156 lines
17 KiB
XML
156 lines
17 KiB
XML
<document id="F230C9C7E73ABAEAFC64D700E4A892BE" ID="10.11646/zootaxa.4175.2.1" ID-DOI="10.11646/zootaxa.4175.2.1" ID-GBIF-Dataset="d7cbc23a-7a4c-4616-83fd-059fb458f7b6" ID-ISSN="1175-5326" ID-Zenodo-Dep="257286" ID-ZooBank="B264C215-000D-42C5-8AC9-B801872CD182" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1476339245459" checkinUser="plazi" docAuthor="Mah, Christopher L." docDate="2016" docId="953787B9FF93FFF45CF0FC398648F946" docLanguage="en" docName="zootaxa.4175.2.1.pdf" docOrigin="Zootaxa 4175 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Sibogaster Doderlein 1924" docType="treatment" docVersion="7" lastPageNumber="129" masterDocId="690EFFC1FF8FFFE85C67FF96857BFFB8" masterDocTitle="Deep-sea (> 1000 m) Goniasteridae (Valvatida; Asteroidea) from the North Pacific, including an overview of Sibogaster, Bathyceramaster n. gen. and three new species" masterLastPageNumber="141" masterPageNumber="101" pageNumber="129" updateTime="1698389676701" updateUser="plazi">
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<mods:mods id="3A39A9C20270B5717D2B1960812916E1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="DF48C06D3459E6D0C19C8EDB77FC2019">
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<mods:title id="680CEF7AB9AB1B10436718534A5E92A5">Deep-sea (> 1000 m) Goniasteridae (Valvatida; Asteroidea) from the North Pacific, including an overview of Sibogaster, Bathyceramaster n. gen. and three new species</mods:title>
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</mods:titleInfo>
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<mods:name id="5AC5EB93B822D68B6B73EB527F8D8B78" type="personal">
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<mods:role id="107712022ADA7BB0F0B0CBD3533857EA">
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<mods:roleTerm id="CAD8FF00C77789638186BA50E52E239D">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A501360AF5B3CDCB9573CCADD252F288">Mah, Christopher L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="3C1429FEDA095F7B9DB568DC3A67973D">text</mods:typeOfResource>
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<mods:titleInfo id="0446AB8D8486FB6AD8447B1645361E07">
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<mods:title id="C555D5C21B4B32262127DBB2785CC1AA">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="FD531F239AF9F9F4B94E40F984483619">
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<mods:date id="BF35AE45EF36BD65084B989074BBE694">2016</mods:date>
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<mods:detail id="8538AAE3F06750EEAC8B80E06FA2D6C6" type="volume">
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<mods:number id="1C9938ABBB8EAE1DDE1E537BD4CFEF84">4175</mods:number>
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</mods:detail>
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<mods:detail id="B016786368926640346C10D57AC239D6" type="issue">
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<mods:number id="85AB634AC9122ED1F0CF0E4109310CB3">2</mods:number>
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<mods:extent id="595120F050FB300D5988979C1D339C16" unit="page">
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<mods:start id="FBA58BF3697E8C930199CE2CB649CA40">101</mods:start>
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<mods:end id="00CBF6573614EEFE1361EE876693965A">141</mods:end>
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<mods:classification id="D8EEE25F42BDC8BDD515CF97E3F605C8">journal article</mods:classification>
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<mods:identifier id="A12A5133A9A553B5A778DD0BE30DEFDF" type="DOI">10.11646/zootaxa.4175.2.1</mods:identifier>
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<mods:identifier id="58CD7C9DFDCA565AB05607C6981AA5D3" type="GBIF-Dataset">d7cbc23a-7a4c-4616-83fd-059fb458f7b6</mods:identifier>
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<mods:identifier id="EC364DBFAF69205F3BBA271DD54D0B0B" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B3B5A685AE3D34229FDF0E01812B542F" type="Zenodo-Dep">257286</mods:identifier>
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<mods:identifier id="9A4CA02CC4D8976C194027500ADD4D9C" type="ZooBank">B264C215-000D-42C5-8AC9-B801872CD182</mods:identifier>
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</mods:mods>
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<treatment id="953787B9FF93FFF45CF0FC398648F946" ID-DOI="http://doi.org/10.5281/zenodo.6055258" ID-GBIF-Taxon="124996945" ID-Zenodo-Dep="6055258" LSID="urn:lsid:plazi:treatment:953787B9FF93FFF45CF0FC398648F946" httpUri="http://treatment.plazi.org/id/953787B9FF93FFF45CF0FC398648F946" lastPageNumber="129" pageId="28" pageNumber="129">
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<subSubSection id="55846524FF93FFF45CF0FC39878FFBB4" pageId="28" pageNumber="129" type="nomenclature">
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<paragraph id="1D2136AFFF93FFF45CF0FC398499FC72" blockId="28.[151,482,943,970]" box="[151,482,943,970]" pageId="28" pageNumber="129">
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<heading id="466981C3FF93FFF45CF0FC398499FC72" bold="true" box="[151,482,943,970]" fontSize="11" level="1" pageId="28" pageNumber="129" reason="1">
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<taxonomicName id="DA9E4D2CFF93FFF45CF0FC398499FC72" ID-CoL="7PZMR" authority="Doderlein, 1924" authorityName="Doderlein" authorityYear="1924" box="[151,482,943,970]" class="Asteroidea" family="Goniasteridae" genus="Sibogaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="genus">
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<emphasis id="2FEAEABDFF93FFF45CF0FC398499FC72" bold="true" box="[151,482,943,970]" pageId="28" pageNumber="129">
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<emphasis id="2FEAEABDFF93FFF45CF0FC39846DFC71" bold="true" box="[151,278,943,969]" italics="true" pageId="28" pageNumber="129">Sibogaster</emphasis>
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<bibRefCitation id="790F4B5EFF93FFF45D7AFC268499FC72" author="Doderlein" box="[285,482,944,970]" pageId="28" pageNumber="129" refString="Doderlein, L. (1924) Die Asteriden der Siboga-Exped. II. Pentagonasteridae. Siboga-Expedition, 46 (2), 49 - 69, pls. 14 - 19." type="journal article" year="1924">Döderlein, 1924</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="1D2136AFFF93FFF45CF0FC60878FFBB4" blockId="28.[151,756,1014,1036]" box="[151,756,1014,1036]" pageId="28" pageNumber="129">
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<bibRefCitation id="790F4B5EFF93FFF45CF0FC608444FBB4" author="Doderlein" box="[151,319,1014,1036]" pageId="28" pageNumber="129" refString="Doderlein, L. (1924) Die Asteriden der Siboga-Exped. II. Pentagonasteridae. Siboga-Expedition, 46 (2), 49 - 69, pls. 14 - 19." type="journal article" year="1924">Döderlein, 1924</bibRefCitation>
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: 63;
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<bibRefCitation id="790F4B5EFF93FFF45D08FC61875BFBB4" author="Clark" box="[367,544,1014,1036]" pageId="28" pageNumber="129" refString="Clark, A. M. (1993) An index of names of recent Asteroidea - Part 2: Valvatida. E chinoderm Studies, 4, 187 - 366." type="journal article" year="1993">A.M. Clark 1993</bibRefCitation>
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: 283 (checklist only)
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</paragraph>
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</subSubSection>
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<subSubSection id="55846524FF93FFF45CF0FBAF845DFACA" pageId="28" pageNumber="129" type="materials_examined">
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<paragraph id="1D2136AFFF93FFF45CF0FBAF871EFBEA" blockId="28.[151,1436,1081,1790]" box="[151,613,1081,1106]" pageId="28" pageNumber="129">
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<emphasis id="2FEAEABDFF93FFF45CF0FBAF8449FBEA" bold="true" box="[151,306,1081,1106]" pageId="28" pageNumber="129">Type species.</emphasis>
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<taxonomicName id="DA9E4D2CFF93FFF45D5EFBAC84CCFBEA" box="[313,439,1081,1106]" class="Asteroidea" family="Goniasteridae" genus="Sibogaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="species" species="digitatus">
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<emphasis id="2FEAEABDFF93FFF45D5EFBAC84CCFBEA" box="[313,439,1081,1106]" italics="true" pageId="28" pageNumber="129">S. digitatus</emphasis>
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</taxonomicName>
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(by monotypy)
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</paragraph>
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<paragraph id="1D2136AFFF93FFF45CA0FBCB86CEFB06" blockId="28.[151,1436,1081,1790]" pageId="28" pageNumber="129">
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<emphasis id="2FEAEABDFF93FFF45CA0FBCB84A8FBCE" bold="true" box="[199,467,1117,1142]" pageId="28" pageNumber="129">Taxonomic comments.</emphasis>
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<taxonomicName id="DA9E4D2CFF93FFF45DBBFBCB872FFBCE" box="[476,596,1117,1142]" class="Asteroidea" family="Goniasteridae" genus="Sibogaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="genus">
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<emphasis id="2FEAEABDFF93FFF45DBBFBCB872FFBCE" box="[476,596,1117,1142]" italics="true" pageId="28" pageNumber="129">Sibogaster</emphasis>
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</taxonomicName>
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has not been collected since its original description by
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<bibRefCitation id="790F4B5EFF93FFF458B5FBCB80E3FBCE" author="Doderlein" box="[1234,1432,1117,1142]" pageId="28" pageNumber="129" refString="Doderlein, L. (1924) Die Asteriden der Siboga-Exped. II. Pentagonasteridae. Siboga-Expedition, 46 (2), 49 - 69, pls. 14 - 19." type="journal article" year="1924">Döderlein (1924)</bibRefCitation>
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. The second species,
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<taxonomicName id="DA9E4D2CFF93FFF45D1CFB14849EFB21" box="[379,485,1154,1177]" class="Asteroidea" family="Goniasteridae" genus="Sibogaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="species" species="nieseni" status="sp. nov.">
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<emphasis id="2FEAEABDFF93FFF45D1CFB14849EFB21" box="[379,485,1154,1177]" italics="true" pageId="28" pageNumber="129">S. nieseni</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="34D957C6FF93FFF45D94FB178749FB22" box="[499,562,1153,1178]" pageId="28" pageNumber="129" rank="species">
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<emphasis id="2FEAEABDFF93FFF45D94FB178749FB22" bold="true" box="[499,562,1153,1178]" pageId="28" pageNumber="129">n. sp.</emphasis>
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</taxonomicNameLabel>
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described below expands the concept of
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<taxonomicName id="DA9E4D2CFF93FFF45F9CFB178108FB22" box="[1019,1139,1153,1178]" class="Asteroidea" family="Goniasteridae" genus="Sibogaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="genus">
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<emphasis id="2FEAEABDFF93FFF45F9CFB178108FB22" box="[1019,1139,1153,1178]" italics="true" pageId="28" pageNumber="129">Sibogaster</emphasis>
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</taxonomicName>
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based on greater character variation, including further differences in glassy bosses among species.
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</paragraph>
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<paragraph id="1D2136AFFF93FFF45CA0FB5F845DFACA" blockId="28.[151,1436,1081,1790]" pageId="28" pageNumber="129">
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The type species,
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<taxonomicName id="DA9E4D2CFF93FFF45DF8FB5C8759FB5A" box="[415,546,1225,1250]" class="Asteroidea" family="Goniasteridae" genus="Sibogaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="species" species="digitatus">
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<emphasis id="2FEAEABDFF93FFF45DF8FB5C8759FB5A" box="[415,546,1225,1250]" italics="true" pageId="28" pageNumber="129">S. digitatus</emphasis>
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</taxonomicName>
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possesses glassy bosses embedded on flattened, abutted abactinal plates, a character which is shared by
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<taxonomicName id="DA9E4D2CFF93FFF45D80FB7B871FFABE" box="[487,612,1261,1286]" class="Asteroidea" family="Goniasteridae" genus="Plinthaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="genus">
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<emphasis id="2FEAEABDFF93FFF45D80FB7B871FFABE" box="[487,612,1261,1286]" italics="true" pageId="28" pageNumber="129">Plinthaster</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="DA9E4D2CFF93FFF45EF9FB788643FABD" box="[670,824,1262,1285]" class="Asteroidea" family="Goniasteridae" genus="Astroceramus" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="genus">
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<emphasis id="2FEAEABDFF93FFF45EF9FB788643FABD" box="[670,824,1262,1285]" italics="true" pageId="28" pageNumber="129">Astroceramus</emphasis>
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</taxonomicName>
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. Both species of
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<taxonomicName id="DA9E4D2CFF93FFF45863FB7B8107FABE" box="[1028,1148,1261,1286]" class="Asteroidea" family="Goniasteridae" genus="Sibogaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="genus">
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<emphasis id="2FEAEABDFF93FFF45863FB7B8107FABE" box="[1028,1148,1261,1286]" italics="true" pageId="28" pageNumber="129">Sibogaster</emphasis>
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</taxonomicName>
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presented herein possess bare abactinal plates with no surficial granules or other accessories. Granules on superomarginal plates in
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<taxonomicName id="DA9E4D2CFF93FFF45CF0FAA38474FAF6" box="[151,271,1333,1358]" class="Asteroidea" family="Goniasteridae" genus="Sibogaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="genus">
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<emphasis id="2FEAEABDFF93FFF45CF0FAA38474FAF6" box="[151,271,1333,1358]" italics="true" pageId="28" pageNumber="129">Sibogaster</emphasis>
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</taxonomicName>
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are either weakly expressed or absent. Superomarginals in both species are elongate (L>W) interradially.
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</paragraph>
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</subSubSection>
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<subSubSection id="55846524FF93FFF45CA0FAEB86E3F99E" pageId="28" pageNumber="129" type="reference_group">
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<paragraph id="1D2136AFFF93FFF45CA0FAEB86E3F99E" blockId="28.[151,1436,1081,1790]" pageId="28" pageNumber="129">
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<taxonomicName id="DA9E4D2CFF93FFF45CA0FAEB8444FA2E" ID-CoL="7PZMR" box="[199,319,1405,1430]" class="Asteroidea" family="Goniasteridae" genus="Sibogaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="genus">
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<emphasis id="2FEAEABDFF93FFF45CA0FAEB8444FA2E" box="[199,319,1405,1430]" italics="true" pageId="28" pageNumber="129">Sibogaster</emphasis>
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</taxonomicName>
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shares a strong affinity with
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<taxonomicName id="DA9E4D2CFF93FFF45EF7FAE88664FA2D" ID-CoL="7PQSX" box="[656,799,1406,1429]" class="Asteroidea" family="Goniasteridae" genus="Pergamaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="genus">
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<emphasis id="2FEAEABDFF93FFF45EF7FAE88664FA2D" box="[656,799,1406,1429]" italics="true" pageId="28" pageNumber="129">Pergamaster</emphasis>
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</taxonomicName>
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particularly
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<taxonomicName id="DA9E4D2CFF93FFF45FD4FAE88040FA2E" ID-CoL="76T2B" authority="Bell, 1908" authorityName="Bell" authorityYear="1908" box="[947,1339,1405,1430]" class="Asteroidea" family="Goniasteridae" genus="Pergamaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="species" species="incertus">
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<emphasis id="2FEAEABDFF93FFF45FD4FAE881DEFA2D" box="[947,1189,1406,1429]" italics="true" pageId="28" pageNumber="129">Pergamaster incertus</emphasis>
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(
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<bibRefCitation id="790F4B5EFF93FFF458D0FAEB8049FA2E" author="Bell" box="[1207,1330,1405,1430]" pageId="28" pageNumber="129" refString="Bell, F. J. (1908) Echinoderma. National Antarctic Expedition, London. Natural History, Zoology, 4, 1 - 16, 5 pls." type="journal article" year="1908">Bell, 1908</bibRefCitation>
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)
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</taxonomicName>
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. Both
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<taxonomicName id="DA9E4D2CFF93FFF459E0FAE8859DFA01" ID-CoL="4X6FF" class="Asteroidea" family="Goniasteridae" genus="Sibogaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="species" species="nieseni">
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<emphasis id="2FEAEABDFF93FFF459E0FAE8859DFA01" italics="true" pageId="28" pageNumber="129">S. nieseni</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="DA9E4D2CFF93FFF45D79FA3484EEFA01" ID-CoL="76T2B" box="[286,405,1442,1465]" class="Asteroidea" family="Goniasteridae" genus="Pergamaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="species" species="incertus">
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<emphasis id="2FEAEABDFF93FFF45D79FA3484EEFA01" box="[286,405,1442,1465]" italics="true" pageId="28" pageNumber="129">P. incertus</emphasis>
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</taxonomicName>
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are devoid of abactinal granules, have similar marginal plate arrangement and morphology and similar overall body shape. They differ in that granule and abactinal plate shapes and size are significantly different in size for similar-sized individuals and in that
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<taxonomicName id="DA9E4D2CFF93FFF45F53FA7C86B8F9B9" ID-CoL="7PQSX" box="[820,963,1514,1537]" class="Asteroidea" family="Goniasteridae" genus="Pergamaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="genus">
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<emphasis id="2FEAEABDFF93FFF45F53FA7C86B8F9B9" box="[820,963,1514,1537]" italics="true" pageId="28" pageNumber="129">Pergamaster</emphasis>
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</taxonomicName>
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displays significantly larger and thicker subambulacral and furrow spines are significantly larger and thicker.
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</paragraph>
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</subSubSection>
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<subSubSection id="55846524FF93FFF45CA0F9A78648F946" pageId="28" pageNumber="129" type="diagnosis">
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<paragraph id="1D2136AFFF93FFF45CA0F9A78648F946" blockId="28.[151,1436,1081,1790]" pageId="28" pageNumber="129">
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<emphasis id="2FEAEABDFF93FFF45CA0F9A7843AF9F2" bold="true" box="[199,321,1585,1610]" pageId="28" pageNumber="129">Diagnosis.</emphasis>
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Abactinal plates smooth, bare (no accessories on plate surface-although embedded glassy bosses present in
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<taxonomicName id="DA9E4D2CFF93FFF45D77F9C084EBF9D6" box="[272,400,1621,1646]" class="Asteroidea" family="Goniasteridae" genus="Sibogaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="species" species="digitatus">
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<emphasis id="2FEAEABDFF93FFF45D77F9C084EBF9D6" box="[272,400,1621,1646]" italics="true" pageId="28" pageNumber="129">S. digitatus</emphasis>
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</taxonomicName>
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), weakly convex, round to polygonal in outline. Marginal plates elongate, mostly smooth, bare (
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<taxonomicName id="DA9E4D2CFF93FFF45CB0F9EC842EF92A" box="[215,341,1657,1682]" class="Asteroidea" family="Goniasteridae" genus="Sibogaster" kingdom="Animalia" order="Valvatida" pageId="28" pageNumber="129" phylum="Echinodermata" rank="species" species="digitatus">
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<emphasis id="2FEAEABDFF93FFF45CB0F9EC842EF92A" box="[215,341,1657,1682]" italics="true" pageId="28" pageNumber="129">S. digitatus</emphasis>
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</taxonomicName>
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has fewer than 6 granules on one or two interradial superomarginals). Inferomarginals adjacent to actinal surface covered with variably abundant granules. Actinal plates covered by granules. Adambulacral plates rectangular in shape. Furrow spines numerous, six to 15. Subambulacrals. thickened relative to furrow spines, remaining granulation identical to others on actinal surface.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |