134 lines
19 KiB
XML
134 lines
19 KiB
XML
<document id="C20646CF812775183FB41C03D821007B" ID-DOI="10.11646/zootaxa.3750.5.7" ID-GBIF-Dataset="d35e62c7-975b-4dff-892e-e8e5c926820e" ID-ISSN="1175-5326" ID-Zenodo-Dep="217852" ID-ZooBank="22C6D62A-D6CA-4BBE-AD0D-29174C2A2769" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460711129622" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2013" docId="7505414FFF917371FF29F8ADFE6E4113" docLanguage="en" docName="zt03750p568.pdf" docOrigin="Zootaxa 3750 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Alloraphes magnus Franz" docType="treatment" docVersion="6" lastPageNumber="568" masterDocId="893C3937FF837362FFBEFFC2FFB14304" masterDocTitle="Revision of the Neotropical genus Alloraphes Franz (Coleoptera: Staphylinidae: Scydmaeninae)" masterLastPageNumber="568" masterPageNumber="549" pageNumber="567" updateTime="1698357997849" updateUser="plazi">
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<mods:title id="4A3D04C3D2386021CAEEFA6E0518B66C">Revision of the Neotropical genus Alloraphes Franz (Coleoptera: Staphylinidae: Scydmaeninae)</mods:title>
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<mods:name id="1CF0462B76A39A150F365D8654C94E2A" type="personal">
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<mods:namePart id="40747C2424A7C0C76841A080BF8E3E1D">Jałoszyński, Paweł</mods:namePart>
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<mods:title id="7641CBCB14B71438D8F36D48EB87F300">Zootaxa</mods:title>
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<mods:part id="CC7B0CBD256A1A3A666AE9DC6CB7F01A">
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<mods:date id="DE9BFB0337C5E52A284960212E85947D">2013</mods:date>
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<mods:number id="6C60636ED9594C25F24E54C76253A37D">3750</mods:number>
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<mods:number id="A49615576D8B1C0939908E22979F2BDA">5</mods:number>
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<mods:classification id="69B55CB311905DFA128AC12036614FCB">journal article</mods:classification>
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<mods:identifier id="0109BE3242BD1BAEF82971161FA8CAA7" type="DOI">10.11646/zootaxa.3750.5.7</mods:identifier>
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<treatment id="7505414FFF917371FF29F8ADFE6E4113" ID-DOI="http://doi.org/10.5281/zenodo.6154539" ID-GBIF-Taxon="119585164" ID-Zenodo-Dep="6154539" LSID="urn:lsid:plazi:treatment:7505414FFF917371FF29F8ADFE6E4113" httpUri="http://treatment.plazi.org/id/7505414FFF917371FF29F8ADFE6E4113" lastPageId="19" lastPageNumber="568" pageId="18" pageNumber="567">
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<subSubSection id="B5B6A3D2FF917370FF29F8ADFDAE44F4" pageId="18" pageNumber="567" type="nomenclature">
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<paragraph id="FD13F059FF917370FF29F8ADFE65448E" blockId="18.[151,468,1903,1963]" box="[151,468,1903,1930]" pageId="18" pageNumber="567">
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<heading id="A65B4735FF917370FF29F8ADFE65448E" bold="true" box="[151,468,1903,1930]" fontSize="11" level="1" pageId="18" pageNumber="567" reason="1">
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<taxonomicName id="3AAC8BDAFF917370FF29F8ADFE65448E" authority="Franz" authorityName="Franz" box="[151,468,1903,1930]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="magnus">
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<emphasis id="CFD82C4BFF917370FF29F8ADFE65448E" bold="true" box="[151,468,1903,1930]" pageId="18" pageNumber="567">
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<emphasis id="CFD82C4BFF917370FF29F8ADFE33448D" bold="true" box="[151,386,1903,1929]" italics="true" pageId="18" pageNumber="567">Alloraphes magnus</emphasis>
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Franz
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="FD13F059FF917370FF29F850FE3044AF" blockId="18.[151,468,1903,1963]" box="[151,385,1938,1963]" pageId="18" pageNumber="567">
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(
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<figureCitation id="6597ECDCFF917370FF21F850FF4D44AF" box="[159,252,1938,1963]" captionStart="FIGURES 1 – 12" captionStartId="3.[151,264,1879,1902]" captionTargetBox="[217,1363,203,1830]" captionTargetId="figure@3.[209,1368,195,1845]" captionTargetPageId="3" captionText="FIGURES 1 – 12. Dorsal (1 – 9, 11 – 12) and laterodorsal (10) habitus of Alloraphes Franz. Alloraphes jamaicae Franz, holotype male (1); A. chiapasensis Franz, holotype male (2); A. yucatani Franz, holotype male (3); A. lenkoi Franz, paratype male (4) and paratype female (5); A. peruanus Franz, holotype male (6) and paratype female (7; but see the Remarks section in the redescription); A. dentatus sp. n., holotype male (8); A. peckorum sp. n., holotype male (9 – 10); A. myrmecophilus Franz, stat. rest., paratype male (11);? Alloraphes magnus Franz, holotype female (12)." httpUri="https://zenodo.org/record/217853/files/figure.png" pageId="18" pageNumber="567">Figs. 12</figureCitation>
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,
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<figureCitation id="6597ECDCFF917370FEB6F850FE9744AF" box="[264,294,1938,1963]" captionStart="FIGURES 22 – 26" captionStartId="7.[151,264,1642,1665]" captionTargetBox="[222,1331,193,1600]" captionTargetId="figure@7.[200,1342,193,1608]" captionTargetPageId="7" captionText="FIGURES 22 – 26. Alloraphes jamaicae Franz, holotype male (22 – 24); A. myrmecophilus Franz stat. rest., paratype male (25);? Alloraphes magnus Franz, holotype female (26). Head in ventral view (22); prothorax in ventral view (23); pterothorax in ventral view (24); body venter (25); and mesothorax in ventral view (26). Abbreviations: aest 2, mesanepisternum; aest 3, metanepisternum; ai, asetose impression; amc, anterior marginal carina of mesocoxal cavity; ar, anterior ridge; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; cx, coxa; el, elytron; epm 3, metepimeron; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; lss, lateral suture of submentum; mc, median carina of mesoventrite; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mesostipes; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; pl, posterior lobe of mesocoxal projection; pre, prepectus; ppf, palpifer; psp, prosternal process; submentum; tm, temple; v 3, metaventrite" httpUri="https://zenodo.org/record/217857/files/figure.png" pageId="18" pageNumber="567">26</figureCitation>
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,
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<figureCitation id="6597ECDCFF917370FE8CF850FEE044AF" box="[306,337,1938,1963]" captionStart="FIGURES 61 – 67" captionStartId="11.[151,264,1809,1832]" captionTargetBox="[161,1423,953,1744]" captionTargetId="figure@11.[151,1436,939,1757]" captionTargetPageId="11" captionText="FIGURES 61 – 67. Original sets of labels pinned under historical type specimens. Alloraphes jamaicae Franz, holotype (61); A. chiapasensis Franz, holotype (62); A. yucatani Franz, holotype (63); A. lenkoi Franz, paratype (64); A. peruanus Franz, holotype (65); A. myrmecophilus Franz, stat. rest., paratype (66);? Alloraphes magnus Franz, holotype (67)." httpUri="https://zenodo.org/record/217860/files/figure.png" pageId="18" pageNumber="567">67</figureCitation>
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,
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<figureCitation id="6597ECDCFF917370FEE3F850FECB44AF" box="[349,378,1938,1963]" captionStart="FIGURE 68" captionStartId="12.[151,250,1290,1312]" captionTargetBox="[151,1418,167,1268]" captionTargetId="figure@12.[151,1436,167,1268]" captionTargetPageId="12" captionText="FIGURE 68. Distribution of Alloraphes: a, A. jamaicae Franz; b, A. chiapasensis Franz; c, A. yucatani Franz; d, A. lenkoi Franz; e, A. peruanus Franz; f, A. dentatus sp. n.; g, A. peckorum sp. n.; h, A. myrmecophilus Franz stat. rest.; i, A. bolivarensis Franz; j,? Alloraphes magnus Franz." httpUri="https://zenodo.org/record/217861/files/figure.png" pageId="18" pageNumber="567">68</figureCitation>
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)
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</paragraph>
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<paragraph id="FD13F059FF917370FF29F818FDAE44F4" blockId="18.[151,543,2010,2032]" box="[151,543,2010,2032]" pageId="18" pageNumber="567">
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<taxonomicName id="3AAC8BDAFF917370FF29F818FDAD44F4" authority="Franz, 1980a: 215" authorityName="Franz" authorityPageNumber="215" authorityYear="1980" box="[151,540,2010,2032]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="567" phylum="Arthropoda" rank="species" species="magnus">
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<emphasis id="CFD82C4BFF917370FF29F818FEEA44F4" box="[151,347,2010,2032]" italics="true" pageId="18" pageNumber="567">Alloraphes magnus</emphasis>
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Franz, 1980a: 215
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B5B6A3D2FF907371FF29FF55FB3F43FC" pageId="19" pageNumber="568" type="materials_examined">
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<paragraph id="FD13F059FF907371FF29FF55FB3F43FC" blockId="19.[151,1437,151,536]" pageId="19" pageNumber="568">
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<emphasis id="CFD82C4BFF907371FF29FF55FD1F43B4" bold="true" box="[151,686,151,176]" pageId="19" pageNumber="568">
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<typeStatus id="22174EFBFF907371FF29FF55FF6343B4" box="[151,210,151,176]" pageId="19" pageNumber="568">Type</typeStatus>
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material.
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<typeStatus id="22174EFBFF907371FEF2FF55FE0C43B4" box="[332,445,151,176]" pageId="19" pageNumber="568" type="holotype">Holotype</typeStatus>
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:
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<collectingCountry id="85BBB0C9FF907371FE74FF55FDA243B4" box="[458,531,151,176]" name="Brazil" pageId="19" pageNumber="568">Brazil</collectingCountry>
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(São Paulo):
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</emphasis>
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♀, three labels (
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<figureCitation id="6597ECDCFF907371FCD7FF55FC0F43B4" box="[873,958,151,176]" captionStart="FIGURES 61 – 67" captionStartId="11.[151,264,1809,1832]" captionTargetBox="[161,1423,953,1744]" captionTargetId="figure@11.[151,1436,939,1757]" captionTargetPageId="11" captionText="FIGURES 61 – 67. Original sets of labels pinned under historical type specimens. Alloraphes jamaicae Franz, holotype (61); A. chiapasensis Franz, holotype (62); A. yucatani Franz, holotype (63); A. lenkoi Franz, paratype (64); A. peruanus Franz, holotype (65); A. myrmecophilus Franz, stat. rest., paratype (66);? Alloraphes magnus Franz, holotype (67)." httpUri="https://zenodo.org/record/217860/files/figure.png" pageId="19" pageNumber="568">Fig. 67</figureCitation>
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): "Barueri, Sao Paulo / Brasij [sic!], lg. K. Lenko" with "No 10" on reverse side [white, printed; reverse handwritten in blue ink]; "
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<taxonomicName id="3AAC8BDAFF907371FBCCFF7EFB5C43D0" box="[1138,1261,188,212]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="568" phylum="Arthropoda" rank="genus">
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<emphasis id="CFD82C4BFF907371FBCCFF7EFB5C43D0" box="[1138,1261,188,212]" italics="true" pageId="19" pageNumber="568">Alloraphes</emphasis>
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</taxonomicName>
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/
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<taxonomicName id="3AAC8BDAFF907371FABDFF7FFAEB43D0" box="[1283,1370,189,212]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="568" phylum="Arthropoda" rank="species" species="magnus">
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<emphasis id="CFD82C4BFF907371FABDFF7FFAEB43D0" box="[1283,1370,189,212]" italics="true" pageId="19" pageNumber="568">magnus</emphasis>
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</taxonomicName>
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/ m. / det. H. Franz" [white, handwritten and printed]; "
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<typeStatus id="22174EFBFF907371FD03FF1DFC8643FC" box="[701,823,223,248]" pageId="19" pageNumber="568" type="holotype">Holotypus</typeStatus>
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" [red, handwritten] (NHMW).
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</paragraph>
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</subSubSection>
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<subSubSection id="B5B6A3D2FF907371FF79FEC6FE6E4113" pageId="19" pageNumber="568" type="discussion">
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<paragraph id="FD13F059FF907371FF79FEC6FE6E4113" blockId="19.[151,1437,151,536]" pageId="19" pageNumber="568">
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<emphasis id="CFD82C4BFF907371FF79FEC6FE8A4219" bold="true" box="[199,315,260,285]" pageId="19" pageNumber="568">Remarks.</emphasis>
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<taxonomicName id="3AAC8BDAFF907371FEFFFEC6FDAB4218" box="[321,538,260,284]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="568" phylum="Arthropoda" rank="species" species="magnus">
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<emphasis id="CFD82C4BFF907371FEFFFEC6FDAB4218" box="[321,538,260,284]" italics="true" pageId="19" pageNumber="568">Alloraphes magnus</emphasis>
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</taxonomicName>
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is known from a single female collected in the southern part of
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<collectingCountry id="85BBB0C9FF907371FB5EFEC6FA954219" box="[1248,1316,260,285]" name="Brazil" pageId="19" pageNumber="568">Brazil</collectingCountry>
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(
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<figureCitation id="6597ECDCFF907371FA8AFEC6FA374219" box="[1332,1414,260,285]" captionStart="FIGURE 68" captionStartId="12.[151,250,1290,1312]" captionTargetBox="[151,1418,167,1268]" captionTargetId="figure@12.[151,1436,167,1268]" captionTargetPageId="12" captionText="FIGURE 68. Distribution of Alloraphes: a, A. jamaicae Franz; b, A. chiapasensis Franz; c, A. yucatani Franz; d, A. lenkoi Franz; e, A. peruanus Franz; f, A. dentatus sp. n.; g, A. peckorum sp. n.; h, A. myrmecophilus Franz stat. rest.; i, A. bolivarensis Franz; j,? Alloraphes magnus Franz." httpUri="https://zenodo.org/record/217861/files/figure.png" pageId="19" pageNumber="568">Fig. 68</figureCitation>
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j). It differs from all species of
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<taxonomicName id="3AAC8BDAFF907371FE63FEEAFDE94244" box="[477,600,296,320]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="568" phylum="Arthropoda" rank="genus">
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<emphasis id="CFD82C4BFF907371FE63FEEAFDE94244" box="[477,600,296,320]" italics="true" pageId="19" pageNumber="568">Alloraphes</emphasis>
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</taxonomicName>
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in the pronotum without even traces of lateral marginal carinae or edges, different shape of the mesoventral intercoxal process and the presence of an additional median carina within the asetose impressions of mesoventrite (
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<figureCitation id="6597ECDCFF907371FD85FEADFD3E428C" box="[571,655,367,392]" captionStart="FIGURES 22 – 26" captionStartId="7.[151,264,1642,1665]" captionTargetBox="[222,1331,193,1600]" captionTargetId="figure@7.[200,1342,193,1608]" captionTargetPageId="7" captionText="FIGURES 22 – 26. Alloraphes jamaicae Franz, holotype male (22 – 24); A. myrmecophilus Franz stat. rest., paratype male (25);? Alloraphes magnus Franz, holotype female (26). Head in ventral view (22); prothorax in ventral view (23); pterothorax in ventral view (24); body venter (25); and mesothorax in ventral view (26). Abbreviations: aest 2, mesanepisternum; aest 3, metanepisternum; ai, asetose impression; amc, anterior marginal carina of mesocoxal cavity; ar, anterior ridge; bst, basisternal part of prosternum; bst, basistipes; cd, cardo; cx, coxa; el, elytron; epm 3, metepimeron; fo, foramen occipitale; gal, galea; gp, gular plate; gs, gular suture; hr, hypostomal ridge; hy, hypomeron; hyr, hypomeral ridge; lac, lacinia; lp, labial palp; lss, lateral suture of submentum; mc, median carina of mesoventrite; mcp, mesocoxal projection; md, mandible; mn, mentum; mscc, mesocoxal cavity; mscs, mesocoxal socket; msff, mesofurcal fovea; mst, mesostipes; msvp, mesoventral intercoxal process; mtap, metaventral anterior process; mtfa, metafurcal arm; mtvp, metaventral intercoxal process; mxp, maxillary palp; nss, pronotosternal suture; occ, occipital constriction; pcc, procoxal cavity; pcs, procoxal socket; pff, profurcal fovea; pl, posterior lobe of mesocoxal projection; pre, prepectus; ppf, palpifer; psp, prosternal process; submentum; tm, temple; v 3, metaventrite" httpUri="https://zenodo.org/record/217857/files/figure.png" pageId="19" pageNumber="568">Fig. 26</figureCitation>
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). All other characters agree with those of
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<taxonomicName id="3AAC8BDAFF907371FBDBFEB2FB51428C" box="[1125,1248,368,392]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="568" phylum="Arthropoda" rank="genus">
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<emphasis id="CFD82C4BFF907371FBDBFEB2FB51428C" box="[1125,1248,368,392]" italics="true" pageId="19" pageNumber="568">Alloraphes</emphasis>
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</taxonomicName>
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, but basing only on a single female specimen it is not possible to clarify the taxonomic placement of this interesting and unusually large (BL
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<quantity id="3A545DBCFF907371FEAEFE75FEC942D4" box="[272,376,439,464]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="19" pageNumber="568" unit="mm" value="1.25">1.25 mm</quantity>
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) species. Examination of males and their genital characters may help solving this problem. Currently
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<taxonomicName id="3AAC8BDAFF907371FEB2FE1EFE3442F0" box="[268,389,476,500]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="568" phylum="Arthropoda" rank="species" species="magnus">
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<emphasis id="CFD82C4BFF907371FEB2FE1EFE3442F0" box="[268,389,476,500]" italics="true" pageId="19" pageNumber="568">A. magnus</emphasis>
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</taxonomicName>
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should be treated as a
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<emphasis id="CFD82C4BFF907371FD28FE1FFCDA42F0" box="[662,875,476,500]" italics="true" pageId="19" pageNumber="568">species inquirenda</emphasis>
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, genus
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<emphasis id="CFD82C4BFF907371FC79FE1FFBD442F0" box="[967,1125,476,500]" italics="true" pageId="19" pageNumber="568">incertae sedis</emphasis>
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within
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<taxonomicName id="3AAC8BDAFF907371FB7FFE1EFADB42F1" box="[1217,1386,476,501]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="568" phylum="Arthropoda" rank="tribe" tribe="Cyrtoscydmini">Cyrtoscydmini</taxonomicName>
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, but certainly close to
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<taxonomicName id="3AAC8BDAFF907371FEE3FDC2FE69411C" box="[349,472,512,536]" class="Insecta" family="Staphylinidae" genus="Alloraphes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="568" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="CFD82C4BFF907371FEE3FDC2FE69411C" box="[349,472,512,536]" italics="true" pageId="19" pageNumber="568">Alloraphes</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |