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<mods:title id="F9FC8DE498A6E7CEDD3D1FC747227137">A comprehensive taxonomic list of brittle stars (Echinodermata: Ophiuroidea) from submarine caves of the Ryukyu Islands, southwestern Japan, with a description of a rare species, Dougaloplus echinatus (Amphiuridae)</mods:title>
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<mods:namePart id="D4544E7A898CB35E2A1DEAA79240CF9B">Okanishi, Masanori</mods:namePart>
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<treatment id="6551470CFFBBFF98FF04DAD8E04AFE93" ID-DOI="http://doi.org/10.5281/zenodo.5924547" ID-GBIF-Taxon="154494653" ID-Zenodo-Dep="5924547" LSID="urn:lsid:plazi:treatment:6551470CFFBBFF98FF04DAD8E04AFE93" httpUri="http://treatment.plazi.org/id/6551470CFFBBFF98FF04DAD8E04AFE93" lastPageId="11" lastPageNumber="84" pageId="10" pageNumber="83">
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<heading id="B60F4176FFBBFF99FF04DAD8E7B1F8FF" bold="true" box="[151,740,1774,1800]" fontSize="11" level="1" pageId="10" pageNumber="83" reason="1">
<taxonomicName id="2AF88D99FFBBFF99FF04DAD8E7B1F8FF" authority="A. M. Clark, 1953" authorityName="A. M. Clark" authorityYear="1953" box="[151,740,1774,1800]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="83" phylum="Echinodermata" rank="subSpecies" species="dubia" subSpecies="amoyensis">
<emphasis id="DF8C2A08FFBBFF99FF04DAD8E7B1F8FF" bold="true" box="[151,740,1774,1800]" pageId="10" pageNumber="83">
<emphasis id="DF8C2A08FFBBFF99FF04DAD8E4AEF8FF" bold="true" box="[151,507,1774,1800]" italics="true" pageId="10" pageNumber="83">Ophionereis dubia amoyensis</emphasis>
<bibRefCitation id="89698BEBFFBBFF99FD90DAD8E7B1F8FF" author="Clark, A. M." box="[515,740,1774,1800]" pageId="10" pageNumber="83" pagination="65 - 94" refId="ref11381" refString="Clark, A. M. (1953) A revision of the genus Ophionereis (Echinodermata, Ophiuroidea). Proceedings of the Zoological Society of London, 123, 65 - 94." type="journal article" year="1953">A. M. Clark, 1953</bibRefCitation>
</emphasis>
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<paragraph id="ED47F61AFFBBFF99FF04DB24E795F8DD" blockId="10.[151,740,1774,1870]" box="[151,704,1809,1834]" pageId="10" pageNumber="83">[New Japanese name: Amoi-amime-kumohitode]</paragraph>
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<paragraph id="ED47F61AFFBBFF99FF04DB03E5ABF8B9" blockId="10.[151,740,1774,1870]" box="[151,254,1845,1870]" pageId="10" pageNumber="83">
(
<figureCitation id="75C3EA9FFFBBFF99FF0CDB03E5A0F8B9" box="[159,245,1845,1870]" captionStart="FIGURE 8" captionStartId="18.[151,247,1881,1903]" captionTargetBox="[179,1436,183,1852]" captionTargetId="figure@18.[150,1436,183,1869]" captionTargetPageId="18" captionText="FIGURE 8. Aboral views of living specimens. A, Ophiothrix purpurea von Martens, 1867 (RUMF-ZE-02042); B, Ophiothrix aff. picteti (RUMF-ZE-02044); C, Ophionereis dubia amoyensis A. M. Clark, 1953 (RUMF-ZE-02046); D, Ophionereis fusca Brock, 1888 (RUMF-ZE-02049); E, Ophionereis porrecta Lyman, 1860 (RUMF-ZE-02054); F, Ophionereis tigris H. L. Clark, 1938 (RUMF-ZE-02045); G, Ophiolepis cavitata Okanishi &amp; Fujita, 2018a (RUMF-ZE-02059); H, Ophiolepis cincta Müller &amp; Troschel, 1842 (RUMF-ZE-02058)." httpUri="https://zenodo.org/record/2605992/files/figure.png" pageId="10" pageNumber="83">Fig. 8C</figureCitation>
)
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<taxonomicName id="2AF88D99FFBBFF99FF04DB4BE736F864" authority="A. M. Clark 1953: 88" authorityName="A. M. Clark" authorityPageNumber="88" authorityYear="1953" box="[151,611,1917,1939]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="83" phylum="Echinodermata" rank="species" species="amoyensis">
<emphasis id="DF8C2A08FFBBFF99FF04DB4BE4D6F864" box="[151,387,1917,1939]" italics="true" pageId="10" pageNumber="83">Ophionereis amoyensis</emphasis>
<treatmentCitation id="6C59D00BFFBBFF99FE1ADB48E736F864" author="Clark, A. M." box="[393,611,1917,1939]" page="88" pageId="10" pageNumber="83" year="1953">
<bibRefCitation id="89698BEBFFBBFF99FE1ADB48E714F864" author="Clark, A. M." box="[393,577,1917,1939]" pageId="10" pageNumber="83" pagination="65 - 94" refId="ref11381" refString="Clark, A. M. (1953) A revision of the genus Ophionereis (Echinodermata, Ophiuroidea). Proceedings of the Zoological Society of London, 123, 65 - 94." type="journal article" year="1953">A. M. Clark 1953</bibRefCitation>
: 88
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–90, text-fig. 11.—
<treatmentCitation id="6C59D00BFFBBFF99FCB5DB4BE108F864" author="Clark, A. M. &amp; Rowe, F. W. E." box="[806,1117,1917,1939]" page="88" pageId="10" pageNumber="83" year="1971">
<bibRefCitation id="89698BEBFFBBFF99FCB5DB4BE16DF864" author="Clark, A. M. &amp; Rowe, F. W. E." box="[806,1080,1917,1939]" pageId="10" pageNumber="83" refId="ref11804" refString="Clark, A. M. &amp; Rowe, F. W. E. (1971) 690 Trustees of the British Museum (Natural History) Monograph of shallow-water Indo- West Pacific Echinoderms. British Museum (Naturak History), London." type="book" year="1971">A. M. Clark &amp; Rowe 1971</bibRefCitation>
: 88
</treatmentCitation>
, 122.—
<treatmentCitation id="6C59D00BFFBBFF99FB23DB4BE0C2F864" author="Clark, A. M." box="[1200,1431,1917,1939]" page="488" pageId="10" pageNumber="83" year="1982">
<bibRefCitation id="89698BEBFFBBFF99FB23DB4BE033F864" author="Clark, A. M." box="[1200,1382,1917,1939]" pageId="10" pageNumber="83" pagination="485 - 501" refId="ref11665" refString="Clark, A. M. (1982) Echinoderms of Hong Kong. In: Morton, B. &amp; Tseng, C. K. (Eds.), Marine Fauna and Flora of Hong Kong. Hong Kong University Press, Hong Kong, pp. 485 - 501." type="book chapter" year="1982">A. M. Clark 1982</bibRefCitation>
: 488
</treatmentCitation>
, 492.—
<treatmentCitation id="6C59D00BFFBBFF99FE9FDBAAE773F845" author="O'Hara, T. D. &amp; Byrne, M." box="[268,550,1948,1970]" page="333" pageId="10" pageNumber="83" year="2017">
<bibRefCitation id="89698BEBFFBBFF99FE9FDBAAE4A6F845" author="O'Hara, T. D. &amp; Byrne, M." box="[268,499,1948,1970]" pageId="10" pageNumber="83" refId="ref15658" refString="O'Hara, T. D. &amp; Byrne, M. (2017) Australian Echinoderms: Biology, Ecology &amp; Evolution. CSIRO Publishing, Clayton, pp. 612." type="book" year="2017">O’Hara &amp; Byrne 2017</bibRefCitation>
: 333
</treatmentCitation>
.
<emphasis id="DF8C2A08FFBBFF99FF04DB8CE49FF827" box="[151,458,1978,2000]" italics="true" pageId="10" pageNumber="83">
<taxonomicName id="2AF88D99FFBBFF99FF04DB8CE493F827" authorityName="A. M. Clark" authorityYear="1953" box="[151,454,1978,2000]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="10" pageNumber="83" phylum="Echinodermata" rank="subSpecies" species="dubia" subSpecies="amoyensis">Ophionereis dubia amoyensis</taxonomicName>
.
</emphasis>
<treatmentCitation id="6C59D00BFFBBFF99FE5CDB8CE652F827" author="Liao, Y. &amp; Clark, A. M." box="[463,775,1978,2000]" page="273" pageId="10" pageNumber="83" year="1995">
<bibRefCitation id="89698BEBFFBBFF99FE5CDB8CE780F827" author="Liao, Y. &amp; Clark, A. M." box="[463,725,1978,2000]" pageId="10" pageNumber="83" refId="ref14200" refString="Liao, Y. &amp; Clark, A. M. (1995) The echinoderms of southern China. Science Press, Beijing, New York." type="book" year="1995">Liao &amp; A. M. Clark 1995</bibRefCitation>
: 273
</treatmentCitation>
, fig. 150.
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</paragraph>
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<subSubSection id="A5E2A591FFBAFF98FF04DCA1E611FF23" pageId="11" pageNumber="84" type="materials_examined">
<paragraph id="ED47F61AFFBAFF98FF04DCA1E611FF23" blockId="11.[151,1437,151,357]" pageId="11" pageNumber="84">
<emphasis id="DF8C2A08FFBAFF98FF04DCA1E4D6FF47" bold="true" box="[151,387,151,176]" pageId="11" pageNumber="84">Material examined.</emphasis>
Ie
<collectingCountry id="95EFB68AFFBAFF98FE3FDCA1E4A5FF47" box="[428,496,151,176]" name="Iceland" pageId="11" pageNumber="84">Island</collectingCountry>
(RUMF-ZE-02046[1], RUMF-ZE-02047[1]): “entrance” of the “Unnamed cave”, under coral rubble, approximately
<quantity id="2A005BFFFFBAFF98FD88DC8AE701FF23" box="[539,596,188,212]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="84" unit="m" value="20.0">20 m</quantity>
depth,
<date id="9946D0DAFFBAFF98FD34DC8AE615FF23" box="[679,832,188,212]" pageId="11" pageNumber="84" value="2017-06-24">24 June 2017</date>
.
</paragraph>
</subSubSection>
<subSubSection id="A5E2A591FFBAFF98FF54DCE9E674FF00" box="[199,801,223,248]" pageId="11" pageNumber="84" type="distribution">
<paragraph id="ED47F61AFFBAFF98FF54DCE9E674FF00" blockId="11.[151,1437,151,357]" box="[199,801,223,248]" pageId="11" pageNumber="84">
<emphasis id="DF8C2A08FFBAFF98FF54DCE9E435FF0F" bold="true" box="[199,352,223,248]" pageId="11" pageNumber="84">Distribution.</emphasis>
<collectingCountry id="95EFB68AFFBAFF98FEF4DCE9E4FFFF0F" box="[359,426,223,248]" name="China" pageId="11" pageNumber="84">China</collectingCountry>
to Southern
<collectingCountry id="95EFB68AFFBAFF98FDA9DCD6E728FF00" box="[570,637,224,247]" name="Japan" pageId="11" pageNumber="84">Japan</collectingCountry>
. Depth
<specimenCount id="FBFE3D93FFBAFF98FD40DCD6E674FF00" box="[723,801,224,248]" pageId="11" pageNumber="84" type="generic">
<quantity id="2A005BFFFFBAFF98FD40DCD6E648FF00" box="[723,797,224,248]" metricMagnitude="2" metricUnit="m" metricValue="1.42" pageId="11" pageNumber="84" unit="m" value="142.0">142 m</quantity>
.
</specimenCount>
</paragraph>
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<subSubSection id="A5E2A591FFBAFF98FF54DD32E04AFE93" pageId="11" pageNumber="84" type="discussion">
<paragraph id="ED47F61AFFBAFF98FF54DD32E04AFE93" blockId="11.[151,1437,151,357]" pageId="11" pageNumber="84">
<emphasis id="DF8C2A08FFBAFF98FF54DD32E46EFEEA" bold="true" box="[199,315,260,285]" pageId="11" pageNumber="84">Remarks.</emphasis>
This species is identified as
<taxonomicName id="2AF88D99FFBAFF98FD03DD32E6B6FEEB" authorityName="A. M. Clark" authorityYear="1953" box="[656,995,260,284]" class="Ophiuroidea" family="Ophionereididae" genus="Ophionereis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ophiurida" pageId="11" pageNumber="84" phylum="Echinodermata" rank="subSpecies" species="dubia" subSpecies="amoyensis">
<emphasis id="DF8C2A08FFBAFF98FD03DD32E6B6FEEB" box="[656,995,260,284]" italics="true" pageId="11" pageNumber="84">Ophionereis dubia amoyensis</emphasis>
</taxonomicName>
by virtue of: lacking genital papillae; having markedly wider dorsal arm plates, three times as side as long scales; and large supplementary dorsal arm plates (e.g.
<bibRefCitation id="89698BEBFFBAFF98FE8BDD7AE71BFE93" author="Clark, A. M. &amp; Rowe, F. W. E." box="[280,590,332,357]" pageId="11" pageNumber="84" refId="ref11804" refString="Clark, A. M. &amp; Rowe, F. W. E. (1971) 690 Trustees of the British Museum (Natural History) Monograph of shallow-water Indo- West Pacific Echinoderms. British Museum (Naturak History), London." type="book" year="1971">A. M. Clark &amp; Rowe, 1971</bibRefCitation>
). This is the first record of the subspecies from Japanese waters.
</paragraph>
</subSubSection>
</treatment>
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