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<document ID-DOI="10.5252/geodiversitas2019v41a15" ID-GBIF-Dataset="1e89d723-7f54-4d6b-85dd-6bcd573eca32" ID-ISSN="1638-9395" ID-Zenodo-Dep="3694209" ID-ZooBank="urn:lsid:zoobank.org:pub:9DD3CC29-3AEA-44B8-8E8F-6AD882DF5B1C" checkinTime="1583172268754" checkinUser="valdenar" docAuthor="Bonis, Louis de, Gardin, Axelle &amp; Blondel, Cécile" docDate="2019" docId="03A487991A70FF8EFF42F9B96D98FA9E" docLanguage="en" docName="Geodiversitas.41.15.601-621.pdf.imf" docOrigin="Geodiversitas 41 (15)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Peignictis pseudamphictis Bonis &amp; Gardin &amp; Blondel 2019, n. sp." docType="treatment" docUuid="C96DF1A3-816B-4BD7-9635-28223CFDA804" docUuidSource="ZooBank" docVersion="1" lastPageNumber="615" masterDocId="FF9DFFE11A60FF9EFFADFF8D6E62FF8E" masterDocTitle="Carnivora from the early Oligocene of the Phosphorites du Quercy in southwestern France" masterLastPageNumber="621" masterPageNumber="601" pageNumber="615" updateTime="1675702286038" updateUser="tatiana" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo>
<mods:title>Carnivora from the early Oligocene of the Phosphorites du Quercy in southwestern France</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Bonis, Louis de</mods:namePart>
<mods:affiliation>Palevoprim: laboratoire de Paléontologie, Évolution, Paléoécosystèmes, Paléoprimatologie, Bâtiment B 35 TSA 51106, 6 rue Michel Brunet F- 86073 Poitiers cedex 9 (France)</mods:affiliation>
<mods:nameIdentifier type="email">louis.debonis@univ-poitiers.fr</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Gardin, Axelle</mods:namePart>
<mods:affiliation>Palevoprim: laboratoire de Paléontologie, Évolution, Paléoécosystèmes, Paléoprimatologie, Bâtiment B 35 TSA 51106, 6 rue Michel Brunet F- 86073 Poitiers cedex 9 (France)</mods:affiliation>
<mods:nameIdentifier type="email">axelle.gardin@univ-poitiers.fr</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Blondel, Cécile</mods:namePart>
<mods:affiliation>Palevoprim: laboratoire de Paléontologie, Évolution, Paléoécosystèmes, Paléoprimatologie, Bâtiment B 35 TSA 51106, 6 rue Michel Brunet F- 86073 Poitiers cedex 9 (France)</mods:affiliation>
<mods:nameIdentifier type="email">cecile.blondel@univ-poitiers.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Geodiversitas</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-09-10</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>41</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>15</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>601</mods:start>
<mods:end>621</mods:end>
</mods:extent>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5252/geodiversitas2019v41a15</mods:identifier>
<mods:identifier type="GBIF-Dataset">1e89d723-7f54-4d6b-85dd-6bcd573eca32</mods:identifier>
<mods:identifier type="ISSN">1638-9395</mods:identifier>
<mods:identifier type="Zenodo-Dep">3694209</mods:identifier>
<mods:identifier type="ZooBank">urn:lsid:zoobank.org:pub:9DD3CC29-3AEA-44B8-8E8F-6AD882DF5B1C</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3703548" ID-GBIF-Taxon="162367513" ID-Zenodo-Dep="3703548" LSID="urn:lsid:zoobank.org:act:C96DF1A3-816B-4BD7-9635-28223CFDA804" httpUri="http://treatment.plazi.org/id/03A487991A70FF8EFF42F9B96D98FA9E" lastPageNumber="615" pageId="16" pageNumber="615">
<subSubSection box="[239,669,1588,1615]" pageId="16" pageNumber="615" type="nomenclature">
<paragraph blockId="16.[239,669,1588,1647]" box="[239,669,1588,1615]" pageId="16" pageNumber="615">
<heading box="[239,669,1588,1615]" pageId="16" pageNumber="615">
<taxonomicName authority="Bonis &amp; Gardin &amp; Blondel, 2019" authorityName="Bonis &amp; Gardin &amp; Blondel" authorityYear="2019" box="[239,515,1588,1615]" class="Mammalia" family="Mustelidae" genus="Peignictis" kingdom="Animalia" order="Carnivora" pageId="16" pageNumber="615" phylum="Chordata" rank="species" species="pseudamphictis" status="n. gen., n. sp.">
