250 lines
30 KiB
XML
250 lines
30 KiB
XML
<document ID-CLB-Dataset="9646" ID-DOI="http://dx.doi.org/10.3897/zookeys.984.53587" ID-GBIF-Dataset="9d291585-91c0-45e5-94d9-ee28795291ba" ID-PMC="PMC7658186" ID-Pensoft-Pub="1313-2970-984-83" ID-Pensoft-UUID="1448149CA44B51ED9EE57DC3A924C430" ID-PubMed="33223870" ID-ZooBank="D8B945AE620949578BB1D96DDBB5AA44" ModsDocID="1313-2970-984-83" checkinTime="1604561368050" checkinUser="pensoft" docAuthor="Irwin, Michael E., Winterton, Shaun L. & Metz, Mark A." docDate="2020" docId="B662223F207950EDB60EF236FB0F33BD" docLanguage="en" docName="ZooKeys 984: 83-132" docOrigin="ZooKeys 984" docSource="http://dx.doi.org/10.3897/zookeys.984.53587" docTitle="Calophytus Irwin & Winterton & Metz 2020, gen. nov." docType="treatment" docUuid="CB02979A-20AF-4C4E-BFB1-B8D05668523C" docUuidSource="ZooBank" docVersion="6" id="1448149CA44B51ED9EE57DC3A924C430" lastPageNumber="83" masterDocId="1448149CA44B51ED9EE57DC3A924C430" masterDocTitle="New stiletto flies from New Caledonia (Therevidae, Agapophytinae)" masterLastPageNumber="132" masterPageNumber="83" pageNumber="83" updateTime="1732742644444" updateUser="ExternalLinkService">
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<mods:mods id="F2FF8767E81A5B78D35F3C27B807BB50" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="60BC1F9982AA85B39F5BFE7BEE7563C7">
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<mods:title id="CBD4FF8FD0177EFADD170A365F607D5B">New stiletto flies from New Caledonia (Therevidae, Agapophytinae)</mods:title>
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<mods:name id="2210F6F888A220823CF73E3A5F50CFCF" type="personal">
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<mods:roleTerm id="2679214D95EFA2E25DED8E3AB581EDB7">Author</mods:roleTerm>
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<mods:namePart id="68A7825173CAA20BC2607B6E75748B9D">Irwin, Michael E.</mods:namePart>
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<mods:affiliation id="0C2166E0C54509771A73AE7373215A2A">Emeritus, University of Illinois, Urbana-Champaign, Illinois, USA & University of Arizona, Tucson, USA</mods:affiliation>
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</mods:name>
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<mods:name id="50BB47757BEA0CD4F6CB46802BB71475" type="personal">
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<mods:roleTerm id="C05405070E644BEB8004E4F8BEC2110C">Author</mods:roleTerm>
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<mods:namePart id="773FBCB0E25B20C05EF10072CAA35835">Winterton, Shaun L.</mods:namePart>
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<mods:affiliation id="5E3786FC1E312A4C76327F00EA13F347">California State Collection of Arthropods, California Department of Food & Agriculture, Sacramento, California, USA</mods:affiliation>
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<mods:nameIdentifier id="B83C44BB240E8AB5DF00866D5742EB8B" type="email">wintertonshaun@gmail.com</mods:nameIdentifier>
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<mods:roleTerm id="4645A6D69BCF1682901BE622BE0F9971">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7DE0606527740141FB08611F882F4285">Metz, Mark A.</mods:namePart>
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<mods:affiliation id="22D2A9E986B21D5F0AA5BDE5520C43A1">USDA, ARS, Systematic Entomology Laboratory, Beltsville, Maryland, USA</mods:affiliation>
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<mods:nameIdentifier id="8AB70EBE5FF42C1F8434871D4DD24FF6" type="orcid">https://orcid.org/0000-0002-3535-535X</mods:nameIdentifier>
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<mods:title id="B08D7FA488F3D01FDEEF5EE1EE89361A">ZooKeys</mods:title>
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<mods:part id="E236B21D15A8447859651D6DEEDB79BE">
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<mods:date id="69548B210733CE01AC5A71452215C2A8">2020</mods:date>
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<mods:detail id="FEF6029168AED3305BFFBFF42020CE8D" type="volume">
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<mods:number id="1CCDC4580E61AAF5213B986ED44568C0">984</mods:number>
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<mods:start id="33DF3FE1B8C29FB4B47928C74F406D7F">83</mods:start>
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<mods:end id="05D65EA2924E9E91FA9706DB7B430D66">132</mods:end>
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<mods:url id="36BB2D79F9A1EB804D573914AED8A17A">http://dx.doi.org/10.3897/zookeys.984.53587</mods:url>
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<mods:classification id="4FD5790D9C6123BA382A0989C70F2ED3">journal article</mods:classification>
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<mods:identifier id="718C01495661ACDEEA3E9BBEC1E29BBC" type="DOI">http://dx.doi.org/10.3897/zookeys.984.53587</mods:identifier>
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<mods:identifier id="E742294E827CC01D6671D962C16DA61C" type="Pensoft-Pub">1313-2970-984-83</mods:identifier>
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<treatment id="B662223F207950EDB60EF236FB0F33BD" ID-GBIF-Taxon="168756101" LSID="urn:lsid:zoobank.org:act:CB02979A-20AF-4C4E-BFB1-B8D05668523C" httpUri="http://treatment.plazi.org/id/B662223F207950EDB60EF236FB0F33BD" lastPageNumber="83" pageId="0" pageNumber="83" scope_family="Therevidae" scope_subFamily="Agapophytinae">
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<subSubSection id="0C7A18AB8706136979B3F8805FFD6B2E" pageId="0" pageNumber="83" type="nomenclature">
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<paragraph id="9956894A0186A18AED4814A15B0F6CEF" pageId="0" pageNumber="83">
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<taxonomicName id="E2F01EBEC07AE8F55404ADA7B511812E" LSID="B662223F-2079-50ED-B60E-F236FB0F33BD" authority="Irwin & Winterton & Metz, 2020" authorityName="Irwin & Winterton & Metz" authorityYear="2020" family="Agapophytinae" genus="Calophytus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Calophytus" order="Therevidae" pageId="0" pageNumber="83" rank="genus" status="gen. nov.">Calophytus</taxonomicName>
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<taxonomicNameLabel id="8FB2AC5A900B7E0E5F8A26A978968A86" pageId="0" pageNumber="83">gen. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="00CA61A226B6381484518AAB22814980" pageId="0" pageNumber="83" type="type species">
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<paragraph id="D203BEF6418EA757715B3E58B368E1CE" pageId="0" pageNumber="83">Type species.</paragraph>
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<paragraph id="5A91BDA6DE138CC462F941560A093C52" pageId="0" pageNumber="83">
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<taxonomicName id="BDEDACD98CE930F8BB38EF1FF742B247" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus chazeaui" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="chazeaui">
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<emphasis id="31520E0E2349F04BC7611AFE494A5D5D" italics="true" pageId="0" pageNumber="83">Calophytus chazeaui</emphasis>
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</taxonomicName>
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sp. nov., here designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="CBFBADB9C89CF8B001963D62BDD07C96" pageId="0" pageNumber="83" type="diagnosis">
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<paragraph id="DEB3B8E75E9A3B6636FF323346F3640E" pageId="0" pageNumber="83">Diagnosis.</paragraph>
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<paragraph id="616BB3699E180B670BEB91F46CCB7BE0" pageId="0" pageNumber="83">Antennae elongate, longer than head, narrow cylindrical; frons wider than ocellar tubercle at narrowest point with only slight sexual dimorphism; dorsocentral macrosetae absent; velutum patches absent on femora; single posteroventral macroseta present midway along hind femur; wing with hyaline areas often free of microtrichia; distinct stripe of dark microtrichia bisecting cell br along wing fold; male genitalia with gonocoxites without velutum patch and medial atrium (gonocoxites proximal medially).</paragraph>
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</subSubSection>
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<subSubSection id="9FC2365F499A0D271012C8C55625F97B" pageId="0" pageNumber="83" type="description">
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<paragraph id="506733E982ACED7E4014405139DBA2F7" pageId="0" pageNumber="83">Description.</paragraph>
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<paragraph id="F914090F38156C78D26A94178DB7F89C" pageId="0" pageNumber="83">
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Antenna longer than head; flagellum cylindrical, slightly tapered distally, shorter than combined scape and pedicel length, rarely longer (in
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<taxonomicName id="BC52BD13EC64BD877013390E03F2CD3E" lsidName="C. webbi" pageId="0" pageNumber="83" rank="species" species="webbi">
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<emphasis id="36B7F32AED37DC45A446806E1A9D55F3" italics="true" pageId="0" pageNumber="83">C. webbi</emphasis>
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</taxonomicName>
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sp. nov.); scape cylindrical, usually elongate; head shape in profile with length and height subequal; frons glossy, slightly protruding anterior to eye around base of antennae; frons wider than ocellar tubercle at narrowest point, only slight sexual dimorphism in frons width with space between eyes slightly narrower in male; parafacial without setae, postocular macrosetae in both sexes arranged in a single row dorsally, rarely scattered dorsomedially on occiput (in
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<taxonomicName id="32721A2A71CDBC0FC6D644C2856F01D7" lsidName="C. grandiosus" pageId="0" pageNumber="83" rank="species" species="grandiosus">
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<emphasis id="A3F48D79FB2EA06F9FD52761CBBA0E5E" italics="true" pageId="0" pageNumber="83">C. grandiosus</emphasis>
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</taxonomicName>
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sp. nov.); prosternum without setae; fore and hind femoral velutum patches absent; femoral macrosetae absent except for single posteroventral macroseta present midway along hind femur; posterior surface of mid coxa without setae; hind femur and tibia relatively longer than that of fore and mid legs; post-spiracular setae absent; scutal chaetotaxy (pairs): notopleural (np), 1-2; supra alar (sa), 1; post alar (pa), 1; dorsocentral (dc), 0; scutellar (sc), 1; wing cell m3 open to margin; wing vein R2+3 smoothly curved or straight to wing margin, wing with markings faint to strongly banded, hyaline areas often free of microtrichia; distinct stripe of dark microtrichia bisecting cell br along wing fold; abdominal tergite II with all setae uniform and regular in length. Male genitalia with dorsal apodeme of aedeagus
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<normalizedToken id="7CC1BD96332B1BEC3D023E99DDD639FC" originalValue="‘T’">'T'</normalizedToken>
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-shaped, distiphallus narrow, straight, ventral apodeme forked; gonocoxite with velutum patch absent, posteromedial margins proximal to each other; inner gonocoxal process (igp) present and articulated; ventral lobe small, rounded apically. Female genitalia with tergite VIII elongate, quadrangular, anteromedial process narrow; tergite VII lacking anteromedial process; acanthophorite setae as two sets (A1 & A2), A1 enlarged and rounded apically, A2 series prominent; sternite VIII emarginate posteromedially, flattened; spermathecal sac present, not lobed; three sac-like spermathecae present, joined to spermathecal sac duct near junction with bursa copulatrix.
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</paragraph>
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</subSubSection>
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<subSubSection id="33D6241E7163FD3BBC4B4B40CA921895" pageId="0" pageNumber="83" type="etymology">
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<paragraph id="5FFA1918F9386E705802DE1DFE2A9B01" pageId="0" pageNumber="83">Etymology.</paragraph>
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<paragraph id="714F522345F1448553C1CF690A7DA751" pageId="0" pageNumber="83">
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Derived from the Greek
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<emphasis id="F17E1C983C691579D1CA1F0E0D89C47C" italics="true" pageId="0" pageNumber="83">kallos</emphasis>
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, beauty; and
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<emphasis id="DAC50630F63F3B1C277527FDE8395106" italics="true" pageId="0" pageNumber="83">phyton</emphasis>
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, plant, and relating to the charismatic appearance of the flies and the natural areas where they are collected in New Caledonia. The name also associates this genus with New Caledonia via the Proto-Celtic root of
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<emphasis id="B346F2B605E75E0ACE6E0D973CBD68F3" italics="true" pageId="0" pageNumber="83">kal</emphasis>
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, hard, pertaining to the Roman name for the original provenance of Caledones in Britannia. Gender is masculine.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="874C3F4842FAABBA6055A29DD704FCEF" pageId="0" pageNumber="83" type="comments">
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<paragraph id="B0807CEA7819D2401A0803F6CCDAE71A" pageId="0" pageNumber="83">Comments.</paragraph>
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<paragraph id="487417405615B9DA17F060D011653AF8" pageId="0" pageNumber="83">
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All species of
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<taxonomicName id="7D81100BAF77B060DAD1A8FA51110313" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="DD072FA62228396A791150E62E0E6777" italics="true" pageId="0" pageNumber="83">Calophytus</emphasis>
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</taxonomicName>
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gen. nov. are medium to large stiletto flies, with heads mostly glossy with striking patches of silver pubescence on the parafacial and pleuron, banded wings and antennae longer than the head. Distinctive to members of
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<taxonomicName id="531407BEBB6D1AC1E8F8470E1993B208" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="6A6E937FC4FD09CEA6D39D028C743361" italics="true" pageId="0" pageNumber="83">Calophytus</emphasis>
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</taxonomicName>
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gen. nov. is the presence of a single posteroventral macroseta midway along the hind femur and a stripe of microtrichia along the wing fold bisecting an otherwise glabrous wing cell br. These two characteristics are also found in the genus
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<taxonomicName id="9FAD30EBB9B92A186A6A51B9606C18C1" authorityName="Krober" authorityYear="1928" class="Insecta" family="Therevidae" genus="Squamopygia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamopygia" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="FBFBF3D3DD1C97427D19F8C55CBDB616" italics="true" pageId="0" pageNumber="83">Squamopygia</emphasis>
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</taxonomicName>
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from Australia and Papua New Guinea and appear unique to these two genera amongst all
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<taxonomicName id="5F260A32164F085CC197FF8BA248EC06" genus="Therevidae" lsidName="Therevidae" pageId="0" pageNumber="83" rank="genus">Therevidae</taxonomicName>
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; they are notably absent in
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<taxonomicName id="1D217C46426FFC3C01C591B0AA650346" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Jeanchazeauia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Jeanchazeauia" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="1D271D77E1996BDD51E238BF64FCE7FF" italics="true" pageId="0" pageNumber="83">Jeanchazeauia</emphasis>
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</taxonomicName>
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gen. nov. While
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<bibRefCitation id="70F4FC66CABB9C4ACB35CE5B85CC9A33" DOI="https://doi.org/10.1111/syen.12147" author="Winterton, SL" journalOrPublisher="Systematic Entomology" pageId="0" pageNumber="83" pagination="144 - 161" refId="B18" refString="Winterton, SL, Hardy, NB, Gaimari, SD, Hauser, M, Hill, HN, Holston, KC, Irwin, ME, Lambkin, CL, Metz, MA, Turco, F, Webb, D, Yang, L, Yeates, DK, Wiegmann, BM, 2016. The phylogeny of stiletto flies (Diptera: Therevidae). Systematic Entomology 41: 144 - 161, DOI: https://doi.org/10.1111/syen.12147" title="The phylogeny of stiletto flies (Diptera: Therevidae)." url="https://doi.org/10.1111/syen.12147" volume="41" year="2016">Winterton et al. (2016)</bibRefCitation>
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recovered a clear sister-group relationship between
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<taxonomicName id="3DF6252656118B4CF2A40D06BFB89A3C" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="03655A39AD597A7D41FDF44D7B82CE3E" italics="true" pageId="0" pageNumber="83">Calophytus</emphasis>
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</taxonomicName>
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gen. nov. and
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<taxonomicName id="A511F38FF70808D3F2C35FD971082FBB" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Jeanchazeauia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Jeanchazeauia" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="297E801ED2EEB886282F483F25BF6E08" italics="true" pageId="0" pageNumber="83">Jeanchazeauia</emphasis>
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</taxonomicName>
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gen. nov. (included in their study as undescribed genus
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<normalizedToken id="F699610D1550B68D4CE358CB0143E541" originalValue="‘NC’">'NC'</normalizedToken>
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) (Fig.
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<figureCitation id="6BF63DA2D3507B3A9036E2901E7FAD59" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phylogenetic placement of Calophytus gen. nov. and Jeanchazeauia gen. nov. in Agapophytinae based on supermatrix analysis of DNA sequence data. Figure modified after Winterton et al. (2016: fig. 3)." figureDoi="10.3897/zookeys.984.53587.figure4" httpUri="https://binary.pensoft.net/fig/471320" pageId="0" pageNumber="83">4</figureCitation>
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), they did not include the rarely collected
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<taxonomicName id="7E89964B40E5A3688E350DD8094B23A6" authorityName="Krober" authorityYear="1928" class="Insecta" family="Therevidae" genus="Squamopygia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamopygia" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="BDA6286A672ED8C6E9BA85F3D63E4CDF" italics="true" pageId="0" pageNumber="83">Squamopygia</emphasis>
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</taxonomicName>
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. The two aforementioned characters found only in
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<taxonomicName id="58B5A339035DBD83254248BE271B4D38" authorityName="Krober" authorityYear="1928" class="Insecta" family="Therevidae" genus="Squamopygia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamopygia" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="38223911FC644FA1992185BE87963120" italics="true" pageId="0" pageNumber="83">Squamopygia</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="2C80C3A6BCF7C0B0020918A15908FCFE" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="1EC1C7E7E3597CEF7EF633D167E97920" italics="true" pageId="0" pageNumber="83">Calophytus</emphasis>
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</taxonomicName>
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gen. nov. also support a likely sister-group relationship between these two genera.
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<taxonomicName id="BD8D51D8DBB00E147DBD756DDEB6623F" authorityName="Krober" authorityYear="1928" class="Insecta" family="Therevidae" genus="Squamopygia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamopygia" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
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<emphasis id="3F4ED782CF6D604B662E3BD9636FB859" italics="true" pageId="0" pageNumber="83">Squamopygia</emphasis>
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</taxonomicName>
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is a monotypic genus with a single species from Northern Australia; two undescribed species are known from southern Australia and Papua New Guinea.
|
||
<taxonomicName id="52004661F583462F52D455F51F4BC6E7" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="30447463A7B6C06252DEAE663D0C4117" italics="true" pageId="0" pageNumber="83">Calophytus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. can be separated from
|
||
<taxonomicName id="AAE93AA2EC2B42753409851BADD9701B" authorityName="Krober" authorityYear="1928" class="Insecta" family="Therevidae" genus="Squamopygia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamopygia" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="552C349FD96CC6EFE43D59B89FC4AFA6" italics="true" pageId="0" pageNumber="83">Squamopygia</emphasis>
|
||
</taxonomicName>
|
||
by the absence of subapical anteroventral macrosetae on the hind femur (present in
|
||
<taxonomicName id="D868571B79FDE016F628CE6FCA32E8A3" authorityName="Krober" authorityYear="1928" class="Insecta" family="Therevidae" genus="Squamopygia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamopygia" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="662A27F7B22152032ACEE314C8182EC3" italics="true" pageId="0" pageNumber="83">Squamopygia</emphasis>
|
||
</taxonomicName>
|
||
), absence of dorsocentral macrosetae (present in
|
||
<taxonomicName id="E8972723FD6F83287626C6B154FC1244" authorityName="Krober" authorityYear="1928" class="Insecta" family="Therevidae" genus="Squamopygia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamopygia" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="77108638F684EFF7AD633D39CF6F84BA" italics="true" pageId="0" pageNumber="83">Squamopygia</emphasis>
|
||
</taxonomicName>
|
||
) and by the width of the male frons. The sexual dimorphism in male frons width is more pronounced in
|
||
<taxonomicName id="08759EBC4ADDDEB64106BBDC040A1472" authorityName="Krober" authorityYear="1928" class="Insecta" family="Therevidae" genus="Squamopygia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamopygia" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="4A7294E8D4A5B8E6EBE1B7EBF1E6C9CA" italics="true" pageId="0" pageNumber="83">Squamopygia</emphasis>
|
||
</taxonomicName>
|
||
with the eyes being contiguous. In
|
||
<taxonomicName id="9E5D6AA756A26B70B79197BE1271AFE5" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E585E09352E9AD9A1AE57BC90D0E31AD" italics="true" pageId="0" pageNumber="83">Calophytus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov. (and
|
||
<taxonomicName id="2DD7E608ABA80BFDF3F2B72BADC8A805" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Jeanchazeauia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Jeanchazeauia" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="D9CE0149772FF12817B4D4CD147B9365" italics="true" pageId="0" pageNumber="83">Jeanchazeauia</emphasis>
|
||
</taxonomicName>
|
||
gen. nov.) the male frons is only slightly narrower than that of the female, with the frons wider than the ocellar tubercle in both sexes.
|
||
<taxonomicName id="74B8CC0FDCF2AB241BA5E75A0088D43D" authorityName="Krober" authorityYear="1928" class="Insecta" family="Therevidae" genus="Squamopygia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamopygia" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="91EAB51393D217CEB01835344F3353A7" italics="true" pageId="0" pageNumber="83">Squamopygia</emphasis>
|
||
</taxonomicName>
|
||
also has banded wings similar to
|
||
<taxonomicName id="29E7EF8F72C3255C0461E1FBEF900499" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="1EF126777284E2B64E1559ABDF540786" italics="true" pageId="0" pageNumber="83">Calophytus</emphasis>
|
||
</taxonomicName>
|
||
gen. nov., but the hyaline areas of the wing membrane retain extensive pale microtrichia. The male and female genitalia of
|
||
<taxonomicName id="1646ED863E708C6D26DBC0136F2C3643" authorityName="Krober" authorityYear="1928" class="Insecta" family="Therevidae" genus="Squamopygia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Squamopygia" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="8C39B848773415B0E39A6F065B8E8E3C" italics="true" pageId="0" pageNumber="83">Squamopygia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="E881F2E181FFC929C691F2B0BA349DD1" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="9F87AB694F7EA115969188D12036822C" italics="true" pageId="0" pageNumber="83">Calophytus</emphasis>
|
||
</taxonomicName>
|
||
are very similar.
|
||
<taxonomicName id="24D5490DF31ACC83F9F75FF5B40FCE6C" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus grandiosus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="grandiosus">
|
||
<emphasis id="23E9663D55441F5B29F71F867420BA7F" italics="true" pageId="0" pageNumber="83">Calophytus grandiosus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is rather distinct from all other
|
||
<taxonomicName id="47F64ED79D6D362A67E35A6FCB1F90EA" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EA05702652F4E66EDAC14A2854C3BE3E" italics="true" pageId="0" pageNumber="83">Calophytus</emphasis>
|
||
</taxonomicName>
|
||
species, differing in the arrangement of postocular macrosetae, wing banding pattern and body size, supporting its position as sister to the rest of the genus (Fig.
|
||
<figureCitation id="F51218CF1F3A76599A11A5A62C5299B7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Phylogenetic placement of Calophytus gen. nov. and Jeanchazeauia gen. nov. in Agapophytinae based on supermatrix analysis of DNA sequence data. Figure modified after Winterton et al. (2016: fig. 3)." figureDoi="10.3897/zookeys.984.53587.figure4" httpUri="https://binary.pensoft.net/fig/471320" pageId="0" pageNumber="83">4</figureCitation>
|
||
).
|
||
<taxonomicName id="F1C52620137E61640B5FBD47DEFE30CF" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="6C669D1D378DEFE64615B38885E70A81" italics="true" pageId="0" pageNumber="83">Calophytus</emphasis>
|
||
</taxonomicName>
|
||
species have been collected in all of the main habitat types throughout New Caledonia, including rainforest, maquis scrub and dry sclerophyll forest (Figs
|
||
<figureCitation id="E0B520F42FDB961CF25C44BA0A7E174D" captionStart="Figure 22" captionStartId="F22" captionText="Figure 22. Distribution of Calophytus spp. collecting records throughout New Caledonia. Provinces are labelled and ecoregions delineated by colour." figureDoi="10.3897/zookeys.984.53587.figure22" httpUri="https://binary.pensoft.net/fig/471332" pageId="0" pageNumber="83">22</figureCitation>
|
||
,
|
||
<figureCitation id="2A2EFE8A32BCD9C81893F8277CC03C26" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Natural habitats of New Caledonian agapophytine stiletto flies A Riviere Bleue Reserve, sclerophyll forest and maquis scrub, habitat of C. chazeaui sp. nov., C. grandiosus sp. nov., C. schlingeri sp. nov., and J. nubilosus sp. nov. B Pindai Peninsula, tropical dry forest, habitat of C. webbi sp. nov. C Mount Dzumac, rainforest, habitat of J. nubilosus sp. nov. D Mount Dore, rainforest and dry sclerophyll forest E Mount Dore, Malaise trap in flyway (photographs by Michael E. Irwin)." figureDoi="10.3897/zookeys.984.53587.figure31" httpUri="https://binary.pensoft.net/fig/471341" pageId="0" pageNumber="83">31</figureCitation>
|
||
,
|
||
<figureCitation id="A3DCC254A200AE3D0882D98F6AB8E8DF" captionStart="Figure 32" captionStartId="F32" captionText="Figure 32. Natural habitats of New Caledonian agapophytine stiletto flies A Mount Aoupinie, rainforest, habitat of C. monteithi sp. nov. B Mount Ningua, rainforest, habitat of J. nubilosus sp. nov. C Mount Mandjelia, habitat of C. grandiosus sp. nov. (photographs by Michael E. Irwin)." figureDoi="10.3897/zookeys.984.53587.figure32" httpUri="https://binary.pensoft.net/fig/471342" pageId="0" pageNumber="83">32</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="4D48AC8A7DE0DDFC61ED871E3B369686" pageId="0" pageNumber="83" type="included species">
|
||
<paragraph id="BD2A5E31CD7B0B5DCE0D94CC7A523401" pageId="0" pageNumber="83">Included species.</paragraph>
|
||
<paragraph id="45BF9BBF83A3C532353EE260824FBCC2" pageId="0" pageNumber="83">
|
||
<taxonomicName id="25422238E7E422C70EAF89457D54C69B" authorityName="Irwin & Winterton & Metz" authorityYear="2020" class="Insecta" family="Therevidae" genus="Calophytus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Calophytus chazeaui" order="Diptera" pageId="0" pageNumber="83" phylum="Arthropoda" rank="species" species="chazeaui">
|
||
<emphasis id="4F62C45D6D7803F1AE5ABC6EDB105F96" italics="true" pageId="0" pageNumber="83">Calophytus chazeaui</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="452B05FAED25F510BA924E7DD097E74E" lsidName="C. grandiosus" pageId="0" pageNumber="83" rank="species" species="grandiosus">
|
||
<emphasis id="FCE19B0A7B5F406389E7AC2AC7C15442" italics="true" pageId="0" pageNumber="83">C. grandiosus</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="6EDBA2F28628335F06172185CA5EACCC" lsidName="C. matilei" pageId="0" pageNumber="83" rank="species" species="matilei">
|
||
<emphasis id="C515207BF43FEC96DA04F22DF5C4E11B" italics="true" pageId="0" pageNumber="83">C. matilei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="304BFB203459E0FB3FF478170DBBC955" lsidName="C. monteithi" pageId="0" pageNumber="83" rank="species" species="monteithi">
|
||
<emphasis id="9ABE348B1EDDB62DE4617218A28CD25F" italics="true" pageId="0" pageNumber="83">C. monteithi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="C6A7A20CD86C79A927478E35E5F98298" lsidName="C. schlingeri" pageId="0" pageNumber="83" rank="species" species="schlingeri">
|
||
<emphasis id="6576D43A94532DDDAB6E8DC4984B9269" italics="true" pageId="0" pageNumber="83">C. schlingeri</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="F261346E61C2AA65AEC86E704A1DFC0B" lsidName="C. webbi" pageId="0" pageNumber="83" rank="species" species="webbi">
|
||
<emphasis id="ED7C2E162D384AED8552D6528B362AD0" italics="true" pageId="0" pageNumber="83">C. webbi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |