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<document id="FA6E012BF54B3D178133848F7055D7DA" ID-CLB-Dataset="270324" ID-DOI="10.5252/geodiversitas2023v45a17" ID-GBIF-Dataset="96784ccf-a749-4494-bfb4-f514eea7e824" ID-ISSN="1638-9395" ID-Zenodo-Dep="8399030" ID-ZooBank="urn:lsid:zoobank.org:pub:EE7AA253-A023-4011-B7E1-5820F8118ED3" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1695890445085" checkinUser="plazi" docAuthor="Béthoux, Olivier &amp; Anderson, John M." docDate="2023" docId="313072224B54FFA9FED1FF3296E3E7D2" docLanguage="en" docName="geodiversitas.2023.45.17.pdf" docOrigin="Geodiversitas 45 (17)" docSource="https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g2023v45a17.pdf" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Reisia guillaumei" docType="treatment" docVersion="2" lastPageNumber="486" masterDocId="CD090A5A4B5DFFA0FFB9FFE59428E17C" masterDocTitle="New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata)" masterLastPageNumber="496" masterPageNumber="479" pageNumber="486" updateTime="1696249958766" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="1F9FA92F71911F4561F391635DD2DBF4">New light shed on Triadophlebiomorpha wing morphology and systematics (Insecta: Odonata)</mods:title>
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<mods:namePart id="55E1D8DD8B437FF1BB655E0D7EE964AB">Béthoux, Olivier</mods:namePart>
<mods:affiliation id="627C349528D9C7C394D479AF5F762506">CR 2 P (CNRS, MNHN, Sorbonne Université), Muséum national dHistoire naturelle, case postale 48, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) obethoux @ mnhn. fr</mods:affiliation>
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<mods:affiliation id="AF452F45480F753840521C0FA97EFD6D">Evolutionary Studies Institute, Witwatersrand University, 1 Jan Smuts Ave., Braamfontein, Johannesburg 2000, South Africa jmanderson. gondwana @ gmail. com</mods:affiliation>
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<taxonomicName id="7E99B8B74B54FFA9FED1FF32960BE18F" authority="(Grauvogel &amp; Laurentiaux, 1952)" box="[360,547,215,243]" class="Insecta" family="Triadotypidae" genus="Reisia" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="486" phylum="Arthropoda" rank="species" species="guillaumei">
<emphasis id="8BED1F264B54FFA9FED1FF32960BE18F" bold="true" box="[360,547,215,243]" italics="true" pageId="9" pageNumber="486">Reisia guillaumei</emphasis>
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(
<bibRefCitation id="DD08BEC54B54FFA9FF66FF129617E06E" author="GRAUVOGEL L. &amp; LAURENTIAUX D." box="[223,575,247,274]" pageId="9" pageNumber="486" pagination="121 - 129" refId="ref14098" refString="GRAUVOGEL L. &amp; LAURENTIAUX D. 1952. - Un protodonate du trias des Vosges. Annales de Paleontologie, Invertebres 38: 121 - 129." type="journal article" year="1952">Grauvogel &amp; Laurentiaux, 1952</bibRefCitation>
), n. comb.
</paragraph>
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<paragraph id="B926C3344B54FFA9FE27FEF295C5E04D" blockId="9.[216,692,247,305]" box="[414,493,279,305]" pageId="9" pageNumber="486">
(
<figureCitation id="21A2DFB14B54FFA9FE1EFEF295CCE04D" box="[423,484,279,305]" captionStart="FIG" captionStartId="10.[224,235,1286,1303]" captionTargetBox="[258,1996,208,1246]" captionTargetId="figure-214@10.[258,1996,553,1240]" captionTargetPageId="10" captionText="FIG. 2. —Reisia guillaumei (Grauvogel &amp; Laurentiaux, 1952),n. comb.,holotype,specimen 7885 (collection Louis Grauvogel at the Staatlisches Museum für Naturkunde Stuttgart,Stuttgart,Germany), left wing: view; A, drawing of venation (large blue arrow indicating the only portion of RA preserved between the pons and the nodus; small green arrows indicating the termination of CuA branches); B, photograph of fragment, dorsal aspect (flipped horizontally); C, photograph of the distal fragment, dorsal aspect (flipped horizontally); D-F, detail of the wing base, location as shown on B; drawing of venation (enlargement from added color-coding); E, F, photographs extracted from RTI file (F, under normals visualisation; large white arrow indicating the end of the preserved section of the adjoined R and MA [or RA and RP+MA]; large row indicating the course of MP below the pons). Scale bars: A-C, 10 mm; D-F, 4 mm." figureDoi="http://doi.org/10.5281/zenodo.8399038" httpUri="https://zenodo.org/record/8399038/files/figure.png" pageId="9" pageNumber="486">Fig. 2</figureCitation>
)
</paragraph>
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<paragraph id="B926C3344B54FFA9FF3DFEB595F4E0FF" blockId="9.[131,775,336,387]" pageId="9" pageNumber="486">
<taxonomicName id="7E99B8B74B54FFA9FF3DFEB596EDE014" authority="Grauvovel &amp; Laurentiaux, 1952: 124" authorityName="Grauvovel &amp; Laurentiaux" authorityPageNumber="124" authorityYear="1952" box="[132,709,336,360]" class="Insecta" family="Triadotypidae" genus="Triadotypus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="486" phylum="Arthropoda" rank="species" species="guillaumei">
<emphasis id="8BED1F264B54FFA9FF3DFEB59572E014" box="[132,346,336,360]" italics="true" pageId="9" pageNumber="486">Triadotypus guillaumei</emphasis>
Grauvovel &amp; Laurentiaux, 1952: 124
</taxonomicName>
, figs 2, 3. —
<bibRefCitation id="DD08BEC54B54FFA9FF04FE8E95ABE0FF" author="NEL A. &amp; BETHOUX O. &amp; BECHLY G. &amp; MARTINEZ-DELCLOS X. &amp; PAPIER F." box="[189,387,363,387]" pageId="9" pageNumber="486" pagination="501 - 525" refId="ref14767" refString="NEL A., BETHOUX O., BECHLY G., MARTINEZ-DELCLOS X. &amp; PAPIER F. 2001. - The Permo-Triassic Odonatoptera of the &quot; protodonate &quot; grade (Insecta: Odonatoptera). Annales de la Societe entomologique de France (N. S.) 37: 501 - 525. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 6137381 x / f 87. image" type="journal article" year="2001">
Nel
<emphasis id="8BED1F264B54FFA9FF5EFE8E953EE0FF" box="[231,278,363,387]" italics="true" pageId="9" pageNumber="486">et al.</emphasis>
2001: 503
</bibRefCitation>
, figs 1-3.
</paragraph>
<paragraph id="B926C3344B54FFA9FF3DFE7E95FFE0B1" blockId="9.[132,774,411,461]" pageId="9" pageNumber="486">
<taxonomicName id="7E99B8B74B54FFA9FF3DFE7E95F2E0CF" authority="Laurentiaux, 1958: 37" authorityName="Laurentiaux" authorityPageNumber="37" authorityYear="1958" baseAuthorityName="Reis" baseAuthorityYear="1909" box="[132,474,411,435]" class="Insecta" family="Triadotypidae" genus="Reisia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="9" pageNumber="486" phylum="Arthropoda" rank="species" species="gelasii">
<emphasis id="8BED1F264B54FFA9FF3DFE7E94D2E0CF" box="[132,250,411,435]" italics="true" pageId="9" pageNumber="486">Reisia gelasii</emphasis>
<bibRefCitation id="DD08BEC54B54FFA9FF46FE7E95F2E0CF" author="LAURENTIAUX D." box="[255,474,411,435]" pageId="9" pageNumber="486" refId="ref14533" refString="LAURENTIAUX D. 1958. - Contribution a l'etude des insectes carboniferes. These Faculte des Sciences de l'Universite de Paris, Paris, 785 p., 46 pls." type="book" year="1958">Laurentiaux, 1958: 37</bibRefCitation>
</taxonomicName>
, text-figs 3, 4, pl. 1, fig. 2 (
<emphasis id="8BED1F264B54FFA9FD66FE7E948DE0B1" italics="true" pageId="9" pageNumber="486">partim</emphasis>
). —
<bibRefCitation id="DD08BEC54B54FFA9FF60FE509552E0B1" author="BECHLY G." box="[217,378,437,461]" pageId="9" pageNumber="486" pagination="31 - 70" refId="ref13178" refString="BECHLY G. 1997. - New fossil odonates from the Upper Triassic of Italy, with a redescription of Italophlebia gervasuttii, and a reclassification of Triassic dragonflies (Insecta: Odonata). Rivista del Museo civico di Scienze Naturali &quot; Enrico Caffi &quot; 19: 31 - 70." type="journal article" year="1997">Bechly 1997: 53</bibRefCitation>
(
<emphasis id="8BED1F264B54FFA9FE30FE5395E1E0B1" box="[393,457,438,461]" italics="true" pageId="9" pageNumber="486">partim</emphasis>
).
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. —
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<materialsCitation id="09F1C9694B54FFA9FE4CFE0095BDE31B" pageId="9" pageNumber="486" specimenCode="7885/7886" specimenCount="1" typeStatus="holotype">
and only known specimen
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(left wing in dorsal and ventral aspect, respectively; also referred to as 57.32 or 5732 in the literature; Louis Grauvogel collection at the Staatliches Museum für Naturkunde Stuttgart, Stuttgart, Germany;
<figureCitation id="21A2DFB14B54FFA9FEF6FDAA95A0E31B" box="[335,392,591,615]" captionStart="FIG" captionStartId="10.[224,235,1286,1303]" captionTargetBox="[258,1996,208,1246]" captionTargetId="figure-214@10.[258,1996,553,1240]" captionTargetPageId="10" captionText="FIG. 2. —Reisia guillaumei (Grauvogel &amp; Laurentiaux, 1952),n. comb.,holotype,specimen 7885 (collection Louis Grauvogel at the Staatlisches Museum für Naturkunde Stuttgart,Stuttgart,Germany), left wing: view; A, drawing of venation (large blue arrow indicating the only portion of RA preserved between the pons and the nodus; small green arrows indicating the termination of CuA branches); B, photograph of fragment, dorsal aspect (flipped horizontally); C, photograph of the distal fragment, dorsal aspect (flipped horizontally); D-F, detail of the wing base, location as shown on B; drawing of venation (enlargement from added color-coding); E, F, photographs extracted from RTI file (F, under normals visualisation; large white arrow indicating the end of the preserved section of the adjoined R and MA [or RA and RP+MA]; large row indicating the course of MP below the pons). Scale bars: A-C, 10 mm; D-F, 4 mm." figureDoi="http://doi.org/10.5281/zenodo.8399038" httpUri="https://zenodo.org/record/8399038/files/figure.png" pageId="9" pageNumber="486">Fig. 2</figureCitation>
).
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</paragraph>
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<collectionCode id="DF885BF14B54FFA9FEE8FD6195F5E3E9" box="[337,477,644,661]" pageId="9" pageNumber="486">STRATIGRAPHY</collectionCode>
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. — Bust (Bas-Rhin,
<collectingCountry id="C18E83A44B54FFA9FD01FD9A96D5E3EB" box="[696,765,639,663]" name="France" pageId="9" pageNumber="486">France</collectingCountry>
); Grès à Voltzia Formation; Anisian, Triassic; Gall &amp; GrauvogelStamm 2005).
</paragraph>
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<paragraph id="B926C3344B54FFA9FF3DFD01972EE24D" blockId="9.[130,775,740,817]" pageId="9" pageNumber="486">
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. — Width opposite the RP1/RP2 fork about
<quantity id="7E616ED14B54FFA9FD57FD019482E26A" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="9" pageNumber="486" unit="mm" value="22.0">22 mm</quantity>
(lower in
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<emphasis id="8BED1F264B54FFA9FEB7FD1A9552E26A" box="[270,378,767,790]" italics="true" pageId="9" pageNumber="486">Reisia nana</emphasis>
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);
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with 4 main posterior branches (in addition to its anterior stem; fewer branches in
<taxonomicName id="7E99B8B74B54FFA9FD8DFCFC9688E24D" authorityName="Béthoux &amp; Anderson" authorityYear="2023" baseAuthorityName="Bethoux, De la Horra, Benito, Barrenechea, Galan &amp; Lopez-Gomez" baseAuthorityYear="2009" box="[564,672,793,817]" class="Insecta" family="Triadotypidae" genus="Reisia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="486" phylum="Arthropoda" rank="species" species="rubra" status="n. comb.">
<emphasis id="8BED1F264B54FFA9FD8DFCFC9688E24D" box="[564,672,793,817]" italics="true" pageId="9" pageNumber="486">Reisia rubra</emphasis>
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).
</paragraph>
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<subSubSection id="F18390BF4B54FFA9FF3DFCBD96E3E7D2" pageId="9" pageNumber="486" type="discussion">
<paragraph id="B926C3344B54FFA9FF3DFCBD94C0E20C" blockId="9.[130,777,856,1711]" box="[132,232,856,882]" pageId="9" pageNumber="486">
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</paragraph>
<paragraph id="B926C3344B54FFA9FF3DFC9296E3E7D2" blockId="9.[130,777,856,1711]" pageId="9" pageNumber="486">
Our investigation revealed a number of discrepancies between previous descriptive accounts and what can be observed in the available material. Most importantly, the part that
<bibRefCitation id="DD08BEC54B54FFA9FD58FC529522E28D" author="NEL A. &amp; BETHOUX O. &amp; BECHLY G. &amp; MARTINEZ-DELCLOS X. &amp; PAPIER F." pageId="9" pageNumber="486" pagination="501 - 525" refId="ref14767" refString="NEL A., BETHOUX O., BECHLY G., MARTINEZ-DELCLOS X. &amp; PAPIER F. 2001. - The Permo-Triassic Odonatoptera of the &quot; protodonate &quot; grade (Insecta: Odonatoptera). Annales de la Societe entomologique de France (N. S.) 37: 501 - 525. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 6137381 x / f 87. image" type="journal article" year="2001">
Nel
<emphasis id="8BED1F264B54FFA9FF3DFC3D9491E28D" box="[132,185,983,1009]" italics="true" pageId="9" pageNumber="486">et al.</emphasis>
(2001)
</bibRefCitation>
regarded as the base of
<collectionCode id="DF885BF14B54FFA9FDB6FC329610E28D" box="[527,568,983,1009]" country="0" name="Mohonk Preserve, Inc." pageId="9" pageNumber="486">MP</collectionCode>
(and
<bibRefCitation id="DD08BEC54B54FFA9FDC3FC329571E56E" author="GRAUVOGEL L. &amp; LAURENTIAUX D." pageId="9" pageNumber="486" pagination="121 - 129" refId="ref14098" refString="GRAUVOGEL L. &amp; LAURENTIAUX D. 1952. - Un protodonate du trias des Vosges. Annales de Paleontologie, Invertebres 38: 121 - 129." type="journal article" year="1952">Grauvogel &amp; Laurentiaux [1952]</bibRefCitation>
as the base of CuP) turns out to be the pons, which is very long. Incidentally, it is strongly convex, which is inconsistent with an identification as (part of)
<collectionCode id="DF885BF14B54FFA9FF3FFBB29486E50D" box="[134,174,1111,1137]" country="0" name="Mohonk Preserve, Inc." pageId="9" pageNumber="486">MP</collectionCode>
. The course of the actual base of
<collectionCode id="DF885BF14B54FFA9FD9DFBB29663E50D" box="[548,587,1111,1137]" country="0" name="Mohonk Preserve, Inc." pageId="9" pageNumber="486">MP</collectionCode>
, although poorly preserved, yet is substantiated by a marked concave groove immediately posterior to the pons (black arrow on
<figureCitation id="21A2DFB14B54FFA9FD0BFB7394BEE5AC" captionStart="FIG" captionStartId="10.[224,235,1286,1303]" captionTargetBox="[258,1996,208,1246]" captionTargetId="figure-214@10.[258,1996,553,1240]" captionTargetPageId="10" captionText="FIG. 2. —Reisia guillaumei (Grauvogel &amp; Laurentiaux, 1952),n. comb.,holotype,specimen 7885 (collection Louis Grauvogel at the Staatlisches Museum für Naturkunde Stuttgart,Stuttgart,Germany), left wing: view; A, drawing of venation (large blue arrow indicating the only portion of RA preserved between the pons and the nodus; small green arrows indicating the termination of CuA branches); B, photograph of fragment, dorsal aspect (flipped horizontally); C, photograph of the distal fragment, dorsal aspect (flipped horizontally); D-F, detail of the wing base, location as shown on B; drawing of venation (enlargement from added color-coding); E, F, photographs extracted from RTI file (F, under normals visualisation; large white arrow indicating the end of the preserved section of the adjoined R and MA [or RA and RP+MA]; large row indicating the course of MP below the pons). Scale bars: A-C, 10 mm; D-F, 4 mm." figureDoi="http://doi.org/10.5281/zenodo.8399038" httpUri="https://zenodo.org/record/8399038/files/figure.png" pageId="9" pageNumber="486">Fig. 2E, F</figureCitation>
). Additionally, there is no evidence of a long RP+MA stem, as suggested by
<bibRefCitation id="DD08BEC54B54FFA9FE89FB3395CBE58C" author="NEL A. &amp; BETHOUX O. &amp; BECHLY G. &amp; MARTINEZ-DELCLOS X. &amp; PAPIER F." box="[304,483,1238,1264]" pageId="9" pageNumber="486" pagination="501 - 525" refId="ref14767" refString="NEL A., BETHOUX O., BECHLY G., MARTINEZ-DELCLOS X. &amp; PAPIER F. 2001. - The Permo-Triassic Odonatoptera of the &quot; protodonate &quot; grade (Insecta: Odonatoptera). Annales de la Societe entomologique de France (N. S.) 37: 501 - 525. https: // gallica. bnf. fr / ark: / 12148 / bpt 6 k 6137381 x / f 87. image" type="journal article" year="2001">
Nel
<emphasis id="8BED1F264B54FFA9FEE7FB3295BBE58C" box="[350,403,1238,1264]" italics="true" pageId="9" pageNumber="486">et al.</emphasis>
(2001)
</bibRefCitation>
. The corresponding area is not preserved in the side preserving the wing in dorsal aspect (
<figureCitation id="21A2DFB14B54FFA9FF34FAF394E4E44C" box="[141,204,1302,1328]" captionStart="FIG" captionStartId="10.[224,235,1286,1303]" captionTargetBox="[258,1996,208,1246]" captionTargetId="figure-214@10.[258,1996,553,1240]" captionTargetPageId="10" captionText="FIG. 2. —Reisia guillaumei (Grauvogel &amp; Laurentiaux, 1952),n. comb.,holotype,specimen 7885 (collection Louis Grauvogel at the Staatlisches Museum für Naturkunde Stuttgart,Stuttgart,Germany), left wing: view; A, drawing of venation (large blue arrow indicating the only portion of RA preserved between the pons and the nodus; small green arrows indicating the termination of CuA branches); B, photograph of fragment, dorsal aspect (flipped horizontally); C, photograph of the distal fragment, dorsal aspect (flipped horizontally); D-F, detail of the wing base, location as shown on B; drawing of venation (enlargement from added color-coding); E, F, photographs extracted from RTI file (F, under normals visualisation; large white arrow indicating the end of the preserved section of the adjoined R and MA [or RA and RP+MA]; large row indicating the course of MP below the pons). Scale bars: A-C, 10 mm; D-F, 4 mm." figureDoi="http://doi.org/10.5281/zenodo.8399038" httpUri="https://zenodo.org/record/8399038/files/figure.png" pageId="9" pageNumber="486">Fig. 2</figureCitation>
): only a very short section of the stem of RA (or R) is preserved, ending opposite the white arrow on
<figureCitation id="21A2DFB14B54FFA9FD21FAD3972CE42C" box="[664,772,1334,1360]" captionStart="FIG" captionStartId="10.[224,235,1286,1303]" captionTargetBox="[258,1996,208,1246]" captionTargetId="figure-214@10.[258,1996,553,1240]" captionTargetPageId="10" captionText="FIG. 2. —Reisia guillaumei (Grauvogel &amp; Laurentiaux, 1952),n. comb.,holotype,specimen 7885 (collection Louis Grauvogel at the Staatlisches Museum für Naturkunde Stuttgart,Stuttgart,Germany), left wing: view; A, drawing of venation (large blue arrow indicating the only portion of RA preserved between the pons and the nodus; small green arrows indicating the termination of CuA branches); B, photograph of fragment, dorsal aspect (flipped horizontally); C, photograph of the distal fragment, dorsal aspect (flipped horizontally); D-F, detail of the wing base, location as shown on B; drawing of venation (enlargement from added color-coding); E, F, photographs extracted from RTI file (F, under normals visualisation; large white arrow indicating the end of the preserved section of the adjoined R and MA [or RA and RP+MA]; large row indicating the course of MP below the pons). Scale bars: A-C, 10 mm; D-F, 4 mm." figureDoi="http://doi.org/10.5281/zenodo.8399038" httpUri="https://zenodo.org/record/8399038/files/figure.png" pageId="9" pageNumber="486">Fig. 2E, F</figureCitation>
. The splitting plane then runs at the level of ScP, which is well exposed, with a short section of MA visible (i.e. RA and RP are embedded in the other side of the specimen). Further on, where preserved, RA (blue arrow on
<figureCitation id="21A2DFB14B54FFA9FDBCFA50965DE4B3" box="[517,629,1461,1487]" captionStart="FIG" captionStartId="8.[132,143,1895,1912]" captionTargetBox="[132,1456,214,1821]" captionTargetId="figure-15@8.[132,1455,675,1821]" captionTargetPageId="8" captionText="FIG. 1. — Reisia gelasii (Reis, 1909), holotype, specimen SNSB-BSPG 1908 I 49 (Bayerische Staatssammlung für Paläontologie und Geologie, Munich, Germany), right wing in dorsal aspect: A, B, overview; A, drawing of venation (small blue arrow, ending of RP3+4; small green arrows indicating the termination of CuA branches); B, photograph; C-F, details, locations as shown on B, photographs extracted from RTI file (D, F, under normals visualisation); C, area posterior to RP3+4, the origins of paired convex and concave intercalaries (I+ and I-) become increasingly concave from the distal to the proximal areas; D, detail of the MP-CuA twilight zone (nature of structures encircled in white discussed in the text); E, F, detail of presumed setal bases (large white arrows indicating the best preserved ones). Scale bars: A, B, 10 mm; C, D, 5 mm; E, F, 2 mm." figureDoi="http://doi.org/10.5281/zenodo.8399036" httpUri="https://zenodo.org/record/8399036/files/figure.png" pageId="9" pageNumber="486">Fig. 1A, B</figureCitation>
), RP and MA are clearly distinct. Quite unfortunately the corresponding area is totally missing in the side preserving the wing in ventral aspect (7886), probably as a consequence of rock splitting at the time of collection. Given our current knowledge of the wing morphology in triadophlebiomorphans, it is rational to assume that RP and MA run next to each other, but distinct, in
<taxonomicName id="7E99B8B74B54FFA9FF18F970957BE7D2" authorityName="Béthoux &amp; Anderson" authorityYear="2023" baseAuthorityName="Grauvogel &amp; Laurentiaux" baseAuthorityYear="1952" box="[161,339,1684,1710]" class="Insecta" family="Triadotypidae" genus="Reisia" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="9" pageNumber="486" phylum="Arthropoda" rank="species" species="guillaumei" status="n. comb.">
<emphasis id="8BED1F264B54FFA9FF18F970957BE7D2" box="[161,339,1684,1710]" italics="true" pageId="9" pageNumber="486">Reisia guillaumei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="90DEA25D4B54FFA9FEE3F9709593E7D2" box="[346,443,1684,1711]" pageId="9" pageNumber="486" rank="species">n. comb.</taxonomicNameLabel>
(
<figureCitation id="21A2DFB14B54FFA9FE73F9719634E7D3" box="[458,540,1684,1711]" captionStart="FIG" captionStartId="10.[224,235,1286,1303]" captionTargetBox="[258,1996,208,1246]" captionTargetId="figure-214@10.[258,1996,553,1240]" captionTargetPageId="10" captionText="FIG. 2. —Reisia guillaumei (Grauvogel &amp; Laurentiaux, 1952),n. comb.,holotype,specimen 7885 (collection Louis Grauvogel at the Staatlisches Museum für Naturkunde Stuttgart,Stuttgart,Germany), left wing: view; A, drawing of venation (large blue arrow indicating the only portion of RA preserved between the pons and the nodus; small green arrows indicating the termination of CuA branches); B, photograph of fragment, dorsal aspect (flipped horizontally); C, photograph of the distal fragment, dorsal aspect (flipped horizontally); D-F, detail of the wing base, location as shown on B; drawing of venation (enlargement from added color-coding); E, F, photographs extracted from RTI file (F, under normals visualisation; large white arrow indicating the end of the preserved section of the adjoined R and MA [or RA and RP+MA]; large row indicating the course of MP below the pons). Scale bars: A-C, 10 mm; D-F, 4 mm." figureDoi="http://doi.org/10.5281/zenodo.8399038" httpUri="https://zenodo.org/record/8399038/files/figure.png" pageId="9" pageNumber="486">Fig. 2A</figureCitation>
; and see above).
</paragraph>
</subSubSection>
</treatment>
</document>