657,1376,1400]" metricMagnitude="-2" metricUnit="m" metricValue="2.715" metricValueMax="2.94" metricValueMin="2.4899999999999998" pageId="9" pageNumber="76" unit="mm" value="27.15" valueMax="29.4" valueMin="24.9">24.929.4 mm</quantity>
), gold green iris (vs. iris copper in the upper and silver in the lower fifth), absent a distinct black supratympanic line (vs. present) and absent black spots on flanks (vs. present).
<taxonomicName id="4C0CC6896828835BFF53FA59FEA8FA24" box="[151,375,1448,1472]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="laui">
<emphasis id="B97861186828835BFF53FA59FEA8FA24" box="[151,375,1448,1472]" italics="true" pageId="9" pageNumber="76">Leptobrachella laui</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0CC6896828835BFE75FA59FE08FA24" box="[433,471,1448,1472]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="liui">
<emphasis id="B97861186828835BFE75FA59FE08FA24" box="[433,471,1448,1472]" italics="true" pageId="9" pageNumber="76">liui</emphasis>
</taxonomicName>
have a relatively widespread distribution in
<collectingCountry id="F31BFD9A6828835BFC13FA56FBC1FA24" box="[983,1054,1447,1472]" name="China" pageId="9" pageNumber="76">China</collectingCountry>
. However,
<taxonomicName id="4C0CC6896828835BFB64FA58FB34FA24" box="[1184,1259,1448,1472]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="laui">
<emphasis id="B97861186828835BFB64FA58FB34FA24" box="[1184,1259,1448,1472]" italics="true" pageId="9" pageNumber="76">L. laui</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C0CC6896828835BFA41FA58FEFAFA00" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFA41FA58FEFAFA00" italics="true" pageId="9" pageNumber="76">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFEE8FA3DFE5AFA01" bold="true" box="[300,389,1484,1509]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFEE8FA3DFE5AFA01" box="[300,389,1484,1509]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having no black supratympanic fold (vs. present), moderate dermal fringes on fingers (vs. absent), throat transparent pink with brown dusting along anterior margin (vs. throat yellowish creamy-white with marble texture) and head slightly longer than wide (vs. HW / HL = 1.15).
<taxonomicName id="4C0CC6896828835BFBCEF9E5FB3AF9C8" box="[1034,1253,1556,1580]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="liui">
<emphasis id="B97861186828835BFBCEF9E5FB3AF9C8" box="[1034,1253,1556,1580]" italics="true" pageId="9" pageNumber="76">Leptobrachella liui</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C0CC6896828835BFA41F9E4FEFAF9B4" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFA41F9E4FEFAF9B4" italics="true" pageId="9" pageNumber="76">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFEEFF9C6FE5BF9B4" bold="true" box="[299,388,1591,1616]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFEEFF9C6FE5BF9B4" box="[299,388,1591,1616]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
by having wide lateral fringes on toes (vs. narrow lateral fringes on toes), creamy-white belly with dark brown spots on chest and margins (vs. yellowish creamy-white belly with marble texture), longitudinal ridges under toes interrupted at the articulations (vs. not interrupted) and greyish-brown dorsal coloration in life (vs. brown or reddish-brown dorsal coloration). From the rest 11 known
<taxonomicName id="4C0CC6896828835BFC26F955FB52F958" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[994,1165,1700,1724]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="genus">
<emphasis id="B97861186828835BFC26F955FB52F958" box="[994,1165,1700,1724]" italics="true" pageId="9" pageNumber="76">Leptobrachella</emphasis>
</taxonomicName>
species from
<collectingCountry id="F31BFD9A6828835BFAF6F955FAA7F959" box="[1330,1400,1700,1725]" name="China" pageId="9" pageNumber="76">China</collectingCountry>
,
<taxonomicName id="4C0CC6896828835BFA41F954FEFAF904" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFA41F954FEFAF904" italics="true" pageId="9" pageNumber="76">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFEEBF936FE53F904" bold="true" box="[303,396,1735,1760]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFEEBF936FE53F904" box="[303,396,1735,1760]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from
<taxonomicName id="4C0CC6896828835BFDEEF938FD42F904" box="[554,669,1736,1760]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="alpinus">
<emphasis id="B97861186828835BFDEEF938FD42F904" box="[554,669,1736,1760]" italics="true" pageId="9" pageNumber="76">L. alpinus</emphasis>
</taxonomicName>
by having a different dominant frequency (6.7 kHz at 16°C in
<taxonomicName id="4C0CC6896828835BFA41F938FF35F8E0" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="alpinus">
<emphasis id="B97861186828835BFA41F938FF35F8E0" italics="true" pageId="9" pageNumber="76">L. alpinus</emphasis>
</taxonomicName>
vs. 5.56.5 kHz at 21.5°C in
<taxonomicName id="4C0CC6896828835BFDFDF91CFD3BF8E0" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[569,740,1772,1796]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFDFDF91CFD3BF8E0" box="[569,740,1772,1796]" italics="true" pageId="9" pageNumber="76">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFD29F91DFC98F8E1" bold="true" box="[749,839,1772,1797]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFD29F91DFC98F8E1" box="[749,839,1772,1797]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and wide lateral fringes on toes (vs. present narrow lateral fringes on toes in
<taxonomicName id="4C0CC6896828835BFE7EF8E0FDB7F8CC" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[442,616,1808,1832]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFE7EF8E0FDB7F8CC" box="[442,616,1808,1832]" italics="true" pageId="9" pageNumber="76">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFDB6F8FEFD11F8CC" bold="true" box="[626,718,1807,1832]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFDB6F8FEFD11F8CC" box="[626,718,1807,1832]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="4C0CC6896828835BFD2DF8E1FC27F8CC" box="[745,1016,1808,1832]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="bourreti">
<emphasis id="B97861186828835BFD2DF8E1FC27F8CC" box="[745,1016,1808,1832]" italics="true" pageId="9" pageNumber="76">Leptobrachella bourreti</emphasis>
</taxonomicName>
exhibits a copper above and golden below iris (vs. iris copper in the upper and silver in the lower fifth), a relatively large body size (male SVL 28.0
<quantity id="4CF410EF6828835BFF53F8A6FF21F894" box="[151,254,1879,1904]" metricMagnitude="-2" metricUnit="m" metricValue="3.62" pageId="9" pageNumber="76" unit="mm" value="36.2">36.2 mm</quantity>
vs. SVL
<quantity id="4CF410EF6828835BFEB5F8A9FDC8F894" box="[369,535,1880,1904]" metricMagnitude="-2" metricUnit="m" metricValue="2.715" metricValueMax="2.94" metricValueMin="2.4899999999999998" pageId="9" pageNumber="76" unit="mm" value="27.15" valueMax="29.4" valueMin="24.9">24.929.4 mm</quantity>
in
<taxonomicName id="4C0CC6896828835BFD84F8A8FD30F894" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[576,751,1880,1904]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFD84F8A8FD30F894" box="[576,751,1880,1904]" italics="true" pageId="9" pageNumber="76">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFD3DF8A6FC8AF894" bold="true" box="[761,853,1879,1904]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFD3DF8A6FC8AF894" box="[761,853,1879,1904]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and poorly distinct dermal ridges under toes (vs. distinct).
<taxonomicName id="4C0CC6896828835BFEC4F88DFE08F870" box="[256,471,1916,1940]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="eos">
<emphasis id="B97861186828835BFEC4F88DFE08F870" box="[256,471,1916,1940]" italics="true" pageId="9" pageNumber="76">Leptobrachella eos</emphasis>
</taxonomicName>
has a relatively large body size (male SVL
<quantity id="4CF410EF6828835BFC79F88DFB80F870" box="[957,1119,1916,1940]" metricMagnitude="-2" metricUnit="m" metricValue="3.39" metricValueMax="3.47" metricValueMin="3.31" pageId="9" pageNumber="76" unit="mm" value="33.9" valueMax="34.7" valueMin="33.1">33.134.7 mm</quantity>
in
<taxonomicName id="4C0CC6896828835BFB45F88CFB1CF870" box="[1153,1219,1917,1940]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="eos">
<emphasis id="B97861186828835BFB45F88CFB1CF870" box="[1153,1219,1917,1940]" italics="true" pageId="9" pageNumber="76">L. eos</emphasis>
</taxonomicName>
vs. SVL
<quantity id="4CF410EF6828835BFAE8F88DFF1CF85C" metricMagnitude="-2" metricUnit="m" metricValue="2.715" metricValueMax="2.94" metricValueMin="2.4899999999999998" pageId="9" pageNumber="76" unit="mm" value="27.15" valueMax="29.4" valueMin="24.9">24.929.4 mm</quantity>
in
<taxonomicName id="4C0CC6896828835BFF23F850FE4DF85C" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[231,402,1952,1976]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFF23F850FE4DF85C" box="[231,402,1952,1976]" italics="true" pageId="9" pageNumber="76">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFE5FF851FE2AF85D" bold="true" box="[411,501,1952,1977]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFE5FF851FE2AF85D" box="[411,501,1952,1977]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), absent black spots on flanks (vs. present), iris orange above and light gold below (vs. iris copper in the upper and silver in the lower fifth in
<taxonomicName id="4C0CC6896828835BFCE8F834FC08F838" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[812,983,1988,2012]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="9" pageNumber="76" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B97861186828835BFCE8F834FC08F838" box="[812,983,1988,2012]" italics="true" pageId="9" pageNumber="76">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B97861186828835BFC1BF835FBE6F839" bold="true" box="[991,1081,1988,2013]" pageId="9" pageNumber="76">
<taxonomicNameLabel id="A24BDC636828835BFC1BF835FBE6F839" box="[991,1081,1988,2013]" pageId="9" pageNumber="76" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), absent a black supratympanic line (vs. present) and wide lateral fringes on toes (vs. present narrow lateral fringes on toes in
<taxonomicName id="4C0CC689682B8358FB02FF68FAADFF54" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[1222,1394,152,176]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FB02FF68FAADFF54" box="[1222,1394,152,176]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FABEFF66FF17FF31" bold="true" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FABEFF66FF17FF31" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="4C0CC689682B8358FF25FF4DFD9FFF30" box="[225,576,188,212]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="mangshanensis">
<emphasis id="B9786118682B8358FF25FF4DFD9FFF30" box="[225,576,188,212]" italics="true" pageId="10" pageNumber="77">Leptobrachella mangshanensis</emphasis>
</taxonomicName>
has a bicolored iris with bright orange upper and greyish cream below (vs. iris copper in the upper and silver in the lower fifth in
<taxonomicName id="4C0CC689682B8358FD33FF10FC7DFF1C" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[759,930,224,248]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FD33FF10FC7DFF1C" box="[759,930,224,248]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FC6DFF2EFBDCFF1C" bold="true" box="[937,1027,223,248]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FC6DFF2EFBDCFF1C" box="[937,1027,223,248]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and nearly smooth dorsal skin with some small, orange, tubercles and irregular, dark brown stripes (vs. nearly smooth dorsal skin with numerous tiny tubercles in
<taxonomicName id="4C0CC689682B8358FEE6FED8FE11FEA4" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[290,462,296,320]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FEE6FED8FE11FEA4" box="[290,462,296,320]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FE13FED6FDEEFEA4" bold="true" box="[471,561,295,320]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FE13FED6FDEEFEA4" box="[471,561,295,320]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="4C0CC689682B8358FD8EFED9FC6DFEA4" box="[586,946,296,320]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="maoershanensis">
<emphasis id="B9786118682B8358FD8EFED9FC6DFEA4" box="[586,946,296,320]" italics="true" pageId="10" pageNumber="77">Leptobrachella maoershanensis</emphasis>
</taxonomicName>
has a bicolored iris with orange above and silver below (vs. iris copper in the upper and silver in the lower fifth in
<taxonomicName id="4C0CC689682B8358FC13FEBCFB5CFE80" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[983,1155,332,356]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FC13FEBCFB5CFE80" box="[983,1155,332,356]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FB49FEBDFB36FE81" bold="true" box="[1165,1257,332,357]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FB49FEBDFB36FE81" box="[1165,1257,332,357]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), creamy-white ventral surface with irregular black spots (vs. yellowish creamy-white ventral surface with marble texture) and shagreened dorsal skin small tubercles and longitudinal folds (vs. nearly smooth dorsal skin with numerous tiny tubercles in
<taxonomicName id="4C0CC689682B8358FEDBFE48FE14FE34" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[287,459,440,464]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FEDBFE48FE14FE34" box="[287,459,440,464]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FE15FE46FDF4FE34" bold="true" box="[465,555,439,464]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FE15FE46FDF4FE34" box="[465,555,439,464]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="4C0CC689682B8358FD86FE49FCB0FE34" box="[578,879,440,464]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="B9786118682B8358FD86FE49FCB0FE34" box="[578,879,440,464]" italics="true" pageId="10" pageNumber="77">Leptobrachella oshanensis</emphasis>
</taxonomicName>
has a relatively low dominant frequency (4.44.6 kHz at 14°C in
<taxonomicName id="4C0CC689682B8358FE83FE2CFE3FFE10" box="[327,480,476,500]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="oshanensis">
<emphasis id="B9786118682B8358FE83FE2CFE3FFE10" box="[327,480,476,500]" italics="true" pageId="10" pageNumber="77">L. oshanensis</emphasis>
</taxonomicName>
vs. 5.56.5 kHz at 21.5°C in
<taxonomicName id="4C0CC689682B8358FCF5FE2CFC03FE10" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[817,988,476,500]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FCF5FE2CFC03FE10" box="[817,988,476,500]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FC21FE2DFBE0FE11" bold="true" box="[997,1087,476,501]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FC21FE2DFBE0FE11" box="[997,1087,476,501]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), absent lateral fringes on toes (vs. present narrow lateral fringes on toes in
<taxonomicName id="4C0CC689682B8358FD4AFDF0FCE6FDFC" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[654,825,512,536]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FD4AFDF0FCE6FDFC" box="[654,825,512,536]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FC85FE0EFC44FDFC" bold="true" box="[833,923,511,536]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FC85FE0EFC44FDFC" box="[833,923,511,536]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and present wide webbing on toes (vs. basal toe webbing in
<taxonomicName id="4C0CC689682B8358FE8CFDD4FE2CFDD8" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[328,499,548,572]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FE8CFDD4FE2CFDD8" box="[328,499,548,572]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FE38FDD5FD88FDD9" bold="true" box="[508,599,548,573]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FE38FDD5FD88FDD9" box="[508,599,548,573]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="4C0CC689682B8358FDABFDD5FC55FDD8" box="[623,906,548,572]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="purpurus">
<emphasis id="B9786118682B8358FDABFDD5FC55FDD8" box="[623,906,548,572]" italics="true" pageId="10" pageNumber="77">Leptobrachella purpurus</emphasis>
</taxonomicName>
has a relatively low dominant frequency (4.3 4.5 kHz at 15°C in
<taxonomicName id="4C0CC689682B8358FEBDFDB8FDDFFD84" box="[377,512,585,608]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="purpurus">
<emphasis id="B9786118682B8358FEBDFDB8FDDFFD84" box="[377,512,585,608]" italics="true" pageId="10" pageNumber="77">L. purpurus</emphasis>
</taxonomicName>
vs. 5.56.5 kHz at 21.5°C in
<taxonomicName id="4C0CC689682B8358FC9FFDB8FBD7FD84" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[859,1032,584,608]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FC9FFDB8FBD7FD84" box="[859,1032,584,608]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FBD6FDB6FBB1FD84" bold="true" box="[1042,1134,583,608]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FBD6FDB6FBB1FD84" box="[1042,1134,583,608]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), a dull white venter with indistinct grey dusting (vs. yellowish creamy-white ventral surface with marble texture in
<taxonomicName id="4C0CC689682B8358FB43FD9CFAEDFD60" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[1159,1330,620,644]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FB43FD9CFAEDFD60" box="[1159,1330,620,644]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FAFDFD9DFA4CFD61" bold="true" box="[1337,1427,620,645]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FAFDFD9DFA4CFD61" box="[1337,1427,620,645]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and a bicolored iris with orange yellow upper half and sliver white lower half (vs. iris copper in the upper and silver in the lower fifth in
<taxonomicName id="4C0CC689682B8358FEBEFD44FDF9FD28" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[378,550,692,716]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FEBEFD44FDF9FD28" box="[378,550,692,716]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FDE9FD45FD58FD29" bold="true" box="[557,647,692,717]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FDE9FD45FD58FD29" box="[557,647,692,717]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="4C0CC689682B8358FD64FD45FC20FD28" box="[672,1023,692,716]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="tengchongensis">
<emphasis id="B9786118682B8358FD64FD45FC20FD28" box="[672,1023,692,716]" italics="true" pageId="10" pageNumber="77">Leptobrachella tengchongensis</emphasis>
</taxonomicName>
has a uniformly dark brown iris (vs. iris copper in the upper and silver in the lower fifth in
<taxonomicName id="4C0CC689682B8358FD3FFD28FC78FD14" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[763,935,728,752]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FD3FFD28FC78FD14" box="[763,935,728,752]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FC69FD26FBD8FD14" bold="true" box="[941,1031,727,752]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FC69FD26FBD8FD14" box="[941,1031,727,752]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), shagreened dorsal skin with small tubercles (vs. nearly smooth dorsal skin with numerous tiny tubercles in
<taxonomicName id="4C0CC689682B8358FC01FD0CFBAFFCF0" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[965,1136,764,788]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FC01FD0CFBAFFCF0" box="[965,1136,764,788]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FBB3FD0DFB0EFCF1" bold="true" box="[1143,1233,764,789]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FBB3FD0DFB0EFCF1" box="[1143,1233,764,789]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and white venter with dark brown blotches (vs. yellowish creamy-white ventral surface with marble texture in
<taxonomicName id="4C0CC689682B8358FB01FCD0FAADFCDC" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[1221,1394,800,824]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FB01FCD0FAADFCDC" box="[1221,1394,800,824]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FABEFCEEFF17FCB9" bold="true" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FABEFCEEFF17FCB9" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="4C0CC689682B8358FF1BFCB5FDEBFCB8" box="[223,564,836,860]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="yingjiangensis">
<emphasis id="B9786118682B8358FF1BFCB5FDEBFCB8" box="[223,564,836,860]" italics="true" pageId="10" pageNumber="77">Leptobrachella yingjiangensis</emphasis>
</taxonomicName>
has a bicolored iris with orange yellow above and sliver white below (vs. iris copper in the upper and silver in the lower fifth in
<taxonomicName id="4C0CC689682B8358FD03FC98FCACFC64" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[711,883,872,896]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FD03FC98FCACFC64" box="[711,883,872,896]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FCBDFC96FC0DFC64" bold="true" box="[889,978,871,896]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FCBDFC96FC0DFC64" box="[889,978,871,896]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), wide lateral fringes on toes (vs. narrow lateral fringes on toes in
<taxonomicName id="4C0CC689682B8358FE7CFC7CFDB9FC40" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[440,614,908,932]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FE7CFC7CFDB9FC40" box="[440,614,908,932]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FDABFC7DFD14FC41" bold="true" box="[623,715,908,933]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FDABFC7DFD14FC41" box="[623,715,908,933]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), shagreened dorsal skin with fine, round brown tubercles (vs. nearly smooth dorsal skin with numerous tiny tubercles in
<taxonomicName id="4C0CC689682B8358FCE6FC40FC12FC2C" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[802,973,944,968]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FCE6FC40FC12FC2C" box="[802,973,944,968]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FC17FC5EFBF4FC2C" bold="true" box="[979,1067,943,968]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FC17FC5EFBF4FC2C" box="[979,1067,943,968]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), ventral surface of body creamy white and scattered with distinct small dark brown flecks on chest and lateral sides of belly (vs. yellowish creamywhite ventral surface with marble texture in
<taxonomicName id="4C0CC689682B8358FD48FC08FCE7FBF4" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[652,824,1016,1040]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FD48FC08FCE7FBF4" box="[652,824,1016,1040]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FCFBFC06FC46FBF4" bold="true" box="[831,921,1015,1040]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FCFBFC06FC46FBF4" box="[831,921,1015,1040]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and a different dominant frequency (5.75.9 kHz at 19°C in
<taxonomicName id="4C0CC689682B8358FE85FBECFDDFFBD0" box="[321,512,1053,1076]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="yingjiangensis">
<emphasis id="B9786118682B8358FE85FBECFDDFFBD0" box="[321,512,1053,1076]" italics="true" pageId="10" pageNumber="77">L. yingjiangensis</emphasis>
</taxonomicName>
vs. 5.56.5 kHz at 21.5°C in
<taxonomicName id="4C0CC689682B8358FC82FBECFC2DFBD0" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[838,1010,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FC82FBECFC2DFBD0" box="[838,1010,1052,1076]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FC3CFBEDFB8EFBD1" bold="true" box="[1016,1105,1052,1077]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FC3CFBEDFB8EFBD1" box="[1016,1105,1052,1077]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="4C0CC689682B8358FBACFBEDFA43FBD0" box="[1128,1436,1052,1076]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="yunkaiensis">
<emphasis id="B9786118682B8358FBACFBEDFA43FBD0" box="[1128,1436,1052,1076]" italics="true" pageId="10" pageNumber="77">Leptobrachella yunkaiensis</emphasis>
</taxonomicName>
has a pinkish surface of belly with distinct or indistinct light dark brown speckling and black tympanum (vs. yellowish creamy-white belly and brown tympanum in
<taxonomicName id="4C0CC689682B8358FCCAFB94FC65FB98" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[782,954,1124,1148]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FCCAFB94FC65FB98" box="[782,954,1124,1148]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FC07FB95FBC1FB99" bold="true" box="[963,1054,1124,1149]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FC07FB95FBC1FB99" box="[963,1054,1124,1149]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and shagreened dorsal skin with skin ridges and raised warts (vs. nearly smooth dorsal skin with numerous tiny tubercles in
<taxonomicName id="4C0CC689682B8358FB06FB78FAAFFB44" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[1218,1392,1160,1184]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FB06FB78FAAFFB44" box="[1218,1392,1160,1184]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FABEFB76FF17FB21" bold="true" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FABEFB76FF17FB21" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
)
</paragraph>
<paragraph id="8BB3BD0A682B8359FF03FB3EFCC8FEA4" blockId="10.[151,1437,151,2013]" lastBlockId="11.[151,1436,151,320]" lastPageId="11" lastPageNumber="78" pageId="10" pageNumber="77">
Besides the above-mentioned species, the remaining ones (35 of 71 species) belong to different phylogenetic clades, exhibit high genetic variation (ranging from 11.9% to 19.0%) and allopatric distribution.
<emphasis id="B9786118682B8358FB35FB05FC36FAB0" italics="true" pageId="10" pageNumber="77">
<taxonomicName id="4C0CC689682B8358FB35FB05FF3CFACB" baseAuthorityName="Matsui" baseAuthorityYear="1997" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="arayai">Leptobrachella arayai</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FF35FAE8FE87FAD4" box="[241,344,1304,1328]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="dringi">L. dringi</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FEA2FAE8FE22FAD4" baseAuthorityName="Dehling &amp; Matsui" baseAuthorityYear="2013" box="[358,509,1304,1328]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="fritinniens">L. fritinniens</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FDCFFAE8FD5CFAD4" box="[523,643,1304,1328]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="gracilis">L. gracilis</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FD56FAE8FCDCFAD4" baseAuthorityName="Matsui" baseAuthorityYear="1997" box="[658,771,1304,1328]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="hamidi">L. hamidi</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FCD5FAE8FC7EFAD4" baseAuthorityName="Boulenger" baseAuthorityYear="1900" box="[785,929,1304,1328]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="heteropus">L. heteropus</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FC6BFAE8FB88FAD4" box="[943,1111,1304,1328]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="kajangensis">L. kajangensis</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FBA1FAE8FB62FAD4" box="[1125,1213,1304,1328]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="kecil">L. kecil</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FB0FFAE8FAA7FAD4" box="[1227,1400,1305,1328]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="marmoratus">L. marmoratus</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FA42FAE8FF31FAB0" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="maurus">L. maurus</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FF3AFACCFE62FAB0" baseAuthorityName="Matsui" baseAuthorityYear="2006" box="[254,445,1340,1364]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="melanoleucus">L. melanoleucus</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FE09FACCFDE9FAB0" box="[461,566,1341,1364]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="pictus">L. pictus</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FD82FACCFCD8FAB0" baseAuthorityName="Dehling" baseAuthorityYear="2012" box="[582,775,1340,1364]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="platycephalus">L. platycephalus</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FCD3FACCFC36FAB0" box="[791,1001,1340,1364]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="sabahmontanus">L. sabahmontanus</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0CC689682B8358FBEDFACCFB59FAB0" baseAuthorityName="Matsui" baseAuthorityYear="2006" box="[1065,1158,1340,1364]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="solus">
<emphasis id="B9786118682B8358FBEDFACCFB59FAB0" box="[1065,1158,1340,1364]" italics="true" pageId="10" pageNumber="77">L. solus</emphasis>
</taxonomicName>
lack supra-axillary and ventrolateral glands (vs. present in
<taxonomicName id="4C0CC689682B8358FDE8FA90FD05FA9C" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[556,730,1376,1400]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FDE8FA90FD05FA9C" box="[556,730,1376,1400]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FD27FAAEFCE0FA9C" bold="true" box="[739,831,1375,1400]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FD27FAAEFCE0FA9C" box="[739,831,1375,1400]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). The following species,
<taxonomicName id="4C0CC689682B8358FBA1FA90FB3AFA9C" baseAuthorityName="Rowley &amp; Cao" baseAuthorityYear="2009" box="[1125,1253,1376,1400]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B9786118682B8358FBA1FA90FB3AFA9C" box="[1125,1253,1376,1400]" italics="true" pageId="10" pageNumber="77">L. applebyi</emphasis>
</taxonomicName>
(reddish brown with white specking),
<taxonomicName id="4C0CC689682B8358FE52FA74FDDEFA78" box="[406,513,1412,1436]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="ardens">
<emphasis id="B9786118682B8358FE52FA74FDDEFA78" box="[406,513,1412,1436]" italics="true" pageId="10" pageNumber="77">L. ardens</emphasis>
</taxonomicName>
(reddish brown with white specking),
<taxonomicName id="4C0CC689682B8358FC7BFA74FBBBFA78" box="[959,1124,1412,1436]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B9786118682B8358FC7BFA74FBBBFA78" box="[959,1124,1412,1436]" italics="true" pageId="10" pageNumber="77">L. bidoupensis</emphasis>
</taxonomicName>
(reddish brown with white specking),
<taxonomicName id="4C0CC689682B8358FED6FA58FE4BFA24" box="[274,404,1448,1472]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="botsfordi">
<emphasis id="B9786118682B8358FED6FA58FE4BFA24" box="[274,404,1448,1472]" italics="true" pageId="10" pageNumber="77">L. botsfordi</emphasis>
</taxonomicName>
(reddish brown with white specking),
<taxonomicName id="4C0CC689682B8358FC82FA58FC65FA24" box="[838,954,1449,1472]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="croceus">
<emphasis id="B9786118682B8358FC82FA58FC65FA24" box="[838,954,1449,1472]" italics="true" pageId="10" pageNumber="77">L. croceus</emphasis>
</taxonomicName>
(bright orange),
<taxonomicName id="4C0CC689682B8358FBBCFA58FAD4FA24" box="[1144,1291,1448,1472]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="B9786118682B8358FBBCFA58FAD4FA24" box="[1144,1291,1448,1472]" italics="true" pageId="10" pageNumber="77">L. maculosus</emphasis>
</taxonomicName>
(brown, less white specking),
<taxonomicName id="4C0CC689682B8358FE93FA3CFE16FA00" box="[343,457,1484,1508]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B9786118682B8358FE93FA3CFE16FA00" box="[343,457,1484,1508]" italics="true" pageId="10" pageNumber="77">L. melicus</emphasis>
</taxonomicName>
(reddish brown with white specking),
<taxonomicName id="4C0CC689682B8358FCBFFA3CFC2DFA00" box="[891,1010,1484,1508]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="pallidus">
<emphasis id="B9786118682B8358FCBFFA3CFC2DFA00" box="[891,1010,1484,1508]" italics="true" pageId="10" pageNumber="77">L. pallidus</emphasis>
</taxonomicName>
(reddish brown with white specking),
<taxonomicName id="4C0CC689682B8358FF53FA00FEE5F9EC" box="[151,314,1520,1544]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="puhoatensis">
<emphasis id="B9786118682B8358FF53FA00FEE5F9EC" box="[151,314,1520,1544]" italics="true" pageId="10" pageNumber="77">L. puhoatensis</emphasis>
</taxonomicName>
(deep reddish brown),
<taxonomicName id="4C0CC689682B8358FDF9FA00FD60F9EC" box="[573,703,1520,1544]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="pyrrhops">
<emphasis id="B9786118682B8358FDF9FA00FD60F9EC" box="[573,703,1520,1544]" italics="true" pageId="10" pageNumber="77">L. pyrrhops</emphasis>
</taxonomicName>
(reddish brown with white specking) and
<taxonomicName id="4C0CC689682B8358FB52FA00FAE5F9EC" box="[1174,1338,1520,1544]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="tadungensis">
<emphasis id="B9786118682B8358FB52FA00FAE5F9EC" box="[1174,1338,1520,1544]" italics="true" pageId="10" pageNumber="77">L. tadungensis</emphasis>
</taxonomicName>
(reddish brown with white specking) have a non-creamy-white venter (vs. yellowish creamy-white ventral surface with marble texture in
<taxonomicName id="4C0CC689682B8358FEA6F9C8FDD1F9B4" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[354,526,1592,1616]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FEA6F9C8FDD1F9B4" box="[354,526,1592,1616]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FDD3F9C6FDACF9B4" bold="true" box="[535,627,1591,1616]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FDD3F9C6FDACF9B4" box="[535,627,1591,1616]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="4C0CC689682B8358FD48F9C9FC7FF9B4" baseAuthorityName="Rowley &amp; Cao" baseAuthorityYear="2009" box="[652,928,1592,1616]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B9786118682B8358FD48F9C9FC7FF9B4" box="[652,928,1592,1616]" italics="true" pageId="10" pageNumber="77">Leptobrachella applebyi</emphasis>
</taxonomicName>
has a small body size (male SVL 19.622.3 vs. SVL
<quantity id="4CF410EF682B8358FF39F9ADFE7DF990" box="[253,418,1628,1652]" metricMagnitude="-2" metricUnit="m" metricValue="2.715" metricValueMax="2.94" metricValueMin="2.4899999999999998" pageId="10" pageNumber="77" unit="mm" value="27.15" valueMax="29.4" valueMin="24.9">24.929.4 mm</quantity>
in
<taxonomicName id="4C0CC689682B8358FE0DF9ACFDAAF990" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[457,629,1628,1652]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FE0DF9ACFDAAF990" box="[457,629,1628,1652]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FDBAF9ADFD05F991" bold="true" box="[638,730,1628,1653]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FDBAF9ADFD05F991" box="[638,730,1628,1653]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
), but
<taxonomicName id="4C0CC689682B8358FCDBF9ACFC0AF990" box="[799,981,1628,1652]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="zhangyapingi">
<emphasis id="B9786118682B8358FCDBF9ACFC0AF990" box="[799,981,1628,1652]" italics="true" pageId="10" pageNumber="77">L. zhangyapingi</emphasis>
</taxonomicName>
has a large body size (male SVL
<quantity id="4CF410EF682B8358FA99F9ADFF22F973" metricMagnitude="-2" metricUnit="m" metricValue="4.915" metricValueMax="5.25" metricValueMin="4.58" pageId="10" pageNumber="77" unit="mm" value="49.15" valueMax="52.5" valueMin="45.8">45.8 52.5 mm</quantity>
vs. SVL
<quantity id="4CF410EF682B8358FEAAF971FDCCF973" box="[366,531,1664,1688]" metricMagnitude="-2" metricUnit="m" metricValue="2.715" metricValueMax="2.94" metricValueMin="2.4899999999999998" pageId="10" pageNumber="77" unit="mm" value="27.15" valueMax="29.4" valueMin="24.9">24.929.4 mm</quantity>
in
<taxonomicName id="4C0CC689682B8358FDFFF970FD37F97C" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[571,744,1664,1688]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FDFFF970FD37F97C" box="[571,744,1664,1688]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FD36F98EFC91F97C" bold="true" box="[754,846,1663,1688]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FD36F98EFC91F97C" box="[754,846,1663,1688]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
). For iris coloration,
<taxonomicName id="4C0CC689682B8358FB83F970FB18F97C" baseAuthorityName="Rowley &amp; Cao" baseAuthorityYear="2009" box="[1095,1223,1664,1688]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B9786118682B8358FB83F970FB18F97C" box="[1095,1223,1664,1688]" italics="true" pageId="10" pageNumber="77">L. applebyi</emphasis>
</taxonomicName>
has a gold iris,
<taxonomicName id="4C0CC689682B8358FA41F970FF3DF958" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="ardens">
<emphasis id="B9786118682B8358FA41F970FF3DF958" italics="true" pageId="10" pageNumber="77">L. ardens</emphasis>
</taxonomicName>
brown,
<taxonomicName id="4C0CC689682B8358FEFAF954FE3DF958" box="[318,482,1700,1724]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B9786118682B8358FEFAF954FE3DF958" box="[318,482,1700,1724]" italics="true" pageId="10" pageNumber="77">L. bidoupensis</emphasis>
</taxonomicName>
reddish above and silver below,
<taxonomicName id="4C0CC689682B8358FC8BF954FC0EF958" box="[847,977,1700,1724]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="botsfordi">
<emphasis id="B9786118682B8358FC8BF954FC0EF958" box="[847,977,1700,1724]" italics="true" pageId="10" pageNumber="77">L. botsfordi</emphasis>
</taxonomicName>
uniformly brownish gold,
<taxonomicName id="4C0CC689682B8358FB39F954FA8DF958" box="[1277,1362,1700,1724]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B9786118682B8358FB39F954FA8DF958" box="[1277,1362,1700,1724]" italics="true" pageId="10" pageNumber="77">L. firthi</emphasis>
</taxonomicName>
bright gold,
<taxonomicName id="4C0CC689682B8358FF1FF938FEADF904" box="[219,370,1736,1760]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="fuliginosis">
<emphasis id="B9786118682B8358FF1FF938FEADF904" box="[219,370,1736,1760]" italics="true" pageId="10" pageNumber="77">L. fuliginosis</emphasis>
</taxonomicName>
reddish orange above and gold below,
<taxonomicName id="4C0CC689682B8358FC82F938FC00F904" box="[838,991,1736,1760]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="kalonensis">
<emphasis id="B9786118682B8358FC82F938FC00F904" box="[838,991,1736,1760]" italics="true" pageId="10" pageNumber="77">L. kalonensis</emphasis>
</taxonomicName>
gold,
<taxonomicName id="4C0CC689682B8358FBEAF938FB5AF904" box="[1070,1157,1736,1760]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="kecil">
<emphasis id="B9786118682B8358FBEAF938FB5AF904" box="[1070,1157,1736,1760]" italics="true" pageId="10" pageNumber="77">L. kecil</emphasis>
</taxonomicName>
dark red,
<taxonomicName id="4C0CC689682B8358FAC0F938FA44F904" box="[1284,1435,1736,1760]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="B9786118682B8358FAC0F938FA44F904" box="[1284,1435,1736,1760]" italics="true" pageId="10" pageNumber="77">L. maculosus</emphasis>
</taxonomicName>
bronze,
<taxonomicName id="4C0CC689682B8358FF31F91CFE6EF8E0" baseAuthorityName="Matsui" baseAuthorityYear="2006" box="[245,433,1772,1796]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="melanoleucus">
<emphasis id="B9786118682B8358FF31F91CFE6EF8E0" box="[245,433,1772,1796]" italics="true" pageId="10" pageNumber="77">L. melanoleucus</emphasis>
</taxonomicName>
orange above and silver below,
<taxonomicName id="4C0CC689682B8358FCF6F91CFC76F8E0" box="[818,937,1772,1796]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B9786118682B8358FCF6F91CFC76F8E0" box="[818,937,1772,1796]" italics="true" pageId="10" pageNumber="77">L. melicus</emphasis>
</taxonomicName>
gold,
<taxonomicName id="4C0CC689682B8358FC3CF91CFBABF8E0" box="[1016,1140,1772,1796]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="pallidus">
<emphasis id="B9786118682B8358FC3CF91CFBABF8E0" box="[1016,1140,1772,1796]" italics="true" pageId="10" pageNumber="77">L. pallidus</emphasis>
</taxonomicName>
dark brown,
<taxonomicName id="4C0CC689682B8358FAD2F91CFA44F8E0" box="[1302,1435,1772,1796]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="pyrrhops">
<emphasis id="B9786118682B8358FAD2F91CFA44F8E0" box="[1302,1435,1772,1796]" italics="true" pageId="10" pageNumber="77">L. pyrrhops</emphasis>
</taxonomicName>
orange,
<taxonomicName id="4C0CC689682B8358FF30F8E0FEA5F8CC" box="[244,378,1808,1832]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="rowleyae">
<emphasis id="B9786118682B8358FF30F8E0FEA5F8CC" box="[244,378,1808,1832]" italics="true" pageId="10" pageNumber="77">L. rowleyae</emphasis>
</taxonomicName>
copper tint in upper half fading to golden in lower third,
<taxonomicName id="4C0CC689682B8358FBDAF8E0FBA7F8CC" baseAuthorityName="Matsui" baseAuthorityYear="2006" box="[1054,1144,1808,1832]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="solus">
<emphasis id="B9786118682B8358FBDAF8E0FBA7F8CC" box="[1054,1144,1808,1832]" italics="true" pageId="10" pageNumber="77">L. solus</emphasis>
</taxonomicName>
dark red above and dark brown below,
<taxonomicName id="4C0CC689682B8358FE8EF8C4FE28F8A8" box="[330,503,1844,1868]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="tadungensis">
<emphasis id="B9786118682B8358FE8EF8C4FE28F8A8" box="[330,503,1844,1868]" italics="true" pageId="10" pageNumber="77">L. tadungensis</emphasis>
</taxonomicName>
gold,
<taxonomicName id="4C0CC689682B8358FD95F8C4FD3AF8A8" box="[593,741,1844,1868]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B9786118682B8358FD95F8C4FD3AF8A8" box="[593,741,1844,1868]" italics="true" pageId="10" pageNumber="77">L. tuberosus</emphasis>
</taxonomicName>
gold and
<taxonomicName id="4C0CC689682B8358FCB5F8C4FBF0F8A8" box="[881,1071,1844,1868]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="zhangyapingi">
<emphasis id="B9786118682B8358FCB5F8C4FBF0F8A8" box="[881,1071,1844,1868]" italics="true" pageId="10" pageNumber="77">L. zhangyapingi</emphasis>
</taxonomicName>
gold above and grey below.
<emphasis id="B9786118682B8358FF53F8A9FCFFF894" box="[151,800,1880,1904]" italics="true" pageId="10" pageNumber="77">
<taxonomicName id="4C0CC689682B8358FF53F8A9FE7DF894" box="[151,418,1880,1904]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="croceus">Leptobrachella croceus</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FE69F8A8FDDAF894" box="[429,517,1880,1904]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="firthi">L. firthi</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FDD4F8A8FD54F894" box="[528,651,1880,1904]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="pallidus">L. pallidus</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FD52F8A8FCFFF894" box="[662,800,1880,1904]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="tuberosus">L. tuberosus</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0CC689682B8358FC92F8A8FBD5F894" box="[854,1034,1880,1904]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="zhangyapingi">
<emphasis id="B9786118682B8358FC92F8A8FBD5F894" box="[854,1034,1880,1904]" italics="true" pageId="10" pageNumber="77">L. zhangyapingi</emphasis>
</taxonomicName>
lack a distinct black supratympanic line (vs. present a black supratympanic line in
<taxonomicName id="4C0CC689682B8358FD72F88CFCBCF870" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[694,867,1916,1940]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FD72F88CFCBCF870" box="[694,867,1916,1940]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8358FCA9F88DFC16F871" bold="true" box="[877,969,1916,1941]" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8358FCA9F88DFC16F871" box="[877,969,1916,1941]" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<emphasis id="B9786118682B8358FC20F88DFA43F85C" italics="true" pageId="10" pageNumber="77">
<taxonomicName id="4C0CC689682B8358FC20F88DFB24F870" baseAuthorityName="Rowley &amp; Cao" baseAuthorityYear="2009" box="[996,1275,1916,1940]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="applebyi">Leptobrachella applebyi</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FACDF88CFAA7F870" box="[1289,1400,1916,1940]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="ardens">L. ardens</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FA42F88CFF3AF85C" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="crocea">L. crocea</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FF36F850FE55F85C" box="[242,394,1952,1976]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="kalonensis">L. kalonensis</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FE53F850FDC7F85C" box="[407,536,1953,1976]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="macrops">L. macrops</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FDE2F850FD61F85C" box="[550,702,1952,1976]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="maculosus">L. maculosus</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FD0FF850FCE7F85C" box="[715,824,1952,1976]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="melica">L. melica</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FC81F850FC1EF85C" box="[837,961,1952,1976]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="pallidus">L. pallidus</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FC0AF850FB89F85C" box="[974,1110,1952,1976]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="pyrrhops">L. pyrrhops</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FBA6F850FB35F85C" box="[1122,1258,1952,1976]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="rowleyae">L. rowleyae</taxonomicName>
,
<taxonomicName id="4C0CC689682B8358FB33F850FA43F85C" box="[1271,1436,1952,1976]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="tadungensis">L. tadungensis</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0CC689682B8358FF0CF834FE95F838" box="[200,330,1988,2012]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="tuberosa">
<emphasis id="B9786118682B8358FF0CF834FE95F838" box="[200,330,1988,2012]" italics="true" pageId="10" pageNumber="77">L. tuberosa</emphasis>
</taxonomicName>
have no lateral fringes on toes (vs. present narrow lateral fringes on toes in
<taxonomicName id="4C0CC689682B8358FB07F834FAAEF838" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[1219,1393,1988,2012]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="10" pageNumber="77" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682B8358FB07F834FAAEF838" box="[1219,1393,1988,2012]" italics="true" pageId="10" pageNumber="77">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682B8359FABEF835FF17FF54" bold="true" lastPageId="11" lastPageNumber="78" pageId="10" pageNumber="77">
<taxonomicNameLabel id="A24BDC63682B8359FABEF835FF17FF54" lastPageId="11" lastPageNumber="78" pageId="10" pageNumber="77" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<emphasis id="B9786118682A8359FF1AFF69FD61FF54" box="[222,702,152,176]" italics="true" pageId="11" pageNumber="78">
<taxonomicName id="4C0CC689682A8359FF1AFF69FE14FF54" box="[222,459,152,176]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="firthi">Leptobrachella firthi</taxonomicName>
,
<taxonomicName id="4C0CC689682A8359FE11FF68FDFFFF4B" box="[469,544,152,176]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="isos">L. isos</taxonomicName>
,
<taxonomicName id="4C0CC689682A8359FDEFFF68FD61FF54" box="[555,702,152,176]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="khasiorum">L. khasiorum</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0CC689682A8359FD37FF68FC78FF54" box="[755,935,152,176]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="zhangyapingi">
<emphasis id="B9786118682A8359FD37FF68FC78FF54" box="[755,935,152,176]" italics="true" pageId="11" pageNumber="78">L. zhangyapingi</emphasis>
</taxonomicName>
have wide lateral fringes on toes (vs. present narrow lateral fringes on toes in
<taxonomicName id="4C0CC689682A8359FDCDFF4CFD6AFF30" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[521,693,188,212]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682A8359FDCDFF4CFD6AFF30" box="[521,693,188,212]" italics="true" pageId="11" pageNumber="78">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682A8359FD79FF4DFCC7FF31" bold="true" box="[701,792,188,213]" pageId="11" pageNumber="78">
<taxonomicNameLabel id="A24BDC63682A8359FD79FF4DFCC7FF31" box="[701,792,188,213]" pageId="11" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<emphasis id="B9786118682A8359FCF4FF4DFDC7FF1C" italics="true" pageId="11" pageNumber="78">
<taxonomicName id="4C0CC689682A8359FCF4FF4DFBEEFF30" box="[816,1073,188,212]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="ardens">Leptobrachella ardens</taxonomicName>
,
<taxonomicName id="4C0CC689682A8359FBFAFF4CFB0AFF30" box="[1086,1237,188,212]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="kalonensis">L. kalonensis</taxonomicName>
,
<taxonomicName id="4C0CC689682A8359FB25FF4CFAA6FF30" box="[1249,1401,188,212]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="maculosus">L. maculosus</taxonomicName>
,
<taxonomicName id="4C0CC689682A8359FA42FF4CFF2BFF1C" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="pallidus">L. pallidus</taxonomicName>
,
<taxonomicName id="4C0CC689682A8359FEC1FF10FEA1FF1C" box="[261,382,225,248]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="petrops">L. petrops</taxonomicName>
,
<taxonomicName id="4C0CC689682A8359FE4BFF10FDC7FF1C" box="[399,536,224,248]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="rowleyae">L. rowleyae</taxonomicName>
</emphasis>
and
<taxonomicName id="4C0CC689682A8359FD9EFF10FCDCFF1C" box="[602,771,224,248]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="tadungensis">
<emphasis id="B9786118682A8359FD9EFF10FCDCFF1C" box="[602,771,224,248]" italics="true" pageId="11" pageNumber="78">L. tadungensis</emphasis>
</taxonomicName>
have no webbing on toes (vs. basal toe webbing in
<taxonomicName id="4C0CC689682A8359FA41FF10FEFAFEF8" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682A8359FA41FF10FEFAFEF8" italics="true" pageId="11" pageNumber="78">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682A8359FEF4FEF5FE50FEF9" bold="true" box="[304,399,260,285]" pageId="11" pageNumber="78">
<taxonomicNameLabel id="A24BDC63682A8359FEF4FEF5FE50FEF9" box="[304,399,260,285]" pageId="11" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
<taxonomicName id="4C0CC689682A8359FE6FFEF5FD18FEF8" box="[427,711,260,284]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="botsfordi">
<emphasis id="B9786118682A8359FE6FFEF5FD18FEF8" box="[427,711,260,284]" italics="true" pageId="11" pageNumber="78">Leptobrachella botsfordi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0CC689682A8359FD1EFEF4FCEBFEF8" box="[730,820,260,284]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="firthi">
<emphasis id="B9786118682A8359FD1EFEF4FCEBFEF8" box="[730,820,260,284]" italics="true" pageId="11" pageNumber="78">L. firthi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0CC689682A8359FC83FEF4FC49FEF8" box="[839,918,261,284]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="isos">
<emphasis id="B9786118682A8359FC83FEF4FC49FEF8" box="[839,918,261,284]" italics="true" pageId="11" pageNumber="78">L. isos</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0CC689682A8359FC6DFEF4FBFAFEF8" box="[937,1061,260,284]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="pallidus">
<emphasis id="B9786118682A8359FC6DFEF4FBFAFEF8" box="[937,1061,260,284]" italics="true" pageId="11" pageNumber="78">L. pallidus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0CC689682A8359FBFCFEF4FB71FEF8" box="[1080,1198,261,284]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="petrops">
<emphasis id="B9786118682A8359FBFCFEF4FB71FEF8" box="[1080,1198,261,284]" italics="true" pageId="11" pageNumber="78">L. petrops</emphasis>
</taxonomicName>
,
<taxonomicName id="4C0CC689682A8359FB05FEF4FA9AFEF8" box="[1217,1349,260,284]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="tuberosa">
<emphasis id="B9786118682A8359FB05FEF4FA9AFEF8" box="[1217,1349,260,284]" italics="true" pageId="11" pageNumber="78">L. tuberosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C0CC689682A8359FA41FEF4FEF1FEA4" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="zhangyapingi">
<emphasis id="B9786118682A8359FA41FEF4FEF1FEA4" italics="true" pageId="11" pageNumber="78">L. zhangyapingi</emphasis>
</taxonomicName>
have no black spots on flanks (vs. present).
</paragraph>
<caption id="DF73ED82682A8359FF53F86AFD97F834" httpUri="https://zenodo.org/record/2600987/files/figure.png" pageId="11" pageNumber="78" startId="11.[151,250,1947,1969]" targetBox="[150,1436,375,1925]" targetPageId="11">
<paragraph id="8BB3BD0A682A8359FF53F86AFD97F834" blockId="11.[151,1436,1947,2000]" pageId="11" pageNumber="78">
<emphasis id="B9786118682A8359FF53F86AFECAF855" bold="true" box="[151,277,1947,1969]" pageId="11" pageNumber="78">FIGURE 5.</emphasis>
(A) Breeding habitat of
<taxonomicName id="4C0CC689682A8359FDD9F86AFC9EF855" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[541,833,1947,1969]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="11" pageNumber="78" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682A8359FDD9F86AFC9EF855" box="[541,833,1947,1969]" italics="true" pageId="11" pageNumber="78">Leptobrachella shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682A8359FC8EF86AFC42F855" bold="true" box="[842,925,1947,1969]" pageId="11" pageNumber="78">
<taxonomicNameLabel id="A24BDC63682A8359FC8EF86AFC42F855" box="[842,925,1947,1969]" pageId="11" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and (B) deforestation in Shiwandashan National Nature Reserve, Guangxi Province, China.
</paragraph>
</caption>
<paragraph id="8BB3BD0A682D835EFF03FF66FD84FE49" blockId="12.[151,1436,151,429]" pageId="12" pageNumber="79">
Finally, calls of
<taxonomicName id="4C0CC689682D835EFEB8FF68FDF8FF54" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[380,551,152,176]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682D835EFEB8FF68FDF8FF54" box="[380,551,152,176]" italics="true" pageId="12" pageNumber="79">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682D835EFDEBFF66FD56FF54" bold="true" box="[559,649,151,176]" pageId="12" pageNumber="79">
<taxonomicNameLabel id="A24BDC63682D835EFDEBFF66FD56FF54" box="[559,649,151,176]" pageId="12" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
consist of a single note with a dominant frequency of 5.56.5 kHz (at 21.5°C). The dominant frequency of 5.56.5 kHz further distinguishes the call of
<taxonomicName id="4C0CC689682D835EFBE7FF4CFB12FF30" authority="Chen &amp; Liao &amp; Zhou &amp; Mo, 2019" authorityName="Chen &amp; Liao &amp; Zhou &amp; Mo" authorityYear="2019" box="[1059,1229,188,212]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="shangsiensis" status="sp. nov.">
<emphasis id="B9786118682D835EFBE7FF4CFB12FF30" box="[1059,1229,188,212]" italics="true" pageId="12" pageNumber="79">L. shangsiensis</emphasis>
</taxonomicName>
<emphasis id="B9786118682D835EFB11FF4DFAF1FF31" bold="true" box="[1237,1326,188,213]" pageId="12" pageNumber="79">
<taxonomicNameLabel id="A24BDC63682D835EFB11FF4DFAF1FF31" box="[1237,1326,188,213]" pageId="12" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
from that of the lower frequency calls of
<taxonomicName id="4C0CC689682D835EFDC7FF10FD5CFF1C" baseAuthorityName="Rowley &amp; Cao" baseAuthorityYear="2009" box="[515,643,224,248]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="applebyi">
<emphasis id="B9786118682D835EFDC7FF10FD5CFF1C" box="[515,643,224,248]" italics="true" pageId="12" pageNumber="79">L. applebyi</emphasis>
</taxonomicName>
(4.04.3 kHz at 21.5°C),
<taxonomicName id="4C0CC689682D835EFC6BFF10FBC5FF1C" box="[943,1050,224,248]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="ardens">
<emphasis id="B9786118682D835EFC6BFF10FBC5FF1C" box="[943,1050,224,248]" italics="true" pageId="12" pageNumber="79">L. ardens</emphasis>
</taxonomicName>
(3.13.4 kHz at 21.424.7°C),
<taxonomicName id="4C0CC689682D835EFA41FF10FEC2FEF8" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="bidoupensis">
<emphasis id="B9786118682D835EFA41FF10FEC2FEF8" italics="true" pageId="12" pageNumber="79">L. bidoupensis</emphasis>
</taxonomicName>
(1.93.8 kHz at 19.021.0°C),
<taxonomicName id="4C0CC689682D835EFD43FEF4FCD4FEF8" box="[647,779,260,284]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="botsfordi">
<emphasis id="B9786118682D835EFD43FEF4FCD4FEF8" box="[647,779,260,284]" italics="true" pageId="12" pageNumber="79">L. botsfordi</emphasis>
</taxonomicName>
(2.63.4 kHz at 14°C),
<taxonomicName id="4C0CC689682D835EFBE5FEF4FB48FEF8" box="[1057,1175,261,284]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="croceus">
<emphasis id="B9786118682D835EFBE5FEF4FB48FEF8" box="[1057,1175,261,284]" italics="true" pageId="12" pageNumber="79">L. croceus</emphasis>
</taxonomicName>
(3.04.3 kHz at 21.6 25.1°C),
<taxonomicName id="4C0CC689682D835EFF3EFED8FE4DFEA4" box="[250,402,296,320]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="fuliginosus">
<emphasis id="B9786118682D835EFF3EFED8FE4DFEA4" box="[250,402,296,320]" italics="true" pageId="12" pageNumber="79">L. fuliginosus</emphasis>
</taxonomicName>
(2.12.8 kHz at 19.319.6°C),
<taxonomicName id="4C0CC689682D835EFD2BFED8FC5DFEA4" box="[751,898,296,320]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="kalonensis">
<emphasis id="B9786118682D835EFD2BFED8FC5DFEA4" box="[751,898,296,320]" italics="true" pageId="12" pageNumber="79">L. kalonensis</emphasis>
</taxonomicName>
(2.8 kHz at 26.4°C),
<taxonomicName id="4C0CC689682D835EFBABFED8FADDFEA4" box="[1135,1282,296,320]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="maculosus">
<emphasis id="B9786118682D835EFBABFED8FADDFEA4" box="[1135,1282,296,320]" italics="true" pageId="12" pageNumber="79">L. maculosus</emphasis>
</taxonomicName>
(2.72.8 kHz at 23.324.1°C),
<taxonomicName id="4C0CC689682D835EFE9CFEBCFE12FE80" box="[344,461,332,356]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="melicus">
<emphasis id="B9786118682D835EFE9CFEBCFE12FE80" box="[344,461,332,356]" italics="true" pageId="12" pageNumber="79">L. melicus</emphasis>
</taxonomicName>
(2.64.0 kHz at 26.126.2°C),
<taxonomicName id="4C0CC689682D835EFCF2FEBCFC70FE80" box="[822,943,332,356]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="pallidus">
<emphasis id="B9786118682D835EFCF2FEBCFC70FE80" box="[822,943,332,356]" italics="true" pageId="12" pageNumber="79">L. pallidus</emphasis>
</taxonomicName>
(2.42.7 kHz at 14.021.4°C),
<taxonomicName id="4C0CC689682D835EFADCFEBCFA44FE80" box="[1304,1435,332,356]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="pyrrhops">
<emphasis id="B9786118682D835EFADCFEBCFA44FE80" box="[1304,1435,332,356]" italics="true" pageId="12" pageNumber="79">L. pyrrhops</emphasis>
</taxonomicName>
(1.92.2 kHz at 25°C),
<taxonomicName id="4C0CC689682D835EFE65FE80FDF9FE6C" box="[417,550,368,392]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="rowleyae">
<emphasis id="B9786118682D835EFE65FE80FDF9FE6C" box="[417,550,368,392]" italics="true" pageId="12" pageNumber="79">L. rowleyae</emphasis>
</taxonomicName>
(3.23.5 kHz at 21.0°C),
<taxonomicName id="4C0CC689682D835EFC8BFE80FC2BFE6C" box="[847,1012,368,392]" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="tadungensis">
<emphasis id="B9786118682D835EFC8BFE80FC2BFE6C" box="[847,1012,368,392]" italics="true" pageId="12" pageNumber="79">L. tadungensis</emphasis>
</taxonomicName>
(2.63.1 kHz at 12.922.3°C) and
<taxonomicName id="4C0CC689682D835EFA41FE80FEDCFE48" class="Amphibia" family="Megophryidae" genus="Leptobrachella" kingdom="Animalia" order="Anura" pageId="12" pageNumber="79" phylum="Chordata" rank="species" species="tuberosus">
<emphasis id="B9786118682D835EFA41FE80FEDCFE48" italics="true" pageId="12" pageNumber="79">L. tuberosus</emphasis>
</taxonomicName>
(2.62.8 kHz at 22.524.5°C).
</paragraph>
</subSubSection>
</treatment>
</document>