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<document id="1EFF850DFECC4A82D5E633299244E66B" ID-DOI="10.11646/zootaxa.4013.1.1" ID-GBIF-Dataset="c9e02c83-1141-4b3b-bd9e-68fac8b5cfc3" ID-ISSN="1175-5326" ID-Zenodo-Dep="289376" ID-ZooBank="EE549B24-9D30-4495-9A39-4C4A4E9E03FF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461244395630" checkinUser="plazi" docAuthor="Cai, Yan-Peng, Zhao, Zong-Yi &amp; Zhou, Hong-Zhang" docDate="2015" docId="7475A014FFB1971CFF3C1F8D0BBDFB67" docLanguage="en" docName="zt04013p026.pdf" docOrigin="Zootaxa 4013" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Quedius (Microsaurus) mukuensis" docType="key" docVersion="7" lastPageNumber="3" masterDocId="884CD86CFFB0971EFFAB192E0E21FFAA" masterDocTitle="Taxonomy of the Quedius mukuensis group (Coleoptera: Staphylinidae: Staphylinini: Quediina) with descriptions of four new species from China" masterLastPageNumber="26" masterPageNumber="1" pageNumber="2" updateTime="1698605968782" updateUser="plazi">
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<mods:title id="D3CD2CB5452FEDF3549CB20E3A28B169">Taxonomy of the Quedius mukuensis group (Coleoptera: Staphylinidae: Staphylinini: Quediina) with descriptions of four new species from China</mods:title>
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<mods:namePart id="8443F3C4F997D180B3ABA026B8BD78FB">Cai, Yan-Peng</mods:namePart>
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<mods:namePart id="55D866B2CDFB165E457A36C57F912ACD">Zhao, Zong-Yi</mods:namePart>
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<paragraph id="FC631102FFB1971FFF3C1F8D0D42F917" blockId="1.[151,867,1698,1725]" box="[151,867,1698,1725]" pageId="1" pageNumber="2">
<heading id="A72BA66EFFB1971FFF3C1F8D0D42F917" bold="true" box="[151,867,1698,1725]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
<emphasis id="CEA8CD10FFB1971FFF3C1F8D0D42F917" bold="true" box="[151,867,1698,1725]" pageId="1" pageNumber="2">
Key to species of
<taxonomicName id="3BDC6A81FFB1971FFEC61F8C0D30F916" ID-CoL="4R2SK" box="[365,785,1698,1725]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="mukuensis" subGenus="Microsaurus">
<emphasis id="CEA8CD10FFB1971FFEC61F8C0FF3F916" bold="true" box="[365,466,1698,1724]" italics="true" pageId="1" pageNumber="2">Quedius</emphasis>
(
<emphasis id="CEA8CD10FFB1971FFE4F1F8C0C5EF916" bold="true" box="[484,639,1698,1724]" italics="true" pageId="1" pageNumber="2">Microsaurus</emphasis>
)
<emphasis id="CEA8CD10FFB1971FFD241F8C0D30F916" bold="true" box="[655,785,1698,1724]" italics="true" pageId="1" pageNumber="2">mukuensis</emphasis>
</taxonomicName>
group
</emphasis>
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1. Wings reduced, only slightly longer than elytra; abdominal tergite VII without or only with inconspicuous whitish apical seam of palisade setae.......................................................................
<taxonomicName id="3BDC6A81FFB1971FFB431E2B0B68F8B1" ID-CoL="4R2S6" box="[1256,1353,1797,1819]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="medius" status="sp. nov.">
<emphasis id="CEA8CD10FFB1971FFB431E2B0B68F8B1" box="[1256,1353,1797,1819]" italics="true" pageId="1" pageNumber="2">Q. medius</emphasis>
</taxonomicName>
<emphasis id="CEA8CD10FFB1971FFAE41E280BBDF8B6" bold="true" box="[1359,1436,1798,1820]" pageId="1" pageNumber="2">
<taxonomicNameLabel id="D59B706BFFB1971FFAE41E280BBDF8B6" box="[1359,1436,1798,1820]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<paragraph id="FC631102FFB1971FFF3C1E0A0BBDF893" blockId="1.[151,1436,1768,2025]" box="[151,1436,1827,1849]" pageId="1" pageNumber="2">
<keyLead id="472DB237FFB1971FFF3C1E0A0BBDF893" box="[151,1436,1827,1849]" pageId="1" pageNumber="2">- Wings fully developed; abdominal tergite VII with distinct whitish apical seam of palisade setae..................... 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="472809A7FFB1971FFF3C1E6E0BBDF8D9" pageId="1" pageNumber="2">
<paragraph id="FC631102FFB1971FFF3C1E6E0BBDF8FC" blockId="1.[151,1436,1768,2025]" box="[151,1436,1856,1878]" pageId="1" pageNumber="2">
<keyLead id="472DB237FFB1971FFF3C1E6E0BBDF8FC" box="[151,1436,1856,1878]" pageId="1" pageNumber="2">
2. Median lobe of aedeagus with two processes at apex facing parameral side (
<figureCitation id="64E70D87FFB1971FFC121E6E0DDAF8FC" box="[953,1019,1856,1878]" captionStart="FIGURE 9" captionStartId="23.[151,250,1877,1899]" captionTargetBox="[193,1433,201,1855]" captionTargetId="figure@23.[151,1436,193,1856]" captionTargetPageId="23" captionText="FIGURE 9. Quedius (Microsaurus) nireus Smetana, 1995: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/289390/files/figure.png" pageId="1" pageNumber="2">Figs. 9</figureCitation>
E; 9-1G)......................... 3
</keyLead>
</paragraph>
<paragraph id="FC631102FFB1971FFF3C1E700BBDF8D9" blockId="1.[151,1436,1768,2025]" box="[151,1436,1885,1907]" pageId="1" pageNumber="2">
<keyLead id="472DB237FFB1971FFF3C1E700BBDF8D9" box="[151,1436,1885,1907]" pageId="1" pageNumber="2">
- Median lobe of aedeagus with one process at apex facing parameral side (e.g.,
<figureCitation id="64E70D87FFB1971FFC781E730A37F8D9" box="[979,1046,1885,1907]" captionStart="FIGURE 2" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[193,1433,216,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 2. Quedius (Microsaurus) indigestus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/289378/files/figure.png" pageId="1" pageNumber="2">Figs. 2</figureCitation>
E; 2-1G)....................... 5
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</paragraph>
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<keyStep id="472809A7FFB1971FFF3C1E550BBAF861" pageId="1" pageNumber="2">
<paragraph id="FC631102FFB1971FFF3C1E550BBDF804" blockId="1.[151,1436,1768,2025]" pageId="1" pageNumber="2">
<keyLead id="472DB237FFB1971FFF3C1E550BBDF804" pageId="1" pageNumber="2">
3. Size distinctly smaller, shorter than
<quantity id="3B24BCE7FFB1971FFD931E550CA5F83B" box="[568,644,1915,1937]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="2" unit="mm" value="6.5">6.5 mm</quantity>
; body bicolored, with elytra reddish brown, other parts black....................................................................................................
<taxonomicName id="3BDC6A81FFB1971FFB7F1EB60BBDF804" ID-CoL="4R2Q5" authority="Smetana" authorityName="Smetana" box="[1236,1436,1944,1966]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="kabateki">
<emphasis id="CEA8CD10FFB1971FFB7F1EB60B62F804" box="[1236,1347,1944,1966]" italics="true" pageId="1" pageNumber="2">Q. kabateki</emphasis>
Smetana
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="FC631102FFB1971FFF3C1E980BBAF861" blockId="1.[151,1436,1768,2025]" box="[151,1435,1973,1995]" pageId="1" pageNumber="2">
<keyLead id="472DB237FFB1971FFF3C1E980BBAF861" box="[151,1435,1973,1995]" pageId="1" pageNumber="2">
- Size distinctly larger, longer than
<quantity id="3B24BCE7FFB1971FFD8F1E9B0C51F861" box="[548,624,1973,1995]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="2" unit="mm" value="6.5">6.5 mm</quantity>
; body unicolored, blackish brown to black............................... 4
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<paragraph id="FC631102FFB1971FFF3C1EFD0BB7F843" blockId="1.[151,1436,1768,2025]" box="[151,1430,2003,2025]" pageId="1" pageNumber="2">
<keyLead id="472DB237FFB1971FFF3C1EFD0BB7F843" box="[151,1430,2003,2025]" pageId="1" pageNumber="2">4. Protonum with three setiferous punctures in each dorsal and sublateral row; male sternite VIII with 23 long setae on each side</keyLead>
</paragraph>
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.................................................................................
<taxonomicName id="3BDC6A81FFB2971CFB1A19B90BBDFF07" ID-CoL="4R2SK" authority="Bernhauer" authorityName="Bernhauer" box="[1201,1436,151,173]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="mukuensis">
<emphasis id="CEA8CD10FFB2971CFB1A19B90B13FF07" box="[1201,1330,151,173]" italics="true" pageId="2" pageNumber="3">Q. mukuensis</emphasis>
Bernhauer
</taxonomicName>
<keyLead id="472DB237FFB2971CFF3C199B0BBDFF4D" pageId="2" pageNumber="3">
- Protonum with four setiferous punctures in each dorsal and sublateral row (
<figureCitation id="64E70D87FFB2971CFC1F199A0DCFFF60" box="[948,1006,180,202]" captionStart="FIGURE 9" captionStartId="23.[151,250,1877,1899]" captionTargetBox="[193,1433,201,1855]" captionTargetId="figure@23.[151,1436,193,1856]" captionTargetPageId="23" captionText="FIGURE 9. Quedius (Microsaurus) nireus Smetana, 1995: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/289390/files/figure.png" pageId="2" pageNumber="3">Fig. 9</figureCitation>
B); male sternite VIII with 34 long setae on each side (
<figureCitation id="64E70D87FFB2971CFEED19FF0FB1FF4D" box="[326,400,209,231]" captionStart="FIGURE 9" captionStartId="23.[151,250,1877,1899]" captionTargetBox="[193,1433,201,1855]" captionTargetId="figure@23.[151,1436,193,1856]" captionTargetPageId="23" captionText="FIGURE 9. Quedius (Microsaurus) nireus Smetana, 1995: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/289390/files/figure.png" pageId="2" pageNumber="3">Fig. 9-1</figureCitation>
A)....................................................................
<taxonomicName id="3BDC6A81FFB2971CFB4019FF0BBDFF4D" ID-CoL="4R2SZ" authority="Smetana" authorityName="Smetana" box="[1259,1436,209,231]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="nireus">
<emphasis id="CEA8CD10FFB2971CFB4019FF0B62FF4D" box="[1259,1347,209,231]" italics="true" pageId="2" pageNumber="3">Q. nireus</emphasis>
Smetana
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="472809A7FFB2971CFF3C19C10BBDFE88" pageId="2" pageNumber="3">
<paragraph id="FC631102FFB2971CFF3C19C10BBDFEAF" blockId="2.[151,1437,151,1229]" box="[151,1436,239,261]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C19C10BBDFEAF" box="[151,1436,239,261]" pageId="2" pageNumber="3">
5. Head with additional setiferous punctures anteriad of posterior frontal setiferous puncture (
<figureCitation id="64E70D87FFB2971CFBE319C10AABFEAF" box="[1096,1162,239,261]" captionStart="FIGURE 7" captionStartId="19.[151,250,1877,1899]" captionTargetBox="[193,1433,206,1855]" captionTargetId="figure@19.[151,1436,193,1856]" captionTargetPageId="19" captionText="FIGURE 7. Quedius (Microsaurus) gongga Smetana, 2006: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/289386/files/figure.png" pageId="2" pageNumber="3">Figs. 7</figureCitation>
A; 8A; 9A)........... 6
</keyLead>
</paragraph>
<paragraph id="FC631102FFB2971CFF3C18230BBDFE88" blockId="2.[151,1437,151,1229]" box="[151,1436,268,290]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C18230BBDFE88" box="[151,1436,268,290]" pageId="2" pageNumber="3">
- Head without additional setiferous punctures anteriad of posterior frontal setiferous puncture (e.g.,
<figureCitation id="64E70D87FFB2971CFB3D18220AEFFE88" box="[1174,1230,268,290]" captionStart="FIGURE 4" captionStartId="11.[151,250,1877,1899]" captionTargetBox="[193,1433,204,1855]" captionTargetId="figure@11.[151,1436,193,1856]" captionTargetPageId="11" captionText="FIGURE 4. Quedius (Microsaurus) purus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/289382/files/figure.png" pageId="2" pageNumber="3">Fig. 4</figureCitation>
A)........... 10
</keyLead>
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<keyStep id="472809A7FFB2971CFF3C18070BBAFE3D" pageId="2" pageNumber="3">
<paragraph id="FC631102FFB2971CFF3C18070BBCFEF7" blockId="2.[151,1437,151,1229]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C18070BBCFEF7" pageId="2" pageNumber="3">
6. Male sternite VIII with six long setae on each side; aedeagus with apex of paramere protruding beyond that of median lobe....................................................................................
<taxonomicName id="3BDC6A81FFB2971CFB6E18690BBCFEF7" ID-CoL="4R2WY" authority="Zheng et al." authorityName="Zheng et al." box="[1221,1437,327,349]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="xiaoae">
<emphasis id="CEA8CD10FFB2971CFB6E18690B03FEF7" box="[1221,1314,327,349]" italics="true" pageId="2" pageNumber="3">Q. xiaoae</emphasis>
Zheng
<emphasis id="CEA8CD10FFB2971CFAC718660BBCFEF7" box="[1388,1437,327,349]" italics="true" pageId="2" pageNumber="3">et al.</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="FC631102FFB2971CFF3C184B0BBAFE3D" blockId="2.[151,1437,151,1229]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C184B0BBAFE3D" pageId="2" pageNumber="3">- Male sternite VIII with fewer than six long setae on each side; aedeagus with apex of paramere not reaching that of median lobe............................................................................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="472809A7FFB2971CFF3C18B10BBAFDA7" pageId="2" pageNumber="3">
<paragraph id="FC631102FFB2971CFF3C18B10BBDFE78" blockId="2.[151,1437,151,1229]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C18B10BBDFE78" pageId="2" pageNumber="3">
7. Median lobe of aedeagus with apical portion strongly dilated laterally into triangular apex (
<figureCitation id="64E70D87FFB2971CFBE018B10AACFE1E" box="[1099,1165,415,437]" captionStart="FIGURE 7" captionStartId="19.[151,250,1877,1899]" captionTargetBox="[193,1433,206,1855]" captionTargetId="figure@19.[151,1436,193,1856]" captionTargetPageId="19" captionText="FIGURE 7. Quedius (Microsaurus) gongga Smetana, 2006: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/289386/files/figure.png" pageId="2" pageNumber="3">Figs. 7</figureCitation>
E; 7-1G)....................................................................................................
<taxonomicName id="3BDC6A81FFB2971CFB7518920BBDFE78" ID-CoL="9F553" authority="Smetana" authorityName="Smetana" box="[1246,1436,444,466]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="gongga">
<emphasis id="CEA8CD10FFB2971CFB7518920B62FE78" box="[1246,1347,444,466]" italics="true" pageId="2" pageNumber="3">Q. gongga</emphasis>
Smetana
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="FC631102FFB2971CFF3C18F40BBAFDA7" blockId="2.[151,1437,151,1229]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C18F40BBAFDA7" pageId="2" pageNumber="3">
- Median lobe of aedeagus with apical portion narrow, nearly parallel-sided, not apreciably dilated laterally, apex narrowly arcu- ate (e.g.,
<figureCitation id="64E70D87FFB2971CFE9218D90F5AFDA7" box="[313,379,503,525]" captionStart="FIGURE 8" captionStartId="21.[151,250,1877,1899]" captionTargetBox="[193,1433,194,1855]" captionTargetId="figure@21.[151,1436,193,1856]" captionTargetPageId="21" captionText="FIGURE 8. Quedius (Microsaurus) haemon Smetana, 1995: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/289388/files/figure.png" pageId="2" pageNumber="3">Figs. 8</figureCitation>
E; 8-1G).............................................................................. 8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="472809A7FFB2971CFF3C1B3A0BBDFDCF" pageId="2" pageNumber="3">
<paragraph id="FC631102FFB2971CFF3C1B3A0BBDFDED" blockId="2.[151,1437,151,1229]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1B3A0BBDFDED" pageId="2" pageNumber="3">
8. Paramere of aedeagus with some sensory peg setae situated above level of subapical setae insertion points (
<figureCitation id="64E70D87FFB2971CFB4D1B3A0B09FD80" box="[1254,1320,532,554]" captionStart="FIGURE 8" captionStartId="21.[151,250,1877,1899]" captionTargetBox="[193,1433,194,1855]" captionTargetId="figure@21.[151,1436,193,1856]" captionTargetPageId="21" captionText="FIGURE 8. Quedius (Microsaurus) haemon Smetana, 1995: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/289388/files/figure.png" pageId="2" pageNumber="3">Figs. 8</figureCitation>
D; 8-1F)......................................................................................
<taxonomicName id="3BDC6A81FFB2971CFB711B1F0BBDFDED" ID-CoL="4R2NZ" authority="Smetana" authorityName="Smetana" box="[1242,1436,561,583]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="haemon">
<emphasis id="CEA8CD10FFB2971CFB711B1F0B62FDED" box="[1242,1347,561,583]" italics="true" pageId="2" pageNumber="3">Q. haemon</emphasis>
Smetana
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="FC631102FFB2971CFF3C1B610BBDFDCF" blockId="2.[151,1437,151,1229]" box="[151,1436,591,613]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1B610BBDFDCF" box="[151,1436,591,613]" pageId="2" pageNumber="3">- Paramere of aedeagus with sensory peg setae far below apex, all below level of subapical setae insertion points......... 9</keyLead>
</paragraph>
</keyStep>
<keyStep id="472809A7FFB2971CFF3C1B420BBDFD70" pageId="2" pageNumber="3">
<paragraph id="FC631102FFB2971CFF3C1B420BBDFD35" blockId="2.[151,1437,151,1229]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1B420BBDFD35" pageId="2" pageNumber="3">
9. Male sternite VIII with four long setae on each side, with medioapical emargination deeper; median lobe of aedeagus with small tooth-shaped process at apex facing parameral side.........................................
<taxonomicName id="3BDC6A81FFB2971CFB5E1BA70BBDFD35" ID-CoL="4R2NY" authority="Smetana" authorityName="Smetana" box="[1269,1436,649,671]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="haan">
<emphasis id="CEA8CD10FFB2971CFB5E1BA70B63FD35" box="[1269,1346,649,671]" italics="true" pageId="2" pageNumber="3">Q. haan</emphasis>
Smetana
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="FC631102FFB2971CFF3C1B890BBDFD70" blockId="2.[151,1437,151,1229]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1B890BBDFD70" pageId="2" pageNumber="3">
- Male sternite VIII with five long setae on each side, with medioapical emargination shallower; median lobe of aedeagus with distinctly developed long medial carina at apex facing parameral side..............................
<taxonomicName id="3BDC6A81FFB2971CFB521BEA0BBDFD70" ID-CoL="6WQ9T" authority="Smetana" authorityName="Smetana" box="[1273,1436,708,730]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="raan">
<emphasis id="CEA8CD10FFB2971CFB521BEA0B62FD70" box="[1273,1347,708,730]" italics="true" pageId="2" pageNumber="3">Q. raan</emphasis>
Smetana
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="472809A7FFB2971CFF3C1BCF0BBAFC98" pageId="2" pageNumber="3">
<paragraph id="FC631102FFB2971CFF3C1BCF0BBDFCBF" blockId="2.[151,1437,151,1229]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1BCF0BBDFCBF" pageId="2" pageNumber="3">
10. Paramere of aedeagus with sensory peg setae not only forming two groups, but also with one other peg seta on each side situ- ated far below them (
<figureCitation id="64E70D87FFB2971CFE061BD10FD1FCBF" box="[429,496,767,789]" captionStart="FIGURE 4" captionStartId="11.[151,250,1877,1899]" captionTargetBox="[193,1433,204,1855]" captionTargetId="figure@11.[151,1436,193,1856]" captionTargetPageId="11" captionText="FIGURE 4. Quedius (Microsaurus) purus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/289382/files/figure.png" pageId="2" pageNumber="3">Figs. 4</figureCitation>
D; 4-1F)........................................................
<taxonomicName id="3BDC6A81FFB2971CFB5F1BD10B66FCBF" ID-CoL="78PY2" box="[1268,1351,767,789]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="purus" status="sp. nov.">
<emphasis id="CEA8CD10FFB2971CFB5F1BD10B66FCBF" box="[1268,1351,767,789]" italics="true" pageId="2" pageNumber="3">Q. purus</emphasis>
</taxonomicName>
<emphasis id="CEA8CD10FFB2971CFAE41BD10BBDFCBF" bold="true" box="[1359,1436,767,789]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="D59B706BFFB2971CFAE41BD10BBDFCBF" box="[1359,1436,767,789]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="FC631102FFB2971CFF3C1A330BBAFC98" blockId="2.[151,1437,151,1229]" box="[151,1435,796,818]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1A330BBAFC98" box="[151,1435,796,818]" pageId="2" pageNumber="3">
- Paramere of aedeagus with sensory peg setae only forming two groups (e.g.,
<figureCitation id="64E70D87FFB2971CFC681A320A25FC98" box="[963,1028,796,818]" captionStart="FIGURE 5" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[193,1433,215,1855]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 5. Quedius (Microsaurus) rectus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/289383/files/figure.png" pageId="2" pageNumber="3">Figs. 5</figureCitation>
D; 5-1F)....................... 11
</keyLead>
</paragraph>
</keyStep>
<keyStep id="472809A7FFB2971CFF3C1A170BBDFC20" pageId="2" pageNumber="3">
<paragraph id="FC631102FFB2971CFF3C1A170BBDFCC7" blockId="2.[151,1437,151,1229]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1A170BBDFCC7" pageId="2" pageNumber="3">
11. Paramere of aedeagus relatively wide, covering entire apical portion of median lobe except for very apex.....................................................................................................
<taxonomicName id="3BDC6A81FFB2971CFB421A790BBDFCC7" ID-CoL="4R2MT" authority="Smetana" authorityName="Smetana" box="[1257,1436,855,877]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="epytus">
<emphasis id="CEA8CD10FFB2971CFB421A790B62FCC7" box="[1257,1347,855,877]" italics="true" pageId="2" pageNumber="3">Q. epytus</emphasis>
Smetana
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="FC631102FFB2971CFF3C1A5B0BBDFC20" blockId="2.[151,1437,151,1229]" box="[151,1436,884,906]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1A5B0BBDFC20" box="[151,1436,884,906]" pageId="2" pageNumber="3">- Paramere of aedeagus relatively narrow, only covering part of apical portion of median lobe........................ 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="472809A7FFB2971CFF3C1ABF0BBAFC6F" pageId="2" pageNumber="3">
<paragraph id="FC631102FFB2971CFF3C1ABF0BBAFC0D" blockId="2.[151,1437,151,1229]" box="[151,1435,913,935]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1ABF0BBAFC0D" box="[151,1435,913,935]" pageId="2" pageNumber="3">12. Male sternite VIII with three long setae on each side....................................................... 13</keyLead>
</paragraph>
<paragraph id="FC631102FFB2971CFF3C1A810BBAFC6F" blockId="2.[151,1437,151,1229]" box="[151,1435,943,965]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1A810BBAFC6F" box="[151,1435,943,965]" pageId="2" pageNumber="3">- Male sternite VIII with four long setae on each side........................................................ 14</keyLead>
</paragraph>
</keyStep>
<keyStep id="472809A7FFB2971CFF3C1AE20BBDFB90" pageId="2" pageNumber="3">
<paragraph id="FC631102FFB2971CFF3C1AE20BBDFC55" blockId="2.[151,1437,151,1229]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1AE20BBDFC55" pageId="2" pageNumber="3">
13. Paramere of aedeagus relatively longer, with apex almost reaching that of median lobe, with about three sensory peg setae on its underside............................................................................
<taxonomicName id="3BDC6A81FFB2971CFB511AC70BBDFC55" ID-CoL="78Q9X" authority="Smetana" authorityName="Smetana" box="[1274,1436,1001,1023]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="tzwu">
<emphasis id="CEA8CD10FFB2971CFB511AC70B62FC55" box="[1274,1347,1001,1023]" italics="true" pageId="2" pageNumber="3">Q. tzwu</emphasis>
Smetana
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="FC631102FFB2971CFF3C1D290BBDFB90" blockId="2.[151,1437,151,1229]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1D290BBDFB90" pageId="2" pageNumber="3">
- Paramere of aedeagus relatively shorter, with apex distinctly not reaching that of median lobe, with about six sensory peg setae on its underside........................................................................
<taxonomicName id="3BDC6A81FFB2971CFB431D0A0BBDFB90" ID-CoL="4R2K9" authority="Smetana" authorityName="Smetana" box="[1256,1436,1060,1082]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="antoni">
<emphasis id="CEA8CD10FFB2971CFB431D0A0B62FB90" box="[1256,1347,1060,1082]" italics="true" pageId="2" pageNumber="3">Q. antoni</emphasis>
Smetana
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="472809A7FFB2971CFF3C1D6F0BBDFBDF" pageId="2" pageNumber="3">
<paragraph id="FC631102FFB2971CFF3C1D6F0BBDFBF2" blockId="2.[151,1437,151,1229]" box="[151,1436,1089,1112]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1D6F0BBDFBF2" box="[151,1436,1089,1112]" pageId="2" pageNumber="3">
14. Paramere of aedeagus straight and parallel-sided (
<figureCitation id="64E70D87FFB2971CFD141D6F0D20FBFD" box="[703,769,1089,1111]" captionStart="FIGURE 5" captionStartId="14.[151,250,1877,1899]" captionTargetBox="[193,1433,215,1855]" captionTargetId="figure@14.[151,1436,193,1856]" captionTargetPageId="14" captionText="FIGURE 5. Quedius (Microsaurus) rectus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/289383/files/figure.png" pageId="2" pageNumber="3">Figs. 5</figureCitation>
D; 5-1F).................................
<taxonomicName id="3BDC6A81FFB2971CFB591D6F0B68FBFD" ID-CoL="78PY5" box="[1266,1353,1089,1111]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="rectus" status="sp. nov.">
<emphasis id="CEA8CD10FFB2971CFB591D6F0B68FBFD" box="[1266,1353,1089,1111]" italics="true" pageId="2" pageNumber="3">Q. rectus</emphasis>
</taxonomicName>
<emphasis id="CEA8CD10FFB2971CFAE41D6C0BBDFBF2" bold="true" box="[1359,1436,1090,1112]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="D59B706BFFB2971CFAE41D6C0BBDFBF2" box="[1359,1436,1090,1112]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="FC631102FFB2971CFF3C1D710BBDFBDF" blockId="2.[151,1437,151,1229]" box="[151,1436,1119,1141]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1D710BBDFBDF" box="[151,1436,1119,1141]" pageId="2" pageNumber="3">- Paramere of aedeagus slightly curved or not parallel-sided................................................... 15</keyLead>
</paragraph>
</keyStep>
<keyStep id="472809A7FFB2971CFF3C1D520BBDFB67" pageId="2" pageNumber="3">
<paragraph id="FC631102FFB2971CFF3C1D520BBAFB05" blockId="2.[151,1437,151,1229]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1D520BBAFB05" pageId="2" pageNumber="3">
15. Paramere of aedeagus straight, gradually narrowed to apex (
<figureCitation id="64E70D87FFB2971CFC8A1D520D45FB38" box="[801,868,1148,1170]" captionStart="FIGURE 6" captionStartId="17.[151,250,1880,1902]" captionTargetBox="[193,1433,212,1858]" captionTargetId="figure@17.[151,1436,195,1858]" captionTargetPageId="17" captionText="FIGURE 6. Quedius (Microsaurus) guoi Zheng &amp; Zheng, 2006: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/289384/files/figure.png" pageId="2" pageNumber="3">Figs. 6</figureCitation>
D; 6-1F).........................................................................................................................
<taxonomicName id="3BDC6A81FFB2971CFB1F1DB70BBAFB05" ID-CoL="4R2NX" authority="Zheng &amp; Zheng" authorityName="Zheng &amp; Zheng" box="[1204,1435,1177,1199]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="guoi">
<emphasis id="CEA8CD10FFB2971CFB1F1DB70ADCFB05" box="[1204,1277,1177,1199]" italics="true" pageId="2" pageNumber="3">Q. guoi</emphasis>
Zheng &amp; Zheng
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="FC631102FFB2971CFF3C1D990BBDFB67" blockId="2.[151,1437,151,1229]" box="[151,1436,1207,1229]" pageId="2" pageNumber="3">
<keyLead id="472DB237FFB2971CFF3C1D990BBDFB67" box="[151,1436,1207,1229]" pageId="2" pageNumber="3">
- Paramere of aedeagus slightly curved toward left, nearly parallel-sided (
<figureCitation id="64E70D87FFB2971CFC3F1D990DF6FB67" box="[916,983,1207,1229]" captionStart="FIGURE 2" captionStartId="4.[151,250,1877,1899]" captionTargetBox="[193,1433,216,1855]" captionTargetId="figure@4.[151,1436,193,1856]" captionTargetPageId="4" captionText="FIGURE 2. Quedius (Microsaurus) indigestus sp. nov.: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/289378/files/figure.png" pageId="2" pageNumber="3">Figs. 2</figureCitation>
D; 2-1F)............
<taxonomicName id="3BDC6A81FFB2971CFB611D990B68FB67" ID-CoL="4R2PM" box="[1226,1353,1207,1229]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="indigestus" status="sp. nov.">
<emphasis id="CEA8CD10FFB2971CFB611D990B68FB67" box="[1226,1353,1207,1229]" italics="true" pageId="2" pageNumber="3">Q. indigestus</emphasis>
</taxonomicName>
<emphasis id="CEA8CD10FFB2971CFAE41D990BBDFB67" bold="true" box="[1359,1436,1207,1229]" pageId="2" pageNumber="3">
<taxonomicNameLabel id="D59B706BFFB2971CFAE41D990BBDFB67" box="[1359,1436,1207,1229]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
</subSubSection>
</treatment>
</document>