146 lines
14 KiB
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146 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.248.4106" ID-GBIF-Dataset="605e7744-e499-4617-bb69-6531a82ddbae" ID-PMC="PMC3520135" ID-Pensoft-Pub="1313-2970-248-1" ID-PubMed="23275757" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-248-1" ModsDocOrigin="ZooKeys 248" ModsDocTitle="A revision of the New World species of Gymnoclasiopa Hendel (Diptera, Ephydridae)" checkinTime="1451247880623" checkinUser="pensoft" docAuthor="Mathis, Wayne N. & Zatwarnicki, Tadeusz" docDate="2012" docId="CD1E45D674E2847A5B862C0EF8F7082E" docLanguage="en" docName="ZooKeys 248: 1-69" docOrigin="ZooKeys 248" docSource="http://dx.doi.org/10.3897/zookeys.248.4106" docTitle="Gymnoclasiopa grecorum Mathis & Zatwarnicki, sp. n." docType="treatment" docVersion="4" lastPageNumber="28" masterDocId="23013B478951FFB1FFF7FFACFFBFFF8F" masterDocTitle="A revision of the New World species of Gymnoclasiopa Hendel (Diptera, Ephydridae)" masterLastPageNumber="69" masterPageNumber="1" pageNumber="25" updateTime="1668154988994" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A revision of the New World species of Gymnoclasiopa Hendel (Diptera, Ephydridae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Mathis, Wayne N.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zatwarnicki, Tadeusz</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2012</mods:date>
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<mods:detail type="volume">
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<mods:number>248</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>69</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.248.4106</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.248.4106</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-248-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152039112" LSID="urn:lsid:zoobank.org:act:29AD8BF1-45F7-41C9-A756-6DD850D7C623" httpUri="http://treatment.plazi.org/id/CD1E45D674E2847A5B862C0EF8F7082E" lastPageId="27" lastPageNumber="28" pageId="24" pageNumber="25">
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<subSubSection pageId="24" pageNumber="25" type="nomenclature">
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<paragraph pageId="24" pageNumber="25">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:29AD8BF1-45F7-41C9-A756-6DD850D7C623" authority="Mathis & Zatwarnicki" class="Insecta" family="Ephydridae" genus="Gymnoclasiopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gymnoclasiopa grecorum" order="Diptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="grecorum">Gymnoclasiopa grecorum Mathis & Zatwarnicki</taxonomicName>
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<taxonomicNameLabel pageId="24" pageNumber="25">sp. n.</taxonomicNameLabel>
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Figs 29-34
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25" type="diagnosis">
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<paragraph pageId="24" pageNumber="25">Diagnosis.</paragraph>
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<paragraph lastPageId="26" lastPageNumber="27" pageId="24" pageNumber="25">
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This species is distinguished from congeners by the following combination of characters: Moderately small to medium-sized shore flies, body length 2.10-3.15 mm; head generally dull, thorax faintly subshiny medially, dull, microtomentose anteriorly and laterally, microtomentose gray, abdomen black with thin investment of whitish microtomentum. Head (Figs 29-30): Frons densely microtomentose, dull, gray to blackish gray. Antenna black; basal flagellomere covered with silvery gray microtomentum, with faint reddish orange coloration mediobasally; arista bearing 5-6 dorsal rays. Face somewhat flat, antennal grooves generally inconspicuous, oriented ventrolaterally to vertical; facial microtomentum generally yellow to golden yellow, slightly grayish to silvery white in antennal grooves; parafacial and gena whitish gray to white; genal height about equal to height of basal flagellomere; gena moderately short; gena-to-eye ratio 0.17-0.20. Maxillary palpus varying from mostly black but with some reddish to yellowish coloration at middle to entirely yellowish. Thorax: Mesonotum and pleural area generally densely microtomentose, gray to tannish gray, dull laterally and anteriorly, mesonotum medially partially subshiny with some faint oliveceous to greenish metallic luster. Wing hyaline; costal vein ratio 0.37-0.43; M vein ratio 0.62-0.74; halter stem tan to yellowish tan; knob yellow to whitish yellow. Coxae gray; femora and tibia concolorous gray to blackish gray, densely microtomentose; tarsi generally yellowish; apical 2-3 tarsomeres darkened, brown to blackish brown. Abdomen: Generally thinly microtomentose; basal tergites
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<pageBreakToken pageId="25" pageNumber="26" start="start">largely</pageBreakToken>
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gray to grayish white; tergite 5 of male subshiny to shiny, less densely microtomentose than anterior tergites. Male terminalia (Figs 31-33): Epandrium in posterior view (Fig. 31) as a broadly rounded, inverted U, width of dorsal portion narrower than expanded apices of lateral arms, lateral arms nearly straight, in lateral view wider ventrally; cercus in posterior view (Fig. 31) unevenly semilunate, much narrower dorsally than ventrally, densely setulose; aedeagus in lateral view (Fig. 33) as a parallelogram, basal margin shallowly emarginate, tapered toward apex, apex narrowly rounded, in ventral view (Fig. 32) expanded slightly laterally from narrow base to wider apex, apical margin bilobed, narrowly incised medially, each lateral lobe slightly narrower than aedeagal base, with thin, wing-like, narrow projections sub-basally; phallapodeme in lateral view (Fig. 33) more or less triangular, in ventral view (Fig. 32) longer than wide, bar-like with basal, sub-basal and apical crossbars, apical margin shallowly emarginate; ejaculatory apodeme in lateral view (Fig. 33) as an irregular comma with less arch; postgonite in lateral view (Fig. 33) narrowly pointed basally, thereafter becoming wider then apex narrowed, tapered, apex pointed, bearing 3-4 tiny setulae along basoposterior margin and 1 setula subapically; pregonite in lateral view (Fig. 33) moderately elongate, tapered, narrowed toward hypandrium, expanded toward aedeagus, aedeagal end truncate; hypandrium in ventral view (Fig. 32) wide, width nearly twice length, broadly and shallowly rounded along anterior margin with anterolateral shallowly and moreroundedly projected, obtusely pointed, shallow projections, angulately and moderately deeply incised along posterior margin
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<pageBreakToken pageId="26" pageNumber="27" start="start">medially</pageBreakToken>
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, in lateral view (Fig. 33) irregularly clavate, very shallowly arched, expanded anteriorly, narrow, parallel sided posteriorly.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="27" type="type material">
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<paragraph pageId="26" pageNumber="27">Type material.</paragraph>
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<paragraph pageId="26" pageNumber="27">
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The holotype male is
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<taxonomicName class="Insecta" family="Ephydridae" genus="Gymnoclasiopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gymnoclasiopa grecorum" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="grecorum">Gymnoclasiopa grecorum</taxonomicName>
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Mathis and Zatwarnicki is labeled "USA. A[las]K[a]. Juneau: Thane Road (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="58.281666">58°16.9'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-134.37334">134°22.4'W</geoCoordinate>
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), 22 July 2011[,] D. & W. N. Mathis/USNM ENT 00117971 [plastic bar code label]/HOLOTYPE ♂
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<taxonomicName class="Insecta" family="Ephydridae" genus="Gymnoclasiopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gymnoclasiopa grecorum" order="Diptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="grecorum">Gymnoclasiopa grecorum</taxonomicName>
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Mathis & Zatwarnicki, USNM [red]." The holotype is double mounted (minuten pin in a block of plastic), is in excellent condition, and is deposited in the USNM. Twenty-four paratypes (13♂, 11♀; USNM) bear the same locality label data as the holotype with dates from 20-22 July 2011. Other paratypes are as follows: UNITED STATES. Alaska. Juneau: Echo Cove (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="58.661667">58°39.7'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-134.90666">134°54.4'W</geoCoordinate>
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), 21 Jul 2011, D. and W. N. Mathis (1♂; USNM); Juneau, Mendenhall Valley, Riverside Rotary Park (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="58.38">58°22.8'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-134.58667">134°35.2'W</geoCoordinate>
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), 21 Jul 2011, D. and W. N. Mathis (1♂; USNM). Matanuska-Susitna: Little Willow Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.81">61°48.6'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-150.09666">150°05.8'W</geoCoordinate>
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; 50 m), 25 Jul 2011, D. and W. N. Mathis (3♂; USNM); Sheep Creek (
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.97167">61°58.3'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="925" value="-150.08333">150°05'W</geoCoordinate>
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; 55 m), 10 Aug 2011, D. and W.N. Mathis (3♀; USNM); Talkeetna (Susitna River;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="61.323334">61°19.4'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-150.12">150°07.2'W</geoCoordinate>
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; 120 m), 10 Aug 2011, D. and W.N. Mathis (3♂, 1♀; USNM).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="26" pageNumber="27" type="type locality">
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<paragraph pageId="26" pageNumber="27">Type locality.</paragraph>
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<paragraph pageId="26" pageNumber="27">
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United States. Alaska. Juneau: Gastineau Channel, Thane Road (S Juneau;
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<geoCoordinate direction="north" orientation="latitude" precision="92" value="58.281666">58°16.9'N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="92" value="-134.37334">134°22.4'W</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27" type="distribution">
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<paragraph pageId="26" pageNumber="27">Distribution</paragraph>
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<paragraph lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
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(Fig. 34). Nearctic: United States (Alaska)
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<pageBreakToken pageId="27" pageNumber="28" start="start">.</pageBreakToken>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="etymology">
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<paragraph pageId="27" pageNumber="28">Etymology.</paragraph>
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<paragraph pageId="27" pageNumber="28">The species epithet, grecorum, is a plural genitive Latin patronym to honor and recognize the generous contributions of Richard Art and Karen Greco (nee Halliday) toward the improvement of mankind, their friendship, and their love of Alaska.</paragraph>
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</subSubSection>
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<subSubSection pageId="27" pageNumber="28" type="remarks">
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<paragraph pageId="27" pageNumber="28">Remarks.</paragraph>
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<paragraph pageId="27" pageNumber="28">
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This species is similar to
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<taxonomicName class="Insecta" family="Ephydridae" genus="Gymnoclasiopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gymnoclasiopa subnubila" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="subnubila">Gymnoclasiopa subnubila</taxonomicName>
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but is distinguished by the generally larger body size and by having the basal flagellomere black and the abdomen dull to subshiny (posterior tergites not as shiny as those of
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<taxonomicName class="Insecta" family="Ephydridae" genus="Gymnoclasiopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gymnoclasiopa subnubila" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="subnubila">Gymnoclasiopa subnubila</taxonomicName>
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).
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</paragraph>
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<caption pageId="27" pageNumber="28">
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<paragraph pageId="27" pageNumber="28">
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Figures 29-30.
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<taxonomicName class="Insecta" family="Ephydridae" genus="Gymnoclasiopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gymnoclasiopa grecorum" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="grecorum">Gymnoclasiopa grecorum</taxonomicName>
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Mathis and Zatwarnicki, new species (USA. Alaska. Juneau: Thane Road). 29 head, anterior view 30 same, lateral view.
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</paragraph>
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</caption>
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<caption pageId="27" pageNumber="28">
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<paragraph pageId="27" pageNumber="28">
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Figures 31-33.
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<taxonomicName class="Insecta" family="Ephydridae" genus="Gymnoclasiopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gymnoclasiopa grecorum" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="grecorum">Gymnoclasiopa grecorum</taxonomicName>
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Mathis and Zatwarnicki, new species (USA. Alaska. Juneau: Thane Road). 31 epandrium and cerci, posterior view 32 internal structures of male terminalia (aedeagus [shaded], phallapodeme, gonite, hypandrium), ventral view 33 same, lateral view. Scale bar = 0.1 mm.
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</paragraph>
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</caption>
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<caption pageId="27" pageNumber="28">
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<paragraph pageId="27" pageNumber="28">
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Figure 34. Distribution map of
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<taxonomicName class="Insecta" family="Ephydridae" genus="Gymnoclasiopa" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Gymnoclasiopa grecorum" order="Diptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="grecorum">Gymnoclasiopa grecorum</taxonomicName>
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Mathis and Zatwarnicki sp. n.
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</paragraph>
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</caption>
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</subSubSection>
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</treatment>
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</document> |