<emphasis bold="true" box="[239,515,1588,1615]" italics="true" pageId="16" pageNumber="615">Peignictis pseudamphictis</emphasis>
</taxonomicName>
n. gen.,
<taxonomicNameLabel box="[609,669,1589,1615]" pageId="16" pageNumber="615" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[402,506,1620,1647]" pageId="16" pageNumber="615" type="description">
<paragraph blockId="16.[239,669,1588,1647]" box="[402,506,1620,1647]" pageId="16" pageNumber="615">
(
<figureCitation box="[411,497,1620,1647]" captionStart="FIG" captionStartId="14.[132,143,1896,1914]" captionTargetBox="[153,1429,215,1875]" captionTargetId="figure@14.[698,1267,764,1331]" captionTargetPageId="14" captionText="FIG. 4. — A, Mustelictis cf. major, right hemi-mandible with p2-p4-m1, UM VD17, in buccal (A1), lingual (A2), and occlusal (A3) views; B, Mustelictis aff. olivieri, left hemi-mandible with p4-m2, UP LPL12, in buccal (B1), lingual (B2), and occlusal (B3) views; C-E, Mustelictis aff. olivieri; C, left m1, UP LPL11, in buccal (C1), and lingual (C2) views; D, right m1, UM VBOA3-4, in buccal (D1) view, occlusal (D2) view, and lingual (D3) views; E, right hemi-mandible with d3-d4, UP MGB6, in lingual (E1), and buccal (E2) views; F, Mustelictis olivieri, left mandible with p3-m1, UP MGB8, in buccal (F1), and lingual (F2) views; G, Mustelictis aff. olivieri, right hemi-mandible with p2-p4, UM VD12, in buccal (G1) view, and lingual (G2) views; H, Peignictis pseudamphictis n. gen., n. sp., right hemimandible with m1, UM VD1, in buccal (H1), lingual (H2), and occlusal (H3) views. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3694217" httpUri="https://zenodo.org/record/3694217/files/figure.png" pageId="16" pageNumber="615">Fig. 4H</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection box="[132,774,1677,1697]" pageId="16" pageNumber="615" type="description">
<paragraph blockId="16.[132,774,1677,1697]" box="[132,774,1677,1697]" pageId="16" pageNumber="615">
<uri box="[132,774,1677,1697]" pageId="16" pageNumber="615">
urn:lsid:zoobank.org:act:
<uuid box="[348,774,1677,1697]" pageId="16" pageNumber="615">C96DF1A3-816B-4BD7-9635-28223CFDA804</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="615" type="materials_examined">
<paragraph blockId="16.[132,775,1741,1792]" pageId="16" pageNumber="615">
<collectionCode box="[132,182,1741,1765]" pageId="16" pageNumber="615">TYPE</collectionCode>
<collectionCode box="[190,285,1746,1763]" pageId="16" pageNumber="615">SPECIMEN</collectionCode>
. —
<materialsCitation ID-GBIF-Occurrence="2572934962" collectionCode="UM" pageId="16" pageNumber="615" specimenCode="VD1" specimenCount="1" typeStatus="holotype">
<typeStatus box="[334,432,1741,1765]" pageId="16" pageNumber="615">Holotype</typeStatus>
: posterior part of hemi-mandible,
<collectionCode box="[132,174,1768,1791]" country="Germany" name="University of Marburg" pageId="16" pageNumber="615">UM</collectionCode>
<specimenCode box="[180,233,1768,1792]" collectionCode="VD" pageId="16" pageNumber="615">VD1</specimenCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection box="[132,454,1820,1845]" pageId="16" pageNumber="615" type="diagnosis">
<paragraph blockId="16.[132,454,1820,1845]" box="[132,454,1820,1845]" pageId="16" pageNumber="615">DIAGNOSIS. — That of the genus</paragraph>
</subSubSection>
<subSubSection box="[132,529,1872,1897]" pageId="16" pageNumber="615" type="materials_examined">
<paragraph blockId="16.[132,529,1872,1897]" box="[132,529,1872,1897]" pageId="16" pageNumber="615">
<typeStatus box="[132,181,1872,1896]" pageId="16" pageNumber="615">
<collectionCode box="[132,181,1872,1896]" pageId="16" pageNumber="615">TYPE</collectionCode>
</typeStatus>
<collectionCode box="[188,279,1877,1894]" pageId="16" pageNumber="615">LOCALITY</collectionCode>
. — Valbro (Lot,
<collectingCountry box="[448,518,1873,1897]" name="France" pageId="16" pageNumber="615">France</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection box="[132,594,1925,1949]" pageId="16" pageNumber="615" type="discussion">
<paragraph blockId="16.[132,594,1925,1949]" box="[132,594,1925,1949]" pageId="16" pageNumber="615">
GEOLOGICAL AGE. — Early Oligocene (
<collectionCode box="[517,555,1925,1949]" country="0" name="Mohonk Preserve, Inc." pageId="16" pageNumber="615">MP</collectionCode>
22)
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="615" type="etymology">
<paragraph blockId="16.[132,774,1977,2028]" pageId="16" pageNumber="615">
ETYMOLOGY. — Parallelism with
<emphasis box="[462,555,1978,2002]" italics="true" pageId="16" pageNumber="615">Amphictis</emphasis>
in the lack of m3 and elongate m2.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="615" type="description">
<paragraph blockId="16.[813,1456,217,1296]" box="[813,957,217,243]" pageId="16" pageNumber="615">DESCRIPTION</paragraph>
<paragraph blockId="16.[813,1456,217,1296]" pageId="16" pageNumber="615">
UM VD1 from Valbro is a piece of hemi-mandible broken in front of m1 and with an unworn m1 and alveoli of m2, (length of m1 =
<quantity box="[1095,1190,313,340]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="16" pageNumber="615" unit="mm" value="5.3">5.3 mm</quantity>
; w of trm1 =
<quantity box="[1355,1451,313,339]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="615" unit="mm" value="2.8">2.8 mm</quantity>
; w of tlm1 = 2; height of the corpus under m1-m2 = 5.1; length of m2 from the alveoli =
<quantity box="[1176,1271,376,403]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="16" pageNumber="615" unit="mm" value="2.5">2.5 mm</quantity>
). The ascending ramus, whose tip is broken off, is broad at the level of the condyle but narrowing upwards because the distal border is mesially oblique (
<figureCitation box="[1123,1215,472,498]" captionStart="FIG" captionStartId="14.[132,143,1896,1914]" captionTargetBox="[153,1429,215,1875]" captionTargetId="figure@14.[698,1267,764,1331]" captionTargetPageId="14" captionText="FIG. 4. — A, Mustelictis cf. major, right hemi-mandible with p2-p4-m1, UM VD17, in buccal (A1), lingual (A2), and occlusal (A3) views; B, Mustelictis aff. olivieri, left hemi-mandible with p4-m2, UP LPL12, in buccal (B1), lingual (B2), and occlusal (B3) views; C-E, Mustelictis aff. olivieri; C, left m1, UP LPL11, in buccal (C1), and lingual (C2) views; D, right m1, UM VBOA3-4, in buccal (D1) view, occlusal (D2) view, and lingual (D3) views; E, right hemi-mandible with d3-d4, UP MGB6, in lingual (E1), and buccal (E2) views; F, Mustelictis olivieri, left mandible with p3-m1, UP MGB8, in buccal (F1), and lingual (F2) views; G, Mustelictis aff. olivieri, right hemi-mandible with p2-p4, UM VD12, in buccal (G1) view, and lingual (G2) views; H, Peignictis pseudamphictis n. gen., n. sp., right hemimandible with m1, UM VD1, in buccal (H1), lingual (H2), and occlusal (H3) views. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3694217" httpUri="https://zenodo.org/record/3694217/files/figure.png" pageId="16" pageNumber="615">Fig. 4H</figureCitation>
<figureCitation box="[1215,1263,472,498]" captionStart="FIG" captionStartId="6.[132,143,1964,1982]" captionTargetBox="[212,1402,209,1931]" captionTargetId="figure@6.[755,1301,780,1349]" captionTargetPageId="6" captionText="FIG. 1. — A, Amphicynodon sp. 1: left hemi-mandible UP VAL2, in buccal (A1), lingual (A2), and occlusal (A3) views; B, Amphicynodon typicus: left mandible UP VAL1, in buccal (B1), lingual (B2), and occlusal (B3) views; C, Amphicynodon sp. 3: right M1, UM VD49, in occlusal view; D, Amphicynodon sp. 2: right M1, UM VBO492, in occlusal view. Scale bars: A, B, 10 mm; C, D, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3694211" httpUri="https://zenodo.org/record/3694211/files/figure.png" pageId="16" pageNumber="615">1-H</figureCitation>
<figureCitation box="[1263,1278,472,498]" captionStart="FIG" captionStartId="10.[132,143,1919,1937]" captionTargetBox="[190,1388,179,1887]" captionTargetId="figure@10.[747,1305,762,1331]" captionTargetPageId="10" captionText="FIG. 3. — A, Pachycynodon sp. 3, left M1-M2, UM VD47, in occlusal view; B, Pachycynodon cf. filholi, UM VD5, left hemi-mandible, in buccal (B1), lingual (B2), and occlusal (B3) views; C-E, Pachycynodon cf. dubius; C, left hemi-mandible UM VD14, in buccal (C1), lingual (C2), and occlusal (C3) views; D, right hemimandible UM VD4, in lingual (D1) and buccal (D2) views; E, right hemimandible VD6, in buccal (E1), and occlusal (E2) views; F, Cephalogale sp., left mandible UP MGB26, in lingual (F1), and buccal (F2) views; G, Pachycynodon sp. 5, left P4 UP VD51, in occlusal (G1), and mesial (G2) views; H, Pachycynodon cf. filholi, right m2, UP LPL14, in occlusal view. Scale bars: 5 mm." figureDoi="http://doi.org/10.5281/zenodo.3694215" httpUri="https://zenodo.org/record/3694215/files/figure.png" pageId="16" pageNumber="615">3</figureCitation>
). The large and deep masseteric fossa extends mesially to the level of m2 and dorsally it diminishes gently without any trace of a boundary. The condyle is situated high (
<quantity box="[1287,1373,568,595]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="16" pageNumber="615" unit="mm" value="38.0">38 mm</quantity>
higher than the base of the angular process compared with
<quantity box="[1376,1456,600,626]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="16" pageNumber="615" unit="mm" value="28.0">28 mm</quantity>
higher in
<taxonomicName authority="Bonis, 1997" authorityName="Bonis" authorityYear="1997" box="[924,1129,632,658]" class="Mammalia" family="Mustelidae" genus="Mustelictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="16" pageNumber="615" phylum="Chordata" rank="species" species="olivieri">
<emphasis box="[924,1129,632,658]" italics="true" pageId="16" pageNumber="615">Mustelictis olivieri</emphasis>
</taxonomicName>
of similar size). The angular process is flat, mesially situated relative to the condyle and not distally projected. The m1trigonid is high with a vertical distal wall, while the protoconid and paraconid form a shearing blade. The pointed metaconid is higher than the paraconid but not clearly reduced nor distally displaced. The talonid basin is flat and surrounded by a low uninterrupted cristid in which we cannot distinguish the cuspids. Buccally, there is a well-marked notch between protoconid and hypocristid. Two alveoli, the mesial being the larger, indicate the length of m2. It is very long for an early Oligocene carnivoran. This is a parallelism with the geologically younger
<emphasis box="[1171,1282,1014,1040]" italics="true" pageId="16" pageNumber="615">Amphictis</emphasis>
. However, the overall morphology of the latter is different in having a lower and less pointed m1 trigonid. Thus, VD1 exhibits some contradictory characters, e.g., a cutting trigonid associated with a grinding talonid. We dont know any evolutionary history of this structure. The closest genus to
<emphasis box="[843,948,1206,1231]" italics="true" pageId="16" pageNumber="615">Peignictis</emphasis>
n. gen. is
<taxonomicName authority="Lange, 1969" authorityName="Lange" authorityYear="1969" box="[1061,1179,1206,1232]" class="Mammalia" family="Mustelidae" genus="Mustelictis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Carnivora" pageId="16" pageNumber="615" phylum="Chordata" rank="genus">
<emphasis box="[1061,1179,1206,1232]" italics="true" pageId="16" pageNumber="615">Mustelictis</emphasis>
</taxonomicName>
, which differs in having a slightly lower m1 trigonid, a more shearing m1 talonid and a shorter m2.
</paragraph>
</subSubSection>
</treatment>
</document>