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<mods:title id="F8971922A94222F291D43FD5AA07D334">Two new species of Leptanilloides Mann, 1823 (Formicidae: Dorylinae) from the Andes of southern Ecuador</mods:title>
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<mods:namePart id="5FDCA39C5FC3EDFDD7A843E86AB466CC">Sonet, Gontran</mods:namePart>
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<mods:affiliation id="C08E1DD1B2FA7EA64FEAF55F388C986F">Universidad Técnica Particular de Loja (UTPL), Departamento de Ciencias Naturales, Museo de Colecciones Biológicas (MUTPL), Loja, Ecuador. &amp; Universidad de Cuenca, Facultad de Ciencias, Agropecuarias, Ave. 12 de Abril s / n, Cuenca, Ecuador &amp; E-mail: david. donosov @ gmail. com &amp; urn: lsid: zoobank. org: author: BD 09 BA 33 - A 9 F 8 - 4 F 77 - A 5 B 3 - F 277062 FB 286</mods:affiliation>
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<paragraph id="904C36A5FFD64072FD9D702108E8FABF" blockId="4.[592,995,1366,1393]" box="[592,995,1366,1393]" pageId="4" pageNumber="5">
<heading id="CB0481C9FFD64072FD9D702108E8FABF" box="[592,995,1366,1393]" centered="true" fontSize="11" level="2" pageId="4" pageNumber="5" reason="2">
Genus 
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<emphasis id="A287EAB7FFD64072FD6E70200845FABF" bold="true" box="[675,846,1366,1393]" italics="true" pageId="4" pageNumber="5">Leptanilloides</emphasis>
Mann, 1823
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<heading id="CB0481C9FFD64072FF7070EE0875FA7C" bold="true" box="[189,894,1432,1459]" fontSize="11" level="3" pageId="4" pageNumber="5" reason="3">
<emphasis id="A287EAB7FFD64072FF7070EE0875FA7C" bold="true" box="[189,894,1432,1459]" pageId="4" pageNumber="5">
Diagnosis of the 
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<emphasis id="A287EAB7FFD64072FE4670EE09C7FA7D" bold="true" box="[395,716,1432,1459]" italics="true" pageId="4" pageNumber="5">Leptanilloides biconstricta</emphasis>
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species-group
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<paragraph id="904C36A5FFD64072FF7070B00ABFF934" blockId="4.[189,1399,1478,1786]" pageId="4" pageNumber="5">
Species from the 
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<emphasis id="A287EAB7FFD64072FE4370B109C1FA2F" box="[398,714,1479,1505]" italics="true" pageId="4" pageNumber="5">Leptanilloides biconstricta</emphasis>
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species-group possess the typical characters of the genus (see diagnosis in 
<bibRefCitation id="F4624B54FFD64072FE49709C09B6F9CA" author="Borowiec M. L. &amp; Longino J. T." box="[388,701,1514,1540]" pageId="4" pageNumber="5" pagination="19 - 48" refId="ref14112" refString="Borowiec M. L. &amp; Longino J. T. 2011. Three new species and reassessment of the rare Neotropical ant genus Leptanilloides (Hymenoptera, Formicidae, Leptanilloidinae). ZooKeys 133: 19 - 48. http: // dx. doi. org / 10.3897 / zookeys. 133.1479" type="journal article" year="2011">Borowiec &amp; Longino 2011</bibRefCitation>
) but can be grouped by the fact that they share the following criteria: promesonotal connection completely unfused and flexible; lateroclypeal tooth (called genal tooth in 
<bibRefCitation id="F4624B54FFD64072FED673460AF4F984" author="Brandao C. R. F. &amp; Diniz J. L. M. &amp; Agosti D. &amp; Delabie J. H. C." box="[283,511,1584,1610]" pageId="4" pageNumber="5" pagination="17 - 36" refId="ref14350" refString="Brandao C. R. F., Diniz J. L. M., Agosti D. &amp; Delabie J. H. C. 1999. Revision of the Neotropical ant subfamily Leptanilloidinae. Systematic Entomology 24: 17 - 36. http: // dx. doi. org / 10.1046 / j. 1365 - 3113.1999.00064. x" type="journal article" year="1999">
Brandão 
<emphasis id="A287EAB7FFD64072FE4973470AB6F984" box="[388,445,1584,1610]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
1999
</bibRefCitation>
, 
<bibRefCitation id="F4624B54FFD64072FDC4734609A6F984" author="Longino J. T." box="[521,685,1584,1610]" pageId="4" pageNumber="5" pagination="1 - 6" refId="ref15223" refString="Longino J. T. 2003. A new Costa Rican species of Leptanilloides (Hymenoptera: Formicidae: doryline section: Leptanilloidinae). Zootaxa 264: 1 - 6." type="journal article" year="2003">Longino 2003</bibRefCitation>
and 
<bibRefCitation id="F4624B54FFD64072FD2F734708B6F985" author="Donoso D. A. &amp; Vieira J. M. &amp; Wild A. L." box="[738,957,1584,1611]" pageId="4" pageNumber="5" pagination="47 - 62" refId="ref14648" refString="Donoso D. A., Vieira J. M. &amp; Wild A. L. 2006. Three new species of Leptanilloides Mann from Andean Ecuador (Formicidade: Leptanilloidinae). Zootaxa 1201: 47 - 62." type="journal article" year="2006">
Donoso 
<emphasis id="A287EAB7FFD64072FC8E73470870F984" box="[835,891,1584,1610]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
2006
</bibRefCitation>
) well-developed (although apparently lacking in 
<taxonomicName id="57F34D26FFD64072FEF4732209BAF9A3" authority="Donoso et al., 2006" authorityName="Donoso" authorityYear="2006" box="[313,689,1619,1646]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="caracola">
<emphasis id="A287EAB7FFD64072FEF473220ACBF9A3" box="[313,448,1619,1645]" italics="true" pageId="4" pageNumber="5">L. caracola</emphasis>
<bibRefCitation id="F4624B54FFD64072FE0A732209BAF9A3" author="Donoso D. A. &amp; Vieira J. M. &amp; Wild A. L." box="[455,689,1619,1646]" pageId="4" pageNumber="5" pagination="47 - 62" refId="ref14648" refString="Donoso D. A., Vieira J. M. &amp; Wild A. L. 2006. Three new species of Leptanilloides Mann from Andean Ecuador (Formicidade: Leptanilloidinae). Zootaxa 1201: 47 - 62." type="journal article" year="2006">
Donoso 
<emphasis id="A287EAB7FFD64072FDE77322096EF9A3" box="[554,613,1619,1645]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
, 2006
</bibRefCitation>
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, a species known only by its 
<typeStatus id="4F488807FFD64072FBCA73250F7AF9A3" box="[1031,1137,1619,1645]" pageId="4" pageNumber="5" type="holotype">holotype</typeStatus>
; because this criterion may be difficult to observe with a stereo microscope, even at high magnification, the absence of a lateroclypeal tooth in 
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<emphasis id="A287EAB7FFD64072FE0973ED0946F97A" box="[452,589,1690,1716]" italics="true" pageId="4" pageNumber="5">L. caracola</emphasis>
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should be confirmed in SEM); abdominal segment III (postpetiole) nearly as high as abdominal segment IV; and postpetiolar spiracle situated proximately to the anterior margin of the tergite.
</paragraph>
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<paragraph id="904C36A5FFD64072FF7072540A17F891" blockId="4.[189,1399,1825,1887]" pageId="4" pageNumber="5">
Moreover, species from this group are generally smaller than species of the 
<taxonomicName id="57F34D26FFD64072FBFE72540FC7F8F2" box="[1075,1228,1826,1852]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="legionaria">
<emphasis id="A287EAB7FFD64072FBFE72540FC7F8F2" box="[1075,1228,1826,1852]" italics="true" pageId="4" pageNumber="5">L. legionaria</emphasis>
</taxonomicName>
species-group (
<figureCitation id="08C82A20FFD64072FF0B72330A00F891" box="[198,267,1861,1887]" captionStart="Fig" captionStartId="5.[190,233,1718,1744]" captionTargetBox="[209,1357,966,1683]" captionTargetId="figure@5.[173,1383,946,1714]" captionTargetPageId="5" captionText="Fig. 1. Relationship between HL and WL among workers in species of Leptanilloides and Asphinctanilloides. Specimens measured are holotypes or paratypes. Measurements are in mm. WL is called Mesosoma Length (ML) in Borowiec &amp; Longino (2011) and Silva et al. (2013). Triangles: the two Leptanilloides described in this paper (L. copalinga Delsinne &amp; Donoso sp. nov. and L. prometea Delsinne &amp; Donoso sp. nov.); diamonds: Leptanilloides biconstricta species-group (La: L. atlantica, Lb: L. biconstricta, Lc: L. caracola, Le: L. erinys, Lf: L. femoralis, Lg: L. gracilis, Li: L. improvisa, Ls: L. sculpturata); dots: Leptanilloides legionaria species-group (Ll: L. legionaria, Lm: L. mckennae, Lno: L. nomada, Lnu: L. nubecula); stars: Asphinctanilloides species (Aa: A. amazon Brandão et al., 1999, Aan: A. anae Brandão et al., 1999, Am: A. manauara Brandão et al., 1999)." figureDoi="http://doi.org/10.5281/zenodo.3787362" httpUri="https://zenodo.org/record/3787362/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="904C36A5FFD64072FF7072F009D1F86E" blockId="4.[189,730,1926,2023]" box="[189,730,1926,1953]" pageId="4" pageNumber="5">This group includes the ten following species:</paragraph>
<paragraph id="904C36A5FFD64072FF7072DC091FF80A" blockId="4.[189,730,1926,2023]" box="[189,532,1961,1988]" pageId="4" pageNumber="5">
<taxonomicName id="57F34D26FFD64072FF7072DC091FF80A" authority="Silva et al., 2013" authorityName="Silva" authorityYear="2013" box="[189,532,1961,1988]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="atlantica">
<emphasis id="A287EAB7FFD64072FF7072DC0A4FF80D" box="[189,324,1961,1987]" italics="true" pageId="4" pageNumber="5">L. atlantica</emphasis>
<bibRefCitation id="F4624B54FFD64072FE8172DF091FF80A" author="Silva R. R. &amp; Feitosa R. M. &amp; Brandao C. R. F. &amp; Freitas A. V. L." box="[332,532,1961,1988]" pageId="4" pageNumber="5" pagination="2039 - 2047" refId="ref15882" refString="Silva R. R., Feitosa R. M., Brandao C. R. F. &amp; Freitas A. V. L. 2013. The first Leptanilloides species (Hymenoptera: Formicidae: Leptanilloidinae) from eastern South America. Journal of Natural History 47 (31 - 32): 2039 - 2047. http: // dx. doi. org / 10.1080 / 00222933.2012.763058" type="journal article" year="2013">
Silva 
<emphasis id="A287EAB7FFD64072FE4272DC0AC1F80D" box="[399,458,1961,1987]" italics="true" pageId="4" pageNumber="5">et al.</emphasis>
, 2013
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</taxonomicName>
</paragraph>
<paragraph id="904C36A5FFD64072FF7072BB0AF5F829" blockId="4.[189,730,1926,2023]" box="[189,510,1997,2023]" pageId="4" pageNumber="5">
<taxonomicName id="57F34D26FFD64072FF7072BB0AF5F829" authority="Mann, 1923" authorityName="Mann" authorityYear="1923" box="[189,510,1997,2023]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="biconstricta">
<emphasis id="A287EAB7FFD64072FF7072BB0A63F829" box="[189,360,1997,2023]" italics="true" pageId="4" pageNumber="5">L. biconstricta</emphasis>
<bibRefCitation id="F4624B54FFD64072FEA272BB0AF5F829" author="Mann W. M." box="[367,510,1997,2023]" pageId="4" pageNumber="5" pagination="13 - 18" refId="ref15255" refString="Mann W. M. 1923. Two new ants from Bolivia. Psyche 30: 13 - 18. http: // dx. doi. org / 10.1155 / 1923 / 58289" type="journal article" year="1923">Mann, 1923</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="904C36A5FFD74073FF70747A0939FEEB" blockId="5.[189,676,267,540]" box="[189,562,267,294]" pageId="5" pageNumber="6">
<taxonomicName id="57F34D26FFD74073FF70747A0939FEEB" authority="Donoso et al., 2006" authorityName="Donoso" authorityYear="2006" box="[189,562,267,294]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="caracola">
<emphasis id="A287EAB7FFD74073FF70747A0A4FFEEB" box="[189,324,267,293]" italics="true" pageId="5" pageNumber="6">L. caracola</emphasis>
<bibRefCitation id="F4624B54FFD74073FE81747A0939FEEB" author="Donoso D. A. &amp; Vieira J. M. &amp; Wild A. L." box="[332,562,267,294]" pageId="5" pageNumber="6" pagination="47 - 62" refId="ref14648" refString="Donoso D. A., Vieira J. M. &amp; Wild A. L. 2006. Three new species of Leptanilloides Mann from Andean Ecuador (Formicidade: Leptanilloidinae). Zootaxa 1201: 47 - 62." type="journal article" year="2006">
Donoso 
<emphasis id="A287EAB7FFD74073FE63747A0AE2FEEB" box="[430,489,267,293]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
, 2006
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</paragraph>
<paragraph id="904C36A5FFD74073FF70745909AFFE87" blockId="5.[189,676,267,540]" box="[189,676,302,329]" pageId="5" pageNumber="6">
<taxonomicName id="57F34D26FFD74073FF707459094AFE87" authority="Delsinne &amp; Donoso" authorityName="Delsinne &amp; Sonet &amp; Donoso" authorityYear="2015" box="[189,577,302,329]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="copalinga" status="sp. nov.">
<emphasis id="A287EAB7FFD74073FF7074590A5AFE87" box="[189,337,303,329]" italics="true" pageId="5" pageNumber="6">L. copalinga</emphasis>
Delsinne &amp; Donoso
</taxonomicName>
<taxonomicNameLabel id="B9B457CCFFD74073FD84745909AFFE87" box="[585,676,303,329]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="904C36A5FFD74073FF7074250964FEA2" blockId="5.[189,676,267,540]" box="[189,623,338,364]" pageId="5" pageNumber="6">
<taxonomicName id="57F34D26FFD74073FF7074250964FEA2" authority="Borowiec &amp; Longino, 2011" authorityName="Borowiec &amp; Longino" authorityYear="2011" box="[189,623,338,364]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="erinys">
<emphasis id="A287EAB7FFD74073FF7074250A2FFEA2" box="[189,292,339,364]" italics="true" pageId="5" pageNumber="6">L. erinys</emphasis>
<bibRefCitation id="F4624B54FFD74073FEE674240964FEA2" author="Borowiec M. L. &amp; Longino J. T." box="[299,623,338,364]" pageId="5" pageNumber="6" pagination="19 - 48" refId="ref14112" refString="Borowiec M. L. &amp; Longino J. T. 2011. Three new species and reassessment of the rare Neotropical ant genus Leptanilloides (Hymenoptera, Formicidae, Leptanilloidinae). ZooKeys 133: 19 - 48. http: // dx. doi. org / 10.3897 / zookeys. 133.1479" type="journal article" year="2011">Borowiec &amp; Longino, 2011</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="904C36A5FFD74073FF707400099DFE41" blockId="5.[189,676,267,540]" box="[189,662,373,399]" pageId="5" pageNumber="6">
<taxonomicName id="57F34D26FFD74073FF707400099DFE41" authority="Borowiec &amp; Longino, 2011" authorityName="Borowiec &amp; Longino" authorityYear="2011" box="[189,662,373,399]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="A287EAB7FFD74073FF7074000A40FE41" box="[189,331,373,399]" italics="true" pageId="5" pageNumber="6">L. femoralis</emphasis>
<bibRefCitation id="F4624B54FFD74073FE9F7403099DFE41" author="Borowiec M. L. &amp; Longino J. T." box="[338,662,373,399]" pageId="5" pageNumber="6" pagination="19 - 48" refId="ref14112" refString="Borowiec M. L. &amp; Longino J. T. 2011. Three new species and reassessment of the rare Neotropical ant genus Leptanilloides (Hymenoptera, Formicidae, Leptanilloidinae). ZooKeys 133: 19 - 48. http: // dx. doi. org / 10.3897 / zookeys. 133.1479" type="journal article" year="2011">Borowiec &amp; Longino, 2011</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="904C36A5FFD74073FF7074EF098AFE7C" blockId="5.[189,676,267,540]" box="[189,641,408,434]" pageId="5" pageNumber="6">
<taxonomicName id="57F34D26FFD74073FF7074EF098AFE7C" authority="Borowiec &amp; Longino, 2011" authorityName="Borowiec &amp; Longino" authorityYear="2011" box="[189,641,408,434]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="A287EAB7FFD74073FF7074EF0A3DFE7C" box="[189,310,408,434]" italics="true" pageId="5" pageNumber="6">L. gracilis</emphasis>
<bibRefCitation id="F4624B54FFD74073FEF074EE098AFE7C" author="Borowiec M. L. &amp; Longino J. T." box="[317,641,408,434]" pageId="5" pageNumber="6" pagination="19 - 48" refId="ref14112" refString="Borowiec M. L. &amp; Longino J. T. 2011. Three new species and reassessment of the rare Neotropical ant genus Leptanilloides (Hymenoptera, Formicidae, Leptanilloidinae). ZooKeys 133: 19 - 48. http: // dx. doi. org / 10.3897 / zookeys. 133.1479" type="journal article" year="2011">Borowiec &amp; Longino, 2011</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="904C36A5FFD74073FF7074CA094CFE1B" blockId="5.[189,676,267,540]" box="[189,583,443,470]" pageId="5" pageNumber="6">
<taxonomicName id="57F34D26FFD74073FF7074CA094CFE1B" authority="Brandao et al., 1999" authorityName="Brandao" authorityYear="1999" box="[189,583,443,470]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="improvisa">
<emphasis id="A287EAB7FFD74073FF7074CA0A5BFE1B" box="[189,336,444,469]" italics="true" pageId="5" pageNumber="6">L. improvisa</emphasis>
<bibRefCitation id="F4624B54FFD74073FE9574CD094CFE1B" author="Brandao C. R. F. &amp; Diniz J. L. M. &amp; Agosti D. &amp; Delabie J. H. C." box="[344,583,443,470]" pageId="5" pageNumber="6" pagination="17 - 36" refId="ref14350" refString="Brandao C. R. F., Diniz J. L. M., Agosti D. &amp; Delabie J. H. C. 1999. Revision of the Neotropical ant subfamily Leptanilloidinae. Systematic Entomology 24: 17 - 36. http: // dx. doi. org / 10.1046 / j. 1365 - 3113.1999.00064. x" type="journal article" year="1999">
Brandão 
<emphasis id="A287EAB7FFD74073FE0F74CA0AF6FE1B" box="[450,509,443,469]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
, 1999
</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="904C36A5FFD74073FF7074A90997FE37" blockId="5.[189,676,267,540]" box="[189,668,478,505]" pageId="5" pageNumber="6">
<taxonomicName id="57F34D26FFD74073FF7074A90931FE37" authority="Delsinne &amp; Donoso" authorityName="Delsinne &amp; Sonet &amp; Donoso" authorityYear="2015" box="[189,570,478,505]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="prometea" status="sp. nov.">
<emphasis id="A287EAB7FFD74073FF7074A90A41FE36" box="[189,330,479,504]" italics="true" pageId="5" pageNumber="6">L. prometea</emphasis>
Delsinne &amp; Donoso
</taxonomicName>
<taxonomicNameLabel id="B9B457CCFFD74073FD8C74A90997FE37" box="[577,668,479,505]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
<paragraph id="904C36A5FFD74073FF7077750953FDD2" blockId="5.[189,676,267,540]" box="[189,600,514,540]" pageId="5" pageNumber="6">
<taxonomicName id="57F34D26FFD74073FF7077750953FDD2" authority="Brandao et al., 1999" authorityName="Brandao" authorityYear="1999" box="[189,600,514,540]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sculpturata">
<emphasis id="A287EAB7FFD74073FF7077750A69FDD2" box="[189,354,514,540]" italics="true" pageId="5" pageNumber="6">L. sculpturata</emphasis>
<bibRefCitation id="F4624B54FFD74073FEA477740953FDD2" author="Brandao C. R. F. &amp; Diniz J. L. M. &amp; Agosti D. &amp; Delabie J. H. C." box="[361,600,514,540]" pageId="5" pageNumber="6" pagination="17 - 36" refId="ref14350" refString="Brandao C. R. F., Diniz J. L. M., Agosti D. &amp; Delabie J. H. C. 1999. Revision of the Neotropical ant subfamily Leptanilloidinae. Systematic Entomology 24: 17 - 36. http: // dx. doi. org / 10.1046 / j. 1365 - 3113.1999.00064. x" type="journal article" year="1999">
Brandão 
<emphasis id="A287EAB7FFD74073FE1977750905FDD2" box="[468,526,514,540]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
, 1999
</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D8E9652EFFD74073FF7077350E46FD3B" pageId="5" pageNumber="6" type="diagnosis">
<paragraph id="904C36A5FFD74073FF7077350867FD93" blockId="5.[189,876,579,606]" box="[189,876,579,606]" pageId="5" pageNumber="6">
<heading id="CB0481C9FFD74073FF7077350867FD93" bold="true" box="[189,876,579,606]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="A287EAB7FFD74073FF7077350867FD93" bold="true" box="[189,876,579,606]" pageId="5" pageNumber="6">
Diagnosis of the 
<taxonomicName id="57F34D26FFD74073FE46773509B1FD90" box="[395,698,579,606]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="legionaria">
<emphasis id="A287EAB7FFD74073FE46773509B1FD90" bold="true" box="[395,698,579,606]" italics="true" pageId="5" pageNumber="6">Leptanilloides legionaria</emphasis>
</taxonomicName>
species-group
</emphasis>
</heading>
</paragraph>
<paragraph id="904C36A5FFD74073FF7077040E46FD3B" blockId="5.[189,1399,626,758]" pageId="5" pageNumber="6">
Species from the 
<taxonomicName id="57F34D26FFD74073FE57770409C2FD42" box="[410,713,626,652]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="legionaria">
<emphasis id="A287EAB7FFD74073FE57770409C2FD42" box="[410,713,626,652]" italics="true" pageId="5" pageNumber="6">Leptanilloides legionaria</emphasis>
</taxonomicName>
species-group possess the characters of the genus (see diagnosis in 
<bibRefCitation id="F4624B54FFD74073FE9477E309A9FD61" author="Borowiec M. L. &amp; Longino J. T." box="[345,674,661,687]" pageId="5" pageNumber="6" pagination="19 - 48" refId="ref14112" refString="Borowiec M. L. &amp; Longino J. T. 2011. Three new species and reassessment of the rare Neotropical ant genus Leptanilloides (Hymenoptera, Formicidae, Leptanilloidinae). ZooKeys 133: 19 - 48. http: // dx. doi. org / 10.3897 / zookeys. 133.1479" type="journal article" year="2011">Borowiec &amp; Longino 2011</bibRefCitation>
) but are grouped because they share the following criteria: promesonotal connection at least partially fused; lateroclypeal tooth absent or reduced; postpetiole reduced and isolated; and postpetiolar spiracle shifted posteriad on anteromedian side of the tergite.
</paragraph>
</subSubSection>
<subSubSection id="D8E9652EFFD74070FF70766B093DFE86" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="discussion">
<paragraph id="904C36A5FFD74073FF70766B09ECFCF9" blockId="5.[189,743,797,893]" box="[189,743,797,823]" pageId="5" pageNumber="6">This group includes the following four species:</paragraph>
<paragraph id="904C36A5FFD74073FF7076370946FC94" blockId="5.[189,743,797,893]" box="[189,589,832,858]" pageId="5" pageNumber="6">
<taxonomicName id="57F34D26FFD74073FF7076370946FC94" authority="Brandao et al., 1999" authorityName="Brandao" authorityYear="1999" box="[189,589,832,858]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="legionaria">
<emphasis id="A287EAB7FFD74073FF7076370A5DFC94" box="[189,342,832,858]" italics="true" pageId="5" pageNumber="6">L. legionaria</emphasis>
<bibRefCitation id="F4624B54FFD74073FE9376360946FC94" author="Brandao C. R. F. &amp; Diniz J. L. M. &amp; Agosti D. &amp; Delabie J. H. C." box="[350,589,832,858]" pageId="5" pageNumber="6" pagination="17 - 36" refId="ref14350" refString="Brandao C. R. F., Diniz J. L. M., Agosti D. &amp; Delabie J. H. C. 1999. Revision of the Neotropical ant subfamily Leptanilloidinae. Systematic Entomology 24: 17 - 36. http: // dx. doi. org / 10.1046 / j. 1365 - 3113.1999.00064. x" type="journal article" year="1999">
Brandão 
<emphasis id="A287EAB7FFD74073FE0576370908FC94" box="[456,515,832,858]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
, 1999
</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="904C36A5FFD74073FF707612090DFCB3" blockId="5.[189,743,797,893]" box="[189,518,867,893]" pageId="5" pageNumber="6">
<taxonomicName id="57F34D26FFD74073FF707612090DFCB3" authority="Longino, 2003" authorityName="Longino" authorityYear="2003" box="[189,518,867,893]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mckennae">
<emphasis id="A287EAB7FFD74073FF7076120A5AFCB3" box="[189,337,867,893]" italics="true" pageId="5" pageNumber="6">L. mckennae</emphasis>
<bibRefCitation id="F4624B54FFD74073FE947615090DFCB3" author="Longino J. T." box="[345,518,867,893]" pageId="5" pageNumber="6" pagination="1 - 6" refId="ref15223" refString="Longino J. T. 2003. A new Costa Rican species of Leptanilloides (Hymenoptera: Formicidae: doryline section: Leptanilloidinae). Zootaxa 264: 1 - 6." type="journal article" year="2003">Longino, 2003</bibRefCitation>
</taxonomicName>
</paragraph>
<caption id="C48C662DFFD74073FF7373C00F75F824" ID-DOI="http://doi.org/10.5281/zenodo.3787362" ID-Zenodo-Dep="3787362" httpUri="https://zenodo.org/record/3787362/files/figure.png" pageId="5" pageNumber="6" startId="5.[190,233,1718,1744]" targetBox="[209,1357,966,1683]" targetPageId="5">
<paragraph id="904C36A5FFD74073FF7373C00F75F824" blockId="5.[190,1399,1718,2026]" pageId="5" pageNumber="6">
<emphasis id="A287EAB7FFD74073FF7373C00A05F91E" bold="true" box="[190,270,1718,1744]" pageId="5" pageNumber="6">Fig. 1.</emphasis>
Relationship between HL and WL among workers in species of 
<taxonomicName id="57F34D26FFD74073FBED73C00FC3F91E" authorityName="Mann" authorityYear="1823" box="[1056,1224,1718,1744]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="A287EAB7FFD74073FBED73C00FC3F91E" box="[1056,1224,1718,1744]" italics="true" pageId="5" pageNumber="6">Leptanilloides</emphasis>
</taxonomicName>
and 
<taxonomicName id="57F34D26FFD74073FACA73C00A39F93D" class="Insecta" family="Formicidae" genus="Asphinctanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="A287EAB7FFD74073FACA73C00A39F93D" italics="true" pageId="5" pageNumber="6">Asphinctanilloides</emphasis>
</taxonomicName>
. Specimens measured are holotypes or paratypes. Measurements are in mm. WL is called Mesosoma Length (ML) in 
<bibRefCitation id="F4624B54FFD74073FDDF738B0860F8D9" author="Borowiec M. L. &amp; Longino J. T." box="[530,875,1789,1815]" pageId="5" pageNumber="6" pagination="19 - 48" refId="ref14112" refString="Borowiec M. L. &amp; Longino J. T. 2011. Three new species and reassessment of the rare Neotropical ant genus Leptanilloides (Hymenoptera, Formicidae, Leptanilloidinae). ZooKeys 133: 19 - 48. http: // dx. doi. org / 10.3897 / zookeys. 133.1479" type="journal article" year="2011">Borowiec &amp; Longino (2011)</bibRefCitation>
and 
<bibRefCitation id="F4624B54FFD74073FC61738B0F85F8D9" author="Silva R. R. &amp; Feitosa R. M. &amp; Brandao C. R. F. &amp; Freitas A. V. L." box="[940,1166,1789,1815]" pageId="5" pageNumber="6" pagination="2039 - 2047" refId="ref15882" refString="Silva R. R., Feitosa R. M., Brandao C. R. F. &amp; Freitas A. V. L. 2013. The first Leptanilloides species (Hymenoptera: Formicidae: Leptanilloidinae) from eastern South America. Journal of Natural History 47 (31 - 32): 2039 - 2047. http: // dx. doi. org / 10.1080 / 00222933.2012.763058" type="journal article" year="2013">
Silva 
<emphasis id="A287EAB7FFD74073FC3E738B0F3AF8D9" box="[1011,1073,1789,1815]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
(2013)
</bibRefCitation>
. Triangles: the two 
<taxonomicName id="57F34D26FFD74073FF7372560A6DF8F4" authorityName="Mann" authorityYear="1823" box="[190,358,1824,1850]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="A287EAB7FFD74073FF7372560A6DF8F4" box="[190,358,1824,1850]" italics="true" pageId="5" pageNumber="6">Leptanilloides</emphasis>
</taxonomicName>
described in this paper (
<taxonomicName id="57F34D26FFD74073FD5272570F3CF8F4" authority="Delsinne &amp; Donoso" authorityName="Delsinne &amp; Sonet &amp; Donoso" authorityYear="2015" box="[671,1079,1824,1850]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="copalinga" status="sp. nov.">
<emphasis id="A287EAB7FFD74073FD5272570833F8F4" box="[671,824,1824,1850]" italics="true" pageId="5" pageNumber="6">L. copalinga</emphasis>
Delsinne &amp; Donoso
</taxonomicName>
<taxonomicNameLabel id="B9B457CCFFD74073FB8972560FA8F8F4" box="[1092,1187,1824,1850]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="57F34D26FFD74073FB2872570AA7F890" authority="Delsinne &amp; Donoso" authorityName="Delsinne &amp; Sonet &amp; Donoso" authorityYear="2015" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="prometea" status="sp. nov.">
<emphasis id="A287EAB7FFD74073FB2872570E7CF8F4" box="[1253,1399,1825,1850]" italics="true" pageId="5" pageNumber="6">L. prometea</emphasis>
Delsinne &amp; Donoso
</taxonomicName>
<taxonomicNameLabel id="B9B457CCFFD74073FE7A72320913F890" box="[439,536,1860,1886]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
); diamonds: 
<taxonomicName id="57F34D26FFD74073FD78723508F8F893" authorityName="Mann" authorityYear="1923" box="[693,1011,1859,1885]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="biconstricta">
<emphasis id="A287EAB7FFD74073FD78723508F8F893" box="[693,1011,1859,1885]" italics="true" pageId="5" pageNumber="6">Leptanilloides biconstricta</emphasis>
</taxonomicName>
species-group (La: 
<taxonomicName id="57F34D26FFD74073FB2B72320BE8F84E" authority=", Lb" authorityName="Lb" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="atlantica">
<emphasis id="A287EAB7FFD74073FB2B72320E64F893" box="[1254,1391,1859,1885]" italics="true" pageId="5" pageNumber="6">L. atlantica</emphasis>
, Lb
</taxonomicName>
: 
<taxonomicName id="57F34D26FFD74073FF3D72110ADAF84F" authority=", Lc" authorityName="Lc" box="[240,465,1894,1921]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="biconstricta">
<emphasis id="A287EAB7FFD74073FF3D72110A96F84E" box="[240,413,1894,1920]" italics="true" pageId="5" pageNumber="6">L. biconstricta</emphasis>
, Lc
</taxonomicName>
: 
<taxonomicName id="57F34D26FFD74073FE1372110997F84F" authority=", Le" authorityName="Le" box="[478,668,1894,1921]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="caracola">
<emphasis id="A287EAB7FFD74073FE1372110963F84E" box="[478,616,1894,1920]" italics="true" pageId="5" pageNumber="6">L. caracola</emphasis>
, Le
</taxonomicName>
: 
<taxonomicName id="57F34D26FFD74073FD6472110848F84E" authority=", Lf" authorityName="Lf" box="[681,835,1894,1921]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="erinys">
<emphasis id="A287EAB7FFD74073FD6472110819F84E" box="[681,786,1895,1920]" italics="true" pageId="5" pageNumber="6">L. erinys</emphasis>
, Lf
</taxonomicName>
: 
<taxonomicName id="57F34D26FFD74073FC9D72110F1DF84F" authority=", Lg" authorityName="Lg" box="[848,1046,1894,1921]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="A287EAB7FFD74073FC9D721108EBF84E" box="[848,992,1894,1920]" italics="true" pageId="5" pageNumber="6">L. femoralis</emphasis>
, Lg
</taxonomicName>
: 
<taxonomicName id="57F34D26FFD74073FBEE72110FC6F84E" authority=", Li" authorityName="Li" box="[1059,1229,1894,1921]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="A287EAB7FFD74073FBEE72110F95F84E" box="[1059,1182,1894,1920]" italics="true" pageId="5" pageNumber="6">L. gracilis</emphasis>
, Li
</taxonomicName>
: 
<taxonomicName id="57F34D26FFD74073FB1772110BD4F86A" authority=", Ls" authorityName="Ls" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="improvisa">
<emphasis id="A287EAB7FFD74073FB1772110E64F84E" box="[1242,1391,1895,1920]" italics="true" pageId="5" pageNumber="6">L. improvisa</emphasis>
, Ls
</taxonomicName>
: 
<taxonomicName id="57F34D26FFD74073FF2672FC0A9BF86D" box="[235,400,1929,1955]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="sculpturata">
<emphasis id="A287EAB7FFD74073FF2672FC0A9BF86D" box="[235,400,1929,1955]" italics="true" pageId="5" pageNumber="6">L. sculpturata</emphasis>
</taxonomicName>
); dots: 
<taxonomicName id="57F34D26FFD74073FE2772FF081FF86D" box="[490,788,1929,1955]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="legionaria">
<emphasis id="A287EAB7FFD74073FE2772FF081FF86D" box="[490,788,1929,1955]" italics="true" pageId="5" pageNumber="6">Leptanilloides legionaria</emphasis>
</taxonomicName>
species-group (Ll: 
<taxonomicName id="57F34D26FFD74073FC3772FC0FDBF86A" authority=", Lm" authorityName="Lm" box="[1018,1232,1929,1956]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="legionaria">
<emphasis id="A287EAB7FFD74073FC3772FC0F98F86D" box="[1018,1171,1929,1955]" italics="true" pageId="5" pageNumber="6">L. legionaria</emphasis>
, Lm
</taxonomicName>
: 
<taxonomicName id="57F34D26FFD74073FB1672FC0BF9F809" authority=", Lno" authorityName="Lno" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="mckennae">
<emphasis id="A287EAB7FFD74073FB1672FC0E64F86D" box="[1243,1391,1929,1955]" italics="true" pageId="5" pageNumber="6">L. mckennae</emphasis>
, Lno
</taxonomicName>
: 
<taxonomicName id="57F34D26FFD74073FF3372DB0AC8F809" authority=", Lnu" authorityName="Lnu" box="[254,451,1965,1991]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nomada">
<emphasis id="A287EAB7FFD74073FF3372DB0A74F809" box="[254,383,1965,1991]" italics="true" pageId="5" pageNumber="6">L. nomada</emphasis>
, Lnu
</taxonomicName>
: 
<taxonomicName id="57F34D26FFD74073FE1D72DB0956F809" box="[464,605,1965,1991]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="nubecula">
<emphasis id="A287EAB7FFD74073FE1D72DB0956F809" box="[464,605,1965,1991]" italics="true" pageId="5" pageNumber="6">L. nubecula</emphasis>
</taxonomicName>
); stars: 
<taxonomicName id="57F34D26FFD74073FD7272DB0892F809" box="[703,921,1965,1991]" class="Insecta" family="Formicidae" genus="Asphinctanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">
<emphasis id="A287EAB7FFD74073FD7272DB0892F809" box="[703,921,1965,1991]" italics="true" pageId="5" pageNumber="6">Asphinctanilloides</emphasis>
</taxonomicName>
species (Aa: 
<taxonomicName id="57F34D26FFD74073FBF272DB0BF0F824" authority="Brandao et al., 1999" authorityName="Brandao" authorityYear="1999" class="Insecta" family="Formicidae" genus="Asphinctanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="amazon">
<emphasis id="A287EAB7FFD74073FBF272DB0FB7F808" box="[1087,1212,1965,1991]" italics="true" pageId="5" pageNumber="6">A. amazon</emphasis>
<bibRefCitation id="F4624B54FFD74073FB0B72DB0BF0F824" author="Brandao C. R. F. &amp; Diniz J. L. M. &amp; Agosti D. &amp; Delabie J. H. C." pageId="5" pageNumber="6" pagination="17 - 36" refId="ref14350" refString="Brandao C. R. F., Diniz J. L. M., Agosti D. &amp; Delabie J. H. C. 1999. Revision of the Neotropical ant subfamily Leptanilloidinae. Systematic Entomology 24: 17 - 36. http: // dx. doi. org / 10.1046 / j. 1365 - 3113.1999.00064. x" type="journal article" year="1999">
Brandão 
<emphasis id="A287EAB7FFD74073FAFE72DB0E7BF809" box="[1331,1392,1965,1991]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
, 1999
</bibRefCitation>
</taxonomicName>
, Aan: 
<taxonomicName id="57F34D26FFD74073FE8B72A60993F824" authority="Brandao et al., 1999" authorityName="Brandao" authorityYear="1999" box="[326,664,2000,2026]" class="Insecta" family="Formicidae" genus="Asphinctanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="anae">
<emphasis id="A287EAB7FFD74073FE8B72A60A94F824" box="[326,415,2000,2026]" italics="true" pageId="5" pageNumber="6">A. anae</emphasis>
<bibRefCitation id="F4624B54FFD74073FE6A72A60993F824" author="Brandao C. R. F. &amp; Diniz J. L. M. &amp; Agosti D. &amp; Delabie J. H. C." box="[423,664,2000,2026]" pageId="5" pageNumber="6" pagination="17 - 36" refId="ref14350" refString="Brandao C. R. F., Diniz J. L. M., Agosti D. &amp; Delabie J. H. C. 1999. Revision of the Neotropical ant subfamily Leptanilloidinae. Systematic Entomology 24: 17 - 36. http: // dx. doi. org / 10.1046 / j. 1365 - 3113.1999.00064. x" type="journal article" year="1999">
Brandão 
<emphasis id="A287EAB7FFD74073FDDC72A70947F824" box="[529,588,2000,2026]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
, 1999
</bibRefCitation>
</taxonomicName>
, Am: 
<taxonomicName id="57F34D26FFD74073FD1372A60F7AF824" authority="Brandao et al., 1999" authorityName="Brandao" authorityYear="1999" box="[734,1137,2000,2026]" class="Insecta" family="Formicidae" genus="Asphinctanilloides" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="manauara">
<emphasis id="A287EAB7FFD74073FD1372A6087CF824" box="[734,887,2000,2026]" italics="true" pageId="5" pageNumber="6">A. manauara</emphasis>
<bibRefCitation id="F4624B54FFD74073FCB372A60F7AF824" author="Brandao C. R. F. &amp; Diniz J. L. M. &amp; Agosti D. &amp; Delabie J. H. C." box="[894,1137,2000,2026]" pageId="5" pageNumber="6" pagination="17 - 36" refId="ref14350" refString="Brandao C. R. F., Diniz J. L. M., Agosti D. &amp; Delabie J. H. C. 1999. Revision of the Neotropical ant subfamily Leptanilloidinae. Systematic Entomology 24: 17 - 36. http: // dx. doi. org / 10.1046 / j. 1365 - 3113.1999.00064. x" type="journal article" year="1999">
Brandão 
<emphasis id="A287EAB7FFD74073FC2472A70F2FF824" box="[1001,1060,2000,2026]" italics="true" pageId="5" pageNumber="6">et al.</emphasis>
, 1999
</bibRefCitation>
</taxonomicName>
).
</paragraph>
</caption>
<paragraph id="904C36A5FFD44070FF70747A0922FEEB" blockId="6.[189,566,267,329]" box="[189,553,267,294]" pageId="6" pageNumber="7">
<taxonomicName id="57F34D26FFD44070FF70747A0922FEEB" authority="Donoso et al., 2006" authorityName="Donoso" authorityYear="2006" box="[189,553,267,294]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nomada">
<emphasis id="A287EAB7FFD44070FF70747A0A30FEEB" box="[189,315,267,293]" italics="true" pageId="6" pageNumber="7">L. nomada</emphasis>
<bibRefCitation id="F4624B54FFD44070FE8F747A0922FEEB" author="Donoso D. A. &amp; Vieira J. M. &amp; Wild A. L." box="[322,553,267,294]" pageId="6" pageNumber="7" pagination="47 - 62" refId="ref14648" refString="Donoso D. A., Vieira J. M. &amp; Wild A. L. 2006. Three new species of Leptanilloides Mann from Andean Ecuador (Formicidade: Leptanilloidinae). Zootaxa 1201: 47 - 62." type="journal article" year="2006">
Donoso 
<emphasis id="A287EAB7FFD44070FE69747A0AD4FEEB" box="[420,479,267,293]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
, 2006
</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="904C36A5FFD44070FF707459093DFE86" blockId="6.[189,566,267,329]" box="[189,566,302,329]" pageId="6" pageNumber="7">
<taxonomicName id="57F34D26FFD44070FF707459093DFE86" authority="Donoso et al., 2006" authorityName="Donoso" authorityYear="2006" box="[189,566,302,329]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nubecula">
<emphasis id="A287EAB7FFD44070FF7074590A43FE87" box="[189,328,303,329]" italics="true" pageId="6" pageNumber="7">L. nubecula</emphasis>
<bibRefCitation id="F4624B54FFD44070FE827459093DFE86" author="Donoso D. A. &amp; Vieira J. M. &amp; Wild A. L." box="[335,566,302,329]" pageId="6" pageNumber="7" pagination="47 - 62" refId="ref14648" refString="Donoso D. A., Vieira J. M. &amp; Wild A. L. 2006. Three new species of Leptanilloides Mann from Andean Ecuador (Formicidade: Leptanilloidinae). Zootaxa 1201: 47 - 62." type="journal article" year="2006">
Donoso 
<emphasis id="A287EAB7FFD44070FE7C74590AE7FE87" box="[433,492,303,329]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
, 2006
</bibRefCitation>
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="D8E9652EFFD44070FF7074060944FE45" box="[189,591,368,395]" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="904C36A5FFD44070FF7074060944FE45" blockId="6.[189,591,368,395]" box="[189,591,368,395]" pageId="6" pageNumber="7">
<heading id="CB0481C9FFD44070FF7074060944FE45" bold="true" box="[189,591,368,395]" fontSize="11" level="3" pageId="6" pageNumber="7" reason="3">
<emphasis id="A287EAB7FFD44070FF7074060944FE45" bold="true" box="[189,591,368,395]" pageId="6" pageNumber="7">
Key to workers of 
<taxonomicName id="57F34D26FFD44070FE6974060944FE45" ID-CoL="5CRS" authorityName="Mann" authorityYear="1823" box="[420,591,368,395]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="A287EAB7FFD44070FE6974060944FE45" bold="true" box="[420,591,368,395]" italics="true" pageId="6" pageNumber="7">Leptanilloides</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D8E9652EFFD44071FF7074E90E7CF9CE" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="key">
<key id="E4626194FFD44071FF7074E90E7CF9CE" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<keyStep id="2B072E00FFD44070FF7074E90E7CFDA7" pageId="6" pageNumber="7">
<paragraph id="904C36A5FFD44070FF7074E90E7CFDCE" blockId="6.[189,1399,414,617]" pageId="6" pageNumber="7">
<keyLead id="2B029590FFD44070FF7074E90E7CFDCE" pageId="6" pageNumber="7">
1. Abdominal segment III (postpetiole) in lateral view much smaller than adjoining fourth abdominal segment. Spiracle of segment III shifted posteriad on anteromedian side of tergite. Body size relatively large, HL 0.68–0.75 (
<figureCitation id="08C82A20FFD44070FD3E74930831FE31" box="[755,826,485,511]" captionStart="Fig" captionStartId="5.[190,233,1718,1744]" captionTargetBox="[209,1357,966,1683]" captionTargetId="figure@5.[173,1383,946,1714]" captionTargetPageId="5" captionText="Fig. 1. Relationship between HL and WL among workers in species of Leptanilloides and Asphinctanilloides. Specimens measured are holotypes or paratypes. Measurements are in mm. WL is called Mesosoma Length (ML) in Borowiec &amp; Longino (2011) and Silva et al. (2013). Triangles: the two Leptanilloides described in this paper (L. copalinga Delsinne &amp; Donoso sp. nov. and L. prometea Delsinne &amp; Donoso sp. nov.); diamonds: Leptanilloides biconstricta species-group (La: L. atlantica, Lb: L. biconstricta, Lc: L. caracola, Le: L. erinys, Lf: L. femoralis, Lg: L. gracilis, Li: L. improvisa, Ls: L. sculpturata); dots: Leptanilloides legionaria species-group (Ll: L. legionaria, Lm: L. mckennae, Lno: L. nomada, Lnu: L. nubecula); stars: Asphinctanilloides species (Aa: A. amazon Brandão et al., 1999, Aan: A. anae Brandão et al., 1999, Am: A. manauara Brandão et al., 1999)." figureDoi="http://doi.org/10.5281/zenodo.3787362" httpUri="https://zenodo.org/record/3787362/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
) ………………2 (
<taxonomicName id="57F34D26FFD44070FBEE74900FB4FE31" ID-CoL="3THNK" box="[1059,1215,485,511]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="legionaria">
<emphasis id="A287EAB7FFD44070FBEE74900FB4FE31" box="[1059,1215,485,511]" italics="true" pageId="6" pageNumber="7">L. legionaria</emphasis>
</taxonomicName>
species-group)
</keyLead>
</paragraph>
<paragraph id="904C36A5FFD44070FF70777F0E7CFDA7" blockId="6.[189,1399,414,617]" pageId="6" pageNumber="7">
<keyLead id="2B029590FFD44070FF70777F0E7CFDA7" pageId="6" pageNumber="7">
– Abdominal segment III (postpetiole) in lateral view nearly as high as abdominal segment IV. Spiracle of segment III situated forward on the tergite. Body size relatively small, HL 0.31–0.62 (
<figureCitation id="08C82A20FFD44070FF3677380A34FDA7" box="[251,319,590,617]" captionStart="Fig" captionStartId="5.[190,233,1718,1744]" captionTargetBox="[209,1357,966,1683]" captionTargetId="figure@5.[173,1383,946,1714]" captionTargetPageId="5" captionText="Fig. 1. Relationship between HL and WL among workers in species of Leptanilloides and Asphinctanilloides. Specimens measured are holotypes or paratypes. Measurements are in mm. WL is called Mesosoma Length (ML) in Borowiec &amp; Longino (2011) and Silva et al. (2013). Triangles: the two Leptanilloides described in this paper (L. copalinga Delsinne &amp; Donoso sp. nov. and L. prometea Delsinne &amp; Donoso sp. nov.); diamonds: Leptanilloides biconstricta species-group (La: L. atlantica, Lb: L. biconstricta, Lc: L. caracola, Le: L. erinys, Lf: L. femoralis, Lg: L. gracilis, Li: L. improvisa, Ls: L. sculpturata); dots: Leptanilloides legionaria species-group (Ll: L. legionaria, Lm: L. mckennae, Lno: L. nomada, Lnu: L. nubecula); stars: Asphinctanilloides species (Aa: A. amazon Brandão et al., 1999, Aan: A. anae Brandão et al., 1999, Am: A. manauara Brandão et al., 1999)." figureDoi="http://doi.org/10.5281/zenodo.3787362" httpUri="https://zenodo.org/record/3787362/files/figure.png" pageId="6" pageNumber="7">Fig. 1</figureCitation>
) ……………………………………………………………5 (
<taxonomicName id="57F34D26FFD44070FBDB77390FC9FDA7" ID-CoL="3THNH" box="[1046,1218,591,617]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="biconstricta">
<emphasis id="A287EAB7FFD44070FBDB77390FC9FDA7" box="[1046,1218,591,617]" italics="true" pageId="6" pageNumber="7">L. biconstricta</emphasis>
</taxonomicName>
species-group)
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B072E00FFD44070FF7077E60E7CFCF9" pageId="6" pageNumber="7">
<paragraph id="904C36A5FFD44070FF7077E60E7CFD3E" blockId="6.[189,1399,656,823]" pageId="6" pageNumber="7">
<keyLead id="2B029590FFD44070FF7077E60E7CFD3E" pageId="6" pageNumber="7">
2. Head subquadrate, CI 85–88; lateral margin nearly straight and parallel. Propodeal declivity short and vertical, propodeum with dorsal and posterior faces clearly differentiated (
<collectingCountry id="E8E47635FFD44070FB6A77C50E07FD00" box="[1191,1292,691,718]" name="Ecuador" pageId="6" pageNumber="7">Ecuador</collectingCountry>
) ……… ……………………………………………………………………… 
<taxonomicName id="57F34D26FFD44070FBC777A10E7CFD3E" authority="Donoso et al., 2006" authorityName="Donoso" authorityYear="2006" box="[1034,1399,726,753]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nomada">
<emphasis id="A287EAB7FFD44070FBC777A10F87FD3F" bold="true" box="[1034,1164,726,753]" italics="true" pageId="6" pageNumber="7">L. nomada</emphasis>
<bibRefCitation id="F4624B54FFD44070FB5F77A10E7CFD3E" author="Donoso D. A. &amp; Vieira J. M. &amp; Wild A. L." box="[1170,1399,726,753]" pageId="6" pageNumber="7" pagination="47 - 62" refId="ref14648" refString="Donoso D. A., Vieira J. M. &amp; Wild A. L. 2006. Three new species of Leptanilloides Mann from Andean Ecuador (Formicidade: Leptanilloidinae). Zootaxa 1201: 47 - 62." type="journal article" year="2006">
Donoso 
<emphasis id="A287EAB7FFD44070FB3977A10E25FD3E" box="[1268,1326,726,752]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
, 2006
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="904C36A5FFD44070FF70778C0E7CFCF9" blockId="6.[189,1399,656,823]" pageId="6" pageNumber="7">
<keyLead id="2B029590FFD44070FF70778C0E7CFCF9" pageId="6" pageNumber="7">– Head subrectangular, CI 75–83; lateral margin convex. Propodeal declivity usually rounded without clear distinction between dorsal and posterior faces ……………………………………3</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B072E00FFD44070FF7076280E7DFBCB" pageId="6" pageNumber="7">
<paragraph id="904C36A5FFD44070FF7076280E7CFC71" blockId="6.[189,1399,862,1030]" pageId="6" pageNumber="7">
<keyLead id="2B029590FFD44070FF7076280E7CFC71" pageId="6" pageNumber="7">
3. Head sculpture less dense, at most 10–12 shallow foveolae across face at midlength. Lateral margin of head distinctly convex. Lateroclypeal tooth present. Posterior margin of head slightly concave (
<collectingCountry id="E8E47635FFD44070FF3776D30A7AFC0E" box="[250,369,933,960]" name="Colombia" pageId="6" pageNumber="7">Colombia</collectingCountry>
) …………………………………………………… 
<taxonomicName id="57F34D26FFD44070FC0276D30E7CFC71" ID-CoL="3THNK" authority="Brandao et al., 1999" authorityName="Brandao" authorityYear="1999" box="[975,1399,932,959]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="legionaria">
<emphasis id="A287EAB7FFD44070FC0276D30F7BFC71" bold="true" box="[975,1136,932,959]" italics="true" pageId="6" pageNumber="7">L. legionaria</emphasis>
<bibRefCitation id="F4624B54FFD44070FBB676D30E7CFC71" author="Brandao C. R. F. &amp; Diniz J. L. M. &amp; Agosti D. &amp; Delabie J. H. C." box="[1147,1399,933,959]" pageId="6" pageNumber="7" pagination="17 - 36" refId="ref14350" refString="Brandao C. R. F., Diniz J. L. M., Agosti D. &amp; Delabie J. H. C. 1999. Revision of the Neotropical ant subfamily Leptanilloidinae. Systematic Entomology 24: 17 - 36. http: // dx. doi. org / 10.1046 / j. 1365 - 3113.1999.00064. x" type="journal article" year="1999">
Brandão 
<emphasis id="A287EAB7FFD44070FB2776D30E22FC71" box="[1258,1321,933,959]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
, 1999
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="904C36A5FFD44070FF7076BE0E7DFBCB" blockId="6.[189,1399,862,1030]" pageId="6" pageNumber="7">
<keyLead id="2B029590FFD44070FF7076BE0E7DFBCB" pageId="6" pageNumber="7">– Head sculpture denser, with at least 15 foveolae across face at midlength. Lateral margin of head slightly convex. Lateroclypeal tooth absent. Posterior margin of head deeply concave …………4</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B072E00FFD44070FF70715B0E7CFB7E" pageId="6" pageNumber="7">
<paragraph id="904C36A5FFD44070FF70715B0E7CFBA7" blockId="6.[189,1399,1068,1201]" pageId="6" pageNumber="7">
<keyLead id="2B029590FFD44070FF70715B0E7CFBA7" pageId="6" pageNumber="7">
4. Legs shorter, HW/HTL×100&gt; 78. Hypostomal tooth present (
<collectingCountry id="E8E47635FFD44070FC17715A0F34FB89" box="[986,1087,1068,1095]" name="Ecuador" pageId="6" pageNumber="7">Ecuador</collectingCountry>
) ………………………… ………………………………………………………………… 
<taxonomicName id="57F34D26FFD44070FC0271260E7CFBA7" authority="Donoso et al., 2006" authorityName="Donoso" authorityYear="2006" box="[975,1399,1103,1130]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="nubecula">
<emphasis id="A287EAB7FFD44070FC0271260F63FBA4" bold="true" box="[975,1128,1103,1130]" italics="true" pageId="6" pageNumber="7">L. nubecula</emphasis>
<bibRefCitation id="F4624B54FFD44070FBB571260E7CFBA7" author="Donoso D. A. &amp; Vieira J. M. &amp; Wild A. L." box="[1144,1399,1103,1130]" pageId="6" pageNumber="7" pagination="47 - 62" refId="ref14648" refString="Donoso D. A., Vieira J. M. &amp; Wild A. L. 2006. Three new species of Leptanilloides Mann from Andean Ecuador (Formicidade: Leptanilloidinae). Zootaxa 1201: 47 - 62." type="journal article" year="2006">
Donoso 
<emphasis id="A287EAB7FFD44070FB2F71260E15FBA4" box="[1250,1310,1104,1130]" italics="true" pageId="6" pageNumber="7">et al</emphasis>
., 2006
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="904C36A5FFD44070FF7071050E7CFB7E" blockId="6.[189,1399,1068,1201]" pageId="6" pageNumber="7">
<keyLead id="2B029590FFD44070FF7071050E7CFB7E" pageId="6" pageNumber="7">
– Legs longer, HW/HTL×100 &lt;78. Hypostomal tooth absent (
<collectingCountry id="E8E47635FFD44070FC1871050F57FB40" box="[981,1116,1139,1166]" name="Costa Rica" pageId="6" pageNumber="7">Costa Rica</collectingCountry>
) ……………………… ………………………………………………………………………… 
<taxonomicName id="57F34D26FFD44070FBEA71E00E7CFB7E" authority="Longino, 2003" authorityName="Longino" authorityYear="2003" box="[1063,1399,1174,1201]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="mckennae">
<emphasis id="A287EAB7FFD44070FBEA71E00FC8FB7F" bold="true" box="[1063,1219,1174,1201]" italics="true" pageId="6" pageNumber="7">L. mckennae</emphasis>
<bibRefCitation id="F4624B54FFD44070FB0771E00E7CFB7E" author="Longino J. T." box="[1226,1399,1174,1200]" pageId="6" pageNumber="7" pagination="1 - 6" refId="ref15223" refString="Longino J. T. 2003. A new Costa Rican species of Leptanilloides (Hymenoptera: Formicidae: doryline section: Leptanilloidinae). Zootaxa 264: 1 - 6." type="journal article" year="2003">Longino, 2003</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B072E00FFD44070FF7071A10E7CFA95" pageId="6" pageNumber="7">
<paragraph id="904C36A5FFD44070FF7071A10E7CFADB" blockId="6.[189,1399,1239,1372]" pageId="6" pageNumber="7">
<keyLead id="2B029590FFD44070FF7071A10E7CFADB" pageId="6" pageNumber="7">
5. Lateroclypeal tooth absent. Masticatory margin of mandible edentate (
<collectingCountry id="E8E47635FFD44070FBFE71A10F93FB3C" box="[1075,1176,1239,1266]" name="Ecuador" pageId="6" pageNumber="7">Ecuador</collectingCountry>
) ………………… …………………………………………………………………… 
<taxonomicName id="57F34D26FFD44070FC20718D0E7CFADB" authority="Donoso et al., 2006" authorityName="Donoso" authorityYear="2006" box="[1005,1399,1274,1301]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="caracola">
<emphasis id="A287EAB7FFD44070FC20718D0F72FADB" bold="true" box="[1005,1145,1274,1301]" italics="true" pageId="6" pageNumber="7">L. caracola</emphasis>
<bibRefCitation id="F4624B54FFD44070FB49718D0E7CFADB" author="Donoso D. A. &amp; Vieira J. M. &amp; Wild A. L." box="[1156,1399,1275,1301]" pageId="6" pageNumber="7" pagination="47 - 62" refId="ref14648" refString="Donoso D. A., Vieira J. M. &amp; Wild A. L. 2006. Three new species of Leptanilloides Mann from Andean Ecuador (Formicidade: Leptanilloidinae). Zootaxa 1201: 47 - 62." type="journal article" year="2006">
Donoso 
<emphasis id="A287EAB7FFD44070FB26718A0E22FADB" box="[1259,1321,1275,1301]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
, 2006
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="904C36A5FFD44070FF7070680E7CFA95" blockId="6.[189,1399,1239,1372]" pageId="6" pageNumber="7">
<keyLead id="2B029590FFD44070FF7070680E7CFA95" pageId="6" pageNumber="7">– Lateroclypeal tooth present. Masticatory margin of mandible dentate (at least minute denticles present) ……………………………………………………………………………………………6</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B072E00FFD44070FF7070F40E7CF983" pageId="6" pageNumber="7">
<paragraph id="904C36A5FFD44070FF7070F40E7CFA2D" blockId="6.[189,1399,1410,1613]" pageId="6" pageNumber="7">
<keyLead id="2B029590FFD44070FF7070F40E7CFA2D" pageId="6" pageNumber="7">6. In lateral view, abdominal segment IV narrowly attached to preceding segment III and broadly to succeeding segment V, with contrast between widths of anterior and posterior articulations of segment IV in lateral view …………………………………………………………………7</keyLead>
</paragraph>
<paragraph id="904C36A5FFD44070FF70709B0E7CF983" blockId="6.[189,1399,1410,1613]" pageId="6" pageNumber="7">
<keyLead id="2B029590FFD44070FF70709B0E7CF983" pageId="6" pageNumber="7">– In lateral view, abdominal segment IV relatively broadly attached to preceding segment III, with little contrast between widths of anterior and posterior articulations of segment IV in lateral view ………………………………………………………………………………………8</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B072E00FFD44070FF7073020E7CF805" pageId="6" pageNumber="7">
<paragraph id="904C36A5FFD44070FF7073020E7CF936" blockId="6.[189,1399,1652,1996]" pageId="6" pageNumber="7">
<keyLead id="2B029590FFD44070FF7073020E7CF936" pageId="6" pageNumber="7">
7. In lateral view, sternite of abdominal segment III evenly rounded, making sternal and tergal portions subequal. Petiole as long as postpetiole. Reticulation on mesopleuron and metapleuron uninterrupted. Head densely foveolate, with 
<emphasis id="A287EAB7FFD44070FCC873CD0812F91A" box="[773,793,1723,1748]" italics="true" pageId="6" pageNumber="7">c.</emphasis>
20–30 foveolae covering a straight transverse line at head midlength. HL ≤ 0.36 (
<collectingCountry id="E8E47635FFD44070FD9373AB09A2F936" box="[606,681,1757,1784]" name="Brazil" pageId="6" pageNumber="7">Brazil</collectingCountry>
) ……………………………… 
<taxonomicName id="57F34D26FFD44070FBD773A80E7CF936" authority="Silva et al., 2013" authorityName="Silva" authorityYear="2013" box="[1050,1399,1757,1784]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="atlantica">
<emphasis id="A287EAB7FFD44070FBD773A80FAEF936" bold="true" box="[1050,1189,1757,1784]" italics="true" pageId="6" pageNumber="7">L. atlantica</emphasis>
<bibRefCitation id="F4624B54FFD44070FB6073AB0E7CF936" author="Silva R. R. &amp; Feitosa R. M. &amp; Brandao C. R. F. &amp; Freitas A. V. L." box="[1197,1399,1757,1784]" pageId="6" pageNumber="7" pagination="2039 - 2047" refId="ref15882" refString="Silva R. R., Feitosa R. M., Brandao C. R. F. &amp; Freitas A. V. L. 2013. The first Leptanilloides species (Hymenoptera: Formicidae: Leptanilloidinae) from eastern South America. Journal of Natural History 47 (31 - 32): 2039 - 2047. http: // dx. doi. org / 10.1080 / 00222933.2012.763058" type="journal article" year="2013">
Silva 
<emphasis id="A287EAB7FFD44070FB3C73A80E26F936" box="[1265,1325,1758,1784]" italics="true" pageId="6" pageNumber="7">et al.</emphasis>
, 2013
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="904C36A5FFD44070FF7072770E7CF805" blockId="6.[189,1399,1652,1996]" pageId="6" pageNumber="7">
<keyLead id="2B029590FFD44070FF7072770E7CF805" pageId="6" pageNumber="7">
– In lateral view, sternite of abdominal segment III (postpetiole) distinctly bulging anteriorly, making sternal portion of segment deeper than tergite (
<bibRefCitation id="F4624B54FFD44070FCB972520FBCF8F0" author="Borowiec M. L. &amp; Longino J. T." box="[884,1207,1828,1854]" pageId="6" pageNumber="7" pagination="19 - 48" refId="ref14112" refString="Borowiec M. L. &amp; Longino J. T. 2011. Three new species and reassessment of the rare Neotropical ant genus Leptanilloides (Hymenoptera, Formicidae, Leptanilloidinae). ZooKeys 133: 19 - 48. http: // dx. doi. org / 10.3897 / zookeys. 133.1479" type="journal article" year="2011">Borowiec &amp; Longino 2011</bibRefCitation>
: fig. 2d). Petiole longer than postpetiole (
<bibRefCitation id="F4624B54FFD44070FDD672310861F8AF" author="Borowiec M. L. &amp; Longino J. T." box="[539,874,1863,1889]" pageId="6" pageNumber="7" pagination="19 - 48" refId="ref14112" refString="Borowiec M. L. &amp; Longino J. T. 2011. Three new species and reassessment of the rare Neotropical ant genus Leptanilloides (Hymenoptera, Formicidae, Leptanilloidinae). ZooKeys 133: 19 - 48. http: // dx. doi. org / 10.3897 / zookeys. 133.1479" type="journal article" year="2011">Borowiec &amp; Longino 2011</bibRefCitation>
: fig. 2d). Reticulation on mesopleuron and metapleuron superficial and interrupted. Head sculpture less dense, with only 10–20 foveolae covering a straight transverse line at head midlength. HL = 
<quantity id="570B9B40FFD44070FC2872F80F37F869" box="[997,1084,1933,1960]" metricMagnitude="-2" metricUnit="m" metricValue="1.2446" pageId="6" pageNumber="7" unit="in" value="0.49">0.49 in</quantity>
only measured specimen (
<collectingCountry id="E8E47635FFD44070FF3772C70A5EF805" box="[250,341,1969,1995]" name="Bolivia" pageId="6" pageNumber="7">Bolivia</collectingCountry>
, 
<collectingCountry id="E8E47635FFD44070FE9372C70ADEF805" box="[350,469,1969,1995]" name="Colombia" pageId="6" pageNumber="7">Colombia</collectingCountry>
, 
<collectingCountry id="E8E47635FFD44070FE1372C70952F802" box="[478,601,1969,1996]" name="Venezuela" pageId="6" pageNumber="7">Venezuela</collectingCountry>
) ………………………………………… 
<taxonomicName id="57F34D26FFD44070FBF772C70E7CF805" ID-CoL="3THNH" authority="Mann, 1823" authorityName="Mann" authorityYear="1823" box="[1082,1399,1968,1995]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="biconstricta">
<emphasis id="A287EAB7FFD44070FBF772C70FEDF805" bold="true" box="[1082,1254,1968,1995]" italics="true" pageId="6" pageNumber="7">L. biconstricta</emphasis>
Mann, 1823
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B072E00FFD54071FF70747D0E7DFE87" pageId="7" pageNumber="8">
<paragraph id="904C36A5FFD54071FF70747D0E7CFEEB" blockId="7.[189,1399,267,329]" box="[189,1399,267,294]" pageId="7" pageNumber="8">
<keyLead id="2B029590FFD54071FF70747D0E7CFEEB" box="[189,1399,267,294]" pageId="7" pageNumber="8">8. In lateral view, petiolar sternite distinctly bulging medially ……………………………………9</keyLead>
</paragraph>
<paragraph id="904C36A5FFD54071FF7074590E7DFE87" blockId="7.[189,1399,267,329]" box="[189,1398,302,329]" pageId="7" pageNumber="8">
<keyLead id="2B029590FFD54071FF7074590E7DFE87" box="[189,1398,302,329]" pageId="7" pageNumber="8">– In lateral view, petiolar sternite straight (not bulging) or bulging anteriorly …………………10</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B072E00FFD54071FF7074060E7CFD4E" pageId="7" pageNumber="8">
<paragraph id="904C36A5FFD54071FF7074060E7CFE3A" blockId="7.[189,1399,368,641]" pageId="7" pageNumber="8">
<keyLead id="2B029590FFD54071FF7074060E7CFE3A" pageId="7" pageNumber="8">
9. Hindtibia with two very small, simple spurs, without pectinate spur clearly visible under 50× magnification. Petiolar spiracle opening in an excavation distinctly larger than propodeal spiracle (
<bibRefCitation id="F4624B54FFD54071FF3774C0094EFE1E" author="Borowiec M. L. &amp; Longino J. T." box="[250,581,438,464]" pageId="7" pageNumber="8" pagination="19 - 48" refId="ref14112" refString="Borowiec M. L. &amp; Longino J. T. 2011. Three new species and reassessment of the rare Neotropical ant genus Leptanilloides (Hymenoptera, Formicidae, Leptanilloidinae). ZooKeys 133: 19 - 48. http: // dx. doi. org / 10.3897 / zookeys. 133.1479" type="journal article" year="2011">Borowiec &amp; Longino 2011</bibRefCitation>
: fig. 5g–h). Flange over metapleural gland opening sharply pointed posteriorly (
<collectingCountry id="E8E47635FFD54071FE4E74AF0AEAFE3D" box="[387,481,473,499]" name="Mexico" pageId="7" pageNumber="8">Mexico</collectingCountry>
, 
<collectingCountry id="E8E47635FFD54071FE2274AF097BFE3A" box="[495,624,473,500]" name="Guatemala" pageId="7" pageNumber="8">Guatemala</collectingCountry>
) ………………………… 
<taxonomicName id="57F34D26FFD54071FC6A74AC0E7CFE3A" authority="Borowiec &amp; Longino, 2011" authorityName="Borowiec &amp; Longino" authorityYear="2011" box="[935,1399,473,500]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="A287EAB7FFD54071FC6A74AC0F28FE3A" bold="true" box="[935,1059,473,500]" italics="true" pageId="7" pageNumber="8">L. gracilis</emphasis>
<bibRefCitation id="F4624B54FFD54071FBE074AF0E7CFE3A" author="Borowiec M. L. &amp; Longino J. T." box="[1069,1399,473,500]" pageId="7" pageNumber="8" pagination="19 - 48" refId="ref14112" refString="Borowiec M. L. &amp; Longino J. T. 2011. Three new species and reassessment of the rare Neotropical ant genus Leptanilloides (Hymenoptera, Formicidae, Leptanilloidinae). ZooKeys 133: 19 - 48. http: // dx. doi. org / 10.3897 / zookeys. 133.1479" type="journal article" year="2011">Borowiec &amp; Longino, 2011</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="904C36A5FFD54071FF70748B0E7CFD4E" blockId="7.[189,1399,368,641]" pageId="7" pageNumber="8">
<keyLead id="2B029590FFD54071FF70748B0E7CFD4E" pageId="7" pageNumber="8">
– Hindtibia with large pectinate spur, clearly discernable under 50× magnification. Petiolar spiracle not in excavation, similar and subequal to or smaller in diameter than propodeal spiracle (
<bibRefCitation id="F4624B54FFD54071FF3777350932FD93" author="Borowiec M. L. &amp; Longino J. T." box="[250,569,579,605]" pageId="7" pageNumber="8" pagination="19 - 48" refId="ref14112" refString="Borowiec M. L. &amp; Longino J. T. 2011. Three new species and reassessment of the rare Neotropical ant genus Leptanilloides (Hymenoptera, Formicidae, Leptanilloidinae). ZooKeys 133: 19 - 48. http: // dx. doi. org / 10.3897 / zookeys. 133.1479" type="journal article" year="2011">Borowiec &amp; Longino 2011</bibRefCitation>
: fig. 4g–h). Flange over metapleural gland opening rounded posteriorly (
<collectingCountry id="E8E47635FFD54071FF3777100A7CFD4F" box="[250,375,614,641]" name="Venezuela" pageId="7" pageNumber="8">Venezuela</collectingCountry>
) ……………………………………………… 
<taxonomicName id="57F34D26FFD54071FC5B77110E7CFD4E" authority="Borowiec &amp; Longino, 2011" authorityName="Borowiec &amp; Longino" authorityYear="2011" box="[918,1399,614,641]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="A287EAB7FFD54071FC5B77110F22FD4F" bold="true" box="[918,1065,614,641]" italics="true" pageId="7" pageNumber="8">L. femoralis</emphasis>
<bibRefCitation id="F4624B54FFD54071FBFC77100E7CFD4E" author="Borowiec M. L. &amp; Longino J. T." box="[1073,1399,614,640]" pageId="7" pageNumber="8" pagination="19 - 48" refId="ref14112" refString="Borowiec M. L. &amp; Longino J. T. 2011. Three new species and reassessment of the rare Neotropical ant genus Leptanilloides (Hymenoptera, Formicidae, Leptanilloidinae). ZooKeys 133: 19 - 48. http: // dx. doi. org / 10.3897 / zookeys. 133.1479" type="journal article" year="2011">Borowiec &amp; Longino, 2011</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B072E00FFD54071FF7077DE0E7DFC81" pageId="7" pageNumber="8">
<paragraph id="904C36A5FFD54071FF7077DE0E7CFCC6" blockId="7.[189,1399,680,847]" pageId="7" pageNumber="8">
<keyLead id="2B029590FFD54071FF7077DE0E7CFCC6" pageId="7" pageNumber="8">
10. Head narrow, HL &lt;0.30, HW &lt;0.20, CI &lt;60. Head dorsum densely foveolate with foveolae separated by less than their diameter, often contiguous (
<collectingCountry id="E8E47635FFD54071FC4377BD0F0EFD28" box="[910,1029,715,742]" name="Colombia" pageId="7" pageNumber="8">Colombia</collectingCountry>
) ……………………………… ………………………………………………………………… 
<taxonomicName id="57F34D26FFD54071FC0277990E7CFCC6" ID-CoL="3THNL" authority="Brandao et al., 1999" authorityName="Brandao" authorityYear="1999" box="[975,1399,750,777]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="sculpturata">
<emphasis id="A287EAB7FFD54071FC0277990F71FCC7" bold="true" box="[975,1146,750,777]" italics="true" pageId="7" pageNumber="8">L. sculpturata</emphasis>
<bibRefCitation id="F4624B54FFD54071FB4E77980E7CFCC6" author="Brandao C. R. F. &amp; Diniz J. L. M. &amp; Agosti D. &amp; Delabie J. H. C." box="[1155,1399,750,777]" pageId="7" pageNumber="8" pagination="17 - 36" refId="ref14350" refString="Brandao C. R. F., Diniz J. L. M., Agosti D. &amp; Delabie J. H. C. 1999. Revision of the Neotropical ant subfamily Leptanilloidinae. Systematic Entomology 24: 17 - 36. http: // dx. doi. org / 10.1046 / j. 1365 - 3113.1999.00064. x" type="journal article" year="1999">
Brandão 
<emphasis id="A287EAB7FFD54071FB2277990E27FCC6" box="[1263,1324,750,776]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
, 1999
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="904C36A5FFD54071FF7076640E7DFC81" blockId="7.[189,1399,680,847]" pageId="7" pageNumber="8">
<keyLead id="2B029590FFD54071FF7076640E7DFC81" pageId="7" pageNumber="8">– Head broader, HL&gt; 0.30, HW&gt; 0.20, CI&gt; 65. Head dorsum less densely foveolate, with foveolae separated by about their diameter or more ……………………………………………11</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B072E00FFD54071FF7076000E7DFC34" pageId="7" pageNumber="8">
<paragraph id="904C36A5FFD54071FF7076000E7DFC7D" blockId="7.[189,1399,886,1018]" pageId="7" pageNumber="8">
<keyLead id="2B029590FFD54071FF7076000E7DFC7D" pageId="7" pageNumber="8">11. Size large, HL ≥ 0.50, HW ≥ 0.38. Flange over metapleural gland opening short, not surpassing propodeum declivity margin in lateral view, and rounded posteriorly ………………………12</keyLead>
</paragraph>
<paragraph id="904C36A5FFD54071FF7076CB0E7DFC34" blockId="7.[189,1399,886,1018]" pageId="7" pageNumber="8">
<keyLead id="2B029590FFD54071FF7076CB0E7DFC34" pageId="7" pageNumber="8">– Smaller, HL &lt;0.45, HW &lt;0.35. Flange over metapleural gland opening long, surpassing propodeum declivity margin in lateral view, and sharply pointed posteriorly ………………13</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B072E00FFD54071FF7071570E7CFB22" pageId="7" pageNumber="8">
<paragraph id="904C36A5FFD54071FF7071570E7CFB4C" blockId="7.[189,1399,1057,1260]" pageId="7" pageNumber="8">
<keyLead id="2B029590FFD54071FF7071570E7CFB4C" pageId="7" pageNumber="8">
12. HL = 0.50 on single known specimen. Masticatory margin of mandible distinctly dentate, with regularly spaced and well-developed teeth (
<bibRefCitation id="F4624B54FFD54071FD32713208FEFB90" author="Brandao C. R. F. &amp; Diniz J. L. M. &amp; Agosti D. &amp; Delabie J. H. C." box="[767,1013,1092,1118]" pageId="7" pageNumber="8" pagination="17 - 36" refId="ref14350" refString="Brandao C. R. F., Diniz J. L. M., Agosti D. &amp; Delabie J. H. C. 1999. Revision of the Neotropical ant subfamily Leptanilloidinae. Systematic Entomology 24: 17 - 36. http: // dx. doi. org / 10.1046 / j. 1365 - 3113.1999.00064. x" type="journal article" year="1999">
Brandão 
<emphasis id="A287EAB7FFD54071FCA3713308A7FB90" box="[878,940,1092,1118]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
1999
</bibRefCitation>
: fig. 16) (
<collectingCountry id="E8E47635FFD54071FBA171320FDAFB91" box="[1132,1233,1092,1119]" name="Ecuador" pageId="7" pageNumber="8">Ecuador</collectingCountry>
) …………… …………………………………………………………………… 
<taxonomicName id="57F34D26FFD54071FC20711E0E7CFB4C" ID-CoL="3THNJ" authority="Brandao et al., 1999" authorityName="Brandao" authorityYear="1999" box="[1005,1399,1127,1154]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="improvisa">
<emphasis id="A287EAB7FFD54071FC20711E0F88FB4F" bold="true" box="[1005,1155,1127,1154]" italics="true" pageId="7" pageNumber="8">L. improvisa</emphasis>
<bibRefCitation id="F4624B54FFD54071FB4771110E7CFB4C" author="Brandao C. R. F. &amp; Diniz J. L. M. &amp; Agosti D. &amp; Delabie J. H. C." box="[1162,1399,1127,1154]" pageId="7" pageNumber="8" pagination="17 - 36" refId="ref14350" refString="Brandao C. R. F., Diniz J. L. M., Agosti D. &amp; Delabie J. H. C. 1999. Revision of the Neotropical ant subfamily Leptanilloidinae. Systematic Entomology 24: 17 - 36. http: // dx. doi. org / 10.1046 / j. 1365 - 3113.1999.00064. x" type="journal article" year="1999">
Brandão 
<emphasis id="A287EAB7FFD54071FB39711E0E25FB4F" box="[1268,1326,1127,1153]" italics="true" pageId="7" pageNumber="8">et al.</emphasis>
, 1999
</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="904C36A5FFD54071FF7071FD0E7CFB22" blockId="7.[189,1399,1057,1260]" pageId="7" pageNumber="8">
<keyLead id="2B029590FFD54071FF7071FD0E7CFB22" pageId="7" pageNumber="8">
– HL ≥ 0.58. Masticatory margin of mandible dentate but teeth extremely minute, blunt and irregularly shaped, difficult to discern even under magnifications of about 100× (
<figureCitation id="08C82A20FFD54071FB1871D80E32FB06" box="[1237,1337,1198,1224]" captionStart="Fig" captionStartId="12.[189,232,1887,1913]" captionTargetBox="[189,1399,944,1851]" captionTargetId="figure@12.[189,1399,944,1851]" captionTargetPageId="12" captionText="Fig. 4. Leptanilloides prometea Delsinne &amp; Donoso sp. nov. – A–C. Paratype worker (specimen code 4052301). A. Habitus, dorsal view. B. Head in full-face view. C. Habitus, lateral view. – D. Apex of hindtibia; reservoir of metatibial gland is visible under translucent cuticle at the base of the tibial spur (paratype, specimen code 4052313)." figureDoi="http://doi.org/10.5281/zenodo.3787372" httpUri="https://zenodo.org/record/3787372/files/figure.png" pageId="7" pageNumber="8">Figs 4B</figureCitation>
, 
<figureCitation id="08C82A20FFD54071FA8771D80E65FB07" box="[1354,1390,1198,1225]" captionStart="Fig" captionStartId="14.[189,232,1825,1851]" captionTargetBox="[189,1399,503,1799]" captionTargetId="figure@14.[189,1399,503,1799]" captionTargetPageId="14" captionText="Fig. 6. Leptanilloides prometea Delsinne &amp; Donoso sp. nov. Paratype worker (specimen code 4052301). A. Head, ventral view. B. Mesotibial pectinate spur. C. Anterior part of the head in full-face view, showing vertical and fused frontal carinae, exposed antennal socket, well-developed lateroclypeal tooth, strongly convex clypeal lamella and subtriangular mandible with minute denticles along its inner margin. D. Lower part of propodeum in lateral view, showing the flange over the metapleural gland opening. E. Petiole in lateral view. F. Simple claw (hindleg)." figureDoi="http://doi.org/10.5281/zenodo.3787376" httpUri="https://zenodo.org/record/3787376/files/figure.png" pageId="7" pageNumber="8">6C</figureCitation>
) (
<collectingCountry id="E8E47635FFD54071FF3771A70A54FB22" box="[250,351,1233,1260]" name="Ecuador" pageId="7" pageNumber="8">Ecuador</collectingCountry>
) ……………………………………………… 
<taxonomicName id="57F34D26FFD54071FC4C71A40E04FB22" authority="Delsinne &amp; Donoso" authorityName="Delsinne &amp; Sonet &amp; Donoso" authorityYear="2015" box="[897,1295,1233,1260]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="prometea" status="sp. nov.">
<emphasis id="A287EAB7FFD54071FC4C71A40F1EFB22" bold="true" box="[897,1045,1234,1260]" italics="true" pageId="7" pageNumber="8">L. prometea</emphasis>
Delsinne &amp; Donoso
</taxonomicName>
<taxonomicNameLabel id="B9B457CCFFD54071FAD471A40E7CFB22" box="[1305,1399,1234,1260]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="2B072E00FFD54071FF7070650E7CF9CE" pageId="7" pageNumber="8">
<paragraph id="904C36A5FFD54071FF7070650E7CFABD" blockId="7.[189,1399,1298,1536]" pageId="7" pageNumber="8">
<keyLead id="2B029590FFD54071FF7070650E7CFABD" pageId="7" pageNumber="8">
13. Ventral margin of petiolar sternite forming an even convexity in lateral view. Reticulation on mesopleuron, metapleuron and lateral side of petiole uninterrupted. In dorsal view, petiole subquadrate, 67 ≤ PI ≤ 80 (
<collectingCountry id="E8E47635FFD54071FDF6702F09ABFABA" box="[571,672,1369,1396]" name="Ecuador" pageId="7" pageNumber="8">Ecuador</collectingCountry>
) ……………………… 
<taxonomicName id="57F34D26FFD54071FC74702F0E7CFABD" authority="Borowiec &amp; Longino, 2011" authorityName="Borowiec &amp; Longino" authorityYear="2011" box="[953,1399,1369,1395]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="erinys">
<emphasis id="A287EAB7FFD54071FC74702F0F2FFABD" bold="true" box="[953,1060,1369,1395]" italics="true" pageId="7" pageNumber="8">L. erinys</emphasis>
<bibRefCitation id="F4624B54FFD54071FBE3702F0E7CFABD" author="Borowiec M. L. &amp; Longino J. T." box="[1070,1399,1369,1395]" pageId="7" pageNumber="8" pagination="19 - 48" refId="ref14112" refString="Borowiec M. L. &amp; Longino J. T. 2011. Three new species and reassessment of the rare Neotropical ant genus Leptanilloides (Hymenoptera, Formicidae, Leptanilloidinae). ZooKeys 133: 19 - 48. http: // dx. doi. org / 10.3897 / zookeys. 133.1479" type="journal article" year="2011">Borowiec &amp; Longino, 2011</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="904C36A5FFD54071FF70700B0E7CF9CE" blockId="7.[189,1399,1298,1536]" pageId="7" pageNumber="8">
<keyLead id="2B029590FFD54071FF70700B0E7CF9CE" pageId="7" pageNumber="8">
– Ventral margin of petiolar sternite relatively straight in lateral view (
<figureCitation id="08C82A20FFD54071FB8E700A0FAFFA58" box="[1091,1188,1404,1430]" captionStart="Fig" captionStartId="10.[189,232,1790,1816]" captionTargetBox="[218,1370,264,1756]" captionTargetId="figure@10.[218,1370,264,1756]" captionTargetPageId="10" captionText="Fig. 3. Leptanilloides copalinga Delsinne &amp; Donoso sp. nov. Paratype worker (specimen code 4006301). A. Anterior part of the head in full-face view, showing vertical and fused frontal carinae, exposed antennal socket, lateroclypeal tooth, strongly convex clypeal lamella and subtriangular mandible with minute denticles along its inner margin. B. Lower part of propodeum in lateral view, showing the long and sharp flange over the metapleural gland opening. C. Petiole in lateral view. D. Propodeum and petiole in dorsal view. E. Abdominal segment III (postpetiole) in lateral view. F. Mesotibial spur, short and pectinate." figureDoi="http://doi.org/10.5281/zenodo.3787370" httpUri="https://zenodo.org/record/3787370/files/figure.png" pageId="7" pageNumber="8">Fig. 3C</figureCitation>
). Reticulation on mesopleuron, metapleuron and lateral side of petiole superficial and interrupted. In dorsal view, petiole slender, rectangular, 52 ≤ PI ≤ 63 (
<figureCitation id="08C82A20FFD54071FD2370B40840FA13" box="[750,843,1474,1501]" captionStart="Fig" captionStartId="10.[189,232,1790,1816]" captionTargetBox="[218,1370,264,1756]" captionTargetId="figure@10.[218,1370,264,1756]" captionTargetPageId="10" captionText="Fig. 3. Leptanilloides copalinga Delsinne &amp; Donoso sp. nov. Paratype worker (specimen code 4006301). A. Anterior part of the head in full-face view, showing vertical and fused frontal carinae, exposed antennal socket, lateroclypeal tooth, strongly convex clypeal lamella and subtriangular mandible with minute denticles along its inner margin. B. Lower part of propodeum in lateral view, showing the long and sharp flange over the metapleural gland opening. C. Petiole in lateral view. D. Propodeum and petiole in dorsal view. E. Abdominal segment III (postpetiole) in lateral view. F. Mesotibial spur, short and pectinate." figureDoi="http://doi.org/10.5281/zenodo.3787370" httpUri="https://zenodo.org/record/3787370/files/figure.png" pageId="7" pageNumber="8">Fig. 3D</figureCitation>
) (
<collectingCountry id="E8E47635FFD54071FCA870B408C1FA13" box="[869,970,1474,1501]" name="Ecuador" pageId="7" pageNumber="8">Ecuador</collectingCountry>
) …………………………………… ………………………………………………………… 
<taxonomicName id="57F34D26FFD54071FCBA70900E06F9CE" authority="Delsinne &amp; Donoso" authorityName="Delsinne &amp; Sonet &amp; Donoso" authorityYear="2015" box="[887,1293,1509,1536]" class="Insecta" family="Formicidae" genus="Leptanilloides" kingdom="Animalia" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="copalinga" status="sp. nov.">
<emphasis id="A287EAB7FFD54071FCBA70900F19F9CE" bold="true" box="[887,1042,1509,1536]" italics="true" pageId="7" pageNumber="8">L. copalinga</emphasis>
Delsinne &amp; Donoso
</taxonomicName>
<taxonomicNameLabel id="B9B457CCFFD54071FAD470900E7CF9CE" box="[1305,1399,1510,1536]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>