227 lines
28 KiB
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227 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/asp.81.e101374" ID-Pensoft-Pub="1864-8312-81-689" ID-Pensoft-UUID="B61E7B03B308528C9FC150520EABBE19" ID-ZooBank="944B0964DA564F9B926A940B392C5AC7" ModsDocID="1864-8312-81-689" checkinTime="1690954509890" checkinUser="pensoft" docAuthor="Rosova, Katerina, Prokop, Jakub, Hammel, Joerg U. & Beutel, Rolf G." docDate="2023" docId="DEDF00746A035664A1B7334CF93F5770" docLanguage="en" docName="ArthropodSystPhyl 81: 689-704" docOrigin="Arthropod Systematics & amp; Phylogeny 81" docPubDate="2023-08-01" docSource="http://dx.doi.org/10.3897/asp.81.e101374" docTitle="Cretomophron Rosova, Prokop & Beutel 2023, gen. nov." docType="treatment" docUuid="B7D03034-E81C-41C9-B407-262A5B21C864" docUuidSource="ZooBank" docVersion="1" id="B61E7B03B308528C9FC150520EABBE19" lastPageNumber="689" masterDocId="B61E7B03B308528C9FC150520EABBE19" masterDocTitle="The earliest evidence of Omophroninae (Coleoptera: Carabidae) from mid-Cretaceous Kachin amber and the description of a larva of a new genus" masterLastPageNumber="704" masterPageNumber="689" pageNumber="689" updateTime="1690954509890" updateUser="pensoft">
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<mods:title id="E9F7D71995E802773556779F00FE2070">The earliest evidence of Omophroninae (Coleoptera: Carabidae) from mid-Cretaceous Kachin amber and the description of a larva of a new genus</mods:title>
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<mods:namePart id="7795289CB26867EAB5733AE4D724C1D7">Rosova, Katerina</mods:namePart>
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<mods:affiliation id="1ABFDC1DF1B190B31959E372E24DC6A7">Charles University, Faculty of Science, Department of Zoology, Vinicna 7, 128 00 Prague 2, Czech Republic</mods:affiliation>
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<mods:namePart id="D813686CFCA59DCBF6AB86F80115453E">Prokop, Jakub</mods:namePart>
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<mods:nameIdentifier id="5E3D09C11AA84773378B4D15A25D5873" type="ORCID">https://orcid.org/0000-0001-6996-7832</mods:nameIdentifier>
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<mods:affiliation id="B432ED6F22800EF574225EB6EA2E9A1C">Charles University, Faculty of Science, Department of Zoology, Vinicna 7, 128 00 Prague 2, Czech Republic</mods:affiliation>
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<mods:name id="838DBAA1C3CBD881C6104787F41B1E18" type="personal">
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<mods:namePart id="371D46B3D386893D9EE470366D1037EB">Hammel, Joerg U.</mods:namePart>
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<mods:nameIdentifier id="FE3B3CD67E67FC38E185FD3CA6FA396F" type="ORCID">https://orcid.org/0000-0002-6744-6811</mods:nameIdentifier>
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<mods:affiliation id="1480A8E8BD9CEEA3B93D15DD222F55A0">Institute of Materials Physics, Helmholtz-Zentrum Hereon, Max-Planck-Strasse 1, 21502 Geesthacht, Germany</mods:affiliation>
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<mods:namePart id="C35BBF35585892EC8EF6422A17E32A02">Beutel, Rolf G.</mods:namePart>
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<mods:nameIdentifier id="34BB4F174DE6CA67BF0A0102A0C9990D" type="ORCID">https://orcid.org/0000-0002-0433-7626</mods:nameIdentifier>
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<mods:affiliation id="521EE9D7FF8E8AE52DADE9B8196E20EC">Institut fuer Zoologie und Evolutionsforschung, Friedrich-Schiller-Universitaet Jena, 07743 Jena, Germany</mods:affiliation>
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<mods:nameIdentifier id="D0078F96B9E8CBC8468E2283A9069E23" type="email">rolf.beutel@uni-jena.de</mods:nameIdentifier>
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<mods:title id="AF6C030FD7DAC2FBB96C9905A965A91C">Arthropod Systematics & amp; Phylogeny</mods:title>
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<mods:date id="29CFA57CB4B27894219FD6008725B600">2023</mods:date>
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<mods:number id="8D555B05B395B3379821565ACD9B9DAE">2023-08-01</mods:number>
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<mods:identifier id="81372D74F6AAC01B49E3EC17689C61AF" type="DOI">http://dx.doi.org/10.3897/asp.81.e101374</mods:identifier>
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<treatment id="DEDF00746A035664A1B7334CF93F5770" LSID="urn:lsid:zoobank.org:act:B7D03034-E81C-41C9-B407-262A5B21C864" httpUri="http://treatment.plazi.org/id/DEDF00746A035664A1B7334CF93F5770" lastPageNumber="689" pageId="0" pageNumber="689">
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<subSubSection id="83A1A17D82D89ACAC23923670B70A8A4" pageId="0" pageNumber="689" type="nomenclature">
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<paragraph id="63FC7EDB3C48DF003553AD440F08C72D" pageId="0" pageNumber="689">
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†
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<taxonomicName id="7ABBF5C7DF661A135C52CF57AFB7AE40" LSID="https://zoobank.org/B7D03034-E81C-41C9-B407-262A5B21C864" authority="Rosova, Prokop & Beutel" authorityName="Rosova, Prokop & Beutel" authorityYear="2023" family="Carabidae" genus="Cretomophron" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Cretomophron" order="Coleoptera" pageId="0" pageNumber="689" rank="genus" status="gen. nov.">Cretomophron Rosova, Prokop & Beutel</taxonomicName>
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<taxonomicNameLabel id="B89EEB640A1969AA86F2C6E4E1E96C5C" pageId="0" pageNumber="689">gen. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="9B3966445DCC97EB199F7FBE26D62A1D" pageId="0" pageNumber="689" type="type species">
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<paragraph id="09328B1177113936DE5C0B5B4D674C9F" pageId="0" pageNumber="689">Type species.</paragraph>
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<paragraph id="7B3AAE3EBD2AFFCF0CD0B3EB370C4F56" pageId="0" pageNumber="689">
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†
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<taxonomicName id="8E19FC9F9211F809B2E0BAFB22096DD9" family="Carabidae" genus="Cretomophron" kingdom="Animalia" lsidName="Cretomophron mutilus" order="Coleoptera" pageId="0" pageNumber="689" rank="species" species="mutilus">
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<emphasis id="A4D7B6B57E7C6961F1175D7ED36DDE59" italics="true" pageId="0" pageNumber="689">Cretomophron mutilus</emphasis>
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</taxonomicName>
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Rosova, Prokop & Beutel, gen. et sp. nov. - sp. nov. by original designation and monotypy. - Included species. Type species only.
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</paragraph>
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</subSubSection>
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<subSubSection id="109A7542A0939D4640D79D4D3BA70252" pageId="0" pageNumber="689" type="diagnosis">
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<paragraph id="A7946E110A72D5250413FA8B62BEAFE6" pageId="0" pageNumber="689">Diagnosis (larva).</paragraph>
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<paragraph id="2DAFA16E287800CB8BB504537A531E86" pageId="0" pageNumber="689">
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The specimen described here differs from larvae of all holometabolan groups outside of
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<taxonomicName id="A3CC18FC78ED4C167402FE97D467D00D" authorityName="Rosova, Prokop & Beutel" authorityYear="2023" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="689" rank="order">Coleoptera</taxonomicName>
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and from
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<taxonomicName id="0DA12A19D36A0427DDB8F7AF250BB738" authorityName="Rosova, Prokop & Beutel" authorityYear="2023" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="0" pageNumber="689" rank="subOrder" subOrder="Archostemata">Archostemata</taxonomicName>
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by the presence of distinctly developed urogomphi on abdominal tergite IX (Figs
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<figureCitation id="80896C6DED9FC50CD17CED6B4D6C5738" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Cretomophron mutilus Rosova, Prokop & Beutel, gen. et sp. nov.: A detail of head and mouthparts, dorsal view; B habitus, dorsal view; C abdominal segment VI, dorsal view. Scale bars: (A) 0.1 mm, (B) 0.8 mm, (C) 0.2 mm. Abbreviations: an - antenna, ga - galea, lc - lacinia, nas - nasale, pl - labial palpus, pmx - maxillary palpus, ste - stemmata." figureDoi="10.3897/asp.81.e101374.figure1" httpUri="https://binary.pensoft.net/fig/886690" pageId="0" pageNumber="689">1B</figureCitation>
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,
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<figureCitation id="8254B5CC53A64523C29BF04F1C90E7DE" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. (A) Habitus of † Cretomophron mutilus Rosova, Prokop & Beutel, gen. et sp. nov., dorsal view. Dashed lines show missing or poorly visible structures. Scale bar 0.8 mm; (B) Habitus of larva of Omophron limbatum (Fabricius, 1776), adapted from Luff 1993, modified." figureDoi="10.3897/asp.81.e101374.figure4" httpUri="https://binary.pensoft.net/fig/886693" pageId="0" pageNumber="689">4</figureCitation>
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). It can be distinguished from all carabid groups outside of
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<taxonomicName id="7FEF28B8A86DF1F4023B272EAB270ECF" lsidName="" pageId="0" pageNumber="689" rank="subFamily" subFamily="Omophroninae">Omophroninae</taxonomicName>
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by a pronouncedly wedge-shaped head, antennae directed upwards (Fig.
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<figureCitation id="7E51A918D8650FB1E2E6A92C3B7E429B" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. † Cretomophron mutilus Rosova, Prokop & Beutel, gen. et sp. nov., volume renders of the segmented SRµCT data: A dorsal view, B lateral view, C ventral view. Scale bar 0.8 mm." figureDoi="10.3897/asp.81.e101374.figure3" httpUri="https://binary.pensoft.net/fig/886692" pageId="0" pageNumber="689">3B</figureCitation>
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), and a laterally oriented terminal 4th antennomere (Figs
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<figureCitation id="587A963C7FD303771A4294D8A1F80731" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. † Cretomophron mutilus Rosova, Prokop & Beutel, gen. et sp. nov.: A head and mouthparts, ventral view; B mesothoracic leg, ventral view; C volume render of prothoracic leg from the segmented SRµCT data, lateral view; D detail of the pretarsus of the metathoracic leg. Scale bars: (A, B) 0.2 mm, (C) 0.8 mm, (D) 0.1 mm. Abbreviations: an - antenna, c - cardo, ga - galea, lc - lacinia, pl - labial palpus, pmt - prementum, pmx - maxillary palpus, ptg - posterior tentorial bridge, sa - sensorial appendage." figureDoi="10.3897/asp.81.e101374.figure2" httpUri="https://binary.pensoft.net/fig/886691" pageId="0" pageNumber="689">2A</figureCitation>
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,
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<figureCitation id="1D4C3495105D5F609803439A67824ECD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. † Cretomophron mutilus Rosova, Prokop & Beutel, gen. et sp. nov.: A head, dorsal view, chaetotaxy simplified; B head, ventral view; C abdominal segment VI, dorsal view. Dashed lines show missing or poorly visible structures. Scale bars 0.4 mm. Abbreviations: an - antenna, c - cardo, co - coronal suture, f - frontal suture, ga - galea, lc - lacinia, md - mandible, nas - nasale, pl - labial palpus, pmt - prementum, pmx - maxillary palpus, ptg - posterior tentorial bridge, sa - sensorial appendage, st - stipes, ste - stemmata." figureDoi="10.3897/asp.81.e101374.figure5" httpUri="https://binary.pensoft.net/fig/886694" pageId="0" pageNumber="689">5A</figureCitation>
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). The prothorax appears proportionally larger than in other carabid larvae, but this is a gradual modification only. Additional unusual diagnostic features are the large triangular nasale (Figs
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<figureCitation id="256E57A06CF63C51D2AC9D2C74C8F2C8" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Cretomophron mutilus Rosova, Prokop & Beutel, gen. et sp. nov.: A detail of head and mouthparts, dorsal view; B habitus, dorsal view; C abdominal segment VI, dorsal view. Scale bars: (A) 0.1 mm, (B) 0.8 mm, (C) 0.2 mm. Abbreviations: an - antenna, ga - galea, lc - lacinia, nas - nasale, pl - labial palpus, pmx - maxillary palpus, ste - stemmata." figureDoi="10.3897/asp.81.e101374.figure1" httpUri="https://binary.pensoft.net/fig/886690" pageId="0" pageNumber="689">1A</figureCitation>
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,
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<figureCitation id="CB9C43927D1AB699B27504FB5E858A9C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. † Cretomophron mutilus Rosova, Prokop & Beutel, gen. et sp. nov.: A head, dorsal view, chaetotaxy simplified; B head, ventral view; C abdominal segment VI, dorsal view. Dashed lines show missing or poorly visible structures. Scale bars 0.4 mm. Abbreviations: an - antenna, c - cardo, co - coronal suture, f - frontal suture, ga - galea, lc - lacinia, md - mandible, nas - nasale, pl - labial palpus, pmt - prementum, pmx - maxillary palpus, ptg - posterior tentorial bridge, sa - sensorial appendage, st - stipes, ste - stemmata." figureDoi="10.3897/asp.81.e101374.figure5" httpUri="https://binary.pensoft.net/fig/886694" pageId="0" pageNumber="689">5A</figureCitation>
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), antennae with the 2nd antennomere markedly elongated, legs with a very short tibia and long setae UN1, UN2 on claws (Fig.
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<figureCitation id="7FDD9F96138CBB16155EC6467E3BE6A3" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. † Cretomophron mutilus Rosova, Prokop & Beutel, gen. et sp. nov.: A head and mouthparts, ventral view; B mesothoracic leg, ventral view; C volume render of prothoracic leg from the segmented SRµCT data, lateral view; D detail of the pretarsus of the metathoracic leg. Scale bars: (A, B) 0.2 mm, (C) 0.8 mm, (D) 0.1 mm. Abbreviations: an - antenna, c - cardo, ga - galea, lc - lacinia, pl - labial palpus, pmt - prementum, pmx - maxillary palpus, ptg - posterior tentorial bridge, sa - sensorial appendage." figureDoi="10.3897/asp.81.e101374.figure2" httpUri="https://binary.pensoft.net/fig/886691" pageId="0" pageNumber="689">2D</figureCitation>
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), and abdominal segment VI with numerous setae collected in transverse rows (Figs
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<figureCitation id="2612EE1D3F0CFE20687620953BE65684" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. † Cretomophron mutilus Rosova, Prokop & Beutel, gen. et sp. nov.: A detail of head and mouthparts, dorsal view; B habitus, dorsal view; C abdominal segment VI, dorsal view. Scale bars: (A) 0.1 mm, (B) 0.8 mm, (C) 0.2 mm. Abbreviations: an - antenna, ga - galea, lc - lacinia, nas - nasale, pl - labial palpus, pmx - maxillary palpus, ste - stemmata." figureDoi="10.3897/asp.81.e101374.figure1" httpUri="https://binary.pensoft.net/fig/886690" pageId="0" pageNumber="689">1A</figureCitation>
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,
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<figureCitation id="034A7B7FCF78EE21264784E940DC7A67" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. † Cretomophron mutilus Rosova, Prokop & Beutel, gen. et sp. nov.: A head and mouthparts, ventral view; B mesothoracic leg, ventral view; C volume render of prothoracic leg from the segmented SRµCT data, lateral view; D detail of the pretarsus of the metathoracic leg. Scale bars: (A, B) 0.2 mm, (C) 0.8 mm, (D) 0.1 mm. Abbreviations: an - antenna, c - cardo, ga - galea, lc - lacinia, pl - labial palpus, pmt - prementum, pmx - maxillary palpus, ptg - posterior tentorial bridge, sa - sensorial appendage." figureDoi="10.3897/asp.81.e101374.figure2" httpUri="https://binary.pensoft.net/fig/886691" pageId="0" pageNumber="689">2B</figureCitation>
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,
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<figureCitation id="C080D529EF22001741C49A08752E6E3E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. (A) Habitus of † Cretomophron mutilus Rosova, Prokop & Beutel, gen. et sp. nov., dorsal view. Dashed lines show missing or poorly visible structures. Scale bar 0.8 mm; (B) Habitus of larva of Omophron limbatum (Fabricius, 1776), adapted from Luff 1993, modified." figureDoi="10.3897/asp.81.e101374.figure4" httpUri="https://binary.pensoft.net/fig/886693" pageId="0" pageNumber="689">4</figureCitation>
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), and large trochanters with distal projections (Figs
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<figureCitation id="74B606222D3B886864392CBFBA1DF6B1" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. † Cretomophron mutilus Rosova, Prokop & Beutel, gen. et sp. nov., volume renders of the segmented SRµCT data: A dorsal view, B lateral view, C ventral view. Scale bar 0.8 mm." figureDoi="10.3897/asp.81.e101374.figure3" httpUri="https://binary.pensoft.net/fig/886692" pageId="0" pageNumber="689">3B, C</figureCitation>
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). Posterior tentorial grooves not shifted to the posterior margin of the ventral head capsule distinguish it from larvae of
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<taxonomicName id="98D80F16EFC56FDBC9404783A35BE39E" authorityName="Latreille" authorityYear="1802" class="Insecta" family="Carabidae" genus="Omophron" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omophron" order="Coleoptera" pageId="0" pageNumber="689" phylum="Arthropoda" rank="genus">
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<emphasis id="3B9007A63BAF6EBD6C07470D4D87F789" italics="true" pageId="0" pageNumber="689">Omophron</emphasis>
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</taxonomicName>
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(Figs
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<figureCitation id="46CAB79569D639C37AB753D42BA98192" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. † Cretomophron mutilus Rosova, Prokop & Beutel, gen. et sp. nov.: A head and mouthparts, ventral view; B mesothoracic leg, ventral view; C volume render of prothoracic leg from the segmented SRµCT data, lateral view; D detail of the pretarsus of the metathoracic leg. Scale bars: (A, B) 0.2 mm, (C) 0.8 mm, (D) 0.1 mm. Abbreviations: an - antenna, c - cardo, ga - galea, lc - lacinia, pl - labial palpus, pmt - prementum, pmx - maxillary palpus, ptg - posterior tentorial bridge, sa - sensorial appendage." figureDoi="10.3897/asp.81.e101374.figure2" httpUri="https://binary.pensoft.net/fig/886691" pageId="0" pageNumber="689">2A</figureCitation>
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,
|
||
<figureCitation id="9C723A90CC5C1F842340DA82EE2491C8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. † Cretomophron mutilus Rosova, Prokop & Beutel, gen. et sp. nov.: A head, dorsal view, chaetotaxy simplified; B head, ventral view; C abdominal segment VI, dorsal view. Dashed lines show missing or poorly visible structures. Scale bars 0.4 mm. Abbreviations: an - antenna, c - cardo, co - coronal suture, f - frontal suture, ga - galea, lc - lacinia, md - mandible, nas - nasale, pl - labial palpus, pmt - prementum, pmx - maxillary palpus, ptg - posterior tentorial bridge, sa - sensorial appendage, st - stipes, ste - stemmata." figureDoi="10.3897/asp.81.e101374.figure5" httpUri="https://binary.pensoft.net/fig/886694" pageId="0" pageNumber="689">5B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="A2072743F1429663575460691E750BD8" doi="10.3897/asp.81.e101374.figure1" httpUri="https://binary.pensoft.net/fig/886690" pageId="0" pageNumber="689" start="Figure 1" startId="F1">
|
||
<paragraph id="BC2425B237FABE23BE071CAD07794541" pageId="0" pageNumber="689">
|
||
<emphasis id="7795AEF0A8138B22E7231A7DF311B006" bold="true" pageId="0" pageNumber="689">Figure 1.</emphasis>
|
||
†
|
||
<taxonomicName id="1EE51B405B77417715B48EE704DB1D73" family="Carabidae" genus="Cretomophron" kingdom="Animalia" lsidName="Cretomophron mutilus" order="Coleoptera" pageId="0" pageNumber="689" rank="species" species="mutilus">
|
||
<emphasis id="DE2E7567A6838C7B3CC947461A212093" italics="true" pageId="0" pageNumber="689">Cretomophron mutilus</emphasis>
|
||
</taxonomicName>
|
||
Rosova, Prokop & Beutel, gen. et sp. nov.:
|
||
<emphasis id="A36CC30864686050087539C10B9B38B8" bold="true" pageId="0" pageNumber="689">A</emphasis>
|
||
detail of head and mouthparts, dorsal view;
|
||
<emphasis id="A8247DAAC1965B63FA5358FE7BF24000" bold="true" pageId="0" pageNumber="689">B</emphasis>
|
||
habitus, dorsal view;
|
||
<emphasis id="69D4E133A2B2ED94E9FADCAA8BB44837" bold="true" pageId="0" pageNumber="689">C</emphasis>
|
||
abdominal segment VI, dorsal view. Scale bars: (A) 0.1 mm, (B) 0.8 mm, (C) 0.2 mm. Abbreviations: an - antenna, ga - galea, lc - lacinia, nas - nasale, pl - labial palpus, pmx - maxillary palpus, ste - stemmata.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="CF726ABEBF9BF64C374C5EB1628B778D" doi="10.3897/asp.81.e101374.figure2" httpUri="https://binary.pensoft.net/fig/886691" pageId="0" pageNumber="689" start="Figure 2" startId="F2">
|
||
<paragraph id="73C944290381CF3242A3E294170A4750" pageId="0" pageNumber="689">
|
||
<emphasis id="EAEF4F3BD56574B5A6302C8EE75355AD" bold="true" pageId="0" pageNumber="689">Figure 2.</emphasis>
|
||
†
|
||
<taxonomicName id="60326B7709FD85485560711CF05FF3C3" family="Carabidae" genus="Cretomophron" kingdom="Animalia" lsidName="Cretomophron mutilus" order="Coleoptera" pageId="0" pageNumber="689" rank="species" species="mutilus">
|
||
<emphasis id="E4DA1B44464438576CFFEAA0FB2FA01F" italics="true" pageId="0" pageNumber="689">Cretomophron mutilus</emphasis>
|
||
</taxonomicName>
|
||
Rosova, Prokop & Beutel, gen. et sp. nov.:
|
||
<emphasis id="3B4CE0C0066AE0CC210D28832BEB864F" bold="true" pageId="0" pageNumber="689">A</emphasis>
|
||
head and mouthparts, ventral view;
|
||
<emphasis id="B4A74250EE24A0906DF2386FC5C90BA4" bold="true" pageId="0" pageNumber="689">B</emphasis>
|
||
mesothoracic leg, ventral view;
|
||
<emphasis id="CB75639B1295FB9803D4F529C171BACF" bold="true" pageId="0" pageNumber="689">C</emphasis>
|
||
volume render of prothoracic leg from the segmented
|
||
<normalizedToken id="60CB1EAEE8B4D294848C1130026CA7BB" originalValue="SRµCT">SRµCT</normalizedToken>
|
||
data, lateral view;
|
||
<emphasis id="D8488C35E6294925D4C78EA8E3271935" bold="true" pageId="0" pageNumber="689">D</emphasis>
|
||
detail of the pretarsus of the metathoracic leg. Scale bars: (A, B) 0.2 mm, (C) 0.8 mm, (D) 0.1 mm. Abbreviations: an - antenna, c - cardo, ga - galea, lc - lacinia, pl - labial palpus, pmt - prementum, pmx - maxillary palpus, ptg - posterior tentorial bridge, sa - sensorial appendage.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="66A75B70D6E4E3A15B7463B9F9F66BBA" doi="10.3897/asp.81.e101374.figure3" httpUri="https://binary.pensoft.net/fig/886692" pageId="0" pageNumber="689" start="Figure 3" startId="F3">
|
||
<paragraph id="463535B3F69CDBD26AAAE71A04B903F9" pageId="0" pageNumber="689">
|
||
<emphasis id="49C6B0BFEE4D7405DFF0823395713A47" bold="true" pageId="0" pageNumber="689">Figure 3.</emphasis>
|
||
†
|
||
<taxonomicName id="2392C3ED4DAF69942611BF4D2BFE617C" family="Carabidae" genus="Cretomophron" kingdom="Animalia" lsidName="Cretomophron mutilus" order="Coleoptera" pageId="0" pageNumber="689" rank="species" species="mutilus">
|
||
<emphasis id="C6E2758D1F4A87DCC4EA0B082DACA952" italics="true" pageId="0" pageNumber="689">Cretomophron mutilus</emphasis>
|
||
</taxonomicName>
|
||
Rosova, Prokop & Beutel, gen. et sp. nov., volume renders of the segmented
|
||
<normalizedToken id="5DDAC146D5DD888A6C57F37290D49894" originalValue="SRµCT">SRµCT</normalizedToken>
|
||
data:
|
||
<emphasis id="AF72BDAE60E6708EC0B25895F654E6B4" bold="true" pageId="0" pageNumber="689">A</emphasis>
|
||
dorsal view,
|
||
<emphasis id="0C3A2BAC46C77BF9B1BE0AE0BE4841F2" bold="true" pageId="0" pageNumber="689">B</emphasis>
|
||
lateral view,
|
||
<emphasis id="88F4FA4D6EC23A34A824B434CA70D5C3" bold="true" pageId="0" pageNumber="689">C</emphasis>
|
||
ventral view. Scale bar 0.8 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="6BF4BB882375083637F87798BDB8EBC5" doi="10.3897/asp.81.e101374.figure4" httpUri="https://binary.pensoft.net/fig/886693" pageId="0" pageNumber="689" start="Figure 4" startId="F4">
|
||
<paragraph id="46BBD1840F6D9133680A4489B5592C17" pageId="0" pageNumber="689">
|
||
<emphasis id="66C44D3607553E1705F9E63D59EFEFC4" bold="true" pageId="0" pageNumber="689">Figure 4.</emphasis>
|
||
(A) Habitus of †
|
||
<taxonomicName id="FC8D945CE078443B03B953563C1400A6" family="Carabidae" genus="Cretomophron" kingdom="Animalia" lsidName="Cretomophron mutilus" order="Coleoptera" pageId="0" pageNumber="689" rank="species" species="mutilus">
|
||
<emphasis id="4C64220100426AB53E59271471C3FBDE" italics="true" pageId="0" pageNumber="689">Cretomophron mutilus</emphasis>
|
||
</taxonomicName>
|
||
Rosova, Prokop & Beutel, gen. et sp. nov., dorsal view. Dashed lines show missing or poorly visible structures. Scale bar 0.8 mm; (B) Habitus of larva of
|
||
<taxonomicName id="C9F78DD89855BED3A8BA4EF60E0D5C77" authorityName="Fabricius" authorityYear="1777" baseAuthorityName="Fabricius" baseAuthorityYear="1777" class="Insecta" family="Carabidae" genus="Omophron" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Omophron limbatum" order="Coleoptera" pageId="0" pageNumber="689" phylum="Arthropoda" rank="species" species="limbatum">
|
||
<emphasis id="FCFDA0AE5691455FC21E324E4C9A8D21" italics="true" pageId="0" pageNumber="689">Omophron limbatum</emphasis>
|
||
</taxonomicName>
|
||
(Fabricius, 1776), adapted from
|
||
<bibRefCitation id="B45DF7DD1F81FDC99B19C3723F6C88DA" author="Luff, ML" journalOrPublisher="Fauna Entomologica Scandinavica, 27, EJ Brill, Leiden, New York, Koeln" pageId="0" pageNumber="689" refId="B51" refString="Luff, ML, 1993. The Carabidae (Coleoptera) larvae of Fenoscandia and Denmark. Fauna Entomologica Scandinavica, 27, EJ Brill, Leiden, New York, Koeln" title="The Carabidae (Coleoptera) larvae of Fenoscandia and Denmark." year="1993">Luff 1993</bibRefCitation>
|
||
, modified.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="3EC4893880E84023FEA97D0DBA2FB079" doi="10.3897/asp.81.e101374.figure5" httpUri="https://binary.pensoft.net/fig/886694" pageId="0" pageNumber="689" start="Figure 5" startId="F5">
|
||
<paragraph id="6A0A25937C2F9F252B1A2827759EB7C6" pageId="0" pageNumber="689">
|
||
<emphasis id="0B66D65D904563AAC5FE7B826EA72310" bold="true" pageId="0" pageNumber="689">Figure 5.</emphasis>
|
||
†
|
||
<taxonomicName id="2031A5C89FC3527B897667503CDDD7FC" family="Carabidae" genus="Cretomophron" kingdom="Animalia" lsidName="Cretomophron mutilus" order="Coleoptera" pageId="0" pageNumber="689" rank="species" species="mutilus">
|
||
<emphasis id="83AAC317382107CEA0047D8F002EE3E8" italics="true" pageId="0" pageNumber="689">Cretomophron mutilus</emphasis>
|
||
</taxonomicName>
|
||
Rosova, Prokop & Beutel, gen. et sp. nov.:
|
||
<emphasis id="ADB437CB2BFD5F47897715380F36598A" bold="true" pageId="0" pageNumber="689">A</emphasis>
|
||
head, dorsal view, chaetotaxy simplified;
|
||
<emphasis id="4D96157851E4E12C2F0446C027BE28C3" bold="true" pageId="0" pageNumber="689">B</emphasis>
|
||
head, ventral view;
|
||
<emphasis id="19338C26E24AAF4F0C2C2AADD816CF11" bold="true" pageId="0" pageNumber="689">C</emphasis>
|
||
abdominal segment VI, dorsal view. Dashed lines show missing or poorly visible structures. Scale bars 0.4 mm. Abbreviations: an - antenna, c - cardo, co - coronal suture, f - frontal suture, ga - galea, lc - lacinia, md - mandible, nas - nasale, pl - labial palpus, pmt - prementum, pmx - maxillary palpus, ptg - posterior tentorial bridge, sa - sensorial appendage, st - stipes, ste - stemmata.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="BB878486F4C5972BAA9EF5D282532AF0" pageId="0" pageNumber="689" type="etymology">
|
||
<paragraph id="3E273E2DD95590B891B1D584250D0D39" pageId="0" pageNumber="689">Etymology.</paragraph>
|
||
<paragraph id="038CCB8A4DF09B93029815F8832A0615" pageId="0" pageNumber="689">
|
||
The generic epithet combines
|
||
<normalizedToken id="00D41BC6D4F524AF0357616C528423CF" originalValue="‘creto-’">'creto-'</normalizedToken>
|
||
(Latin for chalky as in Cretaceous), referring to the geological period, and
|
||
<normalizedToken id="228355F69BD585779CE1127B51B8185D" originalValue="‘omophron’">'omophron'</normalizedToken>
|
||
, the single extant genus of
|
||
<taxonomicName id="929654A52EF59C63844951D1634E8931" lsidName="" pageId="0" pageNumber="689" rank="subFamily" subFamily="Omophroninae">Omophroninae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |