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<mods:title id="90E2252ABE43221A5F9453767D05F576">A revision of the cis-andean species of the genus Brycon Müller &amp; Troschel (Characiformes: Characidae)</mods:title>
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<mods:namePart id="D1F5BF74AD8DD28D8496E6F3A409E5CF">Lima, Flávio C. T.</mods:namePart>
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<taxonomicName id="8F9A1D854F54FFDB4EA4FE8BFD9AFE81" ID-CoL="NH7J" authority="Steindachner, 1877" authorityName="Steindachner" authorityYear="1877" box="[151,584,332,358]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="insignis">
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<emphasis id="7AEEBA144F54FFDB4EA4FE8BFE86FE81" bold="true" box="[151,340,332,358]" italics="true" pageId="35" pageNumber="36">Brycon insignis</emphasis>
<bibRefCitation id="2C0B1BF74F54FFDB4F6FFE8BFD9AFE81" author="Steindachner" box="[348,584,332,358]" pageId="35" pageNumber="36" refString="Steindachner, F. (1877) Die Susswasserfische des sudostlichen Brasilien (III). Sitzungsberichte der Kaiserlichen Akademie der Wissenchaften in Wien, 74 (1), 559 - 694, pls. 1 - 13." type="journal article" year="1877">Steindachner, 1877</bibRefCitation>
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(
<figureCitation id="D0A17A834F54FFDB4EACFEA9FEF4FE60" box="[159,294,366,391]" captionStart-0="FIGURE 22" captionStart-1="FIGURE 23" captionStart-2="FIGURE 24" captionStartId-0="38.[151,250,594,616]" captionStartId-1="38.[151,250,1070,1092]" captionStartId-2="38.[151,250,1614,1636]" captionTargetBox-0="[151,1436,193,573]" captionTargetBox-1="[151,1436,666,1049]" captionTargetBox-2="[151,1436,1142,1593]" captionTargetId-0="figure@38.[151,1436,193,573]" captionTargetId-1="figure@38.[151,1436,666,1049]" captionTargetId-2="figure@38.[151,1436,1142,1593]" captionTargetPageId-0="38" captionTargetPageId-1="38" captionTargetPageId-2="38" captionText-0="FIGURE 22. Brycon insignis, lectotype, NMW 62925: 2, 144.8 mm SL: Brazil, Rio de Janeiro, rio Paraíba do Sul." captionText-1="FIGURE 23. Brycon insignis, MZUSP 103045, 259.0 mm SL: Brazil, Rio de Janeiro, rio Paraíba do Sul." captionText-2="FIGURE 24. Brycon insignis, MZUSP 62746, 310.4 mm SL: Brazil, São Paulo, rio Paraíba do Sul." httpUri-0="https://zenodo.org/record/257791/files/figure.png" httpUri-1="https://zenodo.org/record/257792/files/figure.png" httpUri-2="https://zenodo.org/record/257793/files/figure.png" pageId="35" pageNumber="36">Figs. 22–24</figureCitation>
)
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<taxonomicName id="8F9A1D854F54FFDB4EA4FE7FFDE0FE2B" ID-CoL="NH7J" authority="Steindachner, 1877: 587" authorityName="Steindachner" authorityPageNumber="587" authorityYear="1877" box="[151,562,438,461]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F54FFDB4EA4FE7FFEE7FE2A" box="[151,309,440,461]" italics="true" pageId="35" pageNumber="36">Brycon insignis</emphasis>
<treatmentCitation id="C93B40174F54FFDB4F0FFE71FDE0FE2B" author="Steindachner" box="[316,562,438,460]" page="587" pageId="35" pageNumber="36" year="1877">
<bibRefCitation id="2C0B1BF74F54FFDB4F0FFE71FDD3FE2B" author="Steindachner" box="[316,513,438,460]" pageId="35" pageNumber="36" refString="Steindachner, F. (1877) Die Susswasserfische des sudostlichen Brasilien (III). Sitzungsberichte der Kaiserlichen Akademie der Wissenchaften in Wien, 74 (1), 559 - 694, pls. 1 - 13." type="journal article" year="1877">Steindachner, 1877</bibRefCitation>
: 587
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–591, pl. 4, figs.1, 1a (not 2, 2a; see 
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<bibRefCitation id="2C0B1BF74F54FFDB4D96FE71FBA4FE2B" author="Steindachner" box="[933,1142,438,460]" pageId="35" pageNumber="36" refString="Steindachner, F. (1879 c) Uber einige neue und seltene Fisch-Arten aus den K. K. Zoologischen Museen zu Wien, Stuttgart und Warschau. Denkschriften der Mathematisch-Naturwissenschaftlichen Classe der Akademie der Wissenschaften, Wien, 41, 1 - 52, pls. 1 - 9." type="journal article" year="1879" yearSuffix="c">Steindachner, 1879c</bibRefCitation>
: 50
</treatmentCitation>
, footnote: “Taf. IV, 
<figureCitation id="D0A17A834F54FFDB4B54FE71FA4EFE2B" box="[1383,1436,438,460]" captionStart="FIGURE 1" captionStartId="8.[151,250,1007,1029]" captionTargetBox="[162,1416,203,963]" captionTargetId="figure@8.[151,1436,193,985]" captionTargetPageId="8" captionText="FIGURE 1. Ventral view of premaxillary teeth arrangement in Brycon falcatus, MZUSP 94987, 368.7 mm SL. Scale bar: 5 mm." httpUri="https://zenodo.org/record/257770/files/figure.png" pageId="35" pageNumber="36">Fig.1</figureCitation>
[nicht 
<figureCitation id="D0A17A834F54FFDB4F38FE12FE99FE0C" box="[267,331,469,491]" captionStart="FIGURE 2" captionStartId="8.[151,250,1668,1690]" captionTargetBox="[164,1400,1148,1625]" captionTargetId="figure@8.[151,1436,1109,1647]" captionTargetPageId="8" captionText="FIGURE 2. Ventral view of premaxillary teeth arrangement in Brycon howesi, MZUSP 103685, 207.0 mm SL. Scale bar: 2 mm." httpUri="https://zenodo.org/record/257771/files/figure.png" pageId="35" pageNumber="36">Fig. 2</figureCitation>
…]) (Type locality “Rio Parahyba bei Campos und Mendez; Rio Jequitinhonha”, restricted herein to “Rio Parahyba bei Campos und Mendez”); 
<treatmentCitation id="C93B40174F54FFDB4C7BFE33FCE9FDED" author="Steindachner" box="[584,827,500,522]" page="50" pageId="35" pageNumber="36" year="1879">
<bibRefCitation id="2C0B1BF74F54FFDB4C7BFE33FCCAFDED" author="Steindachner" box="[584,792,500,522]" pageId="35" pageNumber="36" refString="Steindachner, F. (1879 c) Uber einige neue und seltene Fisch-Arten aus den K. K. Zoologischen Museen zu Wien, Stuttgart und Warschau. Denkschriften der Mathematisch-Naturwissenschaftlichen Classe der Akademie der Wissenschaften, Wien, 41, 1 - 52, pls. 1 - 9." type="journal article" year="1879" yearSuffix="c">Steindachner, 1879c</bibRefCitation>
: 50
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(as a possible synonym of 
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<emphasis id="7AEEBA144F54FFDB4A7DFE32FB67FDED" box="[1102,1205,500,522]" italics="true" pageId="35" pageNumber="36">B. devillei</emphasis>
</taxonomicName>
); 
<treatmentCitation id="C93B40174F54FFDB4AF9FE32FAABFDED" author="Howes" box="[1226,1401,500,522]" page="24" pageId="35" pageNumber="36" year="1982">
<bibRefCitation id="2C0B1BF74F54FFDB4AF9FE32FA86FDED" author="Howes" box="[1226,1364,500,522]" pageId="35" pageNumber="36" refString="Howes, G. J. (1982) Review of the genus Brycon (Teleostei: Characoidei). Bulletin of the British Museum (Natural History), Zoology, 43 (1), 1 - 47." type="journal article" year="1982">Howes, 1982</bibRefCitation>
: 24
</treatmentCitation>
, 32 (discussion; possible synonym with 
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<emphasis id="7AEEBA144F54FFDB4C0EFDD4FD77FDCF" box="[573,677,530,552]" italics="true" pageId="35" pageNumber="36">B. devillei</emphasis>
</taxonomicName>
); 
<treatmentCitation id="C93B40174F54FFDB4C8FFDD5FBD9FDCF" author="Almeida-Toledo" box="[700,1035,530,552]" page="36" pageId="35" pageNumber="36" year="1996">
<bibRefCitation id="2C0B1BF74F54FFDB4C8FFDD5FC36FDCF" author="Almeida-Toledo" box="[700,996,530,552]" pageId="35" pageNumber="36" refString="Almeida-Toledo, L. F., Bigoni, A. P., Bernardino, G., Foresti, F. &amp; Toledo-Filho, S. A. (1996) Karyotype and NOR conservatism with heterochromatin reorganization in Neotropical Bryconids. Caryologia, 49 (1), 35 - 43." type="journal article" year="1996">
Almeida-Toledo 
<emphasis id="7AEEBA144F54FFDB4D58FDD4FC4DFDCF" box="[875,927,530,552]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
, 1996
</bibRefCitation>
: 36
</treatmentCitation>
, 38 (karyotype); 
<treatmentCitation id="C93B40174F54FFDB4A8EFDD4FF3EFDA0" author="Menezes" page="110" pageId="35" pageNumber="36" year="2007">
<bibRefCitation id="2C0B1BF74F54FFDB4A8EFDD4FA4AFDCF" author="Menezes" box="[1213,1432,530,552]" pageId="35" pageNumber="36" refString="Menezes, N. A., Weitzman, S. H., Oyakawa, O. T., Lima, F. C. T., Castro, R. M. C. &amp; Weitzman, M. J. (2007) Freshwater fishes of Mata Atlantica. Preliminary list of species and comments on conservation of Neotropical freshwater fishes. Museu de Zoologia da Universidade de Sao Paulo, Sao Paulo, 405 pp." type="book" year="2007">
Menezes 
<emphasis id="7AEEBA144F54FFDB4B2CFDD4FA87FDCF" box="[1311,1365,530,552]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
, 2007
</bibRefCitation>
: 110
</treatmentCitation>
(photograph; distribution, ecology, conservation status); Hilsdorf 
<emphasis id="7AEEBA144F54FFDB4D9CFDF5FC3AFDA0" box="[943,1000,561,583]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
, 2008: 48–50, 277 (photo) (distribution, ecology, conservation); Matsumoto &amp; 
<treatmentCitation id="C93B40174F54FFDB4C53FD97FCE2FD81" author="Hilsdorf" box="[608,816,592,614]" page="395" pageId="35" pageNumber="36" year="2009">
<bibRefCitation id="2C0B1BF74F54FFDB4C53FD97FD2EFD81" author="Hilsdorf" box="[608,764,592,614]" pageId="35" pageNumber="36" refString="Oyakawa, O. T., Menezes, N. A., Shibatta, O. A., Lima, F. C. T., Langeani, F., Pavanelli, C. S., Nielsen, D. T. B. &amp; Hilsdorf, A. W. S. (2009) Peixes de agua doce. In: Bressan, P. M., Kierulff, M. C. M. &amp; Sugieda, A. M. (Orgs.), Fauna ameacada de extincao no estado de Sao Paulo - vertebrados. Fundacao Parque Zoologico de Sao Paulo, Secretaria do Meio Ambiente, Sao Paulo, pp. 349 - 424" type="book chapter" year="2009">Hilsdorf, 2009</bibRefCitation>
: 395
</treatmentCitation>
–402 (Brazil, rio Paraíba do Sul basin; genetic variation); 
<treatmentCitation id="C93B40174F54FFDB4EF4FDA9FE3BFD63" author="Travenzoli" box="[199,489,622,645]" page="6" pageId="35" pageNumber="36" year="2015">
<bibRefCitation id="2C0B1BF74F54FFDB4EF4FDA9FE1AFD63" author="Travenzoli" box="[199,456,622,645]" pageId="35" pageNumber="36" refString="Travenzoli, N. M., Silva, P. C., Santos, U., Zanuncio, J. C., Oliveira, C. &amp; Dergam, J. A. (2015). Cytogenetic and molecular data demonstrate that the Bryconinae (Ostariophysi, Bryconidae) species from southeastern Brazil form a phylogenetic and phylogeographic unit. PLOS ONE, 10 (9), 1 - 18. http: // dx. doi. org / 10.1371 / journal. pone. 0137843" type="journal article" year="2015">
Travenzoli 
<emphasis id="7AEEBA144F54FFDB4F71FDB7FEAFFD63" box="[322,381,622,645]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
, 2015
</bibRefCitation>
: 6
</treatmentCitation>
, 8–11, 14 (Rio Paraíba do Sul, Itaocara, Rio de Janeiro; cytogenetics, phylogenetics relationships, molecular taxonomy)..
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</paragraph>
<paragraph id="482566064F54FFDB4EA4FD6BFE92FD07" blockId="35.[151,1437,438,1074]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="688A41284F54FFDB4EA4FD6BFE92FD07" pageId="35" pageNumber="36">
<taxonomicName id="8F9A1D854F54FFDB4EA4FD6BFD3CFD25" ID-CoL="RQZJ" authority="Eigenmann &amp; Norris, 1900: 358" authorityName="Eigenmann &amp; Norris" authorityPageNumber="358" authorityYear="1900" box="[151,750,684,706]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F54FFDB4EA4FD6BFEAEFD25" box="[151,380,684,706]" italics="true" pageId="35" pageNumber="36">Catabasis acuminatus</emphasis>
<treatmentCitation id="C93B40174F54FFDB4FA0FD6BFD3CFD25" author="Eigenmann" box="[403,750,684,706]" page="358" pageId="35" pageNumber="36" year="1900">
<bibRefCitation id="2C0B1BF74F54FFDB4FA0FD6BFD68FD25" author="Eigenmann" box="[403,698,684,706]" pageId="35" pageNumber="36" refString="Eigenmann, C. H. &amp; Norris, A. A. (1900) Sobre alguns peixes de S. Paulo, Brasil. Revista do Museu Paulista, 4, 349 - 362." type="journal article" year="1900">Eigenmann &amp; Norris, 1900</bibRefCitation>
: 358
</treatmentCitation>
</taxonomicName>
(Type locality “Rio Tieté”); 
<treatmentCitation id="C93B40174F54FFDB4A18FD6BFB24FD25" author="Roberts" box="[1067,1270,684,706]" page="437" pageId="35" pageNumber="36" year="1969">
<bibRefCitation id="2C0B1BF74F54FFDB4A18FD6BFB13FD25" author="Roberts" box="[1067,1217,684,706]" pageId="35" pageNumber="36" refString="Roberts, T. R. (1969) Osteology and relationships of characoid fishes, particularly the genera Hepsetus, Salminus, Hoplias, Ctenolucius, and Acestrorhynchus. Proceedings of the California Academy of Sciences, 36 (15), 391 - 500." type="journal article" year="1969">Roberts, 1969</bibRefCitation>
: 437
</treatmentCitation>
–438 (holotype, discussion).
</treatmentCitationGroup>
</paragraph>
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<treatmentCitationGroup id="688A41284F54FFDB4EA4FD2EFBB3FD18" box="[151,1121,745,767]" pageId="35" pageNumber="36">
<taxonomicName id="8F9A1D854F54FFDB4EA4FD2EFD6AFD18" ID-CoL="7382H" authority="A. Miranda-Ribeiro, 1902: 5" authorityName="A. Miranda-Ribeiro" authorityPageNumber="5" authorityYear="1902" box="[151,696,745,767]" family="Bryconidae" genus="Megalobrycon" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="piabanha">
<emphasis id="7AEEBA144F54FFDB4EA4FD2EFE43FD18" box="[151,401,745,767]" italics="true" pageId="35" pageNumber="36">Megalobrycon piabanha</emphasis>
<treatmentCitation id="C93B40174F54FFDB4FA4FD2DFD6AFD18" author="Miranda-Ribeiro" box="[407,696,745,767]" page="5" pageId="35" pageNumber="36" year="1902">
<bibRefCitation id="2C0B1BF74F54FFDB4FA4FD2DFD70FD18" author="Miranda-Ribeiro" box="[407,674,745,767]" pageId="35" pageNumber="36" refString="Miranda-Ribeiro, A. (1902) Oito especies de peixes do Rio Pomba. A Lavoura, Boletim da Sociedade de Agricultura, 1902, 1 - 8." type="journal article" year="1902">A. Miranda-Ribeiro, 1902</bibRefCitation>
: 5
</treatmentCitation>
</taxonomicName>
(Type locality “Cataguazes, rio Pomba”).
</treatmentCitationGroup>
</paragraph>
<paragraph id="482566064F54FFDB4EA4FCCFFBF5FCF9" blockId="35.[151,1437,438,1074]" box="[151,1063,776,798]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="688A41284F54FFDB4EA4FCCFFBF5FCF9" box="[151,1063,776,798]" pageId="35" pageNumber="36">
<taxonomicName id="8F9A1D854F54FFDB4EA4FCCFFD0AFCF9" authority="A. Miranda-Ribeiro, 1905: 153" authorityName="A. Miranda-Ribeiro" authorityPageNumber="153" authorityYear="1905" box="[151,728,776,798]" family="Bryconidae" genus="Megalobrycon" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="piabagna">
<emphasis id="7AEEBA144F54FFDB4EA4FCCFFE43FCF9" box="[151,401,776,798]" italics="true" pageId="35" pageNumber="36">Megalobrycon piabagna</emphasis>
: 
<treatmentCitation id="C93B40174F54FFDB4FADFCCFFD0AFCF9" author="Miranda-Ribeiro" box="[414,728,776,798]" page="153" pageId="35" pageNumber="36" year="1905">
<bibRefCitation id="2C0B1BF74F54FFDB4FADFCCFFD7BFCF9" author="Miranda-Ribeiro" box="[414,681,776,798]" pageId="35" pageNumber="36" refString="Miranda-Ribeiro, A. (1905) Genus Megalobrycon, Gnthr. Seu enumeratio systematica hujus generis Characinidarum specierum. Arquivos do Museu Nacional, 13, 147 - 154, 1 pl." type="journal article" year="1905">
<collectingRegion id="8A5EA8E44F54FFDB4FADFCCFFE61FCFA" box="[414,435,776,797]" country="Nigeria" name="Rivers" pageId="35" pageNumber="36">A</collectingRegion>
. Miranda-Ribeiro, 1905
</bibRefCitation>
: 153
</treatmentCitation>
</taxonomicName>
–154 (rio Pomba, 
<collectingRegion id="8A5EA8E44F54FFDB4DA3FCCFFBCEFCF9" box="[912,1052,776,798]" country="Brazil" name="Minas Gerais" pageId="35" pageNumber="36">Minas Gerais</collectingRegion>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="482566064F54FFDB4EA4FCEFFA8CFCDB" blockId="35.[151,1437,438,1074]" box="[151,1374,806,829]" pageId="35" pageNumber="36">
<materialsCitation id="F8F26C5B4F54FFDB4EA4FCEFFA8CFCDB" ID-GBIF-Occurrence="1425222565" box="[151,1374,806,829]" collectionCode="MNHN" county="Muller" location="Pellegrin" municipality="Troschel" pageId="35" pageNumber="36" specimenCode="MNHN 1909-214" specimenCount="1" stateProvince="Rio Grande">
<treatmentCitationGroup id="688A41284F54FFDB4EA4FCEFFA8CFCDB" box="[151,1374,806,829]" pageId="35" pageNumber="36">
<taxonomicName id="8F9A1D854F54FFDB4EA4FCEFFEE4FCDB" ID-CoL="NH7B" box="[151,310,806,829]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="falcatus">
<emphasis id="7AEEBA144F54FFDB4EA4FCEFFEE4FCDB" box="[151,310,806,829]" italics="true" pageId="35" pageNumber="36">Brycon falcatus</emphasis>
</taxonomicName>
(non 
<collectingCounty id="A1441E8A4F54FFDB4F42FCE1FE64FCDB" box="[369,438,806,828]" pageId="35" pageNumber="36">Müller</collectingCounty>
&amp; 
<collectingMunicipality id="A841FC7C4F54FFDB4FE4FCE1FDE6FCDB" box="[471,564,806,828]" pageId="35" pageNumber="36">Troschel</collectingMunicipality>
): 
<treatmentCitation id="C93B40174F54FFDB4C77FCE1FCCDFCDB" author="Pellegrin" box="[580,799,806,828]" page="149" pageId="35" pageNumber="36" year="1909">
<bibRefCitation id="2C0B1BF74F54FFDB4C77FCE1FD22FCDB" author="Pellegrin" box="[580,752,806,828]" pageId="35" pageNumber="36" refString="Pellegrin, J. (1909 b) Characinides du Bresil rapportes par M. Jobert. Bulletin du Museum National d'Histoire Naturelle, 15, 147 - 153." type="journal article" year="1909" yearSuffix="b">
<location id="4D4530DD4F54FFDB4C77FCE1FD70FCDB" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F54FFDB4C77FCE1FD70FCDB" box="[580,674,806,828]" county="Muller" municipality="Troschel" name="Pellegrin" pageId="35" pageNumber="36" stateProvince="Rio Grande">Pellegrin</location>
, 1909b
</bibRefCitation>
: 149
</treatmentCitation>
(“ 
<collectingRegion id="8A5EA8E44F54FFDB4D05FCE1FC60FCDB" box="[822,946,806,828]" country="Puerto Rico" name="Rio Grande" pageId="35" pageNumber="36">
<collectingRegion id="8A5EA8E44F54FFDB4D05FCE1FC8FFCDB" box="[822,861,806,828]" country="Nigeria" name="Rivers" pageId="35" pageNumber="36">Rio</collectingRegion>
Grande
</collectingRegion>
”; specimen 
<specimenCode id="183CCE7D4F54FFDB4A1EFCE0FB34FCDB" box="[1069,1254,806,828]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="36">
<collectionCode id="2E8BFEC34F54FFDB4A1EFCE0FBA8FCDB" box="[1069,1146,807,828]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="35" pageNumber="36">MNHN</collectionCode>
1909–214
</specimenCode>
examined).
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F54FFDB4EA4FC82FB62FCBC" box="[151,1200,837,859]" pageId="35" pageNumber="36" type="description">
<paragraph id="482566064F54FFDB4EA4FC82FB62FCBC" blockId="35.[151,1437,438,1074]" box="[151,1200,837,859]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="688A41284F54FFDB4EA4FC82FB62FCBC" box="[151,1200,837,859]" pageId="35" pageNumber="36">
<taxonomicName id="8F9A1D854F54FFDB4EA4FC82FE43FCBC" box="[151,401,837,859]" family="Bryconidae" genus="Megalobrycon" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="piabanga">
<emphasis id="7AEEBA144F54FFDB4EA4FC82FE43FCBC" box="[151,401,837,859]" italics="true" pageId="35" pageNumber="36">Megalobrycon piabanga</emphasis>
</taxonomicName>
(sic): 
<treatmentCitation id="C93B40174F54FFDB4FE2FC82FD63FCBC" author="Magalhaes" box="[465,689,837,859]" page="155" pageId="35" pageNumber="36" year="1931">
<bibRefCitation id="2C0B1BF74F54FFDB4FE2FC82FD50FCBC" author="Magalhaes" box="[465,642,837,859]" pageId="35" pageNumber="36" refString="Magalhaes, A. C. (1931) Monographia brazileira de peixes fluviaes. Graphicars, Sao Paulo, 260 pp." type="book" year="1931">Magalhães, 1931</bibRefCitation>
: 155
</treatmentCitation>
–156, fig. 81 (Rio Paraíba do Sul; natural history).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F54FFDB4EA4FCA2FD2FFC7F" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="482566064F54FFDB4EA4FCA2FC00FC9D" blockId="35.[151,1437,438,1074]" box="[151,978,868,890]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="688A41284F54FFDB4EA4FCA2FC00FC9D" box="[151,978,868,890]" pageId="35" pageNumber="36">
<taxonomicName id="8F9A1D854F54FFDB4EA4FCA2FDDAFC9D" ID-CoL="RQZJ" authority="Howes, 1982: 4" authorityName="Howes" authorityPageNumber="4" authorityYear="1982" box="[151,520,868,890]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F54FFDB4EA4FCA2FE89FC9D" box="[151,347,869,890]" italics="true" pageId="35" pageNumber="36">Brycon acuminatus</emphasis>
: 
<treatmentCitation id="C93B40174F54FFDB4F5BFCA2FDDAFC9D" author="Howes" box="[360,520,868,890]" page="4" pageId="35" pageNumber="36" year="1982">
<bibRefCitation id="2C0B1BF74F54FFDB4F5BFCA2FE23FC9D" author="Howes" box="[360,497,868,890]" pageId="35" pageNumber="36" refString="Howes, G. J. (1982) Review of the genus Brycon (Teleostei: Characoidei). Bulletin of the British Museum (Natural History), Zoology, 43 (1), 1 - 47." type="journal article" year="1982">Howes, 1982</bibRefCitation>
: 4
</treatmentCitation>
</taxonomicName>
–6 (Taubaté, Rio Paraíba do Sul; discussion).
</treatmentCitationGroup>
</paragraph>
<paragraph id="482566064F54FFDB4EA4FC44FD2FFC7F" blockId="35.[151,1437,438,1074]" box="[151,765,898,920]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="688A41284F54FFDB4EA4FC44FD2FFC7F" box="[151,765,898,920]" pageId="35" pageNumber="36">
<taxonomicName id="8F9A1D854F54FFDB4EA4FC44FF33FC7F" ID-CoL="3DKB" box="[151,225,899,920]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="genus">
<emphasis id="7AEEBA144F54FFDB4EA4FC44FF33FC7F" box="[151,225,899,920]" italics="true" pageId="35" pageNumber="36">Brycon</emphasis>
</taxonomicName>
sp.: 
<treatmentCitation id="C93B40174F54FFDB4F23FC45FE11FC7F" author="Bizerril" box="[272,451,898,920]" page="69" pageId="35" pageNumber="36" year="1995">
<bibRefCitation id="2C0B1BF74F54FFDB4F23FC45FE73FC7F" author="Bizerril" box="[272,417,898,920]" pageId="35" pageNumber="36" refString="Bizerril, C. R. S. F. (1995) Estrutura quantitativa de comunidades de peixes em um rio costeiro do sudeste brasileiro. Acta Biologica Leopoldensia, 17 (2), 57 - 80." type="journal article" year="1995">Bizerril, 1995</bibRefCitation>
: 69
</treatmentCitation>
(
<collectingRegion id="8A5EA8E44F54FFDB4FE2FC45FDB5FC7F" box="[465,615,898,920]" country="Brazil" name="Rio de Janeiro" pageId="35" pageNumber="36">
<collectingRegion id="8A5EA8E44F54FFDB4FE2FC45FE25FC7F" box="[465,503,898,920]" country="Nigeria" name="Rivers" pageId="35" pageNumber="36">Rio</collectingRegion>
de Janeiro
</collectingRegion>
, rio São João).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F54FFDB4EA4FC65FB21FC50" box="[151,1267,929,951]" pageId="35" pageNumber="36" type="description">
<paragraph id="482566064F54FFDB4EA4FC65FB21FC50" blockId="35.[151,1437,438,1074]" box="[151,1267,929,951]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="688A41284F54FFDB4EA4FC65FB21FC50" box="[151,1267,929,951]" pageId="35" pageNumber="36">
<taxonomicName id="8F9A1D854F54FFDB4EA4FC65FEE3FC50" ID-CoL="NH77" box="[151,305,929,951]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="devillei">
<emphasis id="7AEEBA144F54FFDB4EA4FC65FEE3FC50" box="[151,305,929,951]" italics="true" pageId="35" pageNumber="36">Brycon devillei</emphasis>
</taxonomicName>
(non Castelnau): 
<treatmentCitation id="C93B40174F54FFDB4FD6FC66FD4DFC50" author="Araujo" box="[485,671,929,951]" page="116" pageId="35" pageNumber="36" year="1996">
<bibRefCitation id="2C0B1BF74F54FFDB4FD6FC66FDBCFC50" author="Araujo" box="[485,622,929,951]" pageId="35" pageNumber="36" refString="Araujo, F. G. (1996) Composicao e estrutura da comunidade de peixes do medio e baixo rio Paraiba do Sul, RJ. Revista Brasileira de Biologia, 56 (1), 111 - 126." type="journal article" year="1996">Araújo, 1996</bibRefCitation>
: 116
</treatmentCitation>
, 119, 121 (Rio de Janeiro, rio Paraíba do Sul, São Fidélis).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F54FFDB4EA4FC06FE6CFBD5" pageId="35" pageNumber="36" type="reference_group">
<paragraph id="482566064F54FFDB4EA4FC06FE6CFBD5" blockId="35.[151,1437,438,1074]" pageId="35" pageNumber="36">
<taxonomicName id="8F9A1D854F54FFDB4EA4FC06FEEDFC31" ID-CoL="NH83" box="[151,319,960,982]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="opalinus">
<emphasis id="7AEEBA144F54FFDB4EA4FC06FEEDFC31" box="[151,319,960,982]" italics="true" pageId="35" pageNumber="36">Brycon opalinus</emphasis>
</taxonomicName>
(non Cuvier): 
<bibRefCitation id="2C0B1BF74F54FFDB4FEAFC07FD19FC31" author="Bizerril" box="[473,715,960,982]" pageId="35" pageNumber="36" refString="Bizerril, C. R. S. F. &amp; Primo, P. B. S. (2001) Peixes de aguas interiores do estado do Rio de Janeiro. Fundacao de Estudos do Mar / Secretaria de Estado de Meio Ambiente e Desenvolvimento Sustentavel, Rio de Janeiro, 417 pp." type="book" year="2001">Bizerril &amp; Primo, 2001</bibRefCitation>
: 54–56, 201, 208, 231, 255, 259, 263, fig. 27 (
<collectingRegion id="8A5EA8E44F54FFDB4A9EFC07FA95FC31" box="[1197,1351,960,982]" country="Brazil" name="Rio de Janeiro" pageId="35" pageNumber="36">
<collectingRegion id="8A5EA8E44F54FFDB4A9EFC07FB01FC31" box="[1197,1235,960,982]" country="Nigeria" name="Rivers" pageId="35" pageNumber="36">Rio</collectingRegion>
de Janeiro
</collectingRegion>
: rio São João; rio Macaé; Lagoa Feia at rio Ururaí and rio Macabu; rio Itabapoana; rio 
<collectingRegion id="8A5EA8E44F54FFDB4DE8FC19FBF5FC13" box="[987,1063,990,1012]" country="Brazil" name="Paraiba" pageId="35" pageNumber="36">Paraíba</collectingRegion>
do 
<collectingRegion id="8A5EA8E44F54FFDB4A7EFC19FBBDFC13" box="[1101,1135,990,1012]" country="Cameroon" name="South" pageId="35" pageNumber="36">Sul</collectingRegion>
and its tributaries rio Muriaé, rio Pomba, and rio Imbé at Lagoa de Cima); 
<bibRefCitation id="2C0B1BF74F54FFDB4CA3FC3AFCB8FBF4" author="Bidegain" box="[656,874,1021,1043]" pageId="35" pageNumber="36" refString="Bidegain, P., Bizerril, C. R. S. F. &amp; Soffiatti, A. (2002) Lagoas do norte fluminense - perfil ambiental. SEMADS, Rio de Janeiro, 148 pp." type="book" year="2002">
Bidegain 
<emphasis id="7AEEBA144F54FFDB4CC0FC39FCF4FBF4" box="[755,806,1021,1043]" italics="true" pageId="35" pageNumber="36">et al.</emphasis>
, 2002
</bibRefCitation>
: 81, 90 (Lagoa Feia system, 
<collectingRegion id="8A5EA8E44F54FFDB4ABDFC3AFAF7FBF4" box="[1166,1317,1021,1043]" country="Brazil" name="Rio de Janeiro" pageId="35" pageNumber="36">
<collectingRegion id="8A5EA8E44F54FFDB4ABDFC3AFB60FBF4" box="[1166,1202,1021,1043]" country="Nigeria" name="Rivers" pageId="35" pageNumber="36">Rio</collectingRegion>
de Janeiro
</collectingRegion>
; occurence, importance in fisheries).
