139 lines
15 KiB
XML
139 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.308.5559" ID-Pensoft-Pub="1313-2970-308-1" ID-Pensoft-UUID="FFA2FFFEFFE74632FFB7FFC7D553CE26" ID-Zenodo-Dep="577662" ID-ZooBank="EEC31850AAAB4081B05AB80A2D944658" ModsDocID="1313-2970-308-1" checkinTime="1451247225725" checkinUser="pensoft" docAuthor="Rodrigues, Taissa & Kellner, Alexander Wilhelm Armin" docDate="2013" docId="6C9CBC4AEFD2CD19341E0E6E0367748E" docLanguage="en" docName="ZooKeys 308: 1-112" docOrigin="ZooKeys 308" docPubDate="2013-06-12" docSource="http://dx.doi.org/10.3897/zookeys.308.5559" docTitle="Ornithocheirus carteri Seeley 1870, (nomen dubium)" docType="treatment" docVersion="5" id="FFA2FFFEFFE74632FFB7FFC7D553CE26" lastPageNumber="42" masterDocId="FFA2FFFEFFE74632FFB7FFC7D553CE26" masterDocTitle="Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England" masterLastPageNumber="112" masterPageNumber="1" pageNumber="42" updateTime="1634706563434" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Taxonomic review of the Ornithocheirus complex (Pterosauria) from the Cretaceous of England</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Rodrigues, Taissa</mods:namePart>
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<mods:affiliation>Department of Biology, Agrarian Sciences Center, Universidade Federal do Espirito Santo. Alto Universitario s / n, Caixa Postal 16, Guararema, CEP 29500 - 000, Alegre, ES, Brazil</mods:affiliation>
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<mods:nameIdentifier type="email">taissa.rodrigues@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kellner, Alexander Wilhelm Armin</mods:namePart>
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<mods:affiliation>Laboratory of Systematics and Taphonomy of Fossil Vertebrates, Department of Geology and Paleontology, Museu Nacional / Universidade Federal do Rio de Janeiro. Quinta da Boa Vista s / n, Sao Cristovao, CEP 20940 - 040, Rio de Janeiro, RJ, Brazil</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2013-06-12</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>308</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>112</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.308.5559</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.308.5559</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-308-1</mods:identifier>
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<mods:identifier type="ZooBank">EEC31850AAAB4081B05AB80A2D944658</mods:identifier>
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<mods:identifier type="Pensoft-UUID">FFA2FFFEFFE74632FFB7FFC7D553CE26</mods:identifier>
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<mods:identifier type="Zenodo-Dep">577662</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152044997" LSID="urn:lsid:plazi:treatment:6C9CBC4AEFD2CD19341E0E6E0367748E" httpUri="http://treatment.plazi.org/id/6C9CBC4AEFD2CD19341E0E6E0367748E" lastPageId="42" lastPageNumber="42" pageId="41" pageNumber="42">
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<subSubSection pageId="41" pageNumber="42" type="nomenclature">
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<paragraph pageId="41" pageNumber="42">
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<taxonomicName LSID="6C9CBC4A-EFD2-CD19-341E-0E6E0367748E" authority="Seeley, 1870" authorityName="Seeley" authorityYear="1870" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="Ornithocheirus carteri" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="carteri" status="(nomen dubium)">Ornithocheirus carteri Seeley, 1870</taxonomicName>
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<taxonomicNameLabel pageId="41" pageNumber="42">(nomen dubium)</taxonomicNameLabel>
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. A-D Ornithocheirus brachyrhinus, holotype CAMSM B 54443 (Albian, Cambridge Greensand), anterior part of the rostrum. A right lateral view B respective line drawing C ventral view D respective line drawing. E-H Ornithocheirus carteri, holotype CAMSM B 54437 (Albian, Cambridge Greensand), anterior part of the rostrum E anterior view F respective line drawing G left lateral view H respective line drawing. Abbreviations: m - maxillae, pm - premaxillae, prid - palatal ridge. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm." httpUri="https://binary.pensoft.net/fig/22084" pageId="41" pageNumber="42">Fig. 16E-H</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="41" pageNumber="42" type="reference_group">
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<paragraph pageId="41" pageNumber="42">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="carteri" status="(nomen dubium)">Ornithocheirus carteri</taxonomicName>
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Seeley:
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B77" refString="Seeley, HG, 1869. Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell and Co., Cambridge, xxiii + 143 pp." title="Index to the fossil remains of Aves, Ornithosauria, and Reptilia, from the Secondary System of Strata arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell and Co., Cambridge, xxiii + 143 pp." year="1869">Seeley 1869</bibRefCitation>
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: p. xvi [disclaimed]
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</paragraph>
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<paragraph pageId="41" pageNumber="42">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="carteri" status="(nomen dubium)">Ornithocheirus carteri</taxonomicName>
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Seeley:
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley 1870</bibRefCitation>
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: p. 128
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</paragraph>
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<paragraph pageId="41" pageNumber="42">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Criorhynchidae" genus="Criorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="carteri" status="(nomen dubium)">Criorhynchus carteri</taxonomicName>
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(Seeley):
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<bibRefCitation author="Hooley, RW" journalOrPublisher="Annals and Magazine of Natural History" pageId="66" pageNumber="67" pagination="529 - 557" publicationUrl="10.1080/00222931408693521" refId="B26" refString="Hooley, RW, 1914. On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge. Annals and Magazine of Natural History 78: 529 - 557, 10.1080/00222931408693521" title="On the ornithosaurian genus Ornithocheirus, with a review of the specimens from the Cambridge Greensand in the Sedgwick Museum, Cambridge." url="10.1080/00222931408693521" volume="78" year="1914">Hooley 1914</bibRefCitation>
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: p. 536
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</paragraph>
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<paragraph pageId="41" pageNumber="42">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Criorhynchidae" genus="Criorhynchus" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="simus" status="(nomen dubium)">Criorhynchus simus</taxonomicName>
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(Owen):
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<bibRefCitation author="Wellnhofer, P" journalOrPublisher="Gustav Fischer Verlag, Stuttgart and New York" pageId="70" pageNumber="71" refId="B101" refString="Wellnhofer, P, 1978. Pterosauria. Handbuch der Palaeoherpetologie, Teil 19. Gustav Fischer Verlag, Stuttgart and New York, 82 pp." title="Pterosauria. Handbuch der Palaeoherpetologie, Teil 19." year="1978">Wellnhofer 1978</bibRefCitation>
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: p. 60
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</paragraph>
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<paragraph pageId="41" pageNumber="42">
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="41" pageNumber="42" phylum="Chordata" rank="species" species="simus" status="(nomen dubium)">Ornithocheirus simus</taxonomicName>
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(Owen):
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<bibRefCitation author="Unwin, DM" journalOrPublisher="Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe" pageId="69" pageNumber="70" pagination="189 - 221" publicationUrl="10.1002/mmng.20010040112" refId="B85" refString="Unwin, DM, 2001. An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England. Mitteilungen aus dem Museum fuer Naturkunde in Berlin, Geowissenschaftliche Reihe 4: 189 - 221, 10.1002/mmng.20010040112" title="An overview of the pterosaur assemblage from the Cambridge Greensand (Cretaceous) of Eastern England." url="10.1002/mmng.20010040112" volume="4" year="2001">Unwin 2001</bibRefCitation>
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: table 1
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</paragraph>
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</subSubSection>
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<subSubSection pageId="41" pageNumber="42" type="holotype">
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<paragraph pageId="41" pageNumber="42">Holotype.</paragraph>
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<paragraph pageId="41" pageNumber="42">
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CAMSM B 54437, anterior portion of the rostrum (
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<figureCitation captionStart="Figure 16" captionStartId="F16" captionText="Figure 16. A-D Ornithocheirus brachyrhinus, holotype CAMSM B 54443 (Albian, Cambridge Greensand), anterior part of the rostrum. A right lateral view B respective line drawing C ventral view D respective line drawing. E-H Ornithocheirus carteri, holotype CAMSM B 54437 (Albian, Cambridge Greensand), anterior part of the rostrum E anterior view F respective line drawing G left lateral view H respective line drawing. Abbreviations: m - maxillae, pm - premaxillae, prid - palatal ridge. Arrows and numbers indicate alveoli or teeth and their respective position. Scale bar = 10 mm." httpUri="https://binary.pensoft.net/fig/22084" pageId="41" pageNumber="42">Fig. 16E-H</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="41" pageNumber="42" type="type locality">
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<paragraph pageId="41" pageNumber="42">Type locality.</paragraph>
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<paragraph pageId="41" pageNumber="42">Cambridge, Cambridgeshire, England.</paragraph>
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</subSubSection>
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<subSubSection pageId="41" pageNumber="42" type="type horizon">
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<paragraph pageId="41" pageNumber="42">Type horizon.</paragraph>
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<paragraph pageId="41" pageNumber="42">Cambridge Greensand (Cenomanian; fossils Albian in age).</paragraph>
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</subSubSection>
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<subSubSection lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42" type="description">
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<paragraph pageId="41" pageNumber="42">Description.</paragraph>
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<paragraph lastPageId="42" lastPageNumber="43" pageId="41" pageNumber="42">
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<bibRefCitation author="Seeley, HG" journalOrPublisher="Zitteliana B" pageId="69" pageNumber="70" refId="B78" refString="Seeley, HG, 1870. The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." title="The Ornithosauria: an elementary study of the bones of pterodactyls, made from fossil remains found in the Cambridge Upper Greensand, and arranged in the Woodwardian Museum of the University of Cambridge. Deighton, Bell, and Co., Cambridge, xii + 135 pp." year="1870">Seeley (1870)</bibRefCitation>
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named this species based on some differences between CAMSM B 54437 and the holotype of
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="simus">
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<emphasis italics="true" pageId="42" pageNumber="43">
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<pageBreakToken pageId="42" pageNumber="43" start="start">Ornithocheirus</pageBreakToken>
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simus
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</emphasis>
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</taxonomicName>
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. He noted that the rostrum is not as high and narrower; the lateral surfaces bear several longitudinal furrows, which he believed to be impressions of blood vessels; and the first pair of teeth are larger, more conical, circular, and separated by a large gap. Although the rostrum is not as tall as in
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="simus">
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<emphasis italics="true" pageId="42" pageNumber="43">Ornithocheirus simus</emphasis>
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</taxonomicName>
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, it is not possible to determine if this size difference is merely ontogenetic. However, the separation between the right and left teeth of the first pair of alveoli is larger than the holotype of
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="simus">
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<emphasis italics="true" pageId="42" pageNumber="43">Ornithocheirus simus</emphasis>
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</taxonomicName>
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. Therefore, we reject referring
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="carteri">
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<emphasis italics="true" pageId="42" pageNumber="43">Ornithocheirus carteri</emphasis>
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</taxonomicName>
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to
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<taxonomicName authorityName="Rodrigues & Kellner" authorityYear="2013" class="Reptilia" family="Anhangueridae" genus="Ornithocheirus" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Pterosauria" pageId="42" pageNumber="43" phylum="Chordata" rank="species" species="simus">
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<emphasis italics="true" pageId="42" pageNumber="43">Ornithocheirus simus</emphasis>
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</taxonomicName>
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and consider it a
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<emphasis italics="true" pageId="42" pageNumber="43">nomen dubium</emphasis>
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.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |