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<document id="77858D3348F4A6F4AB5FCFFD3857DFAD" ID-DOI="10.5281/zenodo.211274" ID-GBIF-Dataset="351898d8-979b-4b2c-999b-aa4e070ad5ef" ID-ISSN="1175-5326" ID-Zenodo-Dep="211274" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460557599315" checkinUser="plazi" docAuthor="Bellis, Glenn &amp; Dyce, Alan" docDate="2012" docId="7508145A667FFFBBFF78FA72FAE727F7" docLanguage="en" docName="zt03566p060.pdf" docOrigin="Zootaxa 3566" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Culicoides bancrofti Lee and Reye" docType="treatment" docVersion="10" lastPageNumber="56" masterDocId="89316C22667CFFBEFFEFFFD6FFCB2378" masterDocTitle="Redescription of the adults of Culicoides bancrofti Lee and Reye and C. hornsbyensis Lee and Reye (Diptera: Ceratopogonidae)" masterLastPageNumber="60" masterPageNumber="51" pageNumber="54" updateTime="1698319605115" updateUser="plazi">
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<mods:title id="7FBB9A388B603BA3F6F623F36170B752">Redescription of the adults of Culicoides bancrofti Lee and Reye and C. hornsbyensis Lee and Reye (Diptera: Ceratopogonidae)</mods:title>
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<mods:namePart id="07E35FE1981DA538C029638B2C44782E">Bellis, Glenn</mods:namePart>
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<mods:namePart id="7051D62957D75D34336A052F9B9BAE9C">Dyce, Alan</mods:namePart>
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<mods:date id="D44DAC5F6D2F811B9508A3AE46053673">2012</mods:date>
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<treatment id="7508145A667FFFBBFF78FA72FAE727F7" ID-DOI="http://doi.org/10.5281/zenodo.5668456" ID-GBIF-Taxon="119556039" ID-Zenodo-Dep="5668456" LSID="urn:lsid:plazi:treatment:7508145A667FFFBBFF78FA72FAE727F7" httpUri="http://treatment.plazi.org/id/7508145A667FFFBBFF78FA72FAE727F7" lastPageId="5" lastPageNumber="56" pageId="3" pageNumber="54">
<subSubSection id="B5BBF6C7667FFFBDFF78FA72FCCA26A7" pageId="3" pageNumber="54" type="nomenclature">
<paragraph id="FD1EA54C667FFFBDFF78FA72FDF726C6" blockId="3.[151,769,1444,1503]" box="[151,572,1444,1470]" pageId="3" pageNumber="54">
<heading id="A6561220667FFFBDFF78FA72FDF726C6" bold="true" box="[151,572,1444,1470]" fontSize="11" level="1" pageId="3" pageNumber="54" reason="1">
<taxonomicName id="3AA1DECF667FFFBDFF78FA72FDF726C6" ID-CoL="32C8N" authority="Lee and Reye" authorityName="Lee and Reye" box="[151,572,1444,1470]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="bancrofti">
<emphasis id="CFD5795E667FFFBDFF78FA72FDF726C6" bold="true" box="[151,572,1444,1470]" pageId="3" pageNumber="54">
<emphasis id="CFD5795E667FFFBDFF78FA72FE4026C6" bold="true" box="[151,395,1444,1470]" italics="true" pageId="3" pageNumber="54">Culicoides bancrofti</emphasis>
Lee and Reye
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="FD1EA54C667FFFBDFF78FA11FCCA26A7" blockId="3.[151,769,1444,1503]" box="[151,769,1479,1503]" pageId="3" pageNumber="54">
(
<figureCitation id="659AB9C9667FFFBDFF70FA11FE9C26A7" box="[159,343,1479,1503]" captionStart="FIGURES 1 10" captionStartId="4.[151,264,1949,1972]" captionTargetBox="[188,1405,272,1876]" captionTargetId="figure@4.[151,1436,265,1928]" captionTargetPageId="4" captionText="FIGURES 1 10. Culicoides Bancrofti group species. C. bancrofti Lee &amp; Reye, 1, female eye separation; 3, female wing; 4, male wing; 7, legs and 9, palpus. C. hornsbyensis Lee &amp; Reye, 2, female eye separation; 5, female wing; 6, male wing; 8, legs and 10, palpus." httpUri="https://zenodo.org/record/211275/files/figure.png" pageId="3" pageNumber="54">Figs 1, 3, 4, 7, 9</figureCitation>
,
<figureCitation id="659AB9C9667FFFBDFE8CFA11FD9826A7" box="[355,595,1479,1503]" captionStart="FIGURES 11 21" captionStartId="6.[151,264,1892,1915]" captionTargetBox="[188,1396,213,1852]" captionTargetId="figure@6.[151,1436,193,1871]" captionTargetPageId="6" captionText="FIGURES 11 21. Culicoides Bancrofti group species. C. bancrofti Lee &amp; Reye, 11, female antenna; 13, female flagellomere 6; 14, male antenna; 16, 17 spermathecae; 18, tibial comb; 20, mandibular teeth. C. hornsbyensis Lee &amp; Reye, 12, female antenna; 15, male antenna; 17, spermathecae; 19, tibial comb; 21 mandibular teeth." httpUri="https://zenodo.org/record/211276/files/figure.png" pageId="3" pageNumber="54">11, 13, 14, 16, 18, 20</figureCitation>
,