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F54FFDB4EA4FB99FF3DF8C4" pageId="35" pageNumber="36" type="diagnosis">
<paragraph id="482566064F54FFDB4EA4FB99FF3DF8C4" blockId="35.[151,1436,1118,2007]" pageId="35" pageNumber="36">
<emphasis id="7AEEBA144F54FFDB4EA4FB99FEC3FB90" bold="true" box="[151,273,1118,1143]" pageId="35" pageNumber="36">Diagnosis.</emphasis>
<taxonomicName id="8F9A1D854F54FFDB4F2AFB98FE1AFB91" box="[281,456,1119,1142]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F54FFDB4F2AFB98FE1AFB91" box="[281,456,1119,1142]" italics="true" pageId="35" pageNumber="36">Brycon insignis</emphasis>
</taxonomicName>
can be diagnosed from all remaining cis-andean 
<taxonomicName id="8F9A1D854F54FFDB4DC7FB98FB97FB91" box="[1012,1093,1119,1142]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="genus">
<emphasis id="7AEEBA144F54FFDB4DC7FB98FB97FB91" box="[1012,1093,1119,1142]" italics="true" pageId="35" pageNumber="36">Brycon</emphasis>
</taxonomicName>
species, with the exception of 
<taxonomicName id="8F9A1D854F54FFDB4EA4FB43FEE3FB7C" box="[151,305,1154,1179]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="stolzmanni">
<emphasis id="7AEEBA144F54FFDB4EA4FB43FEE3FB7C" box="[151,305,1154,1179]" italics="true" pageId="35" pageNumber="36">B. stolzmanni</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F54FFDB4F72FB43FE7BFB7C" box="[321,425,1156,1179]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="coxeyi">
<emphasis id="7AEEBA144F54FFDB4F72FB43FE7BFB7C" box="[321,425,1156,1179]" italics="true" pageId="35" pageNumber="36">B. coxeyi</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F54FFDB4F8AFB43FD9CFB7C" box="[441,590,1156,1179]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="coquenani">
<emphasis id="7AEEBA144F54FFDB4F8AFB43FD9CFB7C" box="[441,590,1156,1179]" italics="true" pageId="35" pageNumber="36">B. coquenani</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F54FFDB4C6DFB43FD35FB7C" box="[606,743,1154,1179]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="vermelha">
<emphasis id="7AEEBA144F54FFDB4C6DFB43FD35FB7C" box="[606,743,1154,1179]" italics="true" pageId="35" pageNumber="36">B. vermelha</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F54FFDB4CC4FB43FCB4FB7C" box="[759,870,1154,1179]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F54FFDB4CC4FB43FCB4FB7C" box="[759,870,1154,1179]" italics="true" pageId="35" pageNumber="36">B. howesi</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F54FFDB4D45FB43FC08FB7C" box="[886,986,1154,1179]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="dulcis">
<emphasis id="7AEEBA144F54FFDB4D45FB43FC08FB7C" box="[886,986,1154,1179]" italics="true" pageId="35" pageNumber="36">B. dulcis</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F54FFDB4DDAFB43FB90FB7C" box="[1001,1090,1155,1179]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="ferox">
<emphasis id="7AEEBA144F54FFDB4DDAFB43FB90FB7C" box="[1001,1090,1155,1179]" italics="true" pageId="35" pageNumber="36">B. ferox</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F54FFDB4A60FB43FB60FB7C" box="[1107,1202,1156,1179]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="vonoi">
<emphasis id="7AEEBA144F54FFDB4A60FB43FB60FB7C" box="[1107,1202,1156,1179]" italics="true" pageId="35" pageNumber="36">B. vonoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F54FFDB4AF2FB43FA90FB7C" box="[1217,1346,1154,1179]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="opalinus">
<emphasis id="7AEEBA144F54FFDB4AF2FB43FA90FB7C" box="[1217,1346,1154,1179]" italics="true" pageId="35" pageNumber="36">B. opalinus</emphasis>
</taxonomicName>
, and 
<taxonomicName id="8F9A1D854F54FFDB4BB7FB43FF2AFB59" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="nattereri">
<emphasis id="7AEEBA144F54FFDB4BB7FB43FF2AFB59" italics="true" pageId="35" pageNumber="36">B. nattereri</emphasis>
</taxonomicName>
by possessing a color pattern consisting in a humeral blotch and a caudal peduncle blotch, without body stripes or other obvious color markings on caudal and anal-fins (vs. body stripes and caudal/anal fin color markings present) (see 
<figureCitation id="D0A17A834F54FFDB4F01FB29FEAAFAE0" box="[306,376,1262,1287]" captionStart="FIGURE 5" captionStartId="11.[151,250,1476,1498]" captionTargetBox="[164,1419,611,1450]" captionTargetId="figure@11.[151,1436,589,1454]" captionTargetPageId="11" captionText="FIGURE 5. Diagrammatic representation of the patterns of dark colour markings found among cis-andean Brycon species, represented by selected species. A. Brycon stolzmanni. B. Brycon coxeyi. C. Brycon insignis. D. Brycon nattereri. E. Brycon hilarii. F. Brycon whitei. G. Brycon amazonicus. H. Brycon gouldingi. I. Brycon melanopterus. J-L. Brycon falcatus." httpUri="https://zenodo.org/record/257774/files/figure.png" pageId="35" pageNumber="36">Fig. 5</figureCitation>
). 
<taxonomicName id="8F9A1D854F54FFDB4FA2FB28FD91FAE1" box="[401,579,1263,1286]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F54FFDB4FA2FB28FD91FAE1" box="[401,579,1263,1286]" italics="true" pageId="35" pageNumber="36">Brycon insignis</emphasis>
</taxonomicName>
can be diagnosed from these species, with the exception of 
<taxonomicName id="8F9A1D854F54FFDB4B30FB28FAA1FAE0" box="[1283,1395,1262,1287]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F54FFDB4B30FB28FAA1FAE0" box="[1283,1395,1262,1287]" italics="true" pageId="35" pageNumber="36">B. howesi</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F54FFDB4BB7FB28FF2DFACC" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="vermelha">
<emphasis id="7AEEBA144F54FFDB4BB7FB28FF2DFACC" italics="true" pageId="35" pageNumber="36">B. vermelha</emphasis>
</taxonomicName>
, and 
<taxonomicName id="8F9A1D854F54FFDB4F7AFAD3FE31FACC" box="[329,483,1300,1323]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="coquenani">
<emphasis id="7AEEBA144F54FFDB4F7AFAD3FE31FACC" box="[329,483,1300,1323]" italics="true" pageId="35" pageNumber="36">B. coquenani</emphasis>
</taxonomicName>
, by possessing a fifth infraorbital bone considerably wider than high (vs. fifth infraorbital bone about as wide as high; see 
<figureCitation id="D0A17A834F54FFDB4C95FAF1FD3CFAA8" box="[678,750,1334,1359]" captionStart="FIGURE 6" captionStartId="12.[151,250,1849,1871]" captionTargetBox="[419,1174,200,1801]" captionTargetId="figure@12.[358,1217,171,1822]" captionTargetPageId="12" captionText="FIGURE 6. Diagrammatic representation of the head profile, maxillary and infraorbitals showing the distinct morphologies found among cis-andean Brycon species, represented by selected species. A. Brycon amazonicus, MZUSP 3815, 152.3 mm SL. B. Brycon insignis, MCZ 21112, 232.2 mm SL. C. Brycon dulcis, MZUSP 1531, 146.6 mm SL. D. Brycon falcatus, MZUSP 3845, 195.1 mm SL. E. Brycon opalinus, MCZ 21123, 210.5 mm SL. F. Brycon orbignyanus, MZUSP 3900, 108.9 mm HL. G. Brycon ferox, MCZ 60934, 310.3 mm SL. H. Brycon orthotaenia, MZUSP 2007, 188.7 mm SL. I. Brycon polylepis, MZUSP 70413, 203.9 mm SL. J. Brycon stolzmanni, UMMZ 185306, 120.0 mm SL." httpUri="https://zenodo.org/record/257775/files/figure.png" pageId="35" pageNumber="36">Fig. 6</figureCitation>
). 
<taxonomicName id="8F9A1D854F54FFDB4D34FAF0FC69FAA9" box="[775,955,1335,1358]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F54FFDB4D34FAF0FC69FAA9" box="[775,955,1335,1358]" italics="true" pageId="35" pageNumber="36">Brycon insignis</emphasis>
</taxonomicName>
can be diagnosed from 
<taxonomicName id="8F9A1D854F54FFDB4AEFFAF0FAA7FAA9" box="[1244,1397,1335,1358]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="coquenani">
<emphasis id="7AEEBA144F54FFDB4AEFFAF0FAA7FAA9" box="[1244,1397,1335,1358]" italics="true" pageId="35" pageNumber="36">B. coquenani</emphasis>
</taxonomicName>
by possessing a higher head (63.1–75.9 % of head length, mean 68.9, vs. 59.8–64.9 %, mean 61.4, in 
<taxonomicName id="8F9A1D854F54FFDB4AC4FA9BFA59FA94" box="[1271,1419,1372,1395]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="coquenani">
<emphasis id="7AEEBA144F54FFDB4AC4FA9BFA59FA94" box="[1271,1419,1372,1395]" italics="true" pageId="35" pageNumber="36">B. coquenani</emphasis>
</taxonomicName>
), by possessing pointed tri- to pentacuspidate dentary teeth, with moderately developed lateral cusps (vs. pointed, trito tetracuspidate dentary teeth, with lateral cusps poorly developed; see, however, the item “Remarks”, below, on individuals formely assigned to 
<taxonomicName id="8F9A1D854F54FFDB4FCDFA01FD24FA39" box="[510,758,1478,1503]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F54FFDB4FCDFA01FD24FA39" box="[510,758,1478,1503]" italics="true" pageId="35" pageNumber="36">Catabasis acuminatus</emphasis>
</taxonomicName>
), and by mature males displaying hooks only in anal fin (vs. mature males displaying hooks in all rayed fins, including caudal fin). 
<taxonomicName id="8F9A1D854F54FFDB4D9BFA2BFB84F9E4" box="[936,1110,1516,1539]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F54FFDB4D9BFA2BFB84F9E4" box="[936,1110,1516,1539]" italics="true" pageId="35" pageNumber="36">Brycon insignis</emphasis>
</taxonomicName>
can be distinguished from 
<taxonomicName id="8F9A1D854F54FFDB4BB7FA2BFF2DF9C0" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="vermelha">
<emphasis id="7AEEBA144F54FFDB4BB7FA2BFF2DF9C0" italics="true" pageId="35" pageNumber="36">B. vermelha</emphasis>
</taxonomicName>
by presenting darkened dorsal, caudal, and anal-fins in both living and preserved specimens (vs. dorsal, caudal, and anal-fins reddish in living, clear in preserved, 
<taxonomicName id="8F9A1D854F54FFDB4D67F9F3FC33F9AC" box="[852,993,1586,1611]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="vermelha">
<emphasis id="7AEEBA144F54FFDB4D67F9F3FC33F9AC" box="[852,993,1586,1611]" italics="true" pageId="35" pageNumber="36">B. vermelha</emphasis>
</taxonomicName>
specimens). 
<taxonomicName id="8F9A1D854F54FFDB4ABFF9F3FA93F9AC" box="[1164,1345,1588,1611]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F54FFDB4ABFF9F3FA93F9AC" box="[1164,1345,1588,1611]" italics="true" pageId="35" pageNumber="36">Brycon insignis</emphasis>
</taxonomicName>
can be distiguished from 
<taxonomicName id="8F9A1D854F54FFDB4F57F990FDD8F988" box="[356,522,1622,1647]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F54FFDB4F57F990FDD8F988" box="[356,522,1622,1647]" italics="true" pageId="35" pageNumber="36">Brycon howesi</emphasis>
</taxonomicName>
by possessing 51–65, modally 58 lateral line scales (vs. 46–50, modally 
<quantity id="8F62CBE34F54FFDB4B70F990FAAFF988" box="[1347,1405,1622,1647]" metricMagnitude="0" metricUnit="m" metricValue="1.2446" pageId="35" pageNumber="36" unit="in" value="49.0">49 in</quantity>
<taxonomicName id="8F9A1D854F54FFDB4BB7F990FF37F974" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F54FFDB4BB7F990FF37F974" italics="true" pageId="35" pageNumber="36">B. howesi</emphasis>
</taxonomicName>
), 9–11, modally 10 horizontal scale rows between the dorsal-fin origin and the lateral line (vs. 7–9, modally 
<quantity id="8F62CBE34F54FFDB4EA4F958FF10F950" box="[151,194,1694,1719]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="35" pageNumber="36" unit="in" value="8.0">8 in</quantity>
<taxonomicName id="8F9A1D854F54FFDB4EFBF958FEE7F950" box="[200,309,1694,1719]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F54FFDB4EFBF958FEE7F950" box="[200,309,1694,1719]" italics="true" pageId="35" pageNumber="36">B. howesi</emphasis>
</taxonomicName>
), 4–7, modally 5 horizontal scales rows between the lateral line and pelvic fin (vs. 3–5, modally 
<quantity id="8F62CBE34F54FFDB4B41F958FA4EF950" box="[1394,1436,1694,1719]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="35" pageNumber="36" unit="in" value="4.0">4 in</quantity>
<taxonomicName id="8F9A1D854F54FFDB4EA4F903FED4F93C" box="[151,262,1730,1755]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F54FFDB4EA4F903FED4F93C" box="[151,262,1730,1755]" italics="true" pageId="35" pageNumber="36">B. howesi</emphasis>
</taxonomicName>
) and a pointed snout in profile in 
<taxonomicName id="8F9A1D854F54FFDB4CA4F903FCDDF93C" box="[663,783,1732,1755]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F54FFDB4CA4F903FCDDF93C" box="[663,783,1732,1755]" italics="true" pageId="35" pageNumber="36">B. insignis</emphasis>
</taxonomicName>
(vs. snout pointed, but rounder, in 
<taxonomicName id="8F9A1D854F54FFDB4A9AF903FACAF93C" box="[1193,1304,1730,1755]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F54FFDB4A9AF903FACAF93C" box="[1193,1304,1730,1755]" italics="true" pageId="35" pageNumber="36">B. howesi</emphasis>
</taxonomicName>
). For more comparisons among 
<taxonomicName id="8F9A1D854F54FFDB4FB4F920FDE9F919" box="[391,571,1767,1790]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F54FFDB4FB4F920FDE9F919" box="[391,571,1767,1790]" italics="true" pageId="35" pageNumber="36">Brycon insignis</emphasis>
</taxonomicName>
, 
<taxonomicName id="8F9A1D854F54FFDB4C7EF920FD0BF918" box="[589,729,1766,1791]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="vermelha">
<emphasis id="7AEEBA144F54FFDB4C7EF920FD0BF918" box="[589,729,1766,1791]" italics="true" pageId="35" pageNumber="36">B. vermelha</emphasis>
</taxonomicName>
, and 
<taxonomicName id="8F9A1D854F54FFDB4D2CF920FC43F918" box="[799,913,1766,1791]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="35" pageNumber="36" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F54FFDB4D2CF920FC43F918" box="[799,913,1766,1791]" italics="true" pageId="35" pageNumber="36">B. howesi</emphasis>
</taxonomicName>
, see the item “Remarks” of these latter two species.
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F54FFDE4EF4F8E9FCCBF92F" lastPageId="38" lastPageNumber="39" pageId="35" pageNumber="36" type="description">
<paragraph id="482566064F54FFDC4EF4F8E9FE13FF10" blockId="35.[151,1436,1118,2007]" lastBlockId="36.[151,1436,151,248]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
<emphasis id="7AEEBA144F54FFDB4EF4F8E9FE8BF8A0" bold="true" box="[199,345,1838,1863]" pageId="35" pageNumber="36">Description.</emphasis>
Morphometric data are presented in 
<tableCitation id="051853BD4F54FFDB4D3FF8E9FCB4F8A0" box="[780,870,1838,1863]" captionStart="TABLE 5" captionStartId="36.[151,239,296,319]" captionTargetBox="[159,1400,379,1351]" captionTargetPageId="36" captionText="TABLE 5. Morphometric data of Brycon insignis (A: lectotype, NMW 62925: 2; B: holotype of Catabasis acuminatus, CAS 11894)." httpUri="http://table.plazi.org/id/1CE5368E4F53FFDC4EA4FEEFFEF1FEB8" pageId="35" pageNumber="36" tableUuid="1CE5368E4F53FFDC4EA4FEEFFEF1FEB8">Table 5</tableCitation>
. Large-sized species, largest examined specimen 493.0 mm SL. Body moderately slender. Largest body height slightly ahead of dorsal-fin origin. Dorsal body profile slightly convex from upper lip margin to vertical through anterior naris, moderately convex from latter point to basis of supraoccipital process, and moderately convex from latter point to dorsal-fin origin, straight along dorsal-fin basis, and straight to slightly convex from dorsal-fin basis to adipose-fin origin. Dorsal profile of caudal peduncle slightly concave. Ventral profile slightly convex from lower lip to pelvic-fin insertion, straight to slightly convex from this point to anal-fin origin and approximately straight along anal-fin base. Ventral profile of caudal peduncle slightly concave.
</paragraph>
<caption id="1CE5368E4F53FFDC4EA4FEEFFEF1FEB8" ID-Table-UUID="1CE5368E4F53FFDC4EA4FEEFFEF1FEB8" httpUri="http://table.plazi.org/id/1CE5368E4F53FFDC4EA4FEEFFEF1FEB8" pageId="36" pageNumber="37" targetBox="[159,1400,379,1351]" targetIsTable="true" targetPageId="36">
<paragraph id="482566064F53FFDC4EA4FEEFFEF1FEB8" blockId="36.[151,1436,295,351]" pageId="36" pageNumber="37">
<emphasis id="7AEEBA144F53FFDC4EA4FEEFFEDBFED8" bold="true" box="[151,265,296,319]" pageId="36" pageNumber="37">TABLE 5.</emphasis>
Morphometric data of 
<taxonomicName id="8F9A1D854F53FFDC4C35FEEFFD7CFED8" box="[518,686,296,319]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F53FFDC4C35FEEFFD7CFED8" box="[518,686,296,319]" italics="true" pageId="36" pageNumber="37">Brycon insignis</emphasis>
</taxonomicName>
(A: lectotype, NMW 62925:2; B: holotype of 
<taxonomicName id="8F9A1D854F53FFDC4A9BFEE0FA47FED8" box="[1192,1429,295,319]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="36" pageNumber="37" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F53FFDC4A9BFEE0FA47FED8" box="[1192,1429,295,319]" italics="true" pageId="36" pageNumber="37">Catabasis acuminatus</emphasis>
</taxonomicName>
, CAS 11894).
</paragraph>
</caption>
<paragraph id="482566064F53FFDC4CDEFEBCFAB8FAA0" pageId="36" pageNumber="37">
<table id="3A9A94A64F5300074EACFEBCFAAAFAA0" box="[159,1400,379,1351]" gridcols="6" gridrows="24" pageId="36" pageNumber="37">
<tr id="F6AA64444F5300074EACFEBCFAAAFE77" box="[159,1400,379,400]" gridrow="0" pageId="36" pageNumber="37" rowspan-0="1">
<th id="B57B0D384F5300074CDEFEBCFCF4FE77" box="[749,806,379,400]" gridcol="1" gridrow="0" pageId="36" pageNumber="37">A</th>
<th id="B57B0D384F5300074D49FEBCFC61FE77" box="[890,947,379,400]" gridcol="2" gridrow="0" pageId="36" pageNumber="37">B</th>
<th id="B57B0D384F5300074A34FEBCFBF3FE77" box="[1031,1057,379,400]" gridcol="3" gridrow="0" pageId="36" pageNumber="37">n</th>
<th id="B57B0D384F5300074A6FFEBCFB08FE77" box="[1116,1242,379,400]" gridcol="4" gridrow="0" pageId="36" pageNumber="37">Range</th>
<th id="B57B0D384F5300074B0DFEBCFAAAFE77" box="[1342,1400,379,400]" gridcol="5" gridrow="0" pageId="36" pageNumber="37">Mean</th>
</tr>
<tr id="F6AA64444F5300074EACFE63FAAAFE5D" box="[159,1400,420,442]" gridrow="1" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFE63FDAFFE5D" box="[159,637,420,442]" gridcol="0" gridrow="1" pageId="36" pageNumber="37">Standard length (SL)</th>
<td id="B57B0D384F5300074CDEFE63FCF4FE5D" box="[749,806,420,442]" gridcol="1" gridrow="1" pageId="36" pageNumber="37">144.8</td>
<td id="B57B0D384F5300074D49FE63FC61FE5D" box="[890,947,420,442]" gridcol="2" gridrow="1" pageId="36" pageNumber="37">135.1</td>
<td id="B57B0D384F5300074A34FE63FBF3FE5D" box="[1031,1057,420,442]" gridcol="3" gridrow="1" pageId="36" pageNumber="37">53</td>
<td id="B57B0D384F5300074A6FFE63FB08FE5D" box="[1116,1242,420,442]" gridcol="4" gridrow="1" pageId="36" pageNumber="37">113.5–365.5</td>
<td id="B57B0D384F5300074B0DFE63FAAAFE5D" box="[1342,1400,420,442]" gridcol="5" gridrow="1" pageId="36" pageNumber="37">-</td>
</tr>
<tr id="F6AA64444F5300074EACFE0AFAAAFE04" box="[159,1400,461,483]" gridrow="2" pageId="36" pageNumber="37" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="B57B0D384F5300074EACFE0AFDAFFE04" box="[159,637,461,483]" gridcol="0" gridrow="2" pageId="36" pageNumber="37">Percentages of standard length</th>
</tr>
<tr id="F6AA64444F5300074EACFE31FAAAFDEB" box="[159,1400,502,524]" gridrow="3" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFE31FDAFFDEB" box="[159,637,502,524]" gridcol="0" gridrow="3" pageId="36" pageNumber="37">Depth at dorsal-fin origin</th>
<td id="B57B0D384F5300074CDEFE31FCF4FDEB" box="[749,806,502,524]" gridcol="1" gridrow="3" pageId="36" pageNumber="37">26.4</td>
<td id="B57B0D384F5300074D49FE31FC61FDEB" box="[890,947,502,524]" gridcol="2" gridrow="3" pageId="36" pageNumber="37">27.5</td>
<td id="B57B0D384F5300074A34FE31FBF3FDEB" box="[1031,1057,502,524]" gridcol="3" gridrow="3" pageId="36" pageNumber="37">31</td>
<td id="B57B0D384F5300074A6FFE31FB08FDEB" box="[1116,1242,502,524]" gridcol="4" gridrow="3" pageId="36" pageNumber="37">25.0–32.0</td>
<td id="B57B0D384F5300074B0DFE31FAAAFDEB" box="[1342,1400,502,524]" gridcol="5" gridrow="3" pageId="36" pageNumber="37">28.1</td>
</tr>
<tr id="F6AA64444F5300074EACFDE7FAAAFDD1" box="[159,1400,544,566]" gridrow="4" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFDE7FDAFFDD1" box="[159,637,544,566]" gridcol="0" gridrow="4" pageId="36" pageNumber="37">Snout to dorsal-fin origin</th>
<td id="B57B0D384F5300074CDEFDE7FCF4FDD1" box="[749,806,544,566]" gridcol="1" gridrow="4" pageId="36" pageNumber="37">54.4</td>
<td id="B57B0D384F5300074D49FDE7FC61FDD1" box="[890,947,544,566]" gridcol="2" gridrow="4" pageId="36" pageNumber="37">52.3</td>
<td id="B57B0D384F5300074A34FDE7FBF3FDD1" box="[1031,1057,544,566]" gridcol="3" gridrow="4" pageId="36" pageNumber="37">53</td>
<td id="B57B0D384F5300074A6FFDE7FB08FDD1" box="[1116,1242,544,566]" gridcol="4" gridrow="4" pageId="36" pageNumber="37">48.7–55.7</td>
<td id="B57B0D384F5300074B0DFDE7FAAAFDD1" box="[1342,1400,544,566]" gridcol="5" gridrow="4" pageId="36" pageNumber="37">53.1</td>
</tr>
<tr id="F6AA64444F5300074EACFD8EFAAAFDB8" box="[159,1400,585,607]" gridrow="5" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFD8EFDAFFDB8" box="[159,637,585,607]" gridcol="0" gridrow="5" pageId="36" pageNumber="37">Dorsal-fin base length</th>
<td id="B57B0D384F5300074CDEFD8EFCF4FDB8" box="[749,806,585,607]" gridcol="1" gridrow="5" pageId="36" pageNumber="37">10.8</td>
<td id="B57B0D384F5300074D49FD8EFC61FDB8" box="[890,947,585,607]" gridcol="2" gridrow="5" pageId="36" pageNumber="37">9.3</td>
<td id="B57B0D384F5300074A34FD8EFBF3FDB8" box="[1031,1057,585,607]" gridcol="3" gridrow="5" pageId="36" pageNumber="37">53</td>
<td id="B57B0D384F5300074A6FFD8EFB08FDB8" box="[1116,1242,585,607]" gridcol="4" gridrow="5" pageId="36" pageNumber="37">9.3–13.3</td>
<td id="B57B0D384F5300074B0DFD8EFAAAFDB8" box="[1342,1400,585,607]" gridcol="5" gridrow="5" pageId="36" pageNumber="37">11.6</td>
</tr>
<tr id="F6AA64444F5300074EACFDB5FAAAFD6F" box="[159,1400,626,648]" gridrow="6" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFDB5FDAFFD6F" box="[159,637,626,648]" gridcol="0" gridrow="6" pageId="36" pageNumber="37">Posterior terminus of dorsal fin to adipose fin</th>
<td id="B57B0D384F5300074CDEFDB5FCF4FD6F" box="[749,806,626,648]" gridcol="1" gridrow="6" pageId="36" pageNumber="37">23.0</td>
<td id="B57B0D384F5300074D49FDB5FC61FD6F" box="[890,947,626,648]" gridcol="2" gridrow="6" pageId="36" pageNumber="37">20.7</td>
<td id="B57B0D384F5300074A34FDB5FBF3FD6F" box="[1031,1057,626,648]" gridcol="3" gridrow="6" pageId="36" pageNumber="37">53</td>
<td id="B57B0D384F5300074A6FFDB5FB08FD6F" box="[1116,1242,626,648]" gridcol="4" gridrow="6" pageId="36" pageNumber="37">20.7–25.7</td>
<td id="B57B0D384F5300074B0DFDB5FAAAFD6F" box="[1342,1400,626,648]" gridcol="5" gridrow="6" pageId="36" pageNumber="37">23.3</td>
</tr>
<tr id="F6AA64444F5300074EACFD5BFAAAFD55" box="[159,1400,668,690]" gridrow="7" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFD5BFDAFFD55" box="[159,637,668,690]" gridcol="0" gridrow="7" pageId="36" pageNumber="37">Posterior terminus of dorsal fin to hypural joint</th>
<td id="B57B0D384F5300074CDEFD5BFCF4FD55" box="[749,806,668,690]" gridcol="1" gridrow="7" pageId="36" pageNumber="37">37.3</td>
<td id="B57B0D384F5300074D49FD5BFC61FD55" box="[890,947,668,690]" gridcol="2" gridrow="7" pageId="36" pageNumber="37">34.7</td>
<td id="B57B0D384F5300074A34FD5BFBF3FD55" box="[1031,1057,668,690]" gridcol="3" gridrow="7" pageId="36" pageNumber="37">53</td>
<td id="B57B0D384F5300074A6FFD5BFB08FD55" box="[1116,1242,668,690]" gridcol="4" gridrow="7" pageId="36" pageNumber="37">32.4–39.9</td>
<td id="B57B0D384F5300074B0DFD5BFAAAFD55" box="[1342,1400,668,690]" gridcol="5" gridrow="7" pageId="36" pageNumber="37">36.3</td>
</tr>
<tr id="F6AA64444F5300074EACFD02FAAAFD3C" box="[159,1400,709,731]" gridrow="8" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFD02FDAFFD3C" box="[159,637,709,731]" gridcol="0" gridrow="8" pageId="36" pageNumber="37">Snout to pelvic-fin insertion</th>
<td id="B57B0D384F5300074CDEFD02FCF4FD3C" box="[749,806,709,731]" gridcol="1" gridrow="8" pageId="36" pageNumber="37">48.2</td>
<td id="B57B0D384F5300074D49FD02FC61FD3C" box="[890,947,709,731]" gridcol="2" gridrow="8" pageId="36" pageNumber="37">50.0</td>
<td id="B57B0D384F5300074A34FD02FBF3FD3C" box="[1031,1057,709,731]" gridcol="3" gridrow="8" pageId="36" pageNumber="37">30</td>
<td id="B57B0D384F5300074A6FFD02FB08FD3C" box="[1116,1242,709,731]" gridcol="4" gridrow="8" pageId="36" pageNumber="37">44.4–51.6</td>
<td id="B57B0D384F5300074B0DFD02FAAAFD3C" box="[1342,1400,709,731]" gridcol="5" gridrow="8" pageId="36" pageNumber="37">47.6</td>
</tr>
<tr id="F6AA64444F5300074EACFD29FAAAFCE3" box="[159,1400,750,772]" gridrow="9" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFD29FDAFFCE3" box="[159,637,750,772]" gridcol="0" gridrow="9" pageId="36" pageNumber="37">Snout to anal-fin origin</th>
<td id="B57B0D384F5300074CDEFD29FCF4FCE3" box="[749,806,750,772]" gridcol="1" gridrow="9" pageId="36" pageNumber="37">66.3</td>
<td id="B57B0D384F5300074D49FD29FC61FCE3" box="[890,947,750,772]" gridcol="2" gridrow="9" pageId="36" pageNumber="37">68.5</td>
<td id="B57B0D384F5300074A34FD29FBF3FCE3" box="[1031,1057,750,772]" gridcol="3" gridrow="9" pageId="36" pageNumber="37">52</td>
<td id="B57B0D384F5300074A6FFD29FB08FCE3" box="[1116,1242,750,772]" gridcol="4" gridrow="9" pageId="36" pageNumber="37">62.6–70.2</td>
<td id="B57B0D384F5300074B0DFD29FAAAFCE3" box="[1342,1400,750,772]" gridcol="5" gridrow="9" pageId="36" pageNumber="37">66.7</td>
</tr>
<tr id="F6AA64444F5300074EACFCDFFAAAFCC9" box="[159,1400,792,814]" gridrow="10" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFCDFFDAFFCC9" box="[159,637,792,814]" gridcol="0" gridrow="10" pageId="36" pageNumber="37">Anal-fin base length</th>
<td id="B57B0D384F5300074CDEFCDFFCF4FCC9" box="[749,806,792,814]" gridcol="1" gridrow="10" pageId="36" pageNumber="37">23.1</td>
<td id="B57B0D384F5300074D49FCDFFC61FCC9" box="[890,947,792,814]" gridcol="2" gridrow="10" pageId="36" pageNumber="37">-</td>
<td id="B57B0D384F5300074A34FCDFFBF3FCC9" box="[1031,1057,792,814]" gridcol="3" gridrow="10" pageId="36" pageNumber="37">50</td>
<td id="B57B0D384F5300074A6FFCDFFB08FCC9" box="[1116,1242,792,814]" gridcol="4" gridrow="10" pageId="36" pageNumber="37">22.5–27.5</td>
<td id="B57B0D384F5300074B0DFCDFFAAAFCC9" box="[1342,1400,792,814]" gridcol="5" gridrow="10" pageId="36" pageNumber="37">24.4</td>
</tr>
<tr id="F6AA64444F5300074EACFC86FAAAFCB0" box="[159,1400,833,855]" gridrow="11" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFC86FDAFFCB0" box="[159,637,833,855]" gridcol="0" gridrow="11" pageId="36" pageNumber="37">Caudal peduncle length</th>
<td id="B57B0D384F5300074CDEFC86FCF4FCB0" box="[749,806,833,855]" gridcol="1" gridrow="11" pageId="36" pageNumber="37">13.9</td>
<td id="B57B0D384F5300074D49FC86FC61FCB0" box="[890,947,833,855]" gridcol="2" gridrow="11" pageId="36" pageNumber="37">10.4</td>
<td id="B57B0D384F5300074A34FC86FBF3FCB0" box="[1031,1057,833,855]" gridcol="3" gridrow="11" pageId="36" pageNumber="37">53</td>
<td id="B57B0D384F5300074A6FFC86FB08FCB0" box="[1116,1242,833,855]" gridcol="4" gridrow="11" pageId="36" pageNumber="37">10.4–16.1</td>
<td id="B57B0D384F5300074B0DFC86FAAAFCB0" box="[1342,1400,833,855]" gridcol="5" gridrow="11" pageId="36" pageNumber="37">13.7</td>
</tr>
<tr id="F6AA64444F5300074EACFCADFAAAFC67" box="[159,1400,874,896]" gridrow="12" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFCADFDAFFC67" box="[159,637,874,896]" gridcol="0" gridrow="12" pageId="36" pageNumber="37">Dorsal-fin height</th>
<td id="B57B0D384F5300074CDEFCADFCF4FC67" box="[749,806,874,896]" gridcol="1" gridrow="12" pageId="36" pageNumber="37">19.6</td>
<td id="B57B0D384F5300074D49FCADFC61FC67" box="[890,947,874,896]" gridcol="2" gridrow="12" pageId="36" pageNumber="37">-</td>
<td id="B57B0D384F5300074A34FCADFBF3FC67" box="[1031,1057,874,896]" gridcol="3" gridrow="12" pageId="36" pageNumber="37">45</td>
<td id="B57B0D384F5300074A6FFCADFB08FC67" box="[1116,1242,874,896]" gridcol="4" gridrow="12" pageId="36" pageNumber="37">16.0–22.1</td>
<td id="B57B0D384F5300074B0DFCADFAAAFC67" box="[1342,1400,874,896]" gridcol="5" gridrow="12" pageId="36" pageNumber="37">20.0</td>
</tr>
<tr id="F6AA64444F5300074EACFC53FAAAFC4D" box="[159,1400,916,938]" gridrow="13" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFC53FDAFFC4D" box="[159,637,916,938]" gridcol="0" gridrow="13" pageId="36" pageNumber="37">Pectoral-fin length</th>
<td id="B57B0D384F5300074CDEFC53FCF4FC4D" box="[749,806,916,938]" gridcol="1" gridrow="13" pageId="36" pageNumber="37">18.1</td>
<td id="B57B0D384F5300074D49FC53FC61FC4D" box="[890,947,916,938]" gridcol="2" gridrow="13" pageId="36" pageNumber="37">-</td>
<td id="B57B0D384F5300074A34FC53FBF3FC4D" box="[1031,1057,916,938]" gridcol="3" gridrow="13" pageId="36" pageNumber="37">48</td>
<td id="B57B0D384F5300074A6FFC53FB08FC4D" box="[1116,1242,916,938]" gridcol="4" gridrow="13" pageId="36" pageNumber="37">15.6–20.9</td>
<td id="B57B0D384F5300074B0DFC53FAAAFC4D" box="[1342,1400,916,938]" gridcol="5" gridrow="13" pageId="36" pageNumber="37">18.2</td>
</tr>
<tr id="F6AA64444F5300074EACFC7AFAAAFC34" box="[159,1400,957,979]" gridrow="14" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFC7AFDAFFC34" box="[159,637,957,979]" gridcol="0" gridrow="14" pageId="36" pageNumber="37">Pelvic-fin length</th>
<td id="B57B0D384F5300074CDEFC7AFCF4FC34" box="[749,806,957,979]" gridcol="1" gridrow="14" pageId="36" pageNumber="37">14.4</td>
<td id="B57B0D384F5300074D49FC7AFC61FC34" box="[890,947,957,979]" gridcol="2" gridrow="14" pageId="36" pageNumber="37">-</td>
<td id="B57B0D384F5300074A34FC7AFBF3FC34" box="[1031,1057,957,979]" gridcol="3" gridrow="14" pageId="36" pageNumber="37">50</td>
<td id="B57B0D384F5300074A6FFC7AFB08FC34" box="[1116,1242,957,979]" gridcol="4" gridrow="14" pageId="36" pageNumber="37">11.5–17.1</td>
<td id="B57B0D384F5300074B0DFC7AFAAAFC34" box="[1342,1400,957,979]" gridcol="5" gridrow="14" pageId="36" pageNumber="37">14.8</td>
</tr>
<tr id="F6AA64444F5300074EACFC21FAAAFC1B" box="[159,1400,998,1020]" gridrow="15" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFC21FDAFFC1B" box="[159,637,998,1020]" gridcol="0" gridrow="15" pageId="36" pageNumber="37">Caudal peduncle depth</th>
<td id="B57B0D384F5300074CDEFC21FCF4FC1B" box="[749,806,998,1020]" gridcol="1" gridrow="15" pageId="36" pageNumber="37">7.7</td>
<td id="B57B0D384F5300074D49FC21FC61FC1B" box="[890,947,998,1020]" gridcol="2" gridrow="15" pageId="36" pageNumber="37">7.6</td>
<td id="B57B0D384F5300074A34FC21FBF3FC1B" box="[1031,1057,998,1020]" gridcol="3" gridrow="15" pageId="36" pageNumber="37">47</td>
<td id="B57B0D384F5300074A6FFC21FB08FC1B" box="[1116,1242,998,1020]" gridcol="4" gridrow="15" pageId="36" pageNumber="37">7.3–9.3</td>
<td id="B57B0D384F5300074B0DFC21FAAAFC1B" box="[1342,1400,998,1020]" gridcol="5" gridrow="15" pageId="36" pageNumber="37">8.2</td>
</tr>
<tr id="F6AA64444F5300074EACFBD7FAAAFBC1" box="[159,1400,1040,1062]" gridrow="16" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFBD7FDAFFBC1" box="[159,637,1040,1062]" gridcol="0" gridrow="16" pageId="36" pageNumber="37">Head length</th>
<td id="B57B0D384F5300074CDEFBD7FCF4FBC1" box="[749,806,1040,1062]" gridcol="1" gridrow="16" pageId="36" pageNumber="37">30.0</td>
<td id="B57B0D384F5300074D49FBD7FC61FBC1" box="[890,947,1040,1062]" gridcol="2" gridrow="16" pageId="36" pageNumber="37">29.2</td>
<td id="B57B0D384F5300074A34FBD7FBF3FBC1" box="[1031,1057,1040,1062]" gridcol="3" gridrow="16" pageId="36" pageNumber="37">53</td>
<td id="B57B0D384F5300074A6FFBD7FB08FBC1" box="[1116,1242,1040,1062]" gridcol="4" gridrow="16" pageId="36" pageNumber="37">25.4–32.5</td>
<td id="B57B0D384F5300074B0DFBD7FAAAFBC1" box="[1342,1400,1040,1062]" gridcol="5" gridrow="16" pageId="36" pageNumber="37">28.0</td>
</tr>
<tr id="F6AA64444F5300074EACFBFEFAAAFBA8" box="[159,1400,1081,1103]" gridrow="17" pageId="36" pageNumber="37" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<th id="B57B0D384F5300074EACFBFEFDAFFBA8" box="[159,637,1081,1103]" gridcol="0" gridrow="17" pageId="36" pageNumber="37">Percentages of head length</th>
</tr>
<tr id="F6AA64444F5300074EACFBA5FAAAFB9F" box="[159,1400,1122,1144]" gridrow="18" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFBA5FDAFFB9F" box="[159,637,1122,1144]" gridcol="0" gridrow="18" pageId="36" pageNumber="37">Head height</th>
<td id="B57B0D384F5300074CDEFBA5FCF4FB9F" box="[749,806,1122,1144]" gridcol="1" gridrow="18" pageId="36" pageNumber="37">65.4</td>
<td id="B57B0D384F5300074D49FBA5FC61FB9F" box="[890,947,1122,1144]" gridcol="2" gridrow="18" pageId="36" pageNumber="37">63.8</td>
<td id="B57B0D384F5300074A34FBA5FBF3FB9F" box="[1031,1057,1122,1144]" gridcol="3" gridrow="18" pageId="36" pageNumber="37">53</td>
<td id="B57B0D384F5300074A6FFBA5FB08FB9F" box="[1116,1242,1122,1144]" gridcol="4" gridrow="18" pageId="36" pageNumber="37">63.1–75.9</td>
<td id="B57B0D384F5300074B0DFBA5FAAAFB9F" box="[1342,1400,1122,1144]" gridcol="5" gridrow="18" pageId="36" pageNumber="37">69.1</td>
</tr>
<tr id="F6AA64444F5300074EACFB4BFAAAFB45" box="[159,1400,1164,1186]" gridrow="19" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFB4BFDAFFB45" box="[159,637,1164,1186]" gridcol="0" gridrow="19" pageId="36" pageNumber="37">Snout length</th>
<td id="B57B0D384F5300074CDEFB4BFCF4FB45" box="[749,806,1164,1186]" gridcol="1" gridrow="19" pageId="36" pageNumber="37">30.6</td>
<td id="B57B0D384F5300074D49FB4BFC61FB45" box="[890,947,1164,1186]" gridcol="2" gridrow="19" pageId="36" pageNumber="37">-</td>
<td id="B57B0D384F5300074A34FB4BFBF3FB45" box="[1031,1057,1164,1186]" gridcol="3" gridrow="19" pageId="36" pageNumber="37">52</td>
<td id="B57B0D384F5300074A6FFB4BFB08FB45" box="[1116,1242,1164,1186]" gridcol="4" gridrow="19" pageId="36" pageNumber="37">25.6–33.4</td>
<td id="B57B0D384F5300074B0DFB4BFAAAFB45" box="[1342,1400,1164,1186]" gridcol="5" gridrow="19" pageId="36" pageNumber="37">29.8</td>
</tr>
<tr id="F6AA64444F5300074EACFB72FAAAFB2C" box="[159,1400,1205,1227]" gridrow="20" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFB72FDAFFB2C" box="[159,637,1205,1227]" gridcol="0" gridrow="20" pageId="36" pageNumber="37">Upper jaw length</th>
<td id="B57B0D384F5300074CDEFB72FCF4FB2C" box="[749,806,1205,1227]" gridcol="1" gridrow="20" pageId="36" pageNumber="37">50.7</td>
<td id="B57B0D384F5300074D49FB72FC61FB2C" box="[890,947,1205,1227]" gridcol="2" gridrow="20" pageId="36" pageNumber="37">45.8</td>
<td id="B57B0D384F5300074A34FB72FBF3FB2C" box="[1031,1057,1205,1227]" gridcol="3" gridrow="20" pageId="36" pageNumber="37">53</td>
<td id="B57B0D384F5300074A6FFB72FB08FB2C" box="[1116,1242,1205,1227]" gridcol="4" gridrow="20" pageId="36" pageNumber="37">42.9–51.2</td>
<td id="B57B0D384F5300074B0DFB72FAAAFB2C" box="[1342,1400,1205,1227]" gridcol="5" gridrow="20" pageId="36" pageNumber="37">48.5</td>
</tr>
<tr id="F6AA64444F5300074EACFB19FAAAFB13" box="[159,1400,1246,1268]" gridrow="21" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFB19FDAFFB13" box="[159,637,1246,1268]" gridcol="0" gridrow="21" pageId="36" pageNumber="37">Horizontal eye diameter</th>
<td id="B57B0D384F5300074CDEFB19FCF4FB13" box="[749,806,1246,1268]" gridcol="1" gridrow="21" pageId="36" pageNumber="37">22.6</td>
<td id="B57B0D384F5300074D49FB19FC61FB13" box="[890,947,1246,1268]" gridcol="2" gridrow="21" pageId="36" pageNumber="37">17.0</td>
<td id="B57B0D384F5300074A34FB19FBF3FB13" box="[1031,1057,1246,1268]" gridcol="3" gridrow="21" pageId="36" pageNumber="37">53</td>
<td id="B57B0D384F5300074A6FFB19FB08FB13" box="[1116,1242,1246,1268]" gridcol="4" gridrow="21" pageId="36" pageNumber="37">17.0–25.6</td>
<td id="B57B0D384F5300074B0DFB19FAAAFB13" box="[1342,1400,1246,1268]" gridcol="5" gridrow="21" pageId="36" pageNumber="37">21.4</td>
</tr>
<tr id="F6AA64444F5300074EACFACFFAAAFAF9" box="[159,1400,1288,1310]" gridrow="22" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFACFFDAFFAF9" box="[159,637,1288,1310]" gridcol="0" gridrow="22" pageId="36" pageNumber="37">Post-orbital length</th>
<td id="B57B0D384F5300074CDEFACFFCF4FAF9" box="[749,806,1288,1310]" gridcol="1" gridrow="22" pageId="36" pageNumber="37">49.3</td>
<td id="B57B0D384F5300074D49FACFFC61FAF9" box="[890,947,1288,1310]" gridcol="2" gridrow="22" pageId="36" pageNumber="37">48.1</td>
<td id="B57B0D384F5300074A34FACFFBF3FAF9" box="[1031,1057,1288,1310]" gridcol="3" gridrow="22" pageId="36" pageNumber="37">53</td>
<td id="B57B0D384F5300074A6FFACFFB08FAF9" box="[1116,1242,1288,1310]" gridcol="4" gridrow="22" pageId="36" pageNumber="37">46.8–54.8</td>
<td id="B57B0D384F5300074B0DFACFFAAAFAF9" box="[1342,1400,1288,1310]" gridcol="5" gridrow="22" pageId="36" pageNumber="37">50.4</td>
</tr>
<tr id="F6AA64444F5300074EACFAF6FAAAFAA0" box="[159,1400,1329,1351]" gridrow="23" pageId="36" pageNumber="37">
<th id="B57B0D384F5300074EACFAF6FDAFFAA0" box="[159,637,1329,1351]" gridcol="0" gridrow="23" pageId="36" pageNumber="37">Least interorbital width</th>
<td id="B57B0D384F5300074CDEFAF6FCF4FAA0" box="[749,806,1329,1351]" gridcol="1" gridrow="23" pageId="36" pageNumber="37">28.3</td>
<td id="B57B0D384F5300074D49FAF6FC61FAA0" box="[890,947,1329,1351]" gridcol="2" gridrow="23" pageId="36" pageNumber="37">28.1</td>
<td id="B57B0D384F5300074A34FAF6FBF3FAA0" box="[1031,1057,1329,1351]" gridcol="3" gridrow="23" pageId="36" pageNumber="37">53</td>
<td id="B57B0D384F5300074A6FFAF6FB08FAA0" box="[1116,1242,1329,1351]" gridcol="4" gridrow="23" pageId="36" pageNumber="37">26.3–37.7</td>
<td id="B57B0D384F5300074B0DFAF6FAAAFAA0" box="[1342,1400,1329,1351]" gridcol="5" gridrow="23" pageId="36" pageNumber="37">32.1</td>
</tr>
</table>
</paragraph>
<paragraph id="482566064F53FFDD4EF4FA40FC15FF33" blockId="36.[151,1437,1415,2017]" lastBlockId="37.[151,1437,151,2013]" lastPageId="37" lastPageNumber="38" pageId="36" pageNumber="37">
Head profile conspicuously acute anteriorly, mouth terminal. Jaws anisognathous, premaxillary projecting slightly relative to dentary, outer row of premaxillary teeth and in some specimens also second row exposed when mouth is closed. Maxillary long, extending posteriorly to slightly posterior to middle of pupil. Adipose eyelid well developed. Premaxillary teeth in three rows; teeth of third row largest. Five (1), 7 (6), 8 (12), 9 (13), 10 (9), 11 (6) ou 12 (3) tricuspidate teeth in outer series. Three (1), 4 (21), 5 (18), 6 (6), or 7 (1) tri- to pentacuspidate teeth in second, inner premaxillary row, plus 1 (14), 2 (23), or 3 (12) tricuspidate teeth between the first and third rows. Two teeth in third premaxillary row, medial teeth largest, symphyseal teeth smaller, slightly tilted towards each other, both pentacuspidate. Maxillary with distal portion distinctly expanded and rounded in profile. Eighteen to 33 maxillary teeth, slightly smaller than teeth of first premaxillary row, anterior teeth tricuspidate, posterior teeth unicuspidate. Dentary with 9 (1), 10 (1), 11 (4), 12 (7), 13 (5), 14 (10), 15 (10), 16 (6), or 17 (3) teeth in main series. Anterior four dentary teeth assymetrical, considerably larger and bulkier than remaining teeth, tetra- to pentacuspidate, each with central cusp distinctly larger than remaining cusps. Remaining dentary teeth progressivelly smaller, tetra, tri- to unicuspidate. Inner (lingual) series consisting of a small, single unicuspid symphyseal tooth, situated immediately posterior to symphyseal dentary teeth of main series, plus row of 23–37 small, unicuspidate teeth, originating on lingual crest of dentary replacement trench at level of the fourth to sixth teeth of main series. Inner symphyseal teeth absent in 6 out of 19 specimens examined for this character. Larger specimen displaying symphyseal teeth 
<quantity id="8F62CBE34F53FFDC4C65F80FFD19F807" box="[598,715,1992,2016]" metricMagnitude="-1" metricUnit="m" metricValue="2.203" pageId="36" pageNumber="37" unit="mm" value="220.3">220.3 mm</quantity>
SL (MZUSP 2091); specimens larger than 
<quantity id="8F62CBE34F53FFDC4AF0F80FFAF1F807" box="[1219,1315,1992,2016]" metricMagnitude="-1" metricUnit="m" metricValue="2.3" pageId="36" pageNumber="37" unit="mm" value="230.0">230 mm</quantity>
SL (MCZ 21112, MZUSP 
<date id="3C2440C64F52FFDD4F60FF5FFE78FF57" box="[339,426,152,176]" pageId="37" pageNumber="38" value="29-10-30">103029</date>
, MZUSP 88515, MZUSP 19128) lacking symphyseal teeth. A single specimen smaller than 
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SL lacking symphyseal teeth (MNRJ 
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SL).
</paragraph>
<paragraph id="482566064F52FFDD4EF4FF18FD22FE13" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="38">
Scales cycloid. Lateral line complete, from supracleithrum to caudal-fin base. Fifty-one (2), 52 (2), 53 (5), 54 (6), 55 (5), 56 (8), 57 (5), 58 (9), 59 (5), 60 (2), 61 (2), 62 (2), 63 (4) ou 65 (1) scales in lateral line series. Laterosensory tube simple in small (&lt;
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SL) specimens, deflected upwards in the first 5–6 scales, downwards in the remaining lateral-line scales. Larger specimens (&gt; 
<quantity id="8F62CBE34F52FFDD4D98FE8BFBD8FE83" box="[939,1034,332,356]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="37" pageNumber="38" unit="mm" value="200.0">200 mm</quantity>
SL) with branched tubules, mostly bifurcated in middle-sized (
<quantity id="8F62CBE34F52FFDD4FFFFEB7FDB1FE60" box="[460,611,368,392]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" metricValueMax="2.7" metricValueMin="2.3" pageId="37" pageNumber="38" unit="mm" value="250.0" valueMax="270.0" valueMin="230.0">230–270 mm</quantity>
SL) specimens but some scales with 3–5 tubules in the larger (&gt; 
<quantity id="8F62CBE34F52FFDD4B0CFEB7FA4FFE60" box="[1343,1437,368,392]" metricMagnitude="-1" metricUnit="m" metricValue="2.9" pageId="37" pageNumber="38" unit="mm" value="290.0">290 mm</quantity>
SL) specimens examined. Horizontal scale rows between dorsal-fin origin and lateral line 9 (9), 10 (25), or 11 (22). Horizontal scale rows between lateral line and pelvic-fin 4 (7), 5 (27), 6 (17), or 7 (3). Circumpeduncular scales 16 (3), 17 (2), 18 (9), 19 (10), 20 (12), 21 (12), or 22 (3).
</paragraph>
<paragraph id="482566064F52FFDD4EF4FE38FDDFFC67" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="38">
Dorsal-fin rays ii, 9. Dorsal fin origin slightly ahead of middle of SL. First dorsal-fin pterygiophore inserting behind neural spine of 13th vertebra. Anal-fin rays iii (not including first, small unbranched ray only visible in the cs specimen), 21 (2), 22 (2), 23 (10), 24 (9), 25 (12), 26 (17), or 27 (5). First anal-fin pterygiophore inserting behind haemal spine of 24th vertebra. Last unbranched and anterior 3–4 branched anal-fin rays longer, remaining rays progressively shorter towards anal-fin end. Anal fin displaying numerous (c. 18–22 per fin-ray main branch) middle-sized hooks on last unbranched and posterior main branch of branched rays 10–20, associated with dense, gelatinous tissue in six specimens (MZUSP 2091, 209.0– 
<quantity id="8F62CBE34F52FFDD4D14FD1FFC49FD08" box="[807,923,728,752]" metricMagnitude="-1" metricUnit="m" metricValue="2.203" pageId="37" pageNumber="38" unit="mm" value="220.3">220.3 mm</quantity>
SL; MZUSP 
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SL; MZUSP 88515, 235.0–247.0 mm SL; MZUSP 107280, 260.0 mm SL). 
<collectingRegion id="8A5EA8E44F52FFDD4D5DFD3BFC50FCF3" box="[878,898,764,788]" country="Nigeria" name="Rivers" pageId="37" pageNumber="38">A</collectingRegion>
single hook per ray segment. Sheath of scales covering basis of anal-fin rays composed of two scale rows, lower scale row formed by 25–30 rectangular scales. Pectoral-fin rays i, 11 (1), 12 (10), 13 (27), 14 (15), or 15 (1). Pelvic-fin rays i,7. Main caudal-fin rays 10/9. Caudal fin forked, lobes slightly pointed.
</paragraph>
<paragraph id="482566064F52FFDD4EF4FC4AFD24FC0B" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="38">Four branchiostegal rays, three on anterior ceratohyal and one on posterior ceratohyal. First branchial arch with 11 (2), 12 (6), 13 (10), 14 (12), 15 (3), or 16 (1) lower, 1 at angle, and 11 (4), 12 (14), 13 (12), 14 (3), or 17 (1) upper gill rakers. Vertebrae 45 (2). Supraneurals 9 (1).</paragraph>
<paragraph id="482566064F52FFDD4EF4FC30FB8DF992" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="38">
<emphasis id="7AEEBA144F52FFDD4EF4FC30FE1FFBF7" bold="true" box="[199,461,1015,1040]" pageId="37" pageNumber="38">Coloration in alcohol.</emphasis>
Description based on wild-caught, recently collected specimens (MZUSP 88515 and MZUSP 
<date id="3C2440C64F52FFDD4EC8FBDBFE86FBD2" box="[251,340,1052,1077]" pageId="37" pageNumber="38" value="45-10-30">103045</date>
); the majority of remaining examined specimens faded, either due to a long period of conservation (&gt; 60 years) or for being originally captive, stocked specimens raised in clear-water, sun-lit ponds. Top of head, snout, supraorbital, sixth infraorbital, upper portion of fifth infraorbital, and dorsal portion of body dark-brown. Second, third, and fourth infraorbitals light brown, with a silvery hue. Opercle mainly silvery, with some lightbrown tinge. Dentary, maxillary, gular area and lower portion of body clear, with a cream tinge. Lateral portion of body light-grey, silvery in specimens retaining a considerable amount of guanine (e.g., MCZ 21112, MZUSP 62746). Scales above lateral line on lateral surfaces of body with dark pigmentation concentrated on scales margins, forming a tenuous reticulated pattern in specimens MZUSP 88515 and MZUSP 
<date id="3C2440C64F52FFDD4AB9FAD0FB33FAD7" box="[1162,1249,1303,1328]" pageId="37" pageNumber="38" value="45-10-30">103045</date>
. Humeral blotch present, moderately conspicuous, approximately rounded in shape, situated immediately above lateral line, its anterior margin at level of second to third, extending longitudinally to posterior margin of fourth to fifth lateral line scales, and vertically less than one scale high. Lateral surfaces of body above lateral line, and posterior to dorsalfin, presenting longitudinal stripes formed by dark pigmentation on central portion of scales in specimens MZUSP 88515 and MZUSP 
<date id="3C2440C64F52FFDD4F4AFA0BFE02FA02" box="[377,464,1484,1509]" pageId="37" pageNumber="38" value="45-10-30">103045</date>
. Large, moderately conspicuous, oval-shaped caudal peduncle blotch, extending along 8–11 last lateral-line scales. All rayed fins with a considerable amount of dark pigmentation at the interradial membranes (discernible even in some severely faded specimens, e.g., MZUSP 2091), imparting a overall dark coloration to these fins. Caudal-fin with a poorly discernible, roughly V-shaped blotch, formed by dark pigmentantion situated on outer caudal-fin rays. Adipose fin light-grey to light-brown.
</paragraph>
<paragraph id="482566064F52FFDD4EF4F9B8FB8EF8CF" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="38">
<emphasis id="7AEEBA144F52FFDD4EF4F9B8FEB3F97F" bold="true" box="[199,353,1663,1688]" pageId="37" pageNumber="38">Color in life.</emphasis>
Description based on a specimen photographed alive at the CESP-Paraibuna fish hatchery and from photographs of specimens MZUSP 
<date id="3C2440C64F52FFDD4C52F963FD6AF95A" box="[609,696,1700,1725]" pageId="37" pageNumber="38" value="45-10-30">103045</date>
, immediately before fixation. Top of head and snout dark. Dentary and maxillary clear. Dorsal portion of body plumbeous. Opercle, infraorbital bones, and lateral portion of body silvery. Humeral blotch inconspicuous; caudal peduncle blotch moderately conspicuous. All fins, except adipose fin, darkened. Caudal fin with outer fin-rays dark, forming a roughly V-shaped blotch.
</paragraph>
<paragraph id="482566064F52FFDD4EF4F8F3FDB5F83B" blockId="37.[151,1437,151,2013]" pageId="37" pageNumber="38">
<emphasis id="7AEEBA144F52FFDD4EF4F8F3FE68F8AA" bold="true" box="[199,442,1844,1869]" pageId="37" pageNumber="38">Sexual dimorphism.</emphasis>
Five specimens (MZUSP 2091, 209.0– 
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SL; MZUSP 
<quantity id="8F62CBE34F52FFDD4AADF8F3FAB8F8AB" box="[1182,1386,1844,1868]" metricMagnitude="4" metricUnit="m" metricValue="7.90921872" pageId="37" pageNumber="38" unit="mm" value="7.90921872E7">79092, 187.2 mm</quantity>
SL; MZUSP 88515, 235.0–247.0 mm SL; MZUSP 107280, 260.0 mm SL) possess anal-fin hooks. Hooks are, however, absent from pelvic- and remaining fins. One specimen (MZUSP 2091) proved to be a male after dissection. At the CESP-Paraibuna fish hatchery, mature males were in fact distinguished from females due to the presence of analfin hooks (L. Girardi, pers. comm, 1999).
</paragraph>
<caption id="1CE5368E4F51FFDE4EA4FD95FACCFD8F" box="[151,1310,594,616]" httpUri="https://zenodo.org/record/257791/files/figure.png" pageId="38" pageNumber="39" targetBox="[151,1436,193,573]" targetPageId="38">
<paragraph id="482566064F51FFDE4EA4FD95FACCFD8F" blockId="38.[151,1310,594,616]" box="[151,1310,594,616]" pageId="38" pageNumber="39">
<emphasis id="7AEEBA144F51FFDE4EA4FD95FEF2FD8F" bold="true" box="[151,288,594,616]" pageId="38" pageNumber="39">FIGURE 22.</emphasis>
<taxonomicName id="8F9A1D854F51FFDE4F16FD94FE16FD8F" box="[293,452,595,616]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F51FFDE4F16FD94FE16FD8F" box="[293,452,595,616]" italics="true" pageId="38" pageNumber="39">Brycon insignis</emphasis>
</taxonomicName>
, lectotype, NMW 62925:2, 144.8 mm SL: Brazil, Rio de Janeiro, rio Paraíba do Sul.
</paragraph>
</caption>
<caption id="1CE5368E4F51FFDE4EA4FBE9FB13FBA3" box="[151,1217,1070,1092]" httpUri="https://zenodo.org/record/257792/files/figure.png" pageId="38" pageNumber="39" targetBox="[151,1436,666,1049]" targetPageId="38">
<paragraph id="482566064F51FFDE4EA4FBE9FB13FBA3" blockId="38.[151,1217,1070,1092]" box="[151,1217,1070,1092]" pageId="38" pageNumber="39">
<emphasis id="7AEEBA144F51FFDE4EA4FBE9FEF2FBA3" bold="true" box="[151,288,1070,1092]" pageId="38" pageNumber="39">FIGURE 23.</emphasis>
<taxonomicName id="8F9A1D854F51FFDE4F16FBE8FE16FBA3" box="[293,452,1071,1092]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F51FFDE4F16FBE8FE16FBA3" box="[293,452,1071,1092]" italics="true" pageId="38" pageNumber="39">Brycon insignis</emphasis>
</taxonomicName>
, MZUSP 103045, 259.0 mm SL: Brazil, Rio de Janeiro, rio Paraíba do Sul.
</paragraph>
</caption>
<caption id="1CE5368E4F51FFDE4EA4F989FB5BF983" box="[151,1161,1614,1636]" httpUri="https://zenodo.org/record/257793/files/figure.png" pageId="38" pageNumber="39" targetBox="[151,1436,1142,1593]" targetPageId="38">
<paragraph id="482566064F51FFDE4EA4F989FB5BF983" blockId="38.[151,1161,1614,1636]" box="[151,1161,1614,1636]" pageId="38" pageNumber="39">
<emphasis id="7AEEBA144F51FFDE4EA4F989FEF2F983" bold="true" box="[151,288,1614,1636]" pageId="38" pageNumber="39">FIGURE 24.</emphasis>
<taxonomicName id="8F9A1D854F51FFDE4F16F988FE16F983" box="[293,452,1615,1636]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F51FFDE4F16F988FE16F983" box="[293,452,1615,1636]" italics="true" pageId="38" pageNumber="39">Brycon insignis</emphasis>
</taxonomicName>
, MZUSP 62746, 310.4 mm SL: Brazil, São Paulo, rio Paraíba do Sul.
</paragraph>
</caption>
<paragraph id="482566064F51FFDE4EF4F94CFCCBF92F" blockId="38.[151,1436,1675,2024]" pageId="38" pageNumber="39">
<emphasis id="7AEEBA144F51FFDE4EF4F94CFE48F943" bold="true" box="[199,410,1675,1700]" pageId="38" pageNumber="39">Common names.</emphasis>
“ 
<taxonomicName id="8F9A1D854F51FFDE4F86F94CFDF1F943" box="[437,547,1675,1700]" family="Bryconidae" genus="Megalobrycon" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="38" pageNumber="184" phylum="Chordata" rank="species" species="piabanha">Piabanha</taxonomicName>
”; “piabanha-vermelha” (
<bibRefCitation id="2C0B1BF74F51FFDE4D78F94CFB72F943" author="Miranda-Ribeiro" box="[843,1184,1675,1700]" pageId="38" pageNumber="39" refString="Miranda-Ribeiro, A. (1902) Oito especies de peixes do Rio Pomba. A Lavoura, Boletim da Sociedade de Agricultura, 1902, 1 - 8." type="journal article" year="1902">Miranda-Ribeiro, 1902: 254</bibRefCitation>
); “biririca” (juvenile specimens, rio Itabapoana, V. L.M. Santos, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F51FFDF4EF4F914FF32FEA7" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39" type="distribution">
<paragraph id="482566064F51FFDE4EF4F914FF3EF847" blockId="38.[151,1436,1675,2024]" pageId="38" pageNumber="39">
<emphasis id="7AEEBA144F51FFDE4EF4F914FE8BF90B" bold="true" box="[199,345,1747,1772]" pageId="38" pageNumber="39">Distribution</emphasis>
. Known from the rio 
<collectingRegion id="8A5EA8E44F51FFDE4C7CF914FD71F90B" box="[591,675,1747,1772]" country="Brazil" name="Paraiba" pageId="38" pageNumber="39">Paraíba</collectingRegion>
do 
<collectingRegion id="8A5EA8E44F51FFDE4CFDF914FD21F90B" box="[718,755,1747,1772]" country="Cameroon" name="South" pageId="38" pageNumber="39">Sul</collectingRegion>
basin at 
<collectingRegion id="8A5EA8E44F51FFDE4D6AF913FC1FF90B" box="[857,973,1747,1772]" country="Brazil" name="Sao Paulo" pageId="38" pageNumber="39">São Paulo</collectingRegion>
, 
<collectingRegion id="8A5EA8E44F51FFDE4DEBF914FBACF90B" box="[984,1150,1747,1772]" country="Brazil" name="Rio de Janeiro" pageId="38" pageNumber="39">
<collectingRegion id="8A5EA8E44F51FFDE4DEBF914FBD3F90B" box="[984,1025,1747,1772]" country="Nigeria" name="Rivers" pageId="38" pageNumber="39">Rio</collectingRegion>
de Janeiro
</collectingRegion>
, and 
<collectingRegion id="8A5EA8E44F51FFDE4A8AF914FA9DF90B" box="[1209,1359,1747,1772]" country="Brazil" name="Minas Gerais" pageId="38" pageNumber="39">Minas Gerais</collectingRegion>
states, plus several smaller coastal river drainages at 
<collectingRegion id="8A5EA8E44F51FFDE4C9BF930FC9CF8F7" box="[680,846,1783,1808]" country="Brazil" name="Rio de Janeiro" pageId="38" pageNumber="39">
<collectingRegion id="8A5EA8E44F51FFDE4C9BF930FD02F8F7" box="[680,720,1783,1808]" country="Nigeria" name="Rivers" pageId="38" pageNumber="39">Rio</collectingRegion>
de Janeiro
</collectingRegion>
state, i.e., rio Guandu, rio Macaé, rio São João, rio Imbé and Lagoa Feia system, and rio Itabapoana, at the border of 
<collectingRegion id="8A5EA8E44F51FFDE4DBFF8DCFBE3F8D3" box="[908,1073,1819,1844]" country="Brazil" name="Rio de Janeiro" pageId="38" pageNumber="39">
<collectingRegion id="8A5EA8E44F51FFDE4DBFF8DCFC66F8D3" box="[908,948,1819,1844]" country="Nigeria" name="Rivers" pageId="38" pageNumber="39">Rio</collectingRegion>
de Janeiro
</collectingRegion>
and Espírito Santos states (
<figureCitation id="D0A17A834F51FFDE4B5CF8DCFF65F8BF" captionStart="FIGURE 25" captionStartId="39.[151,250,1679,1701]" captionTargetBox="[170,1417,392,1639]" captionTargetId="figure@39.[151,1436,373,1658]" captionTargetPageId="39" captionText="FIGURE 25. Map of eastern Brazil, showing known localities of Brycon insignis (red dots), B. vermelha (grey dots), and B. howesi (black dots; dark star indicates type-locality)." httpUri="https://zenodo.org/record/257794/files/figure.png" pageId="38" pageNumber="39">Fig. 25</figureCitation>
). Specimens from the rio Macaé were not examined in the present study, but the species also occurs at this small river system according to 
<bibRefCitation id="2C0B1BF74F51FFDE4F8BF8A4FD1BF89B" author="Bizerril" box="[440,713,1891,1916]" pageId="38" pageNumber="39" refString="Bizerril, C. R. S. F. &amp; Primo, P. B. S. (2001) Peixes de aguas interiores do estado do Rio de Janeiro. Fundacao de Estudos do Mar / Secretaria de Estado de Meio Ambiente e Desenvolvimento Sustentavel, Rio de Janeiro, 417 pp." type="book" year="2001">Bizerril &amp; Primo (2001)</bibRefCitation>
. Extirpated from much of its original range (see “Conservation”, below).
</paragraph>
<paragraph id="482566064F51FFDF4EF4F86CFF32FEA7" blockId="38.[151,1436,1675,2024]" lastBlockId="39.[151,1436,151,320]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">
<emphasis id="7AEEBA144F51FFDE4EF4F86CFE44F823" bold="true" box="[199,406,1963,1988]" pageId="38" pageNumber="39">Ecological notes.</emphasis>
Stomach contents of a dissected specimen (MZUSP 2091) contained a whole, nonmasticated fruit and flowers remains. 
<bibRefCitation id="2C0B1BF74F51FFDE4C08F808FBCEF80F" author="Bizerril" box="[571,1052,1999,2024]" pageId="38" pageNumber="39" refString="Bizerril, C. R. S. F. &amp; Primo, P. B. S. (2001) Peixes de aguas interiores do estado do Rio de Janeiro. Fundacao de Estudos do Mar / Secretaria de Estado de Meio Ambiente e Desenvolvimento Sustentavel, Rio de Janeiro, 417 pp." type="book" year="2001">
Bizerril &amp; Primo (2001: 55, as 
<taxonomicName id="8F9A1D854F51FFDE4DA7F817FBC1F80F" box="[916,1043,1999,2024]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="38" pageNumber="39" phylum="Chordata" rank="species" species="opalinus">
<emphasis id="7AEEBA144F51FFDE4DA7F817FBC1F80F" box="[916,1043,1999,2024]" italics="true" pageId="38" pageNumber="39">B. opalinus</emphasis>
</taxonomicName>
)
</bibRefCitation>
reported insects from stomachs of specimens collected at the rio 
<collectingRegion id="8A5EA8E44F50FFDF4FC2FF50FD97FF57" box="[497,581,151,176]" country="Brazil" name="Paraiba" pageId="39" pageNumber="40">Paraíba</collectingRegion>
do 
<collectingRegion id="8A5EA8E44F50FFDF4C40FF50FD4EFF57" box="[627,668,151,176]" country="Cameroon" name="South" pageId="39" pageNumber="40">Sul</collectingRegion>
. 
<bibRefCitation id="2C0B1BF74F50FFDF4C9BFF50FC7DFF57" author="Magalhaes" box="[680,943,151,176]" pageId="39" pageNumber="40" refString="Magalhaes, A. C. (1931) Monographia brazileira de peixes fluviaes. Graphicars, Sao Paulo, 260 pp." type="book" year="1931">Magalhães (1931: 156)</bibRefCitation>
reported smaller fishes as the main dietary item of 
<taxonomicName id="8F9A1D854F50FFDF4EDCFF7AFEB4FF33" box="[239,358,189,212]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F50FFDF4EDCFF7AFEB4FF33" box="[239,358,189,212]" italics="true" pageId="39" pageNumber="40">B. insignis</emphasis>
</taxonomicName>
. Males achieve maturity at the second year of age, with about 
<quantity id="8F62CBE34F50FFDF4A18FF7BFBA3FF33" box="[1067,1137,188,212]" metricMagnitude="-1" metricUnit="m" metricValue="2.0" pageId="39" pageNumber="40" unit="cm" value="20.0">20 cm</quantity>
TL, while females mature at the third year of age, at the size of 
<quantity id="8F62CBE34F50FFDF4C05FF18FDA9FF10" box="[566,635,223,248]" metricMagnitude="-1" metricUnit="m" metricValue="2.5" pageId="39" pageNumber="40" unit="cm" value="25.0">25 cm</quantity>
TL (
<bibRefCitation id="2C0B1BF74F50FFDF4C80FF18FC45FF1F" author="Salgado" box="[691,919,223,248]" pageId="39" pageNumber="40" refString="Salgado, F. G. A., Chain, M. G., Girardi, L. &amp; Faria, C. A. (1997) O salvamento de uma especie em extincao: a conservacao da piabanha (Brycon insignis) na bacia do rio Paraiba do Sul. Relatorio Tecnico da CESP-Companhia Energetica do Estado de Sao Paulo, 28 pp." type="book" year="1997">
Salgado 
<emphasis id="7AEEBA144F50FFDF4D26FF26FC9CFF1F" box="[789,846,223,248]" italics="true" pageId="39" pageNumber="40">et al.</emphasis>
, 1997
</bibRefCitation>
). Microsatellite anaylsis of populations across the rio 
<collectingRegion id="8A5EA8E44F50FFDF4ED5FEC3FEE8FEFA" box="[230,314,260,285]" country="Brazil" name="Paraiba" pageId="39" pageNumber="40">Paraíba</collectingRegion>
do 
<collectingRegion id="8A5EA8E44F50FFDF4F56FEC3FE58FEFA" box="[357,394,260,285]" country="Cameroon" name="South" pageId="39" pageNumber="40">Sul</collectingRegion>
basin showed significant differentiation among distinct populations (Matsumoto &amp; Hilsdorf, 2008).
</paragraph>
</subSubSection>
<caption id="1CE5368E4F50FFDF4EA4F948FD7DF923" httpUri="https://zenodo.org/record/257794/files/figure.png" pageId="39" pageNumber="40" targetBox="[170,1417,392,1639]" targetPageId="39">
<paragraph id="482566064F50FFDF4EA4F948FD7DF923" blockId="39.[151,1436,1679,1732]" pageId="39" pageNumber="40">
<emphasis id="7AEEBA144F50FFDF4EA4F948FEF3F942" bold="true" box="[151,289,1679,1701]" pageId="39" pageNumber="40">FIGURE 25.</emphasis>
Map of eastern Brazil, showing known localities of 
<taxonomicName id="8F9A1D854F50FFDF4D77F956FC31F941" box="[836,995,1681,1702]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F50FFDF4D77F956FC31F941" box="[836,995,1681,1702]" italics="true" pageId="39" pageNumber="40">Brycon insignis</emphasis>
</taxonomicName>
(red dots), 
<taxonomicName id="8F9A1D854F50FFDF4A68F956FB04F942" box="[1115,1238,1679,1702]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="vermelha">
<emphasis id="7AEEBA144F50FFDF4A68F956FB04F942" box="[1115,1238,1679,1702]" italics="true" pageId="39" pageNumber="40">B. vermelha</emphasis>
</taxonomicName>
(grey dots), and 
<taxonomicName id="8F9A1D854F50FFDF4BB5F956FF0FF923" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F50FFDF4BB5F956FF0FF923" italics="true" pageId="39" pageNumber="40">B. howesi</emphasis>
</taxonomicName>
(black dots; dark star indicates type-locality).
</paragraph>
</caption>
<subSubSection id="0080358D4F50FFD04EF4F92CFA90FCBB" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40" type="description">
<paragraph id="482566064F50FFD04EF4F92CFA90FCBB" blockId="39.[151,1436,1771,2012]" lastBlockId="40.[151,1437,151,2013]" lastPageId="40" lastPageNumber="41" pageId="39" pageNumber="40">
<emphasis id="7AEEBA144F50FFDF4EF4F92CFEBDF8E3" bold="true" box="[199,367,1771,1796]" pageId="39" pageNumber="40">Conservation.</emphasis>
<taxonomicName id="8F9A1D854F50FFDF4F46F92AFDF6F8E3" box="[373,548,1773,1796]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F50FFDF4F46F92AFDF6F8E3" box="[373,548,1773,1796]" italics="true" pageId="39" pageNumber="40">Brycon insignis</emphasis>
</taxonomicName>
was reported as being a common fish in the rio Paraíba do Sul basin in the past (
<bibRefCitation id="2C0B1BF74F50FFDF4EADF8C8FEB1F8CF" author="Magalhaes" box="[158,355,1807,1832]" pageId="39" pageNumber="40" refString="Magalhaes, A. C. (1931) Monographia brazileira de peixes fluviaes. Graphicars, Sao Paulo, 260 pp." type="book" year="1931">Magalhães, 1931</bibRefCitation>
). 
<bibRefCitation id="2C0B1BF74F50FFDF4F44F8C8FD48F8CF" author="Machado" box="[375,666,1807,1832]" pageId="39" pageNumber="40" refString="Machado, C. E. M. &amp; Abreu, H. C. F. (1952) Notas preliminares sobre a caca e a pesca no estado de Sao Paulo-I. A pesca no vale do Paraiba. Boletim de Industria Animal, 13, 145 - 160." type="journal article" year="1952">Machado &amp; Abreu (1952)</bibRefCitation>
recorded that 
<taxonomicName id="8F9A1D854F50FFDF4D08F8D6FC38F8CF" box="[827,1002,1809,1832]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F50FFDF4D08F8D6FC38F8CF" box="[827,1002,1809,1832]" italics="true" pageId="39" pageNumber="40">Brycon insignis</emphasis>
</taxonomicName>
was the second more important fish in the commercial fisheries in the rio Paraíba do Sul basin, with annual landings of 
<quantity id="8F62CBE34F50FFDF4A0EF8F4FB04F8AC" box="[1085,1238,1843,1868]" metricMagnitude="4" metricUnit="kg" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="39" pageNumber="40" unit="t" value="18.5" valueMax="22.0" valueMin="15.0">15–22 tonnes</quantity>
in the São Paulo portion of the basin alone. The steep decline of the populations of the species, already foreseeing by 
<bibRefCitation id="2C0B1BF74F50FFDF4B10F890FF36F873" author="Magalhaes" pageId="39" pageNumber="40" refString="Magalhaes, A. C. (1931) Monographia brazileira de peixes fluviaes. Graphicars, Sao Paulo, 260 pp." type="book" year="1931">Magalhães (1931)</bibRefCitation>
, was due to combination of several factors, the main being water pollution (both sewage and industrial pollution) and deforestation of the riparian forest. The introduction of the dourado, 
<taxonomicName id="8F9A1D854F50FFDF4A74F858FAE5F85F" box="[1095,1335,1951,1976]" class="Actinopterygii" family="Bryconidae" genus="Salminus" kingdom="Animalia" order="Characiformes" pageId="39" pageNumber="40" phylum="Chordata" rank="species" species="brasiliensis">
<emphasis id="7AEEBA144F50FFDF4A74F858FAE5F85F" box="[1095,1335,1951,1976]" italics="true" pageId="39" pageNumber="40">Salminus brasiliensis</emphasis>
</taxonomicName>
, into the rio Paraíba do Sul basin in the mid-1950’s decade is also mentioned by older fishermen as an important cause of decline of the species, probably due to competition and/or predation. The last record of 
<taxonomicName id="8F9A1D854F5FFFD04AB8FF5EFAE9FF57" box="[1163,1339,153,176]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5FFFD04AB8FF5EFAE9FF57" box="[1163,1339,153,176]" italics="true" pageId="40" pageNumber="41">Brycon insignis</emphasis>
</taxonomicName>
into the portion of the rio Paraíba do Sul basin in São Paulo state was during the late 1980’s decade (
<bibRefCitation id="2C0B1BF74F5FFFD04A9AFF7BFA42FF33" author="Salgado" box="[1193,1424,188,213]" pageId="40" pageNumber="41" refString="Salgado, F. G. A., Chain, M. G., Girardi, L. &amp; Faria, C. A. (1997) O salvamento de uma especie em extincao: a conservacao da piabanha (Brycon insignis) na bacia do rio Paraiba do Sul. Relatorio Tecnico da CESP-Companhia Energetica do Estado de Sao Paulo, 28 pp." type="book" year="1997">
Salgado 
<emphasis id="7AEEBA144F5FFFD04B3FFF7AFA97FF32" box="[1292,1349,188,213]" italics="true" pageId="40" pageNumber="41">et al.</emphasis>
, 1997
</bibRefCitation>
). The species is still reported as occurring in the middle/lower stretches of the rio Paraíba do Sul basin at the Rio de Janeiro state (e.g., 
<bibRefCitation id="2C0B1BF74F5FFFD04F5BFEC3FDBDFEFB" author="Bizerril" box="[360,623,260,285]" pageId="40" pageNumber="41" refString="Bizerril, C. R. S. F. &amp; Primo, P. B. S. (2001) Peixes de aguas interiores do estado do Rio de Janeiro. Fundacao de Estudos do Mar / Secretaria de Estado de Meio Ambiente e Desenvolvimento Sustentavel, Rio de Janeiro, 417 pp." type="book" year="2001">Bizerril &amp; Primo, 2001</bibRefCitation>
; G. Sousa 
<emphasis id="7AEEBA144F5FFFD04CD0FEC2FCDAFEFB" box="[739,776,261,284]" italics="true" pageId="40" pageNumber="41">per</emphasis>
A.W. Hilsdorf, pers. comm.), but some recent catastrophic fish killings due to the leakage of pesticides and other pollutants have occurred in the basin, the latter in 
<date id="3C2440C64F5FFFD04B15FEE0FF01FE83" pageId="40" pageNumber="41" value="2008-11">November 2008</date>
. 
<taxonomicName id="8F9A1D854F5FFFD04EEDFE8AFE5CFE83" box="[222,398,333,356]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5FFFD04EEDFE8AFE5CFE83" box="[222,398,333,356]" italics="true" pageId="40" pageNumber="41">Brycon insignis</emphasis>
</taxonomicName>
was apparently extirpated from two small coastal river systems where it once occurred, the rio Guandu and the rio Macaé. The record for the rio Guandu is based on a single specimen collected at the former Imperial farm at Santa Cruz, during the Thayer Expedition, in 1865 (MCZ 21112). This specimen was labelled as being collected at the rio Grande, which Higuchi (1992) identified as being a small river system draining into the Lago da Tijuca, currently within the urban area of Rio de Janeiro city (
<bibRefCitation id="2C0B1BF74F5FFFD04D96FE1BFACCFE12" author="Bizerril" box="[933,1310,476,501]" pageId="40" pageNumber="41" refString="Bizerril, C. R. S. F. &amp; Primo, P. B. S. (2001) Peixes de aguas interiores do estado do Rio de Janeiro. Fundacao de Estudos do Mar / Secretaria de Estado de Meio Ambiente e Desenvolvimento Sustentavel, Rio de Janeiro, 417 pp." type="book" year="2001">Bizerril &amp; Primo, 2001: 135–136</bibRefCitation>
). However, since the latter river lies very far from the Santa Cruz Imperial farm (more than 
<quantity id="8F62CBE34F5FFFD04A3EFDC7FB81FDFF" box="[1037,1107,511,536]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="40" pageNumber="41" unit="km" value="30.0">30 km</quantity>
in a straight line), we believe that the specimen was actually collected in the rio Guandu, a larger river system much closer to the imperial farm and a known locality for other Thayer Expedition specimes (as “Rio Quenda” or “Quendu”). The rio Guandu has been severely altered during the XX century and a continual occurrence of the species in this river system is highly unlikely. As for the rio Macaé, recent collecting efforts in the basin failed in locating the species (M.G. Brito, pers. comm.). 
<taxonomicName id="8F9A1D854F5FFFD04ECDFD72FE7DFD2B" box="[254,431,693,716]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5FFFD04ECDFD72FE7DFD2B" box="[254,431,693,716]" italics="true" pageId="40" pageNumber="41">Brycon insignis</emphasis>
</taxonomicName>
is officially considered as a threatened species in Brazil, under the category “critically endangered” (Hilsdorf, Lima &amp; Matsumoto, 2008). Stocked populations of the species are kept in two fish hatcheries stations at São Paulo and Rio de Janeiro states (CESP-Paraibuna, Paraibuna, and Projeto 
<taxonomicName id="8F9A1D854F5FFFD04B1CFD3BFA4AFCF2" box="[1327,1432,764,789]" family="Bryconidae" genus="Megalobrycon" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="184" phylum="Chordata" rank="species" species="piabanha">Piabanha</taxonomicName>
, Itaocara, respectively). There is still a reasonable genetic variability as shown by microsatellite loci in wild 
<taxonomicName id="8F9A1D854F5FFFD04B78FCE6FF3CFCBB" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5FFFD04B78FCE6FF3CFCBB" italics="true" pageId="40" pageNumber="41">Brycon insignis</emphasis>
</taxonomicName>
, but considerably little variation in the captive stock of the species (Matsumoto &amp; 
<bibRefCitation id="2C0B1BF74F5FFFD04ABDFC83FAE4FCBB" author="Hilsdorf" box="[1166,1334,836,861]" pageId="40" pageNumber="41" refString="Oyakawa, O. T., Menezes, N. A., Shibatta, O. A., Lima, F. C. T., Langeani, F., Pavanelli, C. S., Nielsen, D. T. B. &amp; Hilsdorf, A. W. S. (2009) Peixes de agua doce. In: Bressan, P. M., Kierulff, M. C. M. &amp; Sugieda, A. M. (Orgs.), Fauna ameacada de extincao no estado de Sao Paulo - vertebrados. Fundacao Parque Zoologico de Sao Paulo, Secretaria do Meio Ambiente, Sao Paulo, pp. 349 - 424" type="book chapter" year="2009">Hilsdorf, 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F5FFFD04EF4FCA0FBE0F85E" pageId="40" pageNumber="41" type="discussion">
<paragraph id="482566064F5FFFD04EF4FCA0FBE0F85E" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
<emphasis id="7AEEBA144F5FFFD04EF4FCA0FEE9FC67" bold="true" box="[199,315,871,896]" pageId="40" pageNumber="41">Remarks.</emphasis>
<bibRefCitation id="2C0B1BF74F5FFFD04F7BFCA0FDE3FC67" author="Steindachner" box="[328,561,871,896]" pageId="40" pageNumber="41" refString="Steindachner, F. (1877) Die Susswasserfische des sudostlichen Brasilien (III). Sitzungsberichte der Kaiserlichen Akademie der Wissenchaften in Wien, 74 (1), 559 - 694, pls. 1 - 13." type="journal article" year="1877">Steindachner (1877)</bibRefCitation>
originally described 
<taxonomicName id="8F9A1D854F5FFFD04D01FCAEFC35FC67" box="[818,999,873,896]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5FFFD04D01FCAEFC35FC67" box="[818,999,873,896]" italics="true" pageId="40" pageNumber="41">Brycon insignis</emphasis>
</taxonomicName>
based on an unspecified number of syntypes from the “Rio Parahyba bei Campos und Mendez, Rio Jequitinhonha”. There are some discussion in the literature whether syntypes of Steindachner’s species described based on material collected by the Thayer Expedition should be considered as only the specimens present at the NMW collection or should also encompass material deposited at MCZ, which were examined by Steindachner during his stay at Harvard in several periods between 
<date id="3C2440C64F5FFFD04ECFFBDBFEA5FBD3" box="[252,375,1052,1077]" pageId="40" pageNumber="41" value="1870-04">April 1870</date>
until his final departure from the United States in 
<date id="3C2440C64F5FFFD04D80FBDBFBB4FBD3" box="[947,1126,1052,1077]" pageId="40" pageNumber="41" value="1873-12">December 1873</date>
(
<bibRefCitation id="2C0B1BF74F5FFFD04A46FBDBFA8DFBD2" author="Steindachner" box="[1141,1375,1052,1077]" pageId="40" pageNumber="41" refString="Steindachner, F. (1875 a) Die Susswasserfische des sudostlichen Brasilien. Sitzungsberichte der Kaiserlichen Akademie der Wissenchaften in Wien, 70 (1), 499 - 538, pls. 1 - 6." type="journal article" year="1875" yearSuffix="a">Steindachner, 1875a</bibRefCitation>
: 499; 
<bibRefCitation id="2C0B1BF74F5FFFD04EA4FBF8FE49FBBF" author="Herzig-Straschil" box="[151,411,1087,1112]" pageId="40" pageNumber="41" refString="Herzig-Straschil, B. (1997) Franz Steindachner (1834 - 1919) and other prime contributors to the ichthyological collection of the Naturhistorisches Museum Wien. In: Pietsch, T. W. &amp; Anderson, Jr., W. D. (Eds.), Collection building in ichthyology and herpetology. American Society of Ichthyologists and Herpetologists, Special Publication Number 3, pp. 101 - 108." type="book chapter" year="1997">Herzig-Straschil, 1997</bibRefCitation>
). 
<bibRefCitation id="2C0B1BF74F5FFFD04F80FBF8FD4AFBBF" author="Schaefer" box="[435,664,1087,1112]" pageId="40" pageNumber="41" refString="Schaefer, S. A. (1997) The neotropical cascudinhos: systematics and biogeography of the Otocinclus catfishes (Siluriformes: Loricariidae). Proceedings of the Academy of Natural Sciences of Philadelphia, 148, 1 - 120." type="journal article" year="1997">Schaefer (1997: 49)</bibRefCitation>
considered that only specimens deposited at the NMW should be considered types, an opinion that has not been unanimously accepted. For example, 
<bibRefCitation id="2C0B1BF74F5FFFD04A6BFBA3FAD5FB9B" author="Vari" box="[1112,1287,1124,1149]" pageId="40" pageNumber="41" refString="Vari, R. P. (1992) Systematics of the Neotropical Characiform genus Steindachnerina Fowler (Pisces: Ostariophysi). Smithsonian Contributions to Zoology, 507, 1 - 118." type="journal article" year="1992">Vari (1992: 48)</bibRefCitation>
designated a MCZ specimen as the lectotype of 
<taxonomicName id="8F9A1D854F5FFFD04C16FB40FCB0FB47" box="[549,866,1159,1184]" class="Actinopterygii" family="Curimatidae" genus="Steindachnerina" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="bimaculata">
<emphasis id="7AEEBA144F5FFFD04C16FB40FCB0FB47" box="[549,866,1159,1184]" italics="true" pageId="40" pageNumber="41">Steindachnerina bimaculata</emphasis>
</taxonomicName>
, quoting Steindachner’s description of the species where he refers to “numerous specimens… in the Museum at Cambridge” (Steindachner, 1876: 79). There are in fact a few more more references by Steindachner to material deposited at the MCZ, which points out that he has not necessarily relied exclusively on material brought by him to the NMW in his descriptions of species based totally or partially on material collected by the Thayer Expedition (for example, 
<taxonomicName id="8F9A1D854F5FFFD04DC4FADEFB23FAD7" box="[1015,1265,1305,1328]" class="Actinopterygii" family="Anostomidae" genus="Leporinus" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="mormyrops">
<emphasis id="7AEEBA144F5FFFD04DC4FADEFB23FAD7" box="[1015,1265,1305,1328]" italics="true" pageId="40" pageNumber="41">Leporinus mormyrops</emphasis>
</taxonomicName>
; 
<bibRefCitation id="2C0B1BF74F5FFFD04B37FAD0FF33FAB2" author="Steindachner" pageId="40" pageNumber="41" refString="Steindachner, F. (1875 b) Die Susswasserfische des sudostlichen Brasilien (II). Sitzungsberichte der Kaiserlichen Akademie der Wissenchaften in Wien, 70 (1), 211 - 245, pls. 1 - 6." type="journal article" year="1875" yearSuffix="b">Steindachner, 1875b</bibRefCitation>
: 244). A possible compromise solution to this problem would be to recognize, as 
<bibRefCitation id="2C0B1BF74F5FFFD04A48FAFBFB2EFAB3" author="Vari" box="[1147,1276,1340,1365]" pageId="40" pageNumber="41" refString="Vari, R. P. (1992) Systematics of the Neotropical Characiform genus Steindachnerina Fowler (Pisces: Ostariophysi). Smithsonian Contributions to Zoology, 507, 1 - 118." type="journal article" year="1992">Vari (1992)</bibRefCitation>
did, the MCZ material as typical only in the relatively few descriptions of fishes based on material collected by the Thayer Expedition where Steindachner specifically mentioned specimens from the “Museum zu Cambridge”. With this view in mind, we opted for not considering as belonging to the typical series the specimens deposited at the MCZ belonging to the same original lots from which the material used by 
<bibRefCitation id="2C0B1BF74F5FFFD04DA8FA0BFBADFA03" author="Steindachner" box="[923,1151,1484,1509]" pageId="40" pageNumber="41" refString="Steindachner, F. (1877) Die Susswasserfische des sudostlichen Brasilien (III). Sitzungsberichte der Kaiserlichen Akademie der Wissenchaften in Wien, 74 (1), 559 - 694, pls. 1 - 13." type="journal article" year="1877">Steindachner (1877)</bibRefCitation>
in his descriptions of the two 
<taxonomicName id="8F9A1D854F5FFFD04EFBFA36FECBF9EF" box="[200,281,1521,1544]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="genus">
<emphasis id="7AEEBA144F5FFFD04EFBFA36FECBF9EF" box="[200,281,1521,1544]" italics="true" pageId="40" pageNumber="41">Brycon</emphasis>
</taxonomicName>
species based on material collected by the Thayer Expedition, 
<taxonomicName id="8F9A1D854F5FFFD04DEEFA36FB81F9EF" box="[989,1107,1521,1544]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5FFFD04DEEFA36FB81F9EF" box="[989,1107,1521,1544]" italics="true" pageId="40" pageNumber="41">B. insignis</emphasis>
</taxonomicName>
and 
<taxonomicName id="8F9A1D854F5FFFD04AB9FA36FB30F9EF" box="[1162,1250,1520,1544]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="ferox">
<emphasis id="7AEEBA144F5FFFD04AB9FA36FB30F9EF" box="[1162,1250,1520,1544]" italics="true" pageId="40" pageNumber="41">B. ferox</emphasis>
</taxonomicName>
. We found three lots at the NMW collection which can be assigned as belonging to the syntypical series used by 
<bibRefCitation id="2C0B1BF74F5FFFD04B38F9D3FF34F9B7" author="Steindachner" pageId="40" pageNumber="41" refString="Steindachner, F. (1877) Die Susswasserfische des sudostlichen Brasilien (III). Sitzungsberichte der Kaiserlichen Akademie der Wissenchaften in Wien, 74 (1), 559 - 694, pls. 1 - 13." type="journal article" year="1877">Steindachner (1877)</bibRefCitation>
: NMW 62925 (2), NMW 62926 (
<quantity id="8F62CBE34F5FFFD04C5BF9FFFD27F9B7" box="[616,757,1591,1616]" metricMagnitude="0" metricUnit="m" metricValue="1.1135" pageId="40" pageNumber="41" unit="mm" value="1113.5">1, 113.5 mm</quantity>
SL), and NMW 62927 (1, 
<quantity id="8F62CBE34F5FFFD04A19F9F0FB5DF9B7" box="[1066,1167,1591,1616]" metricMagnitude="-2" metricUnit="m" metricValue="7.159999999999999" pageId="40" pageNumber="41" unit="mm" value="71.6">71.6 mm</quantity>
SL). Label information for the lots NMW 62925 and NMW 62926 states only that they were collected at the “Rio Parahyba”, with no further locality information. The specimen NMW 62927 was collected at the “Rio Jequitinhonha”, and actually belong to the species herein described as 
<taxonomicName id="8F9A1D854F5FFFD04C44F962FCCDF95A" box="[631,799,1700,1725]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="howesi">
<emphasis id="7AEEBA144F5FFFD04C44F962FCCDF95A" box="[631,799,1700,1725]" italics="true" pageId="40" pageNumber="41">Brycon howesi</emphasis>
</taxonomicName>
(see “Remarks” of this species). In order to dispel any doubts concerning the identity of 
<taxonomicName id="8F9A1D854F5FFFD04C23F90EFD6DF907" box="[528,703,1737,1760]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5FFFD04C23F90EFD6DF907" box="[528,703,1737,1760]" italics="true" pageId="40" pageNumber="41">Brycon insignis</emphasis>
</taxonomicName>
, we therefore select the smaller specimen of the lot NMW 62925 (
<quantity id="8F62CBE34F5FFFD04EACF92BFEC0F8E3" box="[159,274,1772,1796]" metricMagnitude="-1" metricUnit="m" metricValue="1.448" pageId="40" pageNumber="41" unit="mm" value="144.8">144.8 mm</quantity>
SL), which is the syntype in better overall condition, as the lectotype of 
<taxonomicName id="8F9A1D854F5FFFD04A77F92AFB21F8E3" box="[1092,1267,1773,1796]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5FFFD04A77F92AFB21F8E3" box="[1092,1267,1773,1796]" italics="true" pageId="40" pageNumber="41">Brycon insignis</emphasis>
</taxonomicName>
, the remaining syntypes becaming, thus, paralectotypes. As noticed by 
<bibRefCitation id="2C0B1BF74F5FFFD04D2BF8C8FB7AF8CF" author="Steindachner" box="[792,1192,1807,1832]" pageId="40" pageNumber="41" refString="Steindachner, F. (1879 c) Uber einige neue und seltene Fisch-Arten aus den K. K. Zoologischen Museen zu Wien, Stuttgart und Warschau. Denkschriften der Mathematisch-Naturwissenschaftlichen Classe der Akademie der Wissenschaften, Wien, 41, 1 - 52, pls. 1 - 9." type="journal article" year="1879" yearSuffix="c">Steindachner (1879c: 50, footnote)</bibRefCitation>
, the indication of the plate in 
<bibRefCitation id="2C0B1BF74F5FFFD04EC2F8F3FDDFF8AA" author="Steindachner" box="[241,525,1844,1869]" pageId="40" pageNumber="41" refString="Steindachner, F. (1877) Die Susswasserfische des sudostlichen Brasilien (III). Sitzungsberichte der Kaiserlichen Akademie der Wissenchaften in Wien, 74 (1), 559 - 694, pls. 1 - 13." type="journal article" year="1877">Steindachner (1877: 135)</bibRefCitation>
is incorrect, 
<taxonomicName id="8F9A1D854F5FFFD04CACF8F2FC9CF8AB" box="[671,846,1845,1868]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5FFFD04CACF8F2FC9CF8AB" box="[671,846,1845,1868]" italics="true" pageId="40" pageNumber="41">Brycon insignis</emphasis>
</taxonomicName>
being actually the species portrayed in figures 1 and 1a, figures 2 and 2b referring to 
<taxonomicName id="8F9A1D854F5FFFD04C37F89EFD8EF897" box="[516,604,1880,1904]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="ferox">
<emphasis id="7AEEBA144F5FFFD04C37F89EFD8EF897" box="[516,604,1880,1904]" italics="true" pageId="40" pageNumber="41">B. ferox</emphasis>
</taxonomicName>
. 
<bibRefCitation id="2C0B1BF74F5FFFD04C59F890FC54F897" author="Steindachner" box="[618,902,1879,1904]" pageId="40" pageNumber="41" refString="Steindachner, F. (1879 c) Uber einige neue und seltene Fisch-Arten aus den K. K. Zoologischen Museen zu Wien, Stuttgart und Warschau. Denkschriften der Mathematisch-Naturwissenschaftlichen Classe der Akademie der Wissenschaften, Wien, 41, 1 - 52, pls. 1 - 9." type="journal article" year="1879" yearSuffix="c">Steindachner (1879c: 50)</bibRefCitation>
considered 
<taxonomicName id="8F9A1D854F5FFFD04A3DF89EFB6FF897" box="[1038,1213,1881,1904]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5FFFD04A3DF89EFB6FF897" box="[1038,1213,1881,1904]" italics="true" pageId="40" pageNumber="41">Brycon insignis</emphasis>
</taxonomicName>
as a synonym of 
<taxonomicName id="8F9A1D854F5FFFD04BB7F89EFF3BF872" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="devillei">
<emphasis id="7AEEBA144F5FFFD04BB7F89EFF3BF872" italics="true" pageId="40" pageNumber="41">B. devillei</emphasis>
</taxonomicName>
, an opinion that prevailed in the literature for a long time (e.g., 
<bibRefCitation id="2C0B1BF74F5FFFD04D9CF8BAFB94F873" author="Howes" box="[943,1094,1916,1940]" pageId="40" pageNumber="41" refString="Howes, G. J. (1982) Review of the genus Brycon (Teleostei: Characoidei). Bulletin of the British Museum (Natural History), Zoology, 43 (1), 1 - 47." type="journal article" year="1982">Howes, 1982</bibRefCitation>
). 
<taxonomicName id="8F9A1D854F5FFFD04A6AF8BAFAD1F872" box="[1113,1283,1916,1941]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="devillei">
<emphasis id="7AEEBA144F5FFFD04A6AF8BAFAD1F872" box="[1113,1283,1916,1941]" italics="true" pageId="40" pageNumber="41">Brycon devillei</emphasis>
</taxonomicName>
is considered herein as a 
<emphasis id="7AEEBA144F5FFFD04F26F866FE35F85E" box="[277,487,1952,1977]" italics="true" pageId="40" pageNumber="41">species inquirenda</emphasis>
(see the item “Remarks” under this species, below).
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F5FFFD14EF4F803FD31FB0F" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41" type="reference_group">
<paragraph id="482566064F5FFFD14EF4F803FA94FED8" blockId="40.[151,1437,151,2013]" lastBlockId="41.[151,1437,151,2013]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<treatmentCitationGroup id="688A41284F5FFFD14EF4F803FA94FED8" infered="true" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
<taxonomicName id="8F9A1D854F5FFFD04EF4F803FC99F83A" ID-CoL="7382H" authority="A. Miranda-Ribeiro (1902: 254)" authorityName="A. Miranda-Ribeiro" authorityPageNumber="254" authorityYear="1902" box="[199,843,1988,2013]" family="Bryconidae" genus="Megalobrycon" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="40" pageNumber="41" phylum="Chordata" rank="species" species="piabanha">
<emphasis id="7AEEBA144F5FFFD04EF4F803FE09F83A" box="[199,475,1988,2013]" italics="true" pageId="40" pageNumber="41">Megalobrycon piabanha</emphasis>
<treatmentCitation id="C93B40174F5FFFD04FD1F803FC99F83A" author="Miranda-Ribeiro" box="[482,843,1988,2013]" page="254" pageId="40" pageNumber="41" year="1902">
<bibRefCitation id="2C0B1BF74F5FFFD04FD1F803FC99F83A" author="Miranda-Ribeiro" box="[482,843,1988,2013]" pageId="40" pageNumber="41" refString="Miranda-Ribeiro, A. (1902) Oito especies de peixes do Rio Pomba. A Lavoura, Boletim da Sociedade de Agricultura, 1902, 1 - 8." type="journal article" year="1902">A. Miranda-Ribeiro (1902: 254)</bibRefCitation>
</treatmentCitation>
</taxonomicName>
was described from the rio Pomba, a tributary of the rio Paraíba do Sul at Minas Gerais state, Brazil, based on a large (
<quantity id="8F62CBE34F5EFFD14DBAFF50FC3BFF48" box="[905,1001,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="5.2" pageId="41" pageNumber="42" unit="mm" value="520.0">520 mm</quantity>
TL) specimen. A little after, the same author (
<bibRefCitation id="2C0B1BF74F5EFFD14EC7FF7BFDF6FF32" author="Miranda-Ribeiro" box="[244,548,188,213]" pageId="41" pageNumber="42" refString="Miranda-Ribeiro, A. (1905) Genus Megalobrycon, Gnthr. Seu enumeratio systematica hujus generis Characinidarum specierum. Arquivos do Museu Nacional, 13, 147 - 154, 1 pl." type="journal article" year="1905">A. Miranda-Ribeiro, 1905</bibRefCitation>
) redescribed and figured the species, unjustifiably ammending its name to 
<taxonomicName id="8F9A1D854F5EFFD14EA4FF18FE61FF1F" box="[151,435,223,248]" family="Bryconidae" genus="Megalobrycon" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="piabagna">
<emphasis id="7AEEBA144F5EFFD14EA4FF18FE61FF1F" box="[151,435,223,248]" italics="true" pageId="41" pageNumber="42">Megalobrycon piabagna</emphasis>
</taxonomicName>
. Searches in the MNRJ collection failed in locate the holotype of 
<taxonomicName id="8F9A1D854F5EFFD14ACAFF18FED3FEFA" family="Bryconidae" genus="Megalobrycon" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="piabanha">
<emphasis id="7AEEBA144F5EFFD14ACAFF18FED3FEFA" italics="true" pageId="41" pageNumber="42">Megalobrycon piabanha</emphasis>
</taxonomicName>
. However, the description and illustration given by A. 
<bibRefCitation id="2C0B1BF74F5EFFD14DBEFEC3FB4AFEFB" author="Miranda-Ribeiro" box="[909,1176,260,285]" pageId="41" pageNumber="42" refString="Miranda-Ribeiro, A. (1902) Oito especies de peixes do Rio Pomba. A Lavoura, Boletim da Sociedade de Agricultura, 1902, 1 - 8." type="journal article" year="1902">Miranda-Ribeiro (1902</bibRefCitation>
, 1904), as well as the provenance of the specimen, leave no doubt that this name represents a junior synonym of 
<taxonomicName id="8F9A1D854F5EFFD14AA3FEEEFAEDFEA7" ID-CoL="NH7J" box="[1168,1343,297,320]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5EFFD14AA3FEEEFAEDFEA7" box="[1168,1343,297,320]" italics="true" pageId="41" pageNumber="42">Brycon insignis</emphasis>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="482566064F5EFFD14EF4FE8BFD31FB0F" blockId="41.[151,1437,151,2013]" pageId="41" pageNumber="42">
<taxonomicName id="8F9A1D854F5EFFD14EF4FE8BFCAFFE82" authority="Eigenmann &amp; Norris" authorityName="Eigenmann &amp; Norris" authorityYear="1900" box="[199,893,332,357]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F5EFFD14EF4FE8BFE12FE83" box="[199,448,332,357]" italics="true" pageId="41" pageNumber="42">Catabasis acuminatus</emphasis>
<bibRefCitation id="2C0B1BF74F5EFFD14FFAFE8BFCAFFE82" author="Eigenmann" box="[457,893,332,357]" pageId="41" pageNumber="42" refString="Eigenmann, C. H. &amp; Norris, A. A. (1900) Sobre alguns peixes de S. Paulo, Brasil. Revista do Museu Paulista, 4, 349 - 362." type="journal article" year="1900">Eigenmann &amp; Norris (1900: 358–359)</bibRefCitation>
</taxonomicName>
was described from the “Rio Tieté”, São Paulo state, Brazil. As discussed under the item “Remarks” of the genus 
<taxonomicName id="8F9A1D854F5EFFD14D4EFEB6FC1CFE6F" ID-CoL="3DKB" box="[893,974,369,392]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="7AEEBA144F5EFFD14D4EFEB6FC1CFE6F" box="[893,974,369,392]" italics="true" pageId="41" pageNumber="42">Brycon</emphasis>
</taxonomicName>
, the genus 
<taxonomicName id="8F9A1D854F5EFFD14A7EFEA8FB6FFE6F" box="[1101,1213,367,392]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="7AEEBA144F5EFFD14A7EFEA8FB6FFE6F" box="[1101,1213,367,392]" italics="true" pageId="41" pageNumber="42">Catabasis</emphasis>
</taxonomicName>
was for a long time considered as possibly related to 
<taxonomicName id="8F9A1D854F5EFFD14C3AFE53FDBDFE4A" ID-CoL="7BWV" box="[521,623,404,429]" class="Actinopterygii" family="Bryconidae" genus="Salminus" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="7AEEBA144F5EFFD14C3AFE53FDBDFE4A" box="[521,623,404,429]" italics="true" pageId="41" pageNumber="42">Salminus</emphasis>
</taxonomicName>
(
<bibRefCitation id="2C0B1BF74F5EFFD14C4DFE53FC66FE4B" author="Eigenmann" box="[638,948,404,429]" pageId="41" pageNumber="42" refString="Eigenmann, C. H. &amp; Norris, A. A. (1900) Sobre alguns peixes de S. Paulo, Brasil. Revista do Museu Paulista, 4, 349 - 362." type="journal article" year="1900">Eigenmann &amp; Norris, 1900</bibRefCitation>
; 
<bibRefCitation id="2C0B1BF74F5EFFD14D8CFE53FB8DFE4A" author="Roberts" box="[959,1119,404,429]" pageId="41" pageNumber="42" refString="Roberts, T. R. (1969) Osteology and relationships of characoid fishes, particularly the genera Hepsetus, Salminus, Hoplias, Ctenolucius, and Acestrorhynchus. Proceedings of the California Academy of Sciences, 36 (15), 391 - 500." type="journal article" year="1969">Roberts, 1969</bibRefCitation>
; 
<bibRefCitation id="2C0B1BF74F5EFFD14A58FE53FB39FE4B" author="Gery" box="[1131,1259,404,428]" pageId="41" pageNumber="42" refString="Gery, J. (1977) Characoids of the world. T. F. H. Publications, Neptune City, 672 pp." type="book" year="1977">Géry, 1977</bibRefCitation>
). 
<bibRefCitation id="2C0B1BF74F5EFFD14ACDFE52FF1EFE37" author="Howes" pageId="41" pageNumber="42" refString="Howes, G. J. (1982) Review of the genus Brycon (Teleostei: Characoidei). Bulletin of the British Museum (Natural History), Zoology, 43 (1), 1 - 47." type="journal article" year="1982">Howes (1982: 4–6)</bibRefCitation>
, based on the examination of a single specimen (MZUSP 1533), was the first to consider 
<taxonomicName id="8F9A1D854F5EFFD14A8AFE70FAFBFE37" box="[1209,1321,439,464]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="7AEEBA144F5EFFD14A8AFE70FAFBFE37" box="[1209,1321,439,464]" italics="true" pageId="41" pageNumber="42">Catabasis</emphasis>
</taxonomicName>
merely as a synonym of 
<taxonomicName id="8F9A1D854F5EFFD14F77FE1AFE47FE13" ID-CoL="3DKB" box="[324,405,477,500]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="7AEEBA144F5EFFD14F77FE1AFE47FE13" box="[324,405,477,500]" italics="true" pageId="41" pageNumber="42">Brycon</emphasis>
</taxonomicName>
. He also considered that 
<taxonomicName id="8F9A1D854F5EFFD14CF5FE1BFC10FE13" box="[710,962,476,501]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F5EFFD14CF5FE1BFC10FE13" box="[710,962,476,501]" italics="true" pageId="41" pageNumber="42">Catabasis acuminatus</emphasis>
</taxonomicName>
was probably a synonym of 
<taxonomicName id="8F9A1D854F5EFFD14B12FE1AFAAEFE13" ID-CoL="NH7C" box="[1313,1404,476,500]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="ferox">
<emphasis id="7AEEBA144F5EFFD14B12FE1AFAAEFE13" box="[1313,1404,476,500]" italics="true" pageId="41" pageNumber="42">B. ferox</emphasis>
</taxonomicName>
, a supposition mainly based on the comparison of the specimen available to 
<bibRefCitation id="2C0B1BF74F5EFFD14DDCFDC7FB43FDFF" author="Howes" box="[1007,1169,512,536]" pageId="41" pageNumber="42" refString="Howes, G. J. (1982) Review of the genus Brycon (Teleostei: Characoidei). Bulletin of the British Museum (Natural History), Zoology, 43 (1), 1 - 47." type="journal article" year="1982">Howes (1982)</bibRefCitation>
with the illustration of 
<taxonomicName id="8F9A1D854F5EFFD14EA4FDE2FDE6FDDB" ID-CoL="NH7C" authority="in Steindachner (1877)" authorityName="in Steindachner" authorityYear="1877" box="[151,564,548,573]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="ferox">
<emphasis id="7AEEBA144F5EFFD14EA4FDE2FEFAFDDB" box="[151,296,548,572]" italics="true" pageId="41" pageNumber="42">Brycon ferox</emphasis>
in 
<bibRefCitation id="2C0B1BF74F5EFFD14F7EFDE3FDE6FDDB" author="Steindachner" box="[333,564,548,573]" pageId="41" pageNumber="42" refString="Steindachner, F. (1877) Die Susswasserfische des sudostlichen Brasilien (III). Sitzungsberichte der Kaiserlichen Akademie der Wissenchaften in Wien, 74 (1), 559 - 694, pls. 1 - 13." type="journal article" year="1877">Steindachner (1877)</bibRefCitation>
</taxonomicName>
. As noted above, the indication of the figures in 
<bibRefCitation id="2C0B1BF74F5EFFD14A6EFDE3FA4EFDDB" author="Steindachner" box="[1117,1436,548,573]" pageId="41" pageNumber="42" refString="Steindachner, F. (1877) Die Susswasserfische des sudostlichen Brasilien (III). Sitzungsberichte der Kaiserlichen Akademie der Wissenchaften in Wien, 74 (1), 559 - 694, pls. 1 - 13." type="journal article" year="1877">Steindachner (1877, plate 4)</bibRefCitation>
are shifted, so what 
<bibRefCitation id="2C0B1BF74F5EFFD14FBBFD8FFD89FD87" author="Howes" box="[392,603,584,608]" pageId="41" pageNumber="42" refString="Howes, G. J. (1982) Review of the genus Brycon (Teleostei: Characoidei). Bulletin of the British Museum (Natural History), Zoology, 43 (1), 1 - 47." type="journal article" year="1982">Howes (1982: 28)</bibRefCitation>
was in fact suggesting was that 
<taxonomicName id="8F9A1D854F5EFFD14DD1FD80FB0CFD87" box="[994,1246,583,608]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F5EFFD14DD1FD80FB0CFD87" box="[994,1246,583,608]" italics="true" pageId="41" pageNumber="42">Catabasis acuminatus</emphasis>
</taxonomicName>
was probably a synonym of 
<taxonomicName id="8F9A1D854F5EFFD14F15FDAAFE05FD63" ID-CoL="NH7J" box="[294,471,621,644]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5EFFD14F15FDAAFE05FD63" box="[294,471,621,644]" italics="true" pageId="41" pageNumber="42">Brycon insignis</emphasis>
</taxonomicName>
. 
<taxonomicName id="8F9A1D854F5EFFD14FD4FDABFD33FD63" box="[487,737,620,645]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F5EFFD14FD4FDABFD33FD63" box="[487,737,620,645]" italics="true" pageId="41" pageNumber="42">Catabasis acuminatus</emphasis>
</taxonomicName>
was noted by its peculiar dentition, consisting exclusively in unicuspid teeth (
<bibRefCitation id="2C0B1BF74F5EFFD14F6AFD48FDD3FD4F" author="Roberts" box="[345,513,655,680]" pageId="41" pageNumber="42" refString="Roberts, T. R. (1969) Osteology and relationships of characoid fishes, particularly the genera Hepsetus, Salminus, Hoplias, Ctenolucius, and Acestrorhynchus. Proceedings of the California Academy of Sciences, 36 (15), 391 - 500." type="journal article" year="1969">Roberts, 1969</bibRefCitation>
: 437–438; 
<bibRefCitation id="2C0B1BF74F5EFFD14CB5FD57FCF0FD4F" author="Howes" box="[646,802,656,680]" pageId="41" pageNumber="42" refString="Howes, G. J. (1982) Review of the genus Brycon (Teleostei: Characoidei). Bulletin of the British Museum (Natural History), Zoology, 43 (1), 1 - 47." type="journal article" year="1982">Howes, 1982</bibRefCitation>
: 4). The only other 
<taxonomicName id="8F9A1D854F5EFFD14A26FD56FBB4FD4F" ID-CoL="3DKB" box="[1045,1126,657,680]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="7AEEBA144F5EFFD14A26FD56FBB4FD4F" box="[1045,1126,657,680]" italics="true" pageId="41" pageNumber="42">Brycon</emphasis>
</taxonomicName>
species known to possess exclusively unicuspid teeth is 
<taxonomicName id="8F9A1D854F5EFFD14FD4FD72FD66FD2B" ID-CoL="NH73" box="[487,692,693,716]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="coquenani">
<emphasis id="7AEEBA144F5EFFD14FD4FD72FD66FD2B" box="[487,692,693,716]" italics="true" pageId="41" pageNumber="42">Brycon coquenani</emphasis>
</taxonomicName>
(see above). However, from the examination of both the specimen identified by 
<bibRefCitation id="2C0B1BF74F5EFFD14F1EFD1FFE1CFD17" author="Howes" box="[301,462,728,752]" pageId="41" pageNumber="42" refString="Howes, G. J. (1982) Review of the genus Brycon (Teleostei: Characoidei). Bulletin of the British Museum (Natural History), Zoology, 43 (1), 1 - 47." type="journal article" year="1982">Howes (1982)</bibRefCitation>
as 
<taxonomicName id="8F9A1D854F5EFFD14FC7FD1EFD1CFD17" ID-CoL="RQZJ" box="[500,718,729,752]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F5EFFD14FC7FD1EFD1CFD17" box="[500,718,729,752]" italics="true" pageId="41" pageNumber="42">Brycon acuminatus</emphasis>
</taxonomicName>
, as well as the holotype of 
<taxonomicName id="8F9A1D854F5EFFD14A36FD10FB2CFD17" box="[1029,1278,727,752]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F5EFFD14A36FD10FB2CFD17" box="[1029,1278,727,752]" italics="true" pageId="41" pageNumber="42">Catabasis acuminatus</emphasis>
</taxonomicName>
(CAS 11894) we were unable to identify any other character differentiating this purported species from 
<taxonomicName id="8F9A1D854F5EFFD14A4EFD3AFB21FCF3" ID-CoL="NH7J" box="[1149,1267,765,788]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5EFFD14A4EFD3AFB21FCF3" box="[1149,1267,765,788]" italics="true" pageId="41" pageNumber="42">B. insignis</emphasis>
</taxonomicName>
. Similar-sized, typical 
<taxonomicName id="8F9A1D854F5EFFD14EC3FCE6FE76FCDF" ID-CoL="NH7J" box="[240,420,801,824]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5EFFD14EC3FCE6FE76FCDF" box="[240,420,801,824]" italics="true" pageId="41" pageNumber="42">Brycon insignis</emphasis>
</taxonomicName>
possess teeth with small, poorly developed lateral cusps, a condition not completely dissimilar to the condition found in the specimens assigned to 
<taxonomicName id="8F9A1D854F5EFFD14D6EFC83FB84FCBB" box="[861,1110,836,861]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F5EFFD14D6EFC83FB84FCBB" box="[861,1110,836,861]" italics="true" pageId="41" pageNumber="42">Catabasis acuminatus</emphasis>
</taxonomicName>
. We consider the absence of cusps in the specimens assigned to 
<taxonomicName id="8F9A1D854F5EFFD14C7BFCA0FC95FC67" box="[584,839,871,896]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F5EFFD14C7BFCA0FC95FC67" box="[584,839,871,896]" italics="true" pageId="41" pageNumber="42">Catabasis acuminatus</emphasis>
</taxonomicName>
should be interpreted as constituting intraspecific variation within 
<taxonomicName id="8F9A1D854F5EFFD14F6BFC4AFDDEFC43" ID-CoL="NH7J" box="[344,524,909,932]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5EFFD14F6BFC4AFDDEFC43" box="[344,524,909,932]" italics="true" pageId="41" pageNumber="42">Brycon insignis</emphasis>
</taxonomicName>
, and, thus, that 
<taxonomicName id="8F9A1D854F5EFFD14CFCFC4BFC19FC43" box="[719,971,908,933]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F5EFFD14CFCFC4BFC19FC43" box="[719,971,908,933]" italics="true" pageId="41" pageNumber="42">Catabasis acuminatus</emphasis>
</taxonomicName>
should be considered a synonym of 
<taxonomicName id="8F9A1D854F5EFFD14BB7FC4AFF3CFC2F" ID-CoL="NH7J" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5EFFD14BB7FC4AFF3CFC2F" italics="true" pageId="41" pageNumber="42">B. insignis</emphasis>
</taxonomicName>
. As for the type-locality of 
<taxonomicName id="8F9A1D854F5EFFD14C0AFC68FCE6FC2F" box="[569,820,943,968]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F5EFFD14C0AFC68FCE6FC2F" box="[569,820,943,968]" italics="true" pageId="41" pageNumber="42">Catabasis acuminatus</emphasis>
</taxonomicName>
, said to be at an unspecified locality in the rio Tietê system (rio Paraná basin), no further records for 
<taxonomicName id="8F9A1D854F5EFFD14CF5FC12FCA4FC0B" ID-CoL="NH7J" box="[710,886,981,1004]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5EFFD14CF5FC12FCA4FC0B" box="[710,886,981,1004]" italics="true" pageId="41" pageNumber="42">Brycon insignis</emphasis>
</taxonomicName>
for the rio Paraná system exist. The rio Tietê is now a highly disturbed river system, its upper reaches heavily polluted by sewage and industrial discharges (
<bibRefCitation id="2C0B1BF74F5EFFD14B64FC3FFF01FBD2" author="Jorge" pageId="41" pageNumber="42" refString="Jorge, J. (2006) Tiete, o rio que a cidade perdeu. Sao Paulo, 1890 - 1940. Alameda Casa Editorial, Sao Paulo, 232 pp." type="book chapter" year="2006">Jorge, 2006</bibRefCitation>
), and its middle and lower reaches dammed by a series of hydroelectric reservoirs. Meagre information is available on the ichthyofauna of the rio Tietê prior to the drastic environmental deterioration of this river system (e.g., 
<bibRefCitation id="2C0B1BF74F5EFFD14EEAFBA3FEA5FB9B" author="Ihering" box="[217,375,1124,1149]" pageId="41" pageNumber="42" refString="Ihering, R. (1929) Da vida dos peixes. Melhoramentos, Sao Paulo, 149 pp." type="book" year="1929">Ihering, 1929</bibRefCitation>
). However, it is quite unlikely that a large fish species, targeted by fishermen like a 
<taxonomicName id="8F9A1D854F5EFFD14B78FBA2FA4EFB9B" ID-CoL="3DKB" box="[1355,1436,1125,1148]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="genus">
<emphasis id="7AEEBA144F5EFFD14B78FBA2FA4EFB9B" box="[1355,1436,1125,1148]" italics="true" pageId="41" pageNumber="42">Brycon</emphasis>
</taxonomicName>
species would not have been reported again if really occurring in the rio Tietê basin. We consider that the reported type-locality of 
<taxonomicName id="8F9A1D854F5EFFD14F62FB6BFD9FFB23" box="[337,589,1196,1221]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F5EFFD14F62FB6BFD9FFB23" box="[337,589,1196,1221]" italics="true" pageId="41" pageNumber="42">Catabasis acuminatus</emphasis>
</taxonomicName>
is incorrect, and that the holotype of this nominal species was probably collected somewhere in the rio Paraíba do Sul basin.
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F5EFFD14EF4FB33FC4AF9B7" pageId="41" pageNumber="42" type="materials_examined">
<paragraph id="482566064F5EFFD14EF4FB33FA41FA9F" blockId="41.[151,1437,151,2013]" pageId="41" pageNumber="42">
<materialsCitation id="F8F26C5B4F5EFFD14EF4FB33FBA1FAD7" ID-GBIF-Occurrence="1425222570" collectingDate="1874-01" collectionCode="NMW" location="Rio Parahyba" pageId="41" pageNumber="42" specimenCount="2" stateProvince="Rio Parahyba" typeStatus="lectotype">
<emphasis id="7AEEBA144F5EFFD14EF4FB33FD57FAEA" bold="true" box="[199,645,1268,1293]" pageId="41" pageNumber="42">
Material examined. 
<typeStatus id="9721D8A44F5EFFD14FF8FB33FDD4FAEA" box="[459,518,1268,1293]" pageId="41" pageNumber="42">Type</typeStatus>
material:
</emphasis>
<collectionCode id="2E8BFEC34F5EFFD14CA5FB32FD0FFAEB" box="[662,733,1269,1292]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="41" pageNumber="42">NMW</collectionCode>
62925:2 (
<quantity id="8F62CBE34F5EFFD14D55FB33FBD4FAEB" box="[870,1030,1268,1292]" metricMagnitude="0" metricUnit="m" metricValue="1.1447999999999998" pageId="41" pageNumber="42" unit="mm" value="1144.8">1, 144.8 mm</quantity>
SL): &quot; 
<collectingRegion id="8A5EA8E44F5EFFD14A50FB33FADBFAEA" box="[1123,1289,1268,1293]" isEnumeration="true" pageId="41" pageNumber="42">Rio Parahyba</collectingRegion>
, 
<collectingDate id="2C60B92E4F5EFFD14B2CFB32FA4AFAEB" box="[1311,1432,1268,1292]" pageId="41" pageNumber="42" value="1874-01">
<date id="3C2440C64F5EFFD14B2CFB32FA4AFAEB" box="[1311,1432,1268,1292]" pageId="41" pageNumber="42" value="1874-01">Jan. 1874</date>
</collectingDate>
; Steindachner don.&quot; 
<typeStatus id="9721D8A44F5EFFD14F4BFADFFE3AFAD7" box="[376,488,1304,1328]" pageId="41" pageNumber="42" type="lectotype">Lectotype</typeStatus>
of 
<taxonomicName id="8F9A1D854F5EFFD14C21FADEFCB2FAD7" authority="Steindachner" authorityName="Steindachner" box="[530,864,1303,1328]" class="Actinopterygii" family="Bryconidae" genus="Brycon" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="insignis">
<emphasis id="7AEEBA144F5EFFD14C21FADEFD11FAD7" box="[530,707,1305,1328]" italics="true" pageId="41" pageNumber="42">Brycon insignis</emphasis>
Steindachner
</taxonomicName>
, by present designation
</materialsCitation>
. NMW 62925:1 (
<quantity id="8F62CBE34F5EFFD14B72FADFFF11FAB3" metricMagnitude="0" metricUnit="m" metricValue="1.1677" pageId="41" pageNumber="42" unit="mm" value="1167.7">1, 167.7 mm</quantity>
SL); 
<materialsCitation id="F8F26C5B4F5EFFD14F37FAFAFBE0FAB2" ID-GBIF-Occurrence="1425222555" box="[260,1074,1340,1365]" collectingDate="1874-01" collectionCode="NMW" location="Rio Parahyba" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Rio Parahyba" typeStatus="paralectotype">
<collectionCode id="2E8BFEC34F5EFFD14F37FAFAFE9EFAB3" box="[260,332,1341,1364]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="41" pageNumber="42">NMW</collectionCode>
62926 (
<quantity id="8F62CBE34F5EFFD14F9DFAFBFDE9FAB3" box="[430,571,1340,1365]" metricMagnitude="0" metricUnit="m" metricValue="1.1135" pageId="41" pageNumber="42" unit="mm" value="1113.5">1, 113.5 mm</quantity>
SL): same data as lectotype. 
<typeStatus id="9721D8A44F5EFFD14DB8FAFBFBE0FAB2" box="[907,1074,1340,1365]" pageId="41" pageNumber="42" type="paralectotype">Paralectotypes</typeStatus>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5EFFD14A0DFAFBFA41FA9F" ID-GBIF-Occurrence="1425222557" collectingDate="1901" collectingDateMax="1905" collectingDateMin="1901" collectionCode="CAS" collectorName="H. von Ihering" country="Brazil" location="rio Tiete" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
<collectionCode id="2E8BFEC34F5EFFD14A0DFAFBFBA6FAB3" box="[1086,1140,1340,1364]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="41" pageNumber="42">CAS</collectionCode>
11894 (
<quantity id="8F62CBE34F5EFFD14AE6FAFBFAB1FAB3" box="[1237,1379,1340,1365]" metricMagnitude="0" metricUnit="m" metricValue="1.1351" pageId="41" pageNumber="42" unit="mm" value="1135.1">1, 135.1 mm</quantity>
SL), 
<collectingCountry id="308D26964F5EFFD14EA4FA98FF0CFA9F" box="[151,222,1375,1400]" name="Brazil" pageId="41" pageNumber="42">Brazil</collectingCountry>
, 
<collectingRegion id="8A5EA8E44F5EFFD14EDAFAA7FE8FFA9F" box="[233,349,1375,1400]" country="Brazil" name="Sao Paulo" pageId="41" pageNumber="42">São Paulo</collectingRegion>
, 
<location id="4D4530DD4F5EFFD14F5BFA98FE18FA9F" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5EFFD14F5BFA98FE18FA9F" box="[360,458,1375,1400]" country="Brazil" name="rio Tiete" pageId="41" pageNumber="42" stateProvince="Sao Paulo">rio Tietê</location>
; 
<collectorName id="E56F03D04F5EFFD14FE5FAA7FDAFFA9F" box="[470,637,1375,1400]" pageId="41" pageNumber="42">H. von Ihering</collectorName>
, 
<collectingDate id="2C60B92E4F5EFFD14CB4FA98FCDBFA9F" box="[647,777,1375,1400]" pageId="41" pageNumber="42" value="1901" valueMax="1905">
<date id="3C2440C64F5EFFD14CB4FA98FCDBFA9F" box="[647,777,1375,1400]" pageId="41" pageNumber="42" value="1901" valueMax="1905">1901–1905</date>
</collectingDate>
. 
<typeStatus id="9721D8A44F5EFFD14D27FA98FCAEFA9F" box="[788,892,1375,1400]" pageId="41" pageNumber="42" type="holotype">Holotype</typeStatus>
of 
<taxonomicName id="8F9A1D854F5EFFD14D92FA98FA41FA9F" authority="Eigenmann &amp; Norris." authorityName="Eigenmann &amp; Norris." box="[929,1427,1375,1400]" family="Bryconidae" genus="Catabasis" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="acuminatus">
<emphasis id="7AEEBA144F5EFFD14D92FA98FB4BFA9F" box="[929,1177,1375,1400]" italics="true" pageId="41" pageNumber="42">Catabasis acuminatus</emphasis>
Eigenmann &amp; Norris.
</taxonomicName>
</materialsCitation>
</paragraph>
<paragraph id="482566064F5EFFD14EF4FA43FC4AF9B7" blockId="41.[151,1437,151,2013]" pageId="41" pageNumber="42">
<materialsCitation id="F8F26C5B4F5EFFD14EF4FA43FC66FA7B" ID-GBIF-Occurrence="1425222549" box="[199,948,1412,1437]" collectionCode="MZUSP" country="Brazil" location="Sao Paulo" pageId="41" pageNumber="42" specimenCode="MZUSP 51309" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
<emphasis id="7AEEBA144F5EFFD14EF4FA43FDF8FA7A" bold="true" box="[199,554,1412,1437]" pageId="41" pageNumber="42">
Non 
<typeStatus id="9721D8A44F5EFFD14F32FA43FE91FA7A" box="[257,323,1412,1437]" pageId="41" pageNumber="42">types</typeStatus>
: 
<collectingCountry id="308D26964F5EFFD14F63FA43FE49FA7A" box="[336,411,1412,1437]" name="Brazil" pageId="41" pageNumber="42">Brazil</collectingCountry>
, 
<collectingRegion id="8A5EA8E44F5EFFD14F9BFA43FDF7FA7A" box="[424,549,1412,1437]" country="Brazil" name="Sao Paulo" pageId="41" pageNumber="42">São Paulo</collectingRegion>
:
</emphasis>
<specimenCode id="183CCE7D4F5EFFD14C01FA43FD0FFA7A" box="[562,733,1412,1437]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="42">
<collectionCode id="2E8BFEC34F5EFFD14C01FA43FD5DFA7B" box="[562,655,1412,1436]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="42">MZUSP</collectionCode>
51309
</specimenCode>
(
<quantity id="8F62CBE34F5EFFD14CC2FA43FCADFA7B" box="[753,895,1412,1437]" metricMagnitude="0" metricUnit="m" metricValue="1.2537" pageId="41" pageNumber="42" unit="mm" value="1253.7">1, 253.7 mm</quantity>
SL)
</materialsCitation>
; MZUSP 62746 (2, 217.9– 
<quantity id="8F62CBE34F5EFFD14AC3FA43FAB0FA7B" box="[1264,1378,1412,1436]" metricMagnitude="-1" metricUnit="m" metricValue="3.104" pageId="41" pageNumber="42" unit="mm" value="310.4">310.4 mm</quantity>
SL); MZUSP 42045 (4, 1 cs, 
<quantity id="8F62CBE34F5EFFD14F98FA60FDB9FA58" box="[427,619,1447,1472]" metricMagnitude="-1" metricUnit="m" metricValue="1.6945000000000001" metricValueMax="1.9580000000000002" metricValueMin="1.4309999999999998" pageId="41" pageNumber="42" unit="mm" value="169.45" valueMax="195.8" valueMin="143.1">143.1–195.8 mm</quantity>
SL); 
<materialsCitation id="F8F26C5B4F5EFFD14C9FFA6FFBAEFA02" ID-GBIF-Occurrence="1425222578" collectionCode="MZUSP" country="Brazil" latitude="-23.1" location="Cacapava" longLatPrecision="1257" longitude="-45.716667" pageId="41" pageNumber="42" specimenCode="MZUSP 104939" specimenCount="1" stateProvince="Paraiba" typeStatus="holotype">
<specimenCode id="183CCE7D4F5EFFD14C9FFA6FFCB7FA27" box="[684,869,1448,1472]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="42">
<collectionCode id="2E8BFEC34F5EFFD14C9FFA6FFCDBFA27" box="[684,777,1448,1472]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="42">MZUSP</collectionCode>
104939
</specimenCode>
(1, 493.0 mm SL): CESP-Paraibuna fish hatchery (stocked from specimens collected at 
<location id="4D4530DD4F5EFFD14C0DFA0BFD7CFA03" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5EFFD14C0DFA0BFD7CFA03" box="[574,686,1484,1508]" country="Brazil" latitude="-23.1" longLatPrecision="1257" longitude="-45.716667" name="Cacapava" pageId="41" pageNumber="42" stateProvince="Paraiba">Caçapava</location>
, rio 
<collectingRegion id="8A5EA8E44F5EFFD14CECFA0BFCE1FA02" box="[735,819,1484,1509]" country="Brazil" name="Paraiba" pageId="41" pageNumber="42">Paraíba</collectingRegion>
do Sul, c. 
<geoCoordinate id="2DAE00C14F5EFFD14D98FA0BFC29FA02" box="[939,1019,1484,1509]" direction="south" orientation="latitude" pageId="41" pageNumber="42" precision="925" value="-23.1">23°6’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5EFFD14A3BFA0BFBA1FA02" box="[1032,1139,1484,1509]" direction="west" orientation="longitude" pageId="41" pageNumber="42" precision="925" value="-45.716667">45°43’W</geoCoordinate>
)
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5EFFD14AB5FA0BFC13F9EF" ID-GBIF-Occurrence="1425222590" collectionCode="MZUSP" country="Brazil" latitude="-22.65" location="Cachoeira Paulista" longLatPrecision="1259" longitude="-45.0" pageId="41" pageNumber="42" specimenCode="MZUSP 19128" specimenCount="1" stateProvince="Paraiba" typeStatus="holotype">
<specimenCode id="183CCE7D4F5EFFD14AB5FA0BFAE2FA03" box="[1158,1328,1484,1508]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="42">
<collectionCode id="2E8BFEC34F5EFFD14AB5FA0BFB31FA03" box="[1158,1251,1484,1508]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="42">MZUSP</collectionCode>
19128
</specimenCode>
(
<quantity id="8F62CBE34F5EFFD14B70FA0BFF11F9E0" metricMagnitude="0" metricUnit="m" metricValue="1.3655" pageId="41" pageNumber="42" unit="mm" value="1365.5">1, 365.5 mm</quantity>
SL), Rio 
<collectingRegion id="8A5EA8E44F5EFFD14F05FA28FE58F9EF" box="[310,394,1519,1544]" country="Brazil" name="Paraiba" pageId="41" pageNumber="42">Paraíba</collectingRegion>
do Sul, 
<location id="4D4530DD4F5EFFD14FDEFA28FD16F9EF" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5EFFD14FDEFA28FD16F9EF" box="[493,708,1519,1544]" country="Brazil" latitude="-22.65" longLatPrecision="1259" longitude="-45.0" name="Cachoeira Paulista" pageId="41" pageNumber="42" stateProvince="Paraiba">Cachoeira Paulista</location>
, c. 
<geoCoordinate id="2DAE00C14F5EFFD14CDEFA37FC99F9EF" box="[749,843,1520,1544]" direction="south" orientation="latitude" pageId="41" pageNumber="42" precision="925" value="-22.65">22°39’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5EFFD14D6BFA28FC13F9EF" box="[856,961,1519,1544]" direction="west" orientation="longitude" pageId="41" pageNumber="42" precision="925" value="-45.0">45°00’W</geoCoordinate>
</materialsCitation>
; 
<materialsCitation id="F8F26C5B4F5EFFD14DFCFA37FB18F9EF" ID-GBIF-Occurrence="1425222589" box="[975,1226,1520,1544]" collectingDate="1981-05" country="Brazil" location="Braga" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Braga" typeStatus="holotype">
J.T. 
<collectingRegion id="8A5EA8E44F5EFFD14A33FA37FB94F9EF" box="[1024,1094,1520,1544]" country="Portugal" name="Braga" pageId="41" pageNumber="42">Braga</collectingRegion>
, 
<collectingDate id="2C60B92E4F5EFFD14A60FA37FB18F9EF" box="[1107,1226,1520,1544]" pageId="41" pageNumber="42" value="1981-05">
<date id="3C2440C64F5EFFD14A60FA37FB18F9EF" box="[1107,1226,1520,1544]" pageId="41" pageNumber="42" value="1981-05">May 1981</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5EFFD14AE4FA37FC3FF9CA" ID-GBIF-Occurrence="1425222575" collectionCode="MZUSP" country="Brazil" latitude="-22.966667" location="Taubate" longLatPrecision="1258" longitude="-45.6" pageId="41" pageNumber="42" specimenCode="MZUSP 1553" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
<specimenCode id="183CCE7D4F5EFFD14AE4FA37FAA7F9EF" box="[1239,1397,1519,1544]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="42">
<collectionCode id="2E8BFEC34F5EFFD14AE4FA37FAE6F9EF" box="[1239,1332,1520,1544]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="42">MZUSP</collectionCode>
1553
</specimenCode>
(
<quantity id="8F62CBE34F5EFFD14BBAFA37FEDAF9CB" metricMagnitude="0" metricUnit="m" metricValue="1.1431" pageId="41" pageNumber="42" unit="mm" value="1143.1">1, 143.1 mm</quantity>
SL), 
<collectingRegion id="8A5EA8E44F5EFFD14F75F9D3FE68F9CA" box="[326,442,1556,1581]" country="Brazil" name="Sao Paulo" pageId="41" pageNumber="42">São Paulo</collectingRegion>
, 
<location id="4D4530DD4F5EFFD14FF6F9D3FDF2F9CA" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5EFFD14FF6F9D3FDF2F9CA" box="[453,544,1556,1581]" country="Brazil" latitude="-22.966667" longLatPrecision="1258" longitude="-45.6" name="Taubate" pageId="41" pageNumber="42" stateProvince="Sao Paulo">Taubaté</location>
, rio 
<collectingRegion id="8A5EA8E44F5EFFD14C63F9D3FD76F9CA" box="[592,676,1556,1581]" country="Brazil" name="Paraiba" pageId="41" pageNumber="42">Paraíba</collectingRegion>
do Sul, c. 
<geoCoordinate id="2DAE00C14F5EFFD14D28F9D3FCA8F9CA" box="[795,890,1556,1581]" direction="south" orientation="latitude" pageId="41" pageNumber="42" precision="925" value="-22.966667">22°58’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5EFFD14DB7F9D3FC3FF9CA" box="[900,1005,1556,1581]" direction="west" orientation="longitude" pageId="41" pageNumber="42" precision="925" value="-45.6">45°36’W</geoCoordinate>
</materialsCitation>
; 
<materialsCitation id="F8F26C5B4F5EFFD14DCAF9D2FC4AF9B7" ID-GBIF-Occurrence="1425222585" collectionCode="CAS" collectorName="E. Garbe" country="Brazil" location="Sao Paulo" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
<collectorName id="E56F03D04F5EFFD14DCAF9D2FB8CF9CA" box="[1017,1118,1556,1581]" pageId="41" pageNumber="42">E. Garbe</collectorName>
, 1911. 
<collectionCode id="2E8BFEC34F5EFFD14A9DF9D3FB36F9CB" box="[1198,1252,1556,1580]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="41" pageNumber="42">CAS</collectionCode>
68912 (
<quantity id="8F62CBE34F5EFFD14B77F9D3FF11F9A8" metricMagnitude="0" metricUnit="m" metricValue="1.1612" pageId="41" pageNumber="42" unit="mm" value="1161.2">1, 161.2 mm</quantity>
SL), “ 
<collectingRegion id="8A5EA8E44F5EFFD14F3FF9FFFE53F9B7" box="[268,385,1591,1616]" country="Brazil" name="Sao Paulo" pageId="41" pageNumber="42">São Paulo</collectingRegion>
” (no precise locality); collector unknown, 1902
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="0080358D4F5EFFD24EF4F99BFC07FCF3" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
<paragraph id="482566064F5EFFD24EF4F99BFC07FCF3" blockId="41.[151,1437,151,2013]" lastBlockId="42.[151,1437,151,789]" lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
<materialsCitation id="F8F26C5B4F5EFFD14EF4F99BFBC8F993" ID-GBIF-Occurrence="1425222577" box="[199,1050,1628,1653]" collectionCode="MCZ" location="Rio de Janeiro" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Rio de Janeiro">
<emphasis id="7AEEBA144F5EFFD14EF4F99BFD61F992" bold="true" box="[199,691,1628,1653]" pageId="41" pageNumber="42">
<collectingRegion id="8A5EA8E44F5EFFD14EF4F99BFEAAF992" box="[199,376,1628,1653]" country="Brazil" name="Rio de Janeiro" pageId="41" pageNumber="42">
<collectingRegion id="8A5EA8E44F5EFFD14EF4F99BFF23F992" box="[199,241,1628,1653]" country="Nigeria" name="Rivers" pageId="41" pageNumber="42">Rio</collectingRegion>
de Janeiro
</collectingRegion>
, rio 
<collectingRegion id="8A5EA8E44F5EFFD14F9CF99BFDDCF992" box="[431,526,1628,1653]" country="Brazil" name="Paraiba" pageId="41" pageNumber="42">Paraíba</collectingRegion>
do 
<collectingRegion id="8A5EA8E44F5EFFD14C0FF99BFDB1F992" box="[572,611,1628,1653]" country="Cameroon" name="South" pageId="41" pageNumber="42">Sul</collectingRegion>
basin:
</emphasis>
<collectionCode id="2E8BFEC34F5EFFD14C88F99BFD2AF993" box="[699,760,1628,1652]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42">MCZ</collectionCode>
21114 (
<quantity id="8F62CBE34F5EFFD14D64F99BFC37F993" box="[855,997,1628,1653]" metricMagnitude="0" metricUnit="m" metricValue="1.1353" pageId="41" pageNumber="42" unit="mm" value="1135.3">1, 135.3 mm</quantity>
SL)
</materialsCitation>
; 
<materialsCitation id="F8F26C5B4F5EFFD14A15F99BFBCEF970" ID-GBIF-Occurrence="1425222566" collectionCode="MCZ" latitude="-22.533333" location="Mendes" longLatPrecision="1259" longitude="-43.733334" municipality="Mendez" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Paraiba">
<collectionCode id="2E8BFEC34F5EFFD14A15F99BFBB1F993" box="[1062,1123,1628,1652]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42">MCZ</collectionCode>
21115 (4, 130.5– 
<quantity id="8F62CBE34F5EFFD14B19F99BFA4EF993" box="[1322,1436,1628,1653]" metricMagnitude="-1" metricUnit="m" metricValue="1.498" pageId="41" pageNumber="42" unit="mm" value="149.8">149.8 mm</quantity>
SL), “ 
<collectingMunicipality id="A841FC7C4F5EFFD14EE8F9B8FEE8F97F" box="[219,314,1663,1688]" pageId="41" pageNumber="42">Mendez</collectingMunicipality>
” [rio 
<collectingRegion id="8A5EA8E44F5EFFD14F4DF9B8FE00F97F" box="[382,466,1663,1688]" country="Brazil" name="Paraiba" pageId="41" pageNumber="42">Paraíba</collectingRegion>
do 
<collectingRegion id="8A5EA8E44F5EFFD14FCCF9B8FDF6F97F" box="[511,548,1663,1688]" country="Cameroon" name="South" pageId="41" pageNumber="42">Sul</collectingRegion>
or tributary at 
<location id="4D4530DD4F5EFFD14CE7F9B8FCE2F97F" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5EFFD14CE7F9B8FCE2F97F" box="[724,816,1663,1688]" latitude="-22.533333" longLatPrecision="1259" longitude="-43.733334" municipality="Mendez" name="Mendes" pageId="41" pageNumber="42" stateProvince="Paraiba">Mendes</location>
, 
<geoCoordinate id="2DAE00C14F5EFFD14D0FF947FC44F97F" box="[828,918,1664,1688]" direction="south" orientation="latitude" pageId="41" pageNumber="42" precision="925" value="-22.533333">22°32'S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5EFFD14D97F947FBC7F970" box="[932,1045,1664,1688]" direction="west" orientation="longitude" pageId="41" pageNumber="42" precision="925" value="-43.733334">43°44' W</geoCoordinate>
]
</materialsCitation>
; 
<materialsCitation id="F8F26C5B4F5EFFD14A1AF947FEB4F907" ID-GBIF-Occurrence="1425222594" collectionCode="MCZ" collectorName="C. F. Hartt &amp; E. Copeland" county="Muriahe" latitude="-21.133333" location="Muriae" longLatPrecision="1265" longitude="-42.383335" municipality="rio Muriae" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Paraiba">
<collectorName id="E56F03D04F5EFFD14A1AF947FB48F970" box="[1065,1178,1664,1688]" pageId="41" pageNumber="42">C.F. Hartt</collectorName>
&amp; 
<collectorName id="E56F03D04F5EFFD14AF2F947FA82F97F" box="[1217,1360,1663,1688]" pageId="41" pageNumber="42">E. Copeland</collectorName>
, 1865. 
<collectionCode id="2E8BFEC34F5EFFD14EA4F963FF06F95B" box="[151,212,1700,1724]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42">MCZ</collectionCode>
21110 (7, 154.0– 
<quantity id="8F62CBE34F5EFFD14F9EF963FDF7F95B" box="[429,549,1700,1725]" metricMagnitude="-1" metricUnit="m" metricValue="1.785" pageId="41" pageNumber="42" unit="mm" value="178.5">178.5 mm</quantity>
SL): “ 
<collectingCounty id="A1441E8A4F5EFFD14C4FF963FD0DF95A" box="[636,735,1700,1725]" pageId="41" pageNumber="42">Muriahé</collectingCounty>
” [
<collectingMunicipality id="A841FC7C4F5EFFD14CCDF963FCADF95A" box="[766,895,1700,1725]" pageId="41" pageNumber="42">rio Muriaé</collectingMunicipality>
at 
<location id="4D4530DD4F5EFFD14D9CF963FBD1F95A" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5EFFD14D9CF963FBD1F95A" box="[943,1027,1700,1725]" county="Muriahe" latitude="-21.133333" longLatPrecision="1265" longitude="-42.383335" municipality="rio Muriae" name="Muriae" pageId="41" pageNumber="42" stateProvince="Paraiba">Muriaé</location>
, 
<geoCoordinate id="2DAE00C14F5EFFD14A26F963FBB2F95B" box="[1045,1120,1700,1724]" direction="south" orientation="latitude" pageId="41" pageNumber="42" precision="925" value="-21.133333">21°8'S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5EFFD14A40F963FB3BF95B" box="[1139,1257,1700,1724]" direction="west" orientation="longitude" pageId="41" pageNumber="42" precision="925" value="-42.383335">42° 23'W</geoCoordinate>
]; 
<collectorName id="E56F03D04F5EFFD14B30F963FAABF95B" box="[1283,1401,1700,1724]" pageId="41" pageNumber="42">C.F. Hartt</collectorName>
&amp; 
<collectorName id="E56F03D04F5EFFD14EA4F900FECFF907" box="[151,285,1735,1760]" pageId="41" pageNumber="42">E.Copeland</collectorName>
, 1865
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5EFFD14F47F90FFC60F8E2" ID-GBIF-Occurrence="1425222576" collectionCode="MCZ" collectorName="Campos dos Goytacazes &amp; C. F. Hartt &amp; E. Copeland" latitude="-21.716667" location="Campos" longLatPrecision="1262" longitude="-41.366665" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Rio">
<collectionCode id="2E8BFEC34F5EFFD14F47F90FFE62F907" box="[372,432,1736,1760]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="41" pageNumber="42">MCZ</collectionCode>
21117 (5, 148.8– 
<quantity id="8F62CBE34F5EFFD14C48F900FD23F938" box="[635,753,1735,1760]" metricMagnitude="-1" metricUnit="m" metricValue="2.316" pageId="41" pageNumber="42" unit="mm" value="231.6">231.6 mm</quantity>
SL), “ 
<location id="4D4530DD4F5EFFD14D0CF90FFC4DF907" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5EFFD14D0CF90FFC4DF907" box="[831,927,1736,1760]" latitude="-21.716667" longLatPrecision="1262" longitude="-41.366665" name="Campos" pageId="41" pageNumber="42" stateProvince="Rio">Campos</location>
” [
<collectingRegion id="8A5EA8E44F5EFFD14D8AF900FC36F907" box="[953,996,1735,1760]" country="Nigeria" name="Rivers" pageId="41" pageNumber="42">Rio</collectingRegion>
Muriaé at 
<collectorName id="E56F03D04F5EFFD14A54F90FFA52F907" box="[1127,1408,1735,1760]" pageId="41" pageNumber="42">Campos dos Goytacazes</collectorName>
, 
<quantity id="8F62CBE34F5EFFD14BBDF90FFF06F8E2" metricMagnitude="3" metricUnit="m" metricValue="4.828032" pageId="41" pageNumber="42" unit="mi" value="3.0">3 miles</quantity>
from town, 
<geoCoordinate id="2DAE00C14F5EFFD14F6CF92BFE6CF8E3" box="[351,446,1772,1796]" direction="south" orientation="latitude" pageId="41" pageNumber="42" precision="925" value="-21.716667">21°43’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5EFFD14FF9F92BFDE5F8E3" box="[458,567,1772,1796]" direction="west" orientation="longitude" pageId="41" pageNumber="42" precision="925" value="-41.366665">41°22’W</geoCoordinate>
]; 
<collectorName id="E56F03D04F5EFFD14C79F92BFD6AF8E3" box="[586,696,1772,1796]" pageId="41" pageNumber="42">C.F. Hartt</collectorName>
&amp; 
<collectorName id="E56F03D04F5EFFD14CEDF92AFCBEF8E2" box="[734,876,1772,1797]" pageId="41" pageNumber="42">E. Copeland</collectorName>
, 1865
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5EFFD14D8DF92BFB8AF8CF" ID-GBIF-Occurrence="1425222592" collectingDate="1865" collectingDateMax="1866" collectingDateMin="1865" collectionCode="CAS" collectorName="Bourget" location="Paraiba" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Paraiba">
<collectionCode id="2E8BFEC34F5EFFD14D8DF92BFC26F8E3" box="[958,1012,1772,1796]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="41" pageNumber="42">CAS</collectionCode>
68832 (1, 258.0 mm SL), rio 
<collectingRegion id="8A5EA8E44F5EFFD14B7BF92BFA4EF8E2" box="[1352,1436,1772,1797]" country="Brazil" name="Paraiba" pageId="41" pageNumber="42">Paraíba</collectingRegion>
do 
<collectingRegion id="8A5EA8E44F5EFFD14E88F8C8FF32F8CF" box="[187,224,1807,1832]" country="Cameroon" name="South" pageId="41" pageNumber="42">Sul</collectingRegion>
drainage [precise locality uncertain]; 
<collectorName id="E56F03D04F5EFFD14CBEF8D7FD38F8CF" box="[653,746,1808,1832]" pageId="41" pageNumber="42">Bourget</collectorName>
, date not specified [
<collectingDate id="2C60B92E4F5EFFD14DF8F8C8FB80F8CF" box="[971,1106,1807,1832]" pageId="41" pageNumber="42" value="1865" valueMax="1866">
<date id="3C2440C64F5EFFD14DF8F8C8FB80F8CF" box="[971,1106,1807,1832]" pageId="41" pageNumber="42" value="1865" valueMax="1866">1865–1866</date>
</collectingDate>
]
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5EFFD14A57F8D7FAD5F897" ID-GBIF-Occurrence="1425222572" collectingDate="1990-02-22" collectionCode="MNRJ" collectorName="Valao de Sao Domingos &amp; D. F. Moraes Jr." county="Itaperuna" latitude="-21.266666" location="rio Muriae" longLatPrecision="1264" longitude="-41.766666" municipality="Valao de Sao Domingos" pageId="41" pageNumber="42" specimenCode="MNRJ 18246, MNRJ 11226" specimenCount="1" stateProvince="Paraiba">
<specimenCode id="183CCE7D4F5EFFD14A57F8D7FB2EF8CF" box="[1124,1276,1807,1832]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="41" pageNumber="42">
<collectionCode id="2E8BFEC34F5EFFD14A57F8D7FB7DF8C0" box="[1124,1199,1808,1831]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="41" pageNumber="42">MNRJ</collectionCode>
18246
</specimenCode>
(
<quantity id="8F62CBE34F5EFFD14B3CF8D7FA4FF8C0" box="[1295,1437,1808,1832]" metricMagnitude="0" metricUnit="m" metricValue="1.1987999999999999" pageId="41" pageNumber="42" unit="mm" value="1198.8">1, 198.8 mm</quantity>
SL), 
<collectingCounty id="A1441E8A4F5EFFD14EE3F8F2FEEFF8AB" box="[208,317,1845,1868]" pageId="41" pageNumber="42">Itaperuna</collectingCounty>
, 
<collectingMunicipality id="A841FC7C4F5EFFD14F7BF8F3FD8BF8AA" box="[328,601,1844,1869]" pageId="41" pageNumber="42">
<collectorName id="E56F03D04F5EFFD14F7BF8F3FD8BF8AA" box="[328,601,1844,1869]" pageId="41" pageNumber="42">Valão de São Domingos</collectorName>
</collectingMunicipality>
(tributary of 
<location id="4D4530DD4F5EFFD14CC2F8F3FCBEF8AA" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5EFFD14CC2F8F3FCBEF8AA" box="[753,876,1844,1869]" county="Itaperuna" latitude="-21.266666" longLatPrecision="1264" longitude="-41.766666" municipality="Valao de Sao Domingos" name="rio Muriae" pageId="41" pageNumber="42" stateProvince="Paraiba">rio Muriaé</location>
, rio 
<collectingRegion id="8A5EA8E44F5EFFD14DADF8F3FC20F8AA" box="[926,1010,1844,1869]" country="Brazil" name="Paraiba" pageId="41" pageNumber="42">Paraíba</collectingRegion>
do 
<collectingRegion id="8A5EA8E44F5EFFD14A2DF8F3FB91F8AA" box="[1054,1091,1844,1869]" country="Cameroon" name="South" pageId="41" pageNumber="42">Sul</collectingRegion>
basin, 
<geoCoordinate id="2DAE00C14F5EFFD14AA6F8F3FB3DF8AA" box="[1173,1263,1844,1869]" direction="south" orientation="latitude" pageId="41" pageNumber="42" precision="925" value="-21.266666">21°16'S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5EFFD14AC8F8F3FAB2F8AA" box="[1275,1376,1844,1869]" direction="west" orientation="longitude" pageId="41" pageNumber="42" precision="925" value="-41.766666">41°46'W</geoCoordinate>
; 
<collectorName id="E56F03D04F5EFFD14B5EF8F2FEDCF888" pageId="41" pageNumber="42">D.F. Moraes Jr.</collectorName>
<emphasis id="7AEEBA144F5EFFD14F25F89EFE82F897" box="[278,336,1879,1904]" italics="true" pageId="41" pageNumber="42">et al.</emphasis>
, 
<collectingDate id="2C60B92E4F5EFFD14F6CF89FFE21F897" box="[351,499,1879,1904]" pageId="41" pageNumber="42" value="1990-02-22">
<date id="3C2440C64F5EFFD14F6CF89FFE21F897" box="[351,499,1879,1904]" pageId="41" pageNumber="42" value="1990-02-22">22 Feb 1990</date>
</collectingDate>
. 
<specimenCode id="183CCE7D4F5EFFD14FCCF89FFD45F897" box="[511,663,1879,1904]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="41" pageNumber="42">
<collectionCode id="2E8BFEC34F5EFFD14FCCF89FFD98F888" box="[511,586,1880,1903]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="41" pageNumber="42">MNRJ</collectionCode>
11226
</specimenCode>
(
<quantity id="8F62CBE34F5EFFD14C98F89FFCE9F888" box="[683,827,1880,1904]" metricMagnitude="0" metricUnit="m" metricValue="1.1018" pageId="41" pageNumber="42" unit="mm" value="1101.8">1, 101.8 mm</quantity>
SL), Itaperuna, córrego do Toiama, trib
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5EFFD14B20F890FD9CF85F" ID-GBIF-Occurrence="1425222591" collectingDate="1982-01-15" collectorName="Bom Jesus do Itabapoana-Santo Antonio de Padua &amp; D. F. Moraes Jr. &amp; G. W. Nunan &amp; L. C. Alvarenga" latitude="-21.233334" location="Rio" longLatPrecision="21" longitude="-41.925003" pageId="41" pageNumber="42" specimenCount="1" stateProvince="Rio">
<collectingRegion id="8A5EA8E44F5EFFD14B20F890FAE9F897" box="[1299,1339,1879,1904]" country="Nigeria" name="Rivers" pageId="41" pageNumber="42">Rio</collectingRegion>
Muriaé, road BR-393 (
<collectorName id="E56F03D04F5EFFD14F05F8BBFCA0F872" box="[310,882,1916,1941]" pageId="41" pageNumber="42">Bom Jesus do Itabapoana-Santo Antônio de Pádua</collectorName>
), c. 
<geoCoordinate id="2DAE00C14F5EFFD14DADF8BBFBDFF873" box="[926,1037,1916,1940]" direction="south" orientation="latitude" pageId="41" pageNumber="42" precision="15" value="-21.233334">21°14'0''S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5EFFD14A2AF8BBFB73F872" box="[1049,1185,1916,1941]" direction="west" orientation="longitude" pageId="41" pageNumber="42" precision="15" value="-41.925003">41°55'30''W</geoCoordinate>
; 
<collectorName id="E56F03D04F5EFFD14A9EF8BAFA88F873" box="[1197,1370,1917,1940]" pageId="41" pageNumber="42">D.F. Moraes Jr.</collectorName>
, 
<collectorName id="E56F03D04F5EFFD14B56F8BBFF30F85F" pageId="41" pageNumber="42">G.W. Nunan</collectorName>
&amp; 
<collectorName id="E56F03D04F5EFFD14F35F867FE65F85E" box="[262,439,1952,1977]" pageId="41" pageNumber="42">L.C. Alvarenga</collectorName>
, 
<date id="3C2440C64F5EFFD14FF1F867FD9CF85F" box="[450,590,1952,1977]" pageId="41" pageNumber="42" value="1982-01-15">
<collectingDate id="2C60B92E4F5EFFD14FF1F867FD9CF85F" box="[450,590,1952,1977]" pageId="41" pageNumber="42" value="1982-01-15">15 Jan 1982</collectingDate>
</date>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5EFFD14C6BF866FD17F83A" ID-GBIF-Occurrence="1425222583" collectionCode="MNRJ" collectorName="Fazenda da Salgada" location="Itaperuna" pageId="41" pageNumber="42" specimenCode="MNRJ 4761" specimenCount="1" stateProvince="A">
<specimenCode id="183CCE7D4F5EFFD14C6BF866FD38F85E" box="[600,746,1952,1977]" collectionCode="MNRJ" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="41" pageNumber="42">
<collectionCode id="2E8BFEC34F5EFFD14C6BF866FD71F85F" box="[600,675,1953,1976]" country="Brazil" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="41" pageNumber="42">MNRJ</collectionCode>
4761
</specimenCode>
(
<quantity id="8F62CBE34F5EFFD14CC1F867FCA8F85F" box="[754,890,1952,1977]" metricMagnitude="0" metricUnit="m" metricValue="1.1382999999999999" pageId="41" pageNumber="42" unit="mm" value="1138.3">1, 138.3 mm</quantity>
SL), 
<collectorName id="E56F03D04F5EFFD14D8AF867FB4AF85E" box="[953,1176,1952,1977]" pageId="41" pageNumber="42">Fazenda da Salgada</collectorName>
and São José, between 
<location id="4D4530DD4F5EFFD14EA4F802FED3F83B" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5EFFD14EA4F802FED3F83B" box="[151,257,1989,2012]" name="Itaperuna" pageId="41" pageNumber="42" stateProvince="A">Itaperuna</location>
and Miracema; 
<collectingRegion id="8A5EA8E44F5EFFD14F8FF803FE06F83B" box="[444,468,1988,2012]" country="Nigeria" name="Rivers" pageId="41" pageNumber="42">A</collectingRegion>
.L. Carvalho, no date
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5EFFD24CE2F803FC32FF57" ID-GBIF-Occurrence="1425222584" collectingDate="1941-07" collectionCode="MZUSP" collectorName="O. Pinto" lastPageId="42" lastPageNumber="43" latitude="-21.483334" location="rio Muriae" longLatPrecision="1264" longitude="-41.6" municipality="Cardoso Moreira" pageId="41" pageNumber="42" specimenCode="MZUSP 3043" specimenCount="1" stateProvince="Paraiba">
<specimenCode id="183CCE7D4F5EFFD14CE2F803FCBDF83B" box="[721,879,1988,2012]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="42">
<collectionCode id="2E8BFEC34F5EFFD14CE2F803FCFCF83B" box="[721,814,1988,2012]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="41" pageNumber="42">MZUSP</collectionCode>
3043
</specimenCode>
(
<quantity id="8F62CBE34F5EFFD14DB0F803FBC0F83B" box="[899,1042,1988,2013]" metricMagnitude="0" metricUnit="m" metricValue="1.2368" pageId="41" pageNumber="42" unit="mm" value="1236.8">1, 236.8 mm</quantity>
SL), 
<collectingMunicipality id="A841FC7C4F5EFFD14A67F803FAC5F83A" box="[1108,1303,1988,2013]" pageId="41" pageNumber="42">Cardoso Moreira</collectingMunicipality>
, 
<location id="4D4530DD4F5EFFD14B10F803FA4EF83A" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5EFFD14B10F803FA4EF83A" box="[1315,1436,1988,2013]" latitude="-21.483334" longLatPrecision="1264" longitude="-41.6" municipality="Cardoso Moreira" name="rio Muriae" pageId="41" pageNumber="42" stateProvince="Paraiba">rio Muriaé</location>
(tributary of rio 
<collectingRegion id="8A5EA8E44F5DFFD24F67FF50FE7AFF57" box="[340,424,151,176]" country="Brazil" name="Paraiba" pageId="42" pageNumber="43">Paraíba</collectingRegion>
do 
<collectingRegion id="8A5EA8E44F5DFFD24FEBFF50FDD3FF57" box="[472,513,151,176]" country="Cameroon" name="South" pageId="42" pageNumber="43">Sul</collectingRegion>
), 
<geoCoordinate id="2DAE00C14F5DFFD24C24FF5FFDA4FF57" box="[535,630,152,176]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-21.483334">21°29’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5DFFD24CB7FF50FD3FFF57" box="[644,749,151,176]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-41.6">41°36’W</geoCoordinate>
; 
<collectorName id="E56F03D04F5DFFD24CCFFF5FFC8DFF57" box="[764,863,151,176]" pageId="42" pageNumber="43">O. Pinto</collectorName>
, 
<collectingDate id="2C60B92E4F5DFFD24D5EFF50FC32FF57" box="[877,992,151,176]" pageId="42" pageNumber="43" value="1941-07">
<date id="3C2440C64F5DFFD24D5EFF50FC32FF57" box="[877,992,151,176]" pageId="42" pageNumber="43" value="1941-07">July 1941</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5DFFD24DDDFF5FFBE7FF33" ID-GBIF-Occurrence="1425222598" collectionCode="MZUSP" collectorName="Campos dos Goytacazes" latitude="-21.7" location="Rio" longLatPrecision="1262" longitude="-42.516666" pageId="42" pageNumber="43" specimenCode="MZUSP 103045" specimenCount="1" stateProvince="Rio">
<specimenCode id="183CCE7D4F5DFFD24DDDFF5FFB7BFF57" box="[1006,1193,151,176]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="43">
<collectionCode id="2E8BFEC34F5DFFD24DDDFF5FFB99FF57" box="[1006,1099,152,176]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="43">MZUSP</collectionCode>
<date id="3C2440C64F5DFFD24A66FF50FB7BFF57" box="[1109,1193,151,176]" pageId="42" pageNumber="43" value="45-10-30">103045</date>
</specimenCode>
(4, 194.5–259.0 mm SL): 
<collectorName id="E56F03D04F5DFFD24EE1FF7BFE3EFF33" box="[210,492,188,213]" pageId="42" pageNumber="43">Campos dos Goytacazes</collectorName>
, 
<collectingRegion id="8A5EA8E44F5DFFD24FCAFF7BFDF0FF32" box="[505,546,188,213]" country="Nigeria" name="Rivers" pageId="42" pageNumber="43">Rio</collectingRegion>
Muriaé, near its mouth, c. 
<geoCoordinate id="2DAE00C14F5DFFD24D6CFF7BFC6FFF33" box="[863,957,188,212]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-21.7">21°42’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5DFFD24DF8FF7BFBE7FF33" box="[971,1077,188,212]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-42.516666">42°31’W</geoCoordinate>
</materialsCitation>
; 
<materialsCitation id="F8F26C5B4F5DFFD24A71FF7BFAF7FF1F" ID-GBIF-Occurrence="1425222588" collectionCode="MNHN" latitude="-21.616667" location="Rio Grande" longLatPrecision="1263" longitude="-41.816666" pageId="42" pageNumber="43" specimenCode="MNHN 1909-214" specimenCount="1" stateProvince="Rio Grande">
collector not especified, 2002. 
<specimenCode id="183CCE7D4F5DFFD24EA4FF27FE8DFF1F" box="[151,351,224,248]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="42" pageNumber="43">
<collectionCode id="2E8BFEC34F5DFFD24EA4FF27FF3EFF10" box="[151,236,224,247]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="42" pageNumber="43">MNHN</collectionCode>
1909-214
</specimenCode>
(
<quantity id="8F62CBE34F5DFFD24F42FF27FE2FFF10" box="[369,509,224,248]" metricMagnitude="0" metricUnit="m" metricValue="1.2877" pageId="42" pageNumber="43" unit="mm" value="1287.7">1, 287.7 mm</quantity>
SL): 
<collectingRegion id="8A5EA8E44F5DFFD24C0EFF18FD6DFF1F" box="[573,703,223,248]" country="Puerto Rico" name="Rio Grande" pageId="42" pageNumber="43">
<collectingRegion id="8A5EA8E44F5DFFD24C0EFF18FDB4FF1F" box="[573,614,223,248]" country="Nigeria" name="Rivers" pageId="42" pageNumber="43">Rio</collectingRegion>
Grande
</collectingRegion>
(tributary of rio 
<collectingRegion id="8A5EA8E44F5DFFD24D48FF18FC1DFF1F" box="[891,975,223,248]" country="Brazil" name="Paraiba" pageId="42" pageNumber="43">Paraíba</collectingRegion>
do 
<collectingRegion id="8A5EA8E44F5DFFD24DC9FF18FBF0FF1F" box="[1018,1058,223,248]" country="Cameroon" name="South" pageId="42" pageNumber="43">Sul</collectingRegion>
), c. 
<geoCoordinate id="2DAE00C14F5DFFD24A62FF27FB7DFF1F" box="[1105,1199,224,248]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-21.616667">21°37’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5DFFD24A88FF27FAF7FF1F" box="[1211,1317,224,248]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-41.816666">41°49’W</geoCoordinate>
</materialsCitation>
; C. Jobert, c. 1878. MZUSP 79088 (2, 141.7– 
<quantity id="8F62CBE34F5DFFD24C05FEC3FD7EFEFB" box="[566,684,260,284]" metricMagnitude="-1" metricUnit="m" metricValue="1.483" pageId="42" pageNumber="43" unit="mm" value="148.3">148.3 mm</quantity>
SL); 
<materialsCitation id="F8F26C5B4F5DFFD24CC5FEC3FBAFFEA7" ID-GBIF-Occurrence="1425222602" collectionCode="MZUSP" county="Fish" location="Itaocara" municipality="Projeto" pageId="42" pageNumber="43" specimenCode="MZUSP 79091" specimenCount="1" stateProvince="Paraiba">
<specimenCode id="183CCE7D4F5DFFD24CC5FEC3FC77FEFB" box="[758,933,260,284]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="43">
<collectionCode id="2E8BFEC34F5DFFD24CC5FEC3FC81FEFB" box="[758,851,260,284]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="43">MZUSP</collectionCode>
79091
</specimenCode>
(
<quantity id="8F62CBE34F5DFFD24D8EFEC3FB81FEFB" box="[957,1107,260,285]" metricMagnitude="0" metricUnit="m" metricValue="1.1465" pageId="42" pageNumber="43" unit="mm" value="1146.5">1, 146.5 mm</quantity>
SL): 
<collectingCounty id="A1441E8A4F5DFFD24AAEFEC3FB1FFEFA" box="[1181,1229,260,285]" pageId="42" pageNumber="43">Fish</collectingCounty>
hatchery, 
<collectingMunicipality id="A841FC7C4F5DFFD24B78FEC3FA4EFEFA" box="[1355,1436,260,285]" pageId="42" pageNumber="43">Projeto</collectingMunicipality>
<taxonomicName id="8F9A1D854F5DFFD24EA4FEE0FED3FEA7" box="[151,257,295,320]" family="Bryconidae" genus="Megalobrycon" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="42" pageNumber="184" phylum="Chordata" rank="species" species="piabanha">Piabanha</taxonomicName>
, 
<location id="4D4530DD4F5DFFD24F3EFEEFFEB5FED8" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5DFFD24F3EFEEFFEB5FED8" box="[269,359,296,319]" county="Fish" municipality="Projeto" name="Itaocara" pageId="42" pageNumber="43" stateProvince="Paraiba">Itaocara</location>
(stocked from specimens collected at the middle rio 
<collectingRegion id="8A5EA8E44F5DFFD24DFAFEE0FBCFFEA7" box="[969,1053,295,320]" country="Brazil" name="Paraiba" pageId="42" pageNumber="43">Paraíba</collectingRegion>
do 
<collectingRegion id="8A5EA8E44F5DFFD24A79FEE0FBA1FEA7" box="[1098,1139,295,320]" country="Cameroon" name="South" pageId="42" pageNumber="43">Sul</collectingRegion>
)
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5DFFD24ABBFEE0FD72FE83" ID-GBIF-Occurrence="1425222580" collectionCode="MCZ" location="Rio de Janeiro" pageId="42" pageNumber="43" specimenCount="1" stateProvince="Rio de Janeiro">
<emphasis id="7AEEBA144F5DFFD24ABBFEE0FEECFE82" bold="true" pageId="42" pageNumber="43">
<collectingRegion id="8A5EA8E44F5DFFD24ABBFEE0FAE9FEA7" box="[1160,1339,295,320]" country="Brazil" name="Rio de Janeiro" pageId="42" pageNumber="43">
<collectingRegion id="8A5EA8E44F5DFFD24ABBFEE0FB60FEA7" box="[1160,1202,295,320]" country="Nigeria" name="Rivers" pageId="42" pageNumber="43">Rio</collectingRegion>
de Janeiro
</collectingRegion>
, coastal river systems:
</emphasis>
<collectionCode id="2E8BFEC34F5DFFD24F76FE8BFE50FE83" box="[325,386,332,356]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="42" pageNumber="43">MCZ</collectionCode>
21112 (
<quantity id="8F62CBE34F5DFFD24FD3FE8BFDBFFE83" box="[480,621,332,356]" metricMagnitude="0" metricUnit="m" metricValue="1.2322" pageId="42" pageNumber="43" unit="mm" value="1232.2">1, 232.2 mm</quantity>
SL)
</materialsCitation>
, 
<materialsCitation id="F8F26C5B4F5DFFD24C9FFE8BFBFFFE6F" ID-GBIF-Occurrence="1425222574" collectorName="D. Pedro &amp; D. Bourget &amp; April" latitude="-22.883333" location="Fazenda Santa Cruz" longLatPrecision="1258" longitude="-43.716667" municipality="Arroio Fundo" pageId="42" pageNumber="43" specimenCount="1" stateProvince="Santa Cruz">
<collectingRegion id="8A5EA8E44F5DFFD24C9FFE8BFCF5FE83" box="[684,807,332,356]" country="Bolivia" name="Santa Cruz" pageId="42" pageNumber="43">Santa Cruz</collectingRegion>
[
<collectingRegion id="8A5EA8E44F5DFFD24D05FE8BFC60FE82" box="[822,946,332,357]" country="Puerto Rico" name="Rio Grande" pageId="42" pageNumber="43">rio Grande</collectingRegion>
(
<collectingMunicipality id="A841FC7C4F5DFFD24DF3FE8BFBB2FE82" box="[960,1120,332,357]" pageId="42" pageNumber="43">Arroio Fundo</collectingMunicipality>
), in urban 
<collectingRegion id="8A5EA8E44F5DFFD24AE4FE8BFAAFFE82" box="[1239,1405,332,357]" country="Brazil" name="Rio de Janeiro" pageId="42" pageNumber="43">
<collectingRegion id="8A5EA8E44F5DFFD24AE4FE8BFB2DFE82" box="[1239,1279,332,357]" country="Nigeria" name="Rivers" pageId="42" pageNumber="43">Rio</collectingRegion>
de Janeiro
</collectingRegion>
, at 
<location id="4D4530DD4F5DFFD24EA4FEA8FEA8FE6F" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5DFFD24EA4FEA8FEA8FE6F" box="[151,378,367,392]" latitude="-22.883333" longLatPrecision="1258" longitude="-43.716667" municipality="Arroio Fundo" name="Fazenda Santa Cruz" pageId="42" pageNumber="43" stateProvince="Santa Cruz">
Fazenda 
<collectingRegion id="8A5EA8E44F5DFFD24ECFFEB7FEA8FE6F" box="[252,378,368,392]" country="Bolivia" name="Santa Cruz" pageId="42" pageNumber="43">Santa Cruz</collectingRegion>
</location>
, 
<geoCoordinate id="2DAE00C14F5DFFD24FB5FEA8FE32FE6F" box="[390,480,367,392]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-22.883333">22°53'S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5DFFD24FDFFEB7FD87FE6F" box="[492,597,368,392]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-43.716667">43°43'W</geoCoordinate>
]; 
<collectorName id="E56F03D04F5DFFD24C54FEB7FD19FE6F" box="[615,715,367,392]" pageId="42" pageNumber="43">D. Pedro</collectorName>
II, leg. 
<collectorName id="E56F03D04F5DFFD24D16FEB7FC76FE6F" box="[805,932,368,392]" pageId="42" pageNumber="43">D. Bourget</collectorName>
, 
<date id="3C2440C64F5DFFD24D9CFEA8FBFFFE6F" box="[943,1069,367,392]" pageId="42" pageNumber="43" value="1865-04">
<collectorName id="E56F03D04F5DFFD24D9CFEA8FC38FE6F" box="[943,1002,367,392]" pageId="42" pageNumber="43">April</collectorName>
1865
</date>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5DFFD24A0BFEB7FED1FE37" ID-GBIF-Occurrence="1425222579" collectingDate="2005-10" collectionCode="MZUSP" collectorName="Lagoa de Juturnaiba" latitude="-22.566668" location="Lagoa de Juturnaiba" longLatPrecision="1259" longitude="-42.3" municipality="Silva Jardim" pageId="42" pageNumber="43" specimenCode="MZUSP 88515" specimenCount="1" stateProvince="Rio">
<specimenCode id="183CCE7D4F5DFFD24A0BFEB7FB31FE6F" box="[1080,1251,367,392]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="43">
<collectionCode id="2E8BFEC34F5DFFD24A0BFEB7FB47FE6F" box="[1080,1173,368,392]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="43">MZUSP</collectionCode>
88515
</specimenCode>
(2, 235.0–247.0 mm SL), 
<collectingMunicipality id="A841FC7C4F5DFFD24F33FE53FE5DFE4A" box="[256,399,404,429]" pageId="42" pageNumber="43">Silva Jardim</collectingMunicipality>
, 
<collectingRegion id="8A5EA8E44F5DFFD24FA9FE53FE10FE4A" box="[410,450,404,429]" country="Nigeria" name="Rivers" pageId="42" pageNumber="43">Rio</collectingRegion>
São João, near mouth at 
<collectorName id="E56F03D04F5DFFD24CEAFE52FC6CFE4A" box="[729,958,404,429]" pageId="42" pageNumber="43">
<location id="4D4530DD4F5DFFD24CEAFE52FC6CFE4A" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5DFFD24CEAFE52FC6CFE4A" box="[729,958,404,429]" latitude="-22.566668" longLatPrecision="1259" longitude="-42.3" municipality="Silva Jardim" name="Lagoa de Juturnaiba" pageId="42" pageNumber="43" stateProvince="Rio">Lagoa de Juturnaíba</location>
</collectorName>
, 
<geoCoordinate id="2DAE00C14F5DFFD24DFAFE53FBF1FE4B" box="[969,1059,404,428]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-22.566668">22°34'S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5DFFD24A1DFE53FB40FE4B" box="[1070,1170,404,428]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-42.3">42°18'W</geoCoordinate>
; collector not specified, 
<collectingDate id="2C60B92E4F5DFFD24EA4FE7FFED1FE37" box="[151,259,439,464]" pageId="42" pageNumber="43" value="2005-10">
<date id="3C2440C64F5DFFD24EA4FE7FFED1FE37" box="[151,259,439,464]" pageId="42" pageNumber="43" value="2005-10">Oct 2005</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5DFFD24F3EFE7FFAF4FE37" ID-GBIF-Occurrence="1425222586" box="[269,1318,439,464]" collectionCode="MZUSP" collectorName="Campos dos Goytacazes" latitude="-21.783333" location="rio Imbe" longLatPrecision="1262" longitude="-41.55" pageId="42" pageNumber="43" specimenCode="MZUSP 103029" specimenCount="1" stateProvince="Rio">
<specimenCode id="183CCE7D4F5DFFD24F3EFE7FFE15FE37" box="[269,455,440,464]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="43">
<collectionCode id="2E8BFEC34F5DFFD24F3EFE7FFEB8FE37" box="[269,362,440,464]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="43">MZUSP</collectionCode>
<date id="3C2440C64F5DFFD24F40FE7FFE15FE37" box="[371,455,440,464]" pageId="42" pageNumber="43" value="29-10-30">103029</date>
</specimenCode>
(1, 291.0 mm SL), 
<collectorName id="E56F03D04F5DFFD24C94FE7FFC6FFE37" box="[679,957,439,464]" pageId="42" pageNumber="43">Campos dos Goytacazes</collectorName>
, 
<location id="4D4530DD4F5DFFD24DFAFE70FBF9FE37" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5DFFD24DFAFE70FBF9FE37" box="[969,1067,439,464]" latitude="-21.783333" longLatPrecision="1262" longitude="-41.55" name="rio Imbe" pageId="42" pageNumber="43" stateProvince="Rio">rio Imbé</location>
, c. 
<geoCoordinate id="2DAE00C14F5DFFD24A60FE7FFB63FE37" box="[1107,1201,440,464]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-21.783333">21°47’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5DFFD24A8EFE7FFAF4FE37" box="[1213,1318,440,464]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-41.55">41°33’W</geoCoordinate>
</materialsCitation>
; 
<materialsCitation id="F8F26C5B4F5DFFD24B00FE7FFC71FDFF" ID-GBIF-Occurrence="1425222605" collectionCode="MZUSP" collectorName="G. Sousa &amp; Campos dos Goytacazes" county="Fish" latitude="-21.783333" location="Projeto" longLatPrecision="1262" longitude="-41.55" municipality="Itaocara" pageId="42" pageNumber="43" specimenCode="MZUSP 79092" specimenCount="1" stateProvince="Rio">
<collectorName id="E56F03D04F5DFFD24B00FE7FFA4AFE37" box="[1331,1432,440,464]" pageId="42" pageNumber="43">G. Sousa</collectorName>
, 2000. 
<specimenCode id="183CCE7D4F5DFFD24ED3FE1BFE5FFE13" box="[224,397,476,500]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="43">
<collectionCode id="2E8BFEC34F5DFFD24ED3FE1BFEEFFE13" box="[224,317,476,500]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="43">MZUSP</collectionCode>
79092
</specimenCode>
(
<quantity id="8F62CBE34F5DFFD24F90FE1BFDE6FE13" box="[419,564,476,500]" metricMagnitude="0" metricUnit="m" metricValue="1.1872" pageId="42" pageNumber="43" unit="mm" value="1187.2">1, 187.2 mm</quantity>
SL), 
<collectingCounty id="A1441E8A4F5DFFD24C4AFE1BFD7BFE12" box="[633,681,476,501]" pageId="42" pageNumber="43">Fish</collectingCounty>
hatchery at 
<collectingMunicipality id="A841FC7C4F5DFFD24D0FFE1AFC44FE13" box="[828,918,477,500]" pageId="42" pageNumber="43">Itaocara</collectingMunicipality>
(
<location id="4D4530DD4F5DFFD24D94FE1BFC28FE12" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5DFFD24D94FE1BFC28FE12" box="[935,1018,476,501]" county="Fish" latitude="-21.783333" longLatPrecision="1262" longitude="-41.55" municipality="Itaocara" name="Projeto" pageId="42" pageNumber="43" stateProvince="Rio">Projeto</location>
<taxonomicName id="8F9A1D854F5DFFD24A37FE1BFBBFFE12" box="[1028,1133,476,501]" family="Bryconidae" genus="Megalobrycon" higherTaxonomySource="CoL" kingdom="Animalia" order="Characiformes" pageId="42" pageNumber="184" phylum="Chordata" rank="species" species="piabanha">Piabanha</taxonomicName>
) (stocked from specimens collected at 
<collectorName id="E56F03D04F5DFFD24F13FDC7FDE4FDFF" box="[288,566,511,536]" pageId="42" pageNumber="43">Campos dos Goytacazes</collectorName>
, 
<location id="4D4530DD4F5DFFD24C71FE38FD74FDFF" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5DFFD24C71FE38FD74FDFF" box="[578,678,511,536]" county="Fish" latitude="-21.783333" longLatPrecision="1262" longitude="-41.55" municipality="Itaocara" name="rio Imbe" pageId="42" pageNumber="43" stateProvince="Rio">rio Imbé</location>
, c. 
<geoCoordinate id="2DAE00C14F5DFFD24CFEFDC7FCF5FDFF" box="[717,807,512,536]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-21.783333">21°47'S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5DFFD24D07FDC7FC48FDFF" box="[820,922,512,536]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-41.55">41°33'W</geoCoordinate>
)
</materialsCitation>
; G. Souza, no date. MZUSP 2091 (5, 140.3– 
<quantity id="8F62CBE34F5DFFD24EA4FDE3FED8FDDB" box="[151,266,548,572]" metricMagnitude="-1" metricUnit="m" metricValue="2.203" pageId="42" pageNumber="43" unit="mm" value="220.3">220.3 mm</quantity>
SL); CAS 11817 (1, 228.0 mm SL); 
<materialsCitation id="F8F26C5B4F5DFFD24C87FDE3FE38FD87" ID-GBIF-Occurrence="1425222587" collectingDate="1911-12" collectionCode="CAS" collectorName="E. Garbe" latitude="-21.95" location="Lagoa Feia" longLatPrecision="1262" longitude="-41.366665" municipality="Quicama" pageId="42" pageNumber="43" specimenCount="1" stateProvince="Rio">
<collectionCode id="2E8BFEC34F5DFFD24C87FDE3FD38FDDB" box="[692,746,548,572]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="42" pageNumber="43">CAS</collectionCode>
11816 (1, 213.0 mm SL); 
<collectingMunicipality id="A841FC7C4F5DFFD24A28FDE3FB51FDDA" box="[1051,1155,548,573]" pageId="42" pageNumber="43">Quiçamã</collectingMunicipality>
, 
<location id="4D4530DD4F5DFFD24AA3FDE2FAC3FDDA" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5DFFD24AA3FDE2FAC3FDDA" box="[1168,1297,548,573]" latitude="-21.95" longLatPrecision="1262" longitude="-41.366665" municipality="Quicama" name="Lagoa Feia" pageId="42" pageNumber="43" stateProvince="Rio">Lagoa Feia</location>
, c. 
<geoCoordinate id="2DAE00C14F5DFFD24B0AFDE3FA45FDDA" box="[1337,1431,548,573]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-21.95">21°57’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5DFFD24EA4FD8FFED2FD87" box="[151,256,584,608]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-41.366665">41°22’W</geoCoordinate>
; 
<collectorName id="E56F03D04F5DFFD24F38FD8FFEA3FD87" box="[267,369,583,608]" pageId="42" pageNumber="43">E. Garbe</collectorName>
, 
<collectingDate id="2C60B92E4F5DFFD24F4FFD8FFE38FD87" box="[380,490,584,608]" pageId="42" pageNumber="43" value="1911-12">
<date id="3C2440C64F5DFFD24F4FFD8FFE38FD87" box="[380,490,584,608]" pageId="42" pageNumber="43" value="1911-12">Dec 1911</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5DFFD24FC6FD8FFD29FD63" ID-GBIF-Occurrence="1425222581" collectingDate="2004-09-03" collectionCode="MZUSP" collectorName="Campos dos Goytacazes &amp; O. T. Oyakawa" location="Lagoa Feia" pageId="42" pageNumber="43" specimenCode="MZUSP 107280" specimenCount="1" stateProvince="Rio">
<specimenCode id="183CCE7D4F5DFFD24FC6FD8FFD7CFD87" box="[501,686,584,608]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="43">
<collectionCode id="2E8BFEC34F5DFFD24FC6FD8FFD80FD87" box="[501,594,584,608]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="43">MZUSP</collectionCode>
107280
</specimenCode>
(1, 260.0 mm SL): 
<collectorName id="E56F03D04F5DFFD24DB9FD8FFB4DFD87" box="[906,1183,583,608]" pageId="42" pageNumber="43">Campos dos Goytacazes</collectorName>
, fish market (probably from 
<location id="4D4530DD4F5DFFD24EE4FDAAFE8EFD62" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5DFFD24EE4FDAAFE8EFD62" box="[215,348,620,645]" name="Lagoa Feia" pageId="42" pageNumber="43" stateProvince="Rio">Lagoa Feia</location>
); 
<collectorName id="E56F03D04F5DFFD24F41FDABFDC5FD62" box="[370,535,620,645]" pageId="42" pageNumber="43">O.T. Oyakawa</collectorName>
<emphasis id="7AEEBA144F5DFFD24C13FDAAFD89FD62" box="[544,603,620,645]" italics="true" pageId="42" pageNumber="43">et al.</emphasis>
, 
<collectingDate id="2C60B92E4F5DFFD24C5FFDABFD29FD63" box="[620,763,620,644]" pageId="42" pageNumber="43" value="2004-09-03">
<date id="3C2440C64F5DFFD24C5FFDABFD29FD63" box="[620,763,620,644]" pageId="42" pageNumber="43" value="2004-09-03">3 Sept 2004</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5DFFD24D3BFDABFAD7FD4F" ID-GBIF-Occurrence="1425222573" collectionCode="MCZ" county="rio Itabapoana" latitude="-21.283333" location="Itabapoana" longLatPrecision="1264" longitude="-40.983334" municipality="Itabapoana" pageId="42" pageNumber="43" specimenCount="1" stateProvince="Rio de Janeiro">
<emphasis id="7AEEBA144F5DFFD24D3BFDABFA4EFD62" bold="true" box="[776,1436,620,645]" pageId="42" pageNumber="43">
<collectingRegion id="8A5EA8E44F5DFFD24D3BFDABFC69FD62" box="[776,955,620,645]" country="Brazil" name="Rio de Janeiro" pageId="42" pageNumber="43">
<collectingRegion id="8A5EA8E44F5DFFD24D3BFDABFCE0FD62" box="[776,818,620,645]" country="Nigeria" name="Rivers" pageId="42" pageNumber="43">Rio</collectingRegion>
de Janeiro
</collectingRegion>
/ 
<collectingRegion id="8A5EA8E44F5DFFD24DF7FDABFBA6FD62" box="[964,1140,620,645]" country="Brazil" name="Espirito Santo" pageId="42" pageNumber="43">Espírito Santo</collectingRegion>
, 
<collectingCounty id="A1441E8A4F5DFFD24AB0FDABFAEAFD62" box="[1155,1336,620,645]" pageId="42" pageNumber="43">rio Itabapoana</collectingCounty>
system:
</emphasis>
<collectionCode id="2E8BFEC34F5DFFD24EA4FD57FF06FD4F" box="[151,212,656,680]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="42" pageNumber="43">MCZ</collectionCode>
21116 (9, 144.0– 
<quantity id="8F62CBE34F5DFFD24FABFD57FDDBFD40" box="[408,521,656,680]" metricMagnitude="-1" metricUnit="m" metricValue="1.918" pageId="42" pageNumber="43" unit="mm" value="191.8">191.8 mm</quantity>
SL), 
<collectingMunicipality id="A841FC7C4F5DFFD24C7BFD48FD11FD4F" box="[584,707,655,680]" pageId="42" pageNumber="43">Itabapoana</collectingMunicipality>
[
<collectingRegion id="8A5EA8E44F5DFFD24CE3FD48FD29FD4F" box="[720,763,655,680]" country="Nigeria" name="Rivers" pageId="42" pageNumber="43">Rio</collectingRegion>
Itabapoana at 
<location id="4D4530DD4F5DFFD24DACFD48FBCFFD4F" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5DFFD24DACFD48FBCFFD4F" box="[927,1053,655,680]" county="rio Itabapoana" latitude="-21.283333" longLatPrecision="1264" longitude="-40.983334" municipality="Itabapoana" name="Itabapoana" pageId="42" pageNumber="43" stateProvince="Rio de Janeiro">Itabapoana</location>
, 
<geoCoordinate id="2DAE00C14F5DFFD24A1BFD57FB54FD4F" box="[1064,1158,656,680]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-21.283333">21°17’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5DFFD24AA1FD48FB2DFD4F" box="[1170,1279,655,680]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-40.983334">40°59’W</geoCoordinate>
]
</materialsCitation>
; 
<materialsCitation id="F8F26C5B4F5DFFD24B22FD57FEDFFD17" ID-GBIF-Occurrence="1425222599" collectingDate="2002-08-14" collectionCode="MZUSP" collectorName="C. F. Hartt &amp; E. Copeland &amp; F. Daudt" latitude="-21.283333" location="rio Itabapoana" longLatPrecision="1264" longitude="-40.983334" pageId="42" pageNumber="43" specimenCode="MZUSP 79087" specimenCount="1" stateProvince="Rio de Janeiro">
<collectorName id="E56F03D04F5DFFD24B22FD57FAADFD40" box="[1297,1407,656,680]" pageId="42" pageNumber="43">C.F. Hartt</collectorName>
&amp; 
<collectorName id="E56F03D04F5DFFD24EA4FD72FEF4FD2A" box="[151,294,692,717]" pageId="42" pageNumber="43">E. Copeland</collectorName>
, 1865. 
<specimenCode id="183CCE7D4F5DFFD24F48FD73FDFAFD2B" box="[379,552,692,716]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="43">
<collectionCode id="2E8BFEC34F5DFFD24F48FD73FE0AFD2B" box="[379,472,692,716]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="42" pageNumber="43">MZUSP</collectionCode>
79087
</specimenCode>
(3, 193.5–231.0 mm SL): 
<location id="4D4530DD4F5DFFD24D6FFD73FBD0FD2A" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5DFFD24D6FFD73FBD0FD2A" box="[860,1026,692,717]" latitude="-21.283333" longLatPrecision="1264" longitude="-40.983334" name="rio Itabapoana" pageId="42" pageNumber="43" stateProvince="Rio de Janeiro">rio Itabapoana</location>
, c. 
<geoCoordinate id="2DAE00C14F5DFFD24A1FFD73FB5BFD2B" box="[1068,1161,692,716]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="925" value="-21.283333">21°17’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5DFFD24AABFD73FAD3FD2A" box="[1176,1281,692,717]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="925" value="-40.983334">40°59’W</geoCoordinate>
; 
<collectorName id="E56F03D04F5DFFD24B3DFD72FAA1FD2A" box="[1294,1395,692,717]" pageId="42" pageNumber="43">F. Daudt</collectorName>
, 
<collectingDate id="2C60B92E4F5DFFD24BB3FD73FEDFFD17" pageId="42" pageNumber="43" value="2002-08-14">
<date id="3C2440C64F5DFFD24BB3FD73FEDFFD17" pageId="42" pageNumber="43" value="2002-08-14">14 Aug 2002</date>
</collectingDate>
</materialsCitation>
. 
<materialsCitation id="F8F26C5B4F5DFFD24F2AFD10FC07FCF3" ID-GBIF-Occurrence="1425222619" collectingDate="2011-02" collectionCode="ZUEC" collectorName="Itamarati de Minas &amp; D. M. Rosa" county="Itamarati de Minas" latitude="-21.476667" location="rio Pomba" longLatPrecision="21" longitude="-42.859997" municipality="rio Novo" pageId="42" pageNumber="43" specimenCode="ZUEC 6384" specimenCount="1" stateProvince="Minas Gerais">
<emphasis id="7AEEBA144F5DFFD24F2AFD10FCC6FD17" bold="true" box="[281,788,727,752]" pageId="42" pageNumber="43">
<collectingRegion id="8A5EA8E44F5DFFD24F2AFD10FE6DFD17" box="[281,447,727,752]" country="Brazil" name="Minas Gerais" pageId="42" pageNumber="43">Minas Gerais</collectingRegion>
, rio 
<collectingRegion id="8A5EA8E44F5DFFD24FC9FD10FD8BFD17" box="[506,601,727,752]" country="Brazil" name="Paraiba" pageId="42" pageNumber="43">Paraíba</collectingRegion>
do 
<collectingRegion id="8A5EA8E44F5DFFD24CBAFD10FD62FD17" box="[649,688,727,752]" country="Cameroon" name="South" pageId="42" pageNumber="43">Sul</collectingRegion>
system:
</emphasis>
<specimenCode id="183CCE7D4F5DFFD24D2DFD1FFC7AFD17" box="[798,936,727,752]" collectionCode="ZUEC" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="42" pageNumber="43">
<collectionCode id="2E8BFEC34F5DFFD24D2DFD1FFCB5FD17" box="[798,871,728,752]" country="Brazil" httpUri="http://grbio.org/cool/9pz2-d68t" name="Museu de Zoologia da Universidade Estadual de Campinas" pageId="42" pageNumber="43">ZUEC</collectionCode>
6384
</specimenCode>
(1, 251.0 mm SL): 
<collectorName id="E56F03D04F5DFFD24AA0FD10FABDFD17" box="[1171,1391,727,752]" pageId="42" pageNumber="43">
<collectingCounty id="A1441E8A4F5DFFD24AA0FD10FABDFD17" box="[1171,1391,727,752]" pageId="42" pageNumber="43">Itamarati de Minas</collectingCounty>
</collectorName>
, 
<collectingMunicipality id="A841FC7C4F5DFFD24B4EFD10FF07FCF3" pageId="42" pageNumber="43">rio Novo</collectingMunicipality>
(trib. 
<location id="4D4530DD4F5DFFD24F29FD3BFE40FCF2" LSID="urn:lsid:plazi:treatment:C033D7104F54FFD24EA4FE8BFC07FCF3:4D4530DD4F5DFFD24F29FD3BFE40FCF2" box="[282,402,764,789]" county="Itamarati de Minas" latitude="-21.476667" longLatPrecision="21" longitude="-42.859997" municipality="rio Novo" name="rio Pomba" pageId="42" pageNumber="43" stateProvince="Minas Gerais">rio Pomba</location>
), 
<geoCoordinate id="2DAE00C14F5DFFD24F96FD3BFDE2FCF2" box="[421,560,764,789]" direction="south" orientation="latitude" pageId="42" pageNumber="43" precision="15" value="-21.476667">21°28’36’’S</geoCoordinate>
, 
<geoCoordinate id="2DAE00C14F5DFFD24C0FFD3BFD03FCF2" box="[572,721,764,789]" direction="west" orientation="longitude" pageId="42" pageNumber="43" precision="15" value="-42.859997">42°51’36’’W</geoCoordinate>
; 
<collectorName id="E56F03D04F5DFFD24CEEFD3AFC88FCF3" box="[733,858,765,788]" pageId="42" pageNumber="43">D.M. Rosa</collectorName>
, 
<collectingDate id="2C60B92E4F5DFFD24D56FD3BFC03FCF3" box="[869,977,764,789]" pageId="42" pageNumber="43" value="2011-02">
<date id="3C2440C64F5DFFD24D56FD3BFC03FCF3" box="[869,977,764,789]" pageId="42" pageNumber="43" value="2011-02">Feb 2011</date>
</collectingDate>
.
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>