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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.492.6852" ID-GBIF-Dataset="ecf56b94-b4d1-415c-82cd-e7a098046843" ID-PMC="PMC4389217" ID-Pensoft-Pub="1313-2970-492-123" ID-PubMed="25878544" ID-ZBK="3DE583E567954A77B70F7B085B0A91F0" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-492-123" ModsDocOrigin="ZooKeys 492" ModsDocTitle="Revision of the genus Epimesoplecia Zhang, 2007 (Diptera, Nematocera, Protopleciidae) with five new species" checkinTime="1451244493529" checkinUser="pensoft" docAuthor="Lin, Xiuqin, Shih, Chungkun &amp; Ren, Dong" docDate="2015" docId="9D4CFC8F7635CFFC67705ACE755D1942" docLanguage="en" docName="ZooKeys 492: 123-143" docOrigin="ZooKeys 492" docSource="http://dx.doi.org/10.3897/zookeys.492.6852" docTitle="Epimesoplecia macrostrena Lin, Shih &amp; Ren, 2015, sp. n." docType="treatment" docUuid="8A937A8A-0349-45A6-B5FB-D5287856A37F" docUuidSource="ZooBank" docVersion="4" lastPageNumber="131" masterDocId="FFF9FF85DA6BFFD0FF6DFFDB48680A0A" masterDocTitle="Revision of the genus Epimesoplecia Zhang, 2007 (Diptera, Nematocera, Protopleciidae) with five new species" masterLastPageNumber="143" masterPageNumber="123" pageNumber="129" updateTime="1668160147486" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the genus Epimesoplecia Zhang, 2007 (Diptera, Nematocera, Protopleciidae) with five new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lin, Xiuqin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Shih, Chungkun</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ren, Dong</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2015</mods:date>
<mods:detail type="volume">
<mods:number>492</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>123</mods:start>
<mods:end>143</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.492.6852</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.492.6852</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-492-123</mods:identifier>
<mods:identifier type="ZBK">3DE583E567954A77B70F7B085B0A91F0</mods:identifier>
<mods:identifier type="ZooBank">3DE583E567954A77B70F7B085B0A91F0</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152059024" LSID="urn:lsid:zoobank.org:act:8A937A8A-0349-45A6-B5FB-D5287856A37F" httpUri="http://treatment.plazi.org/id/9D4CFC8F7635CFFC67705ACE755D1942" lastPageId="8" lastPageNumber="131" pageId="6" pageNumber="129">
<subSubSection pageId="6" pageNumber="129" type="multiple">
<paragraph pageId="6" pageNumber="129">Taxon classification Animalia Diptera Protopleciidae</paragraph>
</subSubSection>
<subSubSection pageId="6" pageNumber="129" type="nomenclature">
<paragraph pageId="6" pageNumber="129">
<taxonomicName LSID="http://zoobank.org/8A937A8A-0349-45A6-B5FB-D5287856A37F" class="Insecta" family="Protopleciidae" genus="Epimesoplecia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epimesoplecia macrostrena" order="Diptera" pageId="6" pageNumber="129" phylum="Arthropoda" rank="species" species="macrostrena">Epimesoplecia macrostrena</taxonomicName>
<taxonomicNameLabel pageId="6" pageNumber="129">sp. n.</taxonomicNameLabel>
Figs 7, 8
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="130" type="etymology">
<paragraph pageId="7" pageNumber="130">
<pageBreakToken pageId="7" pageNumber="130" start="start">Etymology</pageBreakToken>
.
</paragraph>
<paragraph pageId="7" pageNumber="130">
The epithet of macrostrena is derived from the Greek prefix
<normalizedToken originalValue="“macro-”">&quot;macro-&quot;</normalizedToken>
, meaning
<normalizedToken originalValue="“large”">&quot;large&quot;</normalizedToken>
, and Greek word
<normalizedToken originalValue="“strenos”">&quot;strenos&quot;</normalizedToken>
, meaning &quot;insolence or excess of strength&quot;, referring to the large wings and strong body of this species. The specific epithet is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="130" type="diagnosis">
<paragraph pageId="7" pageNumber="130">Diagnosis.</paragraph>
<paragraph pageId="7" pageNumber="130">Antennae filiform. Sc very close to the level of r-m; fork of Rs proximad of fork of M1+2; R2+3 distad of crossvein r-m; R2+3, straight, slightly shorter than bRs and dRs combined; bRs about 5 times (4.2-5) as long as dRs, the latter almost equal to r-m; bM1+2 shorter than dM1+2 (0.7 times); bM3+4 shorter than m-cu (barely longer than m-cu); cell bp slightly wider than cell ba terminally.</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="130" type="materials">
<paragraph pageId="7" pageNumber="130">Materials.</paragraph>
<paragraph pageId="7" pageNumber="130">Holotype: sex unknown. NO. CNU-DIP-NN-2013211, in lateral view, a well-preserved specimen with partial antennae, wings and body as preserved (Fig. 7). Paratypes: CNU-DIP-NN-2013206p/c, sex unknown, a well-preserved specimen with almost complete wings and body (Fig. 8A, B), NO. CNU-DIP-NN-2013212, female, in ventral view, a well-preserved specimen with complete wings and body as preserved (Fig. 8E).</paragraph>
<caption pageId="7" pageNumber="130">
<paragraph pageId="7" pageNumber="130">
Figure 7.
<taxonomicName class="Insecta" family="Protopleciidae" genus="Epimesoplecia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epimesoplecia macrostrena" order="Diptera" pageId="7" pageNumber="130" phylum="Arthropoda" rank="species" species="macrostrena">Epimesoplecia macrostrena</taxonomicName>
sp. n. (CNU-DIP-NN2013211). Holotype. A Photograph of habitus B Line drawing of habitus C Photograph of details of head (under alcohol); Line drawings of D Head E Wing. Scale bars = 1 mm.
</paragraph>
</caption>
<caption pageId="7" pageNumber="130">
<paragraph pageId="7" pageNumber="130">
Figure 8.
<taxonomicName class="Insecta" family="Protopleciidae" genus="Epimesoplecia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epimesoplecia macrostrena" order="Diptera" pageId="7" pageNumber="130" phylum="Arthropoda" rank="species" species="macrostrena">Epimesoplecia macrostrena</taxonomicName>
sp. n. (CNU-DIP-NN2013206p/c). Paratype. Photographs of A, B Part and counterpart C Details of head; Line drawing of D Counterpart; (CNU-DIP-NN2013212) Paratype E Photograph of habitus F Line drawing of habitus. Scale bars = 1 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="7" pageNumber="130" type="horizon and locality">
<paragraph pageId="7" pageNumber="130">Horizon and locality.</paragraph>
<paragraph pageId="7" pageNumber="130">All specimens were collected from the Jiulongshan Formation, late Middle Jurassic age (Late Callovian) from Daohugou Village, Ningcheng County, Inner Mongolia Autonomous Region in China.</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="131" pageId="7" pageNumber="130" type="description">
<paragraph pageId="7" pageNumber="130">Description.</paragraph>
<paragraph pageId="7" pageNumber="130">Head (Figs 7C, D and 8C): Oviform and very small in lateral view; antennae filiform, segments 1-9 well-preserved, scape and pedicel slightly compressed; flagellar segments slender and long, becoming thinner and shorter toward apex (moniliform in CNU-DIP-NN-2013206p/c in Fig. 8C); maxillary palpi segments not visible.</paragraph>
<paragraph pageId="7" pageNumber="130">Thorax (Figs 7A, B and 8E): Scutum well-developed, arched convex; scutellum depressed; haltere clearly depressed.</paragraph>
<paragraph pageId="7" pageNumber="130">
Legs (Figs 7A, B and 8E): Forelegs: femur thicker than tibia; almost four-fifths of tibia; the 1st tarsomere longer than half of tibia, the 2
<normalizedToken originalValue="nd">nd-</normalizedToken>
5th greatly thinned than the former. Mid legs: femur slightly thicker than tibia; tibial spurs minute; the 1st tarsomere slightly longer than the 1st tarsomere of forelegs. Hind legs: femur expanded almost equal to tibia; tibial spurs minute; tarsi not preserved.
</paragraph>
<paragraph lastPageId="8" lastPageNumber="131" pageId="7" pageNumber="130">
Wings (Figs 7E, 8D and F): Wing long and narrow (length: 7.1-8 mm, width: 2.3-3.2 mm), apex of wings not reaching the abdominal terminalia. Costal field long and thin, apex of Sc gradually tapering to the end; Rs arising almost from basal one-
<pageBreakToken pageId="8" pageNumber="131" start="start">fifth</pageBreakToken>
of wing length, furcating proximad of fork of M1+2 bRs 4.4-5.6 times as long as r-m; crossvein m-cu slightly shorter than r-m; CuA arched near anal margin; vein A1 nearly straight, reaching posterior margin.
</paragraph>
<paragraph pageId="8" pageNumber="131">Female genitalia (Fig. 8F): In lateral view, genitalia simple, with 2-segmented cerci.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="131" type="dimensions">
<paragraph pageId="8" pageNumber="131">Dimensions</paragraph>
<paragraph pageId="8" pageNumber="131">(in mm). [Measurements for the holotype in brackets]. Body length 7 (as preserved)-11.3 [10.3 as preserved]; antennae 0.9 (as preserved)-1.4 (segments 1-16) [0.9 segments 1-9]. Forelegs: femur 2.1-2.5 [1.8 as preserved]; tibia 2.9-3.1 [2.7 as preserved]. Mid leg: femur 2.4-3.8 as preserved [3.8 as preserved]; tibia 2.7-3.6 as preserved [3.6 as preserved]. Hind leg: femur 3.4-4.7 [4.7], tibia 3.8-4.8 [4.8]. Wing: length 7.1-8 [7.4 as preserved], width 2.3-3.3 [3.3]; R2+3 2.4-3.2 [3.2]; bRs 1.9-2.9 [2.9]; dRs 0.5-0.8 [0.6]; R4+5 3-3.7 [3.7].</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="131" type="remarks">
<paragraph pageId="8" pageNumber="131">Remarks.</paragraph>
<paragraph pageId="8" pageNumber="131">
The new species is similar to
<taxonomicName class="Insecta" family="Protopleciidae" genus="Epimesoplecia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Epimesoplecia shcherbakovi" order="Diptera" pageId="8" pageNumber="131" phylum="Arthropoda" rank="species" species="shcherbakovi">Epimesoplecia shcherbakovi</taxonomicName>
Zhang, 2007, but is differentiated from the latter by having Rs bifurcation proximad of fork of M1+2 (vs. Rs bifurcation at the same level of fork of M1+2); bM1+2 shorter than dM1+2 (vs. bM1+2 longer than dM1+2); dRs as long as r-m (vs. dRs clearly shorter than r-m); dM1+2 long, almost one-third of M2 (vs. dM1+2 short, significantly less than one-third of M2). Comparisons with other species are listed in Table 1.
</paragraph>
</subSubSection>
</treatment>
</document>