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<document id="80136BE63746B7EC14AA877C7DD26CFE" ID-DOI="10.5281/zenodo.12587480" ID-ISSN="2064-2474" ID-Zenodo-Dep="12587480" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719597665362" checkinUser="felipe" docAuthor="Schawaller, W." docDate="2002" docId="742A878A5124FFF4FD8F589AD6B4FDF9" docLanguage="en" docName="ActaZoolAcadSciH.48.3.197-202.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 48 (3)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Malayoscelis gebieni Schawaller, 2002, sp. n." docType="treatment" docVersion="2" lastPageNumber="202" masterDocId="8813FFF25126FFF1FFF9592DD541FFFD" masterDocTitle="Malayoscelis Gen. N., The Third Genus Of The Pycnocerini (Coleoptera: Tenebrionidae) From The Oriental Region" masterLastPageNumber="202" masterPageNumber="197" pageNumber="199" updateTime="1719604834054" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="64705FEC3CB96D6E7845BC9D31B422E3">Malayoscelis Gen. N., The Third Genus Of The Pycnocerini (Coleoptera: Tenebrionidae) From The Oriental Region</mods:title>
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<mods:date id="3C529C228CEC9C84F77EED6D41BC7B72">2002</mods:date>
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<taxonomicName id="3B834D1F5124FFF3FD8F589AD636FE2C" authorityName="Schawaller" authorityYear="2002" box="[630,887,439,465]" class="Insecta" family="Tenebrionidae" genus="Malayoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="gebieni" status="sp. nov.">
<emphasis id="CEF7EA8E5124FFF3FD8F589AD636FE2C" bold="true" box="[630,887,439,465]" pageId="2" pageNumber="199">Malayoscelis gebieni</emphasis>
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(
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,
<figureCitation id="64B82A195124FFF3FCD158F7D622FE09" box="[808,867,474,500]" captionStart="Figs 310" captionStartId="3.[317,360,1745,1767]" captionTargetBox="[346,1250,832,1701]" captionTargetId="figure-172@3.[334,1253,813,1716]" captionTargetPageId="3" captionText="Figs 310. Malayoscelis gen.n. gebieni sp. n. 3: dorsal view; 4: antenna; 5: maxillary palp; 6: anterior leg of male; 7: posterior leg of male; 8: prosternal process; 9: aedeagus; 10: gonostyles and last coxite" figureDoi="http://doi.org/10.5281/zenodo.12587485" httpUri="https://zenodo.org/record/12587485/files/figure.png" pageId="2" pageNumber="199">310</figureCitation>
)
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<typeStatus id="2338883E5124FFF3FE875B33D496FDC9" box="[382,471,542,564]" pageId="2" pageNumber="199" type="holotype">Holotype</typeStatus>
(male):
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,
<location id="F95C60475124FFF3FD6B5B33D617FDC9" LSID="urn:lsid:plazi:treatment:742A878A5124FFF4FD8F589AD6B4FDF9:F95C60475124FFF3FD6B5B33D617FDC9" box="[658,854,542,564]" country="Malaysia" municipality="Tanah Rata" name="Cameron Highlands" pageId="2" pageNumber="199">Cameron Highlands</location>
,
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leg. I.
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:
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data as holotype,
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,
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leg. U.
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leg. S. &amp; S.
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,
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leg. S. &amp; S.
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leg. S.
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leg. S.
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leg. M.
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<emphasis id="CEF7EA8E5124FFF3FEC45FC2D4D8F8F8" bold="true" box="[317,409,1775,1797]" pageId="2" pageNumber="199">Figs 12.</emphasis>
Dorsal view. 1:
<taxonomicName id="3B834D1F5124FFF3FDC75FC2D7FCF8F8" authorityName="Schawaller" authorityYear="2002" box="[574,701,1775,1797]" class="Insecta" family="Tenebrionidae" genus="Malayoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CEF7EA8E5124FFF3FDC75FC2D7FCF8F8" box="[574,701,1775,1797]" italics="true" pageId="2" pageNumber="199">Malayoscelis</emphasis>
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<taxonomicName id="3B834D1F5124FFF3FCF45FC2D615F8F8" authorityName="Schawaller" authorityYear="2002" box="[781,852,1775,1797]" class="Insecta" family="Tenebrionidae" genus="Malayoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="199" phylum="Arthropoda" rank="species" species="gebieni" status="sp. nov.">
<emphasis id="CEF7EA8E5124FFF3FCF45FC2D615F8F8" box="[781,852,1775,1797]" italics="true" pageId="2" pageNumber="199">gebieni</emphasis>
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(Malaysia); 2:
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<emphasis id="CEF7EA8E5124FFF3FBDE5FC2D1CDF8F8" box="[1063,1164,1775,1797]" italics="true" pageId="2" pageNumber="199">Aediotorix</emphasis>
cf.
<emphasis id="CEF7EA8E5124FFF3FB4D5FDDD1B4F8F8" box="[1204,1269,1776,1797]" italics="true" pageId="2" pageNumber="199">petersi</emphasis>
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, 1921
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(Philippines)
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<paragraph id="FC3C369C5125FFF2FE875899D169F8DD" blockId="3.[317,1270,436,775]" lastBlockId="3.[317,1270,1745,1824]" pageId="3" pageNumber="200">
Description (male): Body length 16.017.5 mm. Body and appendages unicoloured dark brown to black, elytra sometimes indistinctly paler; surface of head and pronotum dull, of elytra somewhat shining; without regular setation. Head (
<figureCitation id="64B82A195125FFF2FCD058C3D623FDF9" box="[809,866,494,516]" captionStart="Figs 310" captionStartId="3.[317,360,1745,1767]" captionTargetBox="[346,1250,832,1701]" captionTargetId="figure-172@3.[334,1253,813,1716]" captionTargetPageId="3" captionText="Figs 310. Malayoscelis gen.n. gebieni sp. n. 3: dorsal view; 4: antenna; 5: maxillary palp; 6: anterior leg of male; 7: posterior leg of male; 8: prosternal process; 9: aedeagus; 10: gonostyles and last coxite" figureDoi="http://doi.org/10.5281/zenodo.12587485" httpUri="https://zenodo.org/record/12587485/files/figure.png" pageId="3" pageNumber="200">Fig. 3</figureCitation>
) with coarse, confluent punctation; genae above antennal insertion without punctation and shining; clypeal suture distinctly impressed; clypeus excavated; temples considerably broadened and separated from the narrow neck; proportions of antennomeres see
<figureCitation id="64B82A195125FFF2FE0E5B69D777FDA7" box="[503,566,580,602]" captionStart="Figs 310" captionStartId="3.[317,360,1745,1767]" captionTargetBox="[346,1250,832,1701]" captionTargetId="figure-172@3.[334,1253,813,1716]" captionTargetPageId="3" captionText="Figs 310. Malayoscelis gen.n. gebieni sp. n. 3: dorsal view; 4: antenna; 5: maxillary palp; 6: anterior leg of male; 7: posterior leg of male; 8: prosternal process; 9: aedeagus; 10: gonostyles and last coxite" figureDoi="http://doi.org/10.5281/zenodo.12587485" httpUri="https://zenodo.org/record/12587485/files/figure.png" pageId="3" pageNumber="200">Fig. 4</figureCitation>
, antennomere 2 short, antennomere 3 prolonged, antennomeres 410 subquadrate, antennomere 11 twice aslong asantennomere 10; both mandiblesbifid; maxillary palps (
<figureCitation id="64B82A195125FFF2FEBD5B53D43DFD69" box="[324,380,638,660]" captionStart="Figs 310" captionStartId="3.[317,360,1745,1767]" captionTargetBox="[346,1250,832,1701]" captionTargetId="figure-172@3.[334,1253,813,1716]" captionTargetPageId="3" captionText="Figs 310. Malayoscelis gen.n. gebieni sp. n. 3: dorsal view; 4: antenna; 5: maxillary palp; 6: anterior leg of male; 7: posterior leg of male; 8: prosternal process; 9: aedeagus; 10: gonostyles and last coxite" figureDoi="http://doi.org/10.5281/zenodo.12587485" httpUri="https://zenodo.org/record/12587485/files/figure.png" pageId="3" pageNumber="200">Fig. 5</figureCitation>
) with broadened last segment; mentum pentagonal with an impression on each side and with a weak longitudinal keel medially; underside of head with rounded gular impression. Pronotum (
<figureCitation id="64B82A195125FFF2FB325BB7D408FD30" captionStart="Figs 12" captionStartId="2.[317,360,1775,1797]" captionTargetBox="[375,1214,866,1728]" captionTargetId="figure-169@2.[366,1220,852,1746]" captionTargetPageId="2" captionText="Figs 12. Dorsal view. 1: Malayoscelis gen. n. gebieni sp. n. (Malaysia); 2: Aediotorix cf. petersi GEBIEN, 1921 (Philippines)" figureDoi="http://doi.org/10.5281/zenodo.12587482" httpUri="https://zenodo.org/record/12587482/files/figure.png" pageId="3" pageNumber="200">Figs 1</figureCitation>
,
<figureCitation id="64B82A195125FFF2FEAF5B9AD423FD30" box="[342,354,695,717]" captionStart="Figs 310" captionStartId="3.[317,360,1745,1767]" captionTargetBox="[346,1250,832,1701]" captionTargetId="figure-172@3.[334,1253,813,1716]" captionTargetPageId="3" captionText="Figs 310. Malayoscelis gen.n. gebieni sp. n. 3: dorsal view; 4: antenna; 5: maxillary palp; 6: anterior leg of male; 7: posterior leg of male; 8: prosternal process; 9: aedeagus; 10: gonostyles and last coxite" figureDoi="http://doi.org/10.5281/zenodo.12587485" httpUri="https://zenodo.org/record/12587485/files/figure.png" pageId="3" pageNumber="200">3</figureCitation>
) 1.3 times wider than medially long, with coarse and confluent punctation as on head, surface somewhat uneven; all margins bordered and shining, lateral margin distinctly crenulate; propleures with punctation sparser than on disc. Prosternum (
<figureCitation id="64B82A195125FFF2FCE45BDCD614FCFA" box="[797,853,753,775]" captionStart="Figs 310" captionStartId="3.[317,360,1745,1767]" captionTargetBox="[346,1250,832,1701]" captionTargetId="figure-172@3.[334,1253,813,1716]" captionTargetPageId="3" captionText="Figs 310. Malayoscelis gen.n. gebieni sp. n. 3: dorsal view; 4: antenna; 5: maxillary palp; 6: anterior leg of male; 7: posterior leg of male; 8: prosternal process; 9: aedeagus; 10: gonostyles and last coxite" figureDoi="http://doi.org/10.5281/zenodo.12587485" httpUri="https://zenodo.org/record/12587485/files/figure.png" pageId="3" pageNumber="200">Fig. 8</figureCitation>
) with a rounded, flat prosternal process not lobes of female. Scale line: 10.0 mm (3), 5.0 mm (410)
</paragraph>
<caption id="A8FC66145125FFF2FEC45FFCD1B7F8F9" ID-DOI="http://doi.org/10.5281/zenodo.12587485" ID-Zenodo-Dep="12587485" httpUri="https://zenodo.org/record/12587485/files/figure.png" pageId="3" pageNumber="200" startId="3.[317,360,1745,1767]" targetBox="[346,1250,832,1701]" targetPageId="3" targetType="figure">
<paragraph id="FC3C369C5125FFF2FEC45FFCD1B7F8F9" blockId="3.[317,1270,1745,1824]" pageId="3" pageNumber="200">
<emphasis id="CEF7EA8E5125FFF2FEC45FFCD4E0F91A" bold="true" box="[317,417,1745,1767]" pageId="3" pageNumber="200">Figs 310.</emphasis>
<taxonomicName id="3B834D1F5125FFF2FE5F5FFCD764F91A" authorityName="Schawaller" authorityYear="2002" box="[422,549,1745,1767]" class="Insecta" family="Tenebrionidae" genus="Malayoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="200" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="CEF7EA8E5125FFF2FE5F5FFCD764F91A" box="[422,549,1745,1767]" italics="true" pageId="3" pageNumber="200">Malayoscelis</emphasis>
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<taxonomicNameLabel id="D5C457F55125FFF2FDD05FFCD729F91A" box="[553,616,1745,1767]" pageId="3" pageNumber="200" rank="genus">gen. n.</taxonomicNameLabel>
<taxonomicName id="3B834D1F5125FFF2FD955FFCD7F2F91A" authorityName="Schawaller" authorityYear="2002" box="[620,691,1745,1767]" class="Insecta" family="Tenebrionidae" genus="Malayoscelis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="200" phylum="Arthropoda" rank="species" species="gebieni" status="sp. nov.">
<emphasis id="CEF7EA8E5125FFF2FD955FFCD7F2F91A" box="[620,691,1745,1767]" italics="true" pageId="3" pageNumber="200">gebieni</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D5C457F55125FFF2FD4E5FFCD7A8F91A" box="[695,745,1745,1767]" pageId="3" pageNumber="200" rank="species">sp. n.</taxonomicNameLabel>
3: dorsal view; 4: antenna; 5: maxillary palp; 6: anterior leg of male; 7: posterior leg of male; 8: prosternal process; 9: aedeagus; 10: gonostyles and last coxite
</paragraph>
</caption>
<paragraph id="FC3C369C5122FFF5FEC45899D626FCBD" blockId="4.[317,1270,436,890]" pageId="4" pageNumber="201">
distinctly surpassing posterior margin. Mesoventrite with dull and confluent punctation. Metaventrite medially impunctate and shining, with a distinct longitudinal line medially. Scutellum only with fine and separate punctures. Elytra (
<figureCitation id="64B82A195122FFF5FD3D58C3D640FDF9" box="[708,769,494,516]" captionStart="Figs 12" captionStartId="2.[317,360,1775,1797]" captionTargetBox="[375,1214,866,1728]" captionTargetId="figure-169@2.[366,1220,852,1746]" captionTargetPageId="2" captionText="Figs 12. Dorsal view. 1: Malayoscelis gen. n. gebieni sp. n. (Malaysia); 2: Aediotorix cf. petersi GEBIEN, 1921 (Philippines)" figureDoi="http://doi.org/10.5281/zenodo.12587482" httpUri="https://zenodo.org/record/12587482/files/figure.png" pageId="4" pageNumber="201">Figs 1</figureCitation>
,
<figureCitation id="64B82A195122FFF5FCF558C3D659FDF9" box="[780,792,494,516]" captionStart="Figs 310" captionStartId="3.[317,360,1745,1767]" captionTargetBox="[346,1250,832,1701]" captionTargetId="figure-172@3.[334,1253,813,1716]" captionTargetPageId="3" captionText="Figs 310. Malayoscelis gen.n. gebieni sp. n. 3: dorsal view; 4: antenna; 5: maxillary palp; 6: anterior leg of male; 7: posterior leg of male; 8: prosternal process; 9: aedeagus; 10: gonostyles and last coxite" figureDoi="http://doi.org/10.5281/zenodo.12587485" httpUri="https://zenodo.org/record/12587485/files/figure.png" pageId="4" pageNumber="201">3</figureCitation>
) long and parallel-sided; with traces of 57 keels, these keels shining, between the keels with shallow and confluent punctation, surface with additional microgranules; epipleura regularly narrowed toward apex, without separated constriction, shining and without puncturesbut with microgranulesin the humeral part. Wingsfully developed. Abdomi - nal ventrites unpunctured, but with leather-like microstructure, last visible ventrite unbordered and without distinct modifications. Legs (
<figureCitation id="64B82A195122FFF5FD515B53D7BEFD69" box="[680,767,638,660]" captionStart="Figs 310" captionStartId="3.[317,360,1745,1767]" captionTargetBox="[346,1250,832,1701]" captionTargetId="figure-172@3.[334,1253,813,1716]" captionTargetPageId="3" captionText="Figs 310. Malayoscelis gen.n. gebieni sp. n. 3: dorsal view; 4: antenna; 5: maxillary palp; 6: anterior leg of male; 7: posterior leg of male; 8: prosternal process; 9: aedeagus; 10: gonostyles and last coxite" figureDoi="http://doi.org/10.5281/zenodo.12587485" httpUri="https://zenodo.org/record/12587485/files/figure.png" pageId="4" pageNumber="201">Figs 67</figureCitation>
) with the anterior and middle femora with a single, posterior femora with 2 distinct teeth shortly before apex; anterior tibia bent, posterior tibia straight; all tibiae with crenulate inner side; anterior and middle tibiae with distinctly hooked inner apex; apex of tibiae without spurs and with comb-like setae; tarsal segments not dilated in males; claw segment of all legs longer than basal segments combined; trochanter of all legs without spines or other peculiarities; surface of all legs nearly impunctate and shining. Aedeagus (
<figureCitation id="64B82A195122FFF5FC2A5A23D14DFCD9" box="[979,1036,782,804]" captionStart="Figs 310" captionStartId="3.[317,360,1745,1767]" captionTargetBox="[346,1250,832,1701]" captionTargetId="figure-172@3.[334,1253,813,1716]" captionTargetPageId="3" captionText="Figs 310. Malayoscelis gen.n. gebieni sp. n. 3: dorsal view; 4: antenna; 5: maxillary palp; 6: anterior leg of male; 7: posterior leg of male; 8: prosternal process; 9: aedeagus; 10: gonostyles and last coxite" figureDoi="http://doi.org/10.5281/zenodo.12587485" httpUri="https://zenodo.org/record/12587485/files/figure.png" pageId="4" pageNumber="201">Fig. 9</figureCitation>
) quite small in comparison with the body length, connate parameres finger-like.
</paragraph>
<paragraph id="FC3C369C5122FFF5FE875A6AD6A1FC87" blockId="4.[317,1270,436,890]" pageId="4" pageNumber="201">Sexual dimorphism: Not striking. Females also with armed femora, but anterior tibia somewhat straighter, anterior and middle tibia without hooked inner apex.</paragraph>
<paragraph id="FC3C369C5122FFF5FE875AB3D758FC10" blockId="4.[317,1270,926,1005]" pageId="4" pageNumber="201">
Etymology: Thistaxon isnamed in honour of H
<smallCapsWord id="FADAA0405122FFF5FCBF5A8CD62AFC4F" baselines="942" box="[838,875,929,946]" lowerCaseFontSize="7" mainFontSize="9" normCase="lower" normString="ans" pageId="4" pageNumber="201">ANS</smallCapsWord>
<smallCapsWord id="FADAA0405122FFF5FC885AB3D6F8FC4F" baselines="942,942" box="[881,953,926,948]" lowerCaseFontSize="7" mainFontSize="9" normCase="title" normString="Gebien" pageId="4" pageNumber="201">GEBIEN</smallCapsWord>
(18741947), whose monograph of the
<taxonomicName id="3B834D1F5122FFF5FE7A5A97D4B7FC2D" box="[387,502,954,976]" pageId="4" pageNumber="198" rank="tribe" tribe="Pycnocerini">Pycnocerini</taxonomicName>
(1904) was the first of his numerous and substantial contributions towards tenebrionid taxonomy.
</paragraph>
<caption id="A8FC66145122FFF5FE355FDAD126F8F0" ID-DOI="http://doi.org/10.5281/zenodo.12587487" ID-Zenodo-Dep="12587487" box="[460,1127,1783,1805]" httpUri="https://zenodo.org/record/12587487/files/figure.png" pageId="4" pageNumber="201" startId="4.[460,495,1783,1805]" targetBox="[317,1270,1101,1754]" targetPageId="4" targetType="figure">
<paragraph id="FC3C369C5122FFF5FE355FDAD126F8F0" blockId="4.[460,1127,1783,1805]" box="[460,1127,1783,1805]" pageId="4" pageNumber="201">
<emphasis id="CEF7EA8E5122FFF5FE355FDAD756F8F0" bold="true" box="[460,535,1783,1805]" pageId="4" pageNumber="201">Fig. 11.</emphasis>
Distribution of the known genera of the Oriental
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</paragraph>
</caption>
<paragraph id="FC3C369C5123FFF4FE875899D6B4FDF9" blockId="5.[317,1269,436,516]" pageId="5" pageNumber="202">
Biology: Adult beetleswere collected in standing trunksof dead broadleaved trees, together with larvae and pupae. The rotten, white coloured substrate was said to be extremely dry. A species of
<taxonomicName id="3B834D1F5123FFF4FEC458C3D4E3FDF9" authorityName="BATES" authorityYear="1868" box="[317,418,494,516]" class="Insecta" family="Tenebrionidae" genus="Aediotorix" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="202" phylum="Arthropoda" rank="genus">
<emphasis id="CEF7EA8E5123FFF4FEC458C3D4E3FDF9" box="[317,418,494,516]" italics="true" pageId="5" pageNumber="202">Aediotorix</emphasis>
</taxonomicName>
wasfound syntopically (B EČVÁŘ, personal communication).
</paragraph>
</subSubSection>
</treatment>
</document>