<figureCitation id="659AB9C9667FFFBDFDB1FA11FD3226A7" box="[606,761,1479,1503]" captionStart="FIGURES 22 29" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[187,1398,215,1830]" captionTargetId="figure@7.[151,1436,193,1871]" captionTargetPageId="7" captionText="FIGURES 22 29. Culicoides Bancrofti group species. C. bancrofti Lee &amp; Reye, 22, hypopygium; 23, parameres; 24, aedeagus; 28, distribution based on published records (Lee &amp; Reye 1953; 1955) and specimens represented in ANIC. C. hornsbyensis Lee &amp; Reye., 25, hypopygium; 26, parameres; 27, aedeagus; 29, distribution based on published records (Lee &amp; Reye 1962) and specimens represented in ANIC." httpUri="https://zenodo.org/record/211277/files/figure.png" pageId="3" pageNumber="54">22, 23, 24, 28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="B5BBF6C7667FFFBDFF78F9D9FD94253B" pageId="3" pageNumber="54" type="reference_group">
<paragraph id="FD1EA54C667FFFBDFF78F9D9FDA1255C" blockId="3.[151,1436,1551,1880]" box="[151,618,1551,1573]" pageId="3" pageNumber="54">
<treatmentCitationGroup id="DDB18262667FFFBDFF78F9D9FDA1255C" box="[151,618,1551,1573]" pageId="3" pageNumber="54">
<taxonomicName id="3AA1DECF667FFFBDFF78F9D9FDAD255C" ID-CoL="32C8N" authority="Lee and Reye 1953: 387" authorityName="Lee and Reye" authorityPageNumber="387" authorityYear="1953" box="[151,614,1551,1573]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="bancrofti">
<emphasis id="CFD5795E667FFFBDFF78F9D9FEAE255D" box="[151,357,1551,1573]" italics="true" pageId="3" pageNumber="54">Culicoides bancrofti</emphasis>
<treatmentCitation id="7C00835D667FFFBDFE83F9C6FDAD255C" author="Lee" box="[364,614,1551,1573]" page="387" pageId="3" pageNumber="54" year="1953">
<bibRefCitation id="9930D8BD667FFFBDFE83F9C6FDF8255D" author="Lee" box="[364,563,1551,1573]" pageId="3" pageNumber="54" refString="Lee, D. J. &amp; Reye, E. J. (1953) Australasian Ceratopogonidae (Diptera, Nematocera. Part VI. Australian species of Culicoides. Proceedings of the Linnean Society of NSW, 77, 369 - 394." type="journal article" year="1953">Lee and Reye 1953</bibRefCitation>
: 387
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="FD1EA54C667FFFBDFF78F9F8FD94253B" blockId="3.[151,1436,1551,1880]" box="[151,607,1581,1604]" pageId="3" pageNumber="54">
<bibRefCitation id="9930D8BD667FFFBDFF78F9F8FE89253B" author="Dyce" box="[151,322,1582,1604]" pageId="3" pageNumber="54" refString="Dyce, A. L, Bellis, G. A. &amp; Muller, M. J. (2007) Pictorial atlas of Australasian Culicoides wings (Diptera, Ceratopogonidae). Australian Biological Resources Study, Canberra, 88 pp." type="book" year="2007">
Dyce
<emphasis id="CFD5795E667FFFBDFF3CF9F9FECD253C" box="[211,262,1582,1604]" italics="true" pageId="3" pageNumber="54">et al.</emphasis>
2007
</bibRefCitation>
:34 (female wing illustrated)
</paragraph>
</subSubSection>
<subSubSection id="B5BBF6C7667FFFBDFF78F99BFAD12420" pageId="3" pageNumber="54" type="materials_examined">
<paragraph id="FD1EA54C667FFFBDFF78F99BFA81251A" blockId="3.[151,1436,1551,1880]" box="[151,1354,1612,1635]" pageId="3" pageNumber="54">
<emphasis id="CFD5795E667FFFBDFF78F99BFEF4251A" box="[151,319,1612,1634]" italics="true" pageId="3" pageNumber="54">
<typeStatus id="221A1BEE667FFFBDFF78F99BFF0D251A" box="[151,198,1613,1634]" pageId="3" pageNumber="54">Type</typeStatus>
material ex
</emphasis>
amined:
<emphasis id="CFD5795E667FFFBDFE78F99AFDF7251B" bold="true" box="[407,572,1612,1635]" pageId="3" pageNumber="54">
<collectingCountry id="85B6E5DC667FFFBDFE78F99AFE37251B" box="[407,508,1612,1635]" name="Australia" pageId="3" pageNumber="54">Australia</collectingCountry>
<collectionCode id="9BB03D89667FFFBDFDEDF99AFDF7251B" LSID="urn:lsid:biocol.org:col:15627" box="[514,572,1612,1635]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="3" pageNumber="54">NSW</collectionCode>
</emphasis>
<typeStatus id="221A1BEE667FFFBDFDACF99AFD6F251A" box="[579,676,1612,1634]" pageId="3" pageNumber="54" type="holotype">Holotype</typeStatus>
, Hornsby Gulley,
<collectionCode id="9BB03D89667FFFBDFCB4F99BFC5F251A" LSID="urn:lsid:biocol.org:col:15627" box="[859,916,1613,1634]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="3" pageNumber="54">NSW</collectionCode>
, D.J. Lee,
<date id="891F838C667FFFBDFC10F99AFBB4251A" box="[1023,1151,1612,1634]" pageId="3" pageNumber="54" value="1950-10-29">29.Oct.1950</date>
, 1700 Hrs (female).
</paragraph>
<paragraph id="FD1EA54C667FFFBDFF78F9BAFAD12420" blockId="3.[151,1436,1551,1880]" pageId="3" pageNumber="54">
<emphasis id="CFD5795E667FFFBDFF78F9BAFE0925F9" box="[151,450,1643,1665]" italics="true" pageId="3" pageNumber="54">Non-type material examined.</emphasis>
<emphasis id="CFD5795E667FFFBDFE25F9BCFDB325F9" bold="true" box="[458,632,1642,1665]" pageId="3" pageNumber="54">
<collectingCountry id="85B6E5DC667FFFBDFE25F9BCFDE425F9" box="[458,559,1642,1665]" name="Australia" pageId="3" pageNumber="54">Australia</collectingCountry>
<collectionCode id="9BB03D89667FFFBDFDD8F9BCFDB825F9" LSID="urn:lsid:biocol.org:col:15627" box="[567,627,1642,1665]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15627" name="Royal Botanic Gardens, National Herbarium of New South Wales" pageId="3" pageNumber="54">NSW</collectionCode>
:
</emphasis>
Hornsby,
<date id="891F838C667FFFBDFD0BF9BDFCAF25F9" box="[740,868,1643,1665]" pageId="3" pageNumber="54" value="1968-10-22">22.Oct.1968</date>
, Light Trap, D.J. Lee (
<specimenCount id="EBA76EC5667FFFBDFBB9F9BAFB7925F9" box="[1110,1202,1643,1665]" pageId="3" pageNumber="54" type="female">1 female</specimenCount>
); Hornsby,
<date id="891F838C667FFFBDFAC5F9BDFA5725F9" box="[1322,1436,1643,1665]" pageId="3" pageNumber="54" value="1957-02-07">7.Feb.1957</date>
Light Trap, D.J. Lee (
<specimenCount id="EBA76EC5667FFFBDFE4AF95CFE2625D8" box="[421,493,1674,1696]" pageId="3" pageNumber="54" type="male">1 male</specimenCount>
); Colovale
<date id="891F838C667FFFBDFDB0F95CFD2A25D8" box="[607,737,1674,1696]" pageId="3" pageNumber="54" value="1956-04-24">24.Apr.1956</date>
, Bottom St., 17.3017.45, to man, B.S. Marlow (
<specimenCount id="EBA76EC5667FFFBDFB3BF95CFAFB25D8" box="[1236,1328,1674,1696]" pageId="3" pageNumber="54" type="female">1 female</specimenCount>
); Colovale Stn.,
<date id="891F838C667FFFBDFEE9F97EFEB625C6" box="[262,381,1704,1726]" pageId="3" pageNumber="54" value="1957-03-06">6.Mar.1957</date>
,
<date id="891F838C667FFFBDFE66F97EFE3525C6" box="[393,510,1704,1726]" pageId="3" pageNumber="54" value="1800" valueMax="1815">18001815</date>
, W.W. Wirth (
<specimenCount id="EBA76EC5667FFFBDFD78F97FFD3F25C6" box="[663,756,1704,1726]" pageId="3" pageNumber="54" type="female">1 female</specimenCount>
); Colovale,
<date id="891F838C667FFFBDFC9EF97EFC1B25C6" box="[881,976,1704,1726]" pageId="3" pageNumber="54" value="1954-04">Apr.1954</date>
(
<specimenCount id="EBA76EC5667FFFBDFC0CF97FFB8A25C6" box="[995,1089,1704,1726]" pageId="3" pageNumber="54" type="female">1 female</specimenCount>
); Mogo S.F.,
<date id="891F838C667FFFBDFB21F97EFA8B25C6" box="[1230,1344,1704,1726]" pageId="3" pageNumber="54" value="1981-10-06">6.Oct.1981</date>
, ex light trap, P. Haycock &amp; E. Walter (
<specimenCount id="EBA76EC5667FFFBDFDE8F911FDAF25A5" box="[519,612,1735,1757]" pageId="3" pageNumber="54" type="female">1 female</specimenCount>
); 5 ml. W. Nowra on Yalwal Rd., to man,
<date id="891F838C667FFFBDFBF3F911FB5425A5" box="[1052,1183,1735,1757]" pageId="3" pageNumber="54" value="1966-12-11">11.Dec.1966</date>
, G. Wellings (
<specimenCount id="EBA76EC5667FFFBDFADBF911FA5925A5" box="[1332,1426,1735,1757]" pageId="3" pageNumber="54" type="female">1 female</specimenCount>
); Warrah, lt tp,
<date id="891F838C667FFFBDFEB7F933FE1A2583" box="[344,465,1765,1787]" pageId="3" pageNumber="54" value="1966-11-05">5.Nov.1966</date>
, R. Holst (
<specimenCount id="EBA76EC5667FFFBDFDA9F930FD442583" box="[582,655,1765,1787]" pageId="3" pageNumber="54" type="male">1 male</specimenCount>
); Mt. Dromidary, Lt Tr.,
<date id="891F838C667FFFBDFC7AF933FBD22583" box="[917,1049,1765,1787]" pageId="3" pageNumber="54" value="1965-11-24">24.Nov.1965</date>
, I. Common and M. Upton (
<specimenCount id="EBA76EC5667FFFBDFAA7F930FA5B2583" box="[1352,1424,1765,1787]" pageId="3" pageNumber="54" type="male">1 male</specimenCount>
); Pebbly Beach,
<date id="891F838C667FFFBDFE8FF8D2FE2E2462" box="[352,485,1796,1818]" pageId="3" pageNumber="54" value="1960-11-17">17.Nov.1960</date>
, I.F.B. Common &amp; M. Upton (
<specimenCount id="EBA76EC5667FFFBDFCF1F8D3FCB92462" box="[798,882,1796,1818]" pageId="3" pageNumber="54" type="male">2 males</specimenCount>
).
<emphasis id="CFD5795E667FFFBDFC6CF8D2FC652463" bold="true" box="[899,942,1796,1819]" pageId="3" pageNumber="54">Vic:</emphasis>
Sardine Ck,
<date id="891F838C667FFFBDFBDBF8D2FB632462" box="[1076,1192,1796,1818]" pageId="3" pageNumber="54" value="1966-02-07">7.Feb.1966</date>
, human bait, A.L. Dyce (
<specimenCount id="EBA76EC5667FFFBDFF3EF8F2FEE62441" box="[209,301,1827,1849]" pageId="3" pageNumber="54" type="female">1 female</specimenCount>
); Acheron R.,
<date id="891F838C667FFFBDFE2FF8F5FDF62441" box="[448,573,1827,1849]" pageId="3" pageNumber="54" value="1966-01-22">22.Jan.1966</date>
, aspirator to man A.L. Dyce (
<specimenCount id="EBA76EC5667FFFBDFC80F8F2FC002441" box="[879,971,1827,1849]" pageId="3" pageNumber="54" type="female">1 female</specimenCount>
); Combienbar Riv.,
<quantity id="3A5908A9667FFFBDFB79F8F5FB1B2441" box="[1174,1232,1827,1849]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="3" pageNumber="54" unit="km" value="25.0">25km</quantity>
NE of Club Terace,
<date id="891F838C667FFFBDFF28F894FE81242F" box="[199,330,1858,1879]" pageId="3" pageNumber="54" value="1977-03-17">17.Mar.1977</date>
, A. Neboiss (
<specimenCount id="EBA76EC5667FFFBDFE39F894FDFB2420" box="[470,560,1858,1880]" pageId="3" pageNumber="54" type="female">1 female</specimenCount>
); Britannia Creek,
<quantity id="3A5908A9667FFFBDFD00F894FCE92420" box="[751,802,1858,1880]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="54" unit="km" value="6.0">6 km</quantity>
S. Warburton,
<date id="891F838C667FFFBDFC53F894FBF62420" box="[956,1085,1858,1880]" pageId="3" pageNumber="54" value="1966-02-27">27.Feb.1966</date>
, A. Neboiss (
<specimenCount id="EBA76EC5667FFFBDFB27F894FAC42420" box="[1224,1295,1858,1880]" pageId="3" pageNumber="54" type="male">1 male</specimenCount>
).
</paragraph>
</subSubSection>
<subSubSection id="B5BBF6C7667FFFBAFF78F850FDB723AC" lastPageId="4" lastPageNumber="55" pageId="3" pageNumber="54" type="diagnosis">
<paragraph id="FD1EA54C667FFFBDFF78F850FA572495" blockId="3.[151,1436,1924,2029]" pageId="3" pageNumber="54">
<emphasis id="CFD5795E667FFFBDFF78F850FEDA24E7" bold="true" box="[151,273,1926,1951]" pageId="3" pageNumber="54">Diagnosis.</emphasis>
Apical pale spot in cell r5 ovoid and distant from apex of cell, 3rd palpal segment not markedly broader than 2nd segment, some capitate sensilla present on the surface of the 2nd segment. Female antennae with basal flagellomeres elongate vase-shaped, SCo distribution 1, 913, usually with multiple STc on flagellomeres 18.
</paragraph>
<paragraph id="FD1EA54C6678FFBAFF78FF41FDB723AC" blockId="4.[151,1436,151,212]" pageId="4" pageNumber="55">Male antennae with SCo distribution 1, 1113. Gonocoxite with long, thin ventral root, apicolateral processes very broad, meeting in middle of caudal margin.</paragraph>
</subSubSection>
<caption id="A9DEF5C46678FFBAFF78F84BFEFA2488" httpUri="https://zenodo.org/record/211275/files/figure.png" pageId="4" pageNumber="55" targetBox="[188,1405,272,1876]" targetPageId="4">
<paragraph id="FD1EA54C6678FFBAFF78F84BFEFA2488" blockId="4.[151,1436,1949,2032]" pageId="4" pageNumber="55">
<emphasis id="CFD5795E6678FFBAFF78F84BFE8324CA" bold="true" box="[151,328,1949,1972]" pageId="4" pageNumber="55">FIGURES 110.</emphasis>
<taxonomicName id="3AA1DECF6678FFBAFEBFF84BFDEF24CB" authority="Bancrofti" authorityName="Bancrofti" box="[336,548,1949,1971]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="CFD5795E6678FFBAFEBFF84BFE7724CB" box="[336,444,1949,1971]" italics="true" pageId="4" pageNumber="55">Culicoides</emphasis>
Bancrofti
</taxonomicName>
group species.
<taxonomicName id="3AA1DECF6678FFBAFD29F848FC0224CB" authority="Lee &amp; Reye" authorityName="Lee &amp; Reye" box="[710,969,1949,1971]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="bancrofti">
<emphasis id="CFD5795E6678FFBAFD29F848FC8924CB" box="[710,834,1949,1971]" italics="true" pageId="4" pageNumber="55">C. bancrofti</emphasis>
Lee &amp; Reye
</taxonomicName>
, 1, female eye separation; 3, female wing; 4, male wing; 7, legs and 9, palpus.
<taxonomicName id="3AA1DECF6678FFBAFE03F86BFCD924AA" authority="Lee &amp; Reye" authorityName="Lee &amp; Reye" box="[492,786,1980,2002]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="hornsbyensis">
<emphasis id="CFD5795E6678FFBAFE03F86BFD4524AA" box="[492,654,1980,2002]" italics="true" pageId="4" pageNumber="55">C. hornsbyensis</emphasis>
Lee &amp; Reye
</taxonomicName>
, 2, female eye separation; 5, female wing; 6, male wing; 8, legs and 10, palpus.
</paragraph>
</caption>
<subSubSection id="B5BBF6C76679FFBBFF28FF4EFE052014" pageId="5" pageNumber="56" type="description">
<paragraph id="FD1EA54C6679FFBBFF28FF4EFEEE23C8" blockId="5.[151,1437,152,1168]" box="[199,293,152,176]" pageId="5" pageNumber="56">
<emphasis id="CFD5795E6679FFBBFF28FF4EFEEE23C8" bold="true" box="[199,293,152,176]" pageId="5" pageNumber="56">Female.</emphasis>
</paragraph>
<paragraph id="FD1EA54C6679FFBBFF28FF69FB62223D" blockId="5.[151,1437,152,1168]" pageId="5" pageNumber="56">
<emphasis id="CFD5795E6679FFBBFF28FF69FEC523AF" bold="true" box="[199,270,191,215]" pageId="5" pageNumber="56">Head.</emphasis>
Eyes separated by a distance less than ¾ of one facet (
<figureCitation id="659AB9C96679FFBBFC68FF69FC0823AF" box="[903,963,191,215]" captionStart="FIGURES 1 10" captionStartId="4.[151,264,1949,1972]" captionTargetBox="[188,1405,272,1876]" captionTargetId="figure@4.[151,1436,265,1928]" captionTargetPageId="4" captionText="FIGURES 1 10. Culicoides Bancrofti group species. C. bancrofti Lee &amp; Reye, 1, female eye separation; 3, female wing; 4, male wing; 7, legs and 9, palpus. C. hornsbyensis Lee &amp; Reye, 2, female eye separation; 5, female wing; 6, male wing; 8, legs and 10, palpus." httpUri="https://zenodo.org/record/211275/files/figure.png" pageId="5" pageNumber="56">Fig 1</figureCitation>
). Palpus (
<figureCitation id="659AB9C96679FFBBFBD6FF69FBBC23AF" box="[1081,1143,191,215]" captionStart="FIGURES 1 10" captionStartId="4.[151,264,1949,1972]" captionTargetBox="[188,1405,272,1876]" captionTargetId="figure@4.[151,1436,265,1928]" captionTargetPageId="4" captionText="FIGURES 1 10. Culicoides Bancrofti group species. C. bancrofti Lee &amp; Reye, 1, female eye separation; 3, female wing; 4, male wing; 7, legs and 9, palpus. C. hornsbyensis Lee &amp; Reye, 2, female eye separation; 5, female wing; 6, male wing; 8, legs and 10, palpus." httpUri="https://zenodo.org/record/211275/files/figure.png" pageId="5" pageNumber="56">Fig 9</figureCitation>
) dark brown; 3rd segment cylindrical with numerous capitate sensilla scattered on the surface of the distal 2/3 of the segment, some capitate sensilla also present on the surface of 2nd segment. Flagellomeres (
<figureCitation id="659AB9C96679FFBBFC73FEDFFBEB2259" box="[924,1056,265,289]" captionStart="FIGURES 11 21" captionStartId="6.[151,264,1892,1915]" captionTargetBox="[188,1396,213,1852]" captionTargetId="figure@6.[151,1436,193,1871]" captionTargetPageId="6" captionText="FIGURES 11 21. Culicoides Bancrofti group species. C. bancrofti Lee &amp; Reye, 11, female antenna; 13, female flagellomere 6; 14, male antenna; 16, 17 spermathecae; 18, tibial comb; 20, mandibular teeth. C. hornsbyensis Lee &amp; Reye, 12, female antenna; 15, male antenna; 17, spermathecae; 19, tibial comb; 21 mandibular teeth." httpUri="https://zenodo.org/record/211276/files/figure.png" pageId="5" pageNumber="56">Figs 11, 13</figureCitation>
) 28 elongate, vase-shaped with basal half clothed in microtrichea; 913 subcylindrical, little longer than basal flagellomeres.
</paragraph>
<paragraph id="FD1EA54C6679FFBBFF28FE84FDE022A3" blockId="5.[151,1437,152,1168]" pageId="5" pageNumber="56">
<emphasis id="CFD5795E6679FFBBFF28FE84FEEC2212" bold="true" box="[199,295,338,362]" pageId="5" pageNumber="56">Thorax.</emphasis>
Legs (
<figureCitation id="659AB9C96679FFBBFE9BFE87FE792211" box="[372,434,337,361]" captionStart="FIGURES 1 10" captionStartId="4.[151,264,1949,1972]" captionTargetBox="[188,1405,272,1876]" captionTargetId="figure@4.[151,1436,265,1928]" captionTargetPageId="4" captionText="FIGURES 1 10. Culicoides Bancrofti group species. C. bancrofti Lee &amp; Reye, 1, female eye separation; 3, female wing; 4, male wing; 7, legs and 9, palpus. C. hornsbyensis Lee &amp; Reye, 2, female eye separation; 5, female wing; 6, male wing; 8, legs and 10, palpus." httpUri="https://zenodo.org/record/211275/files/figure.png" pageId="5" pageNumber="56">Fig 7</figureCitation>
) dark brown, hind femora unbanded, mid and fore femora with subapical pale band, that on mid femora less pronounced, tibiae with pale subbasal band, apex unbanded. Wing (
<figureCitation id="659AB9C96679FFBBFB95FEA3FB7C22F5" box="[1146,1207,373,397]" captionStart="FIGURES 1 10" captionStartId="4.[151,264,1949,1972]" captionTargetBox="[188,1405,272,1876]" captionTargetId="figure@4.[151,1436,265,1928]" captionTargetPageId="4" captionText="FIGURES 1 10. Culicoides Bancrofti group species. C. bancrofti Lee &amp; Reye, 1, female eye separation; 3, female wing; 4, male wing; 7, legs and 9, palpus. C. hornsbyensis Lee &amp; Reye, 2, female eye separation; 5, female wing; 6, male wing; 8, legs and 10, palpus." httpUri="https://zenodo.org/record/211275/files/figure.png" pageId="5" pageNumber="56">Fig 3</figureCitation>
) strongly patterned, apical pale spot in cell r5 ovoid and distant from wing margin. Tibial comb (
<figureCitation id="659AB9C96679FFBBFC01FE4AFBF322CC" box="[1006,1080,412,436]" captionStart="FIGURES 11 21" captionStartId="6.[151,264,1892,1915]" captionTargetBox="[188,1396,213,1852]" captionTargetId="figure@6.[151,1436,193,1871]" captionTargetPageId="6" captionText="FIGURES 11 21. Culicoides Bancrofti group species. C. bancrofti Lee &amp; Reye, 11, female antenna; 13, female flagellomere 6; 14, male antenna; 16, 17 spermathecae; 18, tibial comb; 20, mandibular teeth. C. hornsbyensis Lee &amp; Reye, 12, female antenna; 15, male antenna; 17, spermathecae; 19, tibial comb; 21 mandibular teeth." httpUri="https://zenodo.org/record/211276/files/figure.png" pageId="5" pageNumber="56">Fig 18</figureCitation>
) with 56 spines, 2nd from spur longest. Haltere pale to light brown.
</paragraph>
<paragraph id="FD1EA54C6679FFBBFF28FE31FEC02287" blockId="5.[151,1437,152,1168]" box="[199,267,487,511]" pageId="5" pageNumber="56">
<emphasis id="CFD5795E6679FFBBFF28FE31FEC02287" bold="true" box="[199,267,487,511]" pageId="5" pageNumber="56">Male.</emphasis>
</paragraph>
<paragraph id="FD1EA54C6679FFBBFF28FDD8FBDA2132" blockId="5.[151,1437,152,1168]" pageId="5" pageNumber="56">
<emphasis id="CFD5795E6679FFBBFF28FDD8FEC5215E" bold="true" box="[199,270,526,550]" pageId="5" pageNumber="56">Head.</emphasis>
Palpus with subcylindrical 3rd segment bearing few capitate sensilla, these mostly on the inner surface. Antenna (
<figureCitation id="659AB9C96679FFBBFEEBFDE4FE842132" box="[260,335,562,586]" captionStart="FIGURES 11 21" captionStartId="6.[151,264,1892,1915]" captionTargetBox="[188,1396,213,1852]" captionTargetId="figure@6.[151,1436,193,1871]" captionTargetPageId="6" captionText="FIGURES 11 21. Culicoides Bancrofti group species. C. bancrofti Lee &amp; Reye, 11, female antenna; 13, female flagellomere 6; 14, male antenna; 16, 17 spermathecae; 18, tibial comb; 20, mandibular teeth. C. hornsbyensis Lee &amp; Reye, 12, female antenna; 15, male antenna; 17, spermathecae; 19, tibial comb; 21 mandibular teeth." httpUri="https://zenodo.org/record/211276/files/figure.png" pageId="5" pageNumber="56">Fig 14</figureCitation>
) with flagellomeres 11 and 12 each with a basal whorl of SCh.
</paragraph>
<paragraph id="FD1EA54C6679FFBBFF28FD80FD8D21EC" blockId="5.[151,1437,152,1168]" pageId="5" pageNumber="56">
<emphasis id="CFD5795E6679FFBBFF28FD80FEEC2116" bold="true" box="[199,295,598,622]" pageId="5" pageNumber="56">Thorax.</emphasis>
Wing (
<figureCitation id="659AB9C96679FFBBFE90FD80FE752116" box="[383,446,598,622]" captionStart="FIGURES 1 10" captionStartId="4.[151,264,1949,1972]" captionTargetBox="[188,1405,272,1876]" captionTargetId="figure@4.[151,1436,265,1928]" captionTargetPageId="4" captionText="FIGURES 1 10. Culicoides Bancrofti group species. C. bancrofti Lee &amp; Reye, 1, female eye separation; 3, female wing; 4, male wing; 7, legs and 9, palpus. C. hornsbyensis Lee &amp; Reye, 2, female eye separation; 5, female wing; 6, male wing; 8, legs and 10, palpus." httpUri="https://zenodo.org/record/211275/files/figure.png" pageId="5" pageNumber="56">Fig 4</figureCitation>
) patterned similarly to female with pale spots generally more expansive. Tibial comb with 46 spines, 2nd from spur longest.
</paragraph>
<paragraph id="FD1EA54C6679FFBBFF28FD76FB182030" blockId="5.[151,1437,152,1168]" pageId="5" pageNumber="56">
<emphasis id="CFD5795E6679FFBBFF28FD76FEFE21C1" bold="true" box="[199,309,672,697]" pageId="5" pageNumber="56">Genitalia</emphasis>
. Hypopygium (
<figureCitation id="659AB9C96679FFBBFE04FD76FDF321C0" box="[491,568,672,696]" captionStart="FIGURES 22 29" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[187,1398,215,1830]" captionTargetId="figure@7.[151,1436,193,1871]" captionTargetPageId="7" captionText="FIGURES 22 29. Culicoides Bancrofti group species. C. bancrofti Lee &amp; Reye, 22, hypopygium; 23, parameres; 24, aedeagus; 28, distribution based on published records (Lee &amp; Reye 1953; 1955) and specimens represented in ANIC. C. hornsbyensis Lee &amp; Reye., 25, hypopygium; 26, parameres; 27, aedeagus; 29, distribution based on published records (Lee &amp; Reye 1962) and specimens represented in ANIC." httpUri="https://zenodo.org/record/211277/files/figure.png" pageId="5" pageNumber="56">Fig 22</figureCitation>
) with ninth tergite narrowed apically, apicolateral processes triangular with bases very broad, meeting medially forming a very deep median cleft. Ninth sternite with moderately deep caudomedial excavation. Gonocoxite with dorsal root simple; ventral root long, narrowing to a sharp point. Gonostylus slender, distally curving gently to a pointed apex. Aedeagus (
<figureCitation id="659AB9C96679FFBBFCABFCDAFC44205C" box="[836,911,780,804]" captionStart="FIGURES 22 29" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[187,1398,215,1830]" captionTargetId="figure@7.[151,1436,193,1871]" captionTargetPageId="7" captionText="FIGURES 22 29. Culicoides Bancrofti group species. C. bancrofti Lee &amp; Reye, 22, hypopygium; 23, parameres; 24, aedeagus; 28, distribution based on published records (Lee &amp; Reye 1953; 1955) and specimens represented in ANIC. C. hornsbyensis Lee &amp; Reye., 25, hypopygium; 26, parameres; 27, aedeagus; 29, distribution based on published records (Lee &amp; Reye 1962) and specimens represented in ANIC." httpUri="https://zenodo.org/record/211277/files/figure.png" pageId="5" pageNumber="56">Fig 24</figureCitation>
) with moderately deep basal arch extending to about 1/2 of aedeagus length, septum narrow; distal process moderately long with a simple apex.
</paragraph>
<paragraph id="FD1EA54C6679FFBBFF28FC83FE052014" blockId="5.[151,1437,152,1168]" box="[199,462,852,877]" pageId="5" pageNumber="56">
<emphasis id="CFD5795E6679FFBBFF28FC83FE992015" bold="true" box="[199,338,853,877]" pageId="5" pageNumber="56">Immatures.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="B5BBF6C76679FFBBFF28FCAEFD7D20E8" box="[199,694,888,913]" pageId="5" pageNumber="56" type="distribution">
<paragraph id="FD1EA54C6679FFBBFF28FCAEFD7D20E8" blockId="5.[151,1437,152,1168]" box="[199,694,888,913]" pageId="5" pageNumber="56">
<emphasis id="CFD5795E6679FFBBFF28FCAEFEAB20E9" bold="true" box="[199,352,888,913]" pageId="5" pageNumber="56">Distribution.</emphasis>
(
<figureCitation id="659AB9C96679FFBBFE80FCAEFE7120E8" box="[367,442,888,912]" captionStart="FIGURES 22 29" captionStartId="7.[151,264,1892,1915]" captionTargetBox="[187,1398,215,1830]" captionTargetId="figure@7.[151,1436,193,1871]" captionTargetPageId="7" captionText="FIGURES 22 29. Culicoides Bancrofti group species. C. bancrofti Lee &amp; Reye, 22, hypopygium; 23, parameres; 24, aedeagus; 28, distribution based on published records (Lee &amp; Reye 1953; 1955) and specimens represented in ANIC. C. hornsbyensis Lee &amp; Reye., 25, hypopygium; 26, parameres; 27, aedeagus; 29, distribution based on published records (Lee &amp; Reye 1962) and specimens represented in ANIC." httpUri="https://zenodo.org/record/211277/files/figure.png" pageId="5" pageNumber="56">Fig 28</figureCitation>
)
<collectingCountry id="85B6E5DC6679FFBBFE26FCAEFDFF20E8" box="[457,564,888,912]" name="Australia" pageId="5" pageNumber="56">Australia</collectingCountry>
: NSW, Vic.
</paragraph>
</subSubSection>
<subSubSection id="B5BBF6C76679FFBBFF28FC4AFBA42084" pageId="5" pageNumber="56" type="biology_ecology">
<paragraph id="FD1EA54C6679FFBBFF28FC4AFBA42084" blockId="5.[151,1437,152,1168]" pageId="5" pageNumber="56">
<emphasis id="CFD5795E6679FFBBFF28FC4AFEEC20CD" bold="true" box="[199,295,924,949]" pageId="5" pageNumber="56">Biology.</emphasis>
Largely unknown.
<bibRefCitation id="9930D8BD6679FFBBFDE6FC4BFD0020CC" box="[521,715,924,949]" pageId="5" pageNumber="56" refString="Lee, D. J., Reye, E. J. &amp; Dyce, A. L. (1963) &quot; Sandflies &quot; as possible vectors of disease in domesticated animals in Australia. Proceedings of the Linnean Society of New South Wales, 87, 364 - 376." type="journal article">
Lee
<emphasis id="CFD5795E6679FFBBFDD3FC48FDA420CD" box="[572,623,925,949]" italics="true" pageId="5" pageNumber="56">et al</emphasis>
. (1963)
</bibRefCitation>
suggested adults were diurnally active or perhaps crepuscular. They also reported feeding on humans and label data on specimens examined herein supports this finding although this species is relatively uncommon in collections so is unlikely to be an important pest.
</paragraph>
</subSubSection>
<subSubSection id="B5BBF6C76679FFBBFF28FBDFFAE727F7" pageId="5" pageNumber="56" type="discussion">
<paragraph id="FD1EA54C6679FFBBFF28FBDFFAE727F7" blockId="5.[151,1437,152,1168]" pageId="5" pageNumber="56">
<emphasis id="CFD5795E6679FFBBFF28FBDFFEF02759" bold="true" box="[199,315,1033,1057]" pageId="5" pageNumber="56">Remarks.</emphasis>
This species can be distinguished from
<taxonomicName id="3AA1DECF6679FFBBFD11FBDFFC7A2759" box="[766,945,1033,1057]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="species" species="hornsbyensis">
<emphasis id="CFD5795E6679FFBBFD11FBDFFC7A2759" box="[766,945,1033,1057]" italics="true" pageId="5" pageNumber="56">C. hornsbyensis</emphasis>
</taxonomicName>
by the small pale spot in cell r5 of the wing which does not approach the wing margin, the vase-like shape of basal flagellomeres of the female, the broad apicolateral processes of the male or the SCo distribution in both sexes. The presence of capitate sensilla on the second palpal segment is very unusual in the genus
<taxonomicName id="3AA1DECF6679FFBBFD35FBA1FC9927F7" box="[730,850,1143,1167]" class="Insecta" family="Ceratopogonidae" genus="Culicoides" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="CFD5795E6679FFBBFD35FBA1FC9927F7" box="[730,850,1143,1167]" italics="true" pageId="5" pageNumber="56">Culicoides</emphasis>
</taxonomicName>
and unique amongst the Australian fauna.
</paragraph>
</subSubSection>
</treatment>
</document>