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<document id="F81BB96B2E3C7290C8CFDA3AC67BB881" ID-ISSN="1175­5334" ID-ZooBank="6B088E9E-EAD4-4B8B-938B-178ADACF4CC3" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629809046025" checkinUser="felipe" docAuthor="Harzhauser, Mathias &amp; Landau, Bernard" docDate="2012" docId="387287FCC33CFFE8FF0DE5EDE3D2FBA7" docLanguage="en" docName="zt03472p071.pdf" docOrigin="Zootaxa 3472" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Ovilia Jousseaume 1887" docType="treatment" docVersion="3" lastPageNumber="34" masterDocId="C44BFF84C31DFFC9FF9AE57AE767FF93" masterDocTitle="A revision of the Neogene Cancellariid Gastropods of the Paratethys Sea 3472" masterLastPageNumber="71" masterPageNumber="1" pageNumber="34" updateTime="1699115141119" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="C5739EE77D407BD1A1C0543F60B4C1B8">A revision of the Neogene Cancellariid Gastropods of the Paratethys Sea 3472</mods:title>
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<mods:namePart id="9EF5D11D2CEB8CBAFF67E7958BD2BA5C">Harzhauser, Mathias</mods:namePart>
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<mods:namePart id="84576CC5C44EA078890A6049AD005FD0">Landau, Bernard</mods:namePart>
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<mods:date id="8A48D16F63DBBCBABC62919061B54838">2012</mods:date>
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<mods:number id="B62178696E5B58925E7B189667A01241">2012-09-07</mods:number>
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<treatment id="387287FCC33CFFE8FF0DE5EDE3D2FBA7" ID-DOI="http://doi.org/10.5281/zenodo.5257388" ID-Zenodo-Dep="5257388" LSID="urn:lsid:plazi:treatment:387287FCC33CFFE8FF0DE5EDE3D2FBA7" httpUri="http://treatment.plazi.org/id/387287FCC33CFFE8FF0DE5EDE3D2FBA7" lastPageNumber="34" pageId="33" pageNumber="34">
<subSubSection id="F8C16561C33CFFE8FF0DE5EDE67EFF22" box="[151,281,151,177]" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="B06436EAC33CFFE8FF0DE5EDE67EFF22" blockId="33.[151,296,151,211]" box="[151,281,151,177]" pageId="33" pageNumber="34">
<heading id="EB2C8186C33CFFE8FF0DE5EDE67EFF22" bold="true" box="[151,281,151,177]" fontSize="11" level="1" pageId="33" pageNumber="34" reason="1">
<emphasis id="82AFEAF8C33CFFE8FF0DE5EDE67EFF22" bold="true" box="[151,281,151,177]" pageId="33" pageNumber="34">
<taxonomicName id="77DB4D69C33CFFE8FF0DE5EDE787FF22" ID-CoL="84PPQ" authorityName="Jousseaume" authorityYear="1887" box="[151,224,151,177]" class="Gastropoda" family="Cancellariidae" genus="Ovilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
<emphasis id="82AFEAF8C33CFFE8FF0DE5EDE787FF22" bold="true" box="[151,224,151,177]" italics="true" pageId="33" pageNumber="34">Ovilia</emphasis>
</taxonomicName>
? sp.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F8C16561C33CFFE8FF0DE5C0E64FFF40" box="[151,296,186,211]" pageId="33" pageNumber="34" type="description">
<paragraph id="B06436EAC33CFFE8FF0DE5C0E64FFF40" blockId="33.[151,296,151,211]" box="[151,296,186,211]" pageId="33" pageNumber="34">
<figureCitation id="28E02A6FC33CFFE8FF0DE5C0E78AFF41" box="[151,237,186,210]" captionStart="FIGURE 7" captionStartId="29.[151,250,1787,1810]" captionTargetBox="[241,1327,185,1759]" captionTargetId="figure-176@29.[241,1327,185,1759]" captionTargetPageId="29" captionText="FIGURE 7. A1–A3. Merica succineiformis (Boettger, 1906), Coşteiu de Sus (Romania), SMF XII 2264 a, holotype. B1–B3. Ovilia breitenbergerorum nov. sp., holotype, Mörtersdorf (Austria), NHMW 2012/0012/0001. C1–C3. Ovilia excassidea (Sacco, 1894), Gainfarn (Austria), NHMW 1846/0037/0288, holotype. D1–D3. Ovilia tauroaspira (Sacco, 1894), Mörtersdorf (Austria). E1–E2. Ovilia? sp., Mörtersdorf (Austria). F1–F3. Perplicaria mioquadrata (Sacco, 1894), Lăpugiu de Sus (Romania), NHMW 2009z0098/0002, lectotype. G1–G3. Perplicaria mioquadrata (Sacco, 1894), Coşteiu de Sus (Romania), NHMW 2009z0098/0003. H1–H3. Petitina inermis (Pusch, 1837), Enzesfeld (Austria), NHMW 2009z0098/0012. I1–I2. Petitina inermis (Pusch, 1837), Enzesfeld (Austria), NHMW 2009z0098/0011. J1–J3. Petitina inermis (Pusch, 1837), Grund (Austria), NHMW 1855/0045/0733." pageId="33" pageNumber="34">Figs 7E</figureCitation>
<figureCitation id="28E02A6FC33CFFE8FF77E5C1E67CFF41" box="[237,283,187,211]" captionStart="FIGURE 1" captionStartId="5.[151,250,1829,1852]" captionTargetBox="[207,1376,180,1786]" captionTargetId="figure-143@5.[207,1376,180,1786]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, Korneub. B: Korneuburg Basin, E–Sopr B: Eisenstadt–Sopron Basin). Note that this map shows only an Early Langhian situation; at that time, the Korneuburg Basin is already dry and the NAFB is restricted to a small eastern embayment; for further time slices see Rögl (1998) and Popov et al. (2004)." pageId="33" pageNumber="34">1–E</figureCitation>
<figureCitation id="28E02A6FC33CFFE8FE80E5C1E64FFF40" box="[282,296,187,211]" captionStart="FIGURE 2" captionStartId="6.[151,250,1264,1287]" captionTargetBox="[151,1436,193,1243]" captionTargetId="figure-378@6.[151,1436,193,1243]" captionTargetPageId="6" captionText="FIGURE 2. Miocene chronostratigraphy, magnetostratigraphy and biostratigraphy (PF: Planktonic Foraminifera, CN: Calcareous Nannoplankton) after Hilgen et al. (2009) with regional Paratethys stages after Piller et al. (2007). The figure is modified from a chart produced with the Time Scale Creator program, provided by the International Commission on Stratigraphy (available at: http://stratigraphy.org/column). The number of cancellariid species per stage is indicated in the grey bars." pageId="33" pageNumber="34">2</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F8C16561C33CFFE8FF0DE479E41FFED3" pageId="33" pageNumber="34" type="materials_examined">
<paragraph id="B06436EAC33CFFE8FF0DE479E41FFED3" blockId="33.[151,1437,259,1076]" pageId="33" pageNumber="34">
<emphasis id="82AFEAF8C33CFFE8FF0DE479E6EFFE8F" bold="true" box="[151,392,259,284]" inLineHeading="true" pageId="33" pageNumber="34" reason="1">Illustrated material.</emphasis>
<figureCitation id="28E02A6FC33CFFE8FE14E479E680FE88" box="[398,487,259,283]" captionStart="FIGURE 7" captionStartId="29.[151,250,1787,1810]" captionTargetBox="[241,1327,185,1759]" captionTargetId="figure-176@29.[241,1327,185,1759]" captionTargetPageId="29" captionText="FIGURE 7. A1–A3. Merica succineiformis (Boettger, 1906), Coşteiu de Sus (Romania), SMF XII 2264 a, holotype. B1–B3. Ovilia breitenbergerorum nov. sp., holotype, Mörtersdorf (Austria), NHMW 2012/0012/0001. C1–C3. Ovilia excassidea (Sacco, 1894), Gainfarn (Austria), NHMW 1846/0037/0288, holotype. D1–D3. Ovilia tauroaspira (Sacco, 1894), Mörtersdorf (Austria). E1–E2. Ovilia? sp., Mörtersdorf (Austria). F1–F3. Perplicaria mioquadrata (Sacco, 1894), Lăpugiu de Sus (Romania), NHMW 2009z0098/0002, lectotype. G1–G3. Perplicaria mioquadrata (Sacco, 1894), Coşteiu de Sus (Romania), NHMW 2009z0098/0003. H1–H3. Petitina inermis (Pusch, 1837), Enzesfeld (Austria), NHMW 2009z0098/0012. I1–I2. Petitina inermis (Pusch, 1837), Enzesfeld (Austria), NHMW 2009z0098/0011. J1–J3. Petitina inermis (Pusch, 1837), Grund (Austria), NHMW 1855/0045/0733." pageId="33" pageNumber="34">Figs 7H</figureCitation>
<figureCitation id="28E02A6FC33CFFE8FE7DE47EE570FE88" box="[487,535,260,284]" captionStart="FIGURE 1" captionStartId="5.[151,250,1829,1852]" captionTargetBox="[207,1376,180,1786]" captionTargetId="figure-143@5.[207,1376,180,1786]" captionTargetPageId="5" captionText="FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea (for detailed maps with the localities mentioned in the text see Kroh (2005). The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea (modified from Harzhauser et al. 2007) showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, Korneub. B: Korneuburg Basin, E–Sopr B: Eisenstadt–Sopron Basin). Note that this map shows only an Early Langhian situation; at that time, the Korneuburg Basin is already dry and the NAFB is restricted to a small eastern embayment; for further time slices see Rögl (1998) and Popov et al. (2004)." pageId="33" pageNumber="34">1–H</figureCitation>
<figureCitation id="28E02A6FC33CFFE8FD8DE47EE54FFE8F" box="[535,552,260,284]" captionStart="FIGURE 2" captionStartId="6.[151,250,1264,1287]" captionTargetBox="[151,1436,193,1243]" captionTargetId="figure-378@6.[151,1436,193,1243]" captionTargetPageId="6" captionText="FIGURE 2. Miocene chronostratigraphy, magnetostratigraphy and biostratigraphy (PF: Planktonic Foraminifera, CN: Calcareous Nannoplankton) after Hilgen et al. (2009) with regional Paratethys stages after Piller et al. (2007). The figure is modified from a chart produced with the Time Scale Creator program, provided by the International Commission on Stratigraphy (available at: http://stratigraphy.org/column). The number of cancellariid species per stage is indicated in the grey bars." pageId="33" pageNumber="34">2</figureCitation>
: Mörtersdorf (
<collectingCountry id="C8CC767AC33CFFE8FD53E479E446FE88" box="[713,801,259,283]" name="Austria" pageId="33" pageNumber="34">Austria</collectingCountry>
), height:
<quantity id="77239B0FC33CFFE8FC12E479E4BFFE88" box="[904,984,259,283]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="33" pageNumber="34" unit="mm" value="30.0">30 mm</quantity>
, width:
<quantity id="77239B0FC33CFFE8FBABE479E3E5FE88" box="[1073,1154,259,283]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="33" pageNumber="34" unit="mm" value="26.0">26 mm</quantity>
, private collection Anton and Thomas Breitenberger (Bad Vöslau and Pottenstein,
<collectingCountry id="C8CC767AC33CFFE8FC8FE452E400FED3" box="[789,871,296,320]" name="Austria" pageId="33" pageNumber="34">Austria</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="F8C16561C33CFFE8FF59E436E502FD8B" pageId="33" pageNumber="34" type="description">
<paragraph id="B06436EAC33CFFE8FF59E436E502FD8B" blockId="33.[151,1437,259,1076]" pageId="33" pageNumber="34">
<emphasis id="82AFEAF8C33CFFE8FF59E436E633FEF6" bold="true" box="[195,340,332,357]" inLineHeading="true" pageId="33" pageNumber="34" reason="1">Description.</emphasis>
Globular shell with low, gradate spire and large, convex last whorl. A shoulder appears already on early whorls and separates a flat sutural shelf from the flanks. Sculpture consisting of 13 low fold-like axial ribs overrun by about ten flattened spiral cords, with broad, horizontally elongate nodes developed at the intersections. These are most prominent along the shoulder. Up to three wavy secondary spiral threads are intercalated between the adapical five primary cords, whilst only one (or no) threads are intercalated between the cords on the base. Aperture and base obscured by sediment.
</paragraph>
</subSubSection>
<subSubSection id="F8C16561C33CFFE8FF59E75EE66FFC7F" pageId="33" pageNumber="34" type="discussion">
<paragraph id="B06436EAC33CFFE8FF59E75EE4B6FD5F" blockId="33.[151,1437,259,1076]" pageId="33" pageNumber="34">
<emphasis id="82AFEAF8C33CFFE8FF59E75EE62FFDAE" bold="true" box="[195,328,548,573]" inLineHeading="true" pageId="33" pageNumber="34" reason="1">Discussion.</emphasis>
Only a single specimen is available and the aperture is unknown. Therefore, even the generic assignment remains doubtful. Although some specimens of the Early Miocene
<taxonomicName id="77DB4D69C33CFFE8FB96E732E3A1FDF3" baseAuthorityName="Basterot" baseAuthorityYear="1825" box="[1036,1222,584,608]" class="Gastropoda" family="Cancellariidae" genus="Ovilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="doliolaris">
<emphasis id="82AFEAF8C33CFFE8FB96E732E3A1FDF3" box="[1036,1222,584,608]" italics="true" pageId="33" pageNumber="34">Ovilia doliolaris</emphasis>
</taxonomicName>
have tendencies to develop rugose spiral ribs, as evident from the illustration in
<bibRefCitation id="D44A4B1BC33CFFE8FCC2E717E36CFD17" author="Basterot, B." box="[856,1035,620,644]" pageId="33" pageNumber="34" pagination="1 - 100" refId="ref56180" refString="Basterot, B. (1825) Description geologique du bassin tertiaire du SO de la France. Memoire de la Societe de Histoire Naturelle Paris, 11, 1 - 100." type="journal article" year="1825">Basterot (1825)</bibRefCitation>
, none displays such a conspicuous sculpture. The weak suture of the Austrian shell allows a clear separation from the
<taxonomicName id="77DB4D69C33CFFE8FBFCE7EAE396FD3B" baseAuthorityName="Basterot" baseAuthorityYear="1825" box="[1126,1265,656,680]" class="Gastropoda" family="Cancellariidae" genus="Ovilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="doliolaris">
<emphasis id="82AFEAF8C33CFFE8FBFCE7EAE396FD3B" box="[1126,1265,656,680]" italics="true" pageId="33" pageNumber="34">O doliolaris</emphasis>
</taxonomicName>
group and the newly described
<taxonomicName id="77DB4D69C33CFFE8FECEE7CEE52EFD5F" authorityName="Harzhauser &amp; Landau" authorityYear="2012" box="[340,585,692,716]" class="Gastropoda" family="Cancellariidae" genus="Ovilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="breitenbergerorum">
<emphasis id="82AFEAF8C33CFFE8FECEE7CEE52EFD5F" box="[340,585,692,716]" italics="true" pageId="33" pageNumber="34">O. breitenbergerorum</emphasis>
</taxonomicName>
, which occurs at the same locality.
</paragraph>
<paragraph id="B06436EAC33CFFE8FF59E7A2E66FFC7F" blockId="33.[151,1437,259,1076]" pageId="33" pageNumber="34">
A second group of species, which we provisionally include in the genus
<taxonomicName id="77DB4D69C33CFFE8FB94E7A2E332FD63" authorityName="Jousseaume" authorityYear="1887" box="[1038,1109,728,752]" class="Gastropoda" family="Cancellariidae" genus="Ovilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="genus">
<emphasis id="82AFEAF8C33CFFE8FB94E7A2E332FD63" box="[1038,1109,728,752]" italics="true" pageId="33" pageNumber="34">Ovilia</emphasis>
</taxonomicName>
, have axial sculpture, albeit weak, much less conspicuous spiral cords and a less deeply canaliculate suture. This group is characterised by
<taxonomicName id="77DB4D69C33CFFE8FA1AE786E6DFFCAB" authority="(Sacco, 1894)" authorityName="Harzhauser &amp; Landau" authorityYear="2012" baseAuthorityName="Sacco" baseAuthorityYear="1894" class="Gastropoda" family="Cancellariidae" genus="Ovilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="excassidea">
<emphasis id="82AFEAF8C33CFFE8FA1AE786E674FCAB" italics="true" pageId="33" pageNumber="34">O. excassidea</emphasis>
(
<bibRefCitation id="D44A4B1BC33CFFE8FEB8E65AE6D7FCAB" author="Sacco, F." box="[290,432,800,824]" pageId="33" pageNumber="34" pagination="68 - 70" refId="ref61955" refString="Sacco, F. (1894) I molluschi dei terreni terziarii del Piemonte e della Liguria. Parte 16: Fam. Cancellariidae H. e A. Adams 1853. Bollettino dei Musei di Zoologia ed Anatomia comparata della Reale Universita di Torino, 9 (171), 68 - 70 (published consecutively with Part 15) (April 27 1894)." type="journal article" year="1894">Sacco, 1894</bibRefCitation>
)
</taxonomicName>
in the Miocene and
<taxonomicName id="77DB4D69C33CFFE8FD38E65AE484FCAB" authority="(Brocchi, 1814)" baseAuthorityName="Brocchi" baseAuthorityYear="1814" box="[674,995,800,824]" class="Gastropoda" family="Cancellariidae" genus="Ovilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="cassidea">
<emphasis id="82AFEAF8C33CFFE8FD38E65AE440FCAB" box="[674,807,800,824]" italics="true" pageId="33" pageNumber="34">O. cassidea</emphasis>
(
<bibRefCitation id="D44A4B1BC33CFFE8FCACE65AE4BCFCAB" author="Brocchi, G." box="[822,987,800,824]" pageId="33" pageNumber="34" pagination="1 - 2" refId="ref56616" refString="Brocchi, G. (1814) Conchiologia fossile subapennina, con osservazioni geologiche sugli Apennini e sul suolo adiacente. Stamperia Reale, Milano, 1 - 2, 712 pp." type="book chapter" year="1814">Brocchi, 1814</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="77DB4D69C33CFFE8FB80E65AE219FCAB" authority="Landau, 1984" authorityName="Landau" authorityYear="1984" box="[1050,1406,800,824]" class="Gastropoda" family="Cancellariidae" genus="Ovilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="andaluciensis">
<emphasis id="82AFEAF8C33CFFE8FB80E65AE3BDFCAB" box="[1050,1242,800,824]" italics="true" pageId="33" pageNumber="34">O. andaluciensis</emphasis>
Landau,
<quantity id="77239B0FC33CFFE8FADFE65AE219FCAB" box="[1349,1406,800,824]" metricMagnitude="1" metricUnit="m" metricValue="5.03936" pageId="33" pageNumber="34" unit="in" value="1984.0">1984</quantity>
</taxonomicName>
in the Pliocene. The Austrian shell resembles
<taxonomicName id="77DB4D69C33CFFE8FD04E63EE426FCCF" authorityName="Harzhauser &amp; Landau" authorityYear="2012" baseAuthorityName="Sacco" baseAuthorityYear="1894" box="[670,833,836,860]" class="Gastropoda" family="Cancellariidae" genus="Ovilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="excassidea">
<emphasis id="82AFEAF8C33CFFE8FD04E63EE426FCCF" box="[670,833,836,860]" italics="true" pageId="33" pageNumber="34">O. excassidea</emphasis>
</taxonomicName>
in outline and suture
<typeStatus id="6F608848C33CFFE8FBD4E63FE318FCCE" box="[1102,1151,837,861]" pageId="33" pageNumber="34">type</typeStatus>
and might represent an ancestral form of this Middle Miocene species. The sculpture of both species is also comparable. Differences are the blunt nodes and densely spaced spiral ribs of the Early Miocene Austrian shell, as opposed to the much narrower spiral cords of
<taxonomicName id="77DB4D69C33CFFE8FE0BE6CAE555FC5B" authorityName="Harzhauser &amp; Landau" authorityYear="2012" baseAuthorityName="Sacco" baseAuthorityYear="1894" box="[401,562,944,968]" class="Gastropoda" family="Cancellariidae" genus="Ovilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="excassidea">
<emphasis id="82AFEAF8C33CFFE8FE0BE6CAE555FC5B" box="[401,562,944,968]" italics="true" pageId="33" pageNumber="34">O. excassidea</emphasis>
</taxonomicName>
, and the poorly defined axial ribs, whilst
<taxonomicName id="77DB4D69C33CFFE8FB80E6CAE3DDFC5B" authorityName="Harzhauser &amp; Landau" authorityYear="2012" baseAuthorityName="Sacco" baseAuthorityYear="1894" box="[1050,1210,944,968]" class="Gastropoda" family="Cancellariidae" genus="Ovilia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neogastropoda" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="excassidea">
<emphasis id="82AFEAF8C33CFFE8FB80E6CAE3DDFC5B" box="[1050,1210,944,968]" italics="true" pageId="33" pageNumber="34">O. excassidea</emphasis>
</taxonomicName>
has a distinct axial sculpture.
</paragraph>
</subSubSection>
<subSubSection id="F8C16561C33CFFE8FF59E68DE3D2FBA7" pageId="33" pageNumber="34" type="distribution">
<paragraph id="B06436EAC33CFFE8FF59E68DE375FB83" blockId="33.[151,1437,259,1076]" box="[195,1042,1015,1040]" pageId="33" pageNumber="34">
<emphasis id="82AFEAF8C33CFFE8FF59E68DE63BFB83" bold="true" box="[195,348,1015,1040]" inLineHeading="true" pageId="33" pageNumber="34" reason="1">Distribution.</emphasis>
A rare species, which is only known from the Paratethys Sea.
</paragraph>
<paragraph id="B06436EAC33CFFE8FF59E166E3D2FBA7" blockId="33.[151,1437,259,1076]" box="[195,1205,1052,1076]" pageId="33" pageNumber="34">
<emphasis id="82AFEAF8C33CFFE8FF59E166E65BFBA7" box="[195,316,1052,1076]" italics="true" pageId="33" pageNumber="34">Paratethys</emphasis>
—Eggenburgian (Early Burdigalian): Bohemian Massif (
<collectingCountry id="C8CC767AC33CFFE8FC20E166E375FBA7" box="[954,1042,1052,1076]" name="Austria" pageId="33" pageNumber="34">Austria</collectingCountry>
: Mörtersdorf).
</paragraph>
</subSubSection>
</treatment>
</document>
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