343 lines
36 KiB
XML
343 lines
36 KiB
XML
<document id="414F34CCCB21E4D05EA20C70B868A8AF" ID-DOI="10.11646/phytotaxa.345.3.3" ID-ISSN="1179-3163" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725592567195" checkinUser="felipe" docAuthor="Mesterházy, Attila, Jiménez-Mejías, Pedro, Pifkó, Dániel & Barina, Zoltán" docDate="2018" docId="0A10BF03FF85FFD136E5EB90FF6A947F" docLanguage="en" docName="phytotaxa.345.3.3.pdf" docOrigin="Phytotaxa 345 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.345.3.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Carex markgrafii Kük., Ber. Deutsch. Bot." docType="treatment" docVersion="1" lastPageNumber="277" masterDocId="F629C77BFF81FFD4366DE93DFF9C9616" masterDocTitle="Taxonomy, systematics, and typification of Carex markgrafii Kük. (Cyperaceae)" masterLastPageNumber="278" masterPageNumber="272" pageNumber="276" updateTime="1725601406343" updateUser="GgImagineBatch" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="59FDE9E4BE06971E4EDB0B8D9D2D6DA4">Taxonomy, systematics, and typification of Carex markgrafii Kük. (Cyperaceae)</mods:title>
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<mods:name id="0B5B7294E8F6B4D6355007B6D3CE1A91" type="personal">
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<mods:namePart id="63D89F5CECB3C330AD71AC738DD2F12C">Mesterházy, Attila</mods:namePart>
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<mods:affiliation id="E49B5FCD8D2DF2A99CEB5C18D2F5743B">Directorate of Hortobágy National Park H- 4024 Sumen utca 2., Debrecen, Hungary</mods:affiliation>
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<mods:namePart id="63200F8E3F54A3ECC4DF6740DF2BA8DB">Jiménez-Mejías, Pedro</mods:namePart>
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<mods:affiliation id="FF82B82A6CFAFEEE33B7F0D9774DC166">Department of Botany, National Museum of Natural History, Smithsonian Institution, PO Box 37012, Washington, DC 20013 - 7012.</mods:affiliation>
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</mods:name>
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<mods:namePart id="01162AD2A7ED54FA12D0DE5513DE4538">Pifkó, Dániel</mods:namePart>
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<mods:affiliation id="F4441A28C8A4E8441FB234DBE05BBBEE">Department of Botany, Hungarian Natural History Museum, H- 1431 Budapest, Pf. 137, Hungary</mods:affiliation>
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</mods:name>
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<mods:namePart id="BBF327BFEC3B373443BB52C046C16124">Barina, Zoltán</mods:namePart>
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<mods:affiliation id="9117247C79CA91F0318E915B1BC82809">Department of Botany, Hungarian Natural History Museum, H- 1431 Budapest, Pf. 137, Hungary</mods:affiliation>
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<mods:title id="232A9DF64B6D237F8A491DB0248A5E57">Phytotaxa</mods:title>
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<mods:part id="792E0C87D7AC0816F52C6998BBA947A5">
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<mods:date id="FB3DDAB7835121DAA5E2891C371D2941">2018</mods:date>
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<mods:number id="2897F9976E8BD74DBD0EFA3BAB2A90BC">2018-03-27</mods:number>
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<mods:number id="7DE713645EF93BE500BD912447A311DB">345</mods:number>
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<mods:url id="3612063D71B71E4D8D7367F1DEB73FC8">http://dx.doi.org/10.11646/phytotaxa.345.3.3</mods:url>
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<mods:identifier id="B8442019DA27C32399BBA71A4377687C" type="ISSN">1179-3163</mods:identifier>
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<treatment id="0A10BF03FF85FFD136E5EB90FF6A947F" LSID="urn:lsid:plazi:treatment:0A10BF03FF85FFD136E5EB90FF6A947F" httpUri="http://treatment.plazi.org/id/0A10BF03FF85FFD136E5EB90FF6A947F" lastPageId="5" lastPageNumber="277" pageId="4" pageNumber="276">
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<subSubSection id="CAA35D9EFF85FFD036E5EB90FD6C9518" pageId="4" pageNumber="276" type="nomenclature">
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<paragraph id="82060E15FF85FFD036E5EB90FCC794DE" blockId="4.[136,1452,685,854]" box="[136,859,685,712]" pageId="4" pageNumber="276">
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<heading id="D94EB979FF85FFD036E5EB90FCC794DE" bold="true" box="[136,859,685,712]" fontSize="11" level="1" pageId="4" pageNumber="276" reason="1">
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<emphasis id="B0CDD207FF85FFD036E5EB90FCC794DE" bold="true" box="[136,859,685,712]" pageId="4" pageNumber="276">
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<taxonomicName id="45B97596FF85FFD036E5EB90FDE594DE" ID-CoL="5XB3L" authority="Kuk., Ber. Deutsch. Bot." box="[136,633,685,712]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="276" phylum="Tracheophyta" rank="species" species="markgrafii">
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<emphasis id="B0CDD207FF85FFD036E5EB90FECD94DE" bold="true" box="[136,337,685,712]" italics="true" pageId="4" pageNumber="276">Carex markgrafii</emphasis>
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Kük., Ber. Deutsch. Bot.
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</taxonomicName>
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ges. 44: 420 (1926)
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="82060E15FF85FFD036E5EBE8FD6C9518" blockId="4.[136,1452,685,854]" pageId="4" pageNumber="276">
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<materialsCitation id="32D10448FF85FFD036E5EBE8FD6C9518" collectionCode="F" collectorName="Bemerkenswerte & Pflanzenarten & Albanien. Berichte der Deutschen Botanischen Gesellschaft & Tab" country="Albania" location="Markgraf" pageId="4" pageNumber="276" specimenCount="1" typeStatus="lectotype">
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<typeStatus id="5D02B0B7FF85FFD036E5EBE8FF7994FC" box="[136,229,725,746]" pageId="4" pageNumber="276" type="lectotype">Lectotype</typeStatus>
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</emphasis>
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(here designated):―
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<collectingCountry id="FAAE4E85FF85FFD037C0EBE8FD8094FC" box="[429,540,725,746]" name="Albania" pageId="4" pageNumber="276">ALBANIA</collectingCountry>
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.
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<location id="876658CEFF85FFD0344EEBE8FD1C94FC" LSID="urn:lsid:plazi:treatment:0A10BF03FF85FFD136E5EB90FF6A947F:876658CEFF85FFD0344EEBE8FD1C94FC" box="[547,640,725,746]" country="Albania" name="Markgraf" pageId="4" pageNumber="276">Markgraf</location>
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,
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<collectionCode id="E4A896D0FF85FFD034E5EBE8FD0A94FC" box="[648,662,725,746]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="4" pageNumber="276" type="Herbarium">F</collectionCode>
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. (1926):
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<collectorName id="2F4C6BC3FF85FFD03484EBE8FC1694FC" box="[745,906,725,746]" pageId="4" pageNumber="276">Bemerkenswerte</collectorName>
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neue
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<collectorName id="2F4C6BC3FF85FFD035ADEBE8FBD894FC" box="[960,1092,725,746]" pageId="4" pageNumber="276">Pflanzenarten</collectorName>
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aus
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<collectorName id="2F4C6BC3FF85FFD03206EBE8FE2D9518" pageId="4" pageNumber="276">Albanien. Berichte der Deutschen Botanischen Gesellschaft</collectorName>
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44(7): 420–432.
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<collectorName id="2F4C6BC3FF85FFD0343AEBC4FDE79518" box="[599,635,761,782]" pageId="4" pageNumber="276">Tab</collectorName>
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VII/VIII: 1.
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</paragraph>
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<subSubSection id="CAA35D9EFF85FFD036E5EA20FB1890AE" pageId="4" pageNumber="276" type="description">
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<paragraph id="82060E15FF85FFD036E5EA20FAFE9540" blockId="4.[136,1452,685,854]" pageId="4" pageNumber="276">
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<materialsCitation id="32D10448FF85FFD036E5EA20FAC29540" collectingDate="2014-05-07" collectionCode="A, BP, UPOS" collectorName="Mesterhazy & D. Pifko" country="Albania" county="Madh" elevation="1560" latitude="41.355053" location="Mali" longLatPrecision="1" longitude="20.175943" municipality="Maja Noj" pageId="4" pageNumber="276" specimenCount="2" typeStatus="Epitype">
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<emphasis id="B0CDD207FF85FFD036E5EA20FF4D9524" box="[136,209,797,818]" italics="true" pageId="4" pageNumber="276">
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<typeStatus id="5D02B0B7FF85FFD036E5EA20FF4D9524" box="[136,209,797,818]" pageId="4" pageNumber="276" type="epitype">Epitype</typeStatus>
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</emphasis>
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(here designated):―
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<collectingCountry id="FAAE4E85FF85FFD037CEEA20FD8E9524" box="[419,530,797,818]" name="Albania" pageId="4" pageNumber="276">ALBANIA</collectingCountry>
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.
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<location id="876658CEFF85FFD03472EA20FDD09524" LSID="urn:lsid:plazi:treatment:0A10BF03FF85FFD136E5EB90FF6A947F:876658CEFF85FFD03472EA20FDD09524" box="[543,588,797,818]" country="Albania" county="Madh" latitude="41.355053" longLatPrecision="1" longitude="20.175943" municipality="Maja Noj" name="Mali" pageId="4" pageNumber="276">Mali</location>
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me
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<location id="876658CEFF85FFD03411EA20FD269524" LSID="urn:lsid:plazi:treatment:0A10BF03FF85FFD136E5EB90FF6A947F:876658CEFF85FFD03411EA20FD269524" box="[636,698,797,818]" country="Albania" county="Madh" latitude="41.355053" longLatPrecision="1" longitude="20.175943" municipality="Maja Noj" name="Grope" pageId="4" pageNumber="276">Grope</location>
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,
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<location id="876658CEFF85FFD034A5EA20FC9C9524" LSID="urn:lsid:plazi:treatment:0A10BF03FF85FFD136E5EB90FF6A947F:876658CEFF85FFD034A5EA20FC9C9524" box="[712,768,797,818]" country="Albania" county="Madh" latitude="41.355053" longLatPrecision="1" longitude="20.175943" municipality="Maja Noj" name="South" pageId="4" pageNumber="276">South</location>
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||
hillside of
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||
<collectingMunicipality id="6262946FFF85FFD0351AEA20FC4A9524" box="[887,982,797,818]" pageId="4" pageNumber="276">Maja Noj</collectingMunicipality>
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i
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||
<collectingCounty id="6B677699FF85FFD0359DEA20FBB49524" box="[1008,1064,797,818]" pageId="4" pageNumber="276">Madh</collectingCounty>
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(Shen Ndue), closed calcareous rocky grassland, alt.
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||
<quantity id="4541A3F0FF85FFD03725EA7CFE0D9540" box="[328,401,833,854]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="4" pageNumber="276" unit="m" value="1560.0">
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<elevation id="0994E926FF85FFD03725EA7CFE0D9540" box="[328,401,833,854]" metricMagnitude="3" metricUnit="m" metricValue="1.56" pageId="4" pageNumber="276" unit="m" value="1560.0">1560 m</elevation>
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</quantity>
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<geoCoordinate id="E78D68D2FF85FFD037FAEA7CFDA89540" box="[407,564,833,854]" degrees="41" direction="north" minutes="21" orientation="latitude" pageId="4" pageNumber="276" precision="1" seconds="18.2" value="41.355053">N 41° 21’ 18.2’’</geoCoordinate>
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,
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<geoCoordinate id="E78D68D2FF85FFD03453EA7CFD499540" box="[574,725,833,854]" degrees="20" direction="east" minutes="10" orientation="longitude" pageId="4" pageNumber="276" precision="1" seconds="33.4" value="20.175943">E 20° 10’ 33.4”</geoCoordinate>
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,
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<date id="F60728D5FF85FFD034B2EA7CFCD29540" box="[735,846,833,854]" pageId="4" pageNumber="276" value="2014-05-07">
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<collectingDate id="E643D13DFF85FFD034B2EA7CFCD29540" box="[735,846,833,854]" pageId="4" pageNumber="276" value="2014-05-07">05.07.2014</collectingDate>
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</date>
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, leg.
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||
<collectionCode id="E4A896D0FF85FFD03512EA7CFC0F9540" box="[895,915,833,854]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="276" type="Herbarium">A</collectionCode>
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||
.
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||
<collectorName id="2F4C6BC3FF85FFD035F1EA7CFB909540" box="[924,1036,833,854]" pageId="4" pageNumber="276">Mesterházy</collectorName>
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||
&
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<collectorName id="2F4C6BC3FF85FFD03246EA7CFBE09540" box="[1067,1148,833,854]" pageId="4" pageNumber="276">D. Pifkó</collectorName>
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||
(s.n.) (
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||
<collectionCode id="E4A896D0FF85FFD032ACEA7CFB7E9540" box="[1217,1250,833,854]" country="Hungary" lsid="urn:lsid:biocol.org:col:13261" name="Hungarian Natural History Museum" pageId="4" pageNumber="276" type="Herbarium">BP</collectionCode>
|
||
;
|
||
<typeStatus id="5D02B0B7FF85FFD03281EA7CFA8C9540" box="[1260,1296,833,854]" pageId="4" pageNumber="276" type="isotype">iso-</typeStatus>
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||
<collectionCode id="E4A896D0FF85FFD0337BEA7CFAC49540" box="[1302,1368,833,854]" country="Spain" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15339" lsid="urn:lsid:biocol.org:col:15339" name="Universidad Pablo de Olavide" pageId="4" pageNumber="276" type="Herbarium">UPOS</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="82060E15FF85FFD036E5EABAFF7E95D2" blockId="4.[136,1452,903,2080]" pageId="4" pageNumber="276">
|
||
In the absence of the original type collection, we provide a detailed description of the materials designated as
|
||
<typeStatus id="5D02B0B7FF85FFD036E5EA96FF4295D2" box="[136,222,939,964]" pageId="4" pageNumber="276" type="epitype">epitype</typeStatus>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="82060E15FF85FFD036D0EAF2FC3C938E" blockId="4.[136,1452,903,2080]" pageId="4" pageNumber="276">
|
||
Perennial, laxly caespitose, with dark brown fibrous scales on short rhizomes. Stems erect, 20–30(–35) cm tall and
|
||
<quantity id="4541A3F0FF85FFD036D7EACEFF61921A" box="[186,253,1011,1036]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="276" unit="mm" value="1.0">1 mm</quantity>
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||
wide, trigonous, smooth, becoming more or less scabrid towards the apex. Sheaths of basal leaves dark brown, entire, without blade, shiny, fibrous. Leaves
|
||
<quantity id="4541A3F0FF85FFD034BEED2AFCAD9226" box="[723,817,1047,1072]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="4" pageNumber="276" unit="mm" value="3.0" valueMax="4.0" valueMin="2.0">2–4 mm</quantity>
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||
wide, flat, tapering towards apex, shorter than the stem; ligule
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||
<quantity id="4541A3F0FF85FFD036A2ED06FEB79242" box="[207,299,1083,1108]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="4" pageNumber="276" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1–2 mm</quantity>
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||
long, shorter than wide, ovate-lanceolate, obtuse. Inflorescence
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||
<quantity id="4541A3F0FF85FFD03596ED06FBFF9242" box="[1019,1123,1083,1108]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.5" metricValueMin="2.0" pageId="4" pageNumber="276" unit="cm" value="2.75" valueMax="3.5" valueMin="2.0">2–3.5 cm</quantity>
|
||
long, with one terminal male spike, and 2–3 lateral female spikes. Male spike 13–22 ×
|
||
<quantity id="4541A3F0FF85FFD0357AED62FCEB926E" box="[791,887,1119,1144]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="4" pageNumber="276" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">3–5 mm</quantity>
|
||
, cylindrical, sessile. Lowermost bract
|
||
<quantity id="4541A3F0FF85FFD03346ED62FA3B926E" box="[1323,1447,1119,1144]" metricMagnitude="-2" metricUnit="m" metricValue="1.75" metricValueMax="2.5" metricValueMin="1.0" pageId="4" pageNumber="276" unit="mm" value="17.5" valueMax="25.0" valueMin="10.0">10–25 mm</quantity>
|
||
, lanceolate to subulate, rarely longer than its spike. Female spike 8–13 ×
|
||
<quantity id="4541A3F0FF85FFD035D9EDBEFB8F928A" box="[948,1043,1155,1180]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="4" pageNumber="276" unit="mm" value="3.5" valueMax="4.0" valueMin="3.0">3–4 mm</quantity>
|
||
, ovoid to obovoid or cylindrical, the uppermost sessile, arising next to the male spike, the lowermost a bit more distant, with internode
|
||
<quantity id="4541A3F0FF85FFD032BFED9AFAA192D6" box="[1234,1341,1191,1216]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" metricValueMax="10.0" metricValueMin="6.0" pageId="4" pageNumber="276" unit="mm" value="8.0" valueMax="10.0" valueMin="6.0">6–10 mm</quantity>
|
||
long, and with a peduncle
|
||
<quantity id="4541A3F0FF85FFD0372FEDF6FE3292F2" box="[322,430,1227,1252]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="10.0" metricValueMin="3.0" pageId="4" pageNumber="276" unit="mm" value="6.5" valueMax="10.0" valueMin="3.0">3–10 mm</quantity>
|
||
long. Androgynous spike absent. Male glumes 3–5 ×
|
||
<quantity id="4541A3F0FF85FFD0327CEDF6FBC892F2" box="[1041,1108,1227,1252]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="276" unit="mm" value="1.0">1 mm</quantity>
|
||
, elliptical, with subulate apex, dark brown with a pale midrib, without membranous margins. Female glumes 3.2–4.5 (incl.
|
||
<quantity id="4541A3F0FF85FFD032E1EDD2FB61931E" box="[1164,1277,1263,1288]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="4" pageNumber="276" unit="mm" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 mm</quantity>
|
||
awn) ×
|
||
<quantity id="4541A3F0FF85FFD03335EDD2FF28933A" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.8" metricValueMin="1.5" pageId="4" pageNumber="276" unit="mm" value="1.65" valueMax="1.8" valueMin="1.5">1.5–1.8 mm</quantity>
|
||
ovoid to broadly ovoid with long arista, longer than the utricle, dark brown with a pale midrib and 3 (pairs of) inconspicuous lateral veins. Stigmas 3. Utricles 3–4 ×
|
||
<quantity id="4541A3F0FF85FFD03485EC0AFCB79346" box="[744,811,1335,1360]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="276" unit="mm" value="1.0">1 mm</quantity>
|
||
, pyriform, light green to light brown, with inconspicuous veins; beak short or long, dark brown with dense short hairs when young, glabrescent when mature. Nutlet 1.6 ×
|
||
<quantity id="4541A3F0FF85FFD033E5EC66FF2A938E" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="4" pageNumber="276" unit="mm" value="0.8">0.8 mm</quantity>
|
||
, light brown; style-base enlarged forming a ring-like stylopodium.
|
||
</paragraph>
|
||
<paragraph id="82060E15FF85FFD036D0EC9EFA889082" blockId="4.[136,1452,903,2080]" pageId="4" pageNumber="276">
|
||
<emphasis id="B0CDD207FF85FFD036D0EC9EFE7A93AA" bold="true" box="[189,486,1443,1468]" pageId="4" pageNumber="276">Habitat and distribution</emphasis>
|
||
:―
|
||
<taxonomicName id="45B97596FF85FFD03464EC9EFD5193AA" box="[521,717,1443,1468]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="276" phylum="Tracheophyta" rank="species" species="markgrafii">
|
||
<emphasis id="B0CDD207FF85FFD03464EC9EFD5193AA" box="[521,717,1443,1468]" italics="true" pageId="4" pageNumber="276">Carex markgrafii</emphasis>
|
||
</taxonomicName>
|
||
is known only from the
|
||
<typeStatus id="5D02B0B7FF85FFD03587EC9EFB8693AA" box="[1002,1050,1443,1468]" pageId="4" pageNumber="276">type</typeStatus>
|
||
locality in Central
|
||
<collectingCountry id="FAAE4E85FF85FFD03295EC9EFAC893AA" box="[1272,1364,1443,1468]" name="Albania" pageId="4" pageNumber="276">Albania</collectingCountry>
|
||
, east of
|
||
<collectingRegion id="407DC0F7FF85FFD036E5ECFAFF4F93F6" box="[136,211,1479,1504]" country="Albania" name="Tirane" pageId="4" pageNumber="276">Tirana</collectingRegion>
|
||
, on the Shen Ndue (mali i Sënoit) mount (
|
||
<quantity id="4541A3F0FF85FFD034F2ECFAFD6B93F6" box="[671,759,1479,1504]" metricMagnitude="3" metricUnit="m" metricValue="1.848" pageId="4" pageNumber="276" unit="m" value="1848.0">1848 m</quantity>
|
||
), the highest peak of a limestone karst area of circa.
|
||
<quantity id="4541A3F0FF85FFD03359ECFAFA1A93F6" box="[1332,1414,1479,1504]" metricMagnitude="5" metricUnit="m" metricValue="1.1" pageId="4" pageNumber="276" unit="km" value="110.0">110 km</quantity>
|
||
<superScript id="75CCA35DFF85FFD033EBECF8FA1293C5" attach="left" box="[1414,1422,1477,1491]" fontSize="6" pageId="4" pageNumber="276">2</superScript>
|
||
. It seems to be restricted to the southern slopes, in open rocky grasslands dominated by
|
||
<taxonomicName id="45B97596FF85FFD0325CECD6FB579012" box="[1073,1227,1515,1540]" class="Liliopsida" family="Poaceae" genus="Stipa" kingdom="Plantae" order="Poales" pageId="4" pageNumber="276" phylum="Tracheophyta" rank="species" species="pennata">
|
||
<emphasis id="B0CDD207FF85FFD0325CECD6FB579012" box="[1073,1227,1515,1540]" italics="true" pageId="4" pageNumber="276">Stipa pennata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="45B97596FF85FFD03292ECD6FA309012" box="[1279,1452,1515,1540]" class="Liliopsida" family="Poaceae" genus="Bromus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="276" phylum="Tracheophyta" rank="species" species="erectus">
|
||
<emphasis id="B0CDD207FF85FFD03292ECD6FA309012" box="[1279,1452,1515,1540]" italics="true" pageId="4" pageNumber="276">Bromus erectus</emphasis>
|
||
</taxonomicName>
|
||
vegetation (Buzo 1990). Other dominant species of herbs characteristic in this stand are
|
||
<taxonomicName id="45B97596FF85FFD03204EF32FAB5903E" authorityName="Asso" authorityYear="1779" box="[1129,1321,1551,1576]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="276" phylum="Tracheophyta" rank="species" species="halleriana">
|
||
<emphasis id="B0CDD207FF85FFD03204EF32FAB5903E" box="[1129,1321,1551,1576]" italics="true" pageId="4" pageNumber="276">Carex halleriana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="45B97596FF85FFD0335AEF32FE9F905A" class="Magnoliopsida" family="Caryophyllaceae" genus="Saponaria" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="276" phylum="Tracheophyta" rank="species" species="bellidifolia">
|
||
<emphasis id="B0CDD207FF85FFD0335AEF32FE9F905A" italics="true" pageId="4" pageNumber="276">Saponaria bellidifolia</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B0CDD207FF85FFD03778EF0EFB27905A" box="[277,1211,1587,1612]" italics="true" pageId="4" pageNumber="276">
|
||
<taxonomicName id="45B97596FF85FFD03778EF0EFDFE905A" authority=", Asyneuma" authorityName="Asyneuma" box="[277,610,1587,1612]" class="Liliopsida" family="Asphodelaceae" genus="Asphodelus" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="276" phylum="Tracheophyta" rank="species" species="albus">Asphodelus albus, Asyneuma</taxonomicName>
|
||
limonifolium,
|
||
<taxonomicName id="45B97596FF85FFD03560EF0EFC6B905A" box="[781,1015,1587,1612]" class="Liliopsida" family="Cyperaceae" genus="Sedum" kingdom="Plantae" order="Poales" pageId="4" pageNumber="276" phylum="Tracheophyta" rank="species" species="ochroleucum">Sedum ochroleucum</taxonomicName>
|
||
,
|
||
<taxonomicName id="45B97596FF85FFD0326BEF0EFB27905A" box="[1030,1211,1587,1612]" class="Liliopsida" family="Cyperaceae" genus="Carduus" kingdom="Plantae" order="Poales" pageId="4" pageNumber="276" phylum="Tracheophyta" rank="species" species="nutans">Carduus nutans</taxonomicName>
|
||
</emphasis>
|
||
agg.,
|
||
<emphasis id="B0CDD207FF85FFD0336BEF0EFF469066" italics="true" pageId="4" pageNumber="276">
|
||
<taxonomicName id="45B97596FF85FFD0336BEF0EFF499066" class="Magnoliopsida" family="Cistaceae" genus="Helianthemum" kingdom="Plantae" order="Malvales" pageId="4" pageNumber="276" phylum="Tracheophyta" rank="species" species="canum">Helianthemum canum</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
etc. This grassland community is widespread in the mountains of
|
||
<collectingCountry id="FAAE4E85FF85FFD035D7EF6AFB8A9066" box="[954,1046,1623,1648]" name="Albania" pageId="4" pageNumber="276">Albania</collectingCountry>
|
||
, and most of the above cited species are too.
|
||
<taxonomicName id="45B97596FF85FFD0368EEF46FE469082" box="[227,474,1659,1684]" class="Magnoliopsida" family="Caryophyllaceae" genus="Saponaria" kingdom="Plantae" order="Caryophyllales" pageId="4" pageNumber="276" phylum="Tracheophyta" rank="species" species="bellidifolia">
|
||
<emphasis id="B0CDD207FF85FFD0368EEF46FE469082" box="[227,474,1659,1684]" italics="true" pageId="4" pageNumber="276">Saponaria bellidifolia</emphasis>
|
||
</taxonomicName>
|
||
is the rarest plant, and it is an indicator of the richest plots (Barina ined.).
|
||
</paragraph>
|
||
<paragraph id="82060E15FF85FFD036D0EFA2FB1890AE" blockId="4.[136,1452,903,2080]" box="[189,1156,1695,1720]" pageId="4" pageNumber="276">
|
||
<emphasis id="B0CDD207FF85FFD036D0EFA2FEA590AE" bold="true" box="[189,313,1695,1720]" pageId="4" pageNumber="276">Phenology</emphasis>
|
||
:―The species is known in flower and fruit in the months of June and July.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="CAA35D9EFF85FFD136D0EFFEFF6A947F" lastPageId="5" lastPageNumber="277" pageId="4" pageNumber="276" type="etymology">
|
||
<paragraph id="82060E15FF85FFD036D0EFFEFCEE9116" blockId="4.[136,1452,903,2080]" pageId="4" pageNumber="276">
|
||
<emphasis id="B0CDD207FF85FFD036D0EFFEFEA290CA" bold="true" box="[189,318,1731,1756]" pageId="4" pageNumber="276">Etymology</emphasis>
|
||
:―Named by Kükenthal after the collector of the original
|
||
<typeStatus id="5D02B0B7FF85FFD035BAEFFEFB9B90CA" box="[983,1031,1731,1756]" pageId="4" pageNumber="276">type</typeStatus>
|
||
material, Friedrich Markgraf (1897– 1987), German botanist devoted to the study of the Albanian flora.
|
||
</paragraph>
|
||
<paragraph id="82060E15FF85FFD036D0EE36FC649E36" blockId="4.[136,1452,903,2080]" pageId="4" pageNumber="276">
|
||
<emphasis id="B0CDD207FF85FFD036D0EE36FEF99132" bold="true" box="[189,357,1803,1828]" pageId="4" pageNumber="276">Observations:</emphasis>
|
||
―
|
||
<materialsCitation id="32D10448FF85FFD037ECEE36FBA29186" collectionCode="WWII, ICN" collectorName="The" pageId="4" pageNumber="276" specimenCount="1" typeStatus="lectotype">
|
||
The only specimen mentioned in the description of
|
||
<taxonomicName id="45B97596FF85FFD035D2EE36FB1D9132" box="[959,1153,1803,1828]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="276" phylum="Tracheophyta" rank="species" species="markgrafii">
|
||
<emphasis id="B0CDD207FF85FFD035D2EE36FB1D9132" box="[959,1153,1803,1828]" italics="true" pageId="4" pageNumber="276">Carex markgrafii</emphasis>
|
||
</taxonomicName>
|
||
has been destroyed during
|
||
<collectionCode id="E4A896D0FF85FFD036E5EE12FF4F915E" box="[136,211,1839,1864]" pageId="4" pageNumber="276">WWII</collectionCode>
|
||
. In the protologue, a photo is provided, which can be considered as part of the original material according to Art. 9.3 (
|
||
<collectionCode id="E4A896D0FF85FFD036D6EE6EFF73917A" box="[187,239,1875,1900]" country="Colombia" httpUri="http://grbio.org/cool/xsgz-nxkh" name="Instituto de Ciencias Naturales, Museo de Historia Natural" pageId="4" pageNumber="276">ICN</collectionCode>
|
||
,
|
||
<bibRefCitation id="E62873E4FF85FFD03696EE6EFE42917A" author="McNeill, J. & Barrie, F. R. & Buck, W. R. & Demoulin, V. & Greuter, D. L. & Hawksworth, D. L. & Herendeen, P. S. & Knapp, S. & Marhold, K. & Prado, J. & Proud'Homme van Reine, W. F. & Smith, J. F. & Wiersema, J. H." box="[251,478,1875,1900]" pageId="4" pageNumber="276" pagination="1 - 208" refId="ref4731" refString="McNeill, J., Barrie, F. R., Buck, W. R., Demoulin, V., Greuter, D. L., Hawksworth, D. L., Herendeen, P. S., Knapp, S., Marhold, K., Prado, J., Proud'Homme van Reine, W. F., Smith, J. F. & Wiersema, J. H. (Eds.) (2012) International Code of Nomenclature for algae, fungi and plants (Melbourne Code): Adopted by the Eighteenth International Botanical Congress, Melbourne, Australia, July 2011. Regnum Vegetabile 154: 1 - 208." type="journal article" year="2012">
|
||
McNeill
|
||
<emphasis id="B0CDD207FF85FFD0370DEE6EFE0E917A" box="[352,402,1875,1900]" italics="true" pageId="4" pageNumber="276">et al</emphasis>
|
||
. 2012
|
||
</bibRefCitation>
|
||
). Accordingly, we propose to use it for the typification of the name and designate it as
|
||
<typeStatus id="5D02B0B7FF85FFD036E5EE4AFF6E9186" box="[136,242,1911,1936]" pageId="4" pageNumber="276" type="lectotype">lectotype</typeStatus>
|
||
.
|
||
<collectorName id="2F4C6BC3FF85FFD03691EE4AFEB49186" box="[252,296,1911,1936]" pageId="4" pageNumber="276">The</collectorName>
|
||
published photo represents the general appearance of the species well
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="32D10448FF85FFD03224EE4AFC689E36" collectionCode="A, BP" collectorName="Mesterhazy & D. Pifko & In" pageId="4" pageNumber="276" specimenCount="2" typeStatus="lectotype">
|
||
however, details on utricles and glumes are not provided, characters that are essential in
|
||
<taxonomicName id="45B97596FF85FFD03497EEA6FCA191A2" box="[762,829,1947,1972]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="4" pageNumber="276" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B0CDD207FF85FFD03497EEA6FCA191A2" box="[762,829,1947,1972]" italics="true" pageId="4" pageNumber="276">Carex</emphasis>
|
||
</taxonomicName>
|
||
taxonomy. To overcome these problems, we proceed to complete the
|
||
<typeStatus id="5D02B0B7FF85FFD03771EE82FE1F91CE" box="[284,387,1983,2008]" pageId="4" pageNumber="276" type="lectotype">lectotype</typeStatus>
|
||
specimen with an
|
||
<typeStatus id="5D02B0B7FF85FFD0343DEE82FD3991CE" box="[592,677,1983,2008]" pageId="4" pageNumber="276" type="epitype">epitype</typeStatus>
|
||
, a well-preserved complete herbarium specimen collected in the locus classicus (
|
||
<collectionCode id="E4A896D0FF85FFD03691EEDEFE8891EA" box="[252,276,2019,2044]" country="USA" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="276" type="Herbarium">A</collectionCode>
|
||
.
|
||
<collectorName id="2F4C6BC3FF85FFD03772EEDEFE3E91EA" box="[287,418,2019,2044]" pageId="4" pageNumber="276">Mesterházy</collectorName>
|
||
&
|
||
<collectorName id="2F4C6BC3FF85FFD037ABEEDEFDB491EA" box="[454,552,2019,2044]" pageId="4" pageNumber="276">D. Pifkó</collectorName>
|
||
, s.n.;
|
||
<collectionCode id="E4A896D0FF85FFD03407EEDEFD0C91EA" box="[618,656,2019,2044]" country="Hungary" lsid="urn:lsid:biocol.org:col:13261" name="Hungarian Natural History Museum" pageId="4" pageNumber="276" type="Herbarium">BP</collectionCode>
|
||
; iso-UPOS).
|
||
<collectorName id="2F4C6BC3FF85FFD03549EEDEFCA791EA" box="[804,827,2019,2044]" pageId="4" pageNumber="276">In</collectorName>
|
||
addition, the selected voucher contain a plant fragment that is in complete agreement with the information provided in the protologue
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="82060E15FF84FFD136D0E99DFD2297CF" blockId="5.[136,1452,160,617]" pageId="5" pageNumber="277">
|
||
An additional collection, determined as
|
||
<taxonomicName id="45B97596FF84FFD1341EE99DFC9696AF" box="[627,778,160,185]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="5" pageNumber="277" phylum="Tracheophyta" rank="species" species="markgrafii">
|
||
<emphasis id="B0CDD207FF84FFD1341EE99DFC9696AF" box="[627,778,160,185]" italics="true" pageId="5" pageNumber="277">C. markgrafii</emphasis>
|
||
</taxonomicName>
|
||
by Boris Achtarov, was located at SOM herbarium. Collector details are not given on the label, however, based on the date of collection, it could have been collected by Albanian botanists (Kole Paparisto, Xhafer Qosja or their contemporaries). These specimens were collected on
|
||
<date id="F60728D5FF84FFD13364E9D5FA169717" box="[1289,1418,232,257]" pageId="5" pageNumber="277" value="1955-03-06">03.06.1955</date>
|
||
. in
|
||
<collectingCountry id="FAAE4E85FF84FFD136E5E831FF219733" box="[136,189,268,293]" name="Serbia" pageId="5" pageNumber="277">Mali</collectingCountry>
|
||
me Gropë, which is app.
|
||
<quantity id="4541A3F0FF84FFD13785E831FDAD9733" box="[488,561,268,293]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="277" unit="km" value="10.0">10 km</quantity>
|
||
west from the locus classicus of
|
||
<taxonomicName id="45B97596FF84FFD135DCE831FBD09733" box="[945,1100,268,293]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="5" pageNumber="277" phylum="Tracheophyta" rank="species" species="markgrafii">
|
||
<emphasis id="B0CDD207FF84FFD135DCE831FBD09733" box="[945,1100,268,293]" italics="true" pageId="5" pageNumber="277">C. markgrafii</emphasis>
|
||
</taxonomicName>
|
||
. These specimens have obtuse male glumes, with white margins and ovoid spikes, features which do not match
|
||
<taxonomicName id="45B97596FF84FFD1324BE80DFB5D975F" box="[1062,1217,304,329]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="5" pageNumber="277" phylum="Tracheophyta" rank="species" species="markgrafii">
|
||
<emphasis id="B0CDD207FF84FFD1324BE80DFB5D975F" box="[1062,1217,304,329]" italics="true" pageId="5" pageNumber="277">C. markgrafii</emphasis>
|
||
</taxonomicName>
|
||
. In our opinion, this collection represents a deviant plant of
|
||
<taxonomicName id="45B97596FF84FFD1342CE869FD4B977B" authorityName="Asso" authorityYear="1779" box="[577,727,340,365]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="5" pageNumber="277" phylum="Tracheophyta" rank="species" species="halleriana">
|
||
<emphasis id="B0CDD207FF84FFD1342CE869FD4B977B" box="[577,727,340,365]" italics="true" pageId="5" pageNumber="277">C. halleriana</emphasis>
|
||
</taxonomicName>
|
||
, with the lowermost spike pedunculate and its bract leaf-like (as reported in
|
||
<taxonomicName id="45B97596FF84FFD13764E845FD169787" authority="Kuk." authorityName="Kuk." box="[265,650,376,401]" class="Liliopsida" family="Cyperaceae" form="pedunculata" genus="Carex" kingdom="Plantae" order="Poales" pageId="5" pageNumber="277" phylum="Tracheophyta" rank="form" species="halleriana">
|
||
<emphasis id="B0CDD207FF84FFD13764E845FE039787" box="[265,415,376,401]" italics="true" pageId="5" pageNumber="277">C. halleriana</emphasis>
|
||
f.
|
||
<emphasis id="B0CDD207FF84FFD137D3E845FDD49787" box="[446,584,376,401]" italics="true" pageId="5" pageNumber="277">pedunculata</emphasis>
|
||
Kük.
|
||
</taxonomicName>
|
||
), and without the basal, long-pedunculate spikes typical of this species.
|
||
<taxonomicName id="45B97596FF84FFD136E5E8A1FED597A3" authorityName="Asso" authorityYear="1779" box="[136,329,412,437]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="5" pageNumber="277" phylum="Tracheophyta" rank="species" species="halleriana">
|
||
<emphasis id="B0CDD207FF84FFD136E5E8A1FED597A3" box="[136,329,412,437]" italics="true" pageId="5" pageNumber="277">Carex halleriana</emphasis>
|
||
</taxonomicName>
|
||
is actually the most widespread species in the central mountain region of
|
||
<collectingCountry id="FAAE4E85FF84FFD132FEE8A1FB7397A3" box="[1171,1263,412,437]" name="Albania" pageId="5" pageNumber="277">Albania</collectingCountry>
|
||
, which at higher altitudes can be collected at the beginning of June.
|
||
</paragraph>
|
||
<paragraph id="82060E15FF84FFD136D0E8D9FF6A947F" blockId="5.[136,1452,160,617]" pageId="5" pageNumber="277">
|
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<bibRefCitation id="E62873E4FF84FFD136D0E8D9FEC597EB" author="Chater, A. O." box="[189,345,484,509]" pageId="5" pageNumber="277" pagination="290 - 323" refId="ref4128" refString="Chater, A. O. (1980) Carex L. In: Flora Europaea V. Cambridge University Press, Cambridge etc., pp. 290 - 323." type="book chapter" year="1980">Chater (1980)</bibRefCitation>
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reported as a character for
|
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<taxonomicName id="45B97596FF84FFD134E7E8D9FCBF97EB" box="[650,803,484,509]" class="Liliopsida" family="Cyperaceae" genus="Carex" kingdom="Plantae" order="Poales" pageId="5" pageNumber="277" phylum="Tracheophyta" rank="species" species="markgrafii">
|
||
<emphasis id="B0CDD207FF84FFD134E7E8D9FCBF97EB" box="[650,803,484,509]" italics="true" pageId="5" pageNumber="277">C. markgrafii</emphasis>
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</taxonomicName>
|
||
the “lowest bract leaflike, up to as long as inflorescence”. We were unable to confirm such character after the examination of our collections. Indeed, this character was also not mentioned in the protologue (
|
||
<bibRefCitation id="E62873E4FF84FFD137B8EB11FD179453" author="Markgraf, F." box="[469,651,556,581]" pageId="5" pageNumber="277" pagination="1 - 217" refId="ref4702" refString="Markgraf, F. (1926) An den Grenzen des Mittelmeergebiets. Pflanzengeographie von Mittelalbanien. Repertorium Specierum Novarum Regni Vegetabilis Beihefte 45: 1 - 217." type="journal article" year="1926">Markgraf, 1926</bibRefCitation>
|
||
). It points that Chater’s report might be perhaps based on misidentified materials.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
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</document> |