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<document id="19CF83C5E37F84C9986FF1817633AE3C" ID-DOI="10.11646/zootaxa.2751.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5291314" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629712257117" checkinUser="felipe" docAuthor="Gosliner, Terrence" docDate="2011" docId="394D87A1FFFA2343FF5EFDBDF9216D92" docLanguage="en" docName="zt02751p024.pdf" docOrigin="Zootaxa 2751 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2751.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Philinopsis coronata Gosliner 2011, n. sp." docType="treatment" docVersion="4" lastPageNumber="9" masterDocId="C574FFD9FFFC234AFFC9FFD1FB716D41" masterDocTitle="Six new species of aglajid opisthobranch mollusks from the tropical Indo-Pacific" masterLastPageNumber="24" masterPageNumber="1" pageNumber="7" updateTime="1699100175227" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="D2ECBC21B4008B6A7CC9365A29987454">Six new species of aglajid opisthobranch mollusks from the tropical Indo-Pacific</mods:title>
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<mods:namePart id="AD9B4AEFD090ADC024DD18F3B6DF21AA">Gosliner, Terrence</mods:namePart>
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<mods:date id="0DEC181A08E230E88A069ADBE6D90494">2011</mods:date>
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<mods:number id="EE2576556DBB86A34AB96E67580ACAC1">2011-01-28</mods:number>
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<mods:number id="D0F94123B2CF9C228D327265658F1125">2751</mods:number>
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<mods:url id="0AC3871BEF2602A6C62D2FA787F1A841">https://biotaxa.org/Zootaxa/article/view/zootaxa.2751.1.1</mods:url>
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<mods:identifier id="7C9FC374518F15A6F5AFD9E25493E0B5" type="ISSN">1175-5326</mods:identifier>
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<treatment id="394D87A1FFFA2343FF5EFDBDF9216D92" ID-DOI="http://doi.org/10.5281/zenodo.5294914" ID-Zenodo-Dep="5294914" LSID="urn:lsid:plazi:treatment:394D87A1FFFA2343FF5EFDBDF9216D92" httpUri="http://treatment.plazi.org/id/394D87A1FFFA2343FF5EFDBDF9216D92" lastPageId="9" lastPageNumber="9" pageId="6" pageNumber="7">
<subSubSection id="F9FE653CFFFA234CFF5EFDBDFAAB6FC4" box="[151,474,619,646]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="B15B36B7FFFA234CFF5EFDBDFAAB6FC4" blockId="6.[151,474,619,678]" box="[151,474,619,646]" pageId="6" pageNumber="7">
<heading id="EA1381DBFFFA234CFF5EFDBDFAAB6FC4" bold="true" box="[151,474,619,646]" fontSize="11" level="1" pageId="6" pageNumber="7" reason="1">
<emphasis id="8390EAA5FFFA234CFF5EFDBDFAAB6FC4" bold="true" box="[151,474,619,646]" pageId="6" pageNumber="7">
<taxonomicName id="76E44D34FFFA234CFF5EFDBDFAE16FC7" ID-CoL="4G6F7" authority="Gosliner, 2011" authorityName="Gosliner" authorityYear="2011" box="[151,400,620,646]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="6" pageNumber="7" phylum="Mollusca" rank="species" species="coronata" status="sp. nov.">
<emphasis id="8390EAA5FFFA234CFF5EFDBDFAE16FC7" bold="true" box="[151,400,620,646]" italics="true" pageId="6" pageNumber="7">Philinopsis coronata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="98A357DEFFFA234CFE51FDBAFAAB6FC4" box="[408,474,619,645]" pageId="6" pageNumber="7" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F9FE653CFFFA234CFF5EFD5FFAFD6FE7" box="[151,396,654,678]" pageId="6" pageNumber="7" type="description">
<paragraph id="B15B36B7FFFA234CFF5EFD5FFAFD6FE7" blockId="6.[151,474,619,678]" box="[151,396,654,678]" pageId="6" pageNumber="7">
(
<figureCitation id="29DF2A32FFFA234CFF57FD5FFA6C6FE7" box="[158,285,654,678]" captionStart="FIGURE 1" captionStartId="2.[151,250,1808,1831]" captionTargetBox="[165,1409,365,1781]" captionTargetId="figure-217@2.[165,1409,365,1781]" captionTargetPageId="2" captionText="FIGURE 1. Living animals A. Philinopsis falciphallus n. sp., paratype, CASIZ 177678, Mabini, Philippines, arrow indicates location of posterior lobe. B. Philinopsis falciphallus n. sp., CASIZ 177678, Mabini, Philippines, ventral view showing markings on foot. C. Philinopsis coronata n. sp., holotype, CASIZ 182887, Mabini, Philippines. D. Philinopsis ctenophoraphaga n. sp. CASIZ 096245, Mabini, Philippines, E. Chelidonura mandroroa n. sp., holotype, CASIZ 173595, Radama Islands, Madagascar F. Chelidonura alisonae n. sp., CASIZ 174177. G. Odontoglaja mosaica, paratype, CASIZ 173419, Radama Islands, Madagascar. H. Odontoglaja mosaica, paratype, CASIZ 173415, Radama Islands, Madagascar. Photos by T. M. Gosliner." figureDoi="http://doi.org/10.5281/zenodo.5291318" httpUri="https://zenodo.org/record/5291318/files/figure.png" pageId="6" pageNumber="7">Figures 1C</figureCitation>
,
<figureCitation id="29DF2A32FFFA234CFEE3FD5FFA3D6FE7" box="[298,332,654,678]" captionStart="FIGURE 2" captionStartId="3.[151,250,1261,1284]" captionTargetBox="[316,1271,193,1240]" captionTargetId="figure-381@3.[316,1271,193,1240]" captionTargetPageId="3" captionText="FIGURE 2. Shells. A. Philinopsis falciphallus. n. sp., paratype, CASIZ 177678, Mabini, Philippines. B. Philinopsis coronata n. sp., holotype, CASIZ 182887, Mabini, Philippines. C. Chelidonura mandroroa n. sp., paratype, CASIZ 182002, Radama Islands, Madagascar D. Chelidonura alisonae n. sp., paratype, CASIZ 174146, Easter Island." figureDoi="http://doi.org/10.5281/zenodo.5291320" httpUri="https://zenodo.org/record/5291320/files/figure.png" pageId="6" pageNumber="7">2B</figureCitation>
,
<figureCitation id="29DF2A32FFFA234CFE91FD5FFA196FE7" box="[344,360,654,678]" captionStart="FIGURE 5" captionStartId="7.[151,250,1558,1581]" captionTargetBox="[214,1374,201,1533]" captionTargetId="figure-238@7.[206,1381,193,1537]" captionTargetPageId="7" captionText="FIGURE 5. Internal anatomy, Philinopsis coronata n. sp., holotype, CASIZ 182887 A. Digestive and central nervous system anatomy, bb buccal bulb, c cerebral ganglion, cr crop, gg genital ganglion, pe pedal ganglion, pl pleural ganglion, sg salivary gland, sp supraintestinal ganglion, su subintestinal ganglion, v visceral ganglion. B. Posterior reproductive organs, al albumen gland, am ampulla, bc bursa copulatrix, ga gential atrium, me membrane gland, mu1 mucous gland large, mu2 mucous gland small, ot ovotestis, rs receptaculum seminis. C., D. Penis, as anterior penial spine, p penial papilla, pr prostate, ps posterior penial spines." figureDoi="http://doi.org/10.5281/zenodo.5291330" httpUri="https://zenodo.org/record/5291330/files/figure.png" pageId="6" pageNumber="7">5</figureCitation>
,
<figureCitation id="29DF2A32FFFA234CFEBDFD5FFAF26FE7" box="[372,387,654,678]" captionStart="FIGURE 6" captionStartId="8.[151,250,1370,1393]" captionTargetBox="[151,1436,373,1349]" captionTargetId="figure-471@8.[151,1436,373,1350]" captionTargetPageId="8" captionText="FIGURE 6. Philinopsis coronata n. sp., holotype, CASIZ 182887, scanning electron micrographs of penis A. Penial sac showing location of as anterior spines, pp penial papilla, ps posterior spines B. Penial papilla and posterior spines C. Penial papilla showing ring of tubercles on apex. D. Detail of anterior spines. E. Detail of posterior spines." figureDoi="http://doi.org/10.5281/zenodo.5291334" httpUri="https://zenodo.org/record/5291334/files/figure.png" pageId="6" pageNumber="7">6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="F9FE653CFFFA234CFF5EFD07FF496E77" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="B15B36B7FFFA234CFF5EFD07FF496E53" blockId="6.[151,1437,726,2010]" pageId="6" pageNumber="7">
<emphasis id="8390EAA5FFFA234CFF5EFD07FAF06FAE" bold="true" box="[151,385,726,751]" inLineHeading="true" pageId="6" pageNumber="7" reason="1">Material examined.</emphasis>
<materialsCitation id="018C3CEAFFFA234CFE40FD07FF496E53" collectingDate="2010-05-23" collectionCode="CASIZ" collectorName="P. Paleracio." country="Philippines" county="Luzon" elevation="12" latitude="-13.686025" location="Mainit Bubbles" longLatPrecision="1" longitude="120.895164" municipality="Mabini" pageId="6" pageNumber="7" specimenCode="CASIZ 1822887" specimenCount="1" stateProvince="Batangas" typeStatus="holotype">
<typeStatus id="6E5F8815FFFA234CFE40FD07FA846FAF" box="[393,501,726,750]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
:
<specimenCode id="E1429ECCFFFA234CFDC8FD07F9CE6FAF" box="[513,703,726,750]" collectionCode="CASIZ" pageId="6" pageNumber="7">CASIZ 1822887</specimenCode>
, dissected,
<elevation id="3AC9D184FFFA234CFC8BFD07F8B56FAF" box="[834,964,726,750]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="7" unit="m" value="12.0">
<quantity id="761C9B52FFFA234CFC8BFD07F80D6FAF" box="[834,892,726,750]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="6" pageNumber="7" unit="m" value="12.0">12 m</quantity>
depth
</elevation>
,
<location id="B43B606CFFFA234CFC18FD07FFF26FAF" LSID="urn:lsid:plazi:treatment:394D87A1FFFA2343FF5EFDBDF9216D92:B43B606CFFFA234CFC18FD07FFF26FAF" box="[977,1155,726,750]" country="Philippines" county="Luzon" latitude="-13.686025" longLatPrecision="1" longitude="120.895164" municipality="Mabini" name="Mainit Bubbles" pageId="6" pageNumber="7" stateProvince="Batangas">Mainit Bubbles</location>
,
<collectingMunicipality id="513FACCDFFFA234CFB46FD07FF926FAF" box="[1167,1251,726,750]" pageId="6" pageNumber="7">Mabini</collectingMunicipality>
,
<collectingRegion id="7320F855FFFA234CFB26FD06FBB96E53" country="Philippines" name="Batangas" pageId="6" pageNumber="7">Batangas Province</collectingRegion>
,
<collectingCounty id="583A4E3BFFFA234CFF1DFD2AFA6F6E53" box="[212,286,763,786]" pageId="6" pageNumber="7">Luzon</collectingCounty>
,
<collectingCountry id="C9F37627FFFA234CFEE0FD2BFADB6E53" box="[297,426,762,786]" name="Philippines" pageId="6" pageNumber="7">Philippines</collectingCountry>
,
<geoCoordinate id="D4D05070FFFA234CFE7CFD2BF9386E53" box="[437,585,762,786]" degrees="13.686025" direction="south" orientation="latitude" pageId="6" pageNumber="7" precision="1" value="-13.686025">13.686025°S</geoCoordinate>
,
<geoCoordinate id="D4D05070FFFA234CFD9CFD2BF98E6E53" box="[597,767,762,786]" degrees="120.895167" direction="east" orientation="longitude" pageId="6" pageNumber="7" precision="1" value="120.895164">120.895167° E</geoCoordinate>
,
<date id="C55A1077FFFA234CFCC2FD2BF8D26E53" box="[779,931,762,787]" pageId="6" pageNumber="7" value="2010-05-23">
<collectingDate id="D51EE99FFFFA234CFCC2FD2BF8D26E53" box="[779,931,762,787]" pageId="6" pageNumber="7" value="2010-05-23">23 May 2010</collectingDate>
</date>
,
<collectorName id="1C115361FFFA234CFC67FD2AFF496E53" box="[942,1080,762,786]" pageId="6" pageNumber="7">P. Paleracio.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="B15B36B7FFFA234CFF0EFCCFFF496E77" blockId="6.[151,1437,726,2010]" box="[199,1080,798,823]" pageId="6" pageNumber="7">
<emphasis id="8390EAA5FFFA234CFF0EFCCFF9756E76" bold="true" box="[199,516,798,823]" inLineHeading="true" pageId="6" pageNumber="7" reason="1">Geographical distribution.</emphasis>
Known only from the
<collectingCountry id="C9F37627FFFA234CFCCDFCCFF8F36E77" box="[772,898,798,822]" name="Philippines" pageId="6" pageNumber="7">Philippines</collectingCountry>
(present study).
</paragraph>
</subSubSection>
<subSubSection id="F9FE653CFFFA234CFF0EFC92F8036E87" pageId="6" pageNumber="7" type="etymology">
<paragraph id="B15B36B7FFFA234CFF0EFC92F8FE6E3F" blockId="6.[151,1437,726,2010]" pageId="6" pageNumber="7">
<emphasis id="8390EAA5FFFA234CFF0EFC92FA3C6E1A" bold="true" box="[199,333,835,859]" inLineHeading="true" pageId="6" pageNumber="7" reason="1">Etymology.</emphasis>
The name “coronata” comes from the Latin
<emphasis id="8390EAA5FFFA234CFC92FC92F8D86E1B" box="[859,937,835,858]" italics="true" pageId="6" pageNumber="7">corona</emphasis>
, meaning crown. This refers to the ring of rounded tubercles at the apex of the penis, which resemble a crown.
</paragraph>
<paragraph id="B15B36B7FFFA234CFF0EFC5AF8036E87" blockId="6.[151,1437,726,2010]" pageId="6" pageNumber="7">
<emphasis id="8390EAA5FFFA234CFF0EFC5AFAF46EE2" bold="true" box="[199,389,906,931]" inLineHeading="true" pageId="6" pageNumber="7" reason="1">Natural history.</emphasis>
This species is found in the same habitat and at the same locality as the preceding species on coral rubble in
<quantity id="761C9B52FFFA234CFE88FC7FFA0B6E87" box="[321,378,942,966]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="7" unit="m" value="15.0">15 m</quantity>
depth. Little else is known about its biology.
</paragraph>
</subSubSection>
<subSubSection id="F9FE653CFFFA2342FF0EFC03F94D6C7E" lastPageId="8" lastPageNumber="9" pageId="6" pageNumber="7" type="description">
<paragraph id="B15B36B7FFFA234CFF0EFC03FF8C6813" blockId="6.[151,1437,726,2010]" pageId="6" pageNumber="7">
<emphasis id="8390EAA5FFFA234CFF0EFC03FA296EAA" bold="true" box="[199,344,978,1003]" inLineHeading="true" pageId="6" pageNumber="7" reason="1">Description.</emphasis>
<emphasis id="8390EAA5FFFA234CFEA8FC02F9286EAA" box="[353,601,979,1003]" italics="true" pageId="6" pageNumber="7">External morphology.</emphasis>
The living specimen was
<quantity id="761C9B52FFFA234CFC41FC03F8A96EAB" box="[904,984,978,1002]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="6" pageNumber="7" unit="mm" value="28.0">28 mm</quantity>
in length and
<quantity id="761C9B52FFFA234CFB48FC02FFB56EAB" box="[1153,1220,979,1002]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="6" pageNumber="7" unit="mm" value="7.0">7 mm</quantity>
wide. The general body color of the living animal (
<figureCitation id="29DF2A32FFFA234CFDCEFC27F92D694F" box="[519,604,1014,1038]" captionStart="FIGURE 1" captionStartId="2.[151,250,1808,1831]" captionTargetBox="[165,1409,365,1781]" captionTargetId="figure-217@2.[165,1409,365,1781]" captionTargetPageId="2" captionText="FIGURE 1. Living animals A. Philinopsis falciphallus n. sp., paratype, CASIZ 177678, Mabini, Philippines, arrow indicates location of posterior lobe. B. Philinopsis falciphallus n. sp., CASIZ 177678, Mabini, Philippines, ventral view showing markings on foot. C. Philinopsis coronata n. sp., holotype, CASIZ 182887, Mabini, Philippines. D. Philinopsis ctenophoraphaga n. sp. CASIZ 096245, Mabini, Philippines, E. Chelidonura mandroroa n. sp., holotype, CASIZ 173595, Radama Islands, Madagascar F. Chelidonura alisonae n. sp., CASIZ 174177. G. Odontoglaja mosaica, paratype, CASIZ 173419, Radama Islands, Madagascar. H. Odontoglaja mosaica, paratype, CASIZ 173415, Radama Islands, Madagascar. Photos by T. M. Gosliner." figureDoi="http://doi.org/10.5281/zenodo.5291318" httpUri="https://zenodo.org/record/5291318/files/figure.png" pageId="6" pageNumber="7">Fig. 1C</figureCitation>
) is whitish with a “v”-shaped maroon patch present on the head. Maroon spots are present along the parapodial margins. The entire dorsal and lateral surfaces of the body are ornamented with scattered, irregular yellow spots and a few whitish patches. The ventral surface of the single animal is pale pink with a series of large yellowish and maroon spots on the foot. Living animals are elongate and wide. The anterior end of the cephalic shield is indented but blunt and quadrangular. The cephalic shield is roughly rectangular and terminates posteriorly with an elongate, rounded papilla. The posterior shield is slightly rounded anteriorly and terminates in an elongate conical posterior protrusion that is well-elevated from the base of the shield. The two lateral posterior lobes of the posterior shield are elongate and simply rounded. The left posterior lobe is longer than the right one. The parapodia are very short, leaving most of the cephalic and posterior shields visible. The gill is simply plicate consisting of 13 primary folds and is situated on the right posterior side of the animal.
</paragraph>
<paragraph id="B15B36B7FFFA234CFF0EFA8FF8F368A3" blockId="6.[151,1437,726,2010]" pageId="6" pageNumber="7">
<emphasis id="8390EAA5FFFA234CFF0EFA8FFB8E6837" box="[199,255,1374,1398]" italics="true" pageId="6" pageNumber="7">Shell</emphasis>
(
<figureCitation id="29DF2A32FFFA234CFEC6FA8FFA196837" box="[271,360,1374,1398]" captionStart="FIGURE 2" captionStartId="3.[151,250,1261,1284]" captionTargetBox="[316,1271,193,1240]" captionTargetId="figure-381@3.[316,1271,193,1240]" captionTargetPageId="3" captionText="FIGURE 2. Shells. A. Philinopsis falciphallus. n. sp., paratype, CASIZ 177678, Mabini, Philippines. B. Philinopsis coronata n. sp., holotype, CASIZ 182887, Mabini, Philippines. C. Chelidonura mandroroa n. sp., paratype, CASIZ 182002, Radama Islands, Madagascar D. Chelidonura alisonae n. sp., paratype, CASIZ 174146, Easter Island." figureDoi="http://doi.org/10.5281/zenodo.5291320" httpUri="https://zenodo.org/record/5291320/files/figure.png" pageId="6" pageNumber="7">Fig. 2B</figureCitation>
). The shell is relatively thickly calcified and consists of a narrow band that occupies the posterior extreme of the animal. There is a thin membranous periostracum that is slightly more extensive anteriorly than the calcified portion. The area at the base of the shell near the protoconch is more thickly calcified than the rest of the shell and a posterior lobe is present on the left side of the shell.
</paragraph>
<paragraph id="B15B36B7FFFA234CFF0EFA3EFFF06BFB" blockId="6.[151,1437,726,2010]" pageId="6" pageNumber="7">
<emphasis id="8390EAA5FFFA234CFF0EFA3EFAF26B47" box="[199,387,1519,1542]" italics="true" pageId="6" pageNumber="7">Digestive system</emphasis>
(
<figureCitation id="29DF2A32FFFA234CFE5AFA3FFA9A6B47" box="[403,491,1518,1542]" captionStart="FIGURE 5" captionStartId="7.[151,250,1558,1581]" captionTargetBox="[214,1374,201,1533]" captionTargetId="figure-238@7.[206,1381,193,1537]" captionTargetPageId="7" captionText="FIGURE 5. Internal anatomy, Philinopsis coronata n. sp., holotype, CASIZ 182887 A. Digestive and central nervous system anatomy, bb buccal bulb, c cerebral ganglion, cr crop, gg genital ganglion, pe pedal ganglion, pl pleural ganglion, sg salivary gland, sp supraintestinal ganglion, su subintestinal ganglion, v visceral ganglion. B. Posterior reproductive organs, al albumen gland, am ampulla, bc bursa copulatrix, ga gential atrium, me membrane gland, mu1 mucous gland large, mu2 mucous gland small, ot ovotestis, rs receptaculum seminis. C., D. Penis, as anterior penial spine, p penial papilla, pr prostate, ps posterior penial spines." figureDoi="http://doi.org/10.5281/zenodo.5291330" httpUri="https://zenodo.org/record/5291330/files/figure.png" pageId="6" pageNumber="7">Fig. 5A</figureCitation>
). The buccal mass is large, highly muscularized and slightly elongate posteriorly. It occupies the entire length of the cephalic shield. The buccal bulb lacks any vestige of a radula. There is a large ventral oral gland and the small dorsal oral glands were indistinct. At the posterior end of the buccal mass, near the junction with the crop, is a pair of elongate salivary glands. The crop is large and saccate, wider than the buccal bulb. The crop narrows posteriorly and enters the digestive gland. The intestine emerges from the right side of the digestive gland and terminates near the posterior end of the body near the base of the gill.
</paragraph>
<paragraph id="B15B36B7FFFA234CFF0EF917FBA96A9B" blockId="6.[151,1437,726,2010]" pageId="6" pageNumber="7">
<emphasis id="8390EAA5FFFA234CFF0EF917FAA46B9F" box="[199,469,1734,1758]" italics="true" pageId="6" pageNumber="7">Central nervous system</emphasis>
(
<figureCitation id="29DF2A32FFFA234CFE21F917F9366B9F" box="[488,583,1734,1758]" captionStart="FIGURE 5" captionStartId="7.[151,250,1558,1581]" captionTargetBox="[214,1374,201,1533]" captionTargetId="figure-238@7.[206,1381,193,1537]" captionTargetPageId="7" captionText="FIGURE 5. Internal anatomy, Philinopsis coronata n. sp., holotype, CASIZ 182887 A. Digestive and central nervous system anatomy, bb buccal bulb, c cerebral ganglion, cr crop, gg genital ganglion, pe pedal ganglion, pl pleural ganglion, sg salivary gland, sp supraintestinal ganglion, su subintestinal ganglion, v visceral ganglion. B. Posterior reproductive organs, al albumen gland, am ampulla, bc bursa copulatrix, ga gential atrium, me membrane gland, mu1 mucous gland large, mu2 mucous gland small, ot ovotestis, rs receptaculum seminis. C., D. Penis, as anterior penial spine, p penial papilla, pr prostate, ps posterior penial spines." figureDoi="http://doi.org/10.5281/zenodo.5291330" httpUri="https://zenodo.org/record/5291330/files/figure.png" pageId="6" pageNumber="7">Fig. 5A</figureCitation>
): The circumesophageal nerve ring consists of paired cerebral, pedal and pleural ganglia and a single supraintestinal ganglion on the right side. The cerebral and pedal commissures are both elongate with well-separated respective ganglia. Immediately adjacent and posterior to the right pleural ganglion is the supraintestinal ganglion. From its posterior end is the right branch of the visceral loop and the osphradial nerve. The two lateral branches of the visceral loop join posteriorly at the posterior ganglia. The left visceral loop enters the subintestinal ganglion, while the right lateral nerve enters the visceral ganglion. The visceral ganglion is larger than the subintestinal ganglion. From the visceral ganglion is the genital nerve, which has a distinct genital ganglion.
</paragraph>
<caption id="E59B663FFFFB234DFF5EF9C7FA506B87" ID-DOI="http://doi.org/10.5281/zenodo.5291330" ID-Zenodo-Dep="5291330" httpUri="https://zenodo.org/record/5291330/files/figure.png" pageId="7" pageNumber="8" startId="7.[151,250,1558,1581]" targetBox="[214,1374,201,1533]" targetPageId="7">
<paragraph id="B15B36B7FFFB234DFF5EF9C7FA506B87" blockId="7.[151,1436,1558,1734]" pageId="7" pageNumber="8">
<emphasis id="8390EAA5FFFB234DFF5EF9C7FA626B6D" bold="true" box="[151,275,1558,1581]" pageId="7" pageNumber="8">FIGURE 5.</emphasis>
Internal anatomy,
<taxonomicName id="76E44D34FFFB234DFE1AF9C6F9D76B6D" authority="Gosliner, 2011" authorityName="Gosliner" authorityYear="2011" box="[467,678,1559,1581]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="7" pageNumber="8" phylum="Mollusca" rank="species" species="coronata" status="sp. nov.">
<emphasis id="8390EAA5FFFB234DFE1AF9C6F9D76B6D" box="[467,678,1559,1581]" italics="true" pageId="7" pageNumber="8">Philinopsis coronata</emphasis>
</taxonomicName>
<emphasis id="8390EAA5FFFB234DFD67F9C6F9996B6D" bold="true" box="[686,744,1559,1580]" pageId="7" pageNumber="8">
<taxonomicNameLabel id="98A357DEFFFB234DFD67F9C6F9996B6D" box="[686,744,1559,1580]" pageId="7" pageNumber="8" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, CASIZ 182887
<emphasis id="8390EAA5FFFB234DFC36F9C7FF666B6C" bold="true" box="[1023,1047,1558,1581]" pageId="7" pageNumber="8">A.</emphasis>
Digestive and central nervous system anatomy, bb buccal bulb, c cerebral ganglion, cr crop, gg genital ganglion, pe pedal ganglion, pl pleural ganglion, sg salivary gland, sp supraintestinal ganglion, su subintestinal ganglion, v visceral ganglion.
<emphasis id="8390EAA5FFFB234DFC1DF985F89A6B28" bold="true" box="[980,1003,1620,1641]" pageId="7" pageNumber="8">B.</emphasis>
Posterior reproductive organs, al albumen gland, am ampulla, bc bursa copulatrix, ga gential atrium, me membrane gland, mu1 mucous gland large, mu2 mucous gland small, ot ovotestis, rs receptaculum seminis.
<emphasis id="8390EAA5FFFB234DFDA0F940F9D86BE7" bold="true" box="[617,681,1681,1704]" pageId="7" pageNumber="8">C., D.</emphasis>
Penis, as anterior penial spine, p penial papilla, pr prostate, ps posterior penial spines.
</paragraph>
</caption>
<paragraph id="B15B36B7FFFB2342FF0FF922F94D6C7E" blockId="7.[151,1436,1778,2018]" lastBlockId="8.[151,1436,151,319]" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<emphasis id="8390EAA5FFFB234DFF0FF922FADC6A4B" box="[198,429,1779,1803]" italics="true" pageId="7" pageNumber="8">Reproductive system</emphasis>
(
<figureCitation id="29DF2A32FFFB234DFE75F923F9456A4B" box="[444,564,1778,1802]" captionStart="FIGURE 5" captionStartId="7.[151,250,1558,1581]" captionTargetBox="[214,1374,201,1533]" captionTargetId="figure-238@7.[206,1381,193,1537]" captionTargetPageId="7" captionText="FIGURE 5. Internal anatomy, Philinopsis coronata n. sp., holotype, CASIZ 182887 A. Digestive and central nervous system anatomy, bb buccal bulb, c cerebral ganglion, cr crop, gg genital ganglion, pe pedal ganglion, pl pleural ganglion, sg salivary gland, sp supraintestinal ganglion, su subintestinal ganglion, v visceral ganglion. B. Posterior reproductive organs, al albumen gland, am ampulla, bc bursa copulatrix, ga gential atrium, me membrane gland, mu1 mucous gland large, mu2 mucous gland small, ot ovotestis, rs receptaculum seminis. C., D. Penis, as anterior penial spine, p penial papilla, pr prostate, ps posterior penial spines." figureDoi="http://doi.org/10.5281/zenodo.5291330" httpUri="https://zenodo.org/record/5291330/files/figure.png" pageId="7" pageNumber="8">Fig. 5B, C</figureCitation>
,
<figureCitation id="29DF2A32FFFB234DFDF6F923F93E6A4B" box="[575,591,1778,1802]" captionStart="FIGURE 6" captionStartId="8.[151,250,1370,1393]" captionTargetBox="[151,1436,373,1349]" captionTargetId="figure-471@8.[151,1436,373,1350]" captionTargetPageId="8" captionText="FIGURE 6. Philinopsis coronata n. sp., holotype, CASIZ 182887, scanning electron micrographs of penis A. Penial sac showing location of as anterior spines, pp penial papilla, ps posterior spines B. Penial papilla and posterior spines C. Penial papilla showing ring of tubercles on apex. D. Detail of anterior spines. E. Detail of posterior spines." figureDoi="http://doi.org/10.5281/zenodo.5291334" httpUri="https://zenodo.org/record/5291334/files/figure.png" pageId="7" pageNumber="8">6</figureCitation>
). The arrangement of reproductive organs is monaulic (
<figureCitation id="29DF2A32FFFB234DFB76F923FE676A4B" box="[1215,1302,1778,1802]" captionStart="FIGURE 5" captionStartId="7.[151,250,1558,1581]" captionTargetBox="[214,1374,201,1533]" captionTargetId="figure-238@7.[206,1381,193,1537]" captionTargetPageId="7" captionText="FIGURE 5. Internal anatomy, Philinopsis coronata n. sp., holotype, CASIZ 182887 A. Digestive and central nervous system anatomy, bb buccal bulb, c cerebral ganglion, cr crop, gg genital ganglion, pe pedal ganglion, pl pleural ganglion, sg salivary gland, sp supraintestinal ganglion, su subintestinal ganglion, v visceral ganglion. B. Posterior reproductive organs, al albumen gland, am ampulla, bc bursa copulatrix, ga gential atrium, me membrane gland, mu1 mucous gland large, mu2 mucous gland small, ot ovotestis, rs receptaculum seminis. C., D. Penis, as anterior penial spine, p penial papilla, pr prostate, ps posterior penial spines." figureDoi="http://doi.org/10.5281/zenodo.5291330" httpUri="https://zenodo.org/record/5291330/files/figure.png" pageId="7" pageNumber="8">Fig. 5A</figureCitation>
) with a single branch to the albumen and membrane glands. From the large ovotestis, which is intermingled with the digestive gland, emerges the convoluted ampulla. The ampulla narrows into the hermaphroditic duct, curves around the receptaculum seminis and has a single branch to the short, coiled albumen and membrane glands. The larger mucous gland is bilobed with a massive primary lobe and smaller secondary one. The ampulla then joins the duct of the receptaculum seminis and continues to the genital atrium where it joins the duct of the bursa copulatrix. The bursa is large and spherical. Its duct is narrow where it joins the bursa and widens until its widest portion at the highly muscularized genital atrium. From the genital atrium, the open, ciliated sperm groove leads to the cephalic penis. The penis (
<figureCitation id="29DF2A32FFF42342FE94FF6DFAC56D95" box="[349,436,188,212]" captionStart="FIGURE 5" captionStartId="7.[151,250,1558,1581]" captionTargetBox="[214,1374,201,1533]" captionTargetId="figure-238@7.[206,1381,193,1537]" captionTargetPageId="7" captionText="FIGURE 5. Internal anatomy, Philinopsis coronata n. sp., holotype, CASIZ 182887 A. Digestive and central nervous system anatomy, bb buccal bulb, c cerebral ganglion, cr crop, gg genital ganglion, pe pedal ganglion, pl pleural ganglion, sg salivary gland, sp supraintestinal ganglion, su subintestinal ganglion, v visceral ganglion. B. Posterior reproductive organs, al albumen gland, am ampulla, bc bursa copulatrix, ga gential atrium, me membrane gland, mu1 mucous gland large, mu2 mucous gland small, ot ovotestis, rs receptaculum seminis. C., D. Penis, as anterior penial spine, p penial papilla, pr prostate, ps posterior penial spines." figureDoi="http://doi.org/10.5281/zenodo.5291330" httpUri="https://zenodo.org/record/5291330/files/figure.png" pageId="8" pageNumber="9">Fig. 5C</figureCitation>
,
<figureCitation id="29DF2A32FFF42342FE09FF6DFABE6D95" box="[448,463,188,212]" captionStart="FIGURE 6" captionStartId="8.[151,250,1370,1393]" captionTargetBox="[151,1436,373,1349]" captionTargetId="figure-471@8.[151,1436,373,1350]" captionTargetPageId="8" captionText="FIGURE 6. Philinopsis coronata n. sp., holotype, CASIZ 182887, scanning electron micrographs of penis A. Penial sac showing location of as anterior spines, pp penial papilla, ps posterior spines B. Penial papilla and posterior spines C. Penial papilla showing ring of tubercles on apex. D. Detail of anterior spines. E. Detail of posterior spines." figureDoi="http://doi.org/10.5281/zenodo.5291334" httpUri="https://zenodo.org/record/5291334/files/figure.png" pageId="8" pageNumber="9">6</figureCitation>
) consists of a penial sac and a lobate prostate gland that is joined to the penial sac by a narrow duct. Within the penial sac is a large, rigid conical penial papilla (
<figureCitation id="29DF2A32FFF42342FC06FF0EFF326DB6" box="[975,1091,223,247]" captionStart="FIGURE 6" captionStartId="8.[151,250,1370,1393]" captionTargetBox="[151,1436,373,1349]" captionTargetId="figure-471@8.[151,1436,373,1350]" captionTargetPageId="8" captionText="FIGURE 6. Philinopsis coronata n. sp., holotype, CASIZ 182887, scanning electron micrographs of penis A. Penial sac showing location of as anterior spines, pp penial papilla, ps posterior spines B. Penial papilla and posterior spines C. Penial papilla showing ring of tubercles on apex. D. Detail of anterior spines. E. Detail of posterior spines." figureDoi="http://doi.org/10.5281/zenodo.5291334" httpUri="https://zenodo.org/record/5291334/files/figure.png" pageId="8" pageNumber="9">Fig. 6A,B</figureCitation>
). Anterior to the penial papilla and posterior to it are two areas of spines on the folds of the penial sac (
<figureCitation id="29DF2A32FFF42342FC78FED5FF796C5D" box="[945,1032,260,284]" captionStart="FIGURE 5" captionStartId="7.[151,250,1558,1581]" captionTargetBox="[214,1374,201,1533]" captionTargetId="figure-238@7.[206,1381,193,1537]" captionTargetPageId="7" captionText="FIGURE 5. Internal anatomy, Philinopsis coronata n. sp., holotype, CASIZ 182887 A. Digestive and central nervous system anatomy, bb buccal bulb, c cerebral ganglion, cr crop, gg genital ganglion, pe pedal ganglion, pl pleural ganglion, sg salivary gland, sp supraintestinal ganglion, su subintestinal ganglion, v visceral ganglion. B. Posterior reproductive organs, al albumen gland, am ampulla, bc bursa copulatrix, ga gential atrium, me membrane gland, mu1 mucous gland large, mu2 mucous gland small, ot ovotestis, rs receptaculum seminis. C., D. Penis, as anterior penial spine, p penial papilla, pr prostate, ps posterior penial spines." figureDoi="http://doi.org/10.5281/zenodo.5291330" httpUri="https://zenodo.org/record/5291330/files/figure.png" pageId="8" pageNumber="9">Fig. 5C</figureCitation>
,
<figureCitation id="29DF2A32FFF42342FBDAFED5FF496C5D" box="[1043,1080,260,284]" captionStart="FIGURE 6" captionStartId="8.[151,250,1370,1393]" captionTargetBox="[151,1436,373,1349]" captionTargetId="figure-471@8.[151,1436,373,1350]" captionTargetPageId="8" captionText="FIGURE 6. Philinopsis coronata n. sp., holotype, CASIZ 182887, scanning electron micrographs of penis A. Penial sac showing location of as anterior spines, pp penial papilla, ps posterior spines B. Penial papilla and posterior spines C. Penial papilla showing ring of tubercles on apex. D. Detail of anterior spines. E. Detail of posterior spines." figureDoi="http://doi.org/10.5281/zenodo.5291334" httpUri="https://zenodo.org/record/5291334/files/figure.png" pageId="8" pageNumber="9">6A</figureCitation>
). The apex of the papilla is covered with a ring of rounded tubercles.
</paragraph>
</subSubSection>
<caption id="E59B663FFFF42342FF5EFA8BFF4E68EF" ID-DOI="http://doi.org/10.5281/zenodo.5291334" ID-Zenodo-Dep="5291334" httpUri="https://zenodo.org/record/5291334/files/figure.png" pageId="8" pageNumber="9" startId="8.[151,250,1370,1393]" targetBox="[151,1436,373,1349]" targetPageId="8">
<paragraph id="B15B36B7FFF42342FF5EFA8BFF4E68EF" blockId="8.[151,1436,1370,1454]" pageId="8" pageNumber="9">
<emphasis id="8390EAA5FFF42342FF5EFA8BFA636830" bold="true" box="[151,274,1370,1393]" pageId="8" pageNumber="9">FIGURE 6.</emphasis>
<taxonomicName id="76E44D34FFF42342FED1FA8AFA9B6830" authority="Gosliner, 2011" authorityName="Gosliner" authorityYear="2011" box="[280,490,1371,1393]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="coronata" status="sp. nov.">
<emphasis id="8390EAA5FFF42342FED1FA8AFA9B6830" box="[280,490,1371,1393]" italics="true" pageId="8" pageNumber="9">Philinopsis coronata</emphasis>
</taxonomicName>
<emphasis id="8390EAA5FFF42342FE39FA8AF9586831" bold="true" box="[496,553,1371,1392]" pageId="8" pageNumber="9">
<taxonomicNameLabel id="98A357DEFFF42342FE39FA8AF9586831" box="[496,553,1371,1392]" pageId="8" pageNumber="9" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, holotype, CASIZ 182887, scanning electron micrographs of penis
<emphasis id="8390EAA5FFF42342FB1DFA8BFF9C6830" bold="true" box="[1236,1261,1370,1393]" pageId="8" pageNumber="9">A.</emphasis>
Penial sac showing location of as anterior spines, pp penial papilla, ps posterior spines
<emphasis id="8390EAA5FFF42342FCA7FAABF8F468CE" bold="true" box="[878,901,1402,1423]" pageId="8" pageNumber="9">B.</emphasis>
Penial papilla and posterior spines
<emphasis id="8390EAA5FFF42342FB38FAA8FE7868D1" bold="true" box="[1265,1289,1401,1424]" pageId="8" pageNumber="9">C.</emphasis>
Penial papilla showing ring of tubercles on apex.
<emphasis id="8390EAA5FFF42342FE32FA48F96568EF" bold="true" box="[507,532,1433,1454]" pageId="8" pageNumber="9">D.</emphasis>
Detail of anterior spines.
<emphasis id="8390EAA5FFF42342FCD2FA48F84368EF" bold="true" box="[795,818,1433,1454]" pageId="8" pageNumber="9">E.</emphasis>
Detail of posterior spines.
</paragraph>
</caption>
<subSubSection id="F9FE653CFFF42343FF0CFA0AF9216D92" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="discussion">
<paragraph id="B15B36B7FFF42343FF0CFA0AF9216D92" blockId="8.[151,1436,1499,2027]" lastBlockId="9.[151,1436,151,212]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
<emphasis id="8390EAA5FFF42342FF0CFA0AFA4868B2" bold="true" box="[197,313,1499,1523]" inLineHeading="true" pageId="8" pageNumber="9" reason="1">Remarks.</emphasis>
<taxonomicName id="76E44D34FFF42342FE8CFA0AF94568B2" authorityName="Gosliner" authorityYear="2011" box="[325,564,1499,1523]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="coronata">
<emphasis id="8390EAA5FFF42342FE8CFA0AF94568B2" box="[325,564,1499,1523]" italics="true" pageId="8" pageNumber="9">Philinopsis coronata</emphasis>
</taxonomicName>
is very similar in its external and internal appearance to
<taxonomicName id="76E44D34FFF42342FB26FA0DFEE468B2" authorityName="Gosliner" authorityYear="2011" box="[1263,1429,1499,1523]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="falciphallus">
<emphasis id="8390EAA5FFF42342FB26FA0DFEE468B2" box="[1263,1429,1499,1523]" italics="true" pageId="8" pageNumber="9">P. falciphallus</emphasis>
</taxonomicName>
, described above. The external anatomy and color pattern are very similar. In
<taxonomicName id="76E44D34FFF42342FC3DF9D1FF056B56" authorityName="Gosliner" authorityYear="2011" box="[1012,1140,1536,1559]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="coronata">
<emphasis id="8390EAA5FFF42342FC3DF9D1FF056B56" box="[1012,1140,1536,1559]" italics="true" pageId="8" pageNumber="9">P. coronata</emphasis>
</taxonomicName>
, there is more yellow pigment with less maroon and the posterior lobe of the posterior shield is more elongate than in
<taxonomicName id="76E44D34FFF42342FB78F9F5FE206B7A" authorityName="Gosliner" authorityYear="2011" box="[1201,1361,1571,1595]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="falciphallus">
<emphasis id="8390EAA5FFF42342FB78F9F5FE206B7A" box="[1201,1361,1571,1595]" italics="true" pageId="8" pageNumber="9">P. falciphallus</emphasis>
</taxonomicName>
. Also,
<taxonomicName id="76E44D34FFF42342FF5EF999FA696B1E" authorityName="Gosliner" authorityYear="2011" box="[151,280,1608,1631]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="coronata">
<emphasis id="8390EAA5FFF42342FF5EF999FA696B1E" box="[151,280,1608,1631]" italics="true" pageId="8" pageNumber="9">P. coronata</emphasis>
</taxonomicName>
has more elongate posterior extension of the foot than does
<taxonomicName id="76E44D34FFF42342FC03F999FF1A6B1E" authorityName="Gosliner" authorityYear="2011" box="[970,1131,1607,1631]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="falciphallus">
<emphasis id="8390EAA5FFF42342FC03F999FF1A6B1E" box="[970,1131,1607,1631]" italics="true" pageId="8" pageNumber="9">P. falciphallus</emphasis>
</taxonomicName>
. Both species have yellow and maroon on the foot, but in
<taxonomicName id="76E44D34FFF42342FE38F9BDF9016BC2" authorityName="Gosliner" authorityYear="2011" box="[497,624,1644,1667]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="coronata">
<emphasis id="8390EAA5FFF42342FE38F9BDF9016BC2" box="[497,624,1644,1667]" italics="true" pageId="8" pageNumber="9">P. coronata</emphasis>
</taxonomicName>
there is no maroon line, but rather a series of maroon spots. The shell of
<taxonomicName id="76E44D34FFF42342FF5EF941FA686BE6" authorityName="Gosliner" authorityYear="2011" box="[151,281,1680,1703]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="coronata">
<emphasis id="8390EAA5FFF42342FF5EF941FA686BE6" box="[151,281,1680,1703]" italics="true" pageId="8" pageNumber="9">P. coronata</emphasis>
</taxonomicName>
has a more elongate posterior wing on the right side than is found in
<taxonomicName id="76E44D34FFF42342FB82F941FF9C6BE6" authorityName="Gosliner" authorityYear="2011" box="[1099,1261,1679,1703]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="falciphallus">
<emphasis id="8390EAA5FFF42342FB82F941FF9C6BE6" box="[1099,1261,1679,1703]" italics="true" pageId="8" pageNumber="9">P. falciphallus</emphasis>
</taxonomicName>
. Internally, the digestive and central nervous systems of the two species exhibit no apparent differences. The arrangement of the posterior reproductive organs of the two is also quite similar, but the genital atrium of
<taxonomicName id="76E44D34FFF42342FBB3F909FF8A6BAE" authorityName="Gosliner" authorityYear="2011" box="[1146,1275,1752,1775]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="coronata">
<emphasis id="8390EAA5FFF42342FBB3F909FF8A6BAE" box="[1146,1275,1752,1775]" italics="true" pageId="8" pageNumber="9">P. coronata</emphasis>
</taxonomicName>
appears to be more highly muscularized. The most significant difference between these two species is found in the structure of the penis. Both species have a simple penis and prostate connected by a narrow duct and both are the only species of
<taxonomicName id="76E44D34FFF42342FF7CF892FA426A1A" authorityName=", Rudman" authorityYear="1972" box="[181,307,1859,1883]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="genus">
<emphasis id="8390EAA5FFF42342FF7CF892FA426A1A" box="[181,307,1859,1883]" italics="true" pageId="8" pageNumber="9">Philinopsis</emphasis>
</taxonomicName>
with an armed penial papilla. Despite these similarities, the armature of the penis is markedly different in the two species. In
<taxonomicName id="76E44D34FFF42342FE7DF8B9F9256A3E" authorityName="Gosliner" authorityYear="2011" box="[436,596,1895,1919]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="falciphallus">
<emphasis id="8390EAA5FFF42342FE7DF8B9F9256A3E" box="[436,596,1895,1919]" italics="true" pageId="8" pageNumber="9">P. falciphallus</emphasis>
</taxonomicName>
, there is a large chitinous spine and a series of smaller spines on the penial papilla. In
<taxonomicName id="76E44D34FFF42342FED8F85DFAE06AE2" authorityName="Gosliner" authorityYear="2011" box="[273,401,1932,1955]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="coronata">
<emphasis id="8390EAA5FFF42342FED8F85DFAE06AE2" box="[273,401,1932,1955]" italics="true" pageId="8" pageNumber="9">P. coronata</emphasis>
</taxonomicName>
there are two regions of spines, one anterior to the penial papilla and the other one situated posteriorly to the papilla. The apex of the papilla of
<taxonomicName id="76E44D34FFF42342FD10F861F8296A86" authorityName="Gosliner" authorityYear="2011" box="[729,856,1968,1991]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="coronata">
<emphasis id="8390EAA5FFF42342FD10F861F8296A86" box="[729,856,1968,1991]" italics="true" pageId="8" pageNumber="9">P. coronata</emphasis>
</taxonomicName>
has a ring of rounded tubercles that is not present in
<taxonomicName id="76E44D34FFF42342FF5EF805FA476AAA" authorityName="Gosliner" authorityYear="2011" box="[151,310,2003,2027]" class="Gastropoda" family="Aglajidae" genus="Philinopsis" kingdom="Animalia" order="Cephalaspidea" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="falciphallus">
<emphasis id="8390EAA5FFF42342FF5EF805FA476AAA" box="[151,310,2003,2027]" italics="true" pageId="8" pageNumber="9">P. falciphallus</emphasis>
</taxonomicName>
. The specimens are similar in size and maturity so it is likely that the observed penial differences are due to differences in maturity or preservation. These significant differences in penial morphology confirm that these specimens represent distinct taxa.
</paragraph>
</subSubSection>
</treatment>
</document>
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