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<document id="ED392264076914DF172546DAE20A61D5" ID-DOI="10.11646/zootaxa.3750.2.1" ID-GBIF-Dataset="bcdcc503-9dad-4bef-834b-cdc1a07485dd" ID-ISSN="1175-5326" ID-Zenodo-Dep="248024" ID-ZooBank="F01C254C-7CCA-4669-8C30-BF61180BD675" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460709113477" checkinUser="plazi" docAuthor="Tauber, Catherine A., Sosa, Francisco, Albuquerque, Gilberto S. &amp; Tauber, Maurice J." docDate="2013" docId="0129879FF614FFD70BDAFE391912D083" docLanguage="en" docName="zt03750p129.pdf" docOrigin="Zootaxa 3750 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Leucochrysa (Leucochrysa) nigrilabris Banks 1915" docType="treatment" docVersion="9" lastPageNumber="117" masterDocId="FD10FFE7F616FFC70B4DFFE21D14D273" masterDocTitle="Adults and larvae of two Leucochrysa (Leucochrysa) species (Neuroptera: Chrysopidae): descriptions, biological notes, and relationships" masterLastPageNumber="129" masterPageNumber="101" pageNumber="103" updateTime="1698357713548" updateUser="plazi">
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<mods:title id="B8930397BE5058BC298F247717501826">Adults and larvae of two Leucochrysa (Leucochrysa) species (Neuroptera: Chrysopidae): descriptions, biological notes, and relationships</mods:title>
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<taxonomicName id="4E804D0AF614FFC50BDAFE391E0BD385" ID-CoL="6PTNH" authority="Banks, 1915" authorityName="Banks" authorityYear="1915" box="[151,799,475,502]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="nigrilabris" subGenus="Leucochrysa">
<emphasis id="BBF4EA9BF614FFC50BDAFE391E0BD385" bold="true" box="[151,799,475,502]" pageId="2" pageNumber="103">
<emphasis id="BBF4EA9BF614FFC50BDAFE391C26D386" bold="true" box="[151,306,475,501]" italics="true" pageId="2" pageNumber="103">Leucochrysa</emphasis>
(
<emphasis id="BBF4EA9BF614FFC50A0EFE391CCAD386" bold="true" box="[323,478,475,501]" italics="true" pageId="2" pageNumber="103">Leucochrysa</emphasis>
)
<emphasis id="BBF4EA9BF614FFC50AA3FE391F79D386" bold="true" box="[494,621,475,501]" italics="true" pageId="2" pageNumber="103">nigrilabris</emphasis>
(Banks, 1915)
</emphasis>
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<figureCitation id="11BB2A0CF614FFC50BDAFE1C1C1DD064" box="[151,265,510,535]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-10="FIGURE 11" captionStart-11="FIGURE 12" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStart-9="FIGURE 10" captionStartId-0="3.[151,250,1700,1722]" captionStartId-1="4.[151,250,1965,1987]" captionStartId-10="13.[151,250,1017,1039]" captionStartId-11="14.[151,250,1391,1413]" captionStartId-2="5.[151,250,1619,1641]" captionStartId-3="6.[151,250,1654,1676]" captionStartId-4="7.[151,250,1722,1744]" captionStartId-5="8.[151,250,1180,1202]" captionStartId-6="9.[151,250,1392,1414]" captionStartId-7="10.[151,250,1715,1737]" captionStartId-8="12.[151,250,712,734]" captionStartId-9="12.[151,250,1401,1423]" captionTargetBox-0="[162,1436,193,1676]" captionTargetBox-1="[151,1436,337,1893]" captionTargetBox-10="[151,1436,337,995]" captionTargetBox-11="[151,1436,409,1369]" captionTargetBox-2="[151,1436,193,1597]" captionTargetBox-3="[151,1436,193,1632]" captionTargetBox-4="[151,1436,193,1699]" captionTargetBox-5="[151,1436,193,1158]" captionTargetBox-6="[151,1436,409,1370]" captionTargetBox-7="[151,1436,589,1693]" captionTargetBox-8="[151,1436,445,690]" captionTargetBox-9="[151,1436,814,1379]" captionTargetId-0="figure@3.[162,1447,193,1677]" captionTargetId-1="figure@4.[151,1436,337,1944]" captionTargetId-10="figure@13.[151,1436,337,996]" captionTargetId-11="figure@14.[151,1436,409,1370]" captionTargetId-2="figure@5.[151,1436,193,1598]" captionTargetId-3="figure@6.[151,1436,193,1633]" captionTargetId-4="figure@7.[151,1436,193,1700]" captionTargetId-5="figure@8.[151,1436,193,1158]" captionTargetId-6="figure@9.[151,1436,409,1371]" captionTargetId-7="figure@10.[151,1436,589,1694]" captionTargetId-8="figure@12.[151,1436,445,691]" captionTargetId-9="figure@12.[151,1436,814,1379]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-10="13" captionTargetPageId-11="14" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="12" captionTargetPageId-9="12" captionText-0="FIGURE 1. Leucochrysa (L.) nigrilabris external features. A, B. Head, frontal; C. Head, dorsal; D. Body, dorsal E, F. Head and thorax, dorsal [note the variation in the coloration of the mesoprescutum and mesoscutellum]; G. Head, thorax, lateral; H. Head, prothorax, lateral [A, D, State of Yaracuy, Venezuela, female; B, C, F H, State of Rio de Janeiro, Brazil, female; E, State of Aragua, Venezuela, female]." captionText-1="FIGURE 2. Leucochrysa (L.) nigrilabris wings. A, B. Forewing, two specimens to illustrate relatively minor variation; C. Hindwing [A, State of Yaracuy, Venezuela; B, C, State of Aragua, Venezuela, all female]. Scale applies to all three images." captionText-10="FIGURE 11. Leucochrysa (L.) nigrilabris second instar. A. Head, dorsal; B. Head, lateral; C. Thorax, dorsal; D. Abdominal segments 1 to 4, dorsal; E. Abdominal segments 4 10, dorsal; F. Body, lateral [all, State of Rio de Janeiro, Brazil]. Abbreviations: Ax, abdominal segment number; AxRx, setal row number." captionText-11="FIGURE 12. Leucochrysa (L.) nigrilabris third instar. A. Head, dorsal; B. Head, ventral; C. Thorax, dorsal; D. Abdominal segments 1 to 3, dorsal; E. Abdominal segments 3 7, dorsal; F. Abdominal segments 6 - 10, dorsal; G. Head, lateral; H. Body, ventral; I. Body, lateral [all, State of Rio de Janeiro, Brazil]. Abbreviations: Ax, abdominal segment number; AxRx, setal row number." captionText-2="FIGURE 3. Leucochrysa (L.) nigrilabris male abdomen, mature specimen. A. Second and third abdominal segments, lateral [note absence of microtholi from S 2, dense microtholi on S 3]; B. Terminal segments, lateral; C. Terminal segments, dorsal [note large brown spots on T 5, T 6, base of T 7 (arrow)]; D. Terminal segments, dorsal [note dorsally fused T 9 + ectoproct, without suture or distal cleft. The U-shaped darkening is the distal margin of S 8 + 9 below]; E. Callus cerci; F. Fused sternite 8 + 9, ventral, with sclerotized terminal plates extended distally [all, Aragua, Venezuela]. Abbreviations: c. c., callus cerci; d. a., dorsal apodeme of ninth tergite + ectoproct; s. p., sclerotized plate at tip of S 8 + 9; S 2, S 3, second and third sternites; S 8 + 9, fused eighth and ninth sternites (in parentheses, out of focus through dorsal surface); tr, trichobothria; T 5, T 7, T 8, fifth to eighth tergites; T 9 + ect, fused ninth tergite and ectoproct." captionText-3="FIGURE 4. Leucochrysa (L.) nigrilabris male genitalia, mature specimen. A. Gonarcus, posterodorsal (mediuncus removed); B. Gonarcal complex, posterior (view into gonosaccus from behind, through membrane) [note the mediuncal beak and flanges extend forward (away) and downward]; C. Gonarcal complex, dorsal [view into gonosaccus from above, unobscured by membrane, base of mediuncus in focus, mediuncal beak present (extending forward), but out of focus]; D. Gonarcal complex, frontal [note the gonosaccus is below the mediuncal flanges; the gonosetae are within the gonosaccus]; E. Gonarcal complex, lateral; F. Hypandrium internum (comes very reduced or absent) [all, Aragua, Venezuela]. Abbreviations: bk, beak-like tip of mediuncus; fl, mediuncal flange; gc, gonocornu, encased in membrane; g. a., gonarcal apodeme; g. b., gonarcal bridge; g. p., gonarcal process (membrane attachment); gsac, gonosaccus; gse, gonosetae (within gonosaccus); mu, mediuncus (fused rods and beak); mu-b, sclerotized base of mediuncus." captionText-4="FIGURE 5. Leucochrysa (L.) nigrilabris male abdomen, teneral specimen. A. Terminal segments, lateral (arrows mark the large brown spots on T 6, base of T 7); B. Gonarcal complex, frontal; C. Gonarcal complex, lateral [note thorny membrane over gonarcal bridge and gonocornua]; D. Gonarcal complex, lateral (thorny membrane above gonarcus partially removed); E. Tip of sternite 8 + 9, dorsal, with sclerotized terminal plates extended distally, hypandrium internum (attached to small membrane that had been everted beneath the sclerotized plate); F. Sclerotized plate at tip of S 8 + 9, lateral (all, Aragua, Venezuela). Abbreviations: bk, beak-like tip of mediuncus; c, comes; cleft, cleft separating distal margins of T 9 + ect; c. c., callus cerci; d. a., dorsal apodeme of T 9 + ect; fl, mediuncal flange; gc, gonocornu; gs, gonarcus; gsac, gonosaccus with robust gonosetae on bulbous chalazae; g. a., gonarcal apodeme; g. b., gonarcal bridge; h. i., hypandrium internum; mu-b, sclerotized base of mediuncus; mu-d, elongate distal section of mediuncus; s. p., sclerotized plate at tip of S 8 + 9; S 8 + 9, fused eighth and ninth sternites; S 9, ninth sternite (tip); T 7, seventh tergite; T 9 + ect, fused ninth tergite and ectoproct .." captionText-5="FIGURE 6. Leucochrysa (L.) nigrilabris female abdomen. A. Fifth to terminal segments, lateral; B-D. Seventh to terminal segment, lateral [A, B, mature specimens, State of Rio de Janeiro, Brazil; C, D, teneral specimens, State of Aragua, Venezuela]. Abbreviations: g. l., gonapophysis lateralis; sg, subgenitale; sp, spermatheca; S 7, seventh sternite; T 5, T 8, fifth and eighth tergites; T 9 + ect, fused ninth tergite and ectoproct." captionText-6="FIGURE 7. Leucochrysa (L.) nigrilabris female abdominal and genitalic structures. A. Callus cerci; B. Subgenitale, posterior; C. Spermatheca, lateral; D. Genitalia, ventral [A, B, D, State of Rio de Janeiro, Brazil, mature specimen; C, State of Aragua, Venezuela]. Abbreviations: base, sclerotized base of subgenitale; b. c., bursa copulatrix; b. gl., bursal gland; inv, spermathecal invagination; lobe, rounded lobe of subgenitale; sg / b. c., heavy folded membrane [distally (right): dorsum of subgenitale, proximally (left): venter of bursa copulatrix]; sp, spermatheca; sp. d., spermathecal duct; tr, trichobothria; vel, velum." captionText-7="FIGURE 8. Leucochrysa (L.) nigrilabris colleterial complex. A. Colleterial complex, lateral; B. Base of colleterial complex, lateral (pulled posteriorly); C. Transverse sclerite, posterior. [all, State of Rio de Janeiro, Brazil, mature specimen]. Abbreviations: acc. gl., colleterial accessory gland; acc. gl. d., duct of colleterial accessory gland; c. d., common duct (colleterial reservoir and accessory gland to transverse sclerite); col. gl., colleterial gland; col. r., colleterial reservoir; tr. s., transverse sclerite." captionText-8="FIGURE 9. Prothorax of third instar of four Leucochrysa (Leucochrysa) species. A. L. (L.) nigrilabris; B. L. (L.) insularis; C. L. (L.) varia; D. L. (L.) boxi [A, C, D, State of Rio de Janeiro, Brazil; B, North Carolina, United States]." captionText-9="FIGURE 10. Leucochrysa (L.) nigrilabris first instar. A. Head, dorsal; B. Head, ventral; C. Head, lateral; D. Thorax, dorsal; E. Abdominal segments 1 to 5, dorsal; F. Abdominal segments 4 - 10, dorsal; G. Body, lateral; H. Body, ventral [all, State of Rio de Janeiro, Brazil]. Abbreviations: Ax, abdominal segment number; AxRx, setal row number." httpUri-0="https://zenodo.org/record/248025/files/figure.png" httpUri-1="https://zenodo.org/record/248026/files/figure.png" httpUri-10="https://zenodo.org/record/248035/files/figure.png" httpUri-11="https://zenodo.org/record/248036/files/figure.png" httpUri-2="https://zenodo.org/record/248027/files/figure.png" httpUri-3="https://zenodo.org/record/248028/files/figure.png" httpUri-4="https://zenodo.org/record/248029/files/figure.png" httpUri-5="https://zenodo.org/record/248030/files/figure.png" httpUri-6="https://zenodo.org/record/248031/files/figure.png" httpUri-7="https://zenodo.org/record/248032/files/figure.png" httpUri-8="https://zenodo.org/record/248033/files/figure.png" httpUri-9="https://zenodo.org/record/248034/files/figure.png" pageId="2" pageNumber="103">Figs 112</figureCitation>
,
<figureCitation id="11BB2A0CF614FFC50A59FE1D1C24D064" box="[276,304,511,535]" captionStart="FIGURE 19" captionStartId="23.[151,250,1395,1417]" captionTargetBox="[151,1436,319,1361]" captionTargetId="figure@23.[151,1436,319,1362]" captionTargetPageId="23" captionText="FIGURE 19. Species-specific genital characteristics of Leucochrysa (L.) insularis (left column) and L. (L.) nigrilabris (right column). A, B. Gonarcal complex; C, D. Subgenitale [A, C. Dominican Republic; B, D. Aragua, Venezuela]. Abbreviations: bk, beak-like tip of mediuncus; gc, gonocornu; g. a., gonarcal apodeme; g. b., gonarcal bridge; mu, mediuncus (fused rods and beak); sg-bl, bilobed, dorsal process of subgenitale; sg-v, protruding ventral process of subgenitale; sp, spermatheca." httpUri="https://zenodo.org/record/248043/files/figure.png" pageId="2" pageNumber="103">19</figureCitation>
</paragraph>
<paragraph id="893F3689F614FFC50BDAFDA41CA7D008" blockId="2.[151,1436,582,727]" pageId="2" pageNumber="103">
<taxonomicName id="4E804D0AF614FFC50BDAFDA41CD4D02F" authority="Banks" authorityName="Banks" box="[151,448,582,604]" class="Insecta" family="Chrysopidae" genus="Allochrysa" higherTaxonomySource="CoL" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF614FFC50BDAFDA41C63D02F" box="[151,375,582,604]" italics="true" pageId="2" pageNumber="103">Allochrysa nigrilabris</emphasis>
Banks
</taxonomicName>
, [1915] (Banks
<date id="FD3E1049F614FFC5092FFDA41FCCD02F" box="[610,728,582,604]" pageId="2" pageNumber="103" value="1914" valueMax="1915">19141915</date>
: 623) original description: [
<collectingCountry id="F1977619F614FFC508BDFDA4194DD02F" box="[1008,1113,582,604]" name="Colombia" pageId="2" pageNumber="103">Colombia</collectingCountry>
] “St. Antonio,
<collectingCountry id="F1977619F614FFC50FBFFDA4184CD02F" box="[1266,1368,582,604]" name="Colombia" pageId="2" pageNumber="103">Colombia</collectingCountry>
,
<quantity id="4E789B6CF614FFC50E2FFDA41DCBD008" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="103" unit="m" value="1800.0">1,800 m</quantity>
., December (Fassl)”.
</paragraph>
</subSubSection>
<subSubSection id="C19A6502F614FFC50BDAFD661871D0A4" pageId="2" pageNumber="103" type="description">
<paragraph id="893F3689F614FFC50BDAFD661871D0A4" blockId="2.[151,1436,582,727]" pageId="2" pageNumber="103">
<taxonomicName id="4E804D0AF614FFC50BDAFD661CF0D0E9" authority="Banks" authorityName="Banks" box="[151,484,644,666]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF614FFC50BDAFD661C9ED0E9" box="[151,394,644,666]" italics="true" pageId="2" pageNumber="103">Leucochrysa nigrilabris</emphasis>
(Banks)
</taxonomicName>
. Banks (1944: 32) first reference to combination; Penny (1977: 23) species list.
<taxonomicName id="4E804D0AF614FFC50BDAFD401F90D0CB" authority="Banks" authorityName="Banks" box="[151,644,674,696]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="nigrilabris" subGenus="Leucochrysa">
<emphasis id="BBF4EA9BF614FFC50BDAFD401C0CD0CB" box="[151,280,674,696]" italics="true" pageId="2" pageNumber="103">Leucochrysa</emphasis>
(
<emphasis id="BBF4EA9BF614FFC50A64FD401CBED0CB" box="[297,426,674,696]" italics="true" pageId="2" pageNumber="103">Leucochrysa</emphasis>
)
<emphasis id="BBF4EA9BF614FFC50AF1FD401F3CD0CB" box="[444,552,674,696]" italics="true" pageId="2" pageNumber="103">nigrilabris</emphasis>
(Banks)
</taxonomicName>
. Brooks &amp; Barnard (1990: 276) subgeneric designation, species list; Tauber (2004: 1140) possible synonymy with
<taxonomicName id="4E804D0AF614FFC50901FD23193CD0A4" authority="Walker" authorityName="Walker" box="[588,1064,705,727]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="insularis" subGenus="Leucochrysa">
<emphasis id="BBF4EA9BF614FFC50901FD231FD9D0A4" box="[588,717,705,727]" italics="true" pageId="2" pageNumber="103">Leucochrysa</emphasis>
(
<emphasis id="BBF4EA9BF614FFC50991FD231E49D0A4" box="[732,861,705,727]" italics="true" pageId="2" pageNumber="103">Leucochrysa</emphasis>
)
<emphasis id="BBF4EA9BF614FFC50821FD231ED1D0A4" box="[876,965,705,727]" italics="true" pageId="2" pageNumber="103">insularis</emphasis>
(Walker)
</taxonomicName>
; Oswald (2007) catalog listing.
</paragraph>
</subSubSection>
<subSubSection id="C19A6502F614FFC50BDAFCE61FFDD6BE" pageId="2" pageNumber="103" type="materials_examined">
<paragraph id="893F3689F614FFC50BDAFCE61E65D66B" blockId="2.[151,1436,772,2021]" pageId="2" pageNumber="103">
<emphasis id="BBF4EA9BF614FFC50BDAFCE61C5AD16E" bold="true" box="[151,334,772,797]" pageId="2" pageNumber="103">
<typeStatus id="563B882BF614FFC50BDAFCE61DC6D16E" box="[151,210,772,797]" pageId="2" pageNumber="103">Type</typeStatus>
specimen.
</emphasis>
One
<typeStatus id="563B882BF614FFC50AC6FCE71CAAD16E" box="[395,446,773,797]" pageId="2" pageNumber="103">type</typeStatus>
, from San Antonio,
<collectingCountry id="F1977619F614FFC509EEFCE61E00D16E" box="[675,788,772,797]" name="Colombia" pageId="2" pageNumber="103">Colombia</collectingCountry>
, MCZ (male, examined). We assume that the locality “St. Antonio” refers to the city of San Antonio in the Department of Tolima. Banks (
<date id="FD3E1049F614FFC50F0DFCC519D5D133" box="[1088,1217,807,832]" pageId="2" pageNumber="103" value="1914" valueMax="1915">19141915</date>
) did not state how many specimens he used to write his description; it is likely that he had only one. There is only one
<typeStatus id="563B882BF614FFC50E6CFCAF1845D116" box="[1313,1361,845,869]" pageId="2" pageNumber="103">type</typeStatus>
in the MCZ and none in the AMNH, USNM, or the Academy of Natural Sciences of Philadelphia, where Banks occasionally deposited
<typeStatus id="563B882BF614FFC50AECFC771CCBD1DE" box="[417,479,917,941]" pageId="2" pageNumber="103">types</typeStatus>
. Thus, we consider the specimen in the MCZ to be the
<typeStatus id="563B882BF614FFC50F2AFC7619D8D1DE" box="[1127,1228,916,941]" pageId="2" pageNumber="103" type="holotype">holotype</typeStatus>
, by monotypy. Its labels read: (1) “St Antonio /
<collectingCountry id="F1977619F614FFC50AAEFC551F46D1A3" box="[483,594,951,976]" name="Colombia" pageId="2" pageNumber="103">Colombia</collectingCountry>
S. Am” [Banks hand]; (2) “
<quantity id="4E789B6CF614FFC508DBFC5A1EF9D1BC" box="[918,1005,952,976]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="2" pageNumber="103" unit="m" value="1800.0">1800 m</quantity>
/ Dec” [Banks hand]; (3) “Collection / N. Banks”; (4) “
<typeStatus id="563B882BF614FFC50A13FC3E1C8ED187" box="[350,410,988,1012]" pageId="2" pageNumber="103">Type</typeStatus>
” [red, Banks hand]; (5) “
<typeStatus id="563B882BF614FFC5098CFC3E1FEFD187" box="[705,763,988,1012]" pageId="2" pageNumber="103">Type</typeStatus>
/ 12004”; (6) “
<taxonomicName id="4E804D0AF614FFC508EAFC3E1930D186" box="[935,1060,988,1013]" class="Insecta" family="Chrysopidae" genus="Allochrysa" higherTaxonomySource="CoL" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="genus">Allochrysa</taxonomicName>
/
<taxonomicName id="4E804D0AF614FFC50F76FC3E19BAD186" box="[1083,1198,988,1013]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="nigrilabris">nigrilabris</taxonomicName>
/
<typeStatus id="563B882BF614FFC50F8BFC3F19E2D186" box="[1222,1270,989,1013]" pageId="2" pageNumber="103">type</typeStatus>
/ Bks” [white, red border, Banks hand]; (7) “Genitalia / prep P. Adams / 1982”.
</paragraph>
<paragraph id="893F3689F614FFC50B8AFBC61EFAD6F6" blockId="2.[151,1436,772,2021]" pageId="2" pageNumber="103">
The specimen is in fairly good condition; the body (thorax and head) are somewhat flattened, the forewings and one hindwing are missing, and the terminalia are in a vial with glycerine. We took images that are now placed in the MCZ
<typeStatus id="563B882BF614FFC50A6DFB8F1C4CD6F6" box="[288,344,1133,1157]" pageId="2" pageNumber="103">Type</typeStatus>
Database (http://insects.oeb.harvard.edu/MCZ/index.htm).
</paragraph>
<paragraph id="893F3689F614FFC50B8AFB6D1FFDD6BE" blockId="2.[151,1436,772,2021]" pageId="2" pageNumber="103">
<emphasis id="BBF4EA9BF614FFC50B8AFB6D1FF5D6DB" bold="true" box="[199,737,1167,1192]" pageId="2" pageNumber="103">
Known geographical distribution.
<collectingCountry id="F1977619F614FFC5092EFB6D1FC8D6DB" box="[611,732,1167,1192]" name="Colombia" pageId="2" pageNumber="103">Colombia</collectingCountry>
:
</emphasis>
Department of Tolima.
<emphasis id="BBF4EA9BF614FFC508BCFB6D1967D6DB" bold="true" box="[1009,1139,1167,1192]" pageId="2" pageNumber="103">
<collectingCountry id="F1977619F614FFC508BCFB6D197AD6DB" box="[1009,1134,1167,1192]" name="Venezuela" pageId="2" pageNumber="103">Venezuela</collectingCountry>
:
</emphasis>
States of Aragua, Mérida, Portuguesa, Yaracuy.
<emphasis id="BBF4EA9BF614FFC50AC7FB561CC8D6BE" bold="true" box="[394,476,1204,1229]" pageId="2" pageNumber="103">
<collectingCountry id="F1977619F614FFC50AC7FB561CC3D6BE" box="[394,471,1204,1229]" name="Brazil" pageId="2" pageNumber="103">Brazil</collectingCountry>
:
</emphasis>
State of Rio de Janeiro.
</paragraph>
</subSubSection>
<subSubSection id="C19A6502F614FFC50B8AFB3519D1D596" pageId="2" pageNumber="103" type="diagnosis">
<paragraph id="893F3689F614FFC50B8AFB351CCED4D3" blockId="2.[151,1436,772,2021]" pageId="2" pageNumber="103">
<emphasis id="BBF4EA9BF614FFC50B8AFB351CB2D683" bold="true" box="[199,422,1239,1264]" pageId="2" pageNumber="103">Diagnosis (adult).</emphasis>
Externally,
<emphasis id="BBF4EA9BF614FFC5090AFB3B1F4AD683" box="[583,606,1241,1264]" italics="true" pageId="2" pageNumber="103">L.</emphasis>
(
<emphasis id="BBF4EA9BF614FFC50936FB3B1F86D683" box="[635,658,1241,1264]" italics="true" pageId="2" pageNumber="103">L.</emphasis>
)
<taxonomicName id="4E804D0AF614FFC509E2FB351E33D683" box="[687,807,1239,1264]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF614FFC509E2FB351E33D683" box="[687,807,1239,1264]" italics="true" pageId="2" pageNumber="103">nigrilabris</emphasis>
</taxonomicName>
adults are distinguished from many
<taxonomicName id="4E804D0AF614FFC50E40FB351C2DD767" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF614FFC50E40FB351888D683" box="[1293,1436,1239,1264]" italics="true" pageId="2" pageNumber="103">Leucochrysa</emphasis>
(
<emphasis id="BBF4EA9BF614FFC50BEDFB1E1C3BD766" box="[160,303,1276,1301]" italics="true" pageId="2" pageNumber="103">Leucochrysa</emphasis>
)
</taxonomicName>
species by a dark brown to black labrum, white frons, a single, small spot on the forewing (brown suffusion on the membrane around the distal Psm-Pcu crossvein), and all veins mostly green except inner and outer gradates dark brown to black. However, these features are shared with other
<taxonomicName id="4E804D0AF614FFC50F4FFAA6182FD72F" box="[1026,1339,1348,1373]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF614FFC50F4FFAA61985D72E" box="[1026,1169,1348,1373]" italics="true" pageId="2" pageNumber="103">Leucochrysa</emphasis>
(
<emphasis id="BBF4EA9BF614FFC50FEEFAA61826D72E" box="[1187,1330,1348,1373]" italics="true" pageId="2" pageNumber="103">Leucochrysa</emphasis>
)
</taxonomicName>
species, including some
<emphasis id="BBF4EA9BF614FFC50A10FA8B1C60D7F3" box="[349,372,1385,1408]" italics="true" pageId="2" pageNumber="103">L.</emphasis>
(
<emphasis id="BBF4EA9BF614FFC50AC0FA8B1CB0D7F3" box="[397,420,1385,1408]" italics="true" pageId="2" pageNumber="103">L.</emphasis>
) “
<taxonomicName id="4E804D0AF614FFC50A87FA8B1F11D7F3" box="[458,517,1385,1408]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="varia">
<emphasis id="BBF4EA9BF614FFC50A87FA8B1F11D7F3" box="[458,517,1385,1408]" italics="true" pageId="2" pageNumber="103">varia</emphasis>
</taxonomicName>
-like species” and
<emphasis id="BBF4EA9BF614FFC509BDFA8B1E13D7F3" box="[752,775,1385,1408]" italics="true" pageId="2" pageNumber="103">L.</emphasis>
(
<emphasis id="BBF4EA9BF614FFC5086CFA8B1E23D7F3" box="[801,823,1385,1408]" italics="true" pageId="2" pageNumber="103">L.</emphasis>
)
<emphasis id="BBF4EA9BF614FFC5081CFA851EAFD7F3" box="[849,955,1383,1408]" italics="true" pageId="2" pageNumber="103">
<taxonomicName id="4E804D0AF614FFC5081CFA851EA3D7F3" box="[849,951,1383,1408]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="insularis">insularis</taxonomicName>
.
</emphasis>
Thus, a reliable identification requires examination of the genitalia. The males of
<emphasis id="BBF4EA9BF614FFC50934FA6F1F84D7D7" box="[633,656,1421,1444]" italics="true" pageId="2" pageNumber="103">L.</emphasis>
(
<emphasis id="BBF4EA9BF614FFC509EDFA6F1FA2D7D7" box="[672,694,1421,1444]" italics="true" pageId="2" pageNumber="103">L.</emphasis>
)
<taxonomicName id="4E804D0AF614FFC5098AFA6E1E2BD7D6" box="[711,831,1420,1445]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF614FFC5098AFA6E1E2BD7D6" box="[711,831,1420,1445]" italics="true" pageId="2" pageNumber="103">nigrilabris</emphasis>
</taxonomicName>
are distinguished by: (i) dense microtholi on sternites 38; (ii) small genital structures [gonarcal span of
<quantity id="4E789B6CF614FFC50987FA4D1E64D7BB" box="[714,880,1455,1480]" metricMagnitude="-4" metricUnit="m" metricValue="5.25" metricValueMax="6.5" metricValueMin="4.0" pageId="2" pageNumber="103" unit="mm" value="0.525" valueMax="0.65" valueMin="0.4">0.400.65 mm</quantity>
; cf.,
<quantity id="4E789B6CF614FFC508EBFA4D195CD7BB" box="[934,1096,1455,1480]" metricMagnitude="-4" metricUnit="m" metricValue="7.55" metricValueMax="8.4" metricValueMin="6.7" pageId="2" pageNumber="103" unit="mm" value="0.755" valueMax="0.84" valueMin="0.67">0.670.84 mm</quantity>
in
<emphasis id="BBF4EA9BF614FFC50F26FA531996D7BB" box="[1131,1154,1457,1480]" italics="true" pageId="2" pageNumber="103">L.</emphasis>
(
<emphasis id="BBF4EA9BF614FFC50FDFFA5319BDD7BB" box="[1170,1193,1457,1480]" italics="true" pageId="2" pageNumber="103">L.</emphasis>
)
<taxonomicName id="4E804D0AF614FFC50FF5FA5319E7D7BB" box="[1208,1267,1457,1480]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="varia">
<emphasis id="BBF4EA9BF614FFC50FF5FA5319E7D7BB" box="[1208,1267,1457,1480]" italics="true" pageId="2" pageNumber="103">varia</emphasis>
</taxonomicName>
]; (ii) a narrow, elongate mediuncus that is well separated from the gonarcal bridge, and terminates in a long, slender hook and a pair of large, inflated flanges; and (iii) terminus of S8+9 with an attached pair of flat, sclerotized, plate-like projections with toothed mesal margins. Female
<emphasis id="BBF4EA9BF614FFC50984F9FF1FCBD447" box="[713,735,1565,1588]" italics="true" pageId="2" pageNumber="103">L.</emphasis>
(
<emphasis id="BBF4EA9BF614FFC509BEF9FF1E1ED447" box="[755,778,1565,1588]" italics="true" pageId="2" pageNumber="103">L.</emphasis>
)
<taxonomicName id="4E804D0AF614FFC50851F9FE1E80D446" box="[796,916,1564,1589]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF614FFC50851F9FE1E80D446" box="[796,916,1564,1589]" italics="true" pageId="2" pageNumber="103">nigrilabris</emphasis>
</taxonomicName>
have a doughnut-shaped spermatheca with a broad, bent velum, a deep, relatively narrow invagination, a relatively long, robust spermathecal duct, and apparently no bursal duct; the subgenitale has a well-sclerotized ventral process that extends perpendicularly from the distal margin of the base.
</paragraph>
<paragraph id="893F3689F614FFC50B8AF94E19D1D596" blockId="2.[151,1436,772,2021]" pageId="2" pageNumber="103">
<emphasis id="BBF4EA9BF614FFC50B8AF94E1CC4D4B6" bold="true" box="[199,464,1708,1733]" pageId="2" pageNumber="103">Redescription (adult).</emphasis>
<emphasis id="BBF4EA9BF614FFC50A96F94E1F03D4B6" box="[475,535,1708,1733]" italics="true" pageId="2" pageNumber="103">Head</emphasis>
(
<figureCitation id="11BB2A0CF614FFC50961F94E1F66D4B7" box="[556,626,1708,1733]" captionStart="FIGURE 1" captionStartId="3.[151,250,1700,1722]" captionTargetBox="[162,1436,193,1676]" captionTargetId="figure@3.[162,1447,193,1677]" captionTargetPageId="3" captionText="FIGURE 1. Leucochrysa (L.) nigrilabris external features. A, B. Head, frontal; C. Head, dorsal; D. Body, dorsal E, F. Head and thorax, dorsal [note the variation in the coloration of the mesoprescutum and mesoscutellum]; G. Head, thorax, lateral; H. Head, prothorax, lateral [A, D, State of Yaracuy, Venezuela, female; B, C, F H, State of Rio de Janeiro, Brazil, female; E, State of Aragua, Venezuela, female]." httpUri="https://zenodo.org/record/248025/files/figure.png" pageId="2" pageNumber="103">Fig. 1</figureCitation>
):
<quantity id="4E789B6CF614FFC509C3F94E1E0DD4B7" box="[654,793,1708,1732]" metricMagnitude="-3" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="2" pageNumber="103" unit="mm" value="1.7999999999999998" valueMax="1.9" valueMin="1.7">1.71.9 mm</quantity>
wide (including eyes). Frons, clypeus white to creamcolored, frons without red or pink suffusion, clypeus with or without tinge of pink; genae reddish brown; labrum black throughout; maxillary, labial palpi yellowish to cream-colored. Vertex with central area raised, green mesally, golden brown to reddish brown laterally; lateral, frontal areas cream-colored, with prominent, dark red to dark brown, scalloped to V-shaped mark along anterior margin, extending anteriorly to or between mesal margins of scapes, extending laterally to eyes; post-ocular area cream-colored, without spots. Antenna: scape cream-colored ventrally, with reddish brown mark of various sizes on distal, dorsolateral surface; pedicel cream-colored basally, with brown band distally; flagellum cream-colored, with brown bristles; mesal-ventral surface of basal one to seven flagellomeres lightly tinged with brown (variable); dorsal antennal fossae cream-colored.
</paragraph>
</subSubSection>
<caption id="DDFF6601F615FFC40BDAF9461CA3D565" httpUri="https://zenodo.org/record/248025/files/figure.png" pageId="3" pageNumber="104" targetBox="[162,1436,193,1676]" targetPageId="3">
<paragraph id="893F3689F615FFC40BDAF9461CA3D565" blockId="3.[151,1435,1700,1814]" pageId="3" pageNumber="104">
<emphasis id="BBF4EA9BF615FFC40BDAF9461C06D4C9" bold="true" box="[151,274,1700,1722]" pageId="3" pageNumber="104">FIGURE 1.</emphasis>
<taxonomicName id="4E804D0AF615FFC40A55F9461C8ED4C9" box="[280,410,1700,1722]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF615FFC40A55F9461C8ED4C9" box="[280,410,1700,1722]" italics="true" pageId="3" pageNumber="104">Leucochrysa</emphasis>
</taxonomicName>
(
<emphasis id="BBF4EA9BF615FFC40AEAF9441CA8D4C8" box="[423,444,1702,1723]" italics="true" pageId="3" pageNumber="104">L.</emphasis>
)
<taxonomicName id="4E804D0AF615FFC40A87F9461F22D4C9" box="[458,566,1700,1722]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="104" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF615FFC40A87F9461F22D4C9" box="[458,566,1700,1722]" italics="true" pageId="3" pageNumber="104">nigrilabris</emphasis>
</taxonomicName>
external features. A, B. Head, frontal; C. Head, dorsal; D. Body, dorsal E, F. Head and thorax, dorsal [note the variation in the coloration of the mesoprescutum and mesoscutellum]; G. Head, thorax, lateral; H. Head, prothorax, lateral [A, D, State of Yaracuy, Venezuela, female; B, C, FH, State of Rio de Janeiro, Brazil, female; E, State of Aragua, Venezuela, female].
</paragraph>
</caption>
<subSubSection id="C19A6502F615FFD70B8AF8DF1912D083" lastPageId="16" lastPageNumber="117" pageId="3" pageNumber="104" type="description">
<paragraph id="893F3689F615FFC30B8AF8DF1FFAD36E" blockId="3.[151,1436,1853,2023]" lastBlockId="4.[151,1436,151,285]" lastPageId="4" lastPageNumber="105" pageId="3" pageNumber="104">
<emphasis id="BBF4EA9BF615FFC40B8AF8DF1C0CD525" box="[199,280,1853,1878]" italics="true" pageId="3" pageNumber="104">Thorax</emphasis>
(
<figureCitation id="11BB2A0CF615FFC40A6AF8DF1C7ED525" box="[295,362,1853,1878]" captionStart="FIGURE 1" captionStartId="3.[151,250,1700,1722]" captionTargetBox="[162,1436,193,1676]" captionTargetId="figure@3.[162,1447,193,1677]" captionTargetPageId="3" captionText="FIGURE 1. Leucochrysa (L.) nigrilabris external features. A, B. Head, frontal; C. Head, dorsal; D. Body, dorsal E, F. Head and thorax, dorsal [note the variation in the coloration of the mesoprescutum and mesoscutellum]; G. Head, thorax, lateral; H. Head, prothorax, lateral [A, D, State of Yaracuy, Venezuela, female; B, C, F H, State of Rio de Janeiro, Brazil, female; E, State of Aragua, Venezuela, female]." httpUri="https://zenodo.org/record/248025/files/figure.png" pageId="3" pageNumber="104">Figs 1</figureCitation>
DH): Cervical sclerite with large, red mark laterally. Pronotum
<quantity id="4E789B6CF615FFC40F0FF8DC19F1D525" box="[1090,1253,1854,1878]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" metricValueMax="9.8" metricValueMin="9.2" pageId="3" pageNumber="104" unit="mm" value="0.95" valueMax="0.98" valueMin="0.92">0.920.98 mm</quantity>
long,
<quantity id="4E789B6CF615FFC40E61F8DC1DD7D509" metricMagnitude="-3" metricUnit="m" metricValue="1.4249999999999998" metricValueMax="1.44" metricValueMin="1.41" pageId="3" pageNumber="104" unit="mm" value="1.4249999999999998" valueMax="1.44" valueMin="1.41">1.411.44 mm</quantity>
wide, light green, with elongate, dark red stripe on anterolateral margin, diffuse, light brown, sublateral spots anteriorly, posteriorly. Mesonotum, metanotum variable, with two distinct morphs [
<figureCitation id="11BB2A0CF615FFC40F34F86719D0D5ED" box="[1145,1220,1925,1950]" captionStart="FIGURE 1" captionStartId="3.[151,250,1700,1722]" captionTargetBox="[162,1436,193,1676]" captionTargetId="figure@3.[162,1447,193,1677]" captionTargetPageId="3" captionText="FIGURE 1. Leucochrysa (L.) nigrilabris external features. A, B. Head, frontal; C. Head, dorsal; D. Body, dorsal E, F. Head and thorax, dorsal [note the variation in the coloration of the mesoprescutum and mesoscutellum]; G. Head, thorax, lateral; H. Head, prothorax, lateral [A, D, State of Yaracuy, Venezuela, female; B, C, F H, State of Rio de Janeiro, Brazil, female; E, State of Aragua, Venezuela, female]." httpUri="https://zenodo.org/record/248025/files/figure.png" pageId="3" pageNumber="104">Figs 1</figureCitation>
D, E (brown), 1F (white)]. Brown morph: mesonotum, metanotum dark brown throughout, except anterior margin of prescutum cream-colored, with pair of sublateral, dark brown spots; mesoscutellum slightly lighter brown than remainder of notum; metascutellum black throughout. White morph: mesonotum mottled black to brownish black, anterior margin of prescutum cream-colored, with pair of sublateral, dark brown spots, posterior margin with cream-colored areas, mesoscutellum entirely white; metanotum largely dark brown to black, metascutum with large, white posteromesal patch; metascutellum black throughout.
</paragraph>
<caption id="DDFF6601F612FFC30BDAF84F1866D591" httpUri="https://zenodo.org/record/248026/files/figure.png" pageId="4" pageNumber="105" targetBox="[151,1436,337,1893]" targetPageId="4">
<paragraph id="893F3689F612FFC30BDAF84F1866D591" blockId="4.[151,1436,1965,2018]" pageId="4" pageNumber="105">
<emphasis id="BBF4EA9BF612FFC30BDAF84F1C01D5B0" bold="true" box="[151,277,1965,1987]" pageId="4" pageNumber="105">FIGURE 2.</emphasis>
<taxonomicName id="4E804D0AF612FFC30A53F84F1C8BD5B0" box="[286,415,1965,1987]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF612FFC30A53F84F1C8BD5B0" box="[286,415,1965,1987]" italics="true" pageId="4" pageNumber="105">Leucochrysa</emphasis>
</taxonomicName>
(
<emphasis id="BBF4EA9BF612FFC30AFFF84D1CD3D5B7" box="[434,455,1967,1988]" italics="true" pageId="4" pageNumber="105">L.</emphasis>
)
<taxonomicName id="4E804D0AF612FFC30A9AF84F1F57D5B0" box="[471,579,1965,1987]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="4" pageNumber="105" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF612FFC30A9AF84F1F57D5B0" box="[471,579,1965,1987]" italics="true" pageId="4" pageNumber="105">nigrilabris</emphasis>
</taxonomicName>
wings. A, B. Forewing, two specimens to illustrate relatively minor variation; C. Hindwing [A, State of Yaracuy, Venezuela; B, C, State of Aragua, Venezuela, all female]. Scale applies to all three images.
</paragraph>
</caption>
<caption id="DDFF6601F613FFC20BDAF9B11F85D533" httpUri="https://zenodo.org/record/248027/files/figure.png" pageId="5" pageNumber="106" targetBox="[151,1436,193,1597]" targetPageId="5">
<paragraph id="893F3689F613FFC20BDAF9B11F85D533" blockId="5.[151,1436,1619,1856]" pageId="5" pageNumber="106">
<emphasis id="BBF4EA9BF613FFC20BDAF9B11C1AD41A" bold="true" box="[151,270,1619,1641]" pageId="5" pageNumber="106">FIGURE 3</emphasis>
.
<taxonomicName id="4E804D0AF613FFC20A51F9B11C8AD41A" box="[284,414,1619,1641]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF613FFC20A51F9B11C8AD41A" box="[284,414,1619,1641]" italics="true" pageId="5" pageNumber="106">Leucochrysa</emphasis>
</taxonomicName>
(
<emphasis id="BBF4EA9BF613FFC20AE0F9B61CD7D41A" box="[429,451,1620,1641]" italics="true" pageId="5" pageNumber="106">L.</emphasis>
)
<taxonomicName id="4E804D0AF613FFC20A9FF9B11F2AD41A" box="[466,574,1619,1641]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="5" pageNumber="106" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF613FFC20A9FF9B11F2AD41A" box="[466,574,1619,1641]" italics="true" pageId="5" pageNumber="106">nigrilabris</emphasis>
</taxonomicName>
male abdomen, mature specimen. A. Second and third abdominal segments, lateral [note absence of microtholi from S2, dense microtholi on S3]; B. Terminal segments, lateral; C. Terminal segments, dorsal [note large brown spots on T5, T6, base of T7 (arrow)]; D. Terminal segments, dorsal [note dorsally fused T9+ectoproct, without suture or distal cleft. The U-shaped darkening is the distal margin of S8+9 below]; E. Callus cerci; F. Fused sternite 8+9, ventral, with sclerotized terminal plates extended distally [all, Aragua, Venezuela].
<emphasis id="BBF4EA9BF613FFC20F6CF92C19A2D497" box="[1057,1206,1742,1764]" italics="true" pageId="5" pageNumber="106">Abbreviations:</emphasis>
c.c., callus cerci; d.a., dorsal apodeme of ninth tergite+ectoproct; s.p., sclerotized plate at tip of S8+9; S2, S3, second and third sternites; S8+9, fused eighth and ninth sternites (in parentheses, out of focus through dorsal surface); tr, trichobothria; T5, T7, T8, fifth to eighth tergites; T9+ect, fused ninth tergite and ectoproct.
</paragraph>
</caption>
<caption id="DDFF6601F610FFC10BDAF9941F95D5F1" httpUri="https://zenodo.org/record/248028/files/figure.png" pageId="6" pageNumber="107" targetBox="[151,1436,193,1632]" targetPageId="6">
<paragraph id="893F3689F610FFC10BDAF9941F95D5F1" blockId="6.[151,1436,1654,1922]" pageId="6" pageNumber="107">
<emphasis id="BBF4EA9BF610FFC10BDAF9941C18D4FF" bold="true" box="[151,268,1654,1676]" pageId="6" pageNumber="107">FIGURE 4</emphasis>
.
<taxonomicName id="4E804D0AF610FFC10A57F9941C8FD4FF" box="[282,411,1654,1676]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF610FFC10A57F9941C8FD4FF" box="[282,411,1654,1676]" italics="true" pageId="6" pageNumber="107">Leucochrysa</emphasis>
</taxonomicName>
(
<emphasis id="BBF4EA9BF610FFC10AE6F99A1CD4D4FE" box="[427,448,1656,1677]" italics="true" pageId="6" pageNumber="107">L.</emphasis>
)
<taxonomicName id="4E804D0AF610FFC10A82F9941F2FD4FF" box="[463,571,1654,1676]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="6" pageNumber="107" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF610FFC10A82F9941F2FD4FF" box="[463,571,1654,1676]" italics="true" pageId="6" pageNumber="107">nigrilabris</emphasis>
</taxonomicName>
male genitalia, mature specimen. A. Gonarcus, posterodorsal (mediuncus removed); B. Gonarcal complex, posterior (view into gonosaccus from behind, through membrane) [note the mediuncal beak and flanges extend forward (away) and downward]; C. Gonarcal complex, dorsal [view into gonosaccus from above, unobscured by membrane, base of mediuncus in focus, mediuncal beak present (extending forward), but out of focus]; D. Gonarcal complex, frontal [note the gonosaccus is below the mediuncal flanges; the gonosetae are within the gonosaccus]; E. Gonarcal complex, lateral; F. Hypandrium internum (comes very reduced or absent) [all, Aragua, Venezuela].
<emphasis id="BBF4EA9BF610FFC10F77F8F219DBD555" box="[1082,1231,1808,1830]" italics="true" pageId="6" pageNumber="107">Abbreviations:</emphasis>
bk, beak-like tip of mediuncus; fl, mediuncal flange; gc, gonocornu, encased in membrane; g.a., gonarcal apodeme; g.b., gonarcal bridge; g.p., gonarcal process (membrane attachment); gsac, gonosaccus; gse, gonosetae (within gonosaccus); mu, mediuncus (fused rods and beak); mu-b, sclerotized base of mediuncus.
</paragraph>
</caption>
<caption id="DDFF6601F611FFC00BDAF9581943D590" httpUri="https://zenodo.org/record/248029/files/figure.png" pageId="7" pageNumber="108" targetBox="[151,1436,193,1699]" targetPageId="7">
<paragraph id="893F3689F611FFC00BDAF9581943D590" blockId="7.[151,1436,1722,2019]" pageId="7" pageNumber="108">
<emphasis id="BBF4EA9BF611FFC00BDAF9581C01D4A3" bold="true" box="[151,277,1722,1744]" pageId="7" pageNumber="108">FIGURE 5.</emphasis>
<taxonomicName id="4E804D0AF611FFC00A53F9581CB4D4A3" box="[286,416,1722,1744]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF611FFC00A53F9581CB4D4A3" box="[286,416,1722,1744]" italics="true" pageId="7" pageNumber="108">Leucochrysa</emphasis>
</taxonomicName>
(
<emphasis id="BBF4EA9BF611FFC00AFCF9591CD2D4A3" box="[433,454,1723,1744]" italics="true" pageId="7" pageNumber="108">L.</emphasis>
)
<taxonomicName id="4E804D0AF611FFC00A9BF9581F56D4A3" box="[470,578,1722,1744]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="7" pageNumber="108" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF611FFC00A9BF9581F56D4A3" box="[470,578,1722,1744]" italics="true" pageId="7" pageNumber="108">nigrilabris</emphasis>
</taxonomicName>
male abdomen, teneral specimen. A. Terminal segments, lateral (arrows mark the large brown spots on T6, base of T7); B. Gonarcal complex, frontal; C. Gonarcal complex, lateral [note thorny membrane over gonarcal bridge and gonocornua]; D. Gonarcal complex, lateral (thorny membrane above gonarcus partially removed); E. Tip of sternite 8+9, dorsal, with sclerotized terminal plates extended distally, hypandrium internum (attached to small membrane that had been everted beneath the sclerotized plate); F. Sclerotized plate at tip of S8+9, lateral (all, Aragua, Venezuela).
<emphasis id="BBF4EA9BF611FFC00BDAF8B11C38D51A" box="[151,300,1875,1897]" italics="true" pageId="7" pageNumber="108">Abbreviations:</emphasis>
bk, beak-like tip of mediuncus; c, comes; cleft, cleft separating distal margins of T9+ect; c.c., callus cerci; d.a., dorsal apodeme of T9+ect; fl, mediuncal flange; gc, gonocornu; gs, gonarcus; gsac, gonosaccus with robust gonosetae on bulbous chalazae; g.a., gonarcal apodeme; g.b., gonarcal bridge; h.i., hypandrium internum; mu-b, sclerotized base of mediuncus; mu-d, elongate distal section of mediuncus; s.p., sclerotized plate at tip of S8+9; S8+9, fused eighth and ninth sternites; S9, ninth sternite (tip); T7, seventh tergite; T9+ect, fused ninth tergite and ectoproct..
</paragraph>
</caption>
<caption id="DDFF6601F61EFFCF0BDAFB7E1F85D77E" httpUri="https://zenodo.org/record/248030/files/figure.png" pageId="8" pageNumber="109" targetBox="[151,1436,193,1158]" targetPageId="8">
<paragraph id="893F3689F61EFFCF0BDAFB7E1F85D77E" blockId="8.[151,1435,1180,1293]" pageId="8" pageNumber="109">
<emphasis id="BBF4EA9BF61EFFCF0BDAFB7E1C1BD6C1" bold="true" box="[151,271,1180,1202]" pageId="8" pageNumber="109">FIGURE 6</emphasis>
.
<taxonomicName id="4E804D0AF61EFFCF0A53FB7E1CB4D6C1" box="[286,416,1180,1202]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="109" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF61EFFCF0A53FB7E1CB4D6C1" box="[286,416,1180,1202]" italics="true" pageId="8" pageNumber="109">Leucochrysa</emphasis>
</taxonomicName>
(
<emphasis id="BBF4EA9BF61EFFCF0AFDFB7F1CD1D6C1" box="[432,453,1181,1202]" italics="true" pageId="8" pageNumber="109">L.</emphasis>
)
<taxonomicName id="4E804D0AF61EFFCF0A9BFB7E1F56D6C1" box="[470,578,1180,1202]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="8" pageNumber="109" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF61EFFCF0A9BFB7E1F56D6C1" box="[470,578,1180,1202]" italics="true" pageId="8" pageNumber="109">nigrilabris</emphasis>
</taxonomicName>
female abdomen. A. Fifth to terminal segments, lateral; B-D. Seventh to terminal segment, lateral [A, B, mature specimens, State of Rio de Janeiro, Brazil; C, D, teneral specimens, State of Aragua, Venezuela].
<emphasis id="BBF4EA9BF61EFFCF0BDAFB3B1C38D69C" box="[151,300,1241,1263]" italics="true" pageId="8" pageNumber="109">Abbreviations:</emphasis>
g.l., gonapophysis lateralis; sg, subgenitale; sp, spermatheca; S7, seventh sternite; T5, T8, fifth and eighth tergites; T9+ect, fused ninth tergite and ectoproct.
</paragraph>
</caption>
<paragraph id="893F3689F61EFFCF0B8AFAD41F80D551" blockId="8.[151,1436,1332,1970]" pageId="8" pageNumber="109">
<emphasis id="BBF4EA9BF61EFFCF0B8AFAD41C18D73E" box="[199,268,1334,1357]" italics="true" pageId="8" pageNumber="109">Wings</emphasis>
(
<figureCitation id="11BB2A0CF61EFFCF0A50FAD61C76D73E" box="[285,354,1332,1357]" captionStart="FIGURE 2" captionStartId="4.[151,250,1965,1987]" captionTargetBox="[151,1436,337,1893]" captionTargetId="figure@4.[151,1436,337,1944]" captionTargetPageId="4" captionText="FIGURE 2. Leucochrysa (L.) nigrilabris wings. A, B. Forewing, two specimens to illustrate relatively minor variation; C. Hindwing [A, State of Yaracuy, Venezuela; B, C, State of Aragua, Venezuela, all female]. Scale applies to all three images." httpUri="https://zenodo.org/record/248026/files/figure.png" pageId="8" pageNumber="109">Fig. 2</figureCitation>
) Forewing
<quantity id="4E789B6CF61EFFCF0AA5FAD71F99D73E" box="[488,653,1333,1357]" metricMagnitude="-2" metricUnit="m" metricValue="1.935" metricValueMax="1.9899999999999998" metricValueMin="1.88" pageId="8" pageNumber="109" unit="mm" value="19.35" valueMax="19.9" valueMin="18.8">18.819.9 mm</quantity>
long,
<quantity id="4E789B6CF61EFFCF0995FAD61E75D73E" box="[728,865,1332,1357]" metricMagnitude="-3" metricUnit="m" metricValue="6.7" metricValueMax="6.9" metricValueMin="6.5" pageId="8" pageNumber="109" unit="mm" value="6.7" valueMax="6.9" valueMin="6.5">6.56.9 mm</quantity>
wide (at widest point); ratio of length: maximum width = 2.62.7:1. Costal area moderately broad; tallest costal cell (#910)
<quantity id="4E789B6CF61EFFCF08BFFABB196ED702" box="[1010,1146,1369,1394]" metricMagnitude="-3" metricUnit="m" metricValue="1.65" metricValueMax="1.7" metricValueMin="1.6" pageId="8" pageNumber="109" unit="mm" value="1.65" valueMax="1.7" valueMin="1.6">1.61.7 mm</quantity>
tall, 3.03.1 times width, 0.20.3 times width of wing (midwing). First intramedian cell quadrangular, width (anterior margin) 1.41.7 times width (anterior margin) of third median cell, 0.91.1 times length of posterior margin of m3; length of basal vein (= ma, median arculus) 2.16.2 times greater than length of distal vein. First radial crossvein distal to origin of radial sector (Rs); radial area (between Radius and Rs) with single row of 1718 closed cells; tallest cell (#8) 2.32.4 times taller than wide. No crassate veins; 56 b cells (= cells beneath Rs, not including an inner gradate vein). Two series of gradate veins; 1012 inner gradates, 810 outer gradates. Height of fourth gradate cell 4.35.3 times width. Eight to nine b' cells (cells beneath pseudomedia after second intramedian cell). Three intracubital cells (two closed). Membrane clear except stigma opaque, with dark brown mark basally, distal two crossveins of last b cell (distal Psm-Pcu crossveins) surrounded by dark brown suffusion, and (some specimens) small amount of brown suffusion basally. Veins green, except distal two crossveins of last b cell, inner and outer gradates dark brown to black, sometimes anterior tips of basal R-Rs crossveins, base of Rs, basal Rs-m, basal medial crossveins, tips of posterobasal marginal veins lightly darkened.
</paragraph>
<paragraph id="893F3689F61EFFCF0B8AF8CF1FC6D5C1" blockId="8.[151,1436,1332,1970]" pageId="8" pageNumber="109">
Hindwing
<quantity id="4E789B6CF61EFFCF0A0DF8CF1CF0D536" box="[320,484,1837,1862]" metricMagnitude="-2" metricUnit="m" metricValue="1.72" metricValueMax="1.77" metricValueMin="1.67" pageId="8" pageNumber="109" unit="mm" value="17.2" valueMax="17.7" valueMin="16.7">16.717.7 mm</quantity>
long,
<quantity id="4E789B6CF61EFFCF0963F8CF1FA3D536" box="[558,695,1837,1862]" metricMagnitude="-3" metricUnit="m" metricValue="5.65" metricValueMax="5.8" metricValueMin="5.5" pageId="8" pageNumber="109" unit="mm" value="5.65" valueMax="5.8" valueMin="5.5">5.55.8 mm</quantity>
wide. Two series of gradate veins; 910 inner, 89 outer; 1516 radial cells (counted from origin of Radius, not false origin). Five to six b cells (including small b1 cell); seven to nine b' cells beyond second intramedian cell; two intracubital cells (one closed). Membrane clear; stigma with pronounced brown mark basally. Veins light green.
</paragraph>
<paragraph id="893F3689F61FFFCE0B8AFF751C96D316" blockId="9.[151,1436,151,357]" pageId="9" pageNumber="110">
<emphasis id="BBF4EA9BF61FFFCE0B8AFF751C25D2C3" box="[199,305,151,176]" italics="true" pageId="9" pageNumber="110">Abdomen</emphasis>
(
<figureCitation id="11BB2A0CF61FFFCE0A0DFF751CB3D2C3" box="[320,423,151,176]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStart-5="FIGURE 8" captionStartId-0="5.[151,250,1619,1641]" captionStartId-1="6.[151,250,1654,1676]" captionStartId-2="7.[151,250,1722,1744]" captionStartId-3="8.[151,250,1180,1202]" captionStartId-4="9.[151,250,1392,1414]" captionStartId-5="10.[151,250,1715,1737]" captionTargetBox-0="[151,1436,193,1597]" captionTargetBox-1="[151,1436,193,1632]" captionTargetBox-2="[151,1436,193,1699]" captionTargetBox-3="[151,1436,193,1158]" captionTargetBox-4="[151,1436,409,1370]" captionTargetBox-5="[151,1436,589,1693]" captionTargetId-0="figure@5.[151,1436,193,1598]" captionTargetId-1="figure@6.[151,1436,193,1633]" captionTargetId-2="figure@7.[151,1436,193,1700]" captionTargetId-3="figure@8.[151,1436,193,1158]" captionTargetId-4="figure@9.[151,1436,409,1371]" captionTargetId-5="figure@10.[151,1436,589,1694]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionText-0="FIGURE 3. Leucochrysa (L.) nigrilabris male abdomen, mature specimen. A. Second and third abdominal segments, lateral [note absence of microtholi from S 2, dense microtholi on S 3]; B. Terminal segments, lateral; C. Terminal segments, dorsal [note large brown spots on T 5, T 6, base of T 7 (arrow)]; D. Terminal segments, dorsal [note dorsally fused T 9 + ectoproct, without suture or distal cleft. The U-shaped darkening is the distal margin of S 8 + 9 below]; E. Callus cerci; F. Fused sternite 8 + 9, ventral, with sclerotized terminal plates extended distally [all, Aragua, Venezuela]. Abbreviations: c. c., callus cerci; d. a., dorsal apodeme of ninth tergite + ectoproct; s. p., sclerotized plate at tip of S 8 + 9; S 2, S 3, second and third sternites; S 8 + 9, fused eighth and ninth sternites (in parentheses, out of focus through dorsal surface); tr, trichobothria; T 5, T 7, T 8, fifth to eighth tergites; T 9 + ect, fused ninth tergite and ectoproct." captionText-1="FIGURE 4. Leucochrysa (L.) nigrilabris male genitalia, mature specimen. A. Gonarcus, posterodorsal (mediuncus removed); B. Gonarcal complex, posterior (view into gonosaccus from behind, through membrane) [note the mediuncal beak and flanges extend forward (away) and downward]; C. Gonarcal complex, dorsal [view into gonosaccus from above, unobscured by membrane, base of mediuncus in focus, mediuncal beak present (extending forward), but out of focus]; D. Gonarcal complex, frontal [note the gonosaccus is below the mediuncal flanges; the gonosetae are within the gonosaccus]; E. Gonarcal complex, lateral; F. Hypandrium internum (comes very reduced or absent) [all, Aragua, Venezuela]. Abbreviations: bk, beak-like tip of mediuncus; fl, mediuncal flange; gc, gonocornu, encased in membrane; g. a., gonarcal apodeme; g. b., gonarcal bridge; g. p., gonarcal process (membrane attachment); gsac, gonosaccus; gse, gonosetae (within gonosaccus); mu, mediuncus (fused rods and beak); mu-b, sclerotized base of mediuncus." captionText-2="FIGURE 5. Leucochrysa (L.) nigrilabris male abdomen, teneral specimen. A. Terminal segments, lateral (arrows mark the large brown spots on T 6, base of T 7); B. Gonarcal complex, frontal; C. Gonarcal complex, lateral [note thorny membrane over gonarcal bridge and gonocornua]; D. Gonarcal complex, lateral (thorny membrane above gonarcus partially removed); E. Tip of sternite 8 + 9, dorsal, with sclerotized terminal plates extended distally, hypandrium internum (attached to small membrane that had been everted beneath the sclerotized plate); F. Sclerotized plate at tip of S 8 + 9, lateral (all, Aragua, Venezuela). Abbreviations: bk, beak-like tip of mediuncus; c, comes; cleft, cleft separating distal margins of T 9 + ect; c. c., callus cerci; d. a., dorsal apodeme of T 9 + ect; fl, mediuncal flange; gc, gonocornu; gs, gonarcus; gsac, gonosaccus with robust gonosetae on bulbous chalazae; g. a., gonarcal apodeme; g. b., gonarcal bridge; h. i., hypandrium internum; mu-b, sclerotized base of mediuncus; mu-d, elongate distal section of mediuncus; s. p., sclerotized plate at tip of S 8 + 9; S 8 + 9, fused eighth and ninth sternites; S 9, ninth sternite (tip); T 7, seventh tergite; T 9 + ect, fused ninth tergite and ectoproct .." captionText-3="FIGURE 6. Leucochrysa (L.) nigrilabris female abdomen. A. Fifth to terminal segments, lateral; B-D. Seventh to terminal segment, lateral [A, B, mature specimens, State of Rio de Janeiro, Brazil; C, D, teneral specimens, State of Aragua, Venezuela]. Abbreviations: g. l., gonapophysis lateralis; sg, subgenitale; sp, spermatheca; S 7, seventh sternite; T 5, T 8, fifth and eighth tergites; T 9 + ect, fused ninth tergite and ectoproct." captionText-4="FIGURE 7. Leucochrysa (L.) nigrilabris female abdominal and genitalic structures. A. Callus cerci; B. Subgenitale, posterior; C. Spermatheca, lateral; D. Genitalia, ventral [A, B, D, State of Rio de Janeiro, Brazil, mature specimen; C, State of Aragua, Venezuela]. Abbreviations: base, sclerotized base of subgenitale; b. c., bursa copulatrix; b. gl., bursal gland; inv, spermathecal invagination; lobe, rounded lobe of subgenitale; sg / b. c., heavy folded membrane [distally (right): dorsum of subgenitale, proximally (left): venter of bursa copulatrix]; sp, spermatheca; sp. d., spermathecal duct; tr, trichobothria; vel, velum." captionText-5="FIGURE 8. Leucochrysa (L.) nigrilabris colleterial complex. A. Colleterial complex, lateral; B. Base of colleterial complex, lateral (pulled posteriorly); C. Transverse sclerite, posterior. [all, State of Rio de Janeiro, Brazil, mature specimen]. Abbreviations: acc. gl., colleterial accessory gland; acc. gl. d., duct of colleterial accessory gland; c. d., common duct (colleterial reservoir and accessory gland to transverse sclerite); col. gl., colleterial gland; col. r., colleterial reservoir; tr. s., transverse sclerite." httpUri-0="https://zenodo.org/record/248027/files/figure.png" httpUri-1="https://zenodo.org/record/248028/files/figure.png" httpUri-2="https://zenodo.org/record/248029/files/figure.png" httpUri-3="https://zenodo.org/record/248030/files/figure.png" httpUri-4="https://zenodo.org/record/248031/files/figure.png" httpUri-5="https://zenodo.org/record/248032/files/figure.png" pageId="9" pageNumber="110">Figs 38</figureCitation>
): Tergites with mostly short, slender setae throughout, sternites with longer, slender setae; microsetae dense; pleural region with setae small, very sparse, microsetae very small. Tergites narrow, roughly rectangular, with rounded or irregular margins. Spiracles oval externally; atria not enlarged. Sternites S2S5 longer than wide (lateral view). Coloration: mostly green, with diffuse yellow dorsal stripe; tergites T5, T6, base of T7 each with large dark brown to black spot, bordered with yellow and patches of red; callus cerci white; setae, trichobothria golden.
</paragraph>
<caption id="DDFF6601F61FFFCE0BDAFA921820D472" httpUri="https://zenodo.org/record/248031/files/figure.png" pageId="9" pageNumber="110" targetBox="[151,1436,409,1370]" targetPageId="9">
<paragraph id="893F3689F61FFFCE0BDAFA921820D472" blockId="9.[151,1436,1392,1537]" pageId="9" pageNumber="110">
<emphasis id="BBF4EA9BF61FFFCE0BDAFA921C07D7F5" bold="true" box="[151,275,1392,1414]" pageId="9" pageNumber="110">FIGURE 7.</emphasis>
<taxonomicName id="4E804D0AF61FFFCE0A54FA921C8ED7F5" box="[281,410,1392,1414]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF61FFFCE0A54FA921C8ED7F5" box="[281,410,1392,1414]" italics="true" pageId="9" pageNumber="110">Leucochrysa</emphasis>
</taxonomicName>
(
<emphasis id="BBF4EA9BF61FFFCE0AE4FA901CAAD7F4" box="[425,446,1394,1415]" italics="true" pageId="9" pageNumber="110">L.</emphasis>
)
<taxonomicName id="4E804D0AF61FFFCE0A81FA921F2CD7F5" box="[460,568,1392,1414]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="9" pageNumber="110" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF61FFFCE0A81FA921F2CD7F5" box="[460,568,1392,1414]" italics="true" pageId="9" pageNumber="110">nigrilabris</emphasis>
</taxonomicName>
female abdominal and genitalic structures. A. Callus cerci; B. Subgenitale, posterior; C. Spermatheca, lateral; D. Genitalia, ventral [A, B, D, State of Rio de Janeiro, Brazil, mature specimen; C, State of Aragua, Venezuela].
<emphasis id="BBF4EA9BF61FFFCE0A58FA4F1CBED7B0" box="[277,426,1453,1475]" italics="true" pageId="9" pageNumber="110">Abbreviations:</emphasis>
base, sclerotized base of subgenitale; b.c., bursa copulatrix; b.gl., bursal gland; inv, spermathecal invagination; lobe, rounded lobe of subgenitale; sg/b.c., heavy folded membrane [distally (right): dorsum of subgenitale, proximally (left): venter of bursa copulatrix]; sp, spermatheca; sp.d., spermathecal duct; tr, trichobothria; vel, velum.
</paragraph>
</caption>
<paragraph id="893F3689F61FFFCD0B8AF9CA1869D064" blockId="9.[151,1436,1576,2033]" lastBlockId="10.[151,1436,151,536]" lastPageId="10" lastPageNumber="111" pageId="9" pageNumber="110">
<emphasis id="BBF4EA9BF61FFFCE0B8AF9CA1C1CD432" box="[199,264,1576,1601]" italics="true" pageId="9" pageNumber="110">Male.</emphasis>
S6, S7 height
<emphasis id="BBF4EA9BF61FFFCE0AE7F9CB1CD1D433" box="[426,453,1577,1600]" italics="true" pageId="9" pageNumber="110">ca</emphasis>
1.11.2 times length (lateral view). Microtholi dense on S3S8, absent from S1S2, S9. Callus cerci oval (
<emphasis id="BBF4EA9BF61FFFCE0A22F9AC1C9ED416" box="[367,394,1614,1637]" italics="true" pageId="9" pageNumber="110">ca</emphasis>
1.31.5x taller than wide), greatest diameter
<quantity id="4E789B6CF61FFFCE08EFF9AE195FD417" box="[930,1099,1612,1637]" metricMagnitude="-4" metricUnit="m" metricValue="2.4" metricValueMax="2.6" metricValueMin="2.2" pageId="9" pageNumber="110" unit="mm" value="0.24" valueMax="0.26" valueMin="0.22">0.220.26 mm</quantity>
, with 2635 trichobothria of variable length. T9+ectoproct soft, lightly sclerotized, rounded posterodorsally, truncate distally, broadly fused mesally, midline without distal cleft (mature specimen; cleft present in teneral specimen); ventral section tapering proximally, extending beyond midline of S9; dorsal apodeme lightly to moderately sclerotized, straight, extending
<emphasis id="BBF4EA9BF61FFFCE0BDAF93C1DA6D486" box="[151,178,1758,1781]" italics="true" pageId="9" pageNumber="110">ca</emphasis>
half distance along dorsal margin of ectoproct; small, lightly sclerotized apodeme around proximal margin of callus cerci. S8+9 fused, without suture, with clear intersegmental demarcation throughout; dorsal margin with lightly sclerotized, straight apodeme; S8 1.7x taller than long,
<emphasis id="BBF4EA9BF61FFFCE0838F8C41E84D54E" box="[885,912,1830,1853]" italics="true" pageId="9" pageNumber="110">ca</emphasis>
one-half (0.470.51x) length of S8+9; S8+9 (lateral view) with proximal margin relatively straight, dorsal surface straight, sloping ventrally to distal margin; terminus rounded; tip with pair of flat, sclerotized, plate-like projections with mesal edge toothed, lateral edge rounded. Gonarcus arcuate; bridge straight, narrow mesally (
<quantity id="4E789B6CF61FFFCE082DF8731EDBD5DA" box="[864,975,1937,1961]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="110" unit="mm" value="0.19">0.19 mm</quantity>
), curved abruptly at exterior margin of gonocornua; gonarcal bridge, gonocornua covered with robust membrane bearing dense gonocristae; gonocornua short (length,
<quantity id="4E789B6CF61FFFCE0A78F83B1C89D582" box="[309,413,2009,2033]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="9" pageNumber="110" unit="mm" value="0.07">0.07 mm</quantity>
), extending forward from edge of gonarcal bridge, stout basally, tapering to rounded apex; distance between inner bases of gonocornua
<quantity id="4E789B6CF61CFFCD09EDFF7A1E1CD2DC" box="[672,776,152,176]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="10" pageNumber="111" unit="mm" value="0.07">0.07 mm</quantity>
basally, distance between tips
<quantity id="4E789B6CF61CFFCD0F3FFF7A19C8D2DC" box="[1138,1244,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="1.2" pageId="10" pageNumber="111" unit="mm" value="0.12">0.12 mm</quantity>
; gonarcal lateral apodeme long, relatively narrow (
<quantity id="4E789B6CF61CFFCD096FFF5E1F99D2A7" box="[546,653,188,213]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="111" unit="mm" value="0.6">0.60 mm</quantity>
long,
<quantity id="4E789B6CF61CFFCD0996FF5E1E50D2A7" box="[731,836,188,212]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="111" unit="mm" value="0.2">0.20 mm</quantity>
wide), not extending above bridge, curved distally (lateral view); distance between apodemes distally
<quantity id="4E789B6CF61CFFCD0996FF021E50D284" box="[731,836,224,248]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" pageId="10" pageNumber="111" unit="mm" value="0.72">0.72 mm</quantity>
. Gonarcus, between lateral apodeme and gonocornu, with moderately sclerotized, elongate posteroventral projection holding membrane that extends beneath mediuncus. Mediuncus well separated from gonarcus, with basal membranous connection stout; basal section of mediuncus consisting of heavily sclerotized platform with distal projections laterally, distal section consisting of well sclerotized, ventrally-projecting, smooth plate terminating in sharp, curved, elongate, mesal beak, pair of large lateral lobes; heavy membrane from gonarcus extending beneath mediuncus, recurving above beak to form gonosaccus with pair of elongate, lateral fields of robust gonosetae on bulbous chalazae; fields of gonosetae on surface of membrane facing mediuncus; each field with
<emphasis id="BBF4EA9BF61CFFCD0868FE3F1E54D387" box="[805,832,477,500]" italics="true" pageId="10" pageNumber="111">ca</emphasis>
1520 gonosetae. Hypandrium internum broadly Vshaped, with comes not visible (mature specimen); arm
<quantity id="4E789B6CF61CFFCD0844FDE21EB8D064" box="[777,940,512,536]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="10" pageNumber="111" unit="mm" value="0.21" valueMax="0.22" valueMin="0.2">0.200.22 mm</quantity>
long, distal span between arms
<quantity id="4E789B6CF61CFFCD0E5FFDE2186DD064" box="[1298,1401,512,536]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="10" pageNumber="111" unit="mm" value="0.19">0.19 mm</quantity>
.
</paragraph>
<caption id="DDFF6601F61CFFCD0BDAF9511DF3D537" httpUri="https://zenodo.org/record/248032/files/figure.png" pageId="10" pageNumber="111" targetBox="[151,1436,589,1693]" targetPageId="10">
<paragraph id="893F3689F61CFFCD0BDAF9511DF3D537" blockId="10.[151,1436,1715,1860]" pageId="10" pageNumber="111">
<emphasis id="BBF4EA9BF61CFFCD0BDAF9511C00D4BA" bold="true" box="[151,276,1715,1737]" pageId="10" pageNumber="111">FIGURE 8.</emphasis>
<taxonomicName id="4E804D0AF61CFFCD0A51F9511C8AD4BA" box="[284,414,1715,1737]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF61CFFCD0A51F9511C8AD4BA" box="[284,414,1715,1737]" italics="true" pageId="10" pageNumber="111">Leucochrysa</emphasis>
</taxonomicName>
(
<emphasis id="BBF4EA9BF61CFFCD0AE0F9561CD6D4BA" box="[429,450,1716,1737]" italics="true" pageId="10" pageNumber="111">L.</emphasis>
)
<taxonomicName id="4E804D0AF61CFFCD0A9CF9511F29D4BA" box="[465,573,1715,1737]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="10" pageNumber="111" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF61CFFCD0A9CF9511F29D4BA" box="[465,573,1715,1737]" italics="true" pageId="10" pageNumber="111">nigrilabris</emphasis>
</taxonomicName>
colleterial complex. A. Colleterial complex, lateral; B. Base of colleterial complex, lateral (pulled posteriorly); C. Transverse sclerite, posterior. [all, State of Rio de Janeiro, Brazil, mature specimen].
<emphasis id="BBF4EA9BF61CFFCD0BDAF9121C38D575" box="[151,300,1776,1798]" italics="true" pageId="10" pageNumber="111">Abbreviations:</emphasis>
acc. gl., colleterial accessory gland; acc.gl.d., duct of colleterial accessory gland; c.d., common duct (colleterial reservoir and accessory gland to transverse sclerite); col.gl., colleterial gland; col.r., colleterial reservoir; tr.s., transverse sclerite.
</paragraph>
</caption>
<paragraph id="893F3689F61CFFCC0B8AF8921C4DD14B" blockId="10.[151,1436,1904,2037]" lastBlockId="11.[151,1437,151,2013]" lastPageId="11" lastPageNumber="112" pageId="10" pageNumber="111">
<emphasis id="BBF4EA9BF61CFFCD0B8AF8921C36D5FA" box="[199,290,1904,1929]" italics="true" pageId="10" pageNumber="111">Female.</emphasis>
S6 height
<emphasis id="BBF4EA9BF61CFFCD0AE1F8931CD3D5FB" box="[428,455,1905,1928]" italics="true" pageId="10" pageNumber="111">ca</emphasis>
0.670.92 times length, S7 height
<emphasis id="BBF4EA9BF61CFFCD083EF8931E9AD5FB" box="[883,910,1905,1928]" italics="true" pageId="10" pageNumber="111">ca</emphasis>
0.550.65 times length. Callus cerci round, diameter
<emphasis id="BBF4EA9BF61CFFCD0A4EF8741C0AD5DE" box="[259,286,1942,1965]" italics="true" pageId="10" pageNumber="111">ca</emphasis>
<quantity id="4E789B6CF61CFFCD0A6AF8771CDAD5DF" box="[295,462,1941,1965]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" metricValueMax="1.9" metricValueMin="1.7" pageId="10" pageNumber="111" unit="mm" value="0.18" valueMax="0.19" valueMin="0.17">0.170.19 mm</quantity>
, with 2934 trichobothria. T8 roughly elliptical (lateral view) with rounded corners, similar in depth to T6. T9+ectoproct elongate, vertically erect; ventral margin slightly rounded, extending below level of gonapophyses laterales. S7 with dorsal margin straight, distal end tapering abruptly; terminus truncate, unmodified, with terminal (posteroventral) setae slightly more numerous, longer than in other areas. Gonapophysis lateralis rounded throughout, ~0.480.52 height of T9+ectoproct; inner membranous surface slightly expandable, with vertical patch of small, delicate setae arising from slightly swollen membrane. Colleterial complex consisting of membranous gland, connected to colleterial reservoir via broad duct, and large accessory gland, both opening to exterior via narrow common duct above transverse sclerite; colleterial gland elongate, delicate; colleterial reservoir shorter, robust; accessory gland probably bulbous (torn in our specimens), elongate, granular, with multiple forks distally; transverse sclerite broad, flat, bent distally, with relatively dense, elongate, hair-like setae. Spermatheca simple, doughnut-shaped, with broad, flat, L-shaped velum, maximum diameter
<quantity id="4E789B6CF61DFFCC0F14FE761811D3DF" box="[1113,1285,404,429]" metricMagnitude="-4" metricUnit="m" metricValue="2.85" metricValueMax="3.1" metricValueMin="2.6" pageId="11" pageNumber="112" unit="mm" value="0.285" valueMax="0.31" valueMin="0.26">0.260.31 mm</quantity>
, total length (spermatheca + velum)
<emphasis id="BBF4EA9BF61DFFCC0AE9FE5B1CABD3A3" box="[420,447,441,464]" italics="true" pageId="11" pageNumber="112">ca</emphasis>
<quantity id="4E789B6CF61DFFCC0A8BFE551F3BD3BC" box="[454,559,439,464]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="112" unit="mm" value="0.35">0.35 mm</quantity>
; spermathecal invagination deep, narrow, with rounded terminus; velum with elongate slit opening to bursa copulatrix; tip of velum bending slightly into bursa copulatrix; bursal duct very small or absent; spermathecal duct relatively long,
<emphasis id="BBF4EA9BF61DFFCC09C6FDE31FB2D06B" box="[651,678,513,536]" italics="true" pageId="11" pageNumber="112">ca</emphasis>
<quantity id="4E789B6CF61DFFCC09E1FE1D1E45D064" box="[684,849,511,536]" metricMagnitude="-3" metricUnit="m" metricValue="1.3199999999999998" metricValueMax="1.51" metricValueMin="1.13" pageId="11" pageNumber="112" unit="mm" value="1.3199999999999998" valueMax="1.51" valueMin="1.13">1.131.51 mm</quantity>
, originating from side of spermatheca distal to slit in velum, basal half well sclerotized, tapering throughout, with lateral bend, then U-shaped loop; distal
<emphasis id="BBF4EA9BF61DFFCC0E07FDC71871D04F" box="[1354,1381,549,572]" italics="true" pageId="11" pageNumber="112">ca</emphasis>
half narrow, brushy throughout, with curve, followed by distinct swelling mesally, with densely hirsute, swollen terminus. Bursa copulatrix saccular, with heavily textured surface distally (below subgenitale), becoming smoother anteriorly, with pair of elongate, tubular bursal glands attached to distal end, connections appearing unremarkable. Subgenitale with sclerotized surface smooth (unfolded), attached basally to heavy, transversely folded basal membrane (base of subgenitale then top of bursa copulatrix), with two projections, bilobed one on dorsodistal margin, unilobed one on anteroventral margin base withdrawn above tip of S7 [with pressure, the withdrawn lobe can be extruded].
</paragraph>
<paragraph id="893F3689F61DFFCC0B8AFCA61F94D19E" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="112">
<emphasis id="BBF4EA9BF61DFFCC0B8AFCA61F2CD12E" bold="true" box="[199,568,836,861]" pageId="11" pageNumber="112">Intraspecific variation (adult).</emphasis>
<taxonomicName id="4E804D0AF61DFFCC0909FCA61FC7D12E" box="[580,723,836,861]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="112" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF61DFFCC0909FCA61FC7D12E" box="[580,723,836,861]" italics="true" pageId="11" pageNumber="112">Leucochrysa</emphasis>
</taxonomicName>
(
<emphasis id="BBF4EA9BF61DFFCC09AAFCA71FEAD12F" box="[743,766,837,860]" italics="true" pageId="11" pageNumber="112">L.</emphasis>
)
<taxonomicName id="4E804D0AF61DFFCC085FFCA61E9DD12E" box="[786,905,836,861]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="11" pageNumber="112" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF61DFFCC085FFCA61E9DD12E" box="[786,905,836,861]" italics="true" pageId="11" pageNumber="112">nigrilabris</emphasis>
</taxonomicName>
expresses considerable variation in head and mesothoracic coloration, as well as wing size, shape and degree of brown suffusion surrounding various veins. The most striking aspect of the variation is the polymorphism in mesothoracic coloration (
<figureCitation id="11BB2A0CF61DFFCC0FC9FC6E19DED1D7" box="[1156,1226,908,933]" captionStart="FIGURE 1" captionStartId="3.[151,250,1700,1722]" captionTargetBox="[162,1436,193,1676]" captionTargetId="figure@3.[162,1447,193,1677]" captionTargetPageId="3" captionText="FIGURE 1. Leucochrysa (L.) nigrilabris external features. A, B. Head, frontal; C. Head, dorsal; D. Body, dorsal E, F. Head and thorax, dorsal [note the variation in the coloration of the mesoprescutum and mesoscutellum]; G. Head, thorax, lateral; H. Head, prothorax, lateral [A, D, State of Yaracuy, Venezuela, female; B, C, F H, State of Rio de Janeiro, Brazil, female; E, State of Aragua, Venezuela, female]." httpUri="https://zenodo.org/record/248025/files/figure.png" pageId="11" pageNumber="112">Figs 1</figureCitation>
DF). Among the specimens we examined, eight (4♀,
<specimenCount id="9F86FD00F61DFFCC0978FC521F4DD1BB" box="[565,601,944,968]" pageId="11" pageNumber="112" type="generic">4♂</specimenCount>
) had brown mesoscutella, and five (2♀,
<specimenCount id="9F86FD00F61DFFCC0F66FC52195AD1BB" box="[1067,1102,944,968]" pageId="11" pageNumber="112" type="generic">3♂</specimenCount>
) had white; both color forms occurred among the Venezuelan specimens.
</paragraph>
<paragraph id="893F3689F61DFFCC0B8AFC151817D77F" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="112">
The male genitalia show considerable developmental variation (Compare
<figureCitation id="11BB2A0CF61DFFCC0F45FC15195ED663" box="[1032,1098,1015,1040]" captionStart="FIGURE 5" captionStartId="7.[151,250,1722,1744]" captionTargetBox="[151,1436,193,1699]" captionTargetId="figure@7.[151,1436,193,1700]" captionTargetPageId="7" captionText="FIGURE 5. Leucochrysa (L.) nigrilabris male abdomen, teneral specimen. A. Terminal segments, lateral (arrows mark the large brown spots on T 6, base of T 7); B. Gonarcal complex, frontal; C. Gonarcal complex, lateral [note thorny membrane over gonarcal bridge and gonocornua]; D. Gonarcal complex, lateral (thorny membrane above gonarcus partially removed); E. Tip of sternite 8 + 9, dorsal, with sclerotized terminal plates extended distally, hypandrium internum (attached to small membrane that had been everted beneath the sclerotized plate); F. Sclerotized plate at tip of S 8 + 9, lateral (all, Aragua, Venezuela). Abbreviations: bk, beak-like tip of mediuncus; c, comes; cleft, cleft separating distal margins of T 9 + ect; c. c., callus cerci; d. a., dorsal apodeme of T 9 + ect; fl, mediuncal flange; gc, gonocornu; gs, gonarcus; gsac, gonosaccus with robust gonosetae on bulbous chalazae; g. a., gonarcal apodeme; g. b., gonarcal bridge; h. i., hypandrium internum; mu-b, sclerotized base of mediuncus; mu-d, elongate distal section of mediuncus; s. p., sclerotized plate at tip of S 8 + 9; S 8 + 9, fused eighth and ninth sternites; S 9, ninth sternite (tip); T 7, seventh tergite; T 9 + ect, fused ninth tergite and ectoproct .." httpUri="https://zenodo.org/record/248029/files/figure.png" pageId="11" pageNumber="112">Fig. 5</figureCitation>
with
<figureCitation id="11BB2A0CF61DFFCC0FC1FC1519C7D663" box="[1164,1235,1015,1040]" captionStart="FIGURE 3" captionStartId="5.[151,250,1619,1641]" captionTargetBox="[151,1436,193,1597]" captionTargetId="figure@5.[151,1436,193,1598]" captionTargetPageId="5" captionText="FIGURE 3. Leucochrysa (L.) nigrilabris male abdomen, mature specimen. A. Second and third abdominal segments, lateral [note absence of microtholi from S 2, dense microtholi on S 3]; B. Terminal segments, lateral; C. Terminal segments, dorsal [note large brown spots on T 5, T 6, base of T 7 (arrow)]; D. Terminal segments, dorsal [note dorsally fused T 9 + ectoproct, without suture or distal cleft. The U-shaped darkening is the distal margin of S 8 + 9 below]; E. Callus cerci; F. Fused sternite 8 + 9, ventral, with sclerotized terminal plates extended distally [all, Aragua, Venezuela]. Abbreviations: c. c., callus cerci; d. a., dorsal apodeme of ninth tergite + ectoproct; s. p., sclerotized plate at tip of S 8 + 9; S 2, S 3, second and third sternites; S 8 + 9, fused eighth and ninth sternites (in parentheses, out of focus through dorsal surface); tr, trichobothria; T 5, T 7, T 8, fifth to eighth tergites; T 9 + ect, fused ninth tergite and ectoproct." httpUri="https://zenodo.org/record/248027/files/figure.png" pageId="11" pageNumber="112">Figs 3</figureCitation>
,
<figureCitation id="11BB2A0CF61DFFCC0FACFC1A19E6D663" box="[1249,1266,1016,1040]" captionStart="FIGURE 4" captionStartId="6.[151,250,1654,1676]" captionTargetBox="[151,1436,193,1632]" captionTargetId="figure@6.[151,1436,193,1633]" captionTargetPageId="6" captionText="FIGURE 4. Leucochrysa (L.) nigrilabris male genitalia, mature specimen. A. Gonarcus, posterodorsal (mediuncus removed); B. Gonarcal complex, posterior (view into gonosaccus from behind, through membrane) [note the mediuncal beak and flanges extend forward (away) and downward]; C. Gonarcal complex, dorsal [view into gonosaccus from above, unobscured by membrane, base of mediuncus in focus, mediuncal beak present (extending forward), but out of focus]; D. Gonarcal complex, frontal [note the gonosaccus is below the mediuncal flanges; the gonosetae are within the gonosaccus]; E. Gonarcal complex, lateral; F. Hypandrium internum (comes very reduced or absent) [all, Aragua, Venezuela]. Abbreviations: bk, beak-like tip of mediuncus; fl, mediuncal flange; gc, gonocornu, encased in membrane; g. a., gonarcal apodeme; g. b., gonarcal bridge; g. p., gonarcal process (membrane attachment); gsac, gonosaccus; gse, gonosetae (within gonosaccus); mu, mediuncus (fused rods and beak); mu-b, sclerotized base of mediuncus." httpUri="https://zenodo.org/record/248028/files/figure.png" pageId="11" pageNumber="112">4</figureCitation>
). In the teneral male, the gonarcus is weakly sclerotized and narrow; the mediuncal bridge appears wide (in lateral view) and is tightly encased by the gonocristae-bearing membrane; the mediuncus is very narrow; and the hypandrium internum is unexpanded [length of arms, 0.20 mm—as in mature specimens; distal span between arms, 0.06 mm—
<emphasis id="BBF4EA9BF61DFFCC0E11FB871863D60F" box="[1372,1399,1125,1148]" italics="true" pageId="11" pageNumber="112">ca</emphasis>
<date id="FD3E1049F61DFFCC0ECDFB861888D60F" box="[1408,1436,1124,1148]" pageId="11" pageNumber="112">3x</date>
narrower than mature specimens]. Females, too, show considerable variation with age and maturation. For example, in the teneral female, the subgenitale is small, weakly sclerotized, and withdrawn, and the spermatheca is shifted anteriorly from the distal wall; whereas in mature specimens, the subgenitale is large, robust, heavily sclerotized and expanded, so that the spermatheca appears surrounded by the distal wall (
<figureCitation id="11BB2A0CF61DFFCC0F30FB1619D2D77E" box="[1149,1222,1268,1293]" captionStart="FIGURE 6" captionStartId="8.[151,250,1180,1202]" captionTargetBox="[151,1436,193,1158]" captionTargetId="figure@8.[151,1436,193,1158]" captionTargetPageId="8" captionText="FIGURE 6. Leucochrysa (L.) nigrilabris female abdomen. A. Fifth to terminal segments, lateral; B-D. Seventh to terminal segment, lateral [A, B, mature specimens, State of Rio de Janeiro, Brazil; C, D, teneral specimens, State of Aragua, Venezuela]. Abbreviations: g. l., gonapophysis lateralis; sg, subgenitale; sp, spermatheca; S 7, seventh sternite; T 5, T 8, fifth and eighth tergites; T 9 + ect, fused ninth tergite and ectoproct." httpUri="https://zenodo.org/record/248030/files/figure.png" pageId="11" pageNumber="112">Figs 6</figureCitation>
,
<figureCitation id="11BB2A0CF61DFFCC0F9FFB1619F5D77F" box="[1234,1249,1268,1292]" captionStart="FIGURE 7" captionStartId="9.[151,250,1392,1414]" captionTargetBox="[151,1436,409,1370]" captionTargetId="figure@9.[151,1436,409,1371]" captionTargetPageId="9" captionText="FIGURE 7. Leucochrysa (L.) nigrilabris female abdominal and genitalic structures. A. Callus cerci; B. Subgenitale, posterior; C. Spermatheca, lateral; D. Genitalia, ventral [A, B, D, State of Rio de Janeiro, Brazil, mature specimen; C, State of Aragua, Venezuela]. Abbreviations: base, sclerotized base of subgenitale; b. c., bursa copulatrix; b. gl., bursal gland; inv, spermathecal invagination; lobe, rounded lobe of subgenitale; sg / b. c., heavy folded membrane [distally (right): dorsum of subgenitale, proximally (left): venter of bursa copulatrix]; sp, spermatheca; sp. d., spermathecal duct; tr, trichobothria; vel, velum." httpUri="https://zenodo.org/record/248031/files/figure.png" pageId="11" pageNumber="112">7</figureCitation>
B).
</paragraph>
<paragraph id="893F3689F61DFFCC0B8AFAF51818D743" blockId="11.[151,1437,151,2013]" box="[199,1292,1303,1328]" pageId="11" pageNumber="112">
<emphasis id="BBF4EA9BF61DFFCC0B8AFAF51FD0D743" bold="true" box="[199,708,1303,1328]" pageId="11" pageNumber="112">Description (First Instar, Semaphoront A).</emphasis>
<emphasis id="BBF4EA9BF61DFFCC0987FAF51E10D743" box="[714,772,1303,1328]" italics="true" pageId="11" pageNumber="112">Body</emphasis>
(
<figureCitation id="11BB2A0CF61DFFCC085EFAF51E71D743" box="[787,869,1303,1328]" captionStart="FIGURE 10" captionStartId="12.[151,250,1401,1423]" captionTargetBox="[151,1436,814,1379]" captionTargetId="figure@12.[151,1436,814,1379]" captionTargetPageId="12" captionText="FIGURE 10. Leucochrysa (L.) nigrilabris first instar. A. Head, dorsal; B. Head, ventral; C. Head, lateral; D. Thorax, dorsal; E. Abdominal segments 1 to 5, dorsal; F. Abdominal segments 4 - 10, dorsal; G. Body, lateral; H. Body, ventral [all, State of Rio de Janeiro, Brazil]. Abbreviations: Ax, abdominal segment number; AxRx, setal row number." httpUri="https://zenodo.org/record/248034/files/figure.png" pageId="11" pageNumber="112">Fig. 10</figureCitation>
). 2.83.0 mm long. All setae smooth.
</paragraph>
<paragraph id="893F3689F61DFFCC0B8AFADE1C6DD4EB" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="112">
<emphasis id="BBF4EA9BF61DFFCC0B8AFADE1C10D726" box="[199,260,1340,1365]" italics="true" pageId="11" pageNumber="112">Head</emphasis>
(
<figureCitation id="11BB2A0CF61DFFCC0A5FFADE1C7CD727" box="[274,360,1340,1365]" captionStart="FIGURE 10" captionStartId="12.[151,250,1401,1423]" captionTargetBox="[151,1436,814,1379]" captionTargetId="figure@12.[151,1436,814,1379]" captionTargetPageId="12" captionText="FIGURE 10. Leucochrysa (L.) nigrilabris first instar. A. Head, dorsal; B. Head, ventral; C. Head, lateral; D. Thorax, dorsal; E. Abdominal segments 1 to 5, dorsal; F. Abdominal segments 4 - 10, dorsal; G. Body, lateral; H. Body, ventral [all, State of Rio de Janeiro, Brazil]. Abbreviations: Ax, abdominal segment number; AxRx, setal row number." httpUri="https://zenodo.org/record/248034/files/figure.png" pageId="11" pageNumber="112">Figs 10</figureCitation>
AC).
<quantity id="4E789B6CF61DFFCC0AF9FADE1F43D727" box="[436,599,1340,1365]" metricMagnitude="-4" metricUnit="m" metricValue="5.55" metricValueMax="5.6" metricValueMin="5.5" pageId="11" pageNumber="112" unit="mm" value="0.555" valueMax="0.56" valueMin="0.55">0.550.56 mm</quantity>
wide,
<quantity id="4E789B6CF61DFFCC09EEFADE1E52D727" box="[675,838,1340,1364]" metricMagnitude="-4" metricUnit="m" metricValue="3.85" metricValueMax="3.9" metricValueMin="3.8" pageId="11" pageNumber="112" unit="mm" value="0.385" valueMax="0.39" valueMin="0.38">0.380.39 mm</quantity>
long; mandibles
<quantity id="4E789B6CF61DFFCC0F4AFADE19BFD727" box="[1031,1195,1340,1365]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" metricValueMax="6.1" metricValueMin="5.8999999999999995" pageId="11" pageNumber="112" unit="mm" value="0.6" valueMax="0.61" valueMin="0.59">0.590.61 mm</quantity>
long; ratio, mandible length: head width, ~1.09:1. Epicranial marking brown, paired, extending along entire basal margin of cranium, laterally extending to eye, mesally becoming confluent with base of postfrontal marking. Postfrontal marking slightly darker brown than epicranial marking, elongate, extending to base of antenna. Frontal marking V-shaped, arms extending forward through dark tentorial pits becoming confluent with intermandibular marking. Intermandibular marking broken mesally, extending laterally from tentorial pits to base of mandibles, anteriorly to tip of clypeus. Gena with single, elongate, brown stripe extending from base of eye to posterior edge of cranium. Labial palpus cream-colored basally, light brown distally. Mandibles amber to brown. Antenna light brown to cream-colored. Venter cream-colored to white mesally, light brown laterally, with brown transverse stripe distally; cardo, stipes brown.
</paragraph>
<paragraph id="893F3689F61DFFCC0B8AF9461DE8D53E" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="112">All cephalic setae straight, with pointed terminus. Primary setae S1S12 present; S1, S11 long; S2, S3, S5 short. Labial palpus: basal segment with two setae—one lateral, one ventral; midsegment with two distal setaeone lateral, one mesal. Mandible with one basolateral seta; maxillae with basal seta. Palpiger with two basal setae. Scape without setae; pedicel with one small distal seta; flagellar seta slightly longer than one-half length of segment.</paragraph>
<paragraph id="893F3689F61DFFCC0B8AF8B51F4CD5AE" blockId="11.[151,1437,151,2013]" pageId="11" pageNumber="112">
<emphasis id="BBF4EA9BF61DFFCC0B8AF8B51C0CD503" box="[199,280,1879,1904]" italics="true" pageId="11" pageNumber="112">Thorax</emphasis>
(
<figureCitation id="11BB2A0CF61DFFCC0A61F8B51C92D503" box="[300,390,1879,1904]" captionStart="FIGURE 10" captionStartId="12.[151,250,1401,1423]" captionTargetBox="[151,1436,814,1379]" captionTargetId="figure@12.[151,1436,814,1379]" captionTargetPageId="12" captionText="FIGURE 10. Leucochrysa (L.) nigrilabris first instar. A. Head, dorsal; B. Head, ventral; C. Head, lateral; D. Thorax, dorsal; E. Abdominal segments 1 to 5, dorsal; F. Abdominal segments 4 - 10, dorsal; G. Body, lateral; H. Body, ventral [all, State of Rio de Janeiro, Brazil]. Abbreviations: Ax, abdominal segment number; AxRx, setal row number." httpUri="https://zenodo.org/record/248034/files/figure.png" pageId="11" pageNumber="112">Figs 10</figureCitation>
D, G, H). Notum with diffuse, reddish brown, submesal markings; Setae (LS) on lateral tubercles (LTs) dark brown; dorsal setae light brown. Legs mostly light brown to brown; trochanters, tarsi white to cream-colored; claws, empodia brown. LS gently curved throughout, with tips pointed, fine; all other setae straight to gently curved, with terminus pointed.
</paragraph>
<paragraph id="893F3689F61AFFCB0B8AFF7A1F67D28B" blockId="12.[151,1436,151,392]" pageId="12" pageNumber="113">T1: LT with two long LS, without microseta between. First primary sclerite (Sc1) light brown, large, with associated seta (S1Sc1) long, arising from small chalaza; S2Sc1 short, anterior to S1Sc1. S1 long, arising from small chalaza; S2 absent; S3, S4, S5 short.</paragraph>
<paragraph id="893F3689F61AFFCB0B8AFEE61933D3FB" blockId="12.[151,1436,151,392]" pageId="12" pageNumber="113">T2: Spiracles prominent, tubercle-shaped, apex sclerotized, dome-shaped, amber-colored. LT with three long LS. Sc1, Sc2 small; associated setae (S1Sc1, S2Sc1 and S1Sc2) very small; Sc3 intermediate-sized, light brown, with associated seta (S1Sc3) small. Posterior subsegment with pair of short setae mesal to Sc3, row of four setae posterior to Sc3 (mesal pair short, lateral pair long, arising from robust chalazae).</paragraph>
<caption id="DDFF6601F61AFFCB0BDAFD2A19A6D08E" httpUri="https://zenodo.org/record/248033/files/figure.png" pageId="12" pageNumber="113" targetBox="[151,1436,445,690]" targetPageId="12">
<paragraph id="893F3689F61AFFCB0BDAFD2A19A6D08E" blockId="12.[151,1436,712,765]" pageId="12" pageNumber="113">
<emphasis id="BBF4EA9BF61AFFCB0BDAFD2A1C07D0AD" bold="true" box="[151,275,712,734]" pageId="12" pageNumber="113">FIGURE 9.</emphasis>
Prothorax of third instar of four
<taxonomicName id="4E804D0AF61AFFCB092FFD2A1E68D0AD" box="[610,892,712,734]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF61AFFCB092FFD2A1FF7D0AD" box="[610,739,712,734]" italics="true" pageId="12" pageNumber="113">Leucochrysa</emphasis>
(
<emphasis id="BBF4EA9BF61AFFCB09BFFD2A1E67D0AD" box="[754,883,712,734]" italics="true" pageId="12" pageNumber="113">Leucochrysa</emphasis>
)
</taxonomicName>
species. A.
<emphasis id="BBF4EA9BF61AFFCB08B5FD2B1912D0AD" box="[1016,1030,713,734]" italics="true" pageId="12" pageNumber="113">L</emphasis>
. (
<emphasis id="BBF4EA9BF61AFFCB0F51FD2B1925D0AD" box="[1052,1073,713,734]" italics="true" pageId="12" pageNumber="113">L.</emphasis>
)
<taxonomicName id="4E804D0AF61AFFCB0F72FD2A19BFD0AD" box="[1087,1195,712,734]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF61AFFCB0F72FD2A19BFD0AD" box="[1087,1195,712,734]" italics="true" pageId="12" pageNumber="113">nigrilabris</emphasis>
</taxonomicName>
; B.
<emphasis id="BBF4EA9BF61AFFCB0F9AFD2B19F1D0AD" box="[1239,1253,713,734]" italics="true" pageId="12" pageNumber="113">L</emphasis>
. (
<emphasis id="BBF4EA9BF61AFFCB0FB6FD2B1804D0AD" box="[1275,1296,713,734]" italics="true" pageId="12" pageNumber="113">L.</emphasis>
)
<taxonomicName id="4E804D0AF61AFFCB0E53FD2A1863D0AD" box="[1310,1399,712,734]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="insularis">
<emphasis id="BBF4EA9BF61AFFCB0E53FD2A1863D0AD" box="[1310,1399,712,734]" italics="true" pageId="12" pageNumber="113">insularis</emphasis>
</taxonomicName>
; C.
<emphasis id="BBF4EA9BF61AFFCB0BDAFD0A1DB1D08E" box="[151,165,744,765]" italics="true" pageId="12" pageNumber="113">L</emphasis>
. (
<emphasis id="BBF4EA9BF61AFFCB0BF7FD0A1DDBD08E" box="[186,207,744,765]" italics="true" pageId="12" pageNumber="113">L.</emphasis>
)
<taxonomicName id="4E804D0AF61AFFCB0B90FD0A1C06D08E" box="[221,274,744,765]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="varia">
<emphasis id="BBF4EA9BF61AFFCB0B90FD0A1C06D08E" box="[221,274,744,765]" italics="true" pageId="12" pageNumber="113">varia</emphasis>
</taxonomicName>
; D.
<emphasis id="BBF4EA9BF61AFFCB0A72FD0A1C59D08E" box="[319,333,744,765]" italics="true" pageId="12" pageNumber="113">L</emphasis>
. (
<emphasis id="BBF4EA9BF61AFFCB0A2FFD0A1C63D08E" box="[354,375,744,765]" italics="true" pageId="12" pageNumber="113">L.</emphasis>
)
<emphasis id="BBF4EA9BF61AFFCB0AC8FD051CA4D08E" box="[389,432,743,765]" italics="true" pageId="12" pageNumber="113">boxi</emphasis>
[A, C, D, State of Rio de Janeiro, Brazil; B, North Carolina, United States].
</paragraph>
</caption>
<caption id="DDFF6601F61AFFCB0BDAFA9B1938D7BF" httpUri="https://zenodo.org/record/248034/files/figure.png" pageId="12" pageNumber="113" targetBox="[151,1436,814,1379]" targetPageId="12">
<paragraph id="893F3689F61AFFCB0BDAFA9B1938D7BF" blockId="12.[151,1436,1401,1484]" pageId="12" pageNumber="113">
<emphasis id="BBF4EA9BF61AFFCB0BDAFA9B1C0BD7FC" bold="true" box="[151,287,1401,1423]" pageId="12" pageNumber="113">FIGURE 10.</emphasis>
<taxonomicName id="4E804D0AF61AFFCB0A68FA9B1CB2D7FC" box="[293,422,1401,1423]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF61AFFCB0A68FA9B1CB2D7FC" box="[293,422,1401,1423]" italics="true" pageId="12" pageNumber="113">Leucochrysa</emphasis>
</taxonomicName>
(
<emphasis id="BBF4EA9BF61AFFCB0AF8FA981CDED7FC" box="[437,458,1402,1423]" italics="true" pageId="12" pageNumber="113">L.</emphasis>
)
<taxonomicName id="4E804D0AF61AFFCB0A95FA9B1F50D7FC" box="[472,580,1401,1423]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="12" pageNumber="113" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF61AFFCB0A95FA9B1F50D7FC" box="[472,580,1401,1423]" italics="true" pageId="12" pageNumber="113">nigrilabris</emphasis>
</taxonomicName>
first instar. A. Head, dorsal; B. Head, ventral; C. Head, lateral; D. Thorax, dorsal; E. Abdominal segments 1 to 5, dorsal; F. Abdominal segments 4-10, dorsal; G. Body, lateral; H. Body, ventral [all, State of Rio de Janeiro, Brazil].
<emphasis id="BBF4EA9BF61AFFCB0A72FA541CC1D7BF" box="[319,469,1462,1484]" italics="true" pageId="12" pageNumber="113">Abbreviations:</emphasis>
Ax, abdominal segment number; AxRx, setal row number.
</paragraph>
</caption>
<paragraph id="893F3689F61AFFCB0B8AFA171838D442" blockId="12.[151,1436,1524,2017]" pageId="12" pageNumber="113">T3: LT with three long LS. Sc1 small, with S1Sc1 very small; Sc2 light brown, with S1Sc2 small. Anterior region with pair of short setae; posterior region with four long setae arising from robust, fleshy chalazae.</paragraph>
<paragraph id="893F3689F61AFFCA0B8AF9DE1EF9D2A6" blockId="12.[151,1436,1524,2017]" lastBlockId="13.[151,1436,151,285]" lastPageId="13" lastPageNumber="114" pageId="12" pageNumber="113">
<emphasis id="BBF4EA9BF61AFFCB0B8AF9DE1C25D426" box="[199,305,1596,1621]" italics="true" pageId="12" pageNumber="113">Abdomen</emphasis>
(
<figureCitation id="11BB2A0CF61AFFCB0A0FF9DE1C8ED426" box="[322,410,1596,1621]" captionStart="FIGURE 10" captionStartId="12.[151,250,1401,1423]" captionTargetBox="[151,1436,814,1379]" captionTargetId="figure@12.[151,1436,814,1379]" captionTargetPageId="12" captionText="FIGURE 10. Leucochrysa (L.) nigrilabris first instar. A. Head, dorsal; B. Head, ventral; C. Head, lateral; D. Thorax, dorsal; E. Abdominal segments 1 to 5, dorsal; F. Abdominal segments 4 - 10, dorsal; G. Body, lateral; H. Body, ventral [all, State of Rio de Janeiro, Brazil]. Abbreviations: Ax, abdominal segment number; AxRx, setal row number." httpUri="https://zenodo.org/record/248034/files/figure.png" pageId="12" pageNumber="113">Figs 10</figureCitation>
EH). White to cream-colored; dorsum of A1A3 with diffuse, submesal, reddish brown band; dorsum of A4, A6A8 each with pair of reddish brown submesal spots. LTs each with two long LS (A2A7), without microseta between; LS straight, with pointed tip, dark brown; submedian setae (SMS), setae (LDS) on laterodorsal tubercles (A1A5) hooked, light brown. A1, A6A8: Spiracle without associated seta (SSp); A2A5: each spiracle with small to minute SSp. A1A5: Laterodorsal tubercles (LDTs) each with two long, hooked LDS, one microseta between LDS. A1: Single row of four SMS between LDTs. A2A4: Anterior row of four SMS, posterior row of two SMS, both between LDTs. A5: Anterior row of two to three SMS, posterior row of two SMS, both between LDTs. A6: Two short, straight setae anteriorly, two intermediate-length, hooked setae posteriorly; LDT with one long, hooked LDS, one short, straight LDS. A7: LDT with one intermediate-length, straight LDS, two very short LDS; dorsum with pair of minute setae anteriorly. A8: LT with one short, straight LS, two to three small setae, base of LT with spiracular tubercle projecting from anterior surface; spiracular base and sclerotized ring elongate, projecting posteriorly; dorsum with three to four short, straight setae between LTs. A9: Dorsum with three rows of short, straight setae (anterior two minute, middle four short, posterior four slightly longer). A10: Apparently without setae; terminus light brown, with small eversible pouch.
</paragraph>
<paragraph id="893F3689F61BFFCA0B8AFF3D1E5ED36E" blockId="13.[151,1436,151,285]" pageId="13" pageNumber="114">
<emphasis id="BBF4EA9BF61BFFCA0B8AFF3D1E43D28B" bold="true" box="[199,855,223,248]" pageId="13" pageNumber="114">Description (Second &amp; Third Instars, Semaphoront B).</emphasis>
<emphasis id="BBF4EA9BF61BFFCA0810FF3D1E83D28B" box="[861,919,223,248]" italics="true" pageId="13" pageNumber="114">Body</emphasis>
(
<figureCitation id="11BB2A0CF61BFFCA08EBFF3D1EFBD28B" box="[934,1007,223,248]" captionStart="FIGURE 9" captionStartId="12.[151,250,712,734]" captionTargetBox="[151,1436,445,690]" captionTargetId="figure@12.[151,1436,445,691]" captionTargetPageId="12" captionText="FIGURE 9. Prothorax of third instar of four Leucochrysa (Leucochrysa) species. A. L. (L.) nigrilabris; B. L. (L.) insularis; C. L. (L.) varia; D. L. (L.) boxi [A, C, D, State of Rio de Janeiro, Brazil; B, North Carolina, United States]." httpUri="https://zenodo.org/record/248033/files/figure.png" pageId="13" pageNumber="114">Figs 9</figureCitation>
,
<figureCitation id="11BB2A0CF61BFFCA08B6FF021954D28B" box="[1019,1088,224,248]" captionStart-0="FIGURE 11" captionStart-1="FIGURE 12" captionStartId-0="13.[151,250,1017,1039]" captionStartId-1="14.[151,250,1391,1413]" captionTargetBox-0="[151,1436,337,995]" captionTargetBox-1="[151,1436,409,1369]" captionTargetId-0="figure@13.[151,1436,337,996]" captionTargetId-1="figure@14.[151,1436,409,1370]" captionTargetPageId-0="13" captionTargetPageId-1="14" captionText-0="FIGURE 11. Leucochrysa (L.) nigrilabris second instar. A. Head, dorsal; B. Head, lateral; C. Thorax, dorsal; D. Abdominal segments 1 to 4, dorsal; E. Abdominal segments 4 10, dorsal; F. Body, lateral [all, State of Rio de Janeiro, Brazil]. Abbreviations: Ax, abdominal segment number; AxRx, setal row number." captionText-1="FIGURE 12. Leucochrysa (L.) nigrilabris third instar. A. Head, dorsal; B. Head, ventral; C. Thorax, dorsal; D. Abdominal segments 1 to 3, dorsal; E. Abdominal segments 3 7, dorsal; F. Abdominal segments 6 - 10, dorsal; G. Head, lateral; H. Body, ventral; I. Body, lateral [all, State of Rio de Janeiro, Brazil]. Abbreviations: Ax, abdominal segment number; AxRx, setal row number." httpUri-0="https://zenodo.org/record/248035/files/figure.png" httpUri-1="https://zenodo.org/record/248036/files/figure.png" pageId="13" pageNumber="114">1112</figureCitation>
). L2:
<quantity id="4E789B6CF61BFFCA0FC9FF02181AD284" box="[1156,1294,224,248]" metricMagnitude="-3" metricUnit="m" metricValue="4.8" metricValueMax="4.9" metricValueMin="4.7" pageId="13" pageNumber="114" unit="mm" value="4.8" valueMax="4.9" valueMin="4.7">4.74.9 mm</quantity>
; L3: 8.69.0 mm long. Dorsum cream-colored, with dark brown markings.
</paragraph>
<caption id="DDFF6601F61BFFCA0BDAFC1B1E97D63F" httpUri="https://zenodo.org/record/248035/files/figure.png" pageId="13" pageNumber="114" targetBox="[151,1436,337,995]" targetPageId="13">
<paragraph id="893F3689F61BFFCA0BDAFC1B1E97D63F" blockId="13.[151,1436,1017,1100]" pageId="13" pageNumber="114">
<emphasis id="BBF4EA9BF61BFFCA0BDAFC1B1C0BD67C" bold="true" box="[151,287,1017,1039]" pageId="13" pageNumber="114">FIGURE 11.</emphasis>
<taxonomicName id="4E804D0AF61BFFCA0A6AFC1B1CBCD67C" box="[295,424,1017,1039]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF61BFFCA0A6AFC1B1CBCD67C" box="[295,424,1017,1039]" italics="true" pageId="13" pageNumber="114">Leucochrysa</emphasis>
</taxonomicName>
(
<emphasis id="BBF4EA9BF61BFFCA0AF4FC181CD9D67C" box="[441,461,1018,1039]" italics="true" pageId="13" pageNumber="114">L.</emphasis>
)
<taxonomicName id="4E804D0AF61BFFCA0A90FC1B1F5DD67C" box="[477,585,1017,1039]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="13" pageNumber="114" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF61BFFCA0A90FC1B1F5DD67C" box="[477,585,1017,1039]" italics="true" pageId="13" pageNumber="114">nigrilabris</emphasis>
</taxonomicName>
second instar. A. Head, dorsal; B. Head, lateral; C. Thorax, dorsal; D. Abdominal segments 1 to 4, dorsal; E. Abdominal segments 410, dorsal; F. Body, lateral [all, State of Rio de Janeiro, Brazil].
<emphasis id="BBF4EA9BF61BFFCA0BDAFBD41C38D63F" box="[151,300,1078,1100]" italics="true" pageId="13" pageNumber="114">Abbreviations:</emphasis>
Ax, abdominal segment number; AxRx, setal row number.
</paragraph>
</caption>
<paragraph id="893F3689F61BFFCA0B8AFB961EA3D4DB" blockId="13.[151,1437,1140,2029]" pageId="13" pageNumber="114">
<emphasis id="BBF4EA9BF61BFFCA0B8AFB961C10D6FE" box="[199,260,1140,1165]" italics="true" pageId="13" pageNumber="114">Head</emphasis>
(
<figureCitation id="11BB2A0CF61BFFCA0A58FB961C7ED6FF" box="[277,362,1140,1165]" captionStart="FIGURE 11" captionStartId="13.[151,250,1017,1039]" captionTargetBox="[151,1436,337,995]" captionTargetId="figure@13.[151,1436,337,996]" captionTargetPageId="13" captionText="FIGURE 11. Leucochrysa (L.) nigrilabris second instar. A. Head, dorsal; B. Head, lateral; C. Thorax, dorsal; D. Abdominal segments 1 to 4, dorsal; E. Abdominal segments 4 10, dorsal; F. Body, lateral [all, State of Rio de Janeiro, Brazil]. Abbreviations: Ax, abdominal segment number; AxRx, setal row number." httpUri="https://zenodo.org/record/248035/files/figure.png" pageId="13" pageNumber="114">Figs 11</figureCitation>
A, B, 12A, B, G, H). L2:
<quantity id="4E789B6CF61BFFCA09D8FB961E2ED6FF" box="[661,826,1140,1164]" metricMagnitude="-4" metricUnit="m" metricValue="8.200000000000001" metricValueMax="8.299999999999999" metricValueMin="8.100000000000001" pageId="13" pageNumber="114" unit="mm" value="0.82" valueMax="0.83" valueMin="0.81">0.810.83 mm</quantity>
wide,
<quantity id="4E789B6CF61BFFCA08C7FB96193BD6FF" box="[906,1071,1140,1165]" metricMagnitude="-4" metricUnit="m" metricValue="5.549999999999999" metricValueMax="5.8" metricValueMin="5.3" pageId="13" pageNumber="114" unit="mm" value="0.5549999999999999" valueMax="0.58" valueMin="0.53">0.530.58 mm</quantity>
long; mandibles
<quantity id="4E789B6CF61BFFCA0FBAFB961888D6FF" box="[1271,1436,1140,1165]" metricMagnitude="-4" metricUnit="m" metricValue="9.45" metricValueMax="9.6" metricValueMin="9.3" pageId="13" pageNumber="114" unit="mm" value="0.945" valueMax="0.96" valueMin="0.93">0.930.96 mm</quantity>
long; ratio mandible length: head width 1.141.17:1. L3:
<quantity id="4E789B6CF61BFFCA0860FB7A1EA1D6DC" box="[813,949,1176,1200]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="13" pageNumber="114" unit="mm" value="1.35" valueMax="1.4" valueMin="1.3">1.31.4 mm</quantity>
wide,
<quantity id="4E789B6CF61BFFCA0F4FFB7519B3D6DC" box="[1026,1191,1175,1200]" metricMagnitude="-4" metricUnit="m" metricValue="8.85" metricValueMax="9.1" metricValueMin="8.6" pageId="13" pageNumber="114" unit="mm" value="0.885" valueMax="0.91" valueMin="0.86">0.860.91 mm</quantity>
long, mandibles
<quantity id="4E789B6CF61BFFCA0E26FB751DFBD6A7" metricMagnitude="-3" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="13" pageNumber="114" unit="mm" value="1.6" valueMax="1.7" valueMin="1.5">1.5 1.7 mm</quantity>
long; ratio mandible length: head width, 1.191.23:1. Epicranial marking paired, brown to dark brown, covering entire basal portion of cranium, laterally becoming lighter brown, extending to eye, mesally becoming confluent with base of postfrontal marking. Postfrontal marking slightly darker brown than epicranial marking, elongate, extending to base of antenna. Frontal marking V-shaped, arms extending forward through dark tentorial pits, then curving to base of mandibles becoming confluent with intermandibular marking. Intermandibular marking brown, divided into three sections—two lateral sections at mesal base of mandibles, mesal section between, but separated from two lateral sections; clypeolabral region partially covered by intermandibular marking. Gena with single, elongate, brown stripe extending from base of eye to posterior edge of cranium; region around eyes cream-colored; stemmata, except posteroventral margin, surrounded by brown. Labial palpus: basal segment light brown; mesal segment light brown basally, darker distally, with well defined annulations; terminal segment dark brown. Mandibles brown basally to dark brown at tip. Antenna light brown to brown. Ventrolateral cranial surface light brown. Cardo with light brown mark basally, dark brown mark distally, narrow; stipes light brown to tan, narrow, elongate. Mentum white to cream-colored, with tinge of light brown anteriorly; palpiger light brown. Cervix cream-colored, with prominent lateral patches of brown.
</paragraph>
<paragraph id="893F3689F61BFFCA0B8AF9561861D52E" blockId="13.[151,1437,1140,2029]" pageId="13" pageNumber="114">
Anterior margin of head with slightly rough, obtuse lateral edges; posterior section of cranium spiculate. All cephalic setae smooth, straight, with acute tip; S1S12 present; S1, S11 long; S2, S3, S5 short. Labial palpus: basal segment with three setae, midsegment with distal annulation bearing four setae, remaining annulations with
<emphasis id="BBF4EA9BF61BFFCA0E14F91F1860D567" box="[1369,1396,1789,1812]" italics="true" pageId="13" pageNumber="114">ca</emphasis>
six (L2) or seven (L3). Mandible with one basolateral seta; maxillae with basal seta. Palpiger with two basal setae. Scape with one distolateral seta; pedicel with one small distal seta; flagellar seta
<emphasis id="BBF4EA9BF61BFFCA0F52F8A7192ED52F" box="[1055,1082,1861,1884]" italics="true" pageId="13" pageNumber="114">ca</emphasis>
one-half length of segment.
</paragraph>
<paragraph id="893F3689F61BFFC90B8AF8851E98D28B" blockId="13.[151,1437,1140,2029]" lastBlockId="14.[151,1436,151,357]" lastPageId="14" lastPageNumber="115" pageId="13" pageNumber="114">
<emphasis id="BBF4EA9BF61BFFCA0B8AF8851C0CD5F3" box="[199,280,1895,1920]" italics="true" pageId="13" pageNumber="114">Thorax</emphasis>
(
<figureCitation id="11BB2A0CF61BFFCA0A67F8851C63D5F3" box="[298,375,1895,1920]" captionStart="FIGURE 9" captionStartId="12.[151,250,712,734]" captionTargetBox="[151,1436,445,690]" captionTargetId="figure@12.[151,1436,445,691]" captionTargetPageId="12" captionText="FIGURE 9. Prothorax of third instar of four Leucochrysa (Leucochrysa) species. A. L. (L.) nigrilabris; B. L. (L.) insularis; C. L. (L.) varia; D. L. (L.) boxi [A, C, D, State of Rio de Janeiro, Brazil; B, North Carolina, United States]." httpUri="https://zenodo.org/record/248033/files/figure.png" pageId="13" pageNumber="114">Figs 9</figureCitation>
,
<figureCitation id="11BB2A0CF61BFFCA0ACAF88A1CB6D5F3" box="[391,418,1896,1920]" captionStart="FIGURE 11" captionStartId="13.[151,250,1017,1039]" captionTargetBox="[151,1436,337,995]" captionTargetId="figure@13.[151,1436,337,996]" captionTargetPageId="13" captionText="FIGURE 11. Leucochrysa (L.) nigrilabris second instar. A. Head, dorsal; B. Head, lateral; C. Thorax, dorsal; D. Abdominal segments 1 to 4, dorsal; E. Abdominal segments 4 10, dorsal; F. Body, lateral [all, State of Rio de Janeiro, Brazil]. Abbreviations: Ax, abdominal segment number; AxRx, setal row number." httpUri="https://zenodo.org/record/248035/files/figure.png" pageId="13" pageNumber="114">11</figureCitation>
C, F, 12C, H, I). LTs white to cream-colored, with diffuse to very pale, reddish brown markings at dorsal base submesally; LS mostly dark brown; large sclerites Sc1 (prothorax), Sc3 (mesothorax), Sc2 (metathorax) light amber, shiny. Venter white to cream-colored, with small, diffuse, rosy, submesal patches on each segment. Spiracles amber to light amber; setae mostly brown or light brown. Legs: coxa with dark brown marking on dorsal connection to base, pair of brown marks, one anterior, one posterior; trochanter with pair of dark brown marks, one anterior, one posterior; femur brown dorsobasally, light brown distally; tibia brown throughout; tarsus light brown; claws curved, large, dark brown; empodia dark brown.
</paragraph>
<paragraph id="893F3689F618FFC90B8AFEE71E98D316" blockId="14.[151,1436,151,357]" pageId="14" pageNumber="115">LTs smooth; LS smooth, except very fine tips sometimes slightly salebrose, seldom sparsely and finely barbed, slender, gently curved throughout, tips tapering to very fine terminus; tips of distal LS mostly pointed, tips of lateral LS mostly with small hook. All dorsal, ventral setae smooth.</paragraph>
<caption id="DDFF6601F618FFC90BDAFA8D1DFED792" httpUri="https://zenodo.org/record/248036/files/figure.png" pageId="14" pageNumber="115" targetBox="[151,1436,409,1369]" targetPageId="14">
<paragraph id="893F3689F618FFC90BDAFA8D1DFED792" blockId="14.[151,1436,1391,1505]" pageId="14" pageNumber="115">
<emphasis id="BBF4EA9BF618FFC90BDAFA8D1C36D7F6" bold="true" box="[151,290,1391,1413]" pageId="14" pageNumber="115">FIGURE 12.</emphasis>
<taxonomicName id="4E804D0AF618FFC90A67FA8D1CB8D7F6" box="[298,428,1391,1413]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF618FFC90A67FA8D1CB8D7F6" box="[298,428,1391,1413]" italics="true" pageId="14" pageNumber="115">Leucochrysa</emphasis>
</taxonomicName>
(
<emphasis id="BBF4EA9BF618FFC90AF1FA921CC6D7F6" box="[444,466,1392,1413]" italics="true" pageId="14" pageNumber="115">L.</emphasis>
)
<taxonomicName id="4E804D0AF618FFC90AAFFA8D1F5AD7F6" box="[482,590,1391,1413]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF618FFC90AAFFA8D1F5AD7F6" box="[482,590,1391,1413]" italics="true" pageId="14" pageNumber="115">nigrilabris</emphasis>
</taxonomicName>
third instar. A. Head, dorsal; B. Head, ventral; C. Thorax, dorsal; D. Abdominal segments 1 to 3, dorsal; E. Abdominal segments 37, dorsal; F. Abdominal segments 6-10, dorsal; G. Head, lateral; H. Body, ventral; I. Body, lateral [all, State of Rio de Janeiro, Brazil].
<emphasis id="BBF4EA9BF618FFC9084EFA4E1E8CD7B1" box="[771,920,1452,1474]" italics="true" pageId="14" pageNumber="115">Abbreviations:</emphasis>
Ax, abdominal segment number; AxRx, setal row number.
</paragraph>
</caption>
<paragraph id="893F3689F618FFC90B8AF9EC1E6CD4E1" blockId="14.[151,1436,1549,2006]" pageId="14" pageNumber="115">
T1: All dorsal, ventral setae straight; Sc1 large, extending laterally to dorsal surface of LT, with S1Sc1 long, slender, S2Sc1 very short, slender. Sc2 transparent, oval-shaped,
<emphasis id="BBF4EA9BF618FFC90830F9D11E8CD439" box="[893,920,1587,1610]" italics="true" pageId="14" pageNumber="115">ca</emphasis>
one-third length of Sc1. LT (L2) with 1921 LS (
<emphasis id="BBF4EA9BF618FFC90B87F9B51DF1D41D" box="[202,229,1623,1646]" italics="true" pageId="14" pageNumber="115">ca</emphasis>
10 apical,
<emphasis id="BBF4EA9BF618FFC90A28F9B51C94D41D" box="[357,384,1623,1646]" italics="true" pageId="14" pageNumber="115">ca</emphasis>
911 lateral); LT (L3) with
<emphasis id="BBF4EA9BF618FFC9098AF9B51FF6D41D" box="[711,738,1623,1646]" italics="true" pageId="14" pageNumber="115">ca</emphasis>
2730 LS (
<emphasis id="BBF4EA9BF618FFC90820F9B51E9CD41D" box="[877,904,1623,1646]" italics="true" pageId="14" pageNumber="115">ca</emphasis>
10 apical,
<emphasis id="BBF4EA9BF618FFC90F45F9B51936D41D" box="[1032,1058,1623,1646]" italics="true" pageId="14" pageNumber="115">ca</emphasis>
1720 lateral). Primary setae S1 medium-length, S2, S3, S4, S5 short, slender; no secondary setae.
</paragraph>
<paragraph id="893F3689F618FFC90B8AF97C1C2FD5A5" blockId="14.[151,1436,1549,2006]" pageId="14" pageNumber="115">
T2: Anterior subsegment without secondary setae between spiracles; anterior sclerite (Sc1) with three associated setae (S1Sc1, S2Sc1, S3Sc1); spiracles prominent, with opening sclerotized, dome-shaped, ambercolored ring on small tubercle; walls of chamber parallel, not bulbous; SSp absent. Posterior subsegment with anterior sclerite (Sc2) bearing two very small associated setae (S1Sc2, S2Sc2); Sc3 with small associated seta (S1Sc3); anterior row with two intermediate-length, slender, straight setae; posterior row (L2) with
<emphasis id="BBF4EA9BF618FFC90E40F8CD183CD535" box="[1293,1320,1839,1862]" italics="true" pageId="14" pageNumber="115">ca</emphasis>
56 setae (34 long, robust, 12 slender, hooked or straight), arising from large, plump chalazae; posterior row (L3) with
<emphasis id="BBF4EA9BF618FFC90ECCF8B1188FD519" box="[1409,1435,1875,1898]" italics="true" pageId="14" pageNumber="115">ca</emphasis>
1114 setae (68 long, robust, 57 slender, hooked or straight), arising from large, plump chalazae. LT (L2) bearing
<emphasis id="BBF4EA9BF618FFC90BBEF8791C1AD5C1" box="[243,270,1947,1970]" italics="true" pageId="14" pageNumber="115">ca</emphasis>
1719 LS (
<emphasis id="BBF4EA9BF618FFC90AD8F8791CA4D5C1" box="[405,432,1947,1970]" italics="true" pageId="14" pageNumber="115">ca</emphasis>
89 apical,
<emphasis id="BBF4EA9BF618FFC90976F8791F42D5C1" box="[571,598,1947,1970]" italics="true" pageId="14" pageNumber="115">ca</emphasis>
910 lateral, mostly straight); LT (L3) bearing
<emphasis id="BBF4EA9BF618FFC90F3CF8791998D5C1" box="[1137,1164,1947,1970]" italics="true" pageId="14" pageNumber="115">ca</emphasis>
26 LS (
<emphasis id="BBF4EA9BF618FFC90FA4F8791810D5C1" box="[1257,1284,1947,1970]" italics="true" pageId="14" pageNumber="115">ca</emphasis>
10 apical,
<emphasis id="BBF4EA9BF618FFC90ECCF879188FD5C1" box="[1409,1435,1947,1970]" italics="true" pageId="14" pageNumber="115">ca</emphasis>
1517 lateral).
</paragraph>
<paragraph id="893F3689F619FFC80B8AFF7A1E93D316" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="116">
T3: Sclerites (Sc1, Sc2) each with one associated setae (S1Sc1, S1Sc2); sometimes small seta on anterior fold, lateral to Sc1. Anterior row with one to two short (L2) or two to four intermediate-length (L3), slender, straight setae; posterior row (L2) with
<emphasis id="BBF4EA9BF619FFC80AB0FF031F0CD28B" box="[509,536,225,248]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
12 long, robust, straight or hooked setae arising from large, plump chalazae; posterior row (L3) with
<emphasis id="BBF4EA9BF619FFC80AFAFEE71CC5D36F" box="[439,465,261,284]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
1516 setae (
<emphasis id="BBF4EA9BF619FFC80931FEE71F83D36F" box="[636,663,261,284]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
~1214 long, robust, 24 intermediate-length, slender, straight or hooked), arising from large, plump chalazae. LT (L2) bearing
<emphasis id="BBF4EA9BF619FFC80820FECB1E9CD333" box="[877,904,297,320]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
1920 LS (
<emphasis id="BBF4EA9BF619FFC80F5AFECB1926D333" box="[1047,1074,297,320]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
9 apical, 1011 lateral, mostly straight). LT (L3) bearing
<emphasis id="BBF4EA9BF619FFC80AF1FEAF1CC3D317" box="[444,471,333,356]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
2125 LS (
<emphasis id="BBF4EA9BF619FFC80911FEAF1F63D317" box="[604,631,333,356]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
9 apical, 1216 lateral).
</paragraph>
<paragraph id="893F3689F619FFC80B8AFE8D1E2AD386" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="116">
<emphasis id="BBF4EA9BF619FFC80B8AFE8D1C25D3FB" box="[199,305,367,392]" italics="true" pageId="15" pageNumber="116">Abdomen</emphasis>
(
<figureCitation id="11BB2A0CF619FFC80A72FE8D1C9BD3FB" box="[319,399,367,392]" captionStart="FIGURE 11" captionStartId="13.[151,250,1017,1039]" captionTargetBox="[151,1436,337,995]" captionTargetId="figure@13.[151,1436,337,996]" captionTargetPageId="13" captionText="FIGURE 11. Leucochrysa (L.) nigrilabris second instar. A. Head, dorsal; B. Head, lateral; C. Thorax, dorsal; D. Abdominal segments 1 to 4, dorsal; E. Abdominal segments 4 10, dorsal; F. Body, lateral [all, State of Rio de Janeiro, Brazil]. Abbreviations: Ax, abdominal segment number; AxRx, setal row number." httpUri="https://zenodo.org/record/248035/files/figure.png" pageId="15" pageNumber="116">Figs 11</figureCitation>
DF, 12DF, H, I). Dorsum, venter white to cream-colored; anterior segments without dorsal markings; A6A8 with diffuse, reddish dorsal marks; A10 with small, brownish mesal spot distally. Venter of A2 A4 with diffuse, light reddish sublateral patches; A10 with pair of brown, submesal spots. LS on A2A3 light brown, A4A7 dark brown; SMS light brown to dark amber.
</paragraph>
<paragraph id="893F3689F619FFC80B8AFE1D1C92D0F6" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="116">All setae smooth. SMS of A1A6 hooked, slender, arising from plump chalazae; SMS longest, chalazae largest on A1, A2, becoming smaller posteriorly. A1A5: LDT plump, with two long, slender, hooked LDS, one microseta between LDS. A1A8: Spiracle small, slightly raised, dome-like, with minute to very small associated seta (SSp?) anteriorly or mesally.</paragraph>
<paragraph id="893F3689F619FFC80B8AFD72197CD0D4" blockId="15.[151,1437,151,2013]" box="[199,1128,655,680]" pageId="15" pageNumber="116">
A1: Single transverse row of eight (L2) or
<emphasis id="BBF4EA9BF619FFC809EBFD731FD5D0DB" box="[678,705,657,680]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
nine to ten (L3) SMS mesal to LDTs.
</paragraph>
<paragraph id="893F3689F619FFC80B8AFD5619EAD144" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="116">
A2A3: LT broad, plump, fleshy, with seven (L2) or eight (L3) long, robust LS arranged transversely along dorsal surface,
<emphasis id="BBF4EA9BF619FFC80A0FFD3B1C49D083" box="[322,349,729,752]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
two (L2) or four (L3) shorter, straight setae basally. L2: Dorsum with anterior row of four SMS, mesal row of four, posterior row of four (A2) or six (A3) between LDTs. L3: Dorsum with anterior row of
<emphasis id="BBF4EA9BF619FFC80E1AFD1F1865D167" box="[1367,1393,765,788]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
ten (A2) or ~12 (A3) SMS, mesal row of four (A2, A3), posterior row of
<emphasis id="BBF4EA9BF619FFC808EFFCC31EA9D14B" box="[930,957,801,824]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
12 (A2, A3) between LDTs.
</paragraph>
<paragraph id="893F3689F619FFC80B8AFCA61CBFD19F" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="116">
A4A5: LT tuberculiform, with
<emphasis id="BBF4EA9BF619FFC80970FCA71F4CD12F" box="[573,600,837,860]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
five (L2) or 12 (A4, L3), 910 (A5, L3) robust, hooked or straight LS apically, dorsally,
<emphasis id="BBF4EA9BF619FFC80A29FC8B1C6BD1F3" box="[356,383,873,896]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
four (L2, L3) short, straight LS basally. LDS intermediate length, hooked. L2: Dorsum with anterior row of four (L2) intermediate-length SMS, mesal row of four, posterior row of four between LDTs. L3: Dorsum with anterior row of
<emphasis id="BBF4EA9BF619FFC80A90FC531CECD1BB" box="[477,504,945,968]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
12 (A4) or
<emphasis id="BBF4EA9BF619FFC80937FC531F81D1BB" box="[634,661,945,968]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
nine (A5) intermediate-length SMS, mesal row of four, posterior row of
<emphasis id="BBF4EA9BF619FFC80BF8FC371DC4D19F" box="[181,208,981,1004]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
ten between LDTs.
</paragraph>
<paragraph id="893F3689F619FFC80B8AFC151F76D62B" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="116">
A6: LT tuberculiform, with
<emphasis id="BBF4EA9BF619FFC80946FC1B1F32D663" box="[523,550,1017,1040]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
five (L2) or six (L3) robust, straight LS apically,
<emphasis id="BBF4EA9BF619FFC80F23FC1B199DD663" box="[1134,1161,1017,1040]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
three short, straight LS basally. LDT with one long, hooked LDS, two to three short, straight LDS. Anterior row of three (L2) or five (L3) intermediate-length SMS, no other SMS.
</paragraph>
<paragraph id="893F3689F619FFC80B8AFB861FA0D6B6" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="116">
A7: LT tuberculiform with posterior projection, with
<emphasis id="BBF4EA9BF619FFC80865FB871E57D60F" box="[808,835,1125,1148]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
five (L2) or six (L3) robust, straight LS apically,
<emphasis id="BBF4EA9BF619FFC80ECCFB87188FD60F" box="[1409,1435,1125,1148]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
three short, straight LS basally. LDT with one intermediate-length, straight LDS, one to two short, straight LDS. Two pairs of very small setae between spiracles.
</paragraph>
<paragraph id="893F3689F619FFC80B8AFB3219F1D77F" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="116">
A8: LT with four to five short, straight LS (two short, two to three very short). Row of four to five very short setae between spiracles. Posterior margin with
<emphasis id="BBF4EA9BF619FFC809E9FB171FAAD77F" box="[676,702,1269,1292]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
four (L2) or seven to eight (L3) very short setae.
</paragraph>
<paragraph id="893F3689F619FFC80B8AFAFA1E0AD75C" blockId="15.[151,1437,151,2013]" box="[199,798,1303,1328]" pageId="15" pageNumber="116">
A9: Two transverse rows of
<emphasis id="BBF4EA9BF619FFC80949FAFB1F0BD743" box="[516,543,1305,1328]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
four to six short setae.
</paragraph>
<paragraph id="893F3689F619FFC80B8AFADE1FE9D727" blockId="15.[151,1437,151,2013]" box="[199,765,1340,1365]" pageId="15" pageNumber="116">
A10: Dorsum with
<emphasis id="BBF4EA9BF619FFC80AEDFADF1CAFD727" box="[416,443,1341,1364]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
four to five very short setae.
</paragraph>
<paragraph id="893F3689F619FFC80B8AFABD1825D4EB" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="116">
<emphasis id="BBF4EA9BF619FFC80B8AFABD1F3ED70B" bold="true" box="[199,554,1375,1400]" pageId="15" pageNumber="116">Intraspecific variation (larva).</emphasis>
Three variable features warrant notation. (i) As with the larvae of all chrysopids we have studied,
<emphasis id="BBF4EA9BF619FFC80A15FA671C7AD7EF" box="[344,366,1413,1436]" italics="true" pageId="15" pageNumber="116">L.</emphasis>
(
<emphasis id="BBF4EA9BF619FFC80A33FA671C81D7EF" box="[382,405,1413,1436]" italics="true" pageId="15" pageNumber="116">L.</emphasis>
)
<taxonomicName id="4E804D0AF619FFC80AE8FA661F0FD7EE" box="[421,539,1412,1437]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF619FFC80AE8FA661F0FD7EE" box="[421,539,1412,1437]" italics="true" pageId="15" pageNumber="116">nigrilabris</emphasis>
</taxonomicName>
larvae exhibit considerable bilateral asymmetry in the numbers and size of setae across the dorsum of body. Odd numbers of setae are common. (ii) The left side of the first abdominal segment (A1) of one L
<specimenCount id="9F86FD00F619FFC80BBAFA2E1C67D796" box="[247,371,1484,1509]" pageId="15" pageNumber="116" type="generic">3 specimen</specimenCount>
has a fleshy, lateral protuberance (LT?) bearing an elongate, robust, hooked seta. This feature is unusual because LTs rarely occur on the A1 (e.g., in some
<taxonomicName id="4E804D0AF619FFC8086CFA0D1E99D47B" box="[801,909,1519,1544]" class="Insecta" family="Chrysopidae" genus="Chrysopa" kingdom="Animalia" order="Neuroptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF619FFC8086CFA0D1E99D47B" box="[801,909,1519,1544]" italics="true" pageId="15" pageNumber="116">Chrysopa</emphasis>
</taxonomicName>
spp., see
<collectingCountry id="F1977619F619FFC808B7FA0D1946D47B" box="[1018,1106,1519,1544]" name="Sao Tome and Principe" pageId="15" pageNumber="116">Principi</collectingCountry>
1940, Tsukaguchi 1978), and they have not been reported for leucochrysine larvae. Moreover, LTs almost always bear more than one elongate seta. Thus, we do not know whether this structure is homologous to those on the larvas subsequent segments (A2 A7) and/or to those on the
<taxonomicName id="4E804D0AF619FFC80A83F9BE1F2ED406" box="[462,570,1628,1653]" class="Insecta" family="Chrysopidae" genus="Chrysopa" kingdom="Animalia" order="Neuroptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="genus">
<emphasis id="BBF4EA9BF619FFC80A83F9BE1F2ED406" box="[462,570,1628,1653]" italics="true" pageId="15" pageNumber="116">Chrysopa</emphasis>
</taxonomicName>
A1. (iii) Among the first instars, the elongate setae (LS) on the apex of the lateral tubercles are mostly very long, straight, and tapering to pointed tips; occasionally some are hooked.
</paragraph>
<paragraph id="893F3689F619FFC80B8AF9461925D55B" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="116">
<emphasis id="BBF4EA9BF619FFC80B8AF9461C90D4CE" bold="true" box="[199,388,1700,1725]" pageId="15" pageNumber="116">Biological notes</emphasis>
. Adults were collected in disturbed habitats (e.g., coffee plantation, roadside, along trail) at the edges of mature forests. One female specimen, collected in the State of Rio de Janeiro, was maintained in the laboratory for
<emphasis id="BBF4EA9BF619FFC80A73F90F1C4DD577" box="[318,345,1773,1796]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
two weeks. During that time she laid two batches of eggs. Each batch consisted of a hanging cluster of
<emphasis id="BBF4EA9BF619FFC80A4AF8F31C36D55B" box="[263,290,1809,1832]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
15
<specimenCount id="9F86FD00F619FFC80A1CF8ED1C7AD55B" box="[337,366,1807,1832]" pageId="15" pageNumber="116" type="egg">20</specimenCount>
eggs attached to the ends of a series of long intertwined stalks.
</paragraph>
<paragraph id="893F3689F619FFC80B8AF8D61F26D5AE" blockId="15.[151,1437,151,2013]" pageId="15" pageNumber="116">
The
<emphasis id="BBF4EA9BF619FFC80BB6F8D71C05D53F" box="[251,273,1845,1868]" italics="true" pageId="15" pageNumber="116">L.</emphasis>
(
<emphasis id="BBF4EA9BF619FFC80A6EF8D71C2DD53F" box="[291,313,1845,1868]" italics="true" pageId="15" pageNumber="116">L.</emphasis>
)
<taxonomicName id="4E804D0AF619FFC80A06F8D61CD7D53E" box="[331,451,1844,1869]" class="Insecta" family="Chrysopidae" genus="Leucochrysa" kingdom="Animalia" order="Neuroptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="nigrilabris">
<emphasis id="BBF4EA9BF619FFC80A06F8D61CD7D53E" box="[331,451,1844,1869]" italics="true" pageId="15" pageNumber="116">nigrilabris</emphasis>
</taxonomicName>
eggs were maintained at room temperature; hatching occurred within
<emphasis id="BBF4EA9BF619FFC80FACF8D719E8D53F" box="[1249,1276,1845,1868]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
ten days. The larvae applied and carried dense packets of trash [in this case, frozen &amp; thawed eggs of
<taxonomicName id="4E804D0AF619FFC80FF4F8B51DFBD5E6" authority="Zeller" authorityName="Zeller" class="Insecta" family="Pyralidae" genus="Ephestia" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="kuehniella">
<emphasis id="BBF4EA9BF619FFC80FF4F8B5188FD503" box="[1209,1435,1879,1904]" italics="true" pageId="15" pageNumber="116">Ephestia kuehniella</emphasis>
(Zeller)
</taxonomicName>
] on their dorsa. The first moult occurred
<emphasis id="BBF4EA9BF619FFC80986F89F1FF2D5E7" box="[715,742,1917,1940]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
six days after hatching, and the second moult
<emphasis id="BBF4EA9BF619FFC80FB0F89F180CD5E7" box="[1277,1304,1917,1940]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
seven days after the first; cocoons were spun
<emphasis id="BBF4EA9BF619FFC80964F8431F50D5CB" box="[553,580,1953,1976]" italics="true" pageId="15" pageNumber="116">ca</emphasis>
10 days later. Four pharate adults emerged from their cocoons; only one completed ecdysis to the adult stage.
</paragraph>
<paragraph id="893F3689F606FFD70B8AFF751912D083" blockId="16.[151,1436,151,752]" pageId="16" pageNumber="117">
<emphasis id="BBF4EA9BF606FFD70B8AFF751880D2C3" bold="true" box="[199,1428,151,176]" pageId="16" pageNumber="117">
Specimens examined (adults and larvae, in addition to the
<typeStatus id="563B882BF606FFD708D5FF751EDFD2C3" box="[920,971,151,176]" pageId="16" pageNumber="117">type</typeStatus>
listed above).
<collectingCountry id="F1977619F606FFD70F33FF75183AD2C3" box="[1150,1326,151,176]" name="Venezuela" pageId="16" pageNumber="117">VENEZUELA</collectingCountry>
.
<emphasis id="BBF4EA9BF606FFD70E70FF7A1880D2C3" bold="true" box="[1341,1428,152,176]" italics="true" pageId="16" pageNumber="117">Aragua</emphasis>
</emphasis>
: Parque Nacional Henri Pittier (formerly Parque Nacional Rancho Grande),
<quantity id="4E789B6CF606FFD708A8FF5E192FD2A7" box="[997,1083,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="16" pageNumber="117" unit="m" value="1100.0">1100 m</quantity>
,
<date id="FD3E1049F606FFD70F0AFF5E19F0D2A6" box="[1095,1252,188,213]" pageId="16" pageNumber="117" value="1969-02-11" valueMax="1969-02-13" valueMin="1969-02-11">1113.ii.1969</date>
, J. Salcedo (
<specimenCount id="9F86FD00F606FFD70E3EFF5E1883D2A7" box="[1395,1431,188,212]" pageId="16" pageNumber="117" type="generic">1♂</specimenCount>
, MIZA); idem,
<date id="FD3E1049F606FFD70A0CFF3D1CA6D28B" box="[321,434,223,248]" pageId="16" pageNumber="117" value="1967-05-22">22.v.1967</date>
, L. Rodriguez V. (1♀, MIZA, determined by Adams 1979); idem,
<date id="FD3E1049F606FFD70FFEFF3D180AD28B" box="[1203,1310,223,248]" pageId="16" pageNumber="117" value="1990-11-09">9.xi.1990</date>
, V. Savini, trampa de luz (
<specimenCount id="9F86FD00F606FFD70A06FEE61C7BD36F" box="[331,367,260,284]" pageId="16" pageNumber="117" type="generic">1♂</specimenCount>
, MIZA); idem,
<date id="FD3E1049F606FFD70961FEE61F8CD36E" box="[556,664,260,285]" pageId="16" pageNumber="117" value="1958-06-05">5.vi.1958</date>
, F. Fernández Y. (MIZA, determined by Adams 1979]; idem,
<date id="FD3E1049F606FFD70E3FFEE61C1DD333" pageId="16" pageNumber="117" value="1966-10-16" valueMax="1966-10-31" valueMin="1966-10-16">16 31.x.1966</date>
, S. S. &amp; W. D. Duckworth (5♀, USNM); idem,
<date id="FD3E1049F606FFD7087BFEC51EC6D333" box="[822,978,295,320]" pageId="16" pageNumber="117" value="1969-10-10" valueMax="1969-10-21" valueMin="1969-10-10">1021.x.1969</date>
, Duckworth &amp; Dietz (
<specimenCount id="9F86FD00F606FFD70F98FECA19EDD333" box="[1237,1273,296,320]" pageId="16" pageNumber="117" type="generic">1♂</specimenCount>
, 1♀, USNM); idem,
<date id="FD3E1049F606FFD70BAFFEAE1C5AD316" box="[226,334,332,357]" pageId="16" pageNumber="117" value="1967-06-08">8.vi.1967</date>
, R. W. Poole (2♀, CUIC); idem,
<date id="FD3E1049F606FFD709A8FEAE1E75D316" box="[741,865,332,357]" pageId="16" pageNumber="117" value="1967-08-05">5.viii.1967</date>
, R. W. Poole (
<specimenCount id="9F86FD00F606FFD70F55FEAE1928D317" box="[1048,1084,332,356]" pageId="16" pageNumber="117" type="generic">1♂</specimenCount>
, CUIC); idem,
<date id="FD3E1049F606FFD70FBAFEAE1866D316" box="[1271,1394,332,357]" pageId="16" pageNumber="117" value="1973-09-20">20.ix.1973</date>
, B. Villegas (
<specimenCount id="9F86FD00F606FFD70A4AFE921C3FD3FB" box="[263,299,368,392]" pageId="16" pageNumber="117" type="generic">1♂</specimenCount>
, UCD); idem,
<date id="FD3E1049F606FFD70A97FE8D1F64D3FB" box="[474,624,367,392]" pageId="16" pageNumber="117" value="1996-01-24" valueMax="1996-01-25" valueMin="1996-01-24">2425.i.1996</date>
, white light, J. &amp;. A. Skevington (
<specimenCount id="9F86FD00F606FFD70F4AFE921938D3FB" box="[1031,1068,368,392]" pageId="16" pageNumber="117" type="generic">2♂</specimenCount>
, 2♀, DEBU); idem,
<quantity id="4E789B6CF606FFD70E6DFE8D186DD3F4" box="[1312,1401,367,392]" metricMagnitude="3" metricUnit="m" metricValue="1.15" pageId="16" pageNumber="117" unit="m" value="1150.0">1150 m</quantity>
, at lights on bldg,
<date id="FD3E1049F606FFD70A0BFE761F16D3DE" box="[326,514,404,429]" pageId="16" pageNumber="117" value="1996-02-24" valueMax="1996-03-12" valueMin="1996-02-24">24.ii12.iii.1996</date>
, S.
<collectingCountry id="F1977619F606FFD7097DFE761F80D3DE" box="[560,660,404,429]" name="Marshall Islands" pageId="16" pageNumber="117">Marshall</collectingCountry>
(
<specimenCount id="9F86FD00F606FFD709E5FE761FD8D3DF" box="[680,716,404,428]" pageId="16" pageNumber="117" type="generic">1♂</specimenCount>
, 1♀, DEBU); El Limón,
<quantity id="4E789B6CF606FFD708BFFE761929D3DF" box="[1010,1085,404,429]" metricMagnitude="2" metricUnit="m" metricValue="4.5" pageId="16" pageNumber="117" unit="m" value="450.0">450 m</quantity>
,
<date id="FD3E1049F606FFD70F06FE7619D0D3DE" box="[1099,1220,404,429]" pageId="16" pageNumber="117" value="1964-04-20">20.iv.1964</date>
, E. Osuna (MIZA, determined by Adams 1979);
<emphasis id="BBF4EA9BF606FFD70AA9FE551F2CD3A3" bold="true" box="[484,568,439,464]" italics="true" pageId="16" pageNumber="117">Mérida</emphasis>
: Mérida, el. ca
<quantity id="4E789B6CF606FFD709A5FE551E29D3A3" box="[744,829,439,464]" metricMagnitude="3" metricUnit="m" metricValue="1.3716" pageId="16" pageNumber="117" unit="ft" value="4500.0">4500 ft</quantity>
.,
<geoCoordinate id="ECB4504EF606FFD70801FE551EADD3A3" box="[844,953,439,464]" direction="north" orientation="latitude" pageId="16" pageNumber="117" precision="5" value="8.5942">8.5942°N</geoCoordinate>
/
<geoCoordinate id="ECB4504EF606FFD70883FE5A195BD3A3" box="[974,1103,440,464]" direction="west" orientation="longitude" pageId="16" pageNumber="117" precision="5" value="-71.1429">71.1429°W</geoCoordinate>
,
<date id="FD3E1049F606FFD70F17FE5519EDD3A3" box="[1114,1273,439,464]" pageId="16" pageNumber="117" value="1994-02-22" valueMax="1994-02-23" valueMin="1994-02-22">2223.ii.1994</date>
, J. P. Donahue (1♀, LACM);
<emphasis id="BBF4EA9BF606FFD70A09FE3E1CDCD380" bold="true" box="[324,456,476,499]" italics="true" pageId="16" pageNumber="117">Portuguesa</emphasis>
: Córdoba
<quantity id="4E789B6CF606FFD70904FE3E1F83D387" box="[585,663,476,500]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="16" pageNumber="117" unit="m" value="800.0">800 m</quantity>
,
<date id="FD3E1049F606FFD709E5FE3E1E58D386" box="[680,844,476,501]" pageId="16" pageNumber="117" value="2008-11-24" valueMax="2008-11-29" valueMin="2008-11-24">2429.xi.2008</date>
, R. Páz, Malaise Trap (1♀, UCOB);
<emphasis id="BBF4EA9BF606FFD70E41FE3E187ED380" bold="true" box="[1292,1386,476,499]" italics="true" pageId="16" pageNumber="117">Yaracuy</emphasis>
: La Cumbre, Chivacoa Council,
<quantity id="4E789B6CF606FFD70A95FE1D1F24D064" box="[472,560,511,536]" metricMagnitude="3" metricUnit="m" metricValue="1.075" pageId="16" pageNumber="117" unit="m" value="1075.0">1075 m</quantity>
,
<geoCoordinate id="ECB4504EF606FFD70971FDE21F8ED06B" box="[572,666,512,536]" direction="north" orientation="latitude" pageId="16" pageNumber="117" precision="925" value="10.233334">10º14N</geoCoordinate>
/
<geoCoordinate id="ECB4504EF606FFD709FCFE1D1E02D06B" box="[689,790,511,536]" direction="west" orientation="longitude" pageId="16" pageNumber="117" precision="925" value="-68.96667">68º58W</geoCoordinate>
,
<date id="FD3E1049F606FFD7086FFE1D1EB0D06B" box="[802,932,511,536]" pageId="16" pageNumber="117" value="2012-07-22">22.vii.2012</date>
, in
<taxonomicName id="4E804D0AF606FFD70880FDE219CDD06B" authority="F. Sosa" authorityName="F. Sosa" box="[973,1241,511,536]" class="Magnoliopsida" family="Rubiaceae" genus="Coffea" kingdom="Plantae" order="Gentianales" pageId="16" pageNumber="117" phylum="Tracheophyta" rank="species" species="arabica">
<emphasis id="BBF4EA9BF606FFD70880FDE21963D06B" box="[973,1143,511,536]" italics="true" pageId="16" pageNumber="117">Coffea arabica</emphasis>
, F. Sosa
</taxonomicName>
, manual net (1♀, UCOB); idem,
<quantity id="4E789B6CF606FFD70A0EFDC61C8ED04F" box="[323,410,548,572]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="16" pageNumber="117" unit="m" value="1200.0">1200 m</quantity>
,
<geoCoordinate id="ECB4504EF606FFD70AE8FDC61F10D04E" box="[421,516,548,573]" direction="north" orientation="latitude" pageId="16" pageNumber="117" precision="925" value="10.266666">10º16N</geoCoordinate>
/
<geoCoordinate id="ECB4504EF606FFD70957FDC61F6AD04E" box="[538,638,548,573]" direction="west" orientation="longitude" pageId="16" pageNumber="117" precision="925" value="-68.933334">68º56W</geoCoordinate>
,
<date id="FD3E1049F606FFD709C7FDC61FE9D04E" box="[650,765,548,573]" pageId="16" pageNumber="117" value="2013-02-11">11.ii.2013</date>
, in
<taxonomicName id="4E804D0AF606FFD70868FDC6193BD04F" authority="F. Sosa" authorityName="F. Sosa" box="[805,1071,548,573]" class="Magnoliopsida" family="Rubiaceae" genus="Coffea" kingdom="Plantae" order="Gentianales" pageId="16" pageNumber="117" phylum="Tracheophyta" rank="species" species="arabica">
<emphasis id="BBF4EA9BF606FFD70868FDC61ED9D04E" box="[805,973,548,573]" italics="true" pageId="16" pageNumber="117">Coffea arabica</emphasis>
, F. Sosa
</taxonomicName>
, manual net (
<specimenCount id="9F86FD00F606FFD70F8BFDC619FFD04F" box="[1222,1259,548,572]" pageId="16" pageNumber="117" type="generic">1♂</specimenCount>
, UCOB); idem,
<date id="FD3E1049F606FFD70BDAFDA51C1BD013" box="[151,271,583,608]" pageId="16" pageNumber="117" value="2013-04-18">18.iv.2013</date>
, in
<taxonomicName id="4E804D0AF606FFD70A7AFDAA1E59D013" authority="F. Sosa, C. Martins and F. Diaz" authorityName="F. Sosa, C. Martins and F. Diaz" box="[311,845,583,608]" class="Magnoliopsida" family="Rubiaceae" genus="Coffea" kingdom="Plantae" order="Gentianales" pageId="16" pageNumber="117" phylum="Tracheophyta" rank="species" species="arabica">
<emphasis id="BBF4EA9BF606FFD70A7AFDAA1CCBD013" box="[311,479,583,608]" italics="true" pageId="16" pageNumber="117">Coffea arabica</emphasis>
, F. Sosa, C. Martins and F. Díaz
</taxonomicName>
, manual net (
<specimenCount id="9F86FD00F606FFD708AEFDAA191CD013" box="[995,1032,584,608]" pageId="16" pageNumber="117" type="generic">2♂</specimenCount>
, 1♀, UCOB); El Candelo, Cocorote Council,
<quantity id="4E789B6CF606FFD70A4FFD8E1C49D0F7" box="[258,349,620,644]" metricMagnitude="3" metricUnit="m" metricValue="1.309" pageId="16" pageNumber="117" unit="m" value="1309.0">1309 m</quantity>
,
<geoCoordinate id="ECB4504EF606FFD70A20FD8E1CDED0F7" box="[365,458,620,644]" direction="north" orientation="latitude" pageId="16" pageNumber="117" precision="925" value="10.333333">10º20N</geoCoordinate>
/
<geoCoordinate id="ECB4504EF606FFD70AA5FD8E1F59D0F6" box="[488,589,620,645]" direction="west" orientation="longitude" pageId="16" pageNumber="117" precision="925" value="-68.816666">68º49W</geoCoordinate>
,
<date id="FD3E1049F606FFD70910FD8E1FC1D0F6" box="[605,725,620,645]" pageId="16" pageNumber="117" value="2013-04-18">18.iv.2013</date>
, in forest, F. Sosa, C. Martins and F. Díaz, manual net (
<specimenCount id="9F86FD00F606FFD70E3EFD8E1883D0F7" box="[1395,1431,620,644]" pageId="16" pageNumber="117" type="generic">1♂</specimenCount>
, UCOB).
<emphasis id="BBF4EA9BF606FFD70BB6FD6D1F32D0DB" bold="true" box="[251,550,655,680]" pageId="16" pageNumber="117">
<collectingCountry id="F1977619F606FFD70BB6FD6D1C7CD0DB" box="[251,360,655,680]" name="Brazil" pageId="16" pageNumber="117">BRAZIL</collectingCountry>
.
<emphasis id="BBF4EA9BF606FFD70A39FD6D1F09D0DB" bold="true" box="[372,541,655,680]" italics="true" pageId="16" pageNumber="117">Rio de Janeiro</emphasis>
:
</emphasis>
Sta. M. Madalena, Terras Frias,
<date id="FD3E1049F606FFD708DCFD721917D0DB" box="[913,1027,656,680]" pageId="16" pageNumber="117" value="2003-10-28">28.x.2003</date>
, G. S. Albuquerque, M. J. Tauber, C. A. Tauber Expedition,
<date id="FD3E1049F606FFD70AECFD561F42D0BF" box="[417,598,692,716]" pageId="16" pageNumber="117" value="2003-10" valueMax="2003-11-31" valueMin="2003-10-00">Oct.Nov. 2003</date>
[1♀, 3?sex (lab-reared, pharate adults, incompletely emerged),
<specimenCount id="9F86FD00F606FFD70E72FD56188FD0BE" box="[1343,1435,692,717]" pageId="16" pageNumber="117" type="larva">9 larvae</specimenCount>
(Tauber Lot 2003:067, UCB);
<specimenCount id="9F86FD00F606FFD70AA1FD351F47D083" box="[492,595,727,752]" pageId="16" pageNumber="117" type="larva">15 larvae</specimenCount>
(Albuquerque Lot 2003:021, UENF)].
</paragraph>
</subSubSection>
</treatment>
</